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<document id="2867789BC00DD985BF296A6D279BF875" ID-DOI="10.1590/s0031-10492010002300001" ID-ISSN="1807-0205" ID-Zenodo-Dep="10085966" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1622567237596" checkinUser="carolina" docAuthor="De Albuquerque, Nelson R. &amp; Mcdiarmid, Roy W." docDate="2010" docId="038C2E7EFFDBFFA66BC5475EFB3699BC" docLanguage="en" docName="PapAvZool.50.23.375-384.pdf.imf" docOrigin="Papéis Avulsos de Zoologia 50 (23)" docStyle="DocumentStyle:8E93E160036069519CAC16E940FF2271.1:PapAvZool.2014-2017.journal_article" docStyleId="8E93E160036069519CAC16E940FF2271" docStyleName="PapAvZool.2014-2017.journal_article" docStyleVersion="1" docTitle="Leptophis cupreus" docType="treatment" docVersion="2" lastPageNumber="384" masterDocId="FFB55606FFDAFFA36B014329FFA4982C" masterDocTitle="Redescription Of Leptophis cupreus (Cope) (Serpentes, Colubridae), A Rare South American Colubrine Snake" masterLastPageNumber="384" masterPageNumber="375" pageNumber="375" updateTime="1699477781843" updateUser="ExternalLinkService">
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<mods:title id="897FA04921E40C12B723F8AB9687BF48">Redescription Of Leptophis cupreus (Cope) (Serpentes, Colubridae), A Rare South American Colubrine Snake</mods:title>
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<mods:namePart id="6C6022608A6A6930AA7CC0EF5FA4D0B4">De Albuquerque, Nelson R.</mods:namePart>
<mods:affiliation id="C239807B22A28D2EEA1EFEB68FB07D25">Departamento de Ciências do Ambiente, Universidade Federal de Mato Grosso do Sul, Avenida Rio Branco, 1.270, Universitário, 79304 - 020, Corumbá, MS, Brasil.</mods:affiliation>
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<mods:namePart id="C324CB51D0E17903FB1C53655D8D4085">Mcdiarmid, Roy W.</mods:namePart>
<mods:affiliation id="18E493B2121891D1F84C22003E8F5F0D">USGS Patuxent Wildlife Research Center, Smithsonian Institution, PO Box 37012, National Museum of Natural History, Room 378, MRC 111, Washington, DC 20013 - 7012, USA.</mods:affiliation>
<mods:nameIdentifier id="71225CE57EF0E197A0AF92CF0AFB098B" type="email">mcdiarmr@si.edu</mods:nameIdentifier>
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<mods:title id="3003910A58D4CB1BD1AC195884657D77">Papéis Avulsos de Zoologia</mods:title>
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<treatment id="038C2E7EFFDBFFA66BC5475EFB3699BC" LSID="urn:lsid:plazi:treatment:038C2E7EFFDBFFA66BC5475EFB3699BC" httpUri="http://treatment.plazi.org/id/038C2E7EFFDBFFA66BC5475EFB3699BC" lastPageId="5" pageId="1">
<subSubSection id="C33FCCE3FFDBFFA26BC5475EFC25995C" pageId="1" type="nomenclature">
<paragraph id="8B9A9F68FFDBFFA26BC5475EFDB99CBC" blockId="1.[196,541,1143,1168]" box="[196,541,1143,1168]" pageId="1">
<heading id="D0D22804FFDBFFA26BC5475EFDB99CBC" bold="true" box="[196,541,1143,1168]" centered="true" fontSize="9" level="2" pageId="1" reason="2">
<taxonomicName id="4C25E4EBFFDBFFA26BC5475EFDB99CBC" authority="(Cope, 1868)" baseAuthorityName="Cope" baseAuthorityYear="1868" box="[196,541,1143,1168]" class="Reptilia" family="Colubridae" genus="Leptophis" kingdom="Animalia" order="Squamata" pageId="1" phylum="Chordata" rank="species" species="cupreus">
<emphasis id="B951437AFFDBFFA26BC5475EFDB99CBC" bold="true" box="[196,541,1143,1168]" pageId="1">
<emphasis id="B951437AFFDBFFA26BC5475EFE229CA3" bold="true" box="[196,390,1143,1167]" italics="true" pageId="1">Leptophis cupreus</emphasis>
(
<bibRefCitation id="EFB4E299FFDBFFA26A944751FDB19CBC" author="COPE" box="[405,533,1144,1168]" pageId="1" pagination="96 - 140" refId="ref4919" refString="COPE, E. D. 1868. An examination of the Reptilia and Batrachia obtained by the Orton Expedition to Equador and the upper Amazon, with notes on other species. Proceedings of the Academy of Natural Sciences of Philadelphia, 20: 96 - 140." type="journal article" year="1868">Cope, 1868</bibRefCitation>
)
</emphasis>
</taxonomicName>
</heading>
</paragraph>
<paragraph id="8B9A9F68FFDBFFA269A94391FC25995C" blockId="1.[680,1229,184,1136]" pageId="1">
<materialsCitation id="3B4D9535FFDBFFA269A94391FCDA995C" collectionCode="ANSP" pageId="1" specimenCode="USNM 6666" specimenCount="1">
to as “No. 6666” by Cope in the
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description (1868:106). This specimen, originally cataloged as
<specimenCode id="DB833713FFDBFFA269A943D1FC9D993C" box="[680,825,248,272]" collectionCode="USNM" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="1" type="Museum">USNM 6666</specimenCode>
, was apparently recataloged as
<collectionCode id="ED3407ADFFDBFFA26F8A43D1FB68993C" box="[1163,1228,248,272]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34784" lsid="urn:lsid:biocol.org:col:34784" name="Academy of Natural Sciences of Philadelphia" pageId="1">ANSP</collectionCode>
<quantity id="4CDD328DFFDBFFA269A94231FD5F991C" box="[680,763,280,304]" metricMagnitude="2" metricUnit="m" metricValue="1.321308" pageId="1" unit="in" value="5202.0">5202 in</quantity>
the confusion over provenance of specimens in Copes possession at the time of his death in 1897 (see later discussion)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C33FCCE3FFDBFFA269E04251FB699C5C" pageId="1" type="description">
<paragraph id="8B9A9F68FFDBFFA269E04251FB699C5C" blockId="1.[680,1229,184,1136]" pageId="1">
The ventral and subcaudal counts for this specimen are identical to those reported by
<bibRefCitation id="EFB4E299FFDBFFA26F4142B1FB6C999C" author="COPE" box="[1088,1224,408,432]" pageId="1" pagination="96 - 140" refId="ref4919" refString="COPE, E. D. 1868. An examination of the Reptilia and Batrachia obtained by the Orton Expedition to Equador and the upper Amazon, with notes on other species. Proceedings of the Academy of Natural Sciences of Philadelphia, 20: 96 - 140." type="journal article" year="1868">Cope (1868)</bibRefCitation>
; other details of scutellation are the same, and the measurements are extremely similar. The coloration is different in several respects from that presented in Copes description; however, the specimen has lost all of the
<emphasis id="B951437AFFDBFFA269A94110FCFC9A7C" box="[680,856,569,592]" italics="true" pageId="1">stratum corneum,</emphasis>
a condition that had begun when Cope described it, and as a result has not retained its original coloration. This specimen is a small female, possibly juvenile, SVL
<quantity id="4CDD328DFFDBFFA2689541B1FC549A83" box="[916,1008,664,688]" metricMagnitude="-1" metricUnit="m" metricValue="3.13" pageId="1" unit="mm" value="313.0">313 mm</quantity>
, tail length
<quantity id="4CDD328DFFDBFFA26F6D41B1FB6C9A83" box="[1132,1224,664,688]" metricMagnitude="-1" metricUnit="m" metricValue="2.05" pageId="1" unit="mm" value="205.0">205 mm</quantity>
, with bands on the anterior and middle region of the body, similar to those found in juveniles of other species of
<taxonomicName id="4C25E4EBFFDBFFA269EF41D1FCEE9B3C" box="[750,842,760,784]" class="Reptilia" family="Colubridae" genus="Leptophis" kingdom="Animalia" order="Squamata" pageId="1" phylum="Chordata" rank="genus">
<emphasis id="B951437AFFDBFFA269EF41D1FCEE9B3C" box="[750,842,760,784]" italics="true" pageId="1">Leptophis</emphasis>
</taxonomicName>
(
<emphasis id="B951437AFFDBFFA2685841D0FCD69B3C" box="[857,882,761,784]" italics="true" pageId="1">see</emphasis>
<bibRefCitation id="EFB4E299FFDBFFA2687941D1FC589B3C" author="OLIVER" box="[888,1020,760,784]" pageId="1" pagination="157 - 280" refId="ref5641" refString="OLIVER, J. A. 1948. The relationships and zoogeography of the genus Thalerophis Oliver. Bulletin of the American Museum of Natural History, 92: 157 - 280." type="journal article" year="1948">Oliver, 1948</bibRefCitation>
). Head elongate and distinctly broader than neck, narrower than diameter of midbody. Head length from the posterior tip of retroarticular process of the mandible
<quantity id="4CDD328DFFDBFFA26F184071FB2C9B43" box="[1049,1160,856,880]" metricMagnitude="-2" metricUnit="m" metricValue="1.2269999999999999" pageId="1" unit="mm" value="12.27">12.27 mm</quantity>
, 3.9% of SVL. Snout length from tip of snout to anterior margin of orbit
<quantity id="4CDD328DFFDBFFA2685740B1FC189B83" box="[854,956,920,944]" metricMagnitude="-3" metricUnit="m" metricValue="3.8" pageId="1" unit="mm" value="3.8">3.80 mm</quantity>
. Rostral wider than high, visible from above. Nasals undivided; right nasal separated from preocular by broad prefrontal contact with second and third supralabials; left nasal separated from preocular by loreal scale. Prefrontals not contacting orbits; prefrontals slightly larger than internasals. Frontal longer than wide, about twice as long as prefrontals.
</paragraph>
</subSubSection>
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<taxonomicName id="4C25E4EBFFDBFFA26B5E4791FE069CFC" authority="Cope, 1868" authorityName="Cope" authorityYear="1868" box="[95,418,1208,1232]" class="Reptilia" family="Colubridae" genus="Thrasops" kingdom="Animalia" order="Squamata" pageId="1" phylum="Chordata" rank="species" species="cupreus">
<emphasis id="B951437AFFDBFFA26B5E4791FEA89CFC" box="[95,268,1208,1232]" italics="true" pageId="1">Thrasops cupreus</emphasis>
<materialsCitation id="3B4D9535FFDBFFA26A1D4791FE069CFC" box="[284,418,1208,1232]" collectionCode="ANSP" collectorName="James Orton" county="Proceedings" location="Natural Sciences of Philadelphia" municipality="Academy of Natural Sciences" pageId="1" specimenCode="ANSP 5202" specimenCount="1" stateProvince="Napo and Maranon" typeStatus="holotype">
<bibRefCitation id="EFB4E299FFDBFFA26A1D4791FE069CFC" author="COPE" box="[284,418,1208,1232]" pageId="1" pagination="96 - 140" refId="ref4919" refString="COPE, E. D. 1868. An examination of the Reptilia and Batrachia obtained by the Orton Expedition to Equador and the upper Amazon, with notes on other species. Proceedings of the Academy of Natural Sciences of Philadelphia, 20: 96 - 140." type="journal article" year="1868">Cope, 1868</bibRefCitation>
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</taxonomicName>
.
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of the
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Academy of
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of Philadelphia 20:96-140.
<typeStatus id="549E21CAFFDBFFA26A1047D1FED29D3C" box="[273,374,1272,1296]" pageId="1" type="holotype">Holotype</typeStatus>
:
<specimenCode id="DB833713FFDBFFA26A7E47D1FE5B9D3C" box="[383,511,1272,1296]" collectionCode="ANSP" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34784" lsid="urn:lsid:biocol.org:col:34784" name="Academy of Natural Sciences of Philadelphia" pageId="1">ANSP 5202</specimenCode>
, collected by
<collectorName id="26D0FABEFFDBFFA26B964631FEB89D1C" box="[151,284,1304,1328]" pageId="1">James Orton</collectorName>
in late 1867. Type-locality: “from the
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”.
