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<document ID-DOI="http://dx.doi.org/10.3897/zookeys.1086.68015" ID-GBIF-Dataset="24a3af2a-54f9-4077-afd3-7563950c5dbf" ID-PMC="PMC8866340" ID-Pensoft-Pub="1313-2970-1086-84" ID-Pensoft-UUID="6EDA4E3E12955CA7BFCCF667BADB6B90" ID-PubMed="35221746" ID-ZooBank="89728BCE242A49369095E9B544F8B9F7" ModsDocID="1313-2970-1086-84" checkinTime="1645053244411" checkinUser="pensoft" docAuthor="Crespo, Luis C., Silva, Isamberto, Enguidanos, Alba, Cardoso, Pedro &amp; Arnedo, Miquel" docDate="2022" docId="22F537099C4C5ECAB94F40A21CF3195E" docLanguage="en" docName="ZooKeys 1086: 84-135" docOrigin="ZooKeys 1086" docPubDate="2022-02-16" docSource="http://dx.doi.org/10.3897/zookeys.1086.68015" docTitle="Hogna insularum" docType="treatment" docVersion="4" id="6EDA4E3E12955CA7BFCCF667BADB6B90" lastPageNumber="84" masterDocId="6EDA4E3E12955CA7BFCCF667BADB6B90" masterDocTitle="Island hoppers: Integrative taxonomic revision of Hogna wolf spiders (Araneae, Lycosidae) endemic to the Madeira islands with description of a new species" masterLastPageNumber="135" masterPageNumber="84" pageNumber="84" updateTime="1668151492809" updateUser="ExternalLinkService">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>Island hoppers: Integrative taxonomic revision of Hogna wolf spiders (Araneae, Lycosidae) endemic to the Madeira islands with description of a new species</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Crespo, Luis C.</mods:namePart>
<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0002-5388-5661</mods:nameIdentifier>
<mods:affiliation>Department of Evolutionary Biology, Ecology and Environmental Sciences (Arthropods), Biodiversity Research Institute (IRBio), Universitat de Barcelona, Av. Diagonal 643, 08028 Barcelona, Spain &amp; Laboratory for Integrative Biodiversity Research (LIBRe), Finnish Museum of Natural History (LUOMUS), University of Helsinki, P. O. Box 17, 00014 Helsinki, Finland</mods:affiliation>
<mods:nameIdentifier type="email">luiscarloscrespo@gmail.com</mods:nameIdentifier>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Silva, Isamberto</mods:namePart>
<mods:affiliation>Instituto das Florestas e Conservacao da Natureza IP-RAM, Jardim Botanico da Madeira, Caminho do Meio, Bom Sucesso, 9064 - 512, Funchal, Portugal</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Enguidanos, Alba</mods:namePart>
<mods:affiliation>Department of Evolutionary Biology, Ecology and Environmental Sciences (Arthropods), Biodiversity Research Institute (IRBio), Universitat de Barcelona, Av. Diagonal 643, 08028 Barcelona, Spain</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Cardoso, Pedro</mods:namePart>
<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0001-8119-9960</mods:nameIdentifier>
<mods:affiliation>Laboratory for Integrative Biodiversity Research (LIBRe), Finnish Museum of Natural History (LUOMUS), University of Helsinki, P. O. Box 17, 00014 Helsinki, Finland</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Arnedo, Miquel</mods:namePart>
<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0003-1402-4727</mods:nameIdentifier>
<mods:affiliation>Department of Evolutionary Biology, Ecology and Environmental Sciences (Arthropods), Biodiversity Research Institute (IRBio), Universitat de Barcelona, Av. Diagonal 643, 08028 Barcelona, Spain</mods:affiliation>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>ZooKeys</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2022</mods:date>
<mods:detail type="pubDate">
<mods:number>2022-02-16</mods:number>
</mods:detail>
<mods:detail type="volume">
<mods:number>1086</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>84</mods:start>
<mods:end>135</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:location>
<mods:url>http://dx.doi.org/10.3897/zookeys.1086.68015</mods:url>
</mods:location>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.1086.68015</mods:identifier>
<mods:identifier type="Pensoft-Pub">1313-2970-1086-84</mods:identifier>
<mods:identifier type="ZooBank">89728BCE242A49369095E9B544F8B9F7</mods:identifier>
<mods:identifier type="Pensoft-UUID">6EDA4E3E12955CA7BFCCF667BADB6B90</mods:identifier>
</mods:mods>
<treatment ID-GBIF-Taxon="193383535" LSID="urn:lsid:plazi:treatment:22F537099C4C5ECAB94F40A21CF3195E" httpUri="http://treatment.plazi.org/id/22F537099C4C5ECAB94F40A21CF3195E" lastPageNumber="84" pageId="0" pageNumber="84">
<subSubSection pageId="0" pageNumber="84" type="nomenclature">
<paragraph pageId="0" pageNumber="84">
<taxonomicName LSID="22F53709-9C4C-5ECA-B94F-40A21CF3195E" authority="(Kulczynski, 1899)" baseAuthorityName="Kulczynski" baseAuthorityYear="1899" class="Arachnida" family="Lycosidae" genus="Hogna" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hogna insularum" order="Araneae" pageId="0" pageNumber="84" phylum="Arthropoda" rank="species" species="insularum">Hogna insularum (Kulczynski, 1899)</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="84" type="description">
<paragraph pageId="0" pageNumber="84">
<figureCitation captionStart="Figure 15" captionStartId="F15" captionText="Figure 15. Hogna insularum, male pedipalps A male from Porto Santo (CRBALC 0310), left pedipalp, ventral B male from Deserta Grande (CRBALC 0305), left pedipalp, ventral C male from Bugio (CRBALC 0015), left pedipalp, ventral D detail of the median apophysis of male from Deserta Grande (CRBALC 0305), anteroventral E SEM image, right male pedipalp, male from Porto Santo (CRBALC 0310), ventral F idem, retroventral. Abbreviations, male pedipalp: AT - anterior point, C - cymbium, E - embolus, P - palea, R - ridge, T - tegulum, TA - terminal apophysis, TgA - tegular apophysis, VS - ventral spur. Scale bars: 0.5 mm (A, B, C); 0.2 mm (D)." figureDoi="10.3897/zookeys.1086.68015.figure15" httpUri="https://binary.pensoft.net/fig/646995" pageId="0" pageNumber="84">Figures 15</figureCitation>
<figureCitation captionStart="Figure 16" captionStartId="F16" captionText="Figure 16. Hogna insularum, female genitalia A, B female from Bugio (CRBALC 0301): A epigynum, ventral B vulva, dorsal C, D female from Deserta Grande (CRBALC 0308): C epigynum, ventral D vulva, dorsal E, F female from Madeira (CRBALC 0597): E epigynum, ventral F vulva, dorsal G, H female from Porto Santo (CRBALC 0300): G epigynum, ventral H vulva, dorsal. Abbreviations, female genitalia: D - diverticulum, H - epigynal hoods, MS - median septum, S - spermatheca. Scale bars: 0.2 mm." figureDoi="10.3897/zookeys.1086.68015.figure16" httpUri="https://binary.pensoft.net/fig/646996" pageId="0" pageNumber="84">, 16</figureCitation>
<figureCitation captionStart="Figure 17" captionStartId="F17" captionText="Figure 17. Distribution of H. insularum. Circles: present records; black triangles: revised records from literature; white triangles: unconfirmed records from literature." figureDoi="10.3897/zookeys.1086.68015.figure17" httpUri="https://binary.pensoft.net/fig/646997" pageId="0" pageNumber="84">, 17</figureCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="84" type="reference_group">
<paragraph pageId="0" pageNumber="84">
<taxonomicName authorityName="Kulczynski" authorityYear="1899" class="Arachnida" family="Lycosidae" genus="Trochosa" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Trochosa insularum" order="Araneae" pageId="0" pageNumber="84" phylum="Arthropoda" rank="species" species="insularum">Trochosa insularum</taxonomicName>
Kulczynski, 1899: 429, pl. 9, f. 122, 126 (Dmf).
</paragraph>
<paragraph pageId="0" pageNumber="84">
<taxonomicName baseAuthorityName="Kulczynski" baseAuthorityYear="1899" class="Arachnida" family="Lycosidae" genus="Hogna" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hogna insularum" order="Araneae" pageId="0" pageNumber="84" phylum="Arthropoda" rank="species" species="insularum">Hogna insularum</taxonomicName>
Roewer, 1959: 517, fig. 291c, d.
</paragraph>
<paragraph pageId="0" pageNumber="84">
<taxonomicName authorityName="Wunderlich" authorityYear="1992" class="Arachnida" family="Lycosidae" genus="Hogna" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hogna biscoitoi" order="Araneae" pageId="0" pageNumber="84" phylum="Arthropoda" rank="species" species="biscoitoi">Hogna biscoitoi</taxonomicName>
Wunderlich, 1992: 457, figs 708-709. Syn. nov.
</paragraph>
<paragraph pageId="0" pageNumber="84">
<taxonomicName baseAuthorityName="Kulczynski" baseAuthorityYear="1899" class="Arachnida" family="Lycosidae" genus="Hogna" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hogna insularum" order="Araneae" pageId="0" pageNumber="84" phylum="Arthropoda" rank="species" species="insularum">Hogna insularum</taxonomicName>
Wunderlich, 1995: 415, fig. 27 (m).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="84" type="types">
<paragraph pageId="0" pageNumber="84">Types.</paragraph>
<paragraph pageId="0" pageNumber="84">
<taxonomicName authorityName="Wunderlich" authorityYear="1992" class="Arachnida" family="Lycosidae" genus="Hogna" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hogna biscoitoi" order="Araneae" pageId="0" pageNumber="84" phylum="Arthropoda" rank="species" species="biscoitoi">
<emphasis italics="true" pageId="0" pageNumber="84">Hogna biscoitoi</emphasis>
</taxonomicName>
Holotype ♂ without exact locality, Porto Santo; leg. Winkelmayer, stored at MMF, collection number 24551. Not examined.
</paragraph>
<paragraph pageId="0" pageNumber="84">
<emphasis bold="true" italics="true" pageId="0" pageNumber="84">Syntypes</emphasis>
: Madeira • 7 ♀♀ (MIZ217320-217326). Porto Santo • 2 ♂♂ and 14 ♀♀ and 1 juvenile (MIZ217327-217343), leg. Kulczynski, stored at MIZ, collection numbers indicated above. Examined 2 ♂♂ from Porto Santo, 1 ♀ from Madeira.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="84" type="materials_examined">
<paragraph pageId="0" pageNumber="84">Material examined.</paragraph>
<paragraph pageId="0" pageNumber="84">
<materialsCitation collectingDate="2012-06-28" collectorName="I. Silva" latitude="32.41228" location="Bugio" longLatPrecision="1" longitude="-16.47466" municipality="Planalto Sul" specimenCount="2" specimenCount-female="1" specimenCount-male="1">
<location LSID="urn:lsid:plazi:treatment:22F537099C4C5ECAB94F40A21CF3195E:85243936B15B63284FE21FCBDE1511AC" latitude="32.41228" longLatPrecision="1" longitude="-16.47466" municipality="Planalto Sul" name="Bugio">Bugio</location>
<collectingMunicipality>Planalto Sul</collectingMunicipality>
,
<geoCoordinate degrees="32.41228" direction="north" orientation="latitude" precision="1" value="32.41228">32.41228°N</geoCoordinate>
,
<geoCoordinate degrees="16.47466" direction="west" orientation="longitude" precision="1" value="-16.47466">16.47466°W</geoCoordinate>
,
<specimenCount type="male">1 ♂</specimenCount>
(CRBALC0015) and
<specimenCount type="female">1 ♀</specimenCount>
(CRBALC0017),
<collectingDate value="2012-06-28">28.VI.2012</collectingDate>
, hand collecting, leg.
<collectorName>I. Silva</collectorName>
</materialsCitation>
,
<specimenCount type="male">1 ♂</specimenCount>
(CRBALC0316: LC229),
<specimenCount type="female">1 ♀</specimenCount>
(CRBALC0301: LC190) and
<specimenCount type="juvenile">2 juveniles</specimenCount>
(CRBALC0315: LC228, CRBALC0318: LC231),
<date value="2017-04-13">13.IV.2017</date>
, hand collecting, leg. L. Crespo. Deserta Grande •
<materialsCitation collectingDate="2011-04-17" collectingDateMax="2017-04-11" collectingDateMin="2011-04-17" collectorName="I. Silva" latitude="32.50993" location="Eira" longLatPrecision="1" longitude="-16.5024" specimenCount="3" specimenCount-female="1" specimenCount-juvenile="2">
<location LSID="urn:lsid:plazi:treatment:22F537099C4C5ECAB94F40A21CF3195E:2C7A1002A81E6B031A4C532B575923E0" latitude="32.50993" longLatPrecision="1" longitude="-16.5024" name="Eira">Eira</location>
,
<geoCoordinate degrees="32.50993" direction="north" orientation="latitude" precision="1" value="32.50993">32.50993°N</geoCoordinate>
,
<geoCoordinate degrees="16.50240" direction="west" orientation="longitude" precision="1" value="-16.5024">16.50240°W</geoCoordinate>
,
<specimenCount type="juvenile">2 juveniles</specimenCount>
(CRBALC0312: LC282), CRBALC0319: LC232),
<collectingDate value="2017-04-11">11.IV.2017</collectingDate>
,
<specimenCount type="female">1 ♀</specimenCount>
(FMNHhttp://id.luomus.fi/HLA.148894),
<collectingDate value="2011-04-17">17.IV.2011</collectingDate>
, hand collecting, leg.
<collectorName>I. Silva</collectorName>
</materialsCitation>
;
<materialsCitation collectingDate="1981-08-12" collectorName="J. Murphy" location="North" specimenCount="1" specimenCount-male="1">
<location LSID="urn:lsid:plazi:treatment:22F537099C4C5ECAB94F40A21CF3195E:95EE5DBEE4F522ADA2F4FB89294A952F" name="North">North</location>
end,
<specimenCount type="male">1 ♂</specimenCount>
(MMUE G7508.51),
<collectingDate value="1981-08-12">12.VIII.1981</collectingDate>
, under stone, leg.
<collectorName>J. Murphy</collectorName>
</materialsCitation>
;
<materialsCitation collectingDate="2017-04-08" collectorName="L. Crespo, I. Silva" latitude="32.54613" location="Pedregal" longLatPrecision="4" longitude="-16.5234" specimenCount="2" specimenCount-female="1" specimenCount-juvenile="1">
<location LSID="urn:lsid:plazi:treatment:22F537099C4C5ECAB94F40A21CF3195E:3EC5CDE87DE5FA25E49C2514AF225FAC" latitude="32.54613" longLatPrecision="4" longitude="-16.5234" name="Pedregal">Pedregal</location>
(E),
<geoCoordinate degrees="32.54613" direction="north" orientation="latitude" precision="1" value="32.54613">32.54613°N</geoCoordinate>
,
<geoCoordinate degrees="16.5234" direction="west" orientation="longitude" precision="5" value="-16.5234">16.5234°W</geoCoordinate>
,
<specimenCount type="female">1 ♀</specimenCount>
(CRBALC0308: LC197) and
<specimenCount type="juvenile">1 juvenile</specimenCount>
(CRBALC0306: LC195),
<collectingDate value="2017-04-08">8.IV.2017</collectingDate>
, hand collecting, leg.
<collectorName>L. Crespo</collectorName>
&amp;
<collectorName>I. Silva</collectorName>
</materialsCitation>
,
<specimenCount type="juvenile">1 juvenile</specimenCount>
(CRBALC0285: LC185),
<date value="2017-04-09">9.IV.2017</date>
, hand collecting, leg. L. Crespo;
<materialsCitation collectingDate="2017-04-11" collectorName="L. Crespo, I. Silva" latitude="32.50596" location="Planalto Sul" longLatPrecision="1" longitude="-16.49986" specimenCount="1" specimenCount-juvenile="1">
<location LSID="urn:lsid:plazi:treatment:22F537099C4C5ECAB94F40A21CF3195E:8A91BFED9E551C690684DAFC4C6DF80A" latitude="32.50596" longLatPrecision="1" longitude="-16.49986" name="Planalto Sul">Planalto Sul</location>
,
<geoCoordinate degrees="32.50596" direction="north" orientation="latitude" precision="1" value="32.50596">32.50596°N</geoCoordinate>
,
<geoCoordinate degrees="16.49986" direction="west" orientation="longitude" precision="1" value="-16.49986">16.49986°W</geoCoordinate>
,
<specimenCount type="juvenile">1 juvenile</specimenCount>
(CRBALC0413: LC284),
<collectingDate value="2017-04-11">11.IV.2017</collectingDate>
, hand collecting, leg.
<collectorName>L. Crespo</collectorName>
&amp;
<collectorName>I. Silva</collectorName>
</materialsCitation>
;
<materialsCitation collectingDate="2017-04-10" collectorName="L. Crespo, I. Silva" latitude="32.53168" location="Rocha do Barbusano" longLatPrecision="1" longitude="-16.51471" specimenCount="1" specimenCount-juvenile="1">
<location LSID="urn:lsid:plazi:treatment:22F537099C4C5ECAB94F40A21CF3195E:E4F17DFF37E674B1EDE1E2EBC47AFDA4" latitude="32.53168" longLatPrecision="1" longitude="-16.51471" name="Rocha do Barbusano">Rocha do Barbusano</location>
(S),
<geoCoordinate degrees="32.53168" direction="north" orientation="latitude" precision="1" value="32.53168">32.53168°N</geoCoordinate>
,
<geoCoordinate degrees="16.51471" direction="west" orientation="longitude" precision="1" value="-16.51471">16.51471°W</geoCoordinate>
,
<specimenCount type="juvenile">1 juvenile</specimenCount>
(CRBALC0262: LC175),
<collectingDate value="2017-04-10">10.IV.2017</collectingDate>
, hand collecting, leg.
<collectorName>L. Crespo</collectorName>
&amp;
<collectorName>I. Silva</collectorName>
</materialsCitation>
;
<materialsCitation collectingDate="2011-04-23" collectorName="I. Silva" location="Vale da Castanheira" specimenCount="1" specimenCount-male="1">
<location LSID="urn:lsid:plazi:treatment:22F537099C4C5ECAB94F40A21CF3195E:C9CC388F2D6B3157E10E31CDD6DBAB47" name="Vale da Castanheira">Vale da Castanheira</location>
,
<specimenCount type="male">1 ♂</specimenCount>
(FMNHhttp://id.luomus.fi/HLA.148961),
<collectingDate value="2011-04-23">23.IV.2011</collectingDate>
, hand collecting, leg.
<collectorName>I. Silva</collectorName>
</materialsCitation>
<emphasis italics="true" pageId="0" pageNumber="84">et al.</emphasis>
,
<specimenCount type="male">1 ♂</specimenCount>
(FMNHhttp://id.luomus.fi/HLA.148976),
<date value="2011-05-05">5.V.2011</date>
, pitfall trapping, leg. I. Silva
<emphasis italics="true" pageId="0" pageNumber="84">et al.</emphasis>
,
<specimenCount type="female">1 ♀</specimenCount>
(FMNHhttp://id.luomus.fi/HLA.148982),
<specimenCount type="female">2 ♀♀</specimenCount>
(FMNHhttp://id.luomus.fi/HLA.148986),
<date value="2011-04-22">22.IV.2011</date>
, hand collecting, leg. I. Silva;
<materialsCitation collectingDate="2017-04-09" collectorName="I. Silva" latitude="32.5571" location="Vale da Castanheira" longLatPrecision="5" longitude="-16.52963" specimenCount="1" specimenCount-male="1">
<location LSID="urn:lsid:plazi:treatment:22F537099C4C5ECAB94F40A21CF3195E:CCD5A6521E613F74A4B0D70E24EB23D3" latitude="32.5571" longLatPrecision="5" longitude="-16.52963" name="Vale da Castanheira">Vale da Castanheira</location>
(E),
<geoCoordinate degrees="32.5571" direction="north" orientation="latitude" precision="5" value="32.5571">32.5571°N</geoCoordinate>
,
<geoCoordinate degrees="16.52963" direction="west" orientation="longitude" precision="1" value="-16.52963">16.52963°W</geoCoordinate>
,
<specimenCount type="male">1 ♂</specimenCount>
(CRBALC0305: LC194),
<collectingDate value="2017-04-09">9.IV.2017</collectingDate>
, hand collecting, leg.
<collectorName>I. Silva</collectorName>
</materialsCitation>
;
<materialsCitation collectingDate="2017-04-09" collectorName="I. Silva. Ilheu da Cal" latitude="32.55078" location="Vale da Castanheira" longLatPrecision="1" longitude="-16.52541" specimenCount="3" specimenCount-female="1" specimenCount-male="2">
<location LSID="urn:lsid:plazi:treatment:22F537099C4C5ECAB94F40A21CF3195E:4065F638F444E6690EE5F39DF00793CD" latitude="32.55078" longLatPrecision="1" longitude="-16.52541" name="Vale da Castanheira">Vale da Castanheira</location>
(SE),
<geoCoordinate degrees="32.55078" direction="north" orientation="latitude" precision="1" value="32.55078">32.55078°N</geoCoordinate>
,
<geoCoordinate degrees="16.52541" direction="west" orientation="longitude" precision="1" value="-16.52541">16.52541°W</geoCoordinate>
,
<specimenCount type="male">2 ♂♂</specimenCount>
(CRBALC0313: LC226, CRBALC0349: LC241) and
<specimenCount type="female">1 ♀</specimenCount>
(CRBALC0348: LC240),
<collectingDate value="2017-04-09">9.IV.2017</collectingDate>
, hand collecting, leg.
<collectorName>
I. Silva.
<normalizedToken originalValue="Ilhéu">Ilheu</normalizedToken>
da Cal
</collectorName>
</materialsCitation>
<materialsCitation accessionNumber="SMF65693" collectingDate="2022-01-01" collectingDateMax="2022-12-31" collectingDateMin="2022-01-01" location="Groh. Ilheu de Cima" specimenCount="1" specimenCount-female="1">
<specimenCount type="female">1 ♀</specimenCount>
(
<accessionNumber httpUri="http://www.ncbi.nlm.nih.gov/protein/SMF65693">SMF65693</accessionNumber>
), leg. K.
<location LSID="urn:lsid:plazi:treatment:22F537099C4C5ECAB94F40A21CF3195E:5C163B131F19C4A4FE1A86C25590799A" name="Groh. Ilheu de Cima">
Groh.
<normalizedToken originalValue="Ilhéu">Ilheu</normalizedToken>
de Cima
</location>
</materialsCitation>
• top plateau,
<geoCoordinate degrees="33.05556" direction="north" orientation="latitude" precision="1" value="33.05556">33.05556°N</geoCoordinate>
,
<geoCoordinate degrees="16.28097" direction="west" orientation="longitude" precision="1" value="-16.28097">16.28097°W</geoCoordinate>
,
<specimenCount type="female">1 ♀</specimenCount>
(CRBALC0019),
<date value="2012-04-09">9.IV.2012</date>
, hand collecting, leg. I. Silva,
<specimenCount type="male">1 ♂</specimenCount>
(CRBALC0018),
<date value="2011-05-22">22.V.2011</date>
, hand collecting, leg. I. Silva,
<materialsCitation collectingDate="2017-04-19" collectorName="L. Crespo" location="Silva. Ilheu de Ferro" specimenCount="5" specimenCount-female="1" specimenCount-juvenile="4">
<specimenCount type="female">1 ♀</specimenCount>
(CRBALC0302: LC191) and
<specimenCount type="juvenile">4 juveniles</specimenCount>
(CRBALC0284: LC183, CRBALC0311: LC225, CRBALC0320: LC233, CRBALC0321: LC234),
<collectingDate value="2017-04-19">19.IV.2017</collectingDate>
, hand collecting, leg.
<collectorName>L. Crespo</collectorName>
&amp; I.
<location LSID="urn:lsid:plazi:treatment:22F537099C4C5ECAB94F40A21CF3195E:BEA9B8415A0AD90CA9AC2EA570B2B5A5" name="Silva. Ilheu de Ferro">
Silva.
<normalizedToken originalValue="Ilhéu">Ilheu</normalizedToken>
de Ferro
</location>
</materialsCitation>
<materialsCitation collectingDate="2017-04-18" collectorName="L. Crespo" latitude="33.03698" location="South" longLatPrecision="1" longitude="-16.40814" municipality="Silva. Ilheu Do Desembarcadouro" specimenCount="3" specimenCount-female="1" specimenCount-juvenile="2">
<location LSID="urn:lsid:plazi:treatment:22F537099C4C5ECAB94F40A21CF3195E:B2578BED0AD32D4D56C07EC9AAB2CB8C" latitude="33.03698" longLatPrecision="1" longitude="-16.40814" municipality="Silva. Ilheu Do Desembarcadouro" name="South">South</location>
tip,
<geoCoordinate degrees="33.03698" direction="north" orientation="latitude" precision="1" value="33.03698">33.03698°N</geoCoordinate>
,
<geoCoordinate degrees="16.40814" direction="west" orientation="longitude" precision="1" value="-16.40814">16.40814°W</geoCoordinate>
,
<specimenCount type="female">1 ♀</specimenCount>
(CRBALC0317: LC320) and
<specimenCount type="juvenile">2 juveniles</specimenCount>
(CRBALC0265: LC178, CRBALC0266: LC179),
<collectingDate value="2017-04-18">18.IV.2017</collectingDate>
, hand collecting, leg.
<collectorName>L. Crespo</collectorName>
&amp; I.
<collectingMunicipality>
Silva.
<normalizedToken originalValue="Ilhéu">Ilheu</normalizedToken>
Do Desembarcadouro
</collectingMunicipality>
</materialsCitation>
<specimenCount type="female">2 ♀♀</specimenCount>
(MMUE G7508.50),
<date value="1981-08-28">28.VIII.1981</date>
, under stone, leg. J. Murphy. Madeira •
<materialsCitation collectingDate="2017-03-30" collectorName="I. Silva" latitude="32.7419" location="Cais do Sardinha" longLatPrecision="5" longitude="-16.68317" specimenCount="5" specimenCount-juvenile="5">
<location LSID="urn:lsid:plazi:treatment:22F537099C4C5ECAB94F40A21CF3195E:FD6B0A837903515388CD6437FAF2E32C" latitude="32.7419" longLatPrecision="5" longitude="-16.68317" name="Cais do Sardinha">Cais do Sardinha</location>
,
<geoCoordinate degrees="32.7419" direction="north" orientation="latitude" precision="5" value="32.7419">32.7419°N</geoCoordinate>
,
<geoCoordinate degrees="16.68317" direction="west" orientation="longitude" precision="1" value="-16.68317">16.68317°W</geoCoordinate>
,
<specimenCount type="juvenile">5 juveniles</specimenCount>
(CRBALC0504: LC242, CRBALC0505: LC243, CRBALC0506: LC244, CRBALC0507: LC245, CRBALC0508: LC246),
<collectingDate value="2017-03-30">30.III.2017</collectingDate>
, hand collecting, leg.
<collectorName>I. Silva</collectorName>
</materialsCitation>
;
<materialsCitation collectingDate="1973-04-24" collectorName="J. Murphy" location="Canical" specimenCount="1" specimenCount-female="1">
<normalizedToken originalValue="Caniçal">
<location LSID="urn:lsid:plazi:treatment:22F537099C4C5ECAB94F40A21CF3195E:822F37866C77A1DE942180B5CCC48855" name="Canical">Canical</location>
</normalizedToken>
,
<specimenCount type="female">1 ♀</specimenCount>
(MMUE G7572.859),
<collectingDate value="1973-04-24">24.IV.1973</collectingDate>
, leg.
<collectorName>J. Murphy</collectorName>
</materialsCitation>
;
<materialsCitation collectingDate="1981-08-11" collectorName="J. Murphy" location="Canico" specimenCount="1" specimenCount-female="1">
<normalizedToken originalValue="Caniço">
<location LSID="urn:lsid:plazi:treatment:22F537099C4C5ECAB94F40A21CF3195E:A5649C78E03E5C60BDE3762655CF24BB" name="Canico">Canico</location>
</normalizedToken>
,
<specimenCount type="female">1 ♀</specimenCount>
(MMUE G7508.58),
<collectingDate value="1981-08-11">11.VIII.1981</collectingDate>
, under stone, leg.
<collectorName>J. Murphy</collectorName>
</materialsCitation>
;
<materialsCitation collectingDate="1981-07-29" collectingDateMax="1981-08-01" collectingDateMin="1981-07-29" collectorName="J. Murphy" location="Ponta de Sao Lourenco" specimenCount="2" specimenCount-female="1" specimenCount-male="1">
<location LSID="urn:lsid:plazi:treatment:22F537099C4C5ECAB94F40A21CF3195E:E19DE5DE420E4096CDCC092E6E88A5E8" name="Ponta de Sao Lourenco">
Ponta de
<normalizedToken originalValue="São">Sao</normalizedToken>
<normalizedToken originalValue="Lourenço">Lourenco</normalizedToken>
</location>
,
<specimenCount type="male">1 ♂</specimenCount>
(MMUE G7508.54),
<collectingDate value="1981-07-29">29.VII.1981</collectingDate>
,
<specimenCount type="female">1 ♀</specimenCount>
(MMUE G7508.57),
<collectingDate value="1981-08-01">1.VIII.1981</collectingDate>
, under stone, leg.
<collectorName>J. Murphy</collectorName>
</materialsCitation>
,
<specimenCount type="male">4 ♂♂</specimenCount>
and
<specimenCount type="female">5 ♀♀</specimenCount>
(FMNHhttp://id.luomus.fi/HLA.156001),
<date value="2011-05-15">15.V.2011</date>
, pitfall trapping, leg. L. Crespo et al.,
<specimenCount type="male">1 ♂</specimenCount>
and
<specimenCount type="female">4 ♀♀</specimenCount>
(FMNHhttp://id.luomus.fi/HLA.156012),
<date value="2011-05-02">2.V.2011</date>
, hand collecting, leg. L. Crespo et al.,
<specimenCount type="female">2 ♀♀</specimenCount>
(FMNHhttp://id.luomus.fi/HLA.156034),
<date value="2009-09-26">26.IX.2009</date>
, hand collecting, leg. L. Crespo,
<specimenCount type="female">1 ♀</specimenCount>
(CRBALC0597) and
<specimenCount type="juvenile">3 juveniles</specimenCount>
(CRBALC0599, CRBALC0600, CRBALC0651),
<geoCoordinate degrees="32.749965" direction="north" orientation="latitude" precision="1" value="32.749966">32.749965°N</geoCoordinate>
,
<geoCoordinate degrees="16.692817" direction="west" orientation="longitude" precision="1" value="-16.692818">16.692817°W</geoCoordinate>
,
<date value="2018-04-02">2.IV.2018</date>
, hand collecting, leg. L. Crespo;
<materialsCitation collectingDate="2017-03-30" collectorName="I. Silva. Porto Santo" latitude="32.75022" location="Ponta do Rosto" longLatPrecision="1" longitude="-16.70833" specimenCount="4" specimenCount-female="1" specimenCount-juvenile="3">
<location LSID="urn:lsid:plazi:treatment:22F537099C4C5ECAB94F40A21CF3195E:7C6EAC252D351ABE511FA7D5DF60BB59" latitude="32.75022" longLatPrecision="1" longitude="-16.70833" name="Ponta do Rosto">Ponta do Rosto</location>
,
<specimenCount type="female">1 ♀</specimenCount>
(CRBALC0513: LC251) and
<specimenCount type="juvenile">3 juveniles</specimenCount>
(CRBALC0509: LC247, CRBALC0510: LC248, CRBALC512: LC250),
<geoCoordinate degrees="32.75022" direction="north" orientation="latitude" precision="1" value="32.75022">32.75022°N</geoCoordinate>
,
<geoCoordinate degrees="16.70833" direction="west" orientation="longitude" precision="1" value="-16.70833">16.70833°W</geoCoordinate>
,
<collectingDate value="2017-03-30">30.III.2017</collectingDate>
, hand collecting, leg.
<collectorName>I. Silva. Porto Santo</collectorName>
</materialsCitation>
<materialsCitation collectingDate="2017-04-21" collectorName="L. Crespo, I. Silva" latitude="33.07353" location="Rocha de Nossa Senhora" longLatPrecision="4" longitude="-16.3212" specimenCount="2" specimenCount-juvenile="1" specimenCount-male="1">
<location LSID="urn:lsid:plazi:treatment:22F537099C4C5ECAB94F40A21CF3195E:703D1D5976A30228D0ABD518074E1D51" latitude="33.07353" longLatPrecision="4" longitude="-16.3212" name="Rocha de Nossa Senhora">Rocha de Nossa Senhora</location>
,
<geoCoordinate degrees="33.07353" direction="north" orientation="latitude" precision="1" value="33.07353">33.07353°N</geoCoordinate>
,
<geoCoordinate degrees="16.3212" direction="west" orientation="longitude" precision="5" value="-16.3212">16.3212°W</geoCoordinate>
,
<specimenCount type="male">1 ♂</specimenCount>
(CRBALC0290: LC187) and
<specimenCount type="juvenile">1 juvenile</specimenCount>
(CRBALC0291: LC188),
<collectingDate value="2017-04-21">21.IV.2017</collectingDate>
, hand collecting, leg.
<collectorName>L. Crespo</collectorName>
&amp;
<collectorName>I. Silva</collectorName>
</materialsCitation>
;
<materialsCitation accessionNumber="SMF65689" collectingDate="1983-07-07" collectorName="K. Groh" location="Pedras Vermelhas" specimenCount="3" specimenCount-juvenile="1" specimenCount-male="2">
<location LSID="urn:lsid:plazi:treatment:22F537099C4C5ECAB94F40A21CF3195E:38BC9FE8779C4A400108AA43C24DEA08" name="Pedras Vermelhas">Pedras Vermelhas</location>
,
<specimenCount type="male">2 ♂♂</specimenCount>
and
<specimenCount type="juvenile">1 juvenile</specimenCount>
(
<accessionNumber httpUri="http://www.ncbi.nlm.nih.gov/protein/SMF65689">SMF65689</accessionNumber>
),
<collectingDate value="1983-07-07">7.VII.1983</collectingDate>
, leg.
<collectorName>K. Groh</collectorName>
</materialsCitation>
;
<materialsCitation collectingDate="2017-04-20" collectorName="L. Crespo, I. Silva" latitude="33.04728" location="Pico Ana Ferreira" longLatPrecision="1" longitude="-16.37171" specimenCount="7" specimenCount-female="1" specimenCount-juvenile="5" specimenCount-male="1">
<location LSID="urn:lsid:plazi:treatment:22F537099C4C5ECAB94F40A21CF3195E:D667F9B457402454EE2BCC0B74070808" latitude="33.04728" longLatPrecision="1" longitude="-16.37171" name="Pico Ana Ferreira">Pico Ana Ferreira</location>
,
<geoCoordinate degrees="33.04728" direction="north" orientation="latitude" precision="1" value="33.04728">33.04728°N</geoCoordinate>
,
<geoCoordinate degrees="16.37171" direction="west" orientation="longitude" precision="1" value="-16.37171">16.37171°W</geoCoordinate>
,
<specimenCount type="male">1 ♂</specimenCount>
(CRBALC0310: LC224),
<specimenCount type="female">1 ♀</specimenCount>
(CRBALC0327: LC239) and
<specimenCount type="juvenile">5 juveniles</specimenCount>
(CRBALC0303: LC192, CRBALC0307: LC196, CRBALC0326: LC238, CRBALC0309: LC281, CRBALC0430: LC285),
<collectingDate value="2017-04-20">20.IV.2017</collectingDate>
, hand collecting, leg.
<collectorName>L. Crespo</collectorName>
&amp;
<collectorName>I. Silva</collectorName>
</materialsCitation>
;
<materialsCitation collectingDate="2017-04-21" collectorName="L. Crespo, I. Silva" latitude="33.09428" location="Pico Branco" longLatPrecision="1" longitude="-16.30137" specimenCount="1" specimenCount-male="1">
<location LSID="urn:lsid:plazi:treatment:22F537099C4C5ECAB94F40A21CF3195E:ACCD076EE9FDBB3574B00391F47348DA" latitude="33.09428" longLatPrecision="1" longitude="-16.30137" name="Pico Branco">Pico Branco</location>
,
<geoCoordinate degrees="33.09428" direction="north" orientation="latitude" precision="1" value="33.09428">33.09428°N</geoCoordinate>
,
<geoCoordinate degrees="16.30137" direction="west" orientation="longitude" precision="1" value="-16.30137">16.30137°W</geoCoordinate>
,
<specimenCount type="male">1 ♂</specimenCount>
(CRBALC0304: LC193),
<collectingDate value="2017-04-21">21.IV.2017</collectingDate>
, hand collecting, leg.
<collectorName>L. Crespo</collectorName>
&amp;
<collectorName>I. Silva</collectorName>
</materialsCitation>
,
<specimenCount type="male">1 ♂</specimenCount>
(CRBALC0314: LC227),
<date value="2017-04-23">23.IV.2017</date>
, hand collecting, leg. L. Crespo;
<materialsCitation collectingDate="2017-04-24" collectorName="L. Crespo" latitude="33.0927" location="Pico da Juliana" longLatPrecision="1" longitude="-16.32186" specimenCount="1" specimenCount-juvenile="1">
<location LSID="urn:lsid:plazi:treatment:22F537099C4C5ECAB94F40A21CF3195E:2C0896AC76B34C49871F972F5093CC68" latitude="33.0927" longLatPrecision="1" longitude="-16.32186" name="Pico da Juliana">Pico da Juliana</location>
,
<geoCoordinate degrees="33.09270" direction="north" orientation="latitude" precision="1" value="33.0927">33.09270°N</geoCoordinate>
,
<geoCoordinate degrees="16.32186" direction="west" orientation="longitude" precision="1" value="-16.32186">16.32186°W</geoCoordinate>
,
<specimenCount type="juvenile">1 juvenile</specimenCount>
(CRBALC0286: LC186),
<collectingDate value="2017-04-24">24.IV.2017</collectingDate>
, hand collecting, leg.
<collectorName>L. Crespo</collectorName>
</materialsCitation>
;
<materialsCitation collectingDate="2017-04-17" collectorName="L. Crespo, I. Silva" latitude="33.08196" location="Pico do Castelo" longLatPrecision="1" longitude="-16.33277" specimenCount="4" specimenCount-female="2" specimenCount-juvenile="2">
<location LSID="urn:lsid:plazi:treatment:22F537099C4C5ECAB94F40A21CF3195E:8AF37CF8F00B4E105404BD89006191DB" latitude="33.08196" longLatPrecision="1" longitude="-16.33277" name="Pico do Castelo">Pico do Castelo</location>
,
<geoCoordinate degrees="33.08196" direction="north" orientation="latitude" precision="1" value="33.08196">33.08196°N</geoCoordinate>
,
<geoCoordinate degrees="16.33277" direction="west" orientation="longitude" precision="1" value="-16.33277">16.33277°W</geoCoordinate>
,
<specimenCount type="female">2 ♀♀</specimenCount>
(CRBALC0300: LC189, CRBALC0322: LC235) and
<specimenCount type="juvenile">2 juveniles</specimenCount>
(CRBALC0267: LC180, CRBALC0268: 181),
<collectingDate value="2017-04-17">17.IV.2017</collectingDate>
, hand collecting, leg.
<collectorName>L. Crespo</collectorName>
&amp;
<collectorName>I. Silva</collectorName>
</materialsCitation>
,
<specimenCount type="male">1 ♂</specimenCount>
(CRBALC0692),
<date value="2018-04-08">8.IV.2018</date>
, hand collecting, leg. L. Crespo &amp; A. Bellvert;
<materialsCitation accessionNumber="SMF65695" collectingDate="1983-06-29" collectorName="K. Groh" location="Pico do Concelho" specimenCount="1" specimenCount-female="1">
<location LSID="urn:lsid:plazi:treatment:22F537099C4C5ECAB94F40A21CF3195E:82561D2F1A87BF75CE29E6BBD09C6CF1" name="Pico do Concelho">Pico do Concelho</location>
,
<specimenCount type="female">1 ♀</specimenCount>
(
<accessionNumber httpUri="http://www.ncbi.nlm.nih.gov/protein/SMF65695">SMF65695</accessionNumber>
),
<collectingDate value="1983-06-29">29.VI.1983</collectingDate>
, leg.
<collectorName>K. Groh</collectorName>
</materialsCitation>
;
<materialsCitation accessionNumber="SMF65692" collectingDate="1983-07-01" collectorName="K. Groh" location="Pico do Espigao" specimenCount="1" specimenCount-female="1">
<location LSID="urn:lsid:plazi:treatment:22F537099C4C5ECAB94F40A21CF3195E:0D77ABB8E6130C9130896ADE313ABBF7" name="Pico do Espigao">
Pico do
<normalizedToken originalValue="Espigão">Espigao</normalizedToken>
</location>
,
<specimenCount type="female">1 ♀</specimenCount>
(
<accessionNumber httpUri="http://www.ncbi.nlm.nih.gov/protein/SMF65692">SMF65692</accessionNumber>
),
<collectingDate value="1983-07-01">1.VII.1983</collectingDate>
, leg.
<collectorName>K. Groh</collectorName>
</materialsCitation>
;
<materialsCitation accessionNumber="SMF65694" collectingDate="1983-06-28" collectorName="K. Groh" location="Pico do Facho" specimenCount="1" specimenCount-female="1">
<location LSID="urn:lsid:plazi:treatment:22F537099C4C5ECAB94F40A21CF3195E:90C2494E98152EACF7CF284B13BEC7F6" name="Pico do Facho">Pico do Facho</location>
,
<specimenCount type="female">1 ♀</specimenCount>
(
<accessionNumber httpUri="http://www.ncbi.nlm.nih.gov/protein/SMF65694">SMF65694</accessionNumber>
),
<collectingDate value="1983-06-28">28.VI.1983</collectingDate>
, leg.
<collectorName>K. Groh</collectorName>
</materialsCitation>
;
<materialsCitation accessionNumber="SMF65691" collectingDate="1983-07-10" collectorName="Pico dos Magaricos, K. Groh" location="Pico do Macarico" specimenCount="1" specimenCount-female="1">
<location LSID="urn:lsid:plazi:treatment:22F537099C4C5ECAB94F40A21CF3195E:E5C6A121607739CA809963DDC6650BCB" name="Pico do Macarico">
Pico do
<normalizedToken originalValue="Maçarico">Macarico</normalizedToken>
</location>
[the label reads &quot;
<collectorName>Pico dos Magaricos</collectorName>
&quot;, therefore we find it necessary to present the correct locality name],
<specimenCount type="female">1 ♀</specimenCount>
(
<accessionNumber httpUri="http://www.ncbi.nlm.nih.gov/protein/SMF65691">SMF65691</accessionNumber>
),
<collectingDate value="1983-07-10">10.VII.1983</collectingDate>
, leg.
<collectorName>K. Groh</collectorName>
</materialsCitation>
;
<materialsCitation collectingDate="2017-04-21" collectorName="L. Crespo, I. Silva" latitude="33.09447" location="Terra-Cha" longLatPrecision="1" longitude="-16.29839" municipality="Pico Branco" specimenCount="3" specimenCount-juvenile="2" specimenCount-male="1">
<normalizedToken originalValue="Terra-Chã">
<location LSID="urn:lsid:plazi:treatment:22F537099C4C5ECAB94F40A21CF3195E:A00BC0268349EBC039DEB2840B5E7646" latitude="33.09447" longLatPrecision="1" longitude="-16.29839" municipality="Pico Branco" name="Terra-Cha">Terra-Cha</location>
</normalizedToken>
(
<collectingMunicipality>Pico Branco</collectingMunicipality>
),
<geoCoordinate degrees="33.09447" direction="north" orientation="latitude" precision="1" value="33.09447">33.09447°N</geoCoordinate>
,
<geoCoordinate degrees="16.29839" direction="west" orientation="longitude" precision="1" value="-16.29839">16.29839°W</geoCoordinate>
,
<specimenCount type="male">1 ♂</specimenCount>
(CRBALC0323: LC236) and
<specimenCount type="juvenile">2 juveniles</specimenCount>
(CRBALC0324: LC327, CRBALC0396: LC283),
<collectingDate value="2017-04-21">21.IV.2017</collectingDate>
, hand collecting, leg.
<collectorName>L. Crespo</collectorName>
&amp;
<collectorName>I. Silva</collectorName>
</materialsCitation>
,
<specimenCount type="juvenile">4 juveniles</specimenCount>
(CRBALC0627, CRBALC0628, CRBALC0630, CRBALC0700),
<date value="2018-04-10">10.IV.2018</date>
, hand collecting, leg. L. Crespo &amp; A. Bellvert. Unknown location •
<specimenCount type="female">1 ♀</specimenCount>
(NHRS-JUST-000001115),
<specimenCount type="male">1 ♂</specimenCount>
(MMUE G7508.48),
<date value="1981-08-28">28.VIII.1981</date>
, under stone, leg. J. Murphy,
<materialsCitation accessionNumber="SMF34577" collectingDate="2022-01-01" collectingDateMax="2022-12-31" collectingDateMin="2022-01-01" collectorName="G. Schmidt" specimenCount="4" specimenCount-female="1" specimenCount-juvenile="2" specimenCount-male="1">
<specimenCount type="male">1 ♂</specimenCount>
<specimenCount type="female">1 ♀</specimenCount>
and
<specimenCount type="juvenile">2 juveniles</specimenCount>
(
<accessionNumber httpUri="http://www.ncbi.nlm.nih.gov/protein/SMF34577">SMF34577</accessionNumber>
), 1983, leg.
<collectorName>G. Schmidt</collectorName>
</materialsCitation>
,
<materialsCitation accessionNumber="SMF65690" collectingDate="2022-01-01" collectingDateMax="2022-12-31" collectingDateMin="2022-01-01" collectorName="I. Silva" specimenCount="2" specimenCount-female="1" specimenCount-male="1">
<specimenCount type="male">1 ♂</specimenCount>
and
<specimenCount type="female">1 ♀</specimenCount>
(
<accessionNumber httpUri="http://www.ncbi.nlm.nih.gov/protein/SMF65690">SMF65690</accessionNumber>
), hand collecting, leg.
<collectorName>I. Silva</collectorName>
</materialsCitation>
,
<specimenCount type="female">1 ♀</specimenCount>
(NHM 1892.7.9.12.17), leg. W.R.O. Grant.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="84" type="diagnosis">
<paragraph pageId="0" pageNumber="84">Diagnosis.</paragraph>
<paragraph pageId="0" pageNumber="84">
<taxonomicName baseAuthorityName="Kulczynski" baseAuthorityYear="1899" class="Arachnida" family="Lycosidae" genus="Hogna" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hogna insularum" order="Araneae" pageId="0" pageNumber="84" phylum="Arthropoda" rank="species" species="insularum">
<emphasis italics="true" pageId="0" pageNumber="84">Hogna insularum</emphasis>
</taxonomicName>
can be diagnosed from all other Madeiran
<taxonomicName authorityName="Simon" authorityYear="1885" class="Arachnida" family="Lycosidae" genus="Hogna" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hogna" order="Araneae" pageId="0" pageNumber="84" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="84">Hogna</emphasis>
</taxonomicName>
by a combination of the following characters: the small to medium size (prosoma length &lt;10 mm), the aspect of its legs, brown, with black patches (Fig.
<figureCitation captionStart="Figure 27" captionStartId="F27" captionText="Figure 27. Plate with photographs of the lateral view of the leg I for the complex of H. maderiana, H. insularum and their intermediate forms." figureDoi="10.3897/zookeys.1086.68015.figure27" httpUri="https://binary.pensoft.net/fig/647007" pageId="0" pageNumber="84">27C</figureCitation>
),
<normalizedToken originalValue="males">male's</normalizedToken>
embolus thin, with smoothly curved tip (Fig.
<figureCitation captionStart="Figure 15" captionStartId="F15" captionText="Figure 15. Hogna insularum, male pedipalps A male from Porto Santo (CRBALC 0310), left pedipalp, ventral B male from Deserta Grande (CRBALC 0305), left pedipalp, ventral C male from Bugio (CRBALC 0015), left pedipalp, ventral D detail of the median apophysis of male from Deserta Grande (CRBALC 0305), anteroventral E SEM image, right male pedipalp, male from Porto Santo (CRBALC 0310), ventral F idem, retroventral. Abbreviations, male pedipalp: AT - anterior point, C - cymbium, E - embolus, P - palea, R - ridge, T - tegulum, TA - terminal apophysis, TgA - tegular apophysis, VS - ventral spur. Scale bars: 0.5 mm (A, B, C); 0.2 mm (D)." figureDoi="10.3897/zookeys.1086.68015.figure15" httpUri="https://binary.pensoft.net/fig/646995" pageId="0" pageNumber="84">15</figureCitation>
), and female epigyne median septum roughly half as wide (at posterior transverse part) as long (Fig.
<figureCitation captionStart="Figure 16" captionStartId="F16" captionText="Figure 16. Hogna insularum, female genitalia A, B female from Bugio (CRBALC 0301): A epigynum, ventral B vulva, dorsal C, D female from Deserta Grande (CRBALC 0308): C epigynum, ventral D vulva, dorsal E, F female from Madeira (CRBALC 0597): E epigynum, ventral F vulva, dorsal G, H female from Porto Santo (CRBALC 0300): G epigynum, ventral H vulva, dorsal. Abbreviations, female genitalia: D - diverticulum, H - epigynal hoods, MS - median septum, S - spermatheca. Scale bars: 0.2 mm." figureDoi="10.3897/zookeys.1086.68015.figure16" httpUri="https://binary.pensoft.net/fig/646996" pageId="0" pageNumber="84">16A, C, E, G</figureCitation>
). It is most similar to
<taxonomicName class="Arachnida" kingdom="Animalia" lsidName="H. heeri" order="Araneae" pageId="0" pageNumber="84" phylum="Arthropoda" rank="species" species="heeri">
<emphasis italics="true" pageId="0" pageNumber="84">H. heeri</emphasis>
</taxonomicName>
and
<taxonomicName class="Arachnida" kingdom="Animalia" lsidName="H. isambertoi" order="Araneae" pageId="0" pageNumber="84" phylum="Arthropoda" rank="species" species="isambertoi">
<emphasis italics="true" pageId="0" pageNumber="84">H. isambertoi</emphasis>
</taxonomicName>
sp. nov., from which it cannot be somatically differentiated.
</paragraph>
<caption doi="10.3897/zookeys.1086.68015.figure15" httpUri="https://binary.pensoft.net/fig/646995" pageId="0" pageNumber="84" start="Figure 15" startId="F15">
<paragraph pageId="0" pageNumber="84">
<emphasis bold="true" pageId="0" pageNumber="84">Figure 15.</emphasis>
<taxonomicName baseAuthorityName="Kulczynski" baseAuthorityYear="1899" class="Arachnida" family="Lycosidae" genus="Hogna" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hogna insularum" order="Araneae" pageId="0" pageNumber="84" phylum="Arthropoda" rank="species" species="insularum">
<emphasis italics="true" pageId="0" pageNumber="84">Hogna insularum</emphasis>
</taxonomicName>
, male pedipalps
<emphasis bold="true" pageId="0" pageNumber="84">A</emphasis>
male from Porto Santo (CRBALC0310), left pedipalp, ventral
<emphasis bold="true" pageId="0" pageNumber="84">B</emphasis>
male from Deserta Grande (CRBALC0305), left pedipalp, ventral
<emphasis bold="true" pageId="0" pageNumber="84">C</emphasis>
male from Bugio (CRBALC0015), left pedipalp, ventral
<emphasis bold="true" pageId="0" pageNumber="84">D</emphasis>
detail of the median apophysis of male from Deserta Grande (CRBALC0305), anteroventral
<emphasis bold="true" pageId="0" pageNumber="84">E</emphasis>
SEM image, right male pedipalp, male from Porto Santo (CRBALC0310), ventral
<emphasis bold="true" pageId="0" pageNumber="84">F</emphasis>
idem, retroventral. Abbreviations, male pedipalp: AT - anterior point, C - cymbium, E - embolus, P - palea, R - ridge, T - tegulum, TA - terminal apophysis, TgA - tegular apophysis, VS - ventral spur. Scale bars: 0.5 mm (
<emphasis bold="true" pageId="0" pageNumber="84">A, B, C</emphasis>
); 0.2 mm (
<emphasis bold="true" pageId="0" pageNumber="84">D</emphasis>
).
</paragraph>
</caption>
<caption doi="10.3897/zookeys.1086.68015.figure16" httpUri="https://binary.pensoft.net/fig/646996" pageId="0" pageNumber="84" start="Figure 16" startId="F16">
<paragraph pageId="0" pageNumber="84">
<emphasis bold="true" pageId="0" pageNumber="84">Figure 16.</emphasis>
<taxonomicName baseAuthorityName="Kulczynski" baseAuthorityYear="1899" class="Arachnida" family="Lycosidae" genus="Hogna" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hogna insularum" order="Araneae" pageId="0" pageNumber="84" phylum="Arthropoda" rank="species" species="insularum">
<emphasis italics="true" pageId="0" pageNumber="84">Hogna insularum</emphasis>
</taxonomicName>
, female genitalia
<emphasis bold="true" pageId="0" pageNumber="84">A, B</emphasis>
female from Bugio (CRBALC0301):
<emphasis bold="true" pageId="0" pageNumber="84">A</emphasis>
epigynum, ventral
<emphasis bold="true" pageId="0" pageNumber="84">B</emphasis>
vulva, dorsal
<emphasis bold="true" pageId="0" pageNumber="84">C, D</emphasis>
female from Deserta Grande (CRBALC0308):
<emphasis bold="true" pageId="0" pageNumber="84">C</emphasis>
epigynum, ventral
<emphasis bold="true" pageId="0" pageNumber="84">D</emphasis>
vulva, dorsal
<emphasis bold="true" pageId="0" pageNumber="84">E, F</emphasis>
female from Madeira (CRBALC0597):
<emphasis bold="true" pageId="0" pageNumber="84">E</emphasis>
epigynum, ventral
<emphasis bold="true" pageId="0" pageNumber="84">F</emphasis>
vulva, dorsal
<emphasis bold="true" pageId="0" pageNumber="84">G, H</emphasis>
female from Porto Santo (CRBALC0300):
<emphasis bold="true" pageId="0" pageNumber="84">G</emphasis>
epigynum, ventral
<emphasis bold="true" pageId="0" pageNumber="84">H</emphasis>
vulva, dorsal. Abbreviations, female genitalia: D - diverticulum, H - epigynal hoods, MS - median septum, S - spermatheca. Scale bars: 0.2 mm.
</paragraph>
</caption>
</subSubSection>
<subSubSection pageId="0" pageNumber="84" type="redescription">
<paragraph pageId="0" pageNumber="84">Redescription.</paragraph>
<paragraph pageId="0" pageNumber="84">
<emphasis bold="true" pageId="0" pageNumber="84">Male (CRBALC0310)</emphasis>
: (Fig.
<figureCitation captionStart="Figure 15" captionStartId="F15" captionText="Figure 15. Hogna insularum, male pedipalps A male from Porto Santo (CRBALC 0310), left pedipalp, ventral B male from Deserta Grande (CRBALC 0305), left pedipalp, ventral C male from Bugio (CRBALC 0015), left pedipalp, ventral D detail of the median apophysis of male from Deserta Grande (CRBALC 0305), anteroventral E SEM image, right male pedipalp, male from Porto Santo (CRBALC 0310), ventral F idem, retroventral. Abbreviations, male pedipalp: AT - anterior point, C - cymbium, E - embolus, P - palea, R - ridge, T - tegulum, TA - terminal apophysis, TgA - tegular apophysis, VS - ventral spur. Scale bars: 0.5 mm (A, B, C); 0.2 mm (D)." figureDoi="10.3897/zookeys.1086.68015.figure15" httpUri="https://binary.pensoft.net/fig/646995" pageId="0" pageNumber="84">15A, E, F</figureCitation>
). Total length: 7.8; carapace: 4.6 long, 3.3 wide.
</paragraph>
<paragraph pageId="0" pageNumber="84">
<emphasis bold="true" italics="true" pageId="0" pageNumber="84">Colour</emphasis>
: carapace greyish brown, covered with short black setae, with a median cream longitudinal band, anteriorly broadened, covered with short white setae, with suffused greyish brown patches; two yellow marginal bands, with roughly round grey patches, covered with short white setae; four black striae well visible on each flank. Chelicerae brownish orange, with blackish patches, covered in black and white setae. Gnathocoxae greyish yellow, labium overall blackish, with anterior margin greyish yellow; sternum yellow, with a V-shaped grey patch and suffused patches at lateral borders. Legs pale yellow to orange from femora to tibia, with irregular grey suffused patches, metatarsi and tarsi brown. Pedipalps pale yellow except tarsus, brown. Abdomen with a pair of anterolateral black patches, extending laterally into grey flanks, mottled with yellowish patches covered with white setae; a median dark lanceolate patch is bordered by two yellowish longitudinal bands interconnected in anterior half, posteriorly by means of dark chevrons; venter yellowish, with a median dark grey longitudinal band, bordered by small yellowish and grey patches.
</paragraph>
<paragraph pageId="0" pageNumber="84">
<emphasis bold="true" italics="true" pageId="0" pageNumber="84">Eyes</emphasis>
: MOQ: MW = 0.8 PW, MW = 1.1 LMP, MW = 1.2 AW; Cl = 0.3 DAME. Anterior eye row slightly procurved.
</paragraph>
<paragraph pageId="0" pageNumber="84">
<emphasis bold="true" italics="true" pageId="0" pageNumber="84">Legs</emphasis>
: Measurements: Leg I: 13.6, TiI: 3.1; Leg IV: 14.9, TiIV: 3.1; TiIL/D: 5.5. Spination of Leg I: FeI: d1.1.0, p0.0.1-2; TiI: p1s.0.1s, r1s.0.1s, v2l.2l.2s; MtI: p0.1.1, r0.0.1, v2l.2l.1s. MtI with sparse scopulae in basal half and dense scopulae on distal half.
</paragraph>
<paragraph pageId="0" pageNumber="84">
<emphasis bold="true" italics="true" pageId="0" pageNumber="84">Pedipalp</emphasis>
: cymbium with five dark, stout, macrosetae at tip, Fe with two dorsal and an apical row of four spines, Pa with one prolateral spine, Ti with one dorsal and one prolateral spines. Tegular apophysis with ventral spur short, blunt, with a straight ridge leading to a wide apical point (Fig.
<figureCitation captionStart="Figure 15" captionStartId="F15" captionText="Figure 15. Hogna insularum, male pedipalps A male from Porto Santo (CRBALC 0310), left pedipalp, ventral B male from Deserta Grande (CRBALC 0305), left pedipalp, ventral C male from Bugio (CRBALC 0015), left pedipalp, ventral D detail of the median apophysis of male from Deserta Grande (CRBALC 0305), anteroventral E SEM image, right male pedipalp, male from Porto Santo (CRBALC 0310), ventral F idem, retroventral. Abbreviations, male pedipalp: AT - anterior point, C - cymbium, E - embolus, P - palea, R - ridge, T - tegulum, TA - terminal apophysis, TgA - tegular apophysis, VS - ventral spur. Scale bars: 0.5 mm (A, B, C); 0.2 mm (D)." figureDoi="10.3897/zookeys.1086.68015.figure15" httpUri="https://binary.pensoft.net/fig/646995" pageId="0" pageNumber="84">15D</figureCitation>
); terminal apophysis blade-shaped with sharp end (Fig.
<figureCitation captionStart="Figure 15" captionStartId="F15" captionText="Figure 15. Hogna insularum, male pedipalps A male from Porto Santo (CRBALC 0310), left pedipalp, ventral B male from Deserta Grande (CRBALC 0305), left pedipalp, ventral C male from Bugio (CRBALC 0015), left pedipalp, ventral D detail of the median apophysis of male from Deserta Grande (CRBALC 0305), anteroventral E SEM image, right male pedipalp, male from Porto Santo (CRBALC 0310), ventral F idem, retroventral. Abbreviations, male pedipalp: AT - anterior point, C - cymbium, E - embolus, P - palea, R - ridge, T - tegulum, TA - terminal apophysis, TgA - tegular apophysis, VS - ventral spur. Scale bars: 0.5 mm (A, B, C); 0.2 mm (D)." figureDoi="10.3897/zookeys.1086.68015.figure15" httpUri="https://binary.pensoft.net/fig/646995" pageId="0" pageNumber="84">15A-C</figureCitation>
); embolus long and thin, with tip smoothly curved anteriorly (Fig.
<figureCitation captionStart="Figure 15" captionStartId="F15" captionText="Figure 15. Hogna insularum, male pedipalps A male from Porto Santo (CRBALC 0310), left pedipalp, ventral B male from Deserta Grande (CRBALC 0305), left pedipalp, ventral C male from Bugio (CRBALC 0015), left pedipalp, ventral D detail of the median apophysis of male from Deserta Grande (CRBALC 0305), anteroventral E SEM image, right male pedipalp, male from Porto Santo (CRBALC 0310), ventral F idem, retroventral. Abbreviations, male pedipalp: AT - anterior point, C - cymbium, E - embolus, P - palea, R - ridge, T - tegulum, TA - terminal apophysis, TgA - tegular apophysis, VS - ventral spur. Scale bars: 0.5 mm (A, B, C); 0.2 mm (D)." figureDoi="10.3897/zookeys.1086.68015.figure15" httpUri="https://binary.pensoft.net/fig/646995" pageId="0" pageNumber="84">15A-C</figureCitation>
); palea large (Fig.
<figureCitation captionStart="Figure 15" captionStartId="F15" captionText="Figure 15. Hogna insularum, male pedipalps A male from Porto Santo (CRBALC 0310), left pedipalp, ventral B male from Deserta Grande (CRBALC 0305), left pedipalp, ventral C male from Bugio (CRBALC 0015), left pedipalp, ventral D detail of the median apophysis of male from Deserta Grande (CRBALC 0305), anteroventral E SEM image, right male pedipalp, male from Porto Santo (CRBALC 0310), ventral F idem, retroventral. Abbreviations, male pedipalp: AT - anterior point, C - cymbium, E - embolus, P - palea, R - ridge, T - tegulum, TA - terminal apophysis, TgA - tegular apophysis, VS - ventral spur. Scale bars: 0.5 mm (A, B, C); 0.2 mm (D)." figureDoi="10.3897/zookeys.1086.68015.figure15" httpUri="https://binary.pensoft.net/fig/646995" pageId="0" pageNumber="84">15A-C</figureCitation>
).
</paragraph>
<paragraph pageId="0" pageNumber="84">
<emphasis bold="true" pageId="0" pageNumber="84">Female</emphasis>
(
<emphasis italics="true" pageId="0" pageNumber="84">CRBALC0308</emphasis>
): (Fig.
<figureCitation captionStart="Figure 16" captionStartId="F16" captionText="Figure 16. Hogna insularum, female genitalia A, B female from Bugio (CRBALC 0301): A epigynum, ventral B vulva, dorsal C, D female from Deserta Grande (CRBALC 0308): C epigynum, ventral D vulva, dorsal E, F female from Madeira (CRBALC 0597): E epigynum, ventral F vulva, dorsal G, H female from Porto Santo (CRBALC 0300): G epigynum, ventral H vulva, dorsal. Abbreviations, female genitalia: D - diverticulum, H - epigynal hoods, MS - median septum, S - spermatheca. Scale bars: 0.2 mm." figureDoi="10.3897/zookeys.1086.68015.figure16" httpUri="https://binary.pensoft.net/fig/646996" pageId="0" pageNumber="84">16C-D</figureCitation>
). Total length 12.8; carapace: 5.4 long, 4.1 wide.
</paragraph>
<paragraph pageId="0" pageNumber="84">
<emphasis bold="true" italics="true" pageId="0" pageNumber="84">Colour</emphasis>
: overall as in male, but darker in legs, chelicera and prosoma. Sternum with a faint V-shaped grey patch. Abdomen is lighter, with central chevrons and ventral longitudinal dark band faded.
</paragraph>
<paragraph pageId="0" pageNumber="84">
<emphasis bold="true" italics="true" pageId="0" pageNumber="84">Eyes</emphasis>
: MOQ: MW = 0.8 PW, MW = 1.2 LMP, MW = 1.2 AW; Cl = 0.6 DAME. Anterior eye row slightly procurved.
</paragraph>
<paragraph pageId="0" pageNumber="84">
<emphasis bold="true" italics="true" pageId="0" pageNumber="84">Legs</emphasis>
: Measurements: Leg I: 13.8, TiI: 3.1; Leg IV: 16.0, TiIV: 3.3; TiIL/D: 3.7. Spination of Leg I: FeI: d1.1.0, p0.0.2; TiI: p0.1s.0, v2l.2l.2s; MtI: p0.0.1, r0.0.1, v2l.2l.1s. MtI with sparse scopulae in basal half and dense scopulae on distal half.
</paragraph>
<paragraph pageId="0" pageNumber="84">
<emphasis bold="true" italics="true" pageId="0" pageNumber="84">Epigyne</emphasis>
: anterior pockets almost touching, short, with lateral borders parallel (Fig.
<figureCitation captionStart="Figure 16" captionStartId="F16" captionText="Figure 16. Hogna insularum, female genitalia A, B female from Bugio (CRBALC 0301): A epigynum, ventral B vulva, dorsal C, D female from Deserta Grande (CRBALC 0308): C epigynum, ventral D vulva, dorsal E, F female from Madeira (CRBALC 0597): E epigynum, ventral F vulva, dorsal G, H female from Porto Santo (CRBALC 0300): G epigynum, ventral H vulva, dorsal. Abbreviations, female genitalia: D - diverticulum, H - epigynal hoods, MS - median septum, S - spermatheca. Scale bars: 0.2 mm." figureDoi="10.3897/zookeys.1086.68015.figure16" httpUri="https://binary.pensoft.net/fig/646996" pageId="0" pageNumber="84">16C</figureCitation>
); anterior pocket cavities shallow; median septum with narrow posterior transverse part (Fig.
<figureCitation captionStart="Figure 16" captionStartId="F16" captionText="Figure 16. Hogna insularum, female genitalia A, B female from Bugio (CRBALC 0301): A epigynum, ventral B vulva, dorsal C, D female from Deserta Grande (CRBALC 0308): C epigynum, ventral D vulva, dorsal E, F female from Madeira (CRBALC 0597): E epigynum, ventral F vulva, dorsal G, H female from Porto Santo (CRBALC 0300): G epigynum, ventral H vulva, dorsal. Abbreviations, female genitalia: D - diverticulum, H - epigynal hoods, MS - median septum, S - spermatheca. Scale bars: 0.2 mm." figureDoi="10.3897/zookeys.1086.68015.figure16" httpUri="https://binary.pensoft.net/fig/646996" pageId="0" pageNumber="84">16C</figureCitation>
); spermathecae oval or piriform (Fig.
<figureCitation captionStart="Figure 16" captionStartId="F16" captionText="Figure 16. Hogna insularum, female genitalia A, B female from Bugio (CRBALC 0301): A epigynum, ventral B vulva, dorsal C, D female from Deserta Grande (CRBALC 0308): C epigynum, ventral D vulva, dorsal E, F female from Madeira (CRBALC 0597): E epigynum, ventral F vulva, dorsal G, H female from Porto Santo (CRBALC 0300): G epigynum, ventral H vulva, dorsal. Abbreviations, female genitalia: D - diverticulum, H - epigynal hoods, MS - median septum, S - spermatheca. Scale bars: 0.2 mm." figureDoi="10.3897/zookeys.1086.68015.figure16" httpUri="https://binary.pensoft.net/fig/646996" pageId="0" pageNumber="84">16D</figureCitation>
); copulatory ducts with small, stout diverticulum ventrally (Fig.
<figureCitation captionStart="Figure 16" captionStartId="F16" captionText="Figure 16. Hogna insularum, female genitalia A, B female from Bugio (CRBALC 0301): A epigynum, ventral B vulva, dorsal C, D female from Deserta Grande (CRBALC 0308): C epigynum, ventral D vulva, dorsal E, F female from Madeira (CRBALC 0597): E epigynum, ventral F vulva, dorsal G, H female from Porto Santo (CRBALC 0300): G epigynum, ventral H vulva, dorsal. Abbreviations, female genitalia: D - diverticulum, H - epigynal hoods, MS - median septum, S - spermatheca. Scale bars: 0.2 mm." figureDoi="10.3897/zookeys.1086.68015.figure16" httpUri="https://binary.pensoft.net/fig/646996" pageId="0" pageNumber="84">16D</figureCitation>
); fertilisation ducts emerging at the base of copulatory duct (Fig.
<figureCitation captionStart="Figure 16" captionStartId="F16" captionText="Figure 16. Hogna insularum, female genitalia A, B female from Bugio (CRBALC 0301): A epigynum, ventral B vulva, dorsal C, D female from Deserta Grande (CRBALC 0308): C epigynum, ventral D vulva, dorsal E, F female from Madeira (CRBALC 0597): E epigynum, ventral F vulva, dorsal G, H female from Porto Santo (CRBALC 0300): G epigynum, ventral H vulva, dorsal. Abbreviations, female genitalia: D - diverticulum, H - epigynal hoods, MS - median septum, S - spermatheca. Scale bars: 0.2 mm." figureDoi="10.3897/zookeys.1086.68015.figure16" httpUri="https://binary.pensoft.net/fig/646996" pageId="0" pageNumber="84">16D</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="84" type="intraspecific variation">
<paragraph pageId="0" pageNumber="84">Intraspecific variation.</paragraph>
<paragraph pageId="0" pageNumber="84">
Carapace length, males: 4.6-6.4, females: 5.1-7.4. Length of cymbium tip of male pedipalp can vary from shorter to longer than the bulbus. In the single available adult female from Madeira, the anterior pockets of the epigyne show slightly divergent lateral borders (posteriorly) (Fig.
<figureCitation captionStart="Figure 16" captionStartId="F16" captionText="Figure 16. Hogna insularum, female genitalia A, B female from Bugio (CRBALC 0301): A epigynum, ventral B vulva, dorsal C, D female from Deserta Grande (CRBALC 0308): C epigynum, ventral D vulva, dorsal E, F female from Madeira (CRBALC 0597): E epigynum, ventral F vulva, dorsal G, H female from Porto Santo (CRBALC 0300): G epigynum, ventral H vulva, dorsal. Abbreviations, female genitalia: D - diverticulum, H - epigynal hoods, MS - median septum, S - spermatheca. Scale bars: 0.2 mm." figureDoi="10.3897/zookeys.1086.68015.figure16" httpUri="https://binary.pensoft.net/fig/646996" pageId="0" pageNumber="84">16E</figureCitation>
), while specimens from the remaining islands show parallel lateral borders.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="84" type="distribution">
<paragraph pageId="0" pageNumber="84">Distribution.</paragraph>
<paragraph pageId="0" pageNumber="84">
This species is known from many locations on all islands of the archipelago except Madeira island, where it is only present at the southeast coastal region (Fig.
<figureCitation captionStart="Figure 17" captionStartId="F17" captionText="Figure 17. Distribution of H. insularum. Circles: present records; black triangles: revised records from literature; white triangles: unconfirmed records from literature." figureDoi="10.3897/zookeys.1086.68015.figure17" httpUri="https://binary.pensoft.net/fig/646997" pageId="0" pageNumber="84">17</figureCitation>
).
</paragraph>
<caption doi="10.3897/zookeys.1086.68015.figure17" httpUri="https://binary.pensoft.net/fig/646997" pageId="0" pageNumber="84" start="Figure 17" startId="F17">
<paragraph pageId="0" pageNumber="84">
<emphasis bold="true" pageId="0" pageNumber="84">Figure 17.</emphasis>
Distribution of
<taxonomicName class="Arachnida" kingdom="Animalia" lsidName="H. insularum" order="Araneae" pageId="0" pageNumber="84" phylum="Arthropoda" rank="species" species="insularum">
<emphasis italics="true" pageId="0" pageNumber="84">H. insularum</emphasis>
</taxonomicName>
. Circles: present records; black triangles: revised records from literature; white triangles: unconfirmed records from literature.
</paragraph>
</caption>
</subSubSection>
<subSubSection pageId="0" pageNumber="84" type="ecology">
<paragraph pageId="0" pageNumber="84">Ecology.</paragraph>
<paragraph pageId="0" pageNumber="84">
<taxonomicName baseAuthorityName="Kulczynski" baseAuthorityYear="1899" class="Arachnida" family="Lycosidae" genus="Hogna" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hogna insularum" order="Araneae" pageId="0" pageNumber="84" phylum="Arthropoda" rank="species" species="insularum">
<emphasis italics="true" pageId="0" pageNumber="84">Hogna insularum</emphasis>
</taxonomicName>
occurs in a wide variety of habitats, from grasslands,
<taxonomicName class="Magnoliopsida" family="Ericaceae" genus="Erica" kingdom="Plantae" lsidName="Erica" order="Ericales" pageId="0" pageNumber="84" phylum="Tracheophyta" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="84">Erica</emphasis>
</taxonomicName>
shrubland, to secondary forests (in the latter two cases, only in Porto Santo).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="84" type="conservation status">
<paragraph pageId="0" pageNumber="84">Conservation status.</paragraph>
<paragraph pageId="0" pageNumber="84">
<taxonomicName baseAuthorityName="Kulczynski" baseAuthorityYear="1899" class="Arachnida" family="Lycosidae" genus="Hogna" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hogna insularum" order="Araneae" pageId="0" pageNumber="84" phylum="Arthropoda" rank="species" species="insularum">
<emphasis italics="true" pageId="0" pageNumber="84">Hogna insularum</emphasis>
</taxonomicName>
was assessed according to the IUCN Red List criteria, with the status of Least Concern (
<bibRefCitation DOI="https://doi.org/10.2305/IUCN.UK.2018-2.RLTS.T58048609A58061012.en" author="Cardoso, P" journalOrPublisher="Molecular Ecology" pageId="0" pageNumber="84" refId="B17" refString="Cardoso, P, Crespo, LC, Silva, I, Borges, P, Boieiro, M, 2018c. Hogna insularum. The IUCN Red List of Threatened Species 2018. https://doi.org/10.2305/IUCN.UK.2018-2.RLTS.T58048609A58061012.en" title="Hogna insularum. The IUCN Red List of Threatened Species 2018." url="https://doi.org/10.2305/IUCN.UK.2018-2.RLTS.T58048609A58061012.en" year="2018 c">Cardoso et al. 2018c</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="84" type="comments">
<paragraph pageId="0" pageNumber="84">Comments.</paragraph>
<paragraph pageId="0" pageNumber="84">
<taxonomicName baseAuthorityName="Kulczynski" baseAuthorityYear="1899" class="Arachnida" family="Lycosidae" genus="Hogna" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hogna insularum" order="Araneae" pageId="0" pageNumber="84" phylum="Arthropoda" rank="species" species="insularum">
<emphasis italics="true" pageId="0" pageNumber="84">Hogna insularum</emphasis>
</taxonomicName>
displays remarkable intraspecific variation. In males, both the length of the cymbium tip and the position of the terminal apophysis relative to the embolus are variable (Fig.
<figureCitation captionStart="Figure 15" captionStartId="F15" captionText="Figure 15. Hogna insularum, male pedipalps A male from Porto Santo (CRBALC 0310), left pedipalp, ventral B male from Deserta Grande (CRBALC 0305), left pedipalp, ventral C male from Bugio (CRBALC 0015), left pedipalp, ventral D detail of the median apophysis of male from Deserta Grande (CRBALC 0305), anteroventral E SEM image, right male pedipalp, male from Porto Santo (CRBALC 0310), ventral F idem, retroventral. Abbreviations, male pedipalp: AT - anterior point, C - cymbium, E - embolus, P - palea, R - ridge, T - tegulum, TA - terminal apophysis, TgA - tegular apophysis, VS - ventral spur. Scale bars: 0.5 mm (A, B, C); 0.2 mm (D)." figureDoi="10.3897/zookeys.1086.68015.figure15" httpUri="https://binary.pensoft.net/fig/646995" pageId="0" pageNumber="84">15</figureCitation>
). In females, the epigyne usually presents anterior pockets with anteriorly parallel lateral borders, but a specimen from Madeira shows a posteriorly divergent lateral border. At the same time, the shape of spermathecae seem to vary from ovoid (Fig.
<figureCitation captionStart="Figure 16" captionStartId="F16" captionText="Figure 16. Hogna insularum, female genitalia A, B female from Bugio (CRBALC 0301): A epigynum, ventral B vulva, dorsal C, D female from Deserta Grande (CRBALC 0308): C epigynum, ventral D vulva, dorsal E, F female from Madeira (CRBALC 0597): E epigynum, ventral F vulva, dorsal G, H female from Porto Santo (CRBALC 0300): G epigynum, ventral H vulva, dorsal. Abbreviations, female genitalia: D - diverticulum, H - epigynal hoods, MS - median septum, S - spermatheca. Scale bars: 0.2 mm." figureDoi="10.3897/zookeys.1086.68015.figure16" httpUri="https://binary.pensoft.net/fig/646996" pageId="0" pageNumber="84">16A-D</figureCitation>
), to piriform (Fig.
<figureCitation captionStart="Figure 16" captionStartId="F16" captionText="Figure 16. Hogna insularum, female genitalia A, B female from Bugio (CRBALC 0301): A epigynum, ventral B vulva, dorsal C, D female from Deserta Grande (CRBALC 0308): C epigynum, ventral D vulva, dorsal E, F female from Madeira (CRBALC 0597): E epigynum, ventral F vulva, dorsal G, H female from Porto Santo (CRBALC 0300): G epigynum, ventral H vulva, dorsal. Abbreviations, female genitalia: D - diverticulum, H - epigynal hoods, MS - median septum, S - spermatheca. Scale bars: 0.2 mm." figureDoi="10.3897/zookeys.1086.68015.figure16" httpUri="https://binary.pensoft.net/fig/646996" pageId="0" pageNumber="84">16E, F</figureCitation>
), to rounded (Fig.
<figureCitation captionStart="Figure 16" captionStartId="F16" captionText="Figure 16. Hogna insularum, female genitalia A, B female from Bugio (CRBALC 0301): A epigynum, ventral B vulva, dorsal C, D female from Deserta Grande (CRBALC 0308): C epigynum, ventral D vulva, dorsal E, F female from Madeira (CRBALC 0597): E epigynum, ventral F vulva, dorsal G, H female from Porto Santo (CRBALC 0300): G epigynum, ventral H vulva, dorsal. Abbreviations, female genitalia: D - diverticulum, H - epigynal hoods, MS - median septum, S - spermatheca. Scale bars: 0.2 mm." figureDoi="10.3897/zookeys.1086.68015.figure16" httpUri="https://binary.pensoft.net/fig/646996" pageId="0" pageNumber="84">16G, H</figureCitation>
).
<bibRefCitation author="Wunderlich, J" journalOrPublisher="Beitraege zur Araneologie" pageId="0" pageNumber="84" pagination="1 - 619" refId="B76" refString="Wunderlich, J, 1992. Die Spinnen-Fauna der Makaronesischen Inseln: Taxonomie, Oekologie, Biogeographie und Evolution. Beitraege zur Araneologie 1: 1 - 619" title="Die Spinnen-Fauna der Makaronesischen Inseln: Taxonomie, Oekologie, Biogeographie und Evolution." volume="1" year="1992">Wunderlich (1992)</bibRefCitation>
described
<taxonomicName class="Arachnida" kingdom="Animalia" lsidName="H. biscoitoi" order="Araneae" pageId="0" pageNumber="84" phylum="Arthropoda" rank="species" species="biscoitoi">
<emphasis italics="true" pageId="0" pageNumber="84">H. biscoitoi</emphasis>
</taxonomicName>
based on specimens from Porto Santo. To differentiate it from
<taxonomicName class="Arachnida" kingdom="Animalia" lsidName="H. insularum" order="Araneae" pageId="0" pageNumber="84" phylum="Arthropoda" rank="species" species="insularum">
<emphasis italics="true" pageId="0" pageNumber="84">H. insularum</emphasis>
</taxonomicName>
he stated that in males &quot;the sickle-shaped apophysis points more to the tip of the cymbium&quot; while in the former species the same structure
<normalizedToken originalValue="“(…">&quot;(...</normalizedToken>
) is directed more retrolaterally&quot;. For females, although a diagnostic description was provided, the identification key directed to the same image when referring to the epigyne of both
<taxonomicName class="Arachnida" kingdom="Animalia" lsidName="H. insularum" order="Araneae" pageId="0" pageNumber="84" phylum="Arthropoda" rank="species" species="insularum">
<emphasis italics="true" pageId="0" pageNumber="84">H. insularum</emphasis>
</taxonomicName>
and
<taxonomicName class="Arachnida" kingdom="Animalia" lsidName="H. biscoitoi" order="Araneae" pageId="0" pageNumber="84" phylum="Arthropoda" rank="species" species="biscoitoi">
<emphasis italics="true" pageId="0" pageNumber="84">H. biscoitoi</emphasis>
</taxonomicName>
. We collected an array of specimens from different localities in Porto Santo (from Pico Ana Ferreira to Pico Branco) and surrounding islets. We did observe male pedipalps with different degrees of inclination of the terminal apophysis and with cymbium tip shorter than the length of the copulatory bulbus (Fig.
<figureCitation captionStart="Figure 15" captionStartId="F15" captionText="Figure 15. Hogna insularum, male pedipalps A male from Porto Santo (CRBALC 0310), left pedipalp, ventral B male from Deserta Grande (CRBALC 0305), left pedipalp, ventral C male from Bugio (CRBALC 0015), left pedipalp, ventral D detail of the median apophysis of male from Deserta Grande (CRBALC 0305), anteroventral E SEM image, right male pedipalp, male from Porto Santo (CRBALC 0310), ventral F idem, retroventral. Abbreviations, male pedipalp: AT - anterior point, C - cymbium, E - embolus, P - palea, R - ridge, T - tegulum, TA - terminal apophysis, TgA - tegular apophysis, VS - ventral spur. Scale bars: 0.5 mm (A, B, C); 0.2 mm (D)." figureDoi="10.3897/zookeys.1086.68015.figure15" httpUri="https://binary.pensoft.net/fig/646995" pageId="0" pageNumber="84">15A, C</figureCitation>
), but both characters were unlinked. We suspect that these traits may be affected by the time from the last moult (e.g., Fig.
<figureCitation captionStart="Figure 15" captionStartId="F15" captionText="Figure 15. Hogna insularum, male pedipalps A male from Porto Santo (CRBALC 0310), left pedipalp, ventral B male from Deserta Grande (CRBALC 0305), left pedipalp, ventral C male from Bugio (CRBALC 0015), left pedipalp, ventral D detail of the median apophysis of male from Deserta Grande (CRBALC 0305), anteroventral E SEM image, right male pedipalp, male from Porto Santo (CRBALC 0310), ventral F idem, retroventral. Abbreviations, male pedipalp: AT - anterior point, C - cymbium, E - embolus, P - palea, R - ridge, T - tegulum, TA - terminal apophysis, TgA - tegular apophysis, VS - ventral spur. Scale bars: 0.5 mm (A, B, C); 0.2 mm (D)." figureDoi="10.3897/zookeys.1086.68015.figure15" httpUri="https://binary.pensoft.net/fig/646995" pageId="0" pageNumber="84">15B</figureCitation>
was most likely a recently moulted individual, given its overall pale coloration). Furthermore, fixation in ethanol might sometimes cause a displacement of sclerites, even if small. Molecular data does not seem to provide any additional evidence regarding the possibility that the specimens from Porto Santo may belong to a different species that those reported form other islands. Unfortunately, we could not examine the type material of
<taxonomicName class="Arachnida" kingdom="Animalia" lsidName="H. biscoitoi" order="Araneae" pageId="0" pageNumber="84" phylum="Arthropoda" rank="species" species="biscoitoi">
<emphasis italics="true" pageId="0" pageNumber="84">H. biscoitoi</emphasis>
</taxonomicName>
stored at the Funchal Municipal Museum, since it does not loan type material for study. Based on the variability in the supposedly diagnostic features and the lack of genetic divergence, we consider
<taxonomicName class="Arachnida" kingdom="Animalia" lsidName="H. biscoitoi" order="Araneae" pageId="0" pageNumber="84" phylum="Arthropoda" rank="species" species="biscoitoi">
<emphasis italics="true" pageId="0" pageNumber="84">H. biscoitoi</emphasis>
</taxonomicName>
as a junior synonym of
<taxonomicName class="Arachnida" kingdom="Animalia" lsidName="H. insularum" order="Araneae" pageId="0" pageNumber="84" phylum="Arthropoda" rank="species" species="insularum">
<emphasis italics="true" pageId="0" pageNumber="84">H. insularum</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
</document>