treatments-xml/data/43/A1/7C/43A17C80F41D849F5E625F6BAA9E67E9.xml

<document ID-DOI="http://dx.doi.org/10.3897/phytokeys.106.21991" ID-PMC="PMC6070582" ID-Pensoft-Pub="1314-2003-106-1" ID-Pensoft-UUID="FF8BFFC82928FFA84734FFCB2E61E260" ID-PubMed="30072843" ID-Zenodo-Dep="1326005" ModsDocID="1314-2003-106-1" checkinTime="1532592511837" checkinUser="pensoft" docAuthor="Saerkinen, Tiina, Poczai, Peter, Barboza, Gloria E., Weerden, Gerard M. van der, Baden, Maria &amp; Knapp, Sandra" docDate="2018" docId="43A17C80F41D849F5E625F6BAA9E67E9" docLanguage="en" docName="PhytoKeys 106: 1-223" docOrigin="PhytoKeys 106" docPubDate="2018-07-25" docSource="http://dx.doi.org/10.3897/phytokeys.106.21991" docTitle="Solanum umalilaense Manoko, PhytoKeys 16: 67. 2012" docType="treatment" docVersion="7" id="FF8BFFC82928FFA84734FFCB2E61E260" lastPageNumber="1" masterDocId="FF8BFFC82928FFA84734FFCB2E61E260" masterDocTitle="A revision of the Old World Black Nightshades (Morelloid clade of Solanum L., Solanaceae)" masterLastPageNumber="223" masterPageNumber="1" pageNumber="1" updateTime="1668138996586" updateUser="ExternalLinkService">
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<mods:titleInfo>
<mods:title>A revision of the Old World Black Nightshades (Morelloid clade of Solanum L., Solanaceae)</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Saerkinen, Tiina</mods:namePart>
<mods:affiliation>Royal Botanic Garden Edinburgh, 20 A Inverleith Row, Edinburgh EH 3 5 LR, United Kingdom</mods:affiliation>
<mods:nameIdentifier type="email">tsarkinen@rbge.org.uk</mods:nameIdentifier>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Poczai, Peter</mods:namePart>
<mods:affiliation>Botany Unit, Finnish Museum of Natural History, University of Helsinki, P. O. Box 7, FI- 00014 Helsinki, Finland</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Barboza, Gloria E.</mods:namePart>
<mods:affiliation>Instituto Multidisciplinario de Biologia Vegetal (CONICET-Universidad Nacional de Cordoba), Casilla de Correo 495, 5000 Cordoba, Argentina</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Weerden, Gerard M. van der</mods:namePart>
<mods:affiliation>Experimental Garden, Radboud University, Faculty of Science Box 49, P. O. Box 9010, 6500 Nijmegen, The Netherlands</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Baden, Maria</mods:namePart>
<mods:affiliation>Max-Planck Odense Center on the Biodemography of Aging and Department of Biology, University of Southern Denmark, Campusvej 55, DK- 5230 Odense M, Denmark</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Knapp, Sandra</mods:namePart>
<mods:affiliation>Department of Life Sciences, Natural History Museum, Cromwell Road, London SW 7 5 BD, United Kingdom</mods:affiliation>
<mods:nameIdentifier type="email">s.knapp@nhm.ac.uk</mods:nameIdentifier>
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<mods:title>PhytoKeys</mods:title>
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<mods:date>2018</mods:date>
<mods:detail type="pubDate">
<mods:number>2018-07-25</mods:number>
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<mods:detail type="volume">
<mods:number>106</mods:number>
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<mods:start>1</mods:start>
<mods:end>223</mods:end>
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<mods:url>http://dx.doi.org/10.3897/phytokeys.106.21991</mods:url>
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<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.3897/phytokeys.106.21991</mods:identifier>
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<mods:identifier type="Zenodo-Dep">1326005</mods:identifier>
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<treatment ID-GBIF-Taxon="146692537" LSID="urn:lsid:plazi:treatment:43A17C80F41D849F5E625F6BAA9E67E9" httpUri="http://treatment.plazi.org/id/43A17C80F41D849F5E625F6BAA9E67E9" lastPageNumber="1" pageId="0" pageNumber="1">
<subSubSection pageId="0" pageNumber="1" type="nomenclature">
<paragraph pageId="0" pageNumber="1">
18. 
<taxonomicName LSID="43A17C80-F41D-849F-5E62-5F6BAA9E67E9" authority="Manoko, PhytoKeys 16: 67. 2012" authorityName="Manoko, PhytoKeys 16: 67." authorityYear="2012" class="Magnoliopsida" family="Solanaceae" genus="Solanum" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Solanum umalilaense" order="Solanales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="species" species="umalilaense">Solanum umalilaense Manoko, PhytoKeys 16: 67. 2012</taxonomicName>
<figureCitation captionStart="Figure 55" captionStartId="F55" captionText="Figure 55. Solanum umalilaense A Habit B Flower and buds showing calyx shape C Inflorescence and leaves D Infructescence showing detail of winged stem. (A-D NIJ-A 24750133). Photos by G. van der Weerden." figureDoi="10.3897/phytokeys.106.21991.figure55" httpUri="https://binary.pensoft.net/fig/218760" pageId="0" pageNumber="1">Figure 55</figureCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="materials_examined">
<paragraph pageId="0" pageNumber="1">
<typeStatus>Type</typeStatus>
.
</paragraph>
<paragraph pageId="0" pageNumber="1">
<materialsCitation collectingDate="2018-01-01" collectingDateMax="2018-12-31" collectingDateMin="2018-01-01" country="Tanzania" location="Umalila Forest Reserve" specimenCount="1" stateProvince="Mbeya" typeStatus="holotype">
<collectingCountry name="Tanzania">Tanzania</collectingCountry>
. 
<collectingRegion country="Tanzania" name="Mbeya">Mbeya</collectingRegion>
: 
<collectingRegion country="Tanzania" name="Mbeya">Mbeya District</collectingRegion>
, 
<location LSID="urn:lsid:plazi:treatment:43A17C80F41D849F5E625F6BAA9E67E9:31E90BC4D1BFA22EE24F17B2DEF55FFB" country="Tanzania" name="Umalila Forest Reserve" stateProvince="Mbeya">Umalila Forest Reserve</location>
, ca. 
<quantity metricMagnitude="3" metricUnit="m" metricValue="7.0" unit="km" value="7.0">7 km</quantity>
W of
</materialsCitation>
<materialsCitation collectingDate="1992-11-14" collectorName="R. E. Gereau, D. K. Harder, C. J. Kayombo, M. J. Kayombo" country="Ruanda" elevation="2180" latitude="-9.183333" location="Izumbwe" longLatPrecision="1300" longitude="33.3" specimenCount="1" typeStatus="holotype">
<collectingCountry name="Rwanda">Ruanda</collectingCountry>
II on road to 
<location LSID="urn:lsid:plazi:treatment:43A17C80F41D849F5E625F6BAA9E67E9:3B46BF357E48ED05EBC37D3B3B756B6B" country="Ruanda" latitude="-9.183333" longLatPrecision="1300" longitude="33.3" name="Izumbwe">Izumbwe</location>
, 
<locationDeviation location="Mbogo Mtn.">
<quantity metricMagnitude="3" metricUnit="m" metricValue="2.0" unit="km" value="2.0">2 km</quantity>
SSE of Mbogo Mtn.
</locationDeviation>
main peak, 
<geoCoordinate degrees="9" direction="south" minutes="11" orientation="latitude" precision="925" value="-9.183333">9°11'S</geoCoordinate>
, 
<geoCoordinate degrees="33" direction="east" minutes="18" orientation="longitude" precision="925" value="33.3">33°18'E</geoCoordinate>
, 
<quantity metricMagnitude="3" metricUnit="m" metricValue="2.18" unit="m" value="2180.0">
<elevation metricMagnitude="3" metricUnit="m" metricValue="2.18" unit="m" value="2180.0">2180 m</elevation>
</quantity>
, 
<collectingDate value="1992-11-14">14 Nov 1992</collectingDate>
. 
<emphasis italics="true" pageId="0" pageNumber="1">
<collectorName>R.E. Gereau</collectorName>
, 
<collectorName>D.K. Harder</collectorName>
, 
<collectorName>C.J. Kayombo</collectorName>
&amp; 
<collectorName>M.J. Kayombo</collectorName>
5084
</emphasis>
(
<typeStatus>holotype</typeStatus>
: DSM; isotypes: EA, K [K001081869], MO [MO-2293667, acc. # 4281580], NHT)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="description">
<paragraph pageId="0" pageNumber="1">Description.</paragraph>
<paragraph pageId="0" pageNumber="1">
Annual to short-lived erect perennial herbs to 0.5 m tall, erect, predominantly branching from the base. Stems spreading, ridged or winged, dark purple to green, erect, not markedly hollow; new growth puberulent with simple, spreading, uniseriate, translucent, eglandular trichomes, these 2-8-celled, 0.3-0.8 mm long; older stems glabrescent. Sympodial units difoliate, the leaves not geminate. Leaves simple, 1.8-2.8 cm long, 0.1-1.7 (-2.1) cm wide, ovate to elliptic, membranous, green to dark green above and below, without smell; adaxial and abaxial surfaces glabrous or sparsely pubescent with simple uniseriate trichomes like those on the stem mainly along veins; major veins 3-4 pairs, paler green or often purple tinged; base truncate, narrowly winged on to the petiole; margins entire; apex acuminate to acute; petioles 0.6-1.3 cm long, glabrous or sparsely pubescent with simple uniseriate trichomes like those of the stem. Inflorescences 2-6 cm long, internodal, simple or usually branched and often leafy, the flowers spaced along the rhachis, with 2-9(-20) flowers, glabrous or sparsely pubescent with simple uniseriate trichomes like those on the stem; peduncle (0.5-)1.1-4 cm long, erect, ca. 0.5 mm thick at apex, stout; pedicels 5-7(-9) mm long, ca. 0.2 mm in diameter at the base, ca. 0.5 mm in diameter at the apex, stout, pendent, articulated at the base; pedicel scars widely spaced (1-)3-8 mm apart. Buds subglobose, the corolla exserted 1/2 from the calyx tube before anthesis. Flowers 5-merous, perfect. Calyx tube 0.8-1.0 mm long, cup-shaped, the lobes equal, 0.1-0.5 mm long, 0.5-0.7 mm wide, broadly deltate with an acute tip, green or purple-tinged, glabrous or sparsely pubescent with simple uniseriate trichomes like those of the pedicels. Corolla 7-11 mm in diameter, white, margins occasionally tinged with purple, with a yellow basal star, stellate, lobed ca. 1/2 way to the base, the lobes 2.5-3.5 mm long, 2-3 mm wide, spreading or reflexed, densely papillate abaxially. Stamens equal; filament tube to 0.1 mm long; free portion of the filaments 0.5-1.0 mm long, pubescent adaxially with tangled uniseriate simple trichomes; anthers 1.8-2 mm long, 0.5-0.6 mm wide, ellipsoid, yellow, poricidal at the tips, the pores lengthening to slits with age and drying. Ovary rounded, glabrous; style 3.1-3.5(-4.2) mm long, densely pubescent with simple uniseriate trichomes 0.2-0.5 mm long in the basal 1/2-2/3, exserted 1.0-1.5 mm beyond anther cone; stigma clavate, the surface minutely papillate. Fruit a globose berry, 3-4(-5) mm in diameter, pale yellow-brown at maturity, aromatic, the pericarp soft, matte, not transparent; fruiting pedicels 0.8-1.3 cm long, 0.3-0.5 mm in diameter at the base, 0.5-1.0 mm in diameter at the apex, stout and becoming woody, erect or spreading, pale brown, falling with the mature fruit, not persistent; fruiting calyx not accrescent, the tube ca. 1 mm long, the lobes ca. 1 mm long, appressed to the berry. Seeds (5-)6-12 per berry, 1.6-2.1 mm long, 1.3-1.8 mm wide, flattened and tear-drop shaped with a subapical hilum, brownish, the surfaces minutely pitted, thin, the testal cells rectangular to pentagonal in outline. Stone cells 9-22 per berry, ca. 0.5 mm in diameter. Chromosome number: 
<emphasis italics="true" pageId="0" pageNumber="1">2n</emphasis>
=4x=48 (
<bibRefCitation DOI="https://doi.org/10.3897/phytokeys.16.2884" author="Manoko, MLK" journalOrPublisher="PhytoKeys" pageId="0" pageNumber="1" pagination="65 - 74" refId="B232" refString="Manoko, MLK, van der Weerden, GM, van den Berg, RG, Mariani, C, 2012. A new tetraploid species of Solanum L. sect. Solanum (Solanaceae) from Tanzania. PhytoKeys 16 (0): 65 - 74, DOI: https://doi.org/10.3897/phytokeys.16.2884" title="A new tetraploid species of Solanum L. sect. Solanum (Solanaceae) from Tanzania." url="https://doi.org/10.3897/phytokeys.16.2884" volume="16" year="2012">Manoko et al. 2012</bibRefCitation>
).
</paragraph>
<caption ID-Zenodo-Dep="1326131" doi="10.3897/phytokeys.106.21991.figure55" httpUri="https://binary.pensoft.net/fig/218760" pageId="0" pageNumber="1" start="Figure 55" startId="F55">
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" pageId="0" pageNumber="1">Figure 55.</emphasis>
<taxonomicName class="Magnoliopsida" family="Solanaceae" genus="Solanum" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Solanum umalilaense" order="Centrales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="species" species="umalilaense">
<emphasis italics="true" pageId="0" pageNumber="1">Solanum umalilaense</emphasis>
</taxonomicName>
<emphasis bold="true" pageId="0" pageNumber="1">A</emphasis>
Habit 
<emphasis bold="true" pageId="0" pageNumber="1">B</emphasis>
Flower and buds showing calyx shape 
<emphasis bold="true" pageId="0" pageNumber="1">C</emphasis>
Inflorescence and leaves 
<emphasis bold="true" pageId="0" pageNumber="1">D</emphasis>
Infructescence showing detail of winged stem. (
<emphasis bold="true" pageId="0" pageNumber="1">A-D</emphasis>
NIJ-A24750133). Photos by G. van der Weerden.
</paragraph>
</caption>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="distribution">
<paragraph pageId="0" pageNumber="1">Distribution</paragraph>
<paragraph pageId="0" pageNumber="1">
(Figure 
<figureCitation captionStart="Figure 56" captionStartId="F56" captionText="Figure 56. Distribution of Solanum umalilaense." figureDoi="10.3897/phytokeys.106.21991.figure56" httpUri="https://binary.pensoft.net/fig/218761" pageId="0" pageNumber="1">56</figureCitation>
). Endemic to the southern highlands of Tanzania.
</paragraph>
<caption ID-Zenodo-Dep="1326133" doi="10.3897/phytokeys.106.21991.figure56" httpUri="https://binary.pensoft.net/fig/218761" pageId="0" pageNumber="1" start="Figure 56" startId="F56">
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" pageId="0" pageNumber="1">Figure 56.</emphasis>
Distribution of 
<taxonomicName class="Magnoliopsida" family="Solanaceae" genus="Solanum" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Solanum umalilaense" order="Centrales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="species" species="umalilaense">
<emphasis italics="true" pageId="0" pageNumber="1">Solanum umalilaense</emphasis>
</taxonomicName>
.
</paragraph>
</caption>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="ecology">
<paragraph pageId="0" pageNumber="1">Ecology.</paragraph>
<paragraph pageId="0" pageNumber="1">Grows on volcanic soils, frequent on the ash layer in charcoal-burning areas, also commonly cultivated; between 1,700 and 2,200 m elevation.</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="common names">
<paragraph pageId="0" pageNumber="1">Common names.</paragraph>
<paragraph pageId="0" pageNumber="1">
Tanzania: insungwe [Malila people] (
<bibRefCitation author="Schippers, RR" journalOrPublisher="Economic Botany" pageId="0" pageNumber="1" refId="B330" refString="Schippers, RR, 2000. African indigenous vegetables. An overview of the cultivated species. Natural Resources Institute/ACP-EU Technical Centre of Agricultural and Rural Cooperation, Chatham, UK." title="African indigenous vegetables. An overview of the cultivated species. Natural Resources Institute / ACP-EU Technical Centre of Agricultural and Rural Cooperation, Chatham, UK." year="2000">Schippers 2000</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="uses">
<paragraph pageId="0" pageNumber="1">Uses.</paragraph>
<paragraph pageId="0" pageNumber="1">Leaves used as spinach; berries eaten raw.</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="preliminary conservation status">
<paragraph pageId="0" pageNumber="1">Preliminary conservation status</paragraph>
<paragraph pageId="0" pageNumber="1">
(
<bibRefCitation author="IUCN" journalOrPublisher="Euphytica" pageId="0" pageNumber="1" publicationUrl="http://www.iucnredlist.org/documents/RedListGuidelines.pdf" refId="B176" refString="IUCN, 2016. Guidelines for Using the IUCN Red List Categories and Criteria. Version 12. Prepared by the Standards and Petitions Subcommittee. http://www.iucnredlist.org/documents/RedListGuidelines.pdf" title="Guidelines for Using the IUCN Red List Categories and Criteria. Version 12. Prepared by the Standards and Petitions Subcommittee." url="http://www.iucnredlist.org/documents/RedListGuidelines.pdf" year="2016">IUCN 2016</bibRefCitation>
). 
<taxonomicName class="Magnoliopsida" family="Solanaceae" genus="Solanum" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Solanum umalilaense" order="Centrales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="species" species="umalilaense">
<emphasis italics="true" pageId="0" pageNumber="1">Solanum umalilaense</emphasis>
</taxonomicName>
is known mostly from cultivation in the southern part of Tanzania; based on its range (EOO = 2,559 km2; EN) and the number of populations (AOO = 32 km2), it would be assigned a preliminary conservation status of EN (Endangered), but might be better considered DD (Data Deficient). It is protected by local people and its range is poorly known.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="discussion">
<paragraph pageId="0" pageNumber="1">Discussion.</paragraph>
<paragraph pageId="0" pageNumber="1">
<taxonomicName class="Magnoliopsida" family="Solanaceae" genus="Solanum" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Solanum umalilaense" order="Centrales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="species" species="umalilaense">
<emphasis italics="true" pageId="0" pageNumber="1">Solanum umalilaense</emphasis>
</taxonomicName>
can be distinguished from other African species of morelloids by its simple to branched and often leafy inflorescences, flowers with very short rounded calyx lobes and persistent, small, light yellowish-brown fruits. 
<taxonomicName class="Magnoliopsida" family="Solanaceae" genus="Solanum" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Solanum umalilaense" order="Centrales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="species" species="umalilaense">
<emphasis italics="true" pageId="0" pageNumber="1">Solanum umalilaense</emphasis>
</taxonomicName>
produces a large number of inflorescences such that, at full anthesis, the plant appears to be covered with white flowers, strikingly different from other species in the section (
<bibRefCitation DOI="https://doi.org/10.3897/phytokeys.16.2884" author="Manoko, MLK" journalOrPublisher="PhytoKeys" pageId="0" pageNumber="1" pagination="65 - 74" refId="B232" refString="Manoko, MLK, van der Weerden, GM, van den Berg, RG, Mariani, C, 2012. A new tetraploid species of Solanum L. sect. Solanum (Solanaceae) from Tanzania. PhytoKeys 16 (0): 65 - 74, DOI: https://doi.org/10.3897/phytokeys.16.2884" title="A new tetraploid species of Solanum L. sect. Solanum (Solanaceae) from Tanzania." url="https://doi.org/10.3897/phytokeys.16.2884" volume="16" year="2012">Manoko et al. 2012</bibRefCitation>
), except some populations of 
<taxonomicName family="Solanaceae" kingdom="Plantae" lsidName="S. tarderemotum" order="Solanales" pageId="0" pageNumber="1" rank="species" species="tarderemotum">
<emphasis italics="true" pageId="0" pageNumber="1">S. tarderemotum</emphasis>
</taxonomicName>
(the branched-inflorescence form previously recognised as 
<taxonomicName family="Solanaceae" kingdom="Plantae" lsidName="S. florulentum" order="Solanales" pageId="0" pageNumber="1" rank="species" species="florulentum">
<emphasis italics="true" pageId="0" pageNumber="1">S. florulentum</emphasis>
</taxonomicName>
). Like 
<taxonomicName family="Solanaceae" kingdom="Plantae" lsidName="S. tarderemotum" order="Solanales" pageId="0" pageNumber="1" rank="species" species="tarderemotum">
<emphasis italics="true" pageId="0" pageNumber="1">S. tarderemotum</emphasis>
</taxonomicName>
, the berries fall with the pedicels still attached.
</paragraph>
<paragraph pageId="0" pageNumber="1">
<bibRefCitation author="Manoko, MLK" journalOrPublisher="Trends in Ecology &amp; Evolution" pageId="0" pageNumber="1" publicationUrl="http://dare.ubn.kun.nl/bitstream/2066/30032/1/30032.pdf" refId="B227" refString="Manoko, MLK, 2007. A systematic study of African Solanum L. section Solanum (Solanaceae). PhD Thesis, Radboud University, Nijmegen, The Netherlands, http://dare.ubn.kun.nl/bitstream/2066/30032/1/30032.pdf" title="A systematic study of African Solanum L. section Solanum (Solanaceae). PhD Thesis, Radboud University, Nijmegen, The Netherlands," url="http://dare.ubn.kun.nl/bitstream/2066/30032/1/30032.pdf" year="2007">Manoko (2007)</bibRefCitation>
used AFLP molecular markers to assess the relationships of the tetraploid African species and found that 
<taxonomicName family="Solanaceae" kingdom="Plantae" lsidName="S. umalilaense" order="Solanales" pageId="0" pageNumber="1" rank="species" species="umalilaense">
<emphasis italics="true" pageId="0" pageNumber="1">S. umalilaense</emphasis>
</taxonomicName>
(as &quot;Sp. A&quot;) was a member of a cluster including 
<taxonomicName family="Solanaceae" kingdom="Plantae" lsidName="S. retroflexum" order="Solanales" pageId="0" pageNumber="1" rank="species" species="retroflexum">
<emphasis italics="true" pageId="0" pageNumber="1">S. retroflexum</emphasis>
</taxonomicName>
(as 
<taxonomicName family="Solanaceae" kingdom="Plantae" lsidName="S. hirsutum" order="Solanales" pageId="0" pageNumber="1" rank="species" species="hirsutum">
<emphasis italics="true" pageId="0" pageNumber="1">S. hirsutum</emphasis>
</taxonomicName>
(Vahl) Dunal and 
<taxonomicName family="Solanaceae" kingdom="Plantae" lsidName="S. retroflexum" order="Solanales" pageId="0" pageNumber="1" rank="species" species="retroflexum">
<emphasis italics="true" pageId="0" pageNumber="1">S. retroflexum</emphasis>
</taxonomicName>
) that itself clustered with 
<taxonomicName family="Solanaceae" kingdom="Plantae" lsidName="S. tarderemotum" order="Solanales" pageId="0" pageNumber="1" rank="species" species="tarderemotum">
<emphasis italics="true" pageId="0" pageNumber="1">S. tarderemotum</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph pageId="0" pageNumber="1">
More field work is needed in southern Tanzania and adjacent Zambia to determine the true range and morphological variation of 
<taxonomicName family="Solanaceae" kingdom="Plantae" lsidName="S. umalilaense" order="Solanales" pageId="0" pageNumber="1" rank="species" species="umalilaense">
<emphasis italics="true" pageId="0" pageNumber="1">S. umalilaense</emphasis>
</taxonomicName>
; collections analysed to date come from what might be a single population.
</paragraph>
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" pageId="0" pageNumber="1">Specimens examined. Tanzania</emphasis>
. 
<emphasis bold="true" pageId="0" pageNumber="1">Iringa</emphasis>
: Mufindi, Igowole, 10 Mar 1989, 
<emphasis italics="true" pageId="0" pageNumber="1">Kayombo</emphasis>
&amp; 
<emphasis italics="true" pageId="0" pageNumber="1">Kayombo 215</emphasis>
(MO); 
<emphasis bold="true" pageId="0" pageNumber="1">Mbeya</emphasis>
: Maganjo village, Lwindi ward, 8 Jul 2010, 
<emphasis italics="true" pageId="0" pageNumber="1">Manoko 2010-1</emphasis>
(DSM, NIJ, WAG); Isangati village, Iyunga ward, 8 Jul 2010, 
<emphasis italics="true" pageId="0" pageNumber="1">Manoko 2010-2</emphasis>
(DSM, NIJ, WAG); Igala village, Holondo ward, 8 Jul 2010, 
<emphasis italics="true" pageId="0" pageNumber="1">Manoko 2010-8</emphasis>
(DSM, NIJ, WAG); Isangati village, Iyunga Mapinduzi ward, 9 Jul 2010, 
<emphasis italics="true" pageId="0" pageNumber="1">Manoko 2010-11</emphasis>
(DSM, NIJ, WAG); Isangati village, Iyunga Mapinduzi ward, 9 Jul 2010, 
<emphasis italics="true" pageId="0" pageNumber="1">Manoko 2010-12</emphasis>
(DSM, NIJ, WAG); nr Umalila Forest reserve, 9 Jul 2010, 
<emphasis italics="true" pageId="0" pageNumber="1">Manoko 2010-14</emphasis>
(DSM, NIJ, WAG); Uyole, Mbeya, 22 May 1968, 
<emphasis italics="true" pageId="0" pageNumber="1">Mwambunga 6</emphasis>
(DSM).
</paragraph>
</subSubSection>
</treatment>
</document>