244 lines
35 KiB
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244 lines
35 KiB
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<document id="164A7C742384B8E5937AEE5329E38FBA" ID-DOI="10.1080/14772019.2016.1151958" ID-ISSN="1478-0941" ID-PMC="PMC5221741" ID-PubMed="28077930" ID-Zenodo-Dep="10883098" ID-ZooBank="A30E5364-0003-4467-B902-43A41AD456CC" IM.bibliography_approvedBy="diego" IM.illustrations_approvedBy="diego" IM.materialsCitations_approvedBy="diego" IM.metadata_approvedBy="felipe" IM.tables_approvedBy="diego" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="diego" checkinTime="1711483431746" checkinUser="felipe" docAuthor="Schwarzhans, Werner, Mors, Thomas, Engelbrecht, Andrea, Reguero, Marcelo & Kriwet, Jurgen" docDate="2017" docId="041B87CAFFAEFFFBD642F923E227D894" docLanguage="en" docName="JSystPalaeontol.15.2.147-170.pdf" docOrigin="Journal of Systematic Palaeontology 15 (2)" docSource="http://dx.doi.org/10.1080/14772019.2016.1151958" docStyle="DocumentStyle:602474738E3D100F0F13788B53F61545.3:JSystPalaeontol.2016-.journal_article.open" docStyleId="602474738E3D100F0F13788B53F61545" docStyleName="JSystPalaeontol.2016-.journal_article.open" docStyleVersion="3" docTitle="Palimphemus seymourensis Schwarzhans & Mors & Engelbrecht & Reguero & Kriwet 2017, sp. nov." docType="treatment" docVersion="3" lastPageNumber="157" masterDocId="F822FFB2FFA4FFF0D753FFE4E35DDA07" masterDocTitle="Before the freeze: otoliths from the Eocene of Seymour Island, Antarctica, reveal dominance of gadiform fishes (Teleostei)" masterLastPageNumber="170" masterPageNumber="147" pageNumber="156" updateTime="1711973403844" updateUser="diego" zenodo-license-document="CC-BY-4.0">
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<mods:title id="1666FBAE9B028C9866F17DDF459D9F53">Before the freeze: otoliths from the Eocene of Seymour Island, Antarctica, reveal dominance of gadiform fishes (Teleostei)</mods:title>
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<mods:namePart id="46DD462F351A4C57629CFFE5A1DC95A1">Schwarzhans, Werner</mods:namePart>
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<mods:affiliation id="7D3F6469E2B81AFD0025C792F03DF30F">Ahrensburger Weg 103, D- 22359 Hamburg, Germany; & Natural History Museum of Denmark, Zoological Museum, Universitetsparken 15, DK- 2100 Copenhagen, Denmark;</mods:affiliation>
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<mods:roleTerm id="B0C62C0FCB687092DDF7532A0363BAB7">Author</mods:roleTerm>
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<mods:namePart id="168E367DE07425CFC63171C922BDDB39">Mors, Thomas</mods:namePart>
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<mods:affiliation id="36C0927E10043EDD631383E220EC102F">Swedish Museum of Natural History, Department of Palaeobiology, P. O. Box 5007, SE- 10405 Stockholm, Sweden;</mods:affiliation>
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<mods:namePart id="2B316F88F99403093FA806304C5B6B60">Engelbrecht, Andrea</mods:namePart>
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<mods:affiliation id="1E8073638EA83C857F80F6D6E99216A1">University of Vienna, Department of Palaeontology, Althanstrasse 14, 1090 Vienna, Austria;</mods:affiliation>
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<mods:namePart id="6F7B9017DA89DFAAEE7CA0D7C9BCDFFB">Reguero, Marcelo</mods:namePart>
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<mods:affiliation id="0D58C52273AE95057B011C956AFF25E3">Museo de La Plata, Division ́ Paleontolog ́ ıa de Vertebrados, Paseo del Bosque s / n, B 1900 FWA La Plata, Argentina</mods:affiliation>
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<mods:namePart id="3B5DDB9601B45FA5EC7655E9A877CA50">Kriwet, Jurgen</mods:namePart>
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<mods:affiliation id="8540E5EF593EE14DB33D87331F816D9A">University of Vienna, Department of Palaeontology, Althanstrasse 14, 1090 Vienna, Austria;</mods:affiliation>
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<mods:title id="7CF5E315DBD007E0A5C584A621EF73B4">Journal of Systematic Palaeontology</mods:title>
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<mods:date id="1BA957A8A4159C3546E359DB1903CD4D">2017</mods:date>
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<mods:number id="52D940B1AA1A47407B82274C0EF784BC">2016-03-16</mods:number>
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<mods:identifier id="F50B53DA7EB1E70EAA030BFB791551A9" type="DOI">10.1080/14772019.2016.1151958</mods:identifier>
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<mods:identifier id="3306669B9AA06637A3418327DD387555" type="ISSN">1478-0941</mods:identifier>
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<treatment id="041B87CAFFAEFFFBD642F923E227D894" ID-DOI="http://doi.org/10.5281/zenodo.10883152" ID-Zenodo-Dep="10883152" LSID="urn:lsid:plazi:treatment:041B87CAFFAEFFFBD642F923E227D894" httpUri="http://treatment.plazi.org/id/041B87CAFFAEFFFBD642F923E227D894" lastPageId="11" lastPageNumber="157" pageId="10" pageNumber="156">
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<subSubSection id="C4A86557FFAEFFFAD642F923E1C1DCD9" box="[273,668,1735,1758]" pageId="10" pageNumber="156" type="nomenclature">
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<paragraph id="8C0D36DCFFAEFFFAD642F923E1C1DCD9" blockId="10.[273,668,1735,1790]" box="[273,668,1735,1758]" pageId="10" pageNumber="156">
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<heading id="D74581B0FFAEFFFAD642F923E1C1DCD9" box="[273,668,1735,1758]" centered="true" fontSize="10" level="2" pageId="10" pageNumber="156" reason="8">
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<taxonomicName id="4BB24D5FFFAEFFFAD642F923E11FDCD9" authority="Schwarzhans & Mors & Engelbrecht & Reguero & Kriwet, 2017" authorityName="Schwarzhans & Mors & Engelbrecht & Reguero & Kriwet" authorityYear="2017" box="[273,578,1735,1758]" family="Gadidae" genus="Palimphemus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Gadiformes" pageId="10" pageNumber="156" phylum="Chordata" rank="species" species="seymourensis" status="sp. nov.">
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<emphasis id="BEC6EACEFFAEFFFAD642F923E11FDCD9" bold="true" box="[273,578,1735,1758]" italics="true" pageId="10" pageNumber="156">Palimphemus seymourensis</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A5F557B5FFAEFFFAD51BF923E1C1DCD9" box="[584,668,1735,1758]" pageId="10" pageNumber="156" rank="species">sp. nov.</taxonomicNameLabel>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C4A86557FFAEFFFAD63BF903E118DCF9" box="[360,581,1767,1790]" pageId="10" pageNumber="156" type="description">
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<paragraph id="8C0D36DCFFAEFFFAD63BF903E118DCF9" blockId="10.[273,668,1735,1790]" box="[360,581,1767,1790]" pageId="10" pageNumber="156">
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(
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<figureCitation id="14892A59FFAEFFFAD623F903E2A9DCF9" box="[368,500,1767,1790]" captionStart="Figure 4" captionStartId="9.[159,227,1703,1724]" captionTargetBox="[175,1452,183,1671]" captionTargetId="figure-1@9.[175,1452,183,1671]" captionTargetPageId="9" captionText="Figure 4. Drawings of Eocene otoliths from Seymour Island. A, B, Tripterophycis immutatus Schwarzhans, 1980, NRM-PZ P.15969, mirror imaged; A, inner face; B, dorsal view. C—F, Macruronus eastmani sp. nov.; C—E, holotype, NRM-PZ P.15970, mirror imaged; C, inner face; D, outer face; E, ventral view; F, paratype, NRM-PZ P.15971, inner face. G—M, Palimphemus seymourensis sp. nov.; G—I, holotype, NRM-PZ P.15973, mirror imaged; G, inner face; H, ventral view; I, outer face; J, paratype, NRM-PZ P.15975, mirror imaged, inner face; K—M, paratypes, NRM-PZ P.15974; K, inner face, mirror imaged; L, ventral view; M, inner face. N—R, Coelorinchus nordenskjoeldi sp. nov.; N—P, holotype, NRM-PZ P.15978; N, inner face; O, ventral view; P, outer face; Q, R, paratype, NRM-PZ P.15979, mirror imaged; Q, inner face; R, ventral view. S—U, Coelorinchus balushkini sp. nov., holotype, NRM-PZ P.15976; S, inner face; T, ventral view; U, outer face. V, W, Coelorinchus sp., NRM-PZ P.15911; V, inner face (strongly eroded); W, ventral view." figureDoi="http://doi.org/10.5281/zenodo.10883110" httpUri="https://zenodo.org/record/10883110/files/figure.png" pageId="10" pageNumber="156">Figs 4G—M</figureCitation>
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,
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<figureCitation id="14892A59FFAEFFFAD6ADF903E161DCF9" box="[510,572,1767,1790]" captionStart="Figure 6" captionStartId="15.[159,227,1815,1836]" captionTargetBox="[216,1409,183,1783]" captionTargetId="figure-1@15.[216,1410,183,1783]" captionTargetPageId="15" captionText="Figure 6. Eocene otoliths from Seymour Island. A, Argentina antarctica sp. nov., holotype, NRM-PZ P.15964, mirror imaged, inner face. B, C, Diaphus? marambionis sp. nov., holotype, NRM-PZ P.15966; B, inner face; C, ventral view. D—F, Macruronus eastmani sp. nov., holotype, NRM-PZ P.15970, mirror imaged; D, inner face; E, ventral view; F, outer face. G—I, Palimphemus seymourensis sp. nov., holotype, NRM-PZ P.15973, mirror imaged; G, inner face; H, ventral view; I, outer face. J—L, Coelorinchus nordenskjoeldi sp. nov., holotype, NRM-PZ P.15978; J, inner face; K, ventral view; L, outer face. M, N, Coelorinchus balushkini sp. nov., holotype, NRM-PZ P.15976; M, inner face; N, ventral view. O, P, Hoplobrotula? antipoda sp. nov., holotype, NRM-PZ P.15984, mirror imaged; O, inner face; P, ventral view. Q, Notoberyx cionei gen. nov., sp. nov.; holotype, NRM-PZ P.15987, inner face. R, Cepola anderssoni sp. nov., holotype, NRM-PZ P.15996, mirror imaged, inner face." figureDoi="http://doi.org/10.5281/zenodo.10883114" httpUri="https://zenodo.org/record/10883114/files/figure.png" pageId="10" pageNumber="156">6G—I</figureCitation>
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)
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</paragraph>
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||
</subSubSection>
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<subSubSection id="C4A86557FFAEFFFAD7CCF8D3E62BDB13" pageId="10" pageNumber="156" type="materials_examined">
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<paragraph id="8C0D36DCFFAEFFFAD7CCF8D3E2BFDD89" blockId="10.[159,781,1847,1934]" pageId="10" pageNumber="156">
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<emphasis id="BEC6EACEFFAEFFFAD7CCF8D3E257DD49" bold="true" box="[159,266,1847,1870]" pageId="10" pageNumber="156">Material.</emphasis>
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<materialsCitation id="3CDA3C81FFAEFFFAD64BF8D3E3B5DD69" pageId="10" pageNumber="156" specimenCode="NRM-PZ P.15973" specimenCount="1" typeStatus="holotype">
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<typeStatus id="5309887EFFAEFFFAD64BF8D3E2DDDD49" box="[280,384,1847,1870]" pageId="10" pageNumber="156" type="holotype">Holotype</typeStatus>
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:
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<specimenCode id="DC149EA7FFAEFFFAD6CBF8D3E134DD49" box="[408,617,1847,1870]" pageId="10" pageNumber="156">NRM-PZ P.15973</specimenCode>
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(
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<figureCitation id="14892A59FFAEFFFAD5D5F8D3E054DD49" box="[646,777,1847,1870]" captionStart="Figure 4" captionStartId="9.[159,227,1703,1724]" captionTargetBox="[175,1452,183,1671]" captionTargetId="figure-1@9.[175,1452,183,1671]" captionTargetPageId="9" captionText="Figure 4. Drawings of Eocene otoliths from Seymour Island. A, B, Tripterophycis immutatus Schwarzhans, 1980, NRM-PZ P.15969, mirror imaged; A, inner face; B, dorsal view. C—F, Macruronus eastmani sp. nov.; C—E, holotype, NRM-PZ P.15970, mirror imaged; C, inner face; D, outer face; E, ventral view; F, paratype, NRM-PZ P.15971, inner face. G—M, Palimphemus seymourensis sp. nov.; G—I, holotype, NRM-PZ P.15973, mirror imaged; G, inner face; H, ventral view; I, outer face; J, paratype, NRM-PZ P.15975, mirror imaged, inner face; K—M, paratypes, NRM-PZ P.15974; K, inner face, mirror imaged; L, ventral view; M, inner face. N—R, Coelorinchus nordenskjoeldi sp. nov.; N—P, holotype, NRM-PZ P.15978; N, inner face; O, ventral view; P, outer face; Q, R, paratype, NRM-PZ P.15979, mirror imaged; Q, inner face; R, ventral view. S—U, Coelorinchus balushkini sp. nov., holotype, NRM-PZ P.15976; S, inner face; T, ventral view; U, outer face. V, W, Coelorinchus sp., NRM-PZ P.15911; V, inner face (strongly eroded); W, ventral view." figureDoi="http://doi.org/10.5281/zenodo.10883110" httpUri="https://zenodo.org/record/10883110/files/figure.png" pageId="10" pageNumber="156">Figs 4G—I</figureCitation>
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,
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<figureCitation id="14892A59FFAEFFFAD7CCF8B3E382DD69" box="[159,223,1879,1902]" captionStart="Figure 6" captionStartId="15.[159,227,1815,1836]" captionTargetBox="[216,1409,183,1783]" captionTargetId="figure-1@15.[216,1410,183,1783]" captionTargetPageId="15" captionText="Figure 6. Eocene otoliths from Seymour Island. A, Argentina antarctica sp. nov., holotype, NRM-PZ P.15964, mirror imaged, inner face. B, C, Diaphus? marambionis sp. nov., holotype, NRM-PZ P.15966; B, inner face; C, ventral view. D—F, Macruronus eastmani sp. nov., holotype, NRM-PZ P.15970, mirror imaged; D, inner face; E, ventral view; F, outer face. G—I, Palimphemus seymourensis sp. nov., holotype, NRM-PZ P.15973, mirror imaged; G, inner face; H, ventral view; I, outer face. J—L, Coelorinchus nordenskjoeldi sp. nov., holotype, NRM-PZ P.15978; J, inner face; K, ventral view; L, outer face. M, N, Coelorinchus balushkini sp. nov., holotype, NRM-PZ P.15976; M, inner face; N, ventral view. O, P, Hoplobrotula? antipoda sp. nov., holotype, NRM-PZ P.15984, mirror imaged; O, inner face; P, ventral view. Q, Notoberyx cionei gen. nov., sp. nov.; holotype, NRM-PZ P.15987, inner face. R, Cepola anderssoni sp. nov., holotype, NRM-PZ P.15996, mirror imaged, inner face." figureDoi="http://doi.org/10.5281/zenodo.10883114" httpUri="https://zenodo.org/record/10883114/files/figure.png" pageId="10" pageNumber="156">6G—I</figureCitation>
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)
|
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</materialsCitation>
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||
.
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<typeStatus id="5309887EFFAEFFFAD648F8B3E2DADD69" box="[283,391,1879,1902]" pageId="10" pageNumber="156" type="paratype">Paratypes</typeStatus>
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:
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<specimenCount id="9AB4FD55FFAEFFFAD6EBF8B3E12ADD69" box="[440,631,1879,1902]" count="6" pageId="10" pageNumber="156" type="generic">six specimens</specimenCount>
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, NRM-PZ P.15974—15975 (
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<figureCitation id="14892A59FFAEFFFAD60DF893E288DD89" box="[350,469,1911,1934]" captionStart="Figure 4" captionStartId="9.[159,227,1703,1724]" captionTargetBox="[175,1452,183,1671]" captionTargetId="figure-1@9.[175,1452,183,1671]" captionTargetPageId="9" captionText="Figure 4. Drawings of Eocene otoliths from Seymour Island. A, B, Tripterophycis immutatus Schwarzhans, 1980, NRM-PZ P.15969, mirror imaged; A, inner face; B, dorsal view. C—F, Macruronus eastmani sp. nov.; C—E, holotype, NRM-PZ P.15970, mirror imaged; C, inner face; D, outer face; E, ventral view; F, paratype, NRM-PZ P.15971, inner face. G—M, Palimphemus seymourensis sp. nov.; G—I, holotype, NRM-PZ P.15973, mirror imaged; G, inner face; H, ventral view; I, outer face; J, paratype, NRM-PZ P.15975, mirror imaged, inner face; K—M, paratypes, NRM-PZ P.15974; K, inner face, mirror imaged; L, ventral view; M, inner face. N—R, Coelorinchus nordenskjoeldi sp. nov.; N—P, holotype, NRM-PZ P.15978; N, inner face; O, ventral view; P, outer face; Q, R, paratype, NRM-PZ P.15979, mirror imaged; Q, inner face; R, ventral view. S—U, Coelorinchus balushkini sp. nov., holotype, NRM-PZ P.15976; S, inner face; T, ventral view; U, outer face. V, W, Coelorinchus sp., NRM-PZ P.15911; V, inner face (strongly eroded); W, ventral view." figureDoi="http://doi.org/10.5281/zenodo.10883110" httpUri="https://zenodo.org/record/10883110/files/figure.png" pageId="10" pageNumber="156">Fig. 4J—M</figureCitation>
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).
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</paragraph>
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<paragraph id="8C0D36DCFFAEFFFAD41EFF59E62BDB13" blockId="10.[845,1467,189,276]" pageId="10" pageNumber="156">
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<emphasis id="BEC6EACEFFAEFFFAD41EFF59E084DAD3" bold="true" box="[845,985,189,212]" pageId="10" pageNumber="156">Occurrence.</emphasis>
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Telm 5 unit; ‘
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<taxonomicName id="4BB24D5FFFAEFFFAD3DBFF59E792DAD3" authorityName="Scopoli" authorityYear="1777" box="[1160,1231,189,212]" class="Gastropoda" family="Naticidae" genus="Natica" kingdom="Animalia" order="Littorinimorpha" pageId="10" pageNumber="156" phylum="Mollusca" rank="genus">
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<emphasis id="BEC6EACEFFAEFFFAD3DBFF59E792DAD3" box="[1160,1231,189,212]" italics="true" pageId="10" pageNumber="156">Natica</emphasis>
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</taxonomicName>
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horizon’,
|
||
<taxonomicName id="4BB24D5FFFAEFFFAD215FF59E6FBDAD3" box="[1350,1446,189,212]" class="Bivalvia" family="Cucullaeidae" genus="Cucullea" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Arcida" pageId="10" pageNumber="156" phylum="Mollusca" rank="genus">
|
||
<emphasis id="BEC6EACEFFAEFFFAD215FF59E6FBDAD3" box="[1350,1446,189,212]" italics="true" pageId="10" pageNumber="156">Cucullea</emphasis>
|
||
</taxonomicName>
|
||
I member, La Meseta Formation, late Ypresian, early Eocene. Site IAA 2/95, Seymour Island,
|
||
<collectingCountry id="F4A5764CFFAEFFFAD3ACFF19E62FDB13" box="[1279,1394,253,276]" name="Antarctica" pageId="10" pageNumber="156">Antarctica</collectingCountry>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C4A86557FFAEFFFAD41EFEC9E77BDB63" pageId="10" pageNumber="156" type="etymology">
|
||
<paragraph id="8C0D36DCFFAEFFFAD41EFEC9E77BDB63" blockId="10.[845,1466,301,356]" pageId="10" pageNumber="156">
|
||
<emphasis id="BEC6EACEFFAEFFFAD41EFEC9E092DB43" bold="true" box="[845,975,301,324]" pageId="10" pageNumber="156">Etymology.</emphasis>
|
||
Named after the
|
||
<typeStatus id="5309887EFFAEFFFAD3E6FEC9E7BEDB43" box="[1205,1251,301,324]" pageId="10" pageNumber="156">type</typeStatus>
|
||
locality Seymour Island in
|
||
<collectingCountry id="F4A5764CFFAEFFFAD4E3FEA9E77EDB63" box="[944,1059,333,356]" name="Antarctica" pageId="10" pageNumber="156">Antarctica</collectingCountry>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C4A86557FFAEFFFAD41EFE99E637D874" pageId="10" pageNumber="156" type="diagnosis">
|
||
<paragraph id="8C0D36DCFFAEFFFAD41EFE99E637D874" blockId="10.[845,1467,381,627]" pageId="10" pageNumber="156">
|
||
<emphasis id="BEC6EACEFFAEFFFAD41EFE99E09CDB93" bold="true" box="[845,961,381,404]" pageId="10" pageNumber="156">Diagnosis.</emphasis>
|
||
Outline droplet-shaped with broadly rounded anterior rim and slightly elevated predorsal region. OL: OH = 1.8—1.9. Inner face moderately convex. Sulcus narrow, with moderately widened collum and distinct, but short pseudocolliculum. CaL:OsL (measured from centre of collum) = 1.2—1.35. Colliculi often reduced towards outer margin, particularly ostial colliculum. Ventral furrow well developed, close to ventral rim of otolith.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C4A86557FFAEFFFAD41EFD68E75FDF47" pageId="10" pageNumber="156" type="description">
|
||
<paragraph id="8C0D36DCFFAEFFFAD41EFD68E0CED9C5" blockId="10.[845,1467,652,1344]" pageId="10" pageNumber="156">
|
||
<emphasis id="BEC6EACEFFAEFFFAD41EFD68E085D8A4" bold="true" box="[845,984,652,675]" pageId="10" pageNumber="156">Description.</emphasis>
|
||
Moderately elongate and moderately thick droplet-shaped otoliths of up to
|
||
<quantity id="4B4A9B39FFAEFFFAD3E2FD48E659D8C4" box="[1201,1284,684,707]" metricMagnitude="-3" metricUnit="m" metricValue="4.5" pageId="10" pageNumber="156" unit="mm" value="4.5">4.5 mm</quantity>
|
||
in length (
|
||
<typeStatus id="5309887EFFAEFFFAD2D2FD48E026D8E5" pageId="10" pageNumber="156" type="holotype">holotype</typeStatus>
|
||
<quantity id="4B4A9B39FFAEFFFAD4D2FD2FE08FD8E5" box="[897,978,715,738]" metricMagnitude="-3" metricUnit="m" metricValue="4.5" pageId="10" pageNumber="156" unit="mm" value="4.5">4.5 mm</quantity>
|
||
long). OH:OT = 2.1—2.5. Dorsal rim highest anteriorly above ostial collum in a broadly rounded predorsal angle, somewhat undulating, posteriorly regularly declining without prominent angle. Ventral rim smooth, regularly curved, deepest below rear part of ostium. Anterior rim broadly rounded, with its tip mostly below ostium; posterior rim tapering with rounded tip at level of cauda.
|
||
</paragraph>
|
||
<paragraph id="8C0D36DCFFAEFFFAD434FC2EE75FDF47" blockId="10.[845,1467,652,1344]" pageId="10" pageNumber="156">Inner face moderately convex with slightly supramedian, narrow and rather shallow sulcus. Ostium slightly shorter than cauda and slightly narrower. Ostial and caudal colliculi well marked, with relatively wide collum in between and often reduced towards anterior and posterior tips of otolith. Collum somewhat narrowed from ventral with short but distinct pseudocolliculum. Dorsal depression narrow, with indistinct dorsal margin; ventral furrow well developed and close to ventral rim of otolith. Outer face slightly convex to nearly flat, with some radial furrows coalescing in a shallow umbo opposing the collum of the inner face.</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C4A86557FFAEFFFBD41EFABDE227D894" lastPageId="11" lastPageNumber="157" pageId="10" pageNumber="156" type="discussion">
|
||
<paragraph id="8C0D36DCFFAEFFFAD41EFABDE0B8DC28" blockId="10.[845,1467,1369,1934]" pageId="10" pageNumber="156">
|
||
<emphasis id="BEC6EACEFFAEFFFAD41EFABDE0E6DF77" bold="true" box="[845,955,1369,1392]" pageId="10" pageNumber="156">Remarks.</emphasis>
|
||
Otoliths of
|
||
<taxonomicName id="4BB24D5FFFAEFFFAD31DFABDE65CDF77" authorityName="Schwarzhans & Mors & Engelbrecht & Reguero & Kriwet" authorityYear="2017" box="[1102,1281,1369,1392]" family="Gadidae" genus="Palimphemus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Gadiformes" pageId="10" pageNumber="156" phylum="Chordata" rank="species" species="seymourensis">
|
||
<emphasis id="BEC6EACEFFAEFFFAD31DFABDE65CDF77" box="[1102,1281,1369,1392]" italics="true" pageId="10" pageNumber="156">P. seymourensis</emphasis>
|
||
</taxonomicName>
|
||
vary slightly in the expression of the ornamentation of the dorsal rim and the outer face, irrespective of the size of the otoliths. Small specimens often show somewhat reduced colliculi terminating more distantly from the anterior and posterior otolith rims (
|
||
<figureCitation id="14892A59FFAEFFFAD48EFA1CE707DC08" box="[989,1114,1528,1551]" captionStart="Figure 4" captionStartId="9.[159,227,1703,1724]" captionTargetBox="[175,1452,183,1671]" captionTargetId="figure-1@9.[175,1452,183,1671]" captionTargetPageId="9" captionText="Figure 4. Drawings of Eocene otoliths from Seymour Island. A, B, Tripterophycis immutatus Schwarzhans, 1980, NRM-PZ P.15969, mirror imaged; A, inner face; B, dorsal view. C—F, Macruronus eastmani sp. nov.; C—E, holotype, NRM-PZ P.15970, mirror imaged; C, inner face; D, outer face; E, ventral view; F, paratype, NRM-PZ P.15971, inner face. G—M, Palimphemus seymourensis sp. nov.; G—I, holotype, NRM-PZ P.15973, mirror imaged; G, inner face; H, ventral view; I, outer face; J, paratype, NRM-PZ P.15975, mirror imaged, inner face; K—M, paratypes, NRM-PZ P.15974; K, inner face, mirror imaged; L, ventral view; M, inner face. N—R, Coelorinchus nordenskjoeldi sp. nov.; N—P, holotype, NRM-PZ P.15978; N, inner face; O, ventral view; P, outer face; Q, R, paratype, NRM-PZ P.15979, mirror imaged; Q, inner face; R, ventral view. S—U, Coelorinchus balushkini sp. nov., holotype, NRM-PZ P.15976; S, inner face; T, ventral view; U, outer face. V, W, Coelorinchus sp., NRM-PZ P.15911; V, inner face (strongly eroded); W, ventral view." figureDoi="http://doi.org/10.5281/zenodo.10883110" httpUri="https://zenodo.org/record/10883110/files/figure.png" pageId="10" pageNumber="156">Fig. 4K, M</figureCitation>
|
||
) than is the case in some larger ones (
|
||
<figureCitation id="14892A59FFAEFFFAD4DEF9FCE085DC28" box="[909,984,1560,1583]" captionStart="Figure 4" captionStartId="9.[159,227,1703,1724]" captionTargetBox="[175,1452,183,1671]" captionTargetId="figure-1@9.[175,1452,183,1671]" captionTargetPageId="9" captionText="Figure 4. Drawings of Eocene otoliths from Seymour Island. A, B, Tripterophycis immutatus Schwarzhans, 1980, NRM-PZ P.15969, mirror imaged; A, inner face; B, dorsal view. C—F, Macruronus eastmani sp. nov.; C—E, holotype, NRM-PZ P.15970, mirror imaged; C, inner face; D, outer face; E, ventral view; F, paratype, NRM-PZ P.15971, inner face. G—M, Palimphemus seymourensis sp. nov.; G—I, holotype, NRM-PZ P.15973, mirror imaged; G, inner face; H, ventral view; I, outer face; J, paratype, NRM-PZ P.15975, mirror imaged, inner face; K—M, paratypes, NRM-PZ P.15974; K, inner face, mirror imaged; L, ventral view; M, inner face. N—R, Coelorinchus nordenskjoeldi sp. nov.; N—P, holotype, NRM-PZ P.15978; N, inner face; O, ventral view; P, outer face; Q, R, paratype, NRM-PZ P.15979, mirror imaged; Q, inner face; R, ventral view. S—U, Coelorinchus balushkini sp. nov., holotype, NRM-PZ P.15976; S, inner face; T, ventral view; U, outer face. V, W, Coelorinchus sp., NRM-PZ P.15911; V, inner face (strongly eroded); W, ventral view." figureDoi="http://doi.org/10.5281/zenodo.10883110" httpUri="https://zenodo.org/record/10883110/files/figure.png" pageId="10" pageNumber="156">Fig. 4J</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8C0D36DCFFAEFFFBD434F9DCE1C4DBD4" blockId="10.[845,1467,1369,1934]" lastBlockId="11.[159,782,189,659]" lastPageId="11" lastPageNumber="157" pageId="10" pageNumber="156">
|
||
<taxonomicName id="4BB24D5FFFAEFFFAD434F9DCE7DADC48" authorityName="Schwarzhans & Mors & Engelbrecht & Reguero & Kriwet" authorityYear="2017" box="[871,1159,1592,1615]" family="Gadidae" genus="Palimphemus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Gadiformes" pageId="10" pageNumber="156" phylum="Chordata" rank="species" species="seymourensis">
|
||
<emphasis id="BEC6EACEFFAEFFFAD434F9DCE7DADC48" box="[871,1159,1592,1615]" italics="true" pageId="10" pageNumber="156">Palimphemus seymourensis</emphasis>
|
||
</taxonomicName>
|
||
represents the first record of a gadid from
|
||
<collectingCountry id="F4A5764CFFAEFFFAD4A3F9BCE706DC68" box="[1008,1115,1624,1647]" name="Antarctica" pageId="10" pageNumber="156">Antarctica</collectingCountry>
|
||
and the Southern Hemisphere, except for the occurrence of
|
||
<taxonomicName id="4BB24D5FFFAEFFFAD328F99CE09DDCA8" authority="Schwarzhans, 1980" authorityName="Schwarzhans" authorityYear="1980" family="Gadidae" genus="Gadiculus" kingdom="Animalia" order="Gadiformes" pageId="10" pageNumber="156" phylum="Chordata" rank="species" species="antipodus">
|
||
<emphasis id="BEC6EACEFFAEFFFAD328F99CE60CDC88" box="[1147,1361,1656,1679]" italics="true" pageId="10" pageNumber="156">Gadiculus antipodus</emphasis>
|
||
<bibRefCitation id="E8234B2DFFAEFFFAD20AF99CE09DDCA8" author="Schwarzhans, W." pageId="10" pageNumber="156" pagination="1 - 269" refId="ref17706" refString="Schwarzhans, W. 1980. Die terti ¨ are Teleosteer-Fauna Neuseelands, rekonstruiert anhand von Otolithen. Berliner geowissenschaftliche Abhandlungen (A), 26, 1 - 211. [English translation 1984 in New Zealand Geological Survey Report, 113, 1 - 269.]" type="journal article" year="1980">
|
||
Schwarzhans,
|
||
<quantity id="4B4A9B39FFAEFFFAD4DFF97CE083DCA8" box="[908,990,1688,1711]" metricMagnitude="1" metricUnit="m" metricValue="5.0291999999999994" pageId="10" pageNumber="156" unit="in" value="1980.0">1980 in</quantity>
|
||
</bibRefCitation>
|
||
</taxonomicName>
|
||
the early Miocene of
|
||
<collectingCountry id="F4A5764CFFAEFFFAD380F97CE63CDCA8" box="[1235,1377,1688,1711]" name="New Zealand" pageId="10" pageNumber="156">New Zealand</collectingCountry>
|
||
and the Recent
|
||
<taxonomicName id="4BB24D5FFFAEFFFAD4F7F95CE7F5DCC8" authorityName="Norman" authorityYear="1937" box="[932,1192,1720,1743]" family="Gadidae" genus="Micromesistius" kingdom="Animalia" order="Gadiformes" pageId="10" pageNumber="156" phylum="Chordata" rank="species" species="australis">
|
||
<emphasis id="BEC6EACEFFAEFFFAD4F7F95CE7F5DCC8" box="[932,1192,1720,1743]" italics="true" pageId="10" pageNumber="156">Micromesistius australis</emphasis>
|
||
</taxonomicName>
|
||
off the southern tip of South America and off
|
||
<collectingCountry id="F4A5764CFFAEFFFAD331F93CE7A5DCE8" box="[1122,1272,1752,1775]" name="New Zealand" pageId="10" pageNumber="156">New Zealand</collectingCountry>
|
||
. In the Northern Hemisphere,
|
||
<taxonomicName id="4BB24D5FFFAEFFFAD48BF913E739DD09" authorityName="Kner" authorityYear="1862" box="[984,1124,1783,1806]" family="Gadidae" genus="Palimphemus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Gadiformes" pageId="10" pageNumber="156" phylum="Chordata" rank="genus">
|
||
<emphasis id="BEC6EACEFFAEFFFAD48BF913E739DD09" box="[984,1124,1783,1806]" italics="true" pageId="10" pageNumber="156">Palimphemus</emphasis>
|
||
</taxonomicName>
|
||
is well recorded since the early Oligocene. From these species, such as
|
||
<taxonomicName id="4BB24D5FFFAEFFFAD249F8F3E0A3DD49" authority="(Gaemers, 1994)" baseAuthorityName="Gaemers" baseAuthorityYear="1994" family="Gadidae" genus="Palimphemus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Gadiformes" pageId="10" pageNumber="156" phylum="Chordata" rank="species" species="brevicollum">
|
||
<emphasis id="BEC6EACEFFAEFFFAD249F8F3E6E6DD29" box="[1306,1467,1815,1838]" italics="true" pageId="10" pageNumber="156">P. brevicollum</emphasis>
|
||
(Gaemers, 1994)
|
||
</taxonomicName>
|
||
(in
|
||
<bibRefCitation id="E8234B2DFFAEFFFAD363F8D3E7ABDD49" author="Schwarzhans, W." box="[1072,1270,1847,1870]" pageId="10" pageNumber="156" pagination="1 - 248" refId="ref17843" refString="Schwarzhans, W. 1994. Die Fisch-Otolithen aus dem Oberoligoz ¨ an der Niederrheinischen Bucht. Systematik, Palokologie, Palaobiogeographie, Biostratigraphie und Otolithen-Zonierung. Geologisches Jahrbuch, A, 140, 1 - 248." type="journal article" year="1994">Schwarzhans 1994</bibRefCitation>
|
||
), the new species differs in the relatively thin appearance, the wider collum with a longer pseudocolliculum, and the low index CaL: OsL (1.2—1.35 vs. 1.7—2.0). Early species of the related genus
|
||
<taxonomicName id="4BB24D5FFFAFFFFBD7B4FF39E200DAF3" authorityName="Rafinesque" authorityYear="1814" box="[231,349,221,244]" family="Gadidae" genus="Trisopterus" kingdom="Animalia" order="Gadiformes" pageId="11" pageNumber="157" phylum="Chordata" rank="genus">
|
||
<emphasis id="BEC6EACEFFAFFFFBD7B4FF39E200DAF3" box="[231,349,221,244]" italics="true" pageId="11" pageNumber="157">Trisopterus</emphasis>
|
||
</taxonomicName>
|
||
from the Oligocene of Europe show a narrow collum and no pseudocolliculum (
|
||
<bibRefCitation id="E8234B2DFFAFFFFBD5D7FF19E38FDB33" author="Schwarzhans, W." pageId="11" pageNumber="157" pagination="1 - 248" refId="ref17843" refString="Schwarzhans, W. 1994. Die Fisch-Otolithen aus dem Oberoligoz ¨ an der Niederrheinischen Bucht. Systematik, Palokologie, Palaobiogeographie, Biostratigraphie und Otolithen-Zonierung. Geologisches Jahrbuch, A, 140, 1 - 248." type="journal article" year="1994">Schwarzhans 1994</bibRefCitation>
|
||
). The Paleocene and early Eocene of
|
||
<collectingCountry id="F4A5764CFFAFFFFBD538FEF9E193DB33" box="[619,718,285,308]" name="Denmark" pageId="11" pageNumber="157">Denmark</collectingCountry>
|
||
,
|
||
<collectingCountry id="F4A5764CFFAFFFFBD588FEF9E391DB53" name="United Kingdom" pageId="11" pageNumber="157">England</collectingCountry>
|
||
and
|
||
<collectingCountry id="F4A5764CFFAFFFFBD652FED9E231DB53" box="[257,364,317,340]" name="Greenland" pageId="11" pageNumber="157">Greenland</collectingCountry>
|
||
have yielded species of the related fossil otolith-based genus
|
||
<taxonomicName id="4BB24D5FFFAFFFFBD6E5FEB9E05BDB73" authority="Gaemers, 1976" authorityName="Gaemers" authorityYear="1976" box="[438,774,349,372]" family="Gadidae" genus="Protocolliolus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Gadiformes" pageId="11" pageNumber="157" phylum="Chordata" rank="genus">
|
||
<emphasis id="BEC6EACEFFAFFFFBD6E5FEB9E115DB73" box="[438,584,349,372]" italics="true" pageId="11" pageNumber="157">Protocolliolus</emphasis>
|
||
<bibRefCitation id="E8234B2DFFAFFFFBD50EFEB9E05BDB73" author="Gaemers, P. A. M." box="[605,774,349,372]" pageId="11" pageNumber="157" pagination="507 - 537" refId="ref15014" refString="Gaemers, P. A. M. 1976. New gadiform otoliths from the Tertiary of the North Sea Basin and a revision of some fossil and recent species. Leidse Geologische Mededelingen, 49, 507 - 537." type="journal article" year="1976">Gaemers, 1976</bibRefCitation>
|
||
</taxonomicName>
|
||
, which differ from
|
||
<taxonomicName id="4BB24D5FFFAFFFFBD631FE99E2B3DB93" authorityName="Kner" authorityYear="1862" box="[354,494,381,404]" family="Gadidae" genus="Palimphemus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Gadiformes" pageId="11" pageNumber="157" phylum="Chordata" rank="genus">
|
||
<emphasis id="BEC6EACEFFAFFFFBD631FE99E2B3DB93" box="[354,494,381,404]" italics="true" pageId="11" pageNumber="157">Palimphemus</emphasis>
|
||
</taxonomicName>
|
||
otoliths in the slightly narrower collum without a pseudocolliculum, while the index CaL:OsL is similarly low as in
|
||
<taxonomicName id="4BB24D5FFFAFFFFBD6B9FE58E1CFDBD4" authorityName="Schwarzhans & Mors & Engelbrecht & Reguero & Kriwet" authorityYear="2017" box="[490,658,444,467]" family="Gadidae" genus="Palimphemus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Gadiformes" pageId="11" pageNumber="157" phylum="Chordata" rank="species" species="seymourensis">
|
||
<emphasis id="BEC6EACEFFAFFFFBD6B9FE58E1CFDBD4" box="[490,658,444,467]" italics="true" pageId="11" pageNumber="157">P. seymourensis</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8C0D36DCFFAFFFFBD7E9FE38E227D894" blockId="11.[159,782,189,659]" pageId="11" pageNumber="157">
|
||
The discovery of
|
||
<taxonomicName id="4BB24D5FFFAFFFFBD62DFE38E170DBF4" authorityName="Schwarzhans & Mors & Engelbrecht & Reguero & Kriwet" authorityYear="2017" box="[382,557,476,499]" family="Gadidae" genus="Palimphemus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Gadiformes" pageId="11" pageNumber="157" phylum="Chordata" rank="species" species="seymourensis">
|
||
<emphasis id="BEC6EACEFFAFFFFBD62DFE38E170DBF4" box="[382,557,476,499]" italics="true" pageId="11" pageNumber="157">P. seymourensis</emphasis>
|
||
</taxonomicName>
|
||
in the early Eocene of
|
||
<collectingCountry id="F4A5764CFFAFFFFBD791FE18E26CD814" box="[194,305,508,531]" name="Antarctica" pageId="11" pageNumber="157">Antarctica</collectingCountry>
|
||
now represents the earliest record of the genus. It further documents that the
|
||
<taxonomicName id="4BB24D5FFFAFFFFBD560FDF8E1D1D834" authorityName="Rafinesque" authorityYear="1810" box="[563,652,540,563]" family="Gadidae" kingdom="Animalia" order="Gadiformes" pageId="11" pageNumber="157" phylum="Chordata" rank="family">Gadidae</taxonomicName>
|
||
have had a bipolar temperate distribution in the Palaeogene as opposed to the clearly temperate Northern Hemisphere dominance of today.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |