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<document id="B08C5282FBAB47BE89C2323AE04AACD5" ID-CLB-Dataset="263656" ID-DOI="10.1590/S1984-4689.v39.e21037" ID-GBIF-Dataset="dd7e9cd9-bb4a-4032-9dae-ef3a866c21cf" ID-ISSN="1984-4689" ID-Zenodo-Dep="8233436" ID-ZooBank="6D8A329C-51BB-4C36-89F0-13454C313165" IM.bibliography_approvedBy="juliana" IM.illustrations_approvedBy="juliana" IM.materialsCitations_approvedBy="juliana" IM.metadata_approvedBy="juliana" IM.taxonomicNames_approvedBy="juliana" IM.treatmentCitations_approvedBy="juliana" IM.treatments_approvedBy="juliana" checkinTime="1691525684732" checkinUser="felipe" docAuthor="Pinedo-Escatel, J. Adilson, Dietrich, Christopher H. &amp; Zahniser, James N." docDate="2022" docId="03C387DFFFDF234DFF614F6CFA68FC8C" docLanguage="en" docName="Zoologia.39.e21037.pdf" docOrigin="Zoologia (e 21037) 39" docSource="http://dx.doi.org/10.1590/s1984-4689.v39.e21037" docStyle="DocumentStyle:C0DEAC3FDCE12EEF877F5538032D0DC0.2:Zoologia.2021-.journal_article" docStyleId="C0DEAC3FDCE12EEF877F5538032D0DC0" docStyleName="Zoologia.2021-.journal_article" docStyleVersion="2" docTitle="Andanus Linnavuori 1959" docType="key" docVersion="3" lastPageNumber="3" masterDocId="FFFAFFA7FFDD234FFFFE493FFFF5FF99" masterDocTitle="Andanus tambopixunensis sp. nov., a new species of the remarkable leafhopper genus Andanus (Hemiptera: Cicadellidae: Deltocephalinae), generic redescription and two new synonyms with new placements" masterLastPageNumber="10" masterPageNumber="1" pageNumber="3" updateTime="1691669732718" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
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<mods:title id="9BC453F878042A6369014CD51CD3765E">Andanus tambopixunensis sp. nov., a new species of the remarkable leafhopper genus Andanus (Hemiptera: Cicadellidae: Deltocephalinae), generic redescription and two new synonyms with new placements</mods:title>
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<mods:namePart id="A061AE0A52F966335BDDF5FFAA670821">Dietrich, Christopher H.</mods:namePart>
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Key to species of
<taxonomicName id="4C6A4D4AFFDF234DFEBC4F6DFE64F9FF" ID-CoL="8R3X3" ID-ENA="2768526" authorityName="Linnavuori" authorityYear="1959" box="[322,401,1618,1638]" class="Insecta" family="Cicadellidae" genus="Andanus" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="genus">
<emphasis id="B91EEADBFFDF234DFEBC4F6DFE64F9FF" box="[322,401,1618,1638]" italics="true" pageId="2" pageNumber="3">Andanus</emphasis>
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<paragraph id="8BD536C9FFDF234DFF614F49FCFFF96F" blockId="2.[159,779,1618,1782]" pageId="2" pageNumber="3">
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1. Pygofer with ventral margin serrated (
<figureCitation id="13512A4CFFDF234DFDCC4F49FD6AF910" box="[562,671,1654,1674]" captionStart="Figures 2936" captionStartId="7.[151,218,1702,1722]" captionTargetBox="[222,1430,114,1685]" captionTargetId="figure-51@7.[220,1348,197,1686]" captionTargetPageId="7" captionText="Figures 2936. Male of Andanus sordidus comb. nov.: (29) habitus, dorsal view; (30) habitus, lateral view; (31) Face, anterior view; (32) pygofer and segment X, dorsal view; (33) pygofer, lateral view; (34) subgenital plates and valve, ventral view; (35) aedeagus, connective, and styles, ventral view; (36) aedeagus and connective, lateral view." figureDoi="http://doi.org/10.5281/zenodo.8233452" httpUri="https://zenodo.org/record/8233452/files/figure.png" pageId="2" pageNumber="3">Figs 32, 33</figureCitation>
