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<document id="56FDD74456F092AFC5CA8C7547015F58" ID-CLB-Dataset="7471" ID-DOI="10.11646/zootaxa.4948.3.3" ID-GBIF-Dataset="4321d317-546a-43e0-b149-3471384daca0" ID-ISSN="1175-5326" ID-Zenodo-Dep="4629078" ID-ZooBank="6A2B152A-D23A-4051-9D5A-957D9E509149" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1616485226681" checkinUser="plazi" docAuthor="Skelley, Paul E., Leschen, Richard A. B. &amp; Liu, Zhenhua" docDate="2021" docId="5F3187E24B0F4C4CFF63FB1B52FEE149" docLanguage="en" docName="zootaxa.4948.3.3.pdf" docOrigin="Zootaxa 4948 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Echinothallis Skelley, Leschen &amp; Liu 2021, new genus" docType="treatment" docVersion="6" lastPageNumber="365" masterDocId="A308FF9A4B0E4C4EFFF4FF9C506BE430" masterDocTitle="New Australian Erotylinae with notes on Dacnini (Coleoptera: Cucujoidea Erotylidae)" masterLastPageNumber="380" masterPageNumber="363" pageNumber="364" updateTime="1698934459833" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="E0A773B58FE19494D2764A0D5D75B342">New Australian Erotylinae with notes on Dacnini (Coleoptera: Cucujoidea Erotylidae)</mods:title>
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<mods:namePart id="BE6540DCC38E906FD99A7E749725FA02">Skelley, Paul E.</mods:namePart>
<mods:affiliation id="0B4FFCABC680099420AC79DDBF62846B">Florida State Collection of Arthropods, Florida Department of Agriculture and Consumer Services, 1911 SW 34 th Street, Gainesville, FL 32608, USA.</mods:affiliation>
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<mods:namePart id="91F217523CDD1F4ED229C876A9BA7CBA">Leschen, Richard A. B.</mods:namePart>
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<mods:affiliation id="AEB35FD8717A78EB956186A13652524A">New Zealand Arthropod Collection, Manaaki Whenua-Landcare Research, Private Bag 92170, Auckland, NEW ZEALAND leschenr @ landcareresearch. co. nz; https: // orcid. org / 0000 - 0001 - 8549 - 8933</mods:affiliation>
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<mods:namePart id="06E5DFD818C68027C58DA5F55D07F789">Liu, Zhenhua</mods:namePart>
<mods:affiliation id="5515E0A073B89B6A5F65155B990E9C7D">Australian National Insect Collection, CSIRO, GPO Box 1700, Canberra, ACT 2601, AUSTRALIA &amp; Key Laboratory of Biodiversity Dynamics and Conservation of Guangdong Higher Education Institute, The Museum of Biology, School of Life Science, Sun Yat-sen University, Guangzhou 510275, CHINA.</mods:affiliation>
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<treatment id="5F3187E24B0F4C4CFF63FB1B52FEE149" ID-DOI="http://doi.org/10.5281/zenodo.4629062" ID-GBIF-Taxon="180877821" ID-Zenodo-Dep="4629062" LSID="urn:lsid:plazi:treatment:5F3187E24B0F4C4CFF63FB1B52FEE149" httpUri="http://treatment.plazi.org/id/5F3187E24B0F4C4CFF63FB1B52FEE149" lastPageId="2" lastPageNumber="365" pageId="1" pageNumber="364">
<subSubSection id="9F82657F4B0F4C4FFF63FB1B5298E091" box="[151,755,1159,1186]" pageId="1" pageNumber="364" type="nomenclature">
<paragraph id="D72736F44B0F4C4FFF63FB1B5298E091" blockId="1.[151,755,1159,1186]" box="[151,755,1159,1186]" pageId="1" pageNumber="364">
<heading id="8C6F81984B0F4C4FFF63FB1B5298E091" bold="true" box="[151,755,1159,1186]" fontSize="11" level="1" pageId="1" pageNumber="364" reason="1">
<emphasis id="E5ECEAE64B0F4C4FFF63FB1B5298E091" bold="true" box="[151,755,1159,1186]" pageId="1" pageNumber="364">
