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<document id="D212F1247C242AD3C95A33BF5E563389" ID-CLB-Dataset="54981" ID-DOI="10.5252/geodiversitas2022v44a8" ID-GBIF-Dataset="7e93094c-b8b7-4077-9a4e-5b3c0e65f952" ID-ISSN="1638-9395" ID-Zenodo-Dep="6245987" ID-ZooBank="urn:lsid:zoobank.org:pub:E7490671-2900-4DE3-8E2E-D6921D80C6D1" IM.bibliography_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1645530114998" checkinUser="felipe" docAuthor="Stoetzel, Emmanuelle &amp; Pickford, Martin" docDate="2022" docId="DB698781FFE6FFF7FEE1FEB2FB0A2AC1" docLanguage="fr" docName="Geodiversitas.44.8.237-263.pdf" docOrigin="Geodiversitas 44 (8)" docStyle="DocumentStyle:F830B10FF475E64C1F1601E3B32DDC00.4:Geodiversitas.2018-.journal_article" docStyleId="F830B10FF475E64C1F1601E3B32DDC00" docStyleName="Geodiversitas.2018-.journal_article" docStyleVersion="4" docTitle="Ellobius barbarus Pomel 1892" docType="treatment" docVersion="8" lastPageNumber="251" masterDocId="2750FFF9FFF6FFE7FFEDFFE4FFE12D2A" masterDocTitle="Étude d’un assemblage original de microvertébrés du Pléistocène moyen du nord-est de l’Algérie (Ben Kérat, Oued Zenati) et description de deux nouveaux muridés" masterLastPageNumber="263" masterPageNumber="237" pageNumber="251" updateTime="1699316018958" updateUser="ExternalLinkService" zenodo-license-document="CC0-1.0" zenodo-license-figures="CC0-1.0">
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<mods:title id="2ABB1D9A2A1BD762AB033BF319D7D6BE">Étude d’un assemblage original de microvertébrés du Pléistocène moyen du nord-est de l’Algérie (Ben Kérat, Oued Zenati) et description de deux nouveaux muridés</mods:title>
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<treatment id="DB698781FFE6FFF7FEE1FEB2FB0A2AC1" ID-DOI="http://doi.org/10.5281/zenodo.6315542" ID-GBIF-Taxon="194518608" ID-Zenodo-Dep="6315542" LSID="urn:lsid:plazi:treatment:DB698781FFE6FFF7FEE1FEB2FB0A2AC1" httpUri="http://treatment.plazi.org/id/DB698781FFE6FFF7FEE1FEB2FB0A2AC1" lastPageNumber="251" pageId="16" pageNumber="251">
<subSubSection id="1BDA651CFFE6FFF7FEE1FEB2FD9E2C5B" box="[268,639,342,369]" pageId="16" pageNumber="251" type="nomenclature">
<paragraph id="537F3697FFE6FFF7FEE1FEB2FD9E2C5B" blockId="16.[268,639,342,369]" box="[268,639,342,369]" pageId="16" pageNumber="251">
<heading id="083781FBFFE6FFF7FEE1FEB2FD9E2C5B" box="[268,639,342,369]" centered="true" fontSize="11" level="2" pageId="16" pageNumber="251" reason="2">
<taxonomicName id="94C04D14FFE6FFF7FEE1FEB2FD9E2C5B" authority="Pomel, 1892" authorityName="Pomel" authorityYear="1892" box="[268,639,342,369]" class="Mammalia" family="Cricetidae" genus="Ellobius" isUncertain="true" kingdom="Animalia" order="Rodentia" pageId="16" pageNumber="251" phylum="Chordata" rank="species" species="barbarus">
<emphasis id="61B4EA85FFE6FFF7FEE1FEB2FE852C5B" bold="true" box="[268,356,342,369]" italics="true" pageId="16" pageNumber="251">Ellobius</emphasis>
cf.