); aedeagus with pair of sublateroapical flanges with single subdorsoapical spine and single ventromedial spine (
<figureCitation id="13512A4CFFDF234DFD484F93FCF5F926" box="[694,768,1708,1728]" captionStart="Figures 2936" captionStartId="7.[151,218,1702,1722]" captionTargetBox="[222,1430,114,1685]" captionTargetId="figure-51@7.[220,1348,197,1686]" captionTargetPageId="7" captionText="Figures 2936. Male of Andanus sordidus comb. nov.: (29) habitus, dorsal view; (30) habitus, lateral view; (31) Face, anterior view; (32) pygofer and segment X, dorsal view; (33) pygofer, lateral view; (34) subgenital plates and valve, ventral view; (35) aedeagus, connective, and styles, ventral view; (36) aedeagus and connective, lateral view." figureDoi="http://doi.org/10.5281/zenodo.8233452" httpUri="https://zenodo.org/record/8233452/files/figure.png" pageId="2" pageNumber="3">Fig. 36</figureCitation>
); subgenital plates very long, constricted medially, expanded and clublike toward apices (
<figureCitation id="13512A4CFFDF234DFDFB4FDDFDBDF96F" box="[517,584,1762,1782]" captionStart="Figures 2936" captionStartId="7.[151,218,1702,1722]" captionTargetBox="[222,1430,114,1685]" captionTargetId="figure-51@7.[220,1348,197,1686]" captionTargetPageId="7" captionText="Figures 2936. Male of Andanus sordidus comb. nov.: (29) habitus, dorsal view; (30) habitus, lateral view; (31) Face, anterior view; (32) pygofer and segment X, dorsal view; (33) pygofer, lateral view; (34) subgenital plates and valve, ventral view; (35) aedeagus, connective, and styles, ventral view; (36) aedeagus and connective, lateral view." figureDoi="http://doi.org/10.5281/zenodo.8233452" httpUri="https://zenodo.org/record/8233452/files/figure.png" pageId="2" pageNumber="3">Fig. 34</figureCitation>
) ...........
<taxonomicName id="4C6A4D4AFFDF234DFD5A4FDCFCFFF96F" authorityName="Pinedo-Escatel &amp; Dietrich &amp; Zahniser" authorityYear="2022" baseAuthorityName="Lozada" baseAuthorityYear="1998" box="[676,778,1763,1782]" class="Insecta" family="Cicadellidae" genus="Andanus" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="sordidus">
<emphasis id="B91EEADBFFDF234DFD5A4FDCFCFFF96F" box="[676,778,1763,1782]" italics="true" pageId="2" pageNumber="3">A. sordidus</emphasis>
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<paragraph id="8BD536C9FFDF234DFCCE49F9FA69FEB2" blockId="2.[816,1437,198,789]" pageId="2" pageNumber="3">
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1. Pygofer without processes (
<figureCitation id="13512A4CFFDF234DFB8E49F9FB42FF40" box="[1136,1207,198,218]" captionStart="Figures 916" captionStartId="4.[159,226,1697,1717]" captionTargetBox="[251,1354,197,1678]" captionTargetId="figure-52@4.[245,1357,197,1679]" captionTargetPageId="4" captionText="Figures 916. Male of Andanus bimaculatus: (9) habitus, dorsal view; (10) habitus, lateral view; (11) face, anterior view; (13) pygofer, lateral view; (13) apodemes of sternite I, ventral view; (14) subgenital plates, valve, and styles, ventral view; (15) aedeagus and connective, lateral view; (16) apex of aedeagus, caudal view." figureDoi="http://doi.org/10.5281/zenodo.8233442" httpUri="https://zenodo.org/record/8233442/files/figure.png" pageId="2" pageNumber="3">Figs 12</figureCitation>
,
<figureCitation id="13512A4CFFDF234DFB3F49F9FB0AFF40" box="[1217,1279,198,217]" captionStart="Figures 1724" captionStartId="5.[151,218,1701,1721]" captionTargetBox="[165,1413,169,1686]" captionTargetId="figure-51@5.[209,1371,197,1686]" captionTargetPageId="5" captionText="Figures 1724. Male of Andanus raunoi comb. nov.: (17) habitus, dorsal view; (18) habitus, lateral view; (19) face, anterior view; (20) pygofer and segment X, dorsal view; (21) pygofer, lateral view; (22) subgenital plate and valve, ventral view; (23) aedeagus, connective, and style, ventral view; (24) aedeagus and connective, lateral view." figureDoi="http://doi.org/10.5281/zenodo.8233446" httpUri="https://zenodo.org/record/8233446/files/figure.png" pageId="2" pageNumber="3">20, 21</figureCitation>
); aedeagus with pair of apical or dorsopreapical spines and with or without single ventroapical spine; subgenital plates not as above (
<figureCitation id="13512A4CFFDF234DFC524828FC12FEB3" box="[940,999,279,299]" captionStart="Figures 18" captionStartId="3.[151,218,1694,1714]" captionTargetBox="[151,1408,186,1679]" captionTargetId="figure-48@3.[224,1357,197,1679]" captionTargetPageId="3" captionText="Figures 18. Holotype of Andanus tambopixunensis sp. nov.: (1) Habitus, dorsal view; (2) habitus, lateral view; (3) Face, anterior view; (4) pygofer, lateral view; (5) subgenital plate and style, dorsal view; (6) style, ventral view; (7) aedeagus and connective, lateral view; (8) aedeagus and connective, ventral view." figureDoi="http://doi.org/10.5281/zenodo.8233440" httpUri="https://zenodo.org/record/8233440/files/figure.png" pageId="2" pageNumber="3">Figs 5</figureCitation>
,
<figureCitation id="13512A4CFFDF234DFC0C4827FBF9FEB2" box="[1010,1036,280,299]" captionStart="Figures 916" captionStartId="4.[159,226,1697,1717]" captionTargetBox="[251,1354,197,1678]" captionTargetId="figure-52@4.[245,1357,197,1679]" captionTargetPageId="4" captionText="Figures 916. Male of Andanus bimaculatus: (9) habitus, dorsal view; (10) habitus, lateral view; (11) face, anterior view; (13) pygofer, lateral view; (13) apodemes of sternite I, ventral view; (14) subgenital plates, valve, and styles, ventral view; (15) aedeagus and connective, lateral view; (16) apex of aedeagus, caudal view." figureDoi="http://doi.org/10.5281/zenodo.8233442" httpUri="https://zenodo.org/record/8233442/files/figure.png" pageId="2" pageNumber="3">14</figureCitation>
,
<figureCitation id="13512A4CFFDF234DFBE84827FBDAFEB2" box="[1046,1071,280,299]" captionStart="Figures 1724" captionStartId="5.[151,218,1701,1721]" captionTargetBox="[165,1413,169,1686]" captionTargetId="figure-51@5.[209,1371,197,1686]" captionTargetPageId="5" captionText="Figures 1724. Male of Andanus raunoi comb. nov.: (17) habitus, dorsal view; (18) habitus, lateral view; (19) face, anterior view; (20) pygofer and segment X, dorsal view; (21) pygofer, lateral view; (22) subgenital plate and valve, ventral view; (23) aedeagus, connective, and style, ventral view; (24) aedeagus and connective, lateral view." figureDoi="http://doi.org/10.5281/zenodo.8233446" httpUri="https://zenodo.org/record/8233446/files/figure.png" pageId="2" pageNumber="3">22</figureCitation>
) ...................................................... 2
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2. Pygofer with numerous long macrosetae covering entire dorsoapical half (
<figureCitation id="13512A4CFFDF234DFBEF4871FBA0FEF8" box="[1041,1109,334,354]" captionStart="Figures 916" captionStartId="4.[159,226,1697,1717]" captionTargetBox="[251,1354,197,1678]" captionTargetId="figure-52@4.[245,1357,197,1679]" captionTargetPageId="4" captionText="Figures 916. Male of Andanus bimaculatus: (9) habitus, dorsal view; (10) habitus, lateral view; (11) face, anterior view; (13) pygofer, lateral view; (13) apodemes of sternite I, ventral view; (14) subgenital plates, valve, and styles, ventral view; (15) aedeagus and connective, lateral view; (16) apex of aedeagus, caudal view." figureDoi="http://doi.org/10.5281/zenodo.8233442" httpUri="https://zenodo.org/record/8233442/files/figure.png" pageId="2" pageNumber="3">Fig. 12</figureCitation>