<taxonomicName id="10984D774B0F4C4FFF63FB1B520FE091" authority="Skelley, Leschen &amp; Liu" authorityName="Skelley, Leschen &amp; Liu" authorityYear="2021" box="[151,612,1159,1186]" class="Insecta" family="Erotylidae" genus="Echinothallis" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="364" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="E5ECEAE64B0F4C4FFF63FB1B5151E092" bold="true" box="[151,314,1159,1186]" italics="true" pageId="1" pageNumber="364">Echinothallis</emphasis>
Skelley, Leschen &amp; Liu
</taxonomicName>
,
<taxonomicNameLabel id="FEDF579D4B0F4C4FFD85FB1B5298E091" box="[625,755,1159,1185]" pageId="1" pageNumber="364" rank="genus">new genus</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
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<subSubSection id="9F82657F4B0F4C4FFF63FB5054D2E0D4" box="[151,1209,1227,1252]" pageId="1" pageNumber="364" type="type_taxon">
<paragraph id="D72736F44B0F4C4FFF63FB5054D2E0D4" blockId="1.[151,1209,1227,1252]" box="[151,1209,1227,1252]" pageId="1" pageNumber="364">
<emphasis id="E5ECEAE64B0F4C4FFF63FB505148E0D3" bold="true" box="[151,291,1227,1252]" pageId="1" pageNumber="364">
<typeStatus id="082388564B0F4C4FFF63FB5050A7E0D4" box="[151,204,1228,1252]" pageId="1" pageNumber="364">Type</typeStatus>
species.
</emphasis>
<taxonomicName id="10984D774B0F4C4FFEDEFB505356E0D4" authority="Skelley, Leschen, Liu" authorityName="Skelley, Leschen, Liu" authorityYear="2021" box="[298,829,1228,1252]" class="Insecta" family="Erotylidae" genus="Echinothallis" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="364" phylum="Arthropoda" rank="species" species="banderbearella" status="sp. nov.">
<emphasis id="E5ECEAE64B0F4C4FFEDEFB505239E0D4" box="[298,594,1228,1252]" italics="true" pageId="1" pageNumber="364">Echinothallis banderbearella</emphasis>
, Skelley, Leschen, Liu
</taxonomicName>
,
<taxonomicNameLabel id="FEDF579D4B0F4C4FFCB3FB5053AFE0D4" box="[839,964,1228,1252]" pageId="1" pageNumber="364" rank="species">new species</taxonomicNameLabel>
, by present designation.
</paragraph>
</subSubSection>
<subSubSection id="9F82657F4B0F4C4FFF63FA8D556CE1A8" pageId="1" pageNumber="364" type="diagnosis">
<paragraph id="D72736F44B0F4C4FFF63FA8D556CE1A8" blockId="1.[151,1437,1297,2008]" pageId="1" pageNumber="364">
<emphasis id="E5ECEAE64B0F4C4FFF63FA8D5161E11B" bold="true" box="[151,266,1297,1323]" pageId="1" pageNumber="364">Diagnosis</emphasis>
. A member of the
<taxonomicName id="10984D774B0F4C4FFE21FA8E5245E11C" box="[469,558,1298,1324]" class="Insecta" family="Erotylidae" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="364" phylum="Arthropoda" rank="tribe" tribe="Dacnini">Dacnini</taxonomicName>
with prominent anterior pronotal margin and narrow tetramerous tarsi.
<taxonomicName id="10984D774B0F4C4FFAB0FA8E50B6E160" authorityName="Skelley, Leschen &amp; Liu" authorityYear="2021" class="Insecta" family="Erotylidae" genus="Echinothallis" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="364" phylum="Arthropoda" rank="genus">
<emphasis id="E5ECEAE64B0F4C4FFAB0FA8E50B6E160" italics="true" pageId="1" pageNumber="364">Echinothallis</emphasis>
</taxonomicName>
is distinguished from other genera by the coarsely facetted eyes, elongate, somewhat parallel-sided body, posterior pronotum and elytral base with marginal bead, mesocoxae narrowly separated, mental plate triangular, terminal palpomere of maxilla flattened and wider apically, and the heavily punctate and setose body.