<emphasis id="61B4EA85FFE6FFF7FE61FEB2FE0C2C5B" bold="true" box="[396,493,342,369]" italics="true" pageId="16" pageNumber="251">barbarus</emphasis>
Pomel, 1892
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="1BDA651CFFE6FFF7FF69FE7AFED92F20" pageId="16" pageNumber="251" type="materials_examined">
<paragraph id="537F3697FFE6FFF7FF69FE7AFED92F20" blockId="16.[132,775,413,522]" pageId="16" pageNumber="251">
<materialsCitation id="E3A83CCAFFE6FFF7FF69FE7AFED92F20" ID-GBIF-Occurrence="3748611302" pageId="16" pageNumber="251" specimenCount="1">MATÉRIEL. — N = 27+; deux M1 G, une M1 D, deux M3 G, une M3 D, cinq m1 G, trois m1 D, deux m2 G, deux m2 D, six m3 G et trois m3 D isolées (+ fragments d’émail de molaires isolées non identifiables).</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="1BDA651CFFE6FFF7FF69FDD3FB0A2AC1" pageId="16" pageNumber="251" type="description">
<paragraph id="537F3697FFE6FFF7FF69FDD3FE012F65" blockId="16.[131,777,567,2028]" box="[132,480,567,593]" pageId="16" pageNumber="251">DESCRIPTION ET COMPARAISONS</paragraph>
<paragraph id="537F3697FFE6FFF7FF69FDB2FBCF2984" blockId="16.[131,777,567,2028]" lastBlockId="16.[811,1457,789,2028]" pageId="16" pageNumber="251">
<taxonomicName id="94C04D14FFE6FFF7FF69FDB2FE992F5A" authority="Fischer, 1814" authorityName="Fischer" authorityYear="1814" box="[132,376,598,624]" class="Mammalia" family="Cricetidae" genus="Ellobius" kingdom="Animalia" order="Rodentia" pageId="16" pageNumber="251" phylum="Chordata" rank="genus">
<emphasis id="61B4EA85FFE6FFF7FF69FDB2FF382F5A" box="[132,217,598,624]" italics="true" pageId="16" pageNumber="251">Ellobius</emphasis>
Fischer, 1814
</taxonomicName>
est le seul Arvicolinae ayant jamais existé au Maghreb, et seulement au cours du Pléistocène moyen (Stoetzel 2013). Venu du Proche-Orient, son histoire évolutive en Afrique du Nord diffère de celle des populations asiatiques. Se succèdent ainsi plusieurs espèces au cours du Pléistocène moyen:
<taxonomicName id="94C04D14FFE6FFF7FE1DFD12FCE32E25" authority="Jaeger, 1988" authorityName="Jaeger" authorityYear="1988" box="[496,770,757,784]" class="Mammalia" family="Cricetidae" genus="Ellobius" kingdom="Animalia" order="Rodentia" pageId="16" pageNumber="251" phylum="Chordata" rank="species" species="africanus">
<emphasis id="61B4EA85FFE6FFF7FE1DFD12FE1E2E25" box="[496,511,758,783]" italics="true" pageId="16" pageNumber="251">E</emphasis>
.
<emphasis id="61B4EA85FFE6FFF7FDE2FD11FD8E2E25" box="[527,623,757,783]" italics="true" pageId="16" pageNumber="251">africanus</emphasis>
<bibRefCitation id="37514B66FFE6FFF7FD9AFD12FCE32E25" author="JAEGER J. J." box="[631,770,757,784]" pageId="16" pageNumber="251" pagination="3 - 50" refId="ref19224" refString="JAEGER J. J. 1988. - Origine et evolution du genre Ellobius (Mammalia, Rodentia) en Afrique nord-occidentale. Folia Quaternaria 57: 3 - 50." type="journal article" year="1988">Jaeger, 1988</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="94C04D14FFE6FFF7FF68FCF2FE702E05" authority="Jaeger, 1988" authorityName="Jaeger" authorityYear="1988" box="[133,401,789,816]" class="Mammalia" family="Cricetidae" genus="Ellobius" kingdom="Animalia" order="Rodentia" pageId="16" pageNumber="251" phylum="Chordata" rank="species" species="atlanticus">
<emphasis id="61B4EA85FFE6FFF7FF68FCF2FF752E05" box="[133,148,790,815]" italics="true" pageId="16" pageNumber="251">E</emphasis>
.