); aedeagus with two pairs of apical spines directed caudad (
<figureCitation id="13512A4CFFDF234DFB7F4856FB01FEE5" box="[1153,1268,361,381]" captionStart="Figures 916" captionStartId="4.[159,226,1697,1717]" captionTargetBox="[251,1354,197,1678]" captionTargetId="figure-52@4.[245,1357,197,1679]" captionTargetPageId="4" captionText="Figures 916. Male of Andanus bimaculatus: (9) habitus, dorsal view; (10) habitus, lateral view; (11) face, anterior view; (13) pygofer, lateral view; (13) apodemes of sternite I, ventral view; (14) subgenital plates, valve, and styles, ventral view; (15) aedeagus and connective, lateral view; (16) apex of aedeagus, caudal view." figureDoi="http://doi.org/10.5281/zenodo.8233442" httpUri="https://zenodo.org/record/8233442/files/figure.png" pageId="2" pageNumber="3">Figs 15, 16</figureCitation>
), without single spine on ventral surface or long spines along shaft ......... .....................................................................
<taxonomicName id="4C6A4D4AFFDF234DFAEE489FFA68FE2A" authorityName="Linnavuori" authorityYear="1959" box="[1296,1437,416,435]" class="Insecta" family="Cicadellidae" genus="Andanus" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="bimaculatus">
<emphasis id="B91EEADBFFDF234DFAEE489FFA68FE2A" box="[1296,1437,416,435]" italics="true" pageId="2" pageNumber="3">A. bimaculatus</emphasis>
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2. Pygofer with macrosetae limited to smaller patch not extended to apex (
<figureCitation id="13512A4CFFDF234DFBFC48E9FBC9FE70" box="[1026,1084,470,490]" captionStart="Figures 18" captionStartId="3.[151,218,1694,1714]" captionTargetBox="[151,1408,186,1679]" captionTargetId="figure-48@3.[224,1357,197,1679]" captionTargetPageId="3" captionText="Figures 18. Holotype of Andanus tambopixunensis sp. nov.: (1) Habitus, dorsal view; (2) habitus, lateral view; (3) Face, anterior view; (4) pygofer, lateral view; (5) subgenital plate and style, dorsal view; (6) style, ventral view; (7) aedeagus and connective, lateral view; (8) aedeagus and connective, ventral view." figureDoi="http://doi.org/10.5281/zenodo.8233440" httpUri="https://zenodo.org/record/8233440/files/figure.png" pageId="2" pageNumber="3">Figs 4</figureCitation>
,
<figureCitation id="13512A4CFFDF234DFBB848E9FB95FE70" box="[1094,1120,470,489]" captionStart="Figures 1724" captionStartId="5.[151,218,1701,1721]" captionTargetBox="[165,1413,169,1686]" captionTargetId="figure-51@5.[209,1371,197,1686]" captionTargetPageId="5" captionText="Figures 1724. Male of Andanus raunoi comb. nov.: (17) habitus, dorsal view; (18) habitus, lateral view; (19) face, anterior view; (20) pygofer and segment X, dorsal view; (21) pygofer, lateral view; (22) subgenital plate and valve, ventral view; (23) aedeagus, connective, and style, ventral view; (24) aedeagus and connective, lateral view." figureDoi="http://doi.org/10.5281/zenodo.8233446" httpUri="https://zenodo.org/record/8233446/files/figure.png" pageId="2" pageNumber="3">21</figureCitation>
); aedeagus with one pair of long spines along shaft (
<figureCitation id="13512A4CFFDF234DFBD848CEFB97FD9C" box="[1062,1122,497,517]" captionStart="Figures 18" captionStartId="3.[151,218,1694,1714]" captionTargetBox="[151,1408,186,1679]" captionTargetId="figure-48@3.[224,1357,197,1679]" captionTargetPageId="3" captionText="Figures 18. Holotype of Andanus tambopixunensis sp. nov.: (1) Habitus, dorsal view; (2) habitus, lateral view; (3) Face, anterior view; (4) pygofer, lateral view; (5) subgenital plate and style, dorsal view; (6) style, ventral view; (7) aedeagus and connective, lateral view; (8) aedeagus and connective, ventral view." figureDoi="http://doi.org/10.5281/zenodo.8233440" httpUri="https://zenodo.org/record/8233440/files/figure.png" pageId="2" pageNumber="3">Figs 7</figureCitation>
,
<figureCitation id="13512A4CFFDF234DFB9348CEFB7DFD9D" box="[1133,1160,497,516]" captionStart="Figures 1724" captionStartId="5.[151,218,1701,1721]" captionTargetBox="[165,1413,169,1686]" captionTargetId="figure-51@5.[209,1371,197,1686]" captionTargetPageId="5" captionText="Figures 1724. Male of Andanus raunoi comb. nov.: (17) habitus, dorsal view; (18) habitus, lateral view; (19) face, anterior view; (20) pygofer and segment X, dorsal view; (21) pygofer, lateral view; (22) subgenital plate and valve, ventral view; (23) aedeagus, connective, and style, ventral view; (24) aedeagus and connective, lateral view." figureDoi="http://doi.org/10.5281/zenodo.8233446" httpUri="https://zenodo.org/record/8233446/files/figure.png" pageId="2" pageNumber="3">24</figureCitation>
), with or without preapical spines directed anterad and single spine on ventral surface ................................................................................. 3
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<keyStep id="309E2E6CFFDF234DFCCE4B7CFA68FC8C" pageId="2" pageNumber="3">
<paragraph id="8BD536C9FFDF234DFCCE4B7CFA69FD3E" blockId="2.[816,1437,198,789]" pageId="2" pageNumber="3">
<keyLead id="309B95FCFFDF234DFCCE4B7CFA69FD3E" pageId="2" pageNumber="3">
3. Style with lateral lobe broadly rounded near base, apophysis without triangular lateral projection near mid-length (
<figureCitation id="13512A4CFFDF234DFC924B46FC45FD15" box="[876,944,633,653]" captionStart="Figures 1724" captionStartId="5.[151,218,1701,1721]" captionTargetBox="[165,1413,169,1686]" captionTargetId="figure-51@5.[209,1371,197,1686]" captionTargetPageId="5" captionText="Figures 1724. Male of Andanus raunoi comb. nov.: (17) habitus, dorsal view; (18) habitus, lateral view; (19) face, anterior view; (20) pygofer and segment X, dorsal view; (21) pygofer, lateral view; (22) subgenital plate and valve, ventral view; (23) aedeagus, connective, and style, ventral view; (24) aedeagus and connective, lateral view." figureDoi="http://doi.org/10.5281/zenodo.8233446" httpUri="https://zenodo.org/record/8233446/files/figure.png" pageId="2" pageNumber="3">Fig. 23</figureCitation>
); aedeagus without short distal spines (
<figureCitation id="13512A4CFFDF234DFACB4B46FA8DFD15" box="[1333,1400,633,653]" captionStart="Figures 1724" captionStartId="5.[151,218,1701,1721]" captionTargetBox="[165,1413,169,1686]" captionTargetId="figure-51@5.[209,1371,197,1686]" captionTargetPageId="5" captionText="Figures 1724. Male of Andanus raunoi comb. nov.: (17) habitus, dorsal view; (18) habitus, lateral view; (19) face, anterior view; (20) pygofer and segment X, dorsal view; (21) pygofer, lateral view; (22) subgenital plate and valve, ventral view; (23) aedeagus, connective, and style, ventral view; (24) aedeagus and connective, lateral view." figureDoi="http://doi.org/10.5281/zenodo.8233446" httpUri="https://zenodo.org/record/8233446/files/figure.png" pageId="2" pageNumber="3">Fig. 24</figureCitation>
) ... .............................................................................