</paragraph>
<paragraph id="8B9A9F68FFDBFFA26B5E4671FE369EBC" blockId="1.[95,643,1208,1680]" pageId="1">
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<taxonomicName id="4C25E4EBFFDBFFA26B5E4671FEB69D5C" baseAuthorityName="Cope" baseAuthorityYear="1868" box="[95,274,1368,1392]" class="Reptilia" family="Colubridae" genus="Leptophis" kingdom="Animalia" order="Squamata" pageId="1" phylum="Chordata" rank="species" species="cupreus">
<emphasis id="B951437AFFDBFFA26B5E4671FEB69D5C" box="[95,274,1368,1392]" italics="true" pageId="1">Leptophis cupreus</emphasis>
</taxonomicName>
<treatmentCitation id="0A84B979FFDBFFA26A3D4671FDBB9D5C" author="BOULENGER" box="[316,543,1368,1392]" page="109" pageId="1" year="1894">
<bibRefCitation id="EFB4E299FFDBFFA26A3D4671FDBB9D5C" author="BOULENGER" box="[316,543,1368,1392]" pageId="1" refId="ref4816" refString="BOULENGER, G. A. 1894. Catalogue of the Snakes in the British Museum (Natural History). British Museum of Natural History, London, v. II." type="journal volume" year="1894">Boulenger, 1894:109</bibRefCitation>
</treatmentCitation>
;
<treatmentCitation id="0A84B979FFDBFFA269304671FE8A9DBC" author="WERNER" page="102" pageId="1" year="1929">
<bibRefCitation id="EFB4E299FFDBFFA269304671FE8A9DBC" author="WERNER" pageId="1" pagination="1 - 196" refId="ref5923" refString="WERNER, F. 1929. Ubersicht der Gattungen und Arten der Schlangen aus der Familie Colubridae. III. Teil (Colubrinae). Zoologische Jahrbucher Abteilung fur Systematik, 57: 1 - 196." type="journal article" year="1929">Werner, 1929:102-103</bibRefCitation>
</treatmentCitation>
; Peters, 1960:525; Peters &amp; Orcés-V, 1960:139-141; Peters &amp; Orejas-Miranda, 1970:164;
<treatmentCitation id="0A84B979FFDBFFA26A0E4691FDA99DFC" author="MERTENS" box="[271,525,1464,1488]" page="144" pageId="1" year="1973">
<bibRefCitation id="EFB4E299FFDBFFA26A0E4691FDA99DFC" author="MERTENS" box="[271,525,1464,1488]" pageId="1" pagination="141 - 154" refId="ref5495" refString="MERTENS, R. 1973. Bemerkenswerte Schlanknattern der neotropischen Gattung Leptophis. Studies on the Neotropical Fauna, 8: 141 - 154." type="journal article" year="1973">Mertens, 1973:144-145</bibRefCitation>
</treatmentCitation>
; Dixon &amp; Soini, 1977:20, 58;
<treatmentCitation id="0A84B979FFDBFFA26A6E46F1FDDA9DDC" author="DUELLMAN" box="[367,638,1496,1520]" page="249" pageId="1" year="1978">
<bibRefCitation id="EFB4E299FFDBFFA26A6E46F1FDDA9DDC" author="DUELLMAN" box="[367,638,1496,1520]" pageId="1" pagination="1 - 352" refId="ref5160" refString="DUELLMAN, W. E. 1978. The biology of an equatorial herpetofauna in Amazonian Ecuador. Miscellaneous Publication, University of Kansas, Museum of Natural History, 65: 1 - 352." type="journal article" year="1978">Duellman, 1978:249-250</bibRefCitation>
</treatmentCitation>
; Dixon &amp; Soini, 1986:6, 75, 114; Pérez-Santos &amp;
<treatmentCitation id="0A84B979FFDBFFA26BC14531FE669E1C" author="MORENO &amp; Ofidios de Colombia. Museo Regionale di Scienze Naturali &amp; Torino &amp; Monografie VI" box="[192,450,1560,1584]" page="213" pageId="1" year="1988">
<bibRefCitation id="EFB4E299FFDBFFA26BC14531FE669E1C" author="MORENO &amp; Ofidios de Colombia. Museo Regionale di Scienze Naturali &amp; Torino &amp; Monografie VI" box="[192,450,1560,1584]" pageId="1" refId="ref5699" refString="PEREZ-SANTOS, C. &amp; MORENO, A. G. 1988. Ofidios de Colombia. Museo Regionale di Scienze Naturali, Torino, Monografie VI." type="book" year="1988">Moreno, 1988:213-214</bibRefCitation>
</treatmentCitation>
; Pérez-Santos &amp;
<treatmentCitation id="0A84B979FFDBFFA26B964511FE2A9E7C" author="MORENO &amp; Serpientes de Colombia. Museo Regionale di Scienze Naturali &amp; Torino &amp; Monografie XI" box="[151,398,1592,1616]" page="219" pageId="1" year="1991">
<bibRefCitation id="EFB4E299FFDBFFA26B964511FE2A9E7C" author="MORENO &amp; Serpientes de Colombia. Museo Regionale di Scienze Naturali &amp; Torino &amp; Monografie XI" box="[151,398,1592,1616]" pageId="1" refId="ref5727" refString="PEREZ-SANTOS, C. &amp; MORENO, A. G. 1991. Serpientes de Colombia. Museo Regionale di Scienze Naturali, Torino, Monografie XI." type="book" year="1991">Moreno, 1991:219-220</bibRefCitation>
</treatmentCitation>
; Carrillo de Espinoza &amp; Icochea, 1995:16;
<treatmentCitation id="0A84B979FFDBFFA26A594571FD879E5C" author="HARDING" box="[344,547,1624,1648]" page="225" pageId="1" year="1995">
<bibRefCitation id="EFB4E299FFDBFFA26A594571FD879E5C" author="HARDING" box="[344,547,1624,1648]" pageId="1" pagination="221 - 226" refId="ref5285" refString="HARDING, K. A. 1995. A new species of tree snake of the genus Leptophis Bell 1825 from Mount Aripo, Trinidad. Tropical Zoology, 8: 221 - 226." type="journal article" year="1995">Harding, 1995:225</bibRefCitation>
</treatmentCitation>
; Jorge da Silva &amp; Sites, 1995:895.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C33FCCE3FFDBFFA76BB245F1FF6D9A5C" lastPageId="4" pageId="1" type="description">
<paragraph id="8B9A9F68FFDBFFA26BB245F1FD8A9EDC" blockId="1.[179,558,1752,1776]" box="[179,558,1752,1776]" pageId="1">
<emphasis id="B951437AFFDBFFA26BB245F1FD8A9EDC" bold="true" box="[179,558,1752,1776]" pageId="1">
Description of
<typeStatus id="549E21CAFFDBFFA26A5645F1FE139EDC" box="[343,439,1752,1776]" pageId="1" type="holotype">holotype</typeStatus>
(
<figureCitation id="131E83EDFFDBFFA26AC445F1FD829EDC" box="[453,550,1752,1776]" captionStart-0="FIGURE 1" captionStart-1="FIGURE 2" captionStartId-0="1.[680,760,1800,1819]" captionStartId-1="2.[132,212,1851,1870]" captionTargetBox-0="[680,1228,1216,1764]" captionTargetBox-1="[132,1266,757,1815]" captionTargetId-0="figure-598@1.[680,1228,1216,1764]" captionTargetId-1="figure-247@2.[132,1266,757,1815]" captionTargetPageId-0="1" captionTargetPageId-1="2" captionText-0="FIGURE 1: Dorsal view of the holotype of Leptophis cupreus (ANSP 5202), a young female specimen (518 SVL) from the “Napo and Maranon”." captionText-1="FIGURE 2: Dorsal (A) and lateral (B) view of head of the holotype of Leptophis cupreus (ANSP 5202)." figureDoi-0="http://doi.org/10.5281/zenodo.10085968" figureDoi-1="http://doi.org/10.5281/zenodo.10085970" httpUri-0="https://zenodo.org/record/10085968/files/figure.png" httpUri-1="https://zenodo.org/record/10085970/files/figure.png" pageId="1">Figs. 1-2</figureCitation>
)
</emphasis>
</paragraph>
<caption id="DF5ACFE0FFDBFFA269A94421FCC69F63" ID-DOI="http://doi.org/10.5281/zenodo.10085968" ID-Zenodo-Dep="10085968" httpUri="https://zenodo.org/record/10085968/files/figure.png" pageId="1" startId="1.[680,760,1800,1819]" targetBox="[680,1228,1216,1764]" targetPageId="1">
<paragraph id="8B9A9F68FFDBFFA269A94421FCC69F63" blockId="1.[680,1229,1800,1871]" pageId="1">
<emphasis id="B951437AFFDBFFA269A94421FCB79F37" bold="true" box="[680,787,1800,1819]" pageId="1">FIGURE 1:</emphasis>
Dorsal view of the holotype of
<taxonomicName id="4C25E4EBFFDBFFA26F3C4421FB699F37" baseAuthorityName="Cope" baseAuthorityYear="1868" box="[1085,1229,1800,1819]" class="Reptilia" family="Colubridae" genus="Leptophis" kingdom="Animalia" order="Squamata" pageId="1" phylum="Chordata" rank="species" species="cupreus">
<emphasis id="B951437AFFDBFFA26F3C4421FB699F37" box="[1085,1229,1800,1819]" italics="true" pageId="1">Leptophis cupreus</emphasis>
</taxonomicName>
(ANSP 5202), a young female specimen (518 SVL) from the “Napo and Maranon”.
</paragraph>
</caption>
<paragraph id="8B9A9F68FFDBFFA16B964431FB2D99DC" blockId="1.[95,643,1816,1872]" lastBlockId="2.[718,1267,184,496]" lastPageId="2" pageId="1">
As noted by
<bibRefCitation id="EFB4E299FFDBFFA26A174431FE1E9F1C" author="MALNATE" box="[278,442,1816,1840]" pageId="1" pagination="345 - 375" refId="ref5408" refString="MALNATE, E. V. 1971. A catalog of primary types in the herpetological collections of the Academy of Natural Sciences, Philadelphia (ANSP). Proceedings of the Academy of Natural Sciences of Philadelphia, 123: 345 - 375." type="journal article" year="1971">Malnate (1971)</bibRefCitation>
, ANSP 5202 corresponds to the
<typeStatus id="549E21CAFFDBFFA26BF14411FEEE9F7C" box="[240,330,1848,1872]" pageId="1" type="holotype">holotype</typeStatus>
of
<taxonomicName id="4C25E4EBFFDBFFA26A6C4410FE759F7C" baseAuthorityName="Cope" baseAuthorityYear="1868" box="[365,465,1849,1872]" class="Reptilia" family="Colubridae" genus="Leptophis" kingdom="Animalia" order="Squamata" pageId="1" phylum="Chordata" rank="species" species="cupreus">
<emphasis id="B951437AFFDBFFA26A6C4410FE759F7C" box="[365,465,1849,1872]" italics="true" pageId="1">L. cupreus</emphasis>
</taxonomicName>
that was referred Single anterior temporal on each side in contact with parietal, postoculars, and sixth, seventh and eight supralabials; two posterior temporal scales, upper reaching the end of parietal on each side. Eye large, horizontal diameter
<quantity id="4CDD328DFFD8FFA16A094211FECF9963" box="[264,363,312,336]" metricMagnitude="-3" metricUnit="m" metricValue="2.64" pageId="2" unit="mm" value="2.64">2.64 mm</quantity>
, pupil round. Single preocular on each side, in contact with frontal. Two postoculars on each side, the upper about three times higher than lower. Two pairs of elongate chin shields separated by mental groove, with posterior pair distinctly more elongate. Mental not touching anterior chin shields. Anal plate divided; 152 ventrals; 137 paired subcaudals (tail complete); scales of vertebral and paravertebral rows slightly larger than those of adjacent rows; supralabials 8/8, 4-5/
<quantity id="4CDD328DFFD8FFA16BE04171FEBB9A5C" box="[225,287,600,624]" metricMagnitude="-1" metricUnit="m" metricValue="1.143" metricValueMax="1.27" metricValueMin="1.016" pageId="2" unit="in" value="4.5" valueMax="5.0" valueMin="4.0">4-5 in</quantity>
contact with orbit; infralabials 10/10, 1-
<quantity id="4CDD328DFFD8FFA16B994151FF659ABC" box="[152,193,632,656]" metricMagnitude="-1" metricUnit="m" metricValue="1.27" pageId="2" unit="in" value="5.0">5 in</quantity>
contact with anterior chin shields; parietals longer than broad and in contact with upper postocular. Dorsal scale rows 15-15-11; keels on dorsal scale rows 2-14 of trunk (reduced and often indistinct on scales of vertebral row), absent on first dorsal rows; dorsal scales of tail without keels. Single apical pit present on all dorsal scales of trunk, except those in first dorsal row. Narrow black ocular stripe along upper margins of second and third supralabials, covering lower postocular, lower edges of anterior and lower posterior temporals, and upper edges of last three supralabials; stripe disappears two scales posterior to last supralabial.
</paragraph>
<paragraph id="8B9A9F68FFD8FFA168AF4111FBB69A7C" blockId="2.[942,1042,568,592]" box="[942,1042,568,592]" pageId="2">
<emphasis id="B951437AFFD8FFA168AF4111FBB69A7C" bold="true" box="[942,1042,568,592]" pageId="2">Variation</emphasis>
</paragraph>
<paragraph id="8B9A9F68FFD8FFA068064151FC839CC3" blockId="2.[718,1266,632,688]" lastBlockId="3.[680,1229,952,1264]" lastPageId="3" pageId="2">
<materialsCitation id="3B4D9535FFD8FFA068064151FC869CC3" collectionCode="ICN, MCZ, USNM, LACM" lastPageId="3" pageId="2" specimenCode="ICN 8382, MCZ 164915, USNM 197281, 211036-38, 211041-42, LACM 45444, 76811, ICN 390, USNM 562696, ICN 8379" specimenCount="18" typeStatus="holotype">
Additional specimens examined are similar in scutellation to the
<typeStatus id="549E21CAFFD8FFA1688F41B1FC4D9A9C" box="[910,1001,664,688]" pageId="2" type="holotype">holotype</typeStatus>
(
<tableCitation id="C6A7AAD3FFD8FFA168F741B1FBE69A9C" box="[1014,1090,664,688]" captionStart="TABLE 1" captionStartId="3.[95,160,187,206]" captionText="TABLE 1: Morphometric and meristic variation in the specimens of Leptophis cupreus examined. Sl: Supralabials; SlO: Supralabials entering the orbit; Il: Infralabials; Ilg: Infralabials contacting the first genials; Pr/Pos: Preoculars/postoculars; Ta/Tp: Temporal anterior/ temporal posterior; P + VE: number of pre-ventral and ventral scales; SC: Number of subcaudal scales; HL: head length; SL: snout length; SVL: Snout-vent length; TL: Tail length; m: male; f: female; juv: juvenile. All measurements are in mm.*Holotype of Leptophis cupreus." pageId="2">Table 1</tableCitation>
). The preoculars contact the frontal in eight (
<specimenCode id="DB833713FFD9FFA06A744091FE479BFC" box="[373,483,952,976]" collectionCode="ICN" country="Colombia" httpUri="http://grbio.org/cool/xsgz-nxkh" name="Instituto de Ciencias Naturales, Museo de Historia Natural" pageId="3">ICN 8382</specimenCode>
,
<specimenCode id="DB833713FFD9FFA06AEA4091FDDB9BFC" box="[491,639,952,976]" collectionCode="MCZ" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:33791" lsid="urn:lsid:biocol.org:col:33791" name="Museum of Comparative Zoology" pageId="3" type="Museum">MCZ 164915</specimenCode>
,
<specimenCode id="DB833713FFD9FFA06B5F40F1FEA29BDC" box="[94,262,984,1008]" collectionCode="USNM" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="3" type="Museum">USNM 197281</specimenCode>
,
<specimenCode id="DB833713FFD9FFA06A1140F1FE229BDC" box="[272,390,984,1008]" collectionCode="USNM" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="3" type="Museum">211036-38</specimenCode>
,
<specimenCode id="DB833713FFD9FFA06A9140F1FDA29BDC" box="[400,518,984,1008]" collectionCode="USNM" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="3" type="Museum">211041-42</specimenCode>
) of
<specimenCount id="9D2354E1FFD9FFA0692F40F1FF3F9C3C" pageId="3" type="generic">18 specimens</specimenCount>
examined. The dark postocular stripe extends nine scales posteriorly from the last supralabial as a diffuse black stripe in
<specimenCode id="DB833713FFD9FFA06A444711FE109C7C" box="[325,436,1080,1104]" collectionCode="ICN" country="Colombia" httpUri="http://grbio.org/cool/xsgz-nxkh" name="Instituto de Ciencias Naturales, Museo de Historia Natural" pageId="3">ICN 8382</specimenCode>
; in other specimens it is shorter and more diffuse (
<emphasis id="B951437AFFD9FFA06AB04770FE7D9C5C" box="[433,473,1113,1136]" italics="true" pageId="3">e.g.,</emphasis>
<specimenCode id="DB833713FFD9FFA06AE54771FDDB9C5C" box="[484,639,1112,1136]" collectionCode="LACM" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34802" lsid="urn:lsid:biocol.org:col:34802" name="Natural History Museum of Los Angeles County" pageId="3" type="Museum">LACM 45444</specimenCode>
,
<specimenCode id="DB833713FFD9FFA06B5F4751FF069CBC" box="[94,162,1144,1168]" collectionCode="LACM" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34802" lsid="urn:lsid:biocol.org:col:34802" name="Natural History Museum of Los Angeles County" pageId="3" type="Museum">76811</specimenCode>
) (
<figureCitation id="131E83EDFFD9FFA06BBE4751FEA79CBC" box="[191,259,1144,1168]" captionStart="FIGURE 3" captionStartId="3.[95,175,1826,1845]" captionTargetBox="[94,642,1487,1789]" captionTargetId="figure-357@3.[94,642,1487,1789]" captionTargetPageId="3" captionText="FIGURE 3: Leptophis cupreus (LACM 45444).Lateral view of head showing the short and diffuse postocular stripe." figureDoi="http://doi.org/10.5281/zenodo.10085972" httpUri="https://zenodo.org/record/10085972/files/figure.png" pageId="3">Fig. 3</figureCitation>
).
<specimenCode id="DB833713FFD9FFA06A1F4751FE239CBC" box="[286,391,1144,1168]" collectionCode="ICN" country="Colombia" httpUri="http://grbio.org/cool/xsgz-nxkh" name="Instituto de Ciencias Naturales, Museo de Historia Natural" pageId="3">ICN 390</specimenCode>
,
<specimenCode id="DB833713FFD9FFA06A994751FD929CBC" box="[408,566,1144,1168]" collectionCode="LACM" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34802" lsid="urn:lsid:biocol.org:col:34802" name="Natural History Museum of Los Angeles County" pageId="3" type="Museum">LACM 45444</specimenCode>
,
<specimenCode id="DB833713FFD9FFA069474751FF0A9C9C" collectionCode="MCZ" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:33791" lsid="urn:lsid:biocol.org:col:33791" name="Museum of Comparative Zoology" pageId="3" type="Museum">MCZ 164915</specimenCode>
and
<specimenCode id="DB833713FFD9FFA06BE447B1FE2F9C9C" box="[229,395,1176,1200]" collectionCode="USNM" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="3" type="Museum">USNM 562696</specimenCode>
have keels on all dorsal scales of the trunk, except on the first; in some specimens the keels are reduced on scales of the second and fourteenth rows anterior to the reduction from 15 to 11 scale rows and on scales of the vertebral row; dorsal scales on the tail are keeled posterior to the point of reduction. Keels are more prominent in males than females and juveniles, and not visible in a few long preserved specimens. The
<emphasis id="B951437AFFD9FFA06F224090FB699BFC" box="[1059,1229,953,976]" italics="true" pageId="3">stratum corneum</emphasis>
has been lost in
<specimenCode id="DB833713FFD9FFA0685B40F1FC689BDC" box="[858,972,984,1008]" collectionCode="ICN" country="Colombia" httpUri="http://grbio.org/cool/xsgz-nxkh" name="Instituto de Ciencias Naturales, Museo de Historia Natural" pageId="3">ICN 8379</specimenCode>
; a pale stripe appears at vertebral scale 4 (
<quantity id="4CDD328DFFD9FFA0685E40D1FC099C23" box="[863,941,1016,1040]" metricMagnitude="-2" metricUnit="m" metricValue="3.3" pageId="3" unit="mm" value="33.0">33 mm</quantity>
from the tip of the snout), borders paravertebral rows at vertebral 18 (
<quantity id="4CDD328DFFD9FFA06F7D4731FB689C03" box="[1148,1228,1048,1072]" metricMagnitude="-2" metricUnit="m" metricValue="7.8" pageId="3" unit="mm" value="78.0">78 mm</quantity>
from the tip of the snout), and becomes indistinct at vertebral 27 (
<quantity id="4CDD328DFFD9FFA068504771FC089C43" box="[849,940,1112,1136]" metricMagnitude="-1" metricUnit="m" metricValue="1.1" pageId="3" unit="mm" value="110.0">110 mm</quantity>
from the tip of the snout). In life, the dorsal color of the Venezuelan specimen (
<specimenCode id="DB833713FFD9FFA069B147B1FCFC9C9C" box="[688,856,1176,1200]" collectionCode="USNM" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="3" type="Museum">USNM 562696</specimenCode>
) was coppery tan; the chin and first 15 ventral scales were white and the remainder of the ventrals tan
</materialsCitation>
.
</paragraph>
<caption id="DF5ACFE0FFD8FFA16B854412FC729F62" ID-DOI="http://doi.org/10.5281/zenodo.10085970" ID-Zenodo-Dep="10085970" box="[132,982,1851,1871]" httpUri="https://zenodo.org/record/10085970/files/figure.png" pageId="2" startId="2.[132,212,1851,1870]" targetBox="[132,1266,757,1815]" targetPageId="2">
<paragraph id="8B9A9F68FFD8FFA16B854412FC729F62" blockId="2.[132,982,1851,1871]" box="[132,982,1851,1871]" pageId="2">
<emphasis id="B951437AFFD8FFA16B854412FF4D9F62" bold="true" box="[132,233,1851,1870]" pageId="2">FIGURE 2:</emphasis>
Dorsal
<emphasis id="B951437AFFD8FFA16A2A4412FEE39F62" bold="true" box="[299,327,1851,1870]" pageId="2">(A)</emphasis>
and lateral
<emphasis id="B951437AFFD8FFA16AA64412FE679F62" bold="true" box="[423,451,1851,1871]" pageId="2">(B)</emphasis>
view of head of the holotype of
<taxonomicName id="4C25E4EBFFD8FFA169CF4415FCFC9F62" baseAuthorityName="Cope" baseAuthorityYear="1868" box="[718,856,1852,1871]" class="Reptilia" family="Colubridae" genus="Leptophis" kingdom="Animalia" order="Squamata" pageId="2" phylum="Chordata" rank="species" species="cupreus">
<emphasis id="B951437AFFD8FFA169CF4415FCFC9F62" box="[718,856,1852,1871]" italics="true" pageId="2">Leptophis cupreus</emphasis>
</taxonomicName>
(ANSP 5202).
</paragraph>
</caption>
<caption id="DF5ACFE0FFD9FFA06B5E4392FB119937" pageId="3" startId="3.[95,160,187,206]">
<paragraph id="8B9A9F68FFD9FFA06B5E4392FB119937" blockId="3.[94,1229,187,284]" pageId="3">
<emphasis id="B951437AFFD9FFA06B5E4392FF1E98E3" bold="true" box="[95,186,187,207]" pageId="3">TABLE 1:</emphasis>
Morphometric and meristic variation in the specimens of
<taxonomicName id="4C25E4EBFFD9FFA069BB4395FCE398E2" baseAuthorityName="Cope" baseAuthorityYear="1868" box="[698,839,188,207]" class="Reptilia" family="Colubridae" genus="Leptophis" kingdom="Animalia" order="Squamata" pageId="3" phylum="Chordata" rank="species" species="cupreus">
<emphasis id="B951437AFFD9FFA069BB4395FCE398E2" box="[698,839,188,207]" italics="true" pageId="3">Leptophis cupreus</emphasis>
</taxonomicName>
examined. Sl: Supralabials; SlO: Supralabials entering the orbit; Il: Infralabials; Ilg: Infralabials contacting the first genials; Pr/Pos: Preoculars/postoculars; Ta/Tp: Temporal anterior/ temporal posterior; P + VE: number of pre-ventral and ventral scales; SC: Number of subcaudal scales; HL: head length; SL: snout length; SVL: Snout-vent length; TL: Tail length; m: male; f: female; juv: juvenile. All measurements are in mm.*Holotype of
<taxonomicName id="4C25E4EBFFD9FFA06F254221FB159937" baseAuthorityName="Cope" baseAuthorityYear="1868" box="[1060,1201,264,283]" class="Reptilia" family="Colubridae" genus="Leptophis" kingdom="Animalia" order="Squamata" pageId="3" phylum="Chordata" rank="species" species="cupreus">Leptophis cupreus</taxonomicName>
.
</paragraph>
</caption>
<paragraph id="8B9A9F68FFD9FFA06B7D4213FB129B5F" pageId="3">
<table id="F9256DC8FFD9005C6B5F4213FB189B5F" box="[94,1212,314,883]" gridcols="13" gridrows="19" pageId="3">
<tr id="35159D2AFFD9005C6B5F4213FB189961" box="[94,1212,314,333]" gridrow="0" pageId="3">
<th id="76C4F456FFD9005C6B5F4213FEAE9961" box="[94,266,314,333]" gridcol="0" gridrow="0" pageId="3">Specimens (sex)</th>
<th id="76C4F456FFD9005C6A284213FEE29961" box="[297,326,314,333]" gridcol="1" gridrow="0" pageId="3">Sl</th>
<th id="76C4F456FFD9005C6A624213FE079961" box="[355,419,314,333]" gridcol="2" gridrow="0" pageId="3">SlO</th>
<th id="76C4F456FFD9005C6ABE4213FE559961" box="[447,497,314,333]" gridcol="3" gridrow="0" pageId="3">Il</th>
<th id="76C4F456FFD9005C69004213FD979961" box="[513,563,314,333]" gridcol="4" gridrow="0" pageId="3">Ilg</th>
<th id="76C4F456FFD9005C69474213FD229961" box="[582,646,314,333]" gridcol="5" gridrow="0" pageId="3">Pr/Pos</th>
<th id="76C4F456FFD9005C699D4213FD499961" box="[668,749,314,333]" gridcol="6" gridrow="0" pageId="3">Ta/Tp</th>
<th id="76C4F456FFD9005C68044213FCE59961" box="[773,833,314,333]" gridcol="7" gridrow="0" pageId="3">P + VE</th>
<th id="76C4F456FFD9005C68624213FC2A9961" box="[867,910,314,333]" gridcol="8" gridrow="0" pageId="3">SC</th>
<th id="76C4F456FFD9005C68AA4213FC789961" box="[939,988,314,333]" gridcol="9" gridrow="0" pageId="3">HL</th>
<th id="76C4F456FFD9005C68FD4213FB869961" box="[1020,1058,314,333]" gridcol="10" gridrow="0" pageId="3">SL</th>
<th id="76C4F456FFD9005C6F484213FBCA9961" box="[1097,1134,314,333]" gridcol="11" gridrow="0" pageId="3">SVL</th>
<th id="76C4F456FFD9005C6F904213FB189961" box="[1169,1212,314,333]" gridcol="12" gridrow="0" pageId="3">TL</th>
</tr>
<tr id="35159D2AFFD9005C6B5F4271FB189947" box="[94,1212,344,363]" gridrow="1" pageId="3">
<th id="76C4F456FFD9005C6B5F4271FEAE9947" box="[94,266,344,363]" gridcol="0" gridrow="1" pageId="3">ANSP 5202P* (f)</th>
<td id="76C4F456FFD9005C6A284271FEE29947" box="[297,326,344,363]" gridcol="1" gridrow="1" pageId="3">8/8</td>
<td id="76C4F456FFD9005C6A624271FE079947" box="[355,419,344,363]" gridcol="2" gridrow="1" pageId="3">4/5-4/5</td>
<td id="76C4F456FFD9005C6ABE4271FE559947" box="[447,497,344,363]" gridcol="3" gridrow="1" pageId="3">10/10</td>
<td id="76C4F456FFD9005C69004271FD979947" box="[513,563,344,363]" gridcol="4" gridrow="1" pageId="3">5/5</td>
<td id="76C4F456FFD9005C69474271FD229947" box="[582,646,344,363]" gridcol="5" gridrow="1" pageId="3">1/2-1/2</td>
<td id="76C4F456FFD9005C699D4271FD499947" box="[668,749,344,363]" gridcol="6" gridrow="1" pageId="3">1/2-1/2</td>
<td id="76C4F456FFD9005C68044271FCE59947" box="[773,833,344,363]" gridcol="7" gridrow="1" pageId="3">2+152</td>
<td id="76C4F456FFD9005C68624271FC2A9947" box="[867,910,344,363]" gridcol="8" gridrow="1" pageId="3">137</td>
<td id="76C4F456FFD9005C68AA4271FC789947" box="[939,988,344,363]" gridcol="9" gridrow="1" pageId="3">12.27</td>
<td id="76C4F456FFD9005C68FD4271FB869947" box="[1020,1058,344,363]" gridcol="10" gridrow="1" pageId="3">3.80</td>
<td id="76C4F456FFD9005C6F484271FBCA9947" box="[1097,1134,344,363]" gridcol="11" gridrow="1" pageId="3">313</td>
<td id="76C4F456FFD9005C6F904271FB189947" box="[1169,1212,344,363]" gridcol="12" gridrow="1" pageId="3">205</td>
</tr>
<tr id="35159D2AFFD9005C6B5F425EFB1899A6" box="[94,1212,375,394]" gridrow="2" pageId="3">
<th id="76C4F456FFD9005C6B5F425EFEAE99A6" box="[94,266,375,394]" gridcol="0" gridrow="2" pageId="3">ICN 0347 (f)</th>
<td id="76C4F456FFD9005C6A28425EFEE299A6" box="[297,326,375,394]" gridcol="1" gridrow="2" pageId="3">8/8</td>
<td id="76C4F456FFD9005C6A62425EFE0799A6" box="[355,419,375,394]" gridcol="2" gridrow="2" pageId="3">4/5-4/5</td>
<td id="76C4F456FFD9005C6ABE425EFE5599A6" box="[447,497,375,394]" gridcol="3" gridrow="2" pageId="3">10/10</td>
<td id="76C4F456FFD9005C6900425EFD9799A6" box="[513,563,375,394]" gridcol="4" gridrow="2" pageId="3">6/5</td>
<td id="76C4F456FFD9005C6947425EFD2299A6" box="[582,646,375,394]" gridcol="5" gridrow="2" pageId="3">1/2-1/2</td>
<td id="76C4F456FFD9005C699D425EFD4999A6" box="[668,749,375,394]" gridcol="6" gridrow="2" pageId="3">1/2-1/2</td>
<td id="76C4F456FFD9005C6804425EFCE599A6" box="[773,833,375,394]" gridcol="7" gridrow="2" pageId="3">2+161</td>
<td id="76C4F456FFD9005C6862425EFC2A99A6" box="[867,910,375,394]" gridcol="8" gridrow="2" pageId="3">50+</td>
<td id="76C4F456FFD9005C68AA425EFC7899A6" box="[939,988,375,394]" gridcol="9" gridrow="2" pageId="3">17.13</td>
<td id="76C4F456FFD9005C68FD425EFB8699A6" box="[1020,1058,375,394]" gridcol="10" gridrow="2" pageId="3">5.44</td>
<td id="76C4F456FFD9005C6F48425EFBCA99A6" box="[1097,1134,375,394]" gridcol="11" gridrow="2" pageId="3">491</td>
<td id="76C4F456FFD9005C6F90425EFB1899A6" box="[1169,1212,375,394]" gridcol="12" gridrow="2" pageId="3">142+</td>
</tr>
<tr id="35159D2AFFD9005C6B5F42BCFB189985" box="[94,1212,405,425]" gridrow="3" pageId="3">
<th id="76C4F456FFD9005C6B5F42BCFEAE9985" box="[94,266,405,425]" gridcol="0" gridrow="3" pageId="3">ICN 0390 (m)</th>
<td id="76C4F456FFD9005C6A2842BCFEE29985" box="[297,326,405,425]" gridcol="1" gridrow="3" pageId="3">9/9</td>
<td id="76C4F456FFD9005C6A6242BCFE079985" box="[355,419,405,425]" gridcol="2" gridrow="3" pageId="3">5/6-5/6</td>
<td id="76C4F456FFD9005C6ABE42BCFE559985" box="[447,497,405,425]" gridcol="3" gridrow="3" pageId="3">10/10</td>
<td id="76C4F456FFD9005C690042BCFD979985" box="[513,563,405,425]" gridcol="4" gridrow="3" pageId="3">5/5</td>
<td id="76C4F456FFD9005C694742BCFD229985" box="[582,646,405,425]" gridcol="5" gridrow="3" pageId="3">1/2-1/2</td>
<td id="76C4F456FFD9005C699D42BCFD499985" box="[668,749,405,425]" gridcol="6" gridrow="3" pageId="3">1/1-1/1</td>
<td id="76C4F456FFD9005C680442BCFCE59985" box="[773,833,405,425]" gridcol="7" gridrow="3" pageId="3">1+160</td>
<td id="76C4F456FFD9005C686242BCFC2A9985" box="[867,910,405,425]" gridcol="8" gridrow="3" pageId="3">146</td>
<td id="76C4F456FFD9005C68AA42BCFC789985" box="[939,988,405,425]" gridcol="9" gridrow="3" pageId="3">25.46</td>
<td id="76C4F456FFD9005C68FD42BCFB869985" box="[1020,1058,405,425]" gridcol="10" gridrow="3" pageId="3">8.08</td>
<td id="76C4F456FFD9005C6F4842BCFBCA9985" box="[1097,1134,405,425]" gridcol="11" gridrow="3" pageId="3">831</td>
<td id="76C4F456FFD9005C6F9042BCFB189985" box="[1169,1212,405,425]" gridcol="12" gridrow="3" pageId="3">495+</td>
</tr>
<tr id="35159D2AFFD9005C6B5F429DFB1899EB" box="[94,1212,436,455]" gridrow="4" pageId="3">
<th id="76C4F456FFD9005C6B5F429DFEAE99EB" box="[94,266,436,455]" gridcol="0" gridrow="4" pageId="3">ICN 8379 (f)</th>
<td id="76C4F456FFD9005C6A28429DFEE299EB" box="[297,326,436,455]" gridcol="1" gridrow="4" pageId="3">9/9</td>
<td id="76C4F456FFD9005C6A62429DFE0799EB" box="[355,419,436,455]" gridcol="2" gridrow="4" pageId="3">5/6-5/6</td>
<td id="76C4F456FFD9005C6ABE429DFE5599EB" box="[447,497,436,455]" gridcol="3" gridrow="4" pageId="3">11/11</td>
<td id="76C4F456FFD9005C6900429DFD9799EB" box="[513,563,436,455]" gridcol="4" gridrow="4" pageId="3">6/6</td>
<td id="76C4F456FFD9005C6947429DFD2299EB" box="[582,646,436,455]" gridcol="5" gridrow="4" pageId="3">1/2-1/2</td>
<td id="76C4F456FFD9005C699D429DFD4999EB" box="[668,749,436,455]" gridcol="6" gridrow="4" pageId="3">1/2-1/2</td>
<td id="76C4F456FFD9005C6804429DFCE599EB" box="[773,833,436,455]" gridcol="7" gridrow="4" pageId="3">2+171</td>
<td id="76C4F456FFD9005C6862429DFC2A99EB" box="[867,910,436,455]" gridcol="8" gridrow="4" pageId="3">148</td>
<td id="76C4F456FFD9005C68AA429DFC7899EB" box="[939,988,436,455]" gridcol="9" gridrow="4" pageId="3">22.94</td>
<td id="76C4F456FFD9005C68FD429DFB8699EB" box="[1020,1058,436,455]" gridcol="10" gridrow="4" pageId="3">7.14</td>
<td id="76C4F456FFD9005C6F48429DFBCA99EB" box="[1097,1134,436,455]" gridcol="11" gridrow="4" pageId="3">739</td>
<td id="76C4F456FFD9005C6F90429DFB1899EB" box="[1169,1212,436,455]" gridcol="12" gridrow="4" pageId="3">480</td>
</tr>
<tr id="35159D2AFFD9005C6B5F42FBFB1899CA" box="[94,1212,466,486]" gridrow="5" pageId="3">
<th id="76C4F456FFD9005C6B5F42FBFEAE99CA" box="[94,266,466,486]" gridcol="0" gridrow="5" pageId="3">ICN 8382 (f)</th>
<td id="76C4F456FFD9005C6A2842FBFEE299CA" box="[297,326,466,486]" gridcol="1" gridrow="5" pageId="3">9/9</td>
<td id="76C4F456FFD9005C6A6242FBFE0799CA" box="[355,419,466,486]" gridcol="2" gridrow="5" pageId="3">5/6-5/6</td>
<td id="76C4F456FFD9005C6ABE42FBFE5599CA" box="[447,497,466,486]" gridcol="3" gridrow="5" pageId="3">11/11</td>
<td id="76C4F456FFD9005C690042FBFD9799CA" box="[513,563,466,486]" gridcol="4" gridrow="5" pageId="3">6/6</td>
<td id="76C4F456FFD9005C694742FBFD2299CA" box="[582,646,466,486]" gridcol="5" gridrow="5" pageId="3">1/2-1/2</td>
<td id="76C4F456FFD9005C699D42FBFD4999CA" box="[668,749,466,486]" gridcol="6" gridrow="5" pageId="3">1/2-1/2</td>
<td id="76C4F456FFD9005C680442FBFCE599CA" box="[773,833,466,486]" gridcol="7" gridrow="5" pageId="3">2+170</td>
<td id="76C4F456FFD9005C686242FBFC2A99CA" box="[867,910,466,486]" gridcol="8" gridrow="5" pageId="3">166</td>
<td id="76C4F456FFD9005C68AA42FBFC7899CA" box="[939,988,466,486]" gridcol="9" gridrow="5" pageId="3">18.90</td>
<td id="76C4F456FFD9005C68FD42FBFB8699CA" box="[1020,1058,466,486]" gridcol="10" gridrow="5" pageId="3">5.95</td>
<td id="76C4F456FFD9005C6F4842FBFBCA99CA" box="[1097,1134,466,486]" gridcol="11" gridrow="5" pageId="3">559</td>
<td id="76C4F456FFD9005C6F9042FBFB1899CA" box="[1169,1212,466,486]" gridcol="12" gridrow="5" pageId="3">386</td>
</tr>
<tr id="35159D2AFFD9005C6B5F42D8FB189A28" box="[94,1212,497,516]" gridrow="6" pageId="3">
<th id="76C4F456FFD9005C6B5F42D8FEAE9A28" box="[94,266,497,516]" gridcol="0" gridrow="6" pageId="3">LACM 45444 (f)</th>
<td id="76C4F456FFD9005C6A2842D8FEE29A28" box="[297,326,497,516]" gridcol="1" gridrow="6" pageId="3">8/8</td>
<td id="76C4F456FFD9005C6A6242D8FE079A28" box="[355,419,497,516]" gridcol="2" gridrow="6" pageId="3">4/5-4/5</td>
<td id="76C4F456FFD9005C6ABE42D8FE559A28" box="[447,497,497,516]" gridcol="3" gridrow="6" pageId="3">10/10</td>
<td id="76C4F456FFD9005C690042D8FD979A28" box="[513,563,497,516]" gridcol="4" gridrow="6" pageId="3">5/5</td>
<td id="76C4F456FFD9005C694742D8FD229A28" box="[582,646,497,516]" gridcol="5" gridrow="6" pageId="3">1/2-1/2</td>
<td id="76C4F456FFD9005C699D42D8FD499A28" box="[668,749,497,516]" gridcol="6" gridrow="6" pageId="3">1/2-1/2</td>
<td id="76C4F456FFD9005C680442D8FCE59A28" box="[773,833,497,516]" gridcol="7" gridrow="6" pageId="3">2+130</td>
<td id="76C4F456FFD9005C686242D8FC2A9A28" box="[867,910,497,516]" gridcol="8" gridrow="6" pageId="3">123</td>
<td id="76C4F456FFD9005C68AA42D8FC789A28" box="[939,988,497,516]" gridcol="9" gridrow="6" pageId="3">15.9</td>
<td id="76C4F456FFD9005C68FD42D8FB869A28" box="[1020,1058,497,516]" gridcol="10" gridrow="6" pageId="3">5.0</td>
<td id="76C4F456FFD9005C6F4842D8FBCA9A28" box="[1097,1134,497,516]" gridcol="11" gridrow="6" pageId="3">345</td>
<td id="76C4F456FFD9005C6F9042D8FB189A28" box="[1169,1212,497,516]" gridcol="12" gridrow="6" pageId="3">236</td>
</tr>
<tr id="35159D2AFFD9005C6B5F4139FB189A0F" box="[94,1212,528,547]" gridrow="7" pageId="3">
<th id="76C4F456FFD9005C6B5F4139FEAE9A0F" box="[94,266,528,547]" gridcol="0" gridrow="7" pageId="3">LACM 76811 (f)</th>
<td id="76C4F456FFD9005C6A284139FEE29A0F" box="[297,326,528,547]" gridcol="1" gridrow="7" pageId="3">8/8</td>
<td id="76C4F456FFD9005C6A624139FE079A0F" box="[355,419,528,547]" gridcol="2" gridrow="7" pageId="3">4/5-4/5</td>
<td id="76C4F456FFD9005C6ABE4139FE559A0F" box="[447,497,528,547]" gridcol="3" gridrow="7" pageId="3">10/10</td>
<td id="76C4F456FFD9005C69004139FD979A0F" box="[513,563,528,547]" gridcol="4" gridrow="7" pageId="3">5/5</td>
<td id="76C4F456FFD9005C69474139FD229A0F" box="[582,646,528,547]" gridcol="5" gridrow="7" pageId="3">1/2-1/2</td>
<td id="76C4F456FFD9005C699D4139FD499A0F" box="[668,749,528,547]" gridcol="6" gridrow="7" pageId="3">1/2-1/2</td>
<td id="76C4F456FFD9005C68044139FCE59A0F" box="[773,833,528,547]" gridcol="7" gridrow="7" pageId="3">1+154</td>
<td id="76C4F456FFD9005C68624139FC2A9A0F" box="[867,910,528,547]" gridcol="8" gridrow="7" pageId="3">138</td>
<td id="76C4F456FFD9005C68AA4139FC789A0F" box="[939,988,528,547]" gridcol="9" gridrow="7" pageId="3">16.2</td>
<td id="76C4F456FFD9005C68FD4139FB869A0F" box="[1020,1058,528,547]" gridcol="10" gridrow="7" pageId="3">5.1</td>
<td id="76C4F456FFD9005C6F484139FBCA9A0F" box="[1097,1134,528,547]" gridcol="11" gridrow="7" pageId="3">395</td>
<td id="76C4F456FFD9005C6F904139FB189A0F" box="[1169,1212,528,547]" gridcol="12" gridrow="7" pageId="3">259</td>
</tr>
<tr id="35159D2AFFD9005C6B5F4107FB189A6E" box="[94,1212,558,578]" gridrow="8" pageId="3">
<th id="76C4F456FFD9005C6B5F4107FEAE9A6E" box="[94,266,558,578]" gridcol="0" gridrow="8" pageId="3">MCZ 166586 (m)</th>
<td id="76C4F456FFD9005C6A284107FEE29A6E" box="[297,326,558,578]" gridcol="1" gridrow="8" pageId="3">8/8</td>
<td id="76C4F456FFD9005C6A624107FE079A6E" box="[355,419,558,578]" gridcol="2" gridrow="8" pageId="3">4/5-4/5</td>
<td id="76C4F456FFD9005C6ABE4107FE559A6E" box="[447,497,558,578]" gridcol="3" gridrow="8" pageId="3">10/10</td>
<td id="76C4F456FFD9005C69004107FD979A6E" box="[513,563,558,578]" gridcol="4" gridrow="8" pageId="3">5/5</td>
<td id="76C4F456FFD9005C69474107FD229A6E" box="[582,646,558,578]" gridcol="5" gridrow="8" pageId="3">1/2-1/2</td>
<td id="76C4F456FFD9005C699D4107FD499A6E" box="[668,749,558,578]" gridcol="6" gridrow="8" pageId="3">1/2-1/2</td>
<td id="76C4F456FFD9005C68044107FCE59A6E" box="[773,833,558,578]" gridcol="7" gridrow="8" pageId="3">3+144</td>
<td id="76C4F456FFD9005C68624107FC2A9A6E" box="[867,910,558,578]" gridcol="8" gridrow="8" pageId="3">131+</td>
<td id="76C4F456FFD9005C68AA4107FC789A6E" box="[939,988,558,578]" gridcol="9" gridrow="8" pageId="3"></td>
<td id="76C4F456FFD9005C68FD4107FB869A6E" box="[1020,1058,558,578]" gridcol="10" gridrow="8" pageId="3"></td>
<td id="76C4F456FFD9005C6F484107FBCA9A6E" box="[1097,1134,558,578]" gridcol="11" gridrow="8" pageId="3">418</td>
<td id="76C4F456FFD9005C6F904107FB189A6E" box="[1169,1212,558,578]" gridcol="12" gridrow="8" pageId="3">274+</td>
</tr>
<tr id="35159D2AFFD9005C6B5F4164FB189A4C" box="[94,1212,589,608]" gridrow="9" pageId="3">
<th id="76C4F456FFD9005C6B5F4164FEAE9A4C" box="[94,266,589,608]" gridcol="0" gridrow="9" pageId="3">MCZ 164915 (m)</th>
<td id="76C4F456FFD9005C6A284164FEE29A4C" box="[297,326,589,608]" gridcol="1" gridrow="9" pageId="3">8/7</td>
<td id="76C4F456FFD9005C6A624164FE079A4C" box="[355,419,589,608]" gridcol="2" gridrow="9" pageId="3">4/5-4/5</td>
<td id="76C4F456FFD9005C6ABE4164FE559A4C" box="[447,497,589,608]" gridcol="3" gridrow="9" pageId="3">10/10</td>
<td id="76C4F456FFD9005C69004164FD979A4C" box="[513,563,589,608]" gridcol="4" gridrow="9" pageId="3">5/5</td>
<td id="76C4F456FFD9005C69474164FD229A4C" box="[582,646,589,608]" gridcol="5" gridrow="9" pageId="3">1/2-1/2</td>
<td id="76C4F456FFD9005C699D4164FD499A4C" box="[668,749,589,608]" gridcol="6" gridrow="9" pageId="3">1/2-1/2</td>
<td id="76C4F456FFD9005C68044164FCE59A4C" box="[773,833,589,608]" gridcol="7" gridrow="9" pageId="3">1+147</td>
<td id="76C4F456FFD9005C68624164FC2A9A4C" box="[867,910,589,608]" gridcol="8" gridrow="9" pageId="3">121+</td>
<td id="76C4F456FFD9005C68AA4164FC789A4C" box="[939,988,589,608]" gridcol="9" gridrow="9" pageId="3">16.76</td>
<td id="76C4F456FFD9005C68FD4164FB869A4C" box="[1020,1058,589,608]" gridcol="10" gridrow="9" pageId="3">5.31</td>
<td id="76C4F456FFD9005C6F484164FBCA9A4C" box="[1097,1134,589,608]" gridcol="11" gridrow="9" pageId="3">447</td>
<td id="76C4F456FFD9005C6F904164FB189A4C" box="[1169,1212,589,608]" gridcol="12" gridrow="9" pageId="3">282+</td>
</tr>
<tr id="35159D2AFFD9005C6B5F4142FB189A53" box="[94,1212,619,639]" gridrow="10" pageId="3">
<th id="76C4F456FFD9005C6B5F4142FEAE9A53" box="[94,266,619,639]" gridcol="0" gridrow="10" pageId="3">TCWC 42808 (f)</th>
<td id="76C4F456FFD9005C6A284142FEE29A53" box="[297,326,619,639]" gridcol="1" gridrow="10" pageId="3">8/8</td>
<td id="76C4F456FFD9005C6A624142FE079A53" box="[355,419,619,639]" gridcol="2" gridrow="10" pageId="3">4/5-4/5</td>
<td id="76C4F456FFD9005C6ABE4142FE559A53" box="[447,497,619,639]" gridcol="3" gridrow="10" pageId="3">—/10</td>
<td id="76C4F456FFD9005C69004142FD979A53" box="[513,563,619,639]" gridcol="4" gridrow="10" pageId="3">5/5</td>
<td id="76C4F456FFD9005C69474142FD229A53" box="[582,646,619,639]" gridcol="5" gridrow="10" pageId="3">1/2-1/2</td>
<td id="76C4F456FFD9005C699D4142FD499A53" box="[668,749,619,639]" gridcol="6" gridrow="10" pageId="3">1/1-1/1</td>
<td id="76C4F456FFD9005C68044142FCE59A53" box="[773,833,619,639]" gridcol="7" gridrow="10" pageId="3">1+153</td>
<td id="76C4F456FFD9005C68624142FC2A9A53" box="[867,910,619,639]" gridcol="8" gridrow="10" pageId="3">161+</td>
<td id="76C4F456FFD9005C68AA4142FC789A53" box="[939,988,619,639]" gridcol="9" gridrow="10" pageId="3">9.96</td>
<td id="76C4F456FFD9005C68FD4142FB869A53" box="[1020,1058,619,639]" gridcol="10" gridrow="10" pageId="3">2.97</td>
<td id="76C4F456FFD9005C6F484142FBCA9A53" box="[1097,1134,619,639]" gridcol="11" gridrow="10" pageId="3">185</td>
<td id="76C4F456FFD9005C6F904142FB189A53" box="[1169,1212,619,639]" gridcol="12" gridrow="10" pageId="3">57+</td>
</tr>
<tr id="35159D2AFFD9005C6B5F41A3FB189AB1" box="[94,1212,650,669]" gridrow="11" pageId="3">
<th id="76C4F456FFD9005C6B5F41A3FEAE9AB1" box="[94,266,650,669]" gridcol="0" gridrow="11" pageId="3">USNM 197281(juv)</th>
<td id="76C4F456FFD9005C6A2841A3FEE29AB1" box="[297,326,650,669]" gridcol="1" gridrow="11" pageId="3">8/8</td>
<td id="76C4F456FFD9005C6A6241A3FE079AB1" box="[355,419,650,669]" gridcol="2" gridrow="11" pageId="3">4/5-4/5</td>
<td id="76C4F456FFD9005C6ABE41A3FE559AB1" box="[447,497,650,669]" gridcol="3" gridrow="11" pageId="3">10/10</td>
<td id="76C4F456FFD9005C690041A3FD979AB1" box="[513,563,650,669]" gridcol="4" gridrow="11" pageId="3">5/5</td>
<td id="76C4F456FFD9005C694741A3FD229AB1" box="[582,646,650,669]" gridcol="5" gridrow="11" pageId="3">1/2-1/2</td>
<td id="76C4F456FFD9005C699D41A3FD499AB1" box="[668,749,650,669]" gridcol="6" gridrow="11" pageId="3">1/2-1/2</td>
<td id="76C4F456FFD9005C680441A3FCE59AB1" box="[773,833,650,669]" gridcol="7" gridrow="11" pageId="3">1+156</td>
<td id="76C4F456FFD9005C686241A3FC2A9AB1" box="[867,910,650,669]" gridcol="8" gridrow="11" pageId="3">142</td>
<td id="76C4F456FFD9005C68AA41A3FC789AB1" box="[939,988,650,669]" gridcol="9" gridrow="11" pageId="3">11.5</td>
<td id="76C4F456FFD9005C68FD41A3FB869AB1" box="[1020,1058,650,669]" gridcol="10" gridrow="11" pageId="3">3.7</td>
<td id="76C4F456FFD9005C6F4841A3FBCA9AB1" box="[1097,1134,650,669]" gridcol="11" gridrow="11" pageId="3">171</td>
<td id="76C4F456FFD9005C6F9041A3FB189AB1" box="[1169,1212,650,669]" gridcol="12" gridrow="11" pageId="3">107</td>
</tr>
<tr id="35159D2AFFD9005C6B5F4180FB189A90" box="[94,1212,681,700]" gridrow="12" pageId="3">
<th id="76C4F456FFD9005C6B5F4180FEAE9A90" box="[94,266,681,700]" gridcol="0" gridrow="12" pageId="3">USNM 211036 (m)</th>
<td id="76C4F456FFD9005C6A284180FEE29A90" box="[297,326,681,700]" gridcol="1" gridrow="12" pageId="3">8/8</td>
<td id="76C4F456FFD9005C6A624180FE079A90" box="[355,419,681,700]" gridcol="2" gridrow="12" pageId="3">4/5-4/5</td>
<td id="76C4F456FFD9005C6ABE4180FE559A90" box="[447,497,681,700]" gridcol="3" gridrow="12" pageId="3">10/10</td>
<td id="76C4F456FFD9005C69004180FD979A90" box="[513,563,681,700]" gridcol="4" gridrow="12" pageId="3">5/5</td>
<td id="76C4F456FFD9005C69474180FD229A90" box="[582,646,681,700]" gridcol="5" gridrow="12" pageId="3">1/2-1/2</td>
<td id="76C4F456FFD9005C699D4180FD499A90" box="[668,749,681,700]" gridcol="6" gridrow="12" pageId="3">1/2-1/2</td>
<td id="76C4F456FFD9005C68044180FCE59A90" box="[773,833,681,700]" gridcol="7" gridrow="12" pageId="3">2+133</td>
<td id="76C4F456FFD9005C68624180FC2A9A90" box="[867,910,681,700]" gridcol="8" gridrow="12" pageId="3">126</td>
<td id="76C4F456FFD9005C68AA4180FC789A90" box="[939,988,681,700]" gridcol="9" gridrow="12" pageId="3">15.3</td>
<td id="76C4F456FFD9005C68FD4180FB869A90" box="[1020,1058,681,700]" gridcol="10" gridrow="12" pageId="3">4.15</td>
<td id="76C4F456FFD9005C6F484180FBCA9A90" box="[1097,1134,681,700]" gridcol="11" gridrow="12" pageId="3">346</td>
<td id="76C4F456FFD9005C6F904180FB189A90" box="[1169,1212,681,700]" gridcol="12" gridrow="12" pageId="3">230</td>
</tr>
<tr id="35159D2AFFD9005C6B5F41EEFB189AF6" box="[94,1212,711,730]" gridrow="13" pageId="3">
<th id="76C4F456FFD9005C6B5F41EEFEAE9AF6" box="[94,266,711,730]" gridcol="0" gridrow="13" pageId="3">USNM 211037 (m)</th>
<td id="76C4F456FFD9005C6A2841EEFEE29AF6" box="[297,326,711,730]" gridcol="1" gridrow="13" pageId="3">8/8</td>
<td id="76C4F456FFD9005C6A6241EEFE079AF6" box="[355,419,711,730]" gridcol="2" gridrow="13" pageId="3">4/5-4/5</td>
<td id="76C4F456FFD9005C6ABE41EEFE559AF6" box="[447,497,711,730]" gridcol="3" gridrow="13" pageId="3">10/10</td>
<td id="76C4F456FFD9005C690041EEFD979AF6" box="[513,563,711,730]" gridcol="4" gridrow="13" pageId="3">6/6</td>
<td id="76C4F456FFD9005C694741EEFD229AF6" box="[582,646,711,730]" gridcol="5" gridrow="13" pageId="3">1/3-1/3</td>
<td id="76C4F456FFD9005C699D41EEFD499AF6" box="[668,749,711,730]" gridcol="6" gridrow="13" pageId="3">1/2-1/2/2</td>
<td id="76C4F456FFD9005C680441EEFCE59AF6" box="[773,833,711,730]" gridcol="7" gridrow="13" pageId="3">2+150</td>
<td id="76C4F456FFD9005C686241EEFC2A9AF6" box="[867,910,711,730]" gridcol="8" gridrow="13" pageId="3">143</td>
<td id="76C4F456FFD9005C68AA41EEFC789AF6" box="[939,988,711,730]" gridcol="9" gridrow="13" pageId="3">14.15</td>
<td id="76C4F456FFD9005C68FD41EEFB869AF6" box="[1020,1058,711,730]" gridcol="10" gridrow="13" pageId="3">3.65</td>
<td id="76C4F456FFD9005C6F4841EEFBCA9AF6" box="[1097,1134,711,730]" gridcol="11" gridrow="13" pageId="3">315</td>
<td id="76C4F456FFD9005C6F9041EEFB189AF6" box="[1169,1212,711,730]" gridcol="12" gridrow="13" pageId="3">206</td>
</tr>
<tr id="35159D2AFFD9005C6B5F41CFFB189AD5" box="[94,1212,742,761]" gridrow="14" pageId="3">
<th id="76C4F456FFD9005C6B5F41CFFEAE9AD5" box="[94,266,742,761]" gridcol="0" gridrow="14" pageId="3">USNM 211038 (f)</th>
<td id="76C4F456FFD9005C6A2841CFFEE29AD5" box="[297,326,742,761]" gridcol="1" gridrow="14" pageId="3">8/8</td>
<td id="76C4F456FFD9005C6A6241CFFE079AD5" box="[355,419,742,761]" gridcol="2" gridrow="14" pageId="3">4/5-4/5</td>
<td id="76C4F456FFD9005C6ABE41CFFE559AD5" box="[447,497,742,761]" gridcol="3" gridrow="14" pageId="3">10/10</td>
<td id="76C4F456FFD9005C690041CFFD979AD5" box="[513,563,742,761]" gridcol="4" gridrow="14" pageId="3">5/5</td>
<td id="76C4F456FFD9005C694741CFFD229AD5" box="[582,646,742,761]" gridcol="5" gridrow="14" pageId="3">1/2-1/2</td>
<td id="76C4F456FFD9005C699D41CFFD499AD5" box="[668,749,742,761]" gridcol="6" gridrow="14" pageId="3">1/21/2</td>
<td id="76C4F456FFD9005C680441CFFCE59AD5" box="[773,833,742,761]" gridcol="7" gridrow="14" pageId="3">1+147</td>
<td id="76C4F456FFD9005C686241CFFC2A9AD5" box="[867,910,742,761]" gridcol="8" gridrow="14" pageId="3">135</td>
<td id="76C4F456FFD9005C68AA41CFFC789AD5" box="[939,988,742,761]" gridcol="9" gridrow="14" pageId="3">17.95</td>
<td id="76C4F456FFD9005C68FD41CFFB869AD5" box="[1020,1058,742,761]" gridcol="10" gridrow="14" pageId="3">5.3</td>
<td id="76C4F456FFD9005C6F4841CFFBCA9AD5" box="[1097,1134,742,761]" gridcol="11" gridrow="14" pageId="3">408</td>
<td id="76C4F456FFD9005C6F9041CFFB189AD5" box="[1169,1212,742,761]" gridcol="12" gridrow="14" pageId="3">275</td>
</tr>
<tr id="35159D2AFFD9005C6B5F402DFB189B34" box="[94,1212,772,792]" gridrow="15" pageId="3">
<th id="76C4F456FFD9005C6B5F402DFEAE9B34" box="[94,266,772,792]" gridcol="0" gridrow="15" pageId="3">USNM 211039 (m)</th>
<td id="76C4F456FFD9005C6A28402DFEE29B34" box="[297,326,772,792]" gridcol="1" gridrow="15" pageId="3">8/8</td>
<td id="76C4F456FFD9005C6A62402DFE079B34" box="[355,419,772,792]" gridcol="2" gridrow="15" pageId="3">4/5-4/5</td>
<td id="76C4F456FFD9005C6ABE402DFE559B34" box="[447,497,772,792]" gridcol="3" gridrow="15" pageId="3">10/10</td>
<td id="76C4F456FFD9005C6900402DFD979B34" box="[513,563,772,792]" gridcol="4" gridrow="15" pageId="3">5/5</td>
<td id="76C4F456FFD9005C6947402DFD229B34" box="[582,646,772,792]" gridcol="5" gridrow="15" pageId="3">1/2-1/2</td>
<td id="76C4F456FFD9005C699D402DFD499B34" box="[668,749,772,792]" gridcol="6" gridrow="15" pageId="3">1/2-1/1</td>
<td id="76C4F456FFD9005C6804402DFCE59B34" box="[773,833,772,792]" gridcol="7" gridrow="15" pageId="3">1+155</td>
<td id="76C4F456FFD9005C6862402DFC2A9B34" box="[867,910,772,792]" gridcol="8" gridrow="15" pageId="3">135</td>
<td id="76C4F456FFD9005C68AA402DFC789B34" box="[939,988,772,792]" gridcol="9" gridrow="15" pageId="3">14.9</td>
<td id="76C4F456FFD9005C68FD402DFB869B34" box="[1020,1058,772,792]" gridcol="10" gridrow="15" pageId="3">4.7</td>
<td id="76C4F456FFD9005C6F48402DFBCA9B34" box="[1097,1134,772,792]" gridcol="11" gridrow="15" pageId="3">374</td>
<td id="76C4F456FFD9005C6F90402DFB189B34" box="[1169,1212,772,792]" gridcol="12" gridrow="15" pageId="3">230</td>
</tr>
<tr id="35159D2AFFD9005C6B5F400AFB189B1A" box="[94,1212,803,822]" gridrow="16" pageId="3">
<th id="76C4F456FFD9005C6B5F400AFEAE9B1A" box="[94,266,803,822]" gridcol="0" gridrow="16" pageId="3">USNM 211041 (m)</th>
<td id="76C4F456FFD9005C6A28400AFEE29B1A" box="[297,326,803,822]" gridcol="1" gridrow="16" pageId="3">8/8</td>
<td id="76C4F456FFD9005C6A62400AFE079B1A" box="[355,419,803,822]" gridcol="2" gridrow="16" pageId="3">4/5-4/5</td>
<td id="76C4F456FFD9005C6ABE400AFE559B1A" box="[447,497,803,822]" gridcol="3" gridrow="16" pageId="3">10/10</td>
<td id="76C4F456FFD9005C6900400AFD979B1A" box="[513,563,803,822]" gridcol="4" gridrow="16" pageId="3">6/5</td>
<td id="76C4F456FFD9005C6947400AFD229B1A" box="[582,646,803,822]" gridcol="5" gridrow="16" pageId="3">1/2-1/2</td>
<td id="76C4F456FFD9005C699D400AFD499B1A" box="[668,749,803,822]" gridcol="6" gridrow="16" pageId="3">2/2-1/2</td>
<td id="76C4F456FFD9005C6804400AFCE59B1A" box="[773,833,803,822]" gridcol="7" gridrow="16" pageId="3">2+147</td>
<td id="76C4F456FFD9005C6862400AFC2A9B1A" box="[867,910,803,822]" gridcol="8" gridrow="16" pageId="3">145</td>
<td id="76C4F456FFD9005C68AA400AFC789B1A" box="[939,988,803,822]" gridcol="9" gridrow="16" pageId="3">16.6</td>
<td id="76C4F456FFD9005C68FD400AFB869B1A" box="[1020,1058,803,822]" gridcol="10" gridrow="16" pageId="3">5.05</td>
<td id="76C4F456FFD9005C6F48400AFBCA9B1A" box="[1097,1134,803,822]" gridcol="11" gridrow="16" pageId="3">443</td>
<td id="76C4F456FFD9005C6F90400AFB189B1A" box="[1169,1212,803,822]" gridcol="12" gridrow="16" pageId="3">295</td>
</tr>
<tr id="35159D2AFFD9005C6B5F406BFB189B79" box="[94,1212,834,853]" gridrow="17" pageId="3">
<th id="76C4F456FFD9005C6B5F406BFEAE9B79" box="[94,266,834,853]" gridcol="0" gridrow="17" pageId="3">USNM 211042 (m)</th>
<td id="76C4F456FFD9005C6A28406BFEE29B79" box="[297,326,834,853]" gridcol="1" gridrow="17" pageId="3">8/8</td>
<td id="76C4F456FFD9005C6A62406BFE079B79" box="[355,419,834,853]" gridcol="2" gridrow="17" pageId="3">4/5-4/5</td>
<td id="76C4F456FFD9005C6ABE406BFE559B79" box="[447,497,834,853]" gridcol="3" gridrow="17" pageId="3">—/—</td>
<td id="76C4F456FFD9005C6900406BFD979B79" box="[513,563,834,853]" gridcol="4" gridrow="17" pageId="3">—/—</td>
<td id="76C4F456FFD9005C6947406BFD229B79" box="[582,646,834,853]" gridcol="5" gridrow="17" pageId="3">1/2-1/2</td>
<td id="76C4F456FFD9005C699D406BFD499B79" box="[668,749,834,853]" gridcol="6" gridrow="17" pageId="3">1/2-1/2</td>
<td id="76C4F456FFD9005C6804406BFCE59B79" box="[773,833,834,853]" gridcol="7" gridrow="17" pageId="3">1+157</td>
<td id="76C4F456FFD9005C6862406BFC2A9B79" box="[867,910,834,853]" gridcol="8" gridrow="17" pageId="3">137+</td>
<td id="76C4F456FFD9005C68AA406BFC789B79" box="[939,988,834,853]" gridcol="9" gridrow="17" pageId="3">13.6</td>
<td id="76C4F456FFD9005C68FD406BFB869B79" box="[1020,1058,834,853]" gridcol="10" gridrow="17" pageId="3">4.15</td>
<td id="76C4F456FFD9005C6F48406BFBCA9B79" box="[1097,1134,834,853]" gridcol="11" gridrow="17" pageId="3">335</td>
<td id="76C4F456FFD9005C6F90406BFB189B79" box="[1169,1212,834,853]" gridcol="12" gridrow="17" pageId="3">228+</td>
</tr>
<tr id="35159D2AFFD9005C6B5F4049FB189B5F" box="[94,1212,864,883]" gridrow="18" pageId="3">
<th id="76C4F456FFD9005C6B5F4049FEAE9B5F" box="[94,266,864,883]" gridcol="0" gridrow="18" pageId="3">USNM 562696 (f)</th>
<td id="76C4F456FFD9005C6A284049FEE29B5F" box="[297,326,864,883]" gridcol="1" gridrow="18" pageId="3">8/8</td>
<td id="76C4F456FFD9005C6A624049FE079B5F" box="[355,419,864,883]" gridcol="2" gridrow="18" pageId="3">4/5-4/5</td>
<td id="76C4F456FFD9005C6ABE4049FE559B5F" box="[447,497,864,883]" gridcol="3" gridrow="18" pageId="3">10/10</td>
<td id="76C4F456FFD9005C69004049FD979B5F" box="[513,563,864,883]" gridcol="4" gridrow="18" pageId="3">5/6</td>
<td id="76C4F456FFD9005C69474049FD229B5F" box="[582,646,864,883]" gridcol="5" gridrow="18" pageId="3">1/2-1/2</td>
<td id="76C4F456FFD9005C699D4049FD499B5F" box="[668,749,864,883]" gridcol="6" gridrow="18" pageId="3">1/2-1/2</td>