</paragraph>
</subSubSection>
<subSubSection id="9F82657F4B0F4C4CFF33FA3D5124E021" lastPageId="2" lastPageNumber="365" pageId="1" pageNumber="364" type="description">
<paragraph id="D72736F44B0F4C4FFF33FA3D50BAE1D0" blockId="1.[151,1437,1297,2008]" pageId="1" pageNumber="364">
<emphasis id="E5ECEAE64B0F4C4FFF33FA3D5132E18B" bold="true" box="[199,345,1441,1467]" pageId="1" pageNumber="364">Description.</emphasis>
Length
<quantity id="10609B114B0F4C4FFE4AFA3E5220E18C" box="[446,587,1442,1468]" metricMagnitude="-3" metricUnit="m" metricValue="3.75" metricValueMax="3.9" metricValueMin="3.6" pageId="1" pageNumber="364" unit="mm" value="3.75" valueMax="3.9" valueMin="3.6">3.63.9 mm</quantity>
; width
<quantity id="10609B114B0F4C4FFD51FA3E5358E18C" box="[677,819,1442,1468]" metricMagnitude="-3" metricUnit="m" metricValue="1.35" metricValueMax="1.4" metricValueMin="1.3" pageId="1" pageNumber="364" unit="mm" value="1.35" valueMax="1.4" valueMin="1.3">1.31.4 mm</quantity>
. Body elongate, somewhat parallel-sided body (
<figureCitation id="4FA32A714B0F4C4FFA90FA3E50AFE1D0" captionStart="FIGURES 18" captionStartId="3.[151,264,1900,1924]" captionTargetBox="[191,1392,442,1836]" captionTargetId="figure-103@3.[151,1436,417,1876]" captionTargetPageId="3" captionText="FIGURES 18. Echinothallis banderbearella sp. nov.: 13) habitus, holotype; 1) dorsal view; 2) ventral view; 3) lateral view; 4) female genitalia, allotype; 58) male genitalia, paratype; 5) abdominal segments VIIIIX, ventral view; 6) tegmen, dorsal view; 7) penis, lateral view; 8) penis, dorsal view. Scale bars: 13 = 1.0 mm, 48 = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.4629080" httpUri="https://zenodo.org/record/4629080/files/figure.png" pageId="1" pageNumber="364">Figs. 13</figureCitation>
).
</paragraph>
<paragraph id="D72736F44B0F4C4FFF33FA7651FBE2EC" blockId="1.[151,1437,1297,2008]" pageId="1" pageNumber="364">Head with sides narrowing anteriorly; supraocular stria as long as eye, ending near base of antenna. Eyes coarsely faceted; almost oval, prominent; with setae between facets. Stridulatory files on occipital region absent. Mentum with plate broadly triangular, width 2× length, coarsely punctate; median projection small. Submentum short, transverse; surface and groove behind with large, deep punctures. Maxilla with terminal palpomere flattened, length = apical width, narrowed basally, apically prolonged medially, sensory apex truncate. Labium with terminal palpomere narrowed toward base, apically flattened, width 0.5× width of terminal maxillary palpomere; apical sensory area elongate.</paragraph>
<paragraph id="D72736F44B0F4C4FFF33F97A53CCE35C" blockId="1.[151,1437,1297,2008]" pageId="1" pageNumber="364">Pronotum convex, anterior margin lacking marginal bead; anterior angles blunt, not projecting forward, surface explanate at angle; lateral margin gently arched, convergent anteriorly and posteriorly, with fine marginal bead, and fringe of irregularly long setae; posterior angles sharp, projecting posteriorly; posterior margin of pronotum weakly lobed at middle, marginal bead present, obscured in surface sculpture.</paragraph>
<paragraph id="D72736F44B0F4C4FFF33F8EA53AEE3E8" blockId="1.[151,1437,1297,2008]" pageId="1" pageNumber="364">Scutellar shield broadly, roundly pentagonal. Elytra elongate, length 1.8× pronotal length; base with marginal bead present, obscured in surface sculpture; lateral margin evenly arcuate from base to apex, widest at midlength; humeral denticles present, small. Wings apparently lacking, elytra fused.</paragraph>
<paragraph id="D72736F44B0C4C4CFF33FF0B5224E52D" blockId="2.[151,1437,151,1401]" pageId="2" pageNumber="365">Legs slender and densely hirsute. Femora fusiform, somewhat flattened, with two sharp edges on inner surface. Tibiae slender, slightly curved basally; apex with fringe of moderately long spinules. Tarsi elongate, cylindrical, tarsomeres 13 densely setose ventrally but not lobed, tarsomeres 14 more or less of same size; tarsomere 5 slightly shorter than preceding four tarsomeres.</paragraph>
<paragraph id="D72736F44B0C4C4CFF33FEBB5547E5E1" blockId="2.[151,1437,151,1401]" pageId="2" pageNumber="365">Prosternum anterior margin with strong marginal bead; anterior margin of procoxa with line; procoxal lines present between coxa, not extending anteriorly, weakly extending posteriorly onto prosternal process where obscured in dense punctation; prosternal process parallel-sided between coxae, widening posterior of coxae, width between procoxa about = transverse diameter of coxa; posterior margin truncate; pores on prosternal process not observed. Hypomeron with deep coarse punctures bearing long setae, punctures separated by 1 diameter.</paragraph>
<paragraph id="D72736F44B0C4C4CFF33FE4754AEE629" blockId="2.[151,1437,151,1401]" pageId="2" pageNumber="365">Mesoventrite flattened covered with deep seta bearing punctures as large as facets, punctures coalescing; mesometaventrite suture narrow, distance between mesocoxae narrower than the width of mesocoxa.</paragraph>
<paragraph id="D72736F44B0C4C4CFF33FDBF54FEE699" blockId="2.[151,1437,151,1401]" pageId="2" pageNumber="365">Metaventrite moderately short, length between meso- and metacoxa 1.5× distance between mesocoxae; anterior margin distinctly bordered medially; coxal lines extending onto surface 2/3 distance to posterior margin; anterior marginal line obscured in coarse punctures behind mesocoxae; metathoracic discrimen and coxal lines obscured in coarse punctation. Metanepisternum narrow, covered with scattered seta bearing punctures.</paragraph>
<paragraph id="D72736F44B0C4C4CFF33FD2F5484E725" blockId="2.[151,1437,151,1401]" pageId="2" pageNumber="365">Abdomen with ventrite 1 anterior process bluntly rounded; two coxal lines present, first extending from medial margin of metacoxal cavity nearly to posterior margin of ventrite 1, second short line extending posteriorly from lateral 1/3 of mesocoxal cavity; anterior margin of ventrite 1 behind metacoxae distinctly bordered.</paragraph>
<paragraph id="D72736F44B0C4C4CFF33FC83553DE795" blockId="2.[151,1437,151,1401]" pageId="2" pageNumber="365">
Male genitalia (
<figureCitation id="4FA32A714B0C4C4CFE81FC8351B5E709" box="[373,478,798,825]" captionStart="FIGURES 18" captionStartId="3.[151,264,1900,1924]" captionTargetBox="[191,1392,442,1836]" captionTargetId="figure-103@3.[151,1436,417,1876]" captionTargetPageId="3" captionText="FIGURES 18. Echinothallis banderbearella sp. nov.: 13) habitus, holotype; 1) dorsal view; 2) ventral view; 3) lateral view; 4) female genitalia, allotype; 58) male genitalia, paratype; 5) abdominal segments VIIIIX, ventral view; 6) tegmen, dorsal view; 7) penis, lateral view; 8) penis, dorsal view. Scale bars: 13 = 1.0 mm, 48 = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.4629080" httpUri="https://zenodo.org/record/4629080/files/figure.png" pageId="2" pageNumber="365">Figs. 58</figureCitation>
) with penis short, arched; median strut and flagellum long; penis length = 0.3× median strut length; internal sac with weak microsetae in caudal half; flagellum with small elongate oval head, virga long, as long as median strut, hair-like; parameres bluntly rounded apically, with long setae; abdominal sternite IX broad, truncate, sides of spiculum gastrale separated anteriorly by oval sclerite together they form a pregenital ring.
</paragraph>
<paragraph id="D72736F44B0C4C4CFF33FC335124E021" blockId="2.[151,1437,151,1401]" pageId="2" pageNumber="365">
Female genitalia (
<figureCitation id="4FA32A714B0C4C4CFE6CFC3351B4E7F9" box="[408,479,943,969]" captionStart="FIGURES 18" captionStartId="3.[151,264,1900,1924]" captionTargetBox="[191,1392,442,1836]" captionTargetId="figure-103@3.[151,1436,417,1876]" captionTargetPageId="3" captionText="FIGURES 18. Echinothallis banderbearella sp. nov.: 13) habitus, holotype; 1) dorsal view; 2) ventral view; 3) lateral view; 4) female genitalia, allotype; 58) male genitalia, paratype; 5) abdominal segments VIIIIX, ventral view; 6) tegmen, dorsal view; 7) penis, lateral view; 8) penis, dorsal view. Scale bars: 13 = 1.0 mm, 48 = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.4629080" httpUri="https://zenodo.org/record/4629080/files/figure.png" pageId="2" pageNumber="365">Fig. 4</figureCitation>
) with normally sclerotized gonocoxite and gonostylus, gonostylus distinct and set apically on gonocoxite; segment IX lacking denticulate lamellae; head of spermatheca elongate oval, with short sclerotized duct.
</paragraph>
</subSubSection>
<subSubSection id="9F82657F4B0C4C4CFF33FB8753A9E069" pageId="2" pageNumber="365" type="etymology">
<paragraph id="D72736F44B0C4C4CFF33FB8753A9E069" blockId="2.[151,1437,151,1401]" pageId="2" pageNumber="365">
<emphasis id="E5ECEAE64B0C4C4CFF33FB875123E005" bold="true" box="[199,328,1051,1077]" pageId="2" pageNumber="365">Etymology</emphasis>
. The genus name is derived from two roots: “
<emphasis id="E5ECEAE64B0C4C4CFC96FB8753C6E005" box="[866,941,1051,1077]" italics="true" pageId="2" pageNumber="365">echino</emphasis>
” Greek for spiny and “
<taxonomicName id="10984D774B0C4C4CFB34FB875565E005" authorityName="Erichson" authorityYear="1842" box="[1216,1294,1051,1077]" class="Insecta" family="Erotylidae" genus="Thallis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="365" phylum="Arthropoda" rank="genus">
<emphasis id="E5ECEAE64B0C4C4CFB34FB875565E005" box="[1216,1294,1051,1077]" italics="true" pageId="2" pageNumber="365">Thallis</emphasis>
</taxonomicName>
” a genus of
<taxonomicName id="10984D774B0C4C4CFF63FBA3509BE069" box="[151,240,1087,1113]" class="Insecta" family="Erotylidae" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="365" phylum="Arthropoda" rank="tribe" tribe="Dacnini">Dacnini</taxonomicName>
in which it was considered to initially belong. Gender feminine.
</paragraph>
</subSubSection>
<subSubSection id="9F82657F4B0C4C4CFF33FBFF52FEE149" pageId="2" pageNumber="365" type="discussion">
<paragraph id="D72736F44B0C4C4CFF33FBFF52FEE149" blockId="2.[151,1437,151,1401]" pageId="2" pageNumber="365">
<emphasis id="E5ECEAE64B0C4C4CFF33FBFF515FE04D" bold="true" box="[199,308,1123,1149]" pageId="2" pageNumber="365">Remarks</emphasis>
. At first look this new genus and species appeared to be a new species of
<taxonomicName id="10984D774B0C4C4CFB99FBFF554EE04D" authority="Erichson" authorityName="Erichson" authorityYear="1842" box="[1133,1317,1123,1149]" class="Insecta" family="Erotylidae" genus="Thallis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="365" phylum="Arthropoda" rank="genus">
<emphasis id="E5ECEAE64B0C4C4CFB99FBFF54D0E04D" box="[1133,1211,1123,1149]" italics="true" pageId="2" pageNumber="365">Thallis</emphasis>
Erichson
</taxonomicName>
which has a range of species from glabrous to hairy, some already described from
<collectingCountry id="AF8F76644B0C4C4CFC40FB1B5470E091" box="[948,1051,1159,1185]" name="Australia" pageId="2" pageNumber="365">Australia</collectingCountry>
(other species occur in
<collectingCountry id="AF8F76644B0C4C4CFAD0FB1B50BEE0F5" name="New Caledonia" pageId="2" pageNumber="365">New Caledonia</collectingCountry>
and the
<collectingCountry id="AF8F76644B0C4C4CFECEFB3751C7E0F6" box="[314,428,1195,1222]" name="Indonesia" pageId="2" pageNumber="365">Moluccas</collectingCountry>
). However, closer examination show that
<taxonomicName id="10984D774B0C4C4CFC7EFB375474E0F5" authorityName="Skelley, Leschen &amp; Liu" authorityYear="2021" box="[906,1055,1195,1221]" class="Insecta" family="Erotylidae" genus="Echinothallis" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="365" phylum="Arthropoda" rank="genus">
<emphasis id="E5ECEAE64B0C4C4CFC7EFB375474E0F5" box="[906,1055,1195,1221]" italics="true" pageId="2" pageNumber="365">Echinothallis</emphasis>
</taxonomicName>
is unlike typical species of
<taxonomicName id="10984D774B0C4C4CFA94FB3750DAE0D9" authorityName="Erichson" authorityYear="1842" class="Insecta" family="Erotylidae" genus="Thallis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="365" phylum="Arthropoda" rank="genus">
<emphasis id="E5ECEAE64B0C4C4CFA94FB3750DAE0D9" italics="true" pageId="2" pageNumber="365">Thallis</emphasis>
</taxonomicName>
, including the hairy species, in many important characters that are lacking in some other dacnines (see below). Further studies of currently described species of
<taxonomicName id="10984D774B0C4C4CFD4EFB6F5363E13D" authorityName="Erichson" authorityYear="1842" box="[698,776,1267,1293]" class="Insecta" family="Erotylidae" genus="Thallis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="365" phylum="Arthropoda" rank="genus">
<emphasis id="E5ECEAE64B0C4C4CFD4EFB6F5363E13D" box="[698,776,1267,1293]" italics="true" pageId="2" pageNumber="365">Thallis</emphasis>
</taxonomicName>
shows it to be an accumulation of species groups that may be valid genera themselves (see
<bibRefCitation id="B3094B054B0C4C4CFDF1FA8B534AE101" author="Skelley, P. E. &amp; Leschen, R. A. B." box="[517,801,1303,1329]" pageId="2" pageNumber="365" pagination="1 - 58" refId="ref11659" refString="Skelley, P. E. &amp; Leschen, R. A. B. (2007) Erotylinae (Insecta: Coleoptera, Cucujoidea: Erotylidae): taxonomy and biogeography. Fauna of New Zealand, 59, 1 - 58." type="journal article" year="2007">Skelley &amp; Leschen 2007</bibRefCitation>
). A revision of
<taxonomicName id="10984D774B0C4C4CFC3BFA8B5476E101" authorityName="Erichson" authorityYear="1842" box="[975,1053,1303,1329]" class="Insecta" family="Erotylidae" genus="Thallis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="365" phylum="Arthropoda" rank="genus">
<emphasis id="E5ECEAE64B0C4C4CFC3BFA8B5476E101" box="[975,1053,1303,1329]" italics="true" pageId="2" pageNumber="365">Thallis</emphasis>
</taxonomicName>
is needed to resolve that problem which is outside the scope of the present work. Since
<taxonomicName id="10984D774B0C4C4CFD1AFAA753E8E165" authorityName="Skelley, Leschen &amp; Liu" authorityYear="2021" box="[750,899,1339,1365]" class="Insecta" family="Erotylidae" genus="Echinothallis" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="365" phylum="Arthropoda" rank="genus">
<emphasis id="E5ECEAE64B0C4C4CFD1AFAA753E8E165" box="[750,899,1339,1365]" italics="true" pageId="2" pageNumber="365">Echinothallis</emphasis>
</taxonomicName>
falls outside of the current concept of
<taxonomicName id="10984D774B0C4C4CFAC5FAA75514E165" authorityName="Erichson" authorityYear="1842" box="[1329,1407,1339,1365]" class="Insecta" family="Erotylidae" genus="Thallis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="365" phylum="Arthropoda" rank="genus">
<emphasis id="E5ECEAE64B0C4C4CFAC5FAA75514E165" box="[1329,1407,1339,1365]" italics="true" pageId="2" pageNumber="365">Thallis</emphasis>
</taxonomicName>
, it is here described as a prelude to this revision.
</paragraph>
</subSubSection>
</treatment>
</document>