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<mods:title id="006F5AB2A91F0E784347E48EF068ECFB">A new Brazilian species of the genus Philothalpus Kraatz (Coleoptera: Staphylinidae)</mods:title>
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<mods:namePart id="352025799E1CA2E323058AB8E5B3AD9F">Kleman-Junior, Louri</mods:namePart>
<mods:affiliation id="B4C078B9BD925B02F20263C980CE8871">Laboratório de Biodiversidade, Ecologia e Conservação de Ecossistemas Amazônicos, Universidade do Estado do Amazonas, Centro de Estudos Superiores de Itacoatiara, 69100 - 000 Itacoatiara, Amazonas, Brazil. &amp; Programa de Pós-Graduação em Ciência e Tecnologia para Recursos Amazônicos, Instituto de Ciências Exatas e Tecnologia, Universidade Federal do Amazonas, 69100 - 000 Itacoatiara, Amazonas, Brazil.</mods:affiliation>
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<mods:namePart id="6F424BC2C9D67457D15621BF770EF3D7">Barroso, Paulo Ricardo Ramires</mods:namePart>
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<mods:affiliation id="837A13B593805010E7FADE0F4EB32F6E">Programa de Pós-Graduação em Ciência e Tecnologia para Recursos Amazônicos, Instituto de Ciências Exatas e Tecnologia, Universidade Federal do Amazonas, 69100 - 000 Itacoatiara, Amazonas, Brazil.</mods:affiliation>
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<mods:namePart id="15B4D5DE88FD5358B24C836C37D6CA39">Pereira, Luana De Fátima Baraúna</mods:namePart>
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<mods:affiliation id="B4EDC7509F93FAF29A843D0BF3FDBCBF">Programa de Pós-Graduação em Ciência e Tecnologia para Recursos Amazônicos, Instituto de Ciências Exatas e Tecnologia, Universidade Federal do Amazonas, 69100 - 000 Itacoatiara, Amazonas, Brazil.</mods:affiliation>
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<mods:namePart id="5D8A090CEB375814B6DDBF60FE5F80E2">Asenjo, Angélico</mods:namePart>
<mods:affiliation id="4B5D97B8A51582A2C91A7075299DA11D">Instituto Tecnológico VALE. R. Boaventura da Silva 955, Nazaré, 66055 - 090 Belém, PA. Brazil. &amp; Departamento de Entomología, Museo de Historia Natural de la Universidad Nacional Mayor de San Marcos, Av. Arenales 1256, Jesús María, Lima 14, Perú.</mods:affiliation>
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<subSubSection id="2EA66508635F054DF0DB1B5BFF7FFAEE" pageId="5" pageNumber="444" type="nomenclature">
<paragraph id="66033683635F054DF0DB1B5BFF7FFAEE" blockId="5.[151,1436,1338,1400]" pageId="5" pageNumber="444">
<heading id="3D4B81EF635F054DF0DB1B5BFF7FFAEE" bold="true" centered="true" fontSize="11" level="1" pageId="5" pageNumber="444" reason="1">
<emphasis id="54C8EA91635F054DF0DB1B5BFF7FFAEE" bold="true" pageId="5" pageNumber="444">
To include
<taxonomicName id="A1BC4D00635F054DF1651B5BFE36FAC3" authority="Kleman-Junior &amp; Barroso &amp; Pereira &amp; Asenjo, 2024" authorityName="Kleman-Junior &amp; Barroso &amp; Pereira &amp; Asenjo" authorityYear="2024" box="[297,432,1339,1365]" class="Insecta" family="Staphylinidae" genus="Philothalpus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="444" phylum="Arthropoda" rank="species" species="precious" status="sp. nov.">
<emphasis id="54C8EA91635F054DF1651B5BFE36FAC3" bold="true" box="[297,432,1339,1365]" italics="true" pageId="5" pageNumber="444">P. precious</emphasis>
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<taxonomicNameLabel id="4FFB57EA635F054DF1F71B5BFD98FAC3" box="[443,542,1339,1365]" pageId="5" pageNumber="444" rank="species">sp. nov.</taxonomicNameLabel>
, the taxonomic key proposed by
<bibRefCitation id="022D4B72635F054DF39C1B5AFB58FAC2" author="Chatzimanolis, S." box="[976,1246,1338,1364]" pageId="5" pageNumber="444" pagination="129 - 138" refId="ref5533" refString="Chatzimanolis, S. (2020) Even more new species, additional records, and a revised key for Philothalpus Kraatz (Coleoptera: Staphylinidae: Staphylininae). Zootaxa, 4742 (1), 129 - 138. https: // doi. org / 10.11646 / zootaxa. 4742.1.8" type="journal article" year="2020">Chatzimanolis (2020)</bibRefCitation>
is modified as follows:
</emphasis>
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<subSubSection id="2EA66508635F054DF0AE1BC4FDC2F83A" pageId="5" pageNumber="444" type="description">
<paragraph id="66033683635F054DF0AE1BC4FDBEFA2C" blockId="5.[151,1437,1444,1964]" box="[226,568,1444,1466]" pageId="5" pageNumber="444">Couplets 16 without modification.</paragraph>
<paragraph id="66033683635F054DF0DB1BA1FA1BF9B8" blockId="5.[151,1437,1444,1964]" pageId="5" pageNumber="444">
7. Head and pronotum red-brown to dark brown (
<bibRefCitation id="022D4B72635F054DF2E21BA1FC41FA41" author="Chatzimanolis, S. &amp; Ashe, J. S." box="[686,967,1473,1495]" pageId="5" pageNumber="444" pagination="63 - 119" refId="ref5586" refString="Chatzimanolis, S. &amp; Ashe, J. S. (2005) Revision and phylogeny of the neotropical genus Philothalpus (= Eugastus Sharp and Allostenopsis Bernhauer) (Coleoptera: Staphylinidae: Xanthopygina). Insect Systematics &amp; Evolution, 36, 63 - 119. https: // doi. org / 10.1163 / 187631205788912813" type="journal article" year="2005">Chatzimanolis &amp; Ashe 2005</bibRefCitation>
; fig. 5). Elytra metallic green. Abdomen orange except for antero-medial disk of tergites IIV, anterior 2/33/4 of segment VI except at base, anterior 1/2 2/3 of segment VII and posterior 1/2 of segment VIII brown. Legs orange. Aedeagus as in
<bibRefCitation id="022D4B72635F054DF3C71B9CFB23F987" author="Chatzimanolis, S. &amp; Ashe, J. S." box="[907,1189,1531,1554]" pageId="5" pageNumber="444" pagination="63 - 119" refId="ref5586" refString="Chatzimanolis, S. &amp; Ashe, J. S. (2005) Revision and phylogeny of the neotropical genus Philothalpus (= Eugastus Sharp and Allostenopsis Bernhauer) (Coleoptera: Staphylinidae: Xanthopygina). Insect Systematics &amp; Evolution, 36, 63 - 119. https: // doi. org / 10.1163 / 187631205788912813" type="journal article" year="2005">Chatzimanolis &amp; Ashe (2005</bibRefCitation>
; figs. 5356).....................................................................................................
<taxonomicName id="A1BC4D00635F054DF572187AFA1BF9B8" authorityName="Chatzimanolis &amp; Ashe" authorityYear="2005" box="[1342,1437,1561,1583]" class="Insecta" family="Staphylinidae" genus="Philothalpus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="444" phylum="Arthropoda" rank="species" species="bilobus">
<emphasis id="54C8EA91635F054DF572187AFA1BF9B8" box="[1342,1437,1561,1583]" italics="true" pageId="5" pageNumber="444">P. bilobus</emphasis>
</taxonomicName>
</paragraph>
<paragraph id="66033683635F054DF0DB1857FA1BF910" blockId="5.[151,1437,1444,1964]" pageId="5" pageNumber="444">
- Head and pronotum black or dark reddish brown with greenish or greenish golden overtones (
<bibRefCitation id="022D4B72635F054DF4141856FAEFF9DA" author="Chatzimanolis, S. &amp; Ashe, J. S." box="[1112,1385,1590,1612]" pageId="5" pageNumber="444" pagination="63 - 119" refId="ref5586" refString="Chatzimanolis, S. &amp; Ashe, J. S. (2005) Revision and phylogeny of the neotropical genus Philothalpus (= Eugastus Sharp and Allostenopsis Bernhauer) (Coleoptera: Staphylinidae: Xanthopygina). Insect Systematics &amp; Evolution, 36, 63 - 119. https: // doi. org / 10.1163 / 187631205788912813" type="journal article" year="2005">Chatzimanolis &amp; Ashe 2005</bibRefCitation>
; figs. 7, 29, 30) and Figs. (1AC). Elytra black with greenish or greenish golden overtones (
<figureCitation id="FE872A06635F054DF4401834FBD1F9FF" box="[1036,1111,1620,1642]" captionStart="FIGURE 1" captionStartId="3.[151,250,1951,1975]" captionTargetBox="[163,1425,465,1915]" captionTargetId="figure-139@3.[151,1436,453,1927]" captionTargetPageId="3" captionText="FIGURE 1. Holotype of Philothalpus precious sp. nov.. A. Dorsal habitus. B. Head. C. Pronotum. D. Elytra. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.10791919" httpUri="https://zenodo.org/record/10791919/files/figure.png" pageId="5" pageNumber="444">Fig. 1D</figureCitation>
); abdomen overall darker, colored differently; legs darker, at least partly dark brown.......................................................... 7A
</paragraph>
<paragraph id="66033683635F054DF0DB18EEFA1AF96A" blockId="5.[151,1437,1444,1964]" pageId="5" pageNumber="444">
7A Head and pronotum black with greenish overtones (
<bibRefCitation id="022D4B72635F054DF29718EEFC75F932" author="Chatzimanolis, S. &amp; Ashe, J. S." box="[731,1011,1678,1700]" pageId="5" pageNumber="444" pagination="63 - 119" refId="ref5586" refString="Chatzimanolis, S. &amp; Ashe, J. S. (2005) Revision and phylogeny of the neotropical genus Philothalpus (= Eugastus Sharp and Allostenopsis Bernhauer) (Coleoptera: Staphylinidae: Xanthopygina). Insect Systematics &amp; Evolution, 36, 63 - 119. https: // doi. org / 10.1163 / 187631205788912813" type="journal article" year="2005">Chatzimanolis &amp; Ashe 2005</bibRefCitation>
; figs. 7, 29, 30). Elytra black with greenish overtones. Abdomen dark reddish brown except segment II and posterior border of sternites IIIVI, posterior 1/3 of segment VII and anterior 4/5 of segment VIII dark orange. Legs brown. Aedeagus as in
<bibRefCitation id="022D4B72635F054DF39418A9FB61F949" author="Chatzimanolis, S. &amp; Ashe, J. S." box="[984,1255,1737,1759]" pageId="5" pageNumber="444" pagination="63 - 119" refId="ref5586" refString="Chatzimanolis, S. &amp; Ashe, J. S. (2005) Revision and phylogeny of the neotropical genus Philothalpus (= Eugastus Sharp and Allostenopsis Bernhauer) (Coleoptera: Staphylinidae: Xanthopygina). Insect Systematics &amp; Evolution, 36, 63 - 119. https: // doi. org / 10.1163 / 187631205788912813" type="journal article" year="2005">Chatzimanolis &amp; Ashe 2005</bibRefCitation>
(figs. 8285).........................................................................................
<taxonomicName id="A1BC4D00635F054DF4B81887FA1AF96A" authorityName="Chatzimanolis &amp; Ashe" authorityYear="2005" box="[1268,1436,1767,1788]" class="Insecta" family="Staphylinidae" genus="Philothalpus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="444" phylum="Arthropoda" rank="species" species="mauroprasinus">
<emphasis id="54C8EA91635F054DF4B81887FA1AF96A" box="[1268,1436,1767,1788]" italics="true" pageId="5" pageNumber="444">P. mauroprasinus</emphasis>
</taxonomicName>
</paragraph>
<paragraph id="66033683635F054DF0DB1964FDC2F83A" blockId="5.[151,1437,1444,1964]" pageId="5" pageNumber="444">
- Head and pronotum dark reddish brown with greenish golden overtones (
<figureCitation id="FE872A06635F054DF3F11964FBA8F88F" box="[957,1070,1796,1818]" captionStart="FIGURE 1" captionStartId="3.[151,250,1951,1975]" captionTargetBox="[163,1425,465,1915]" captionTargetId="figure-139@3.[151,1436,453,1927]" captionTargetPageId="3" captionText="FIGURE 1. Holotype of Philothalpus precious sp. nov.. A. Dorsal habitus. B. Head. C. Pronotum. D. Elytra. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.10791919" httpUri="https://zenodo.org/record/10791919/files/figure.png" pageId="5" pageNumber="444">Figs 1AC</figureCitation>
). Elytra black with metallic greenish golden overtones (
<figureCitation id="FE872A06635F054DF1D91941FE67F8A0" box="[405,481,1825,1847]" captionStart="FIGURE 1" captionStartId="3.[151,250,1951,1975]" captionTargetBox="[163,1425,465,1915]" captionTargetId="figure-139@3.[151,1436,453,1927]" captionTargetPageId="3" captionText="FIGURE 1. Holotype of Philothalpus precious sp. nov.. A. Dorsal habitus. B. Head. C. Pronotum. D. Elytra. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.10791919" httpUri="https://zenodo.org/record/10791919/files/figure.png" pageId="5" pageNumber="444">Fig. 1D</figureCitation>
). Abdomen dark reddish brown, almost black, except posterior border of segment III, posterior 1/3 of segments IVV, posterior 1/4 of segment VI, posterior 2/5 of segment VII, anterior border of segment VII, and anterior 2/3 of segment VIII, reddish brown. Legs brownish orange with dorsal part of hind femora and apical third of all tibiae dark brown. Aedeagus as in Figures (2AE)...........................................................
<taxonomicName id="A1BC4D00635F054DF4AC191AFACCF819" authority="Kleman-Junior &amp; Barroso &amp; Pereira &amp; Asenjo, 2024" authorityName="Kleman-Junior &amp; Barroso &amp; Pereira &amp; Asenjo" authorityYear="2024" box="[1248,1354,1914,1935]" class="Insecta" family="Staphylinidae" genus="Philothalpus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="444" phylum="Arthropoda" rank="species" species="precious" status="sp. nov.">
<emphasis id="54C8EA91635F054DF4AC191AFACCF819" box="[1248,1354,1914,1935]" italics="true" pageId="5" pageNumber="444">P. precious</emphasis>
</taxonomicName>
<emphasis id="54C8EA91635F054DF51C1919FA1AF818" bold="true" box="[1360,1436,1913,1934]" pageId="5" pageNumber="444">
<taxonomicNameLabel id="4FFB57EA635F054DF51C1919FA1AF818" box="[1360,1436,1913,1934]" pageId="5" pageNumber="444" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
Couplets 826 without modification.
</paragraph>
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