<td id="76C4F456FFD9005C68044049FCE59B5F" box="[773,833,864,883]" gridcol="7" gridrow="18" pageId="3">3+154</td>
<td id="76C4F456FFD9005C68624049FC2A9B5F" box="[867,910,864,883]" gridcol="8" gridrow="18" pageId="3">146</td>
<td id="76C4F456FFD9005C68AA4049FC789B5F" box="[939,988,864,883]" gridcol="9" gridrow="18" pageId="3">17.80</td>
<td id="76C4F456FFD9005C68FD4049FB869B5F" box="[1020,1058,864,883]" gridcol="10" gridrow="18" pageId="3">5.66</td>
<td id="76C4F456FFD9005C6F484049FBCA9B5F" box="[1097,1134,864,883]" gridcol="11" gridrow="18" pageId="3">490</td>
<td id="76C4F456FFD9005C6F904049FB189B5F" box="[1169,1212,864,883]" gridcol="12" gridrow="18" pageId="3">331</td>
</tr>
</table>
</paragraph>
<paragraph id="8B9A9F68FFD9FFA068554611FB859D7C" blockId="3.[852,1057,1336,1360]" box="[852,1057,1336,1360]" pageId="3">
<emphasis id="B951437AFFD9FFA068554611FB859D7C" bold="true" box="[852,1057,1336,1360]" pageId="3">
Hemipenis (
<figureCitation id="131E83EDFFD9FFA068D94611FBBD9D7C" box="[984,1049,1336,1360]" captionStart="FIGURE 4" captionStartId="4.[132,212,1851,1870]" captionTargetBox="[132,1266,681,1815]" captionTargetId="figure-179@4.[132,1266,681,1815]" captionTargetPageId="4" captionText="FIGURE 4: Leptophis cupreus. Left hemipenis of ICN 390: (A) sulcate and (B) asulcate sides." figureDoi="http://doi.org/10.5281/zenodo.10085974" httpUri="https://zenodo.org/record/10085974/files/figure.png" pageId="3">Fig. 4</figureCitation>
)
</emphasis>
</paragraph>
<paragraph id="8B9A9F68FFD9FFA769E04651FEAE993C" blockId="3.[680,1229,1400,1872]" lastBlockId="4.[132,680,184,272]" lastPageId="4" pageId="3">
Left retracted organ extends for length of 7 subcaudals. Everted hemipenis single, noncapitate;
<emphasis id="B951437AFFD9FFA06F9546B1FCBD9DFC" italics="true" pageId="3">sulcus spermaticus</emphasis>
undivided, intrasulcar surface smooth. Basal region bearing numerous spines, distributed in five rows; first row with six spines; spines on first row larger than those in other rows; two spines adjacent to
<emphasis id="B951437AFFD9FFA069A94511FD449E7C" box="[680,736,1592,1616]" italics="true" pageId="3">sulcus</emphasis>
largest. Spines arranged irregularly rather than in transverse rows. Few spinules present, occurring in area adjacent to
<emphasis id="B951437AFFD9FFA068524551FC369EBC" box="[851,914,1656,1680]" italics="true" pageId="3">sulcus.</emphasis>
Small number of papillate calyces with fringing papillae occurs above fifth row of basal spines; papillae decrease in length and number distally and become stouter, as calyces increase in size. Seven papillae occur on calyces in middle of organ, 6-5 between middle of organ and proximal region of lobe, and
<quantity id="4CDD328DFFD9FFA068144411FCE69F7C" box="[789,834,1848,1872]" metricMagnitude="-1" metricUnit="m" metricValue="1.016" pageId="3" unit="in" value="4.0">4 in</quantity>
proximal region. Lobe is completely calyculate. Proximal region of lobe has few, irregularly distributed, papillate calyces. Sulcate side is similar to asulcate side.
</paragraph>
<caption id="DF5ACFE0FFD9FFA06B5E440BFE429F63" ID-DOI="http://doi.org/10.5281/zenodo.10085972" ID-Zenodo-Dep="10085972" httpUri="https://zenodo.org/record/10085972/files/figure.png" pageId="3" startId="3.[95,175,1826,1845]" targetBox="[94,642,1487,1789]" targetPageId="3">
<paragraph id="8B9A9F68FFD9FFA06B5E440BFE429F63" blockId="3.[94,643,1826,1871]" pageId="3">
<emphasis id="B951437AFFD9FFA06B5E440BFF679F19" bold="true" box="[95,195,1826,1845]" pageId="3">FIGURE 3:</emphasis>
<taxonomicName id="4C25E4EBFFD9FFA06BC6440BFEF49F19" baseAuthorityName="Cope" baseAuthorityYear="1868" box="[199,336,1826,1845]" class="Reptilia" family="Colubridae" genus="Leptophis" kingdom="Animalia" order="Squamata" pageId="3" phylum="Chordata" rank="species" species="cupreus">
<emphasis id="B951437AFFD9FFA06BC6440BFEF49F19" box="[199,336,1826,1845]" italics="true" pageId="3">Leptophis cupreus</emphasis>
</taxonomicName>
(LACM 45444).Lateral view of head showing the short and diffuse postocular stripe.
</paragraph>
</caption>
<paragraph id="8B9A9F68FFDEFFA76A614271FE68995C" blockId="4.[352,460,344,368]" box="[352,460,344,368]" pageId="4">
<emphasis id="B951437AFFDEFFA76A614271FE68995C" bold="true" box="[352,460,344,368]" pageId="4">Dentition</emphasis>
</paragraph>
<paragraph id="8B9A9F68FFDEFFA76BBC42B1FF6D9A5C" blockId="4.[132,681,408,624]" pageId="4">ICN 390 has 25/25 recurved maxillary teeth without a diastema, 15/14 palatine teeth, 26/25 pterygoid teeth, and 29/30 dentary teeth. Maxillary teeth increase in size posteriorly. Last three maxillary teeth are ungrooved and enlarged. One specimen from the Iquitos region has 21 maxillary teeth (Dixon &amp; Soini, 1977).</paragraph>
</subSubSection>
<subSubSection id="C33FCCE3FFDEFFA668B44391FF5E997C" lastPageId="5" pageId="4" type="biology_ecology">
<paragraph id="8B9A9F68FFDEFFA768B44391FBA898FC" blockId="4.[949,1036,184,208]" box="[949,1036,184,208]" pageId="4">
<emphasis id="B951437AFFDEFFA768B44391FBA898FC" bold="true" box="[949,1036,184,208]" pageId="4">Ecology</emphasis>
</paragraph>
<paragraph id="8B9A9F68FFDEFFA6680643D1FF5E997C" blockId="4.[718,1267,248,624]" lastBlockId="5.[94,642,184,336]" lastPageId="5" pageId="4">
Little is known about the ecology of
<emphasis id="B951437AFFDEFFA76F8443D0FB57993C" box="[1157,1267,249,272]" italics="true" pageId="4">
<taxonomicName id="4C25E4EBFFDEFFA76F8443D0FB4B993C" baseAuthorityName="Cope" baseAuthorityYear="1868" box="[1157,1263,249,272]" class="Reptilia" family="Colubridae" genus="Leptophis" kingdom="Animalia" order="Squamata" pageId="4" phylum="Chordata" rank="species" species="cupreus">L. cupreus</taxonomicName>
.
</emphasis>
Dixon &amp; Soini (1977) collected
<specimenCount id="9D2354E1FFDEFFA76F374231FB77991C" box="[1078,1235,280,304]" pageId="4" type="generic">two specimens</specimenCount>
in the leaf litter of primary forest at Yanamomo,
<collectingCountry id="F332DFF8FFDEFFA76FBD4211FB4B9963" box="[1212,1263,312,335]" name="Peru" pageId="4">Peru</collectingCountry>
. USNM 562696 was collected at
<quantity id="4CDD328DFFDEFFA76F344271FB2E9943" box="[1077,1162,344,368]" metricMagnitude="3" metricUnit="m" metricValue="1.88" pageId="4" unit="m" value="1880.0">1880 m</quantity>
elevation on Cerro de la Neblina, substantially higher than other records of the genus. It was first seen as it searched among the leaf rosettes of
<emphasis id="B951437AFFDEFFA76F3E4291FB5699FC" box="[1087,1266,440,464]" italics="true" pageId="4">
<taxonomicName id="4C25E4EBFFDEFFA76F3E4291FB4A99FC" authorityName="Maguire" authorityYear="1972" box="[1087,1262,440,464]" class="Magnoliopsida" family="Bonnetiaceae" genus="Neblinaria" higherTaxonomySource="CoL" kingdom="Plantae" order="Malpighiales" pageId="4" phylum="Tracheophyta" rank="species" species="celiae">Neblinaria celiae</taxonomicName>
,
</emphasis>
a bizarre plant restricted to the higher elevations of Cerro de le Neblina (
<bibRefCitation id="EFB4E299FFDEFFA768B142D1FB2C9A3C" author="GIVNISH" box="[944,1160,504,528]" pageId="4" pagination="481 - 485" refId="ref5196" refString="GIVNISH, T. J.; MCDIARMID, R. W. &amp; BUCK, W. R. 1986. Fire adaptation in Neblinaria celiae (Theaceae), a high-elevation rosette shrub endemic to a wet equatorial tepui. Oecologia, Berlin, 70: 481 - 485." type="journal article" year="1986">
Givnish
<emphasis id="B951437AFFDEFFA76F0D42D0FBE29A3C" box="[1036,1094,504,528]" italics="true" pageId="4">et al.,</emphasis>
1986
</bibRefCitation>
); presumably the snake was hunting frogs that frequently hide in the leafy rosettes that can hold up to 100 ml of rainwater. According to notes accompanying
<specimenCount id="9D2354E1FFDEFFA66FCD4171FF6498FC" lastPageId="5" pageId="4" type="generic">one specimen</specimenCount>
(USNM 211042), a frog of the genus
<taxonomicName id="4C25E4EBFFDFFFA66B5F43F1FEAA98DC" authorityName="Dumeril &amp; Bibron" authorityYear="1841" box="[94,270,216,240]" class="Amphibia" family="Eleutherodactylidae" genus="Eleutherodactylus" kingdom="Animalia" order="Anura" pageId="5" phylum="Chordata" rank="genus">
<emphasis id="B951437AFFDFFFA66B5F43F1FEAA98DC" box="[94,270,216,240]" italics="true" pageId="5">Eleutherodactylus</emphasis>
</taxonomicName>
(=
<taxonomicName id="4C25E4EBFFDFFFA66A3543F0FE0B98DC" box="[308,431,217,240]" class="Amphibia" family="Strabomantidae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="5" phylum="Chordata" rank="genus">
<emphasis id="B951437AFFDFFFA66A3543F0FE0B98DC" box="[308,431,217,240]" italics="true" pageId="5">Pristimantis</emphasis>
</taxonomicName>
) was removed from its stomach. Another specimen (USNM 211038) collected in March, 1956 contained two large ova in the oviduct.
</paragraph>
</subSubSection>
<caption id="DF5ACFE0FFDEFFA76B854412FC229F63" ID-DOI="http://doi.org/10.5281/zenodo.10085974" ID-Zenodo-Dep="10085974" box="[132,902,1851,1871]" httpUri="https://zenodo.org/record/10085974/files/figure.png" pageId="4" startId="4.[132,212,1851,1870]" targetBox="[132,1266,681,1815]" targetPageId="4">
<paragraph id="8B9A9F68FFDEFFA76B854412FC229F63" blockId="4.[132,902,1851,1871]" box="[132,902,1851,1871]" pageId="4">
<emphasis id="B951437AFFDEFFA76B854412FF4E9F62" bold="true" box="[132,234,1851,1870]" pageId="4">FIGURE 4:</emphasis>
<emphasis id="B951437AFFDEFFA76BEE4415FE249F62" box="[239,384,1852,1871]" italics="true" pageId="4">
<taxonomicName id="4C25E4EBFFDEFFA76BEE4415FED89F62" baseAuthorityName="Cope" baseAuthorityYear="1868" box="[239,380,1852,1871]" class="Reptilia" family="Colubridae" genus="Leptophis" kingdom="Animalia" order="Squamata" pageId="4" phylum="Chordata" rank="species" species="cupreus">Leptophis cupreus</taxonomicName>
.
</emphasis>
Left hemipenis of ICN 390:
<emphasis id="B951437AFFDEFFA769714412FD289F62" bold="true" box="[624,652,1851,1870]" pageId="4">(A)</emphasis>
sulcate and
<emphasis id="B951437AFFDEFFA769F34412FCAA9F62" bold="true" box="[754,782,1851,1871]" pageId="4">(B)</emphasis>
asulcate sides.
</paragraph>
</caption>
<subSubSection id="C33FCCE3FFDFFFA66A2D42B1FF159AFC" pageId="5" type="distribution">
<paragraph id="8B9A9F68FFDFFFA66A2D42B1FE11999C" blockId="5.[300,437,408,432]" box="[300,437,408,432]" pageId="5">
<emphasis id="B951437AFFDFFFA66A2D42B1FE11999C" bold="true" box="[300,437,408,432]" pageId="5">Distribution</emphasis>
</paragraph>
<paragraph id="8B9A9F68FFDFFFA66B9642F1FF159AFC" blockId="5.[94,643,472,720]" pageId="5">
Though apparently rare, or at least rarely encountered,
<taxonomicName id="4C25E4EBFFDFFFA66BD042D0FE909A3C" baseAuthorityName="Cope" baseAuthorityYear="1868" box="[209,308,505,528]" class="Reptilia" family="Colubridae" genus="Leptophis" kingdom="Animalia" order="Squamata" pageId="5" phylum="Chordata" rank="species" species="cupreus">
<emphasis id="B951437AFFDFFFA66BD042D0FE909A3C" box="[209,308,505,528]" italics="true" pageId="5">L. cupreus</emphasis>
</taxonomicName>
appears to be widely distributed. It is known from the southwestern Guayana Highlands of
<collectingCountry id="F332DFF8FFDFFFA66BB94111FEBA9A7C" box="[184,286,568,592]" name="Venezuela" pageId="5">Venezuela</collectingCountry>
and adjacent
<collectingCountry id="F332DFF8FFDFFFA66AB04111FDBC9A7C" box="[433,536,568,592]" name="Colombia" pageId="5">Colombia</collectingCountry>
(Sierra de La Macarena), the Amazonian lowlands of
<collectingCountry id="F332DFF8FFDFFFA669294171FDDB9A5C" box="[552,639,600,624]" name="Ecuador" pageId="5">Ecuador</collectingCountry>
,
<collectingCountry id="F332DFF8FFDFFFA66B5F4151FF619ABC" box="[94,197,632,656]" name="Colombia" pageId="5">Colombia</collectingCountry>
and
<collectingCountry id="F332DFF8FFDFFFA66BFE4151FE949AA3" box="[255,304,632,655]" name="Peru" pageId="5">Peru</collectingCountry>
, and from two localities on the Pacific versant of the Andes in
<collectingCountry id="F332DFF8FFDFFFA66A9541B1FE5F9A9C" box="[404,507,664,688]" name="Colombia" pageId="5">Colombia</collectingCountry>
and
<collectingCountry id="F332DFF8FFDFFFA6692C41B1FD279A9C" box="[557,643,664,688]" name="Ecuador" pageId="5">Ecuador</collectingCountry>
(
<figureCitation id="131E83EDFFDFFFA66B674191FF009AFC" box="[102,164,696,720]" captionStart="FIGURE 5" captionStartId="5.[95,175,1826,1845]" captionText="FIGURE 5: Map of northern South America showing the distribution of Leptophis cupreus." pageId="5">Fig. 5</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C33FCCE3FFDFFFA66B734031FB3699BC" pageId="5" type="diagnosis">
<paragraph id="8B9A9F68FFDFFFA66B734031FDCB9B1C" blockId="5.[114,623,792,816]" box="[114,623,792,816]" pageId="5">
<emphasis id="B951437AFFDFFFA66B734031FDCB9B1C" bold="true" box="[114,623,792,816]" pageId="5">Diagnosis and comparison with similar species</emphasis>
</paragraph>
<paragraph id="8B9A9F68FFDFFFA66B964071FB3699BC" blockId="5.[94,643,856,1296]" lastBlockId="5.[680,1229,184,400]" pageId="5">
<taxonomicName id="4C25E4EBFFDFFFA66B964071FEE19B5C" baseAuthorityName="Cope" baseAuthorityYear="1868" box="[151,325,856,880]" class="Reptilia" family="Colubridae" genus="Leptophis" kingdom="Animalia" order="Squamata" pageId="5" phylum="Chordata" rank="species" species="cupreus">
<emphasis id="B951437AFFDFFFA66B964071FEE19B5C" box="[151,325,856,880]" italics="true" pageId="5">Leptophis cupreus</emphasis>
</taxonomicName>
is distinguished from its congeners by adults having a uniformly copper dorsum (
<figureCitation id="131E83EDFFDFFFA66B6740B1FF019B9C" box="[102,165,920,944]" captionStart="FIGURE 6" captionStartId="5.[680,760,1800,1819]" captionTargetBox="[94,1228,1342,1790]" captionTargetPageId="5" captionText="FIGURE 6: Leptophis cupreus. A specimen without locality data photographed at the Valencia Aquarium, Valencia, Venezuela. Courtesy of Marco Natera Mumaw." figureDoi="http://doi.org/10.5281/zenodo.10085976" httpUri="https://zenodo.org/record/10085976/files/figure.png" pageId="5">Fig. 6</figureCitation>
) (vs. dorsum uniformly green, dorsum green with each dorsal scale edged with black, or middorsal area green or bronze contrasting in colour with the posterior area of trunk). The posterior venter is also coppery but slightly darker than the dorsum, and has dark brown and white streaks. Further,
<taxonomicName id="4C25E4EBFFDFFFA6691F4710FD269C7C" baseAuthorityName="Cope" baseAuthorityYear="1868" box="[542,642,1081,1104]" class="Reptilia" family="Colubridae" genus="Leptophis" kingdom="Animalia" order="Squamata" pageId="5" phylum="Chordata" rank="species" species="cupreus">
<emphasis id="B951437AFFDFFFA6691F4710FD269C7C" box="[542,642,1081,1104]" italics="true" pageId="5">L. cupreus</emphasis>
</taxonomicName>
differs from the occasionally sympatric
<emphasis id="B951437AFFDFFFA669064770FDF79CBC" italics="true" pageId="5">
<taxonomicName id="4C25E4EBFFDFFFA669064770FF189CBC" authority="ahaetulla" class="Reptilia" family="Colubridae" genus="Leptophis" kingdom="Animalia" order="Squamata" pageId="5" phylum="Chordata" rank="subSpecies" species="ahaetulla" subSpecies="ahaetulla">L. ahaetulla ahaetulla</taxonomicName>
, L. a.
<taxonomicName id="4C25E4EBFFDFFFA66A054750FE0D9CBC" authorityName="Boulenger" authorityYear="1894" box="[260,425,1145,1168]" class="Reptilia" family="Colubridae" genus="Leptophis" kingdom="Animalia" order="Squamata" pageId="5" phylum="Chordata" rank="species" species="nigromarginatus">nigromarginatus</taxonomicName>
, L. a.
<taxonomicName id="4C25E4EBFFDFFFA66AF14750FDEB9CBC" authorityName="Boulenger" authorityYear="1894" box="[496,591,1145,1168]" class="Reptilia" family="Colubridae" genus="Leptophis" kingdom="Animalia" order="Squamata" pageId="5" phylum="Chordata" rank="species" species="urostictus">urostictus</taxonomicName>
,
</emphasis>
and
<taxonomicName id="4C25E4EBFFDFFFA66B5F47B0FF0B9C9C" authorityName="Despax" authorityYear="1910" box="[94,175,1177,1200]" class="Reptilia" family="Colubridae" genus="Leptophis" kingdom="Animalia" order="Squamata" pageId="5" phylum="Chordata" rank="species" species="riveti">
<emphasis id="B951437AFFDFFFA66B5F47B0FF0B9C9C" box="[94,175,1177,1200]" italics="true" pageId="5">L. riveti</emphasis>
</taxonomicName>
(
<emphasis id="B951437AFFDFFFA66BBF47B0FF739C9C" box="[190,215,1177,1200]" italics="true" pageId="5">see</emphasis>
<bibRefCitation id="EFB4E299FFDFFFA66BDF47B1FEC79C9C" author="OLIVER" box="[222,355,1176,1200]" pageId="5" pagination="157 - 280" refId="ref5641" refString="OLIVER, J. A. 1948. The relationships and zoogeography of the genus Thalerophis Oliver. Bulletin of the American Museum of Natural History, 92: 157 - 280." type="journal article" year="1948">Oliver, 1948</bibRefCitation>
;
<bibRefCitation id="EFB4E299FFDFFFA66A6F47B1FD9E9C9C" author="ALBUQUERQUE" box="[366,570,1176,1200]" pageId="5" refId="ref4675" refString="ALBUQUERQUE, N. R. 2008. Revisao taxonomica das subespecies de Leptophis ahaetulla (Linnaeus, 1758) (Serpentes, Colubridae). (Tese de Doutorado). Pontificia Universidade Catolica do Rio Grande do Sul, Porto Alegre." type="book" year="2008">Albuquerque, 2008</bibRefCitation>
,
<bibRefCitation id="EFB4E299FFDFFFA6694447B1FDD89C9C" author="ALBUQUERQUE" box="[581,636,1176,1200]" pageId="5" pagination="293 - 297" refId="ref4719" refString="ALBUQUERQUE, N. R. 2009. New records of Leptophis ahaetulla ahaetulla (Serpentes, Colubridae) for Venezuela, Colombia and the placement of L. a. copei into the synonymy of L. a. ahaetulla. Biota Neotropica, 9: 293 - 297." type="journal article" year="2009">2009</bibRefCitation>
) by the absence of black spots in the center of each parietal scale (vs. present in
<emphasis id="B951437AFFDFFFA66A9547F0FDD09CDC" box="[404,628,1241,1264]" italics="true" pageId="5">
L. a.
<taxonomicName id="4C25E4EBFFDFFFA66AD347F0FDD09CDC" authorityName="Boulenger" authorityYear="1894" box="[466,628,1241,1264]" class="Reptilia" family="Colubridae" genus="Leptophis" kingdom="Animalia" order="Squamata" pageId="5" phylum="Chordata" rank="species" species="nigromarginatus">nigromarginatus</taxonomicName>
</emphasis>
); scales on dorsal surface of head not edged with black (vs. edged in
<emphasis id="B951437AFFDFFFA668364390FBB798FC" box="[823,1043,185,208]" italics="true" pageId="5">
L. a.
<taxonomicName id="4C25E4EBFFDFFFA668704390FBB798FC" authorityName="Boulenger" authorityYear="1894" box="[881,1043,185,208]" class="Reptilia" family="Colubridae" genus="Leptophis" kingdom="Animalia" order="Squamata" pageId="5" phylum="Chordata" rank="species" species="nigromarginatus">nigromarginatus</taxonomicName>
</emphasis>
and
<emphasis id="B951437AFFDFFFA66F4B4390FD6098DC" italics="true" pageId="5">
L. a.
<taxonomicName id="4C25E4EBFFDFFFA66F854390FD6098DC" authorityName="Boulenger" authorityYear="1894" class="Reptilia" family="Colubridae" genus="Leptophis" kingdom="Animalia" order="Squamata" pageId="5" phylum="Chordata" rank="species" species="urostictus">urostictus</taxonomicName>
</emphasis>
); dorsum unstriped in adults (vs. two dorsolateral stripes separated from each other by a pale vertebral stripe in
<emphasis id="B951437AFFDFFFA668074230FC3D991C" box="[774,921,280,304]" italics="true" pageId="5">
L. a.
<taxonomicName id="4C25E4EBFFDFFFA6683F4231FC3D991C" box="[830,921,280,304]" class="Reptilia" family="Colubridae" genus="Leptophis" kingdom="Animalia" order="Squamata" pageId="5" phylum="Chordata" rank="species" species="ahaetulla">ahaetulla</taxonomicName>
</emphasis>
), adult color pattern without dark oblique bands (vs. with dark bands in
<taxonomicName id="4C25E4EBFFDFFFA66F6C4210FB19997C" authorityName="Despax" authorityYear="1910" box="[1133,1213,313,336]" class="Reptilia" family="Colubridae" genus="Leptophis" kingdom="Animalia" order="Squamata" pageId="5" phylum="Chordata" rank="species" species="riveti">
<emphasis id="B951437AFFDFFFA66F6C4210FB19997C" box="[1133,1213,313,336]" italics="true" pageId="5">L. riveti</emphasis>
</taxonomicName>
); and keels absent on the first dorsal scale rows (vs. keels present on all dorsal scales of trunk in
<taxonomicName id="4C25E4EBFFDFFFA66F324250FB2799BC" authorityName="Despax" authorityYear="1910" box="[1075,1155,377,400]" class="Reptilia" family="Colubridae" genus="Leptophis" kingdom="Animalia" order="Squamata" pageId="5" phylum="Chordata" rank="species" species="riveti">
<emphasis id="B951437AFFDFFFA66F324250FB2799BC" box="[1075,1155,377,400]" italics="true" pageId="5">L. riveti</emphasis>
</taxonomicName>
).
</paragraph>
</subSubSection>
</treatment>
</document>