<emphasis id="61B4EA85FFE6FFF7FF4CFCF1FEE72E05" box="[161,262,789,815]" italics="true" pageId="16" pageNumber="251">atlanticus</emphasis>
<bibRefCitation id="37514B66FFE6FFF7FEE6FCF2FE702E05" author="JAEGER J. J." box="[267,401,789,816]" pageId="16" pageNumber="251" pagination="3 - 50" refId="ref19224" refString="JAEGER J. J. 1988. - Origine et evolution du genre Ellobius (Mammalia, Rodentia) en Afrique nord-occidentale. Folia Quaternaria 57: 3 - 50." type="journal article" year="1988">Jaeger, 1988</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="94C04D14FFE6FFF7FE74FCF2FD422E05" authority="Pomel, 1892" authorityName="Pomel" authorityYear="1892" box="[409,675,789,816]" class="Mammalia" family="Cricetidae" genus="Ellobius" kingdom="Animalia" order="Rodentia" pageId="16" pageNumber="251" phylum="Chordata" rank="species" species="barbarus">
<emphasis id="61B4EA85FFE6FFF7FE74FCF2FE482E05" box="[409,425,790,815]" italics="true" pageId="16" pageNumber="251">E</emphasis>
.
<emphasis id="61B4EA85FFE6FFF7FE5AFCF1FDF22E05" box="[439,531,789,815]" italics="true" pageId="16" pageNumber="251">barbarus</emphasis>
Pomel, 1892
</taxonomicName>
,
<taxonomicName id="94C04D14FFE6FFF7FD46FCF2FEEE2E65" authority="Jaeger, 1988" authorityName="Jaeger" authorityYear="1988" class="Mammalia" family="Cricetidae" genus="Ellobius" kingdom="Animalia" order="Rodentia" pageId="16" pageNumber="251" phylum="Chordata" rank="species" species="zimae">
<emphasis id="61B4EA85FFE6FFF7FD46FCF2FD5A2E05" box="[683,699,790,815]" italics="true" pageId="16" pageNumber="251">E</emphasis>
.
<emphasis id="61B4EA85FFE6FFF7FD24FCF2FCE62E05" box="[713,775,790,815]" italics="true" pageId="16" pageNumber="251">zimae</emphasis>
<bibRefCitation id="37514B66FFE6FFF7FF69FCD2FEEE2E65" author="JAEGER J. J." box="[132,271,821,848]" pageId="16" pageNumber="251" pagination="3 - 50" refId="ref19224" refString="JAEGER J. J. 1988. - Origine et evolution du genre Ellobius (Mammalia, Rodentia) en Afrique nord-occidentale. Folia Quaternaria 57: 3 - 50." type="journal article" year="1988">Jaeger, 1988</bibRefCitation>
</taxonomicName>
, avant de disparaître au Pléistocène supérieur. Certains auteurs suggèrent que le genre
<taxonomicName id="94C04D14FFE6FFF7FDABFCB1FD7A2E45" authorityName="Fischer" authorityYear="1814" box="[582,667,853,879]" class="Mammalia" family="Cricetidae" genus="Ellobius" kingdom="Animalia" order="Rodentia" pageId="16" pageNumber="251" phylum="Chordata" rank="genus">
<emphasis id="61B4EA85FFE6FFF7FDABFCB1FD7A2E45" box="[582,667,853,879]" italics="true" pageId="16" pageNumber="251">Ellobius</emphasis>
</taxonomicName>
aurait pu s’éteindre plus tôt au Maghreb occidental (
<collectingCountry id="2BD77607FFE6FFF7FD8CFC91FD4B2EA5" box="[609,682,885,911]" name="Morocco" pageId="16" pageNumber="251">Maroc</collectingCountry>
et ouest de l’Algérie) qu’au Maghreb oriental (
<collectingCountry id="2BD77607FFE6FFF7FDD4FC71FD6E2E9A" box="[569,655,917,944]" name="Tunisia" pageId="16" pageNumber="251">Tunisie</collectingCountry>
) (
<bibRefCitation id="37514B66FFE6FFF7FD47FC71FEC12EE5" author="MEIN P. &amp; PICKFORD M." pageId="16" pageNumber="251" pagination="247 - 253" refId="ref19610" refString="MEIN P. &amp; PICKFORD M. 1992. - Gisements karstiques pleistocenes au Djebel Ressas, Tunisie. Comptes Rendus de l'Academie des Sciences de Paris, Serie II 315: 247 - 253. https: // gallica. bnf. fr / ark: / 12148 / bpt 6 k 6304735 s / f 261. item" type="journal article" year="1992">Mein &amp; Pickford 1992</bibRefCitation>
). De plus, sa présence à Tighennif, dont l’âge serait potentiellement à reculer considérablement d’après des études récentes (