<taxonomicName id="4C6A4D4AFFDF234DFAB84BAAFA69FD3E" authorityName="Pinedo-Escatel &amp; Dietrich &amp; Zahniser" authorityYear="2022" baseAuthorityName="Zanol" baseAuthorityYear="1993" box="[1350,1436,661,680]" class="Insecta" family="Cicadellidae" genus="Andanus" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="raunoi">
<emphasis id="B91EEADBFFDF234DFAB84BAAFA69FD3E" box="[1350,1436,661,680]" italics="true" pageId="2" pageNumber="3">A. raunoi</emphasis>
</taxonomicName>
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</paragraph>
<paragraph id="8BD536C9FFDF234DFCCE4B8FFA68FC8C" blockId="2.[816,1437,198,789]" pageId="2" pageNumber="3">
<keyLead id="309B95FCFFDF234DFCCE4B8FFA68FC8C" pageId="2" pageNumber="3">
3. Style with lateral lobe near base acute, apophysis with triangular lateral projection (
<figureCitation id="13512A4CFFDF234DFB8A4BF5FB5BFD47" box="[1140,1198,714,734]" captionStart="Figures 18" captionStartId="3.[151,218,1694,1714]" captionTargetBox="[151,1408,186,1679]" captionTargetId="figure-48@3.[224,1357,197,1679]" captionTargetPageId="3" captionText="Figures 18. Holotype of Andanus tambopixunensis sp. nov.: (1) Habitus, dorsal view; (2) habitus, lateral view; (3) Face, anterior view; (4) pygofer, lateral view; (5) subgenital plate and style, dorsal view; (6) style, ventral view; (7) aedeagus and connective, lateral view; (8) aedeagus and connective, ventral view." figureDoi="http://doi.org/10.5281/zenodo.8233440" httpUri="https://zenodo.org/record/8233440/files/figure.png" pageId="2" pageNumber="3">Fig. 6</figureCitation>
); aedeagus with pair of short dorsoapical and single ventroapical spine (
<figureCitation id="13512A4CFFDF234DFABA4BD9FA8EFD60" box="[1348,1403,742,762]" captionStart="Figures 18" captionStartId="3.[151,218,1694,1714]" captionTargetBox="[151,1408,186,1679]" captionTargetId="figure-48@3.[224,1357,197,1679]" captionTargetPageId="3" captionText="Figures 18. Holotype of Andanus tambopixunensis sp. nov.: (1) Habitus, dorsal view; (2) habitus, lateral view; (3) Face, anterior view; (4) pygofer, lateral view; (5) subgenital plate and style, dorsal view; (6) style, ventral view; (7) aedeagus and connective, lateral view; (8) aedeagus and connective, ventral view." figureDoi="http://doi.org/10.5281/zenodo.8233440" httpUri="https://zenodo.org/record/8233440/files/figure.png" pageId="2" pageNumber="3">Fig. 7</figureCitation>
) .. ................................................
<taxonomicName id="4C6A4D4AFFDF234DFB6C4A3EFABCFC8D" authority="Pinedo-Escatel &amp; Dietrich &amp; Zahniser, 2022" authorityName="Pinedo-Escatel &amp; Dietrich &amp; Zahniser" authorityYear="2022" box="[1170,1353,769,788]" class="Insecta" family="Cicadellidae" genus="Andanus" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="tambopixunensis" status="sp. nov.">
<emphasis id="B91EEADBFFDF234DFB6C4A3EFABCFC8D" box="[1170,1353,769,788]" italics="true" pageId="2" pageNumber="3">A. tambopixunensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22D57A0FFDF234DFAB14A3DFA68FC8C" box="[1359,1437,770,789]" pageId="2" pageNumber="3" rank="species">sp. nov.</taxonomicNameLabel>
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</treatment>
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