<bibRefCitation id="37514B66FFE6FFF7FEB3FC10FDAA2924" author="SAHNOUNI M. &amp; PARES J. &amp; DUVAL M. &amp; VAN DER MADE J. &amp; HARICHANE Z. &amp; GONZALEZ A. &amp; ABDESSADOK S. &amp; ARNOLD L. &amp; CACERES I. &amp; KANDI N. &amp; CHEHEB R. &amp; BOULAGHRAIF K. &amp; AGUSTI J. &amp; SAIDANI N. &amp; MOUHOUBI Y." box="[350,587,1012,1039]" pageId="16" pageNumber="251" refId="ref20467" refString="SAHNOUNI M., PARES J., DUVAL M., VAN DER MADE J., HARICHANE Z., PEREZ- GONZALEZ A., ABDESSADOK S., ARNOLD L., CACERES I., KANDI N., CHELLI- CHEHEB R., BOULAGHRAIF K., AGUSTI J., SAIDANI N. &amp; MOUHOUBI Y. 2018 b. - Early Homo adaptive and behavioral patterns in North Africa: Perspectives from Ain Hanech and Tighennif (formerly Ternifine) sites in northern Algeria. Communication orale, 15 th PANAF Congress, 10 - 14 September 2018, Rabat, Morocco. Abstract Book, p. 5." type="journal volume" year="2018">
Sahnouni
<emphasis id="61B4EA85FFE6FFF7FE21FC11FE162924" box="[460,503,1012,1038]" italics="true" pageId="16" pageNumber="251">et al</emphasis>
. 2018b
</bibRefCitation>
;
<bibRefCitation id="37514B66FFE6FFF7FDB5FC10FD152925" author="PICKFORD M." box="[600,756,1012,1039]" pageId="16" pageNumber="251" pagination="1 - 66" refId="ref19906" refString="PICKFORD M. 2020. - The Fossil Suidae (Mammalia, Artiodactyla) from Ternifine (Tighenif) Algeria. Munchner Geowissenschaftliche Abhandlungen (A) 50: 1 - 66." type="journal article" year="2020">Pickford 2020</bibRefCitation>
), pourrait également refléter une présence au Maghreb dès le Pléistocène inférieur (à confirmer avec l’analyse du matériel microfaunique des fouilles récentes; Saidani, thèse en cours). Trois de ces espèces représenteraient une lignée anagénétique:
<taxonomicName id="94C04D14FFE6FFF7FF20FB71FEB12984" authorityName="Jaeger" authorityYear="1988" box="[205,336,1172,1198]" class="Mammalia" family="Cricetidae" genus="Ellobius" kingdom="Animalia" order="Rodentia" pageId="16" pageNumber="251" phylum="Chordata" rank="species" species="africanus">
<emphasis id="61B4EA85FFE6FFF7FF20FB71FF3C2984" box="[205,221,1173,1198]" italics="true" pageId="16" pageNumber="251">E</emphasis>
.
<emphasis id="61B4EA85FFE6FFF7FF1DFB70FEB12984" box="[240,336,1172,1198]" italics="true" pageId="16" pageNumber="251">africanus</emphasis>
</taxonomicName>
<emphasis id="61B4EA85FFE6FFF7FEB7FB70FE892984" box="[346,360,1172,1198]" italics="true" pageId="16" pageNumber="251">–</emphasis>
<taxonomicName id="94C04D14FFE6FFF7FE9FFB71FE162984" authorityName="Jaeger" authorityYear="1988" box="[370,503,1172,1198]" class="Mammalia" family="Cricetidae" genus="Ellobius" kingdom="Animalia" order="Rodentia" pageId="16" pageNumber="251" phylum="Chordata" rank="species" species="atlanticus">
<emphasis id="61B4EA85FFE6FFF7FE9FFB71FE602984" box="[370,385,1173,1198]" italics="true" pageId="16" pageNumber="251">E</emphasis>
.
<emphasis id="61B4EA85FFE6FFF7FE7DFB70FE162984" box="[400,503,1172,1198]" italics="true" pageId="16" pageNumber="251">atlanticus</emphasis>
</taxonomicName>
<emphasis id="61B4EA85FFE6FFF7FDEDFB70FDEF2984" box="[512,526,1172,1198]" italics="true" pageId="16" pageNumber="251">–</emphasis>
<taxonomicName id="94C04D14FFE6FFF7FDF5FB71FD722984" authorityName="Pomel" authorityYear="1892" box="[536,659,1172,1198]" class="Mammalia" family="Cricetidae" genus="Ellobius" kingdom="Animalia" order="Rodentia" pageId="16" pageNumber="251" phylum="Chordata" rank="species" species="barbarus">
<emphasis id="61B4EA85FFE6FFF7FDF5FB71FDC62984" box="[536,551,1173,1198]" italics="true" pageId="16" pageNumber="251">E</emphasis>
.
<emphasis id="61B4EA85FFE6FFF7FDDBFB70FD722984" box="[566,659,1172,1198]" italics="true" pageId="16" pageNumber="251">barbarus</emphasis>
</taxonomicName>
. Au cours du temps, on observe une augmentation progressive de la taille des molaires (longueur et largeur), une augmentation de l’indice d’hypsodontie, un développement d’échancrures d’émail, l’angle rentrant lingual 1 des M2 devient plus profond et plus large et le lobe antérieur des m1 se simplifie (Jaeger 1975, 1988).
<taxonomicName id="94C04D14FFE6FFF7FE83FAB0FE2D2847" authorityName="Jaeger" authorityYear="1988" box="[366,460,1364,1390]" class="Mammalia" family="Cricetidae" genus="Ellobius" kingdom="Animalia" order="Rodentia" pageId="16" pageNumber="251" phylum="Chordata" rank="species" species="zimae">
<emphasis id="61B4EA85FFE6FFF7FE83FAB0FE9F2847" box="[366,382,1364,1389]" italics="true" pageId="16" pageNumber="251">E</emphasis>
.
<emphasis id="61B4EA85FFE6FFF7FE60FAB0FE2D2847" box="[397,460,1364,1389]" italics="true" pageId="16" pageNumber="251">zimae</emphasis>
</taxonomicName>
serait issu d’une autre lignée (s’enracinant peut-être chez
<taxonomicName id="94C04D14FFE6FFF7FE5CFA90FDCF28A7" authorityName="Jaeger" authorityYear="1988" box="[433,558,1395,1422]" class="Mammalia" family="Cricetidae" genus="Ellobius" kingdom="Animalia" order="Rodentia" pageId="16" pageNumber="251" phylum="Chordata" rank="species" species="africanus">
<emphasis id="61B4EA85FFE6FFF7FE5CFA90FE2028A7" box="[433,449,1396,1421]" italics="true" pageId="16" pageNumber="251">E</emphasis>
.
<emphasis id="61B4EA85FFE6FFF7FE22FA97FDCF28A7" box="[463,558,1395,1421]" italics="true" pageId="16" pageNumber="251">africanus</emphasis>
</taxonomicName>
) et présente des caractères dentaires un peu distincts, comme la disparition de la racine linguale sur les M1, des m3 réduites, des m1 à deux racines avec un lobe antérieur large et court et un moindre développement des échancrures d’émail (
<bibRefCitation id="37514B66FFE6FFF7FDA1FA17FD2E2B27" author="JAEGER J. J." box="[588,719,1523,1549]" pageId="16" pageNumber="251" pagination="3 - 50" refId="ref19224" refString="JAEGER J. J. 1988. - Origine et evolution du genre Ellobius (Mammalia, Rodentia) en Afrique nord-occidentale. Folia Quaternaria 57: 3 - 50." type="journal article" year="1988">Jaeger 1988</bibRefCitation>
). Au sein de la lignée
<taxonomicName id="94C04D14FFE6FFF7FED2F9F7FE5F2B07" authorityName="Jaeger" authorityYear="1988" box="[319,446,1555,1581]" class="Mammalia" family="Cricetidae" genus="Ellobius" kingdom="Animalia" order="Rodentia" pageId="16" pageNumber="251" phylum="Chordata" rank="species" species="africanus">
<emphasis id="61B4EA85FFE6FFF7FED2F9F7FEAF2B06" box="[319,334,1555,1580]" italics="true" pageId="16" pageNumber="251">E</emphasis>
.
<emphasis id="61B4EA85FFE6FFF7FEB3F9F7FE5F2B07" box="[350,446,1555,1581]" italics="true" pageId="16" pageNumber="251">africanus</emphasis>
</taxonomicName>
<emphasis id="61B4EA85FFE6FFF7FE2AF9F7FE342B07" box="[455,469,1555,1581]" italics="true" pageId="16" pageNumber="251">–</emphasis>
<taxonomicName id="94C04D14FFE6FFF7FE33F9F7FD852B06" authorityName="Jaeger" authorityYear="1988" box="[478,612,1554,1581]" class="Mammalia" family="Cricetidae" genus="Ellobius" kingdom="Animalia" order="Rodentia" pageId="16" pageNumber="251" phylum="Chordata" rank="species" species="atlanticus">
<emphasis id="61B4EA85FFE6FFF7FE33F9F7FE0C2B06" box="[478,493,1555,1580]" italics="true" pageId="16" pageNumber="251">E</emphasis>
.
<emphasis id="61B4EA85FFE6FFF7FE10F9F6FD852B06" box="[509,612,1554,1580]" italics="true" pageId="16" pageNumber="251">atlanticus</emphasis>
</taxonomicName>
<emphasis id="61B4EA85FFE6FFF7FD81F9F7FD9B2B07" box="[620,634,1555,1581]" italics="true" pageId="16" pageNumber="251">–</emphasis>
<taxonomicName id="94C04D14FFE6FFF7FD69F9F7FD1E2B06" authorityName="Pomel" authorityYear="1892" box="[644,767,1554,1581]" class="Mammalia" family="Cricetidae" genus="Ellobius" kingdom="Animalia" order="Rodentia" pageId="16" pageNumber="251" phylum="Chordata" rank="species" species="barbarus">
<emphasis id="61B4EA85FFE6FFF7FD69F9F7FD722B06" box="[644,659,1555,1580]" italics="true" pageId="16" pageNumber="251">E</emphasis>
.
<emphasis id="61B4EA85FFE6FFF7FD4FF9F6FD1E2B06" box="[674,767,1554,1580]" italics="true" pageId="16" pageNumber="251">barbarus</emphasis>
</taxonomicName>
,
<taxonomicName id="94C04D14FFE6FFF7FF69F9D7FEE12B67" authorityName="Jaeger" authorityYear="1988" box="[132,256,1587,1613]" class="Mammalia" family="Cricetidae" genus="Ellobius" kingdom="Animalia" order="Rodentia" pageId="16" pageNumber="251" phylum="Chordata" rank="species" species="africanus">
<emphasis id="61B4EA85FFE6FFF7FF69F9D7FF752B66" box="[132,148,1587,1612]" italics="true" pageId="16" pageNumber="251">E</emphasis>
.
<emphasis id="61B4EA85FFE6FFF7FF4CF9D7FEE12B67" box="[161,256,1587,1613]" italics="true" pageId="16" pageNumber="251">africanus</emphasis>
</taxonomicName>
est le plus petit, l’indice d’hypsodontie est faible, les molaires inférieures sont plus étroites, le lobe antérieur de la m1 est large et complexe avec de nombreux replis d’émails, les échancrures d’émail sont peu développées, les M2 et m2 sont nettement moins longues que les M1 et m1.
<taxonomicName id="94C04D14FFE6FFF7FF6EF937FEE92BC6" authorityName="Jaeger" authorityYear="1988" box="[131,264,1746,1772]" class="Mammalia" family="Cricetidae" genus="Ellobius" kingdom="Animalia" order="Rodentia" pageId="16" pageNumber="251" phylum="Chordata" rank="species" species="atlanticus">
<emphasis id="61B4EA85FFE6FFF7FF6EF937FF722BC6" box="[131,147,1747,1772]" italics="true" pageId="16" pageNumber="251">E</emphasis>
.
<emphasis id="61B4EA85FFE6FFF7FF4CF936FEE92BC6" box="[161,264,1746,1772]" italics="true" pageId="16" pageNumber="251">atlanticus</emphasis>
</taxonomicName>
a une taille et un degré d’hypsodontie intermédiaires entre
<taxonomicName id="94C04D14FFE6FFF7FEE3F917FE6C2A26" authorityName="Jaeger" authorityYear="1988" box="[270,397,1778,1804]" class="Mammalia" family="Cricetidae" genus="Ellobius" kingdom="Animalia" order="Rodentia" pageId="16" pageNumber="251" phylum="Chordata" rank="species" species="africanus">
<emphasis id="61B4EA85FFE6FFF7FEE3F917FEFF2A26" box="[270,286,1779,1804]" italics="true" pageId="16" pageNumber="251">E</emphasis>
.
<emphasis id="61B4EA85FFE6FFF7FEC0F916FE6C2A26" box="[301,397,1778,1804]" italics="true" pageId="16" pageNumber="251">africanus</emphasis>
</taxonomicName>
et
<taxonomicName id="94C04D14FFE6FFF7FE40F917FDC92A26" authorityName="Pomel" authorityYear="1892" box="[429,552,1778,1804]" class="Mammalia" family="Cricetidae" genus="Ellobius" kingdom="Animalia" order="Rodentia" pageId="16" pageNumber="251" phylum="Chordata" rank="species" species="barbarus">
<emphasis id="61B4EA85FFE6FFF7FE40F917FE5C2A26" box="[429,445,1779,1804]" italics="true" pageId="16" pageNumber="251">E</emphasis>
.
<emphasis id="61B4EA85FFE6FFF7FE26F916FDC92A26" box="[459,552,1778,1804]" italics="true" pageId="16" pageNumber="251">barbarus</emphasis>
</taxonomicName>
, le lobe antérieur de la m1 est court mais large (étendu du côté lingual), lisse et dépourvu de plissements d’émail, les échancrures d’émail sont peu développées, les M2 sont presque aussi longues que les M1.
<taxonomicName id="94C04D14FFE6FFF7FF36F896FEB72AA1" authorityName="Pomel" authorityYear="1892" box="[219,342,1905,1932]" class="Mammalia" family="Cricetidae" genus="Ellobius" kingdom="Animalia" order="Rodentia" pageId="16" pageNumber="251" phylum="Chordata" rank="species" species="barbarus">
<emphasis id="61B4EA85FFE6FFF7FF36F896FF0A2AA1" box="[219,235,1906,1931]" italics="true" pageId="16" pageNumber="251">E</emphasis>
.
<emphasis id="61B4EA85FFE6FFF7FF14F895FEB72AA1" box="[249,342,1905,1931]" italics="true" pageId="16" pageNumber="251">barbarus</emphasis>
</taxonomicName>
est le plus grand (molaires plus longues et plus larges), l’indice d’hypsodontie est plus important que chez les autres espèces fossiles (mais inférieur aux espèces actuelles), les échancrures d’émail sont relativement élevées, le lobe antérieur de la m1 est simple, étroit et allongé mais porte parfois un ou deux plissements d’émail sur le rebord antéro-labial, la racine linguale de la M1 est réduite, les M3 sont plus réduites (
<bibRefCitation id="37514B66FFE6FFF7FC19FC91FB902EA5" author="JAEGER J. J." box="[1012,1137,885,911]" pageId="16" pageNumber="251" pagination="3 - 50" refId="ref19224" refString="JAEGER J. J. 1988. - Origine et evolution du genre Ellobius (Mammalia, Rodentia) en Afrique nord-occidentale. Folia Quaternaria 57: 3 - 50." type="journal article" year="1988">Jaeger 1988</bibRefCitation>
;
<bibRefCitation id="37514B66FFE6FFF7FB90FC91FAF02EA5" author="GERAADS D." box="[1149,1297,885,911]" pageId="16" pageNumber="251" pagination="147 - 172" refId="ref18174" refString="GERAADS D. 1994. - Rongeurs et Lagomorphes du Pleistocene moyen de la &quot; Grotte des Rhinoceros &quot;, carriere Oulad Hamida 1, a Casablanca, Maroc. Neues Jahrbuch fur Geologie und Paleontologie, Abh 191 (2): 147 - 172." type="journal article" year="1994">Geraads 1994</bibRefCitation>
,
<bibRefCitation id="37514B66FFE6FFF7FACDFC91FA822EA5" author="GERAADS D." box="[1312,1379,885,911]" pageId="16" pageNumber="251" pagination="95 - 104" refId="ref18357" refString="GERAADS D. 2016 a. - La faune de vertebres du Pleistocene moyen de la Grotte des Rhinoceros, Casablanca, Maroc: 4 - Rodentia et Lagomorpha, in RAYNAL J. P. &amp; MOHIB A. (eds), Prehistoire de Casablanca. 1 - La Grotte des Rhinoceros (fouilles 1991 et 1996). Villes et Sites Archeologiques du Maroc, Vol. VI, Rabat: 95 - 104." type="book chapter" year="2016">2016a</bibRefCitation>
). Chez les représentant asiatiques actuels (
<taxonomicName id="94C04D14FFE6FFF7FB75FC72FA4B2E85" authority="Thomas, 1897" authorityName="Thomas" authorityYear="1897" box="[1176,1450,917,943]" class="Mammalia" family="Cricetidae" genus="Ellobius" kingdom="Animalia" order="Rodentia" pageId="16" pageNumber="251" phylum="Chordata" rank="species" species="lutescens">
<emphasis id="61B4EA85FFE6FFF7FB75FC72FB492E85" box="[1176,1192,918,943]" italics="true" pageId="16" pageNumber="251">E</emphasis>
.
<emphasis id="61B4EA85FFE6FFF7FB59FC71FAEB2E85" box="[1204,1290,917,943]" italics="true" pageId="16" pageNumber="251">lutescens</emphasis>
Thomas, 1897
</taxonomicName>
,
<taxonomicName id="94C04D14FFE6FFF7FCC6FC51FBB12EE5" authority="Blyth, 1843" authorityName="Blyth" authorityYear="1843" box="[811,1104,949,975]" class="Mammalia" family="Cricetidae" genus="Ellobius" kingdom="Animalia" order="Rodentia" pageId="16" pageNumber="251" phylum="Chordata" rank="species" species="fuscocapillus">
<emphasis id="61B4EA85FFE6FFF7FCC6FC51FCDA2EE4" box="[811,827,949,974]" italics="true" pageId="16" pageNumber="251">E</emphasis>
.
<emphasis id="61B4EA85FFE6FFF7FCA4FC51FC292EE5" box="[841,968,949,975]" italics="true" pageId="16" pageNumber="251">fuscocapillus</emphasis>
Blyth, 1843
</taxonomicName>
,
<taxonomicName id="94C04D14FFE6FFF7FBBAFC51FAB02EE5" authority="Pallas, 1770" authorityName="Pallas" authorityYear="1770" box="[1111,1361,949,975]" class="Mammalia" family="Cricetidae" genus="Ellobius" kingdom="Animalia" order="Rodentia" pageId="16" pageNumber="251" phylum="Chordata" rank="species" species="talpinus">
<emphasis id="61B4EA85FFE6FFF7FBBAFC51FB862EE4" box="[1111,1127,949,974]" italics="true" pageId="16" pageNumber="251">E</emphasis>
.
<emphasis id="61B4EA85FFE6FFF7FB98FC51FB292EE5" box="[1141,1224,949,975]" italics="true" pageId="16" pageNumber="251">talpinus</emphasis>
Pallas, 1770
</taxonomicName>
), la taille est plus réduite, le degré d’hypsodontie est plus important, les échancrures d’émail sont plus élevées, la proportion des M2 et m2 est moindre, sur la m1 le lobe antérieur est large et plus simple (pas d’incisions labiales ou linguales) et des différences existent également au niveau de l’ouverture des angles rentrants linguaux et de la position respective des triangles (
<bibRefCitation id="37514B66FFE6FFF7FC76FB70FBFC2984" author="JAEGER J. J." box="[923,1053,1172,1198]" pageId="16" pageNumber="251" pagination="3 - 50" refId="ref19224" refString="JAEGER J. J. 1988. - Origine et evolution du genre Ellobius (Mammalia, Rodentia) en Afrique nord-occidentale. Folia Quaternaria 57: 3 - 50." type="journal article" year="1988">Jaeger 1988</bibRefCitation>
).
</paragraph>
<caption id="07BF661FFFE6FFF7FCC0FD44FB8C2FF5" ID-DOI="http://doi.org/10.5281/zenodo.6246039" ID-Zenodo-Dep="6246039" httpUri="https://zenodo.org/record/6246039/files/figure.png" pageId="16" pageNumber="251" startId="16.[813,824,672,690]" targetBox="[827,1441,219,615]" targetPageId="16">
<paragraph id="537F3697FFE6FFF7FCC0FD44FB8C2FF5" blockId="16.[813,1455,672,735]" pageId="16" pageNumber="251">
FIG. 9. — Exemples de molaires de
<taxonomicName id="94C04D14FFE6FFF7FBB4FD44FA902F98" authority="de Ben Kerat" authorityName="de Ben Kerat" box="[1113,1393,672,690]" class="Mammalia" family="Muridae" genus="Praomys" isUncertain="true" kingdom="Animalia" order="Rodentia" pageId="16" pageNumber="251" phylum="Chordata" rank="species" species="pomeli">
<emphasis id="61B4EA85FFE6FFF7FBB4FD44FB432F98" box="[1113,1186,672,690]" italics="true" pageId="16" pageNumber="251">Praomys</emphasis>
cf.
<emphasis id="61B4EA85FFE6FFF7FB2EFD44FB1A2F98" box="[1219,1275,672,690]" italics="true" pageId="16" pageNumber="251">pomeli</emphasis>
de Ben Kérat
</taxonomicName>
en vue occlusale:
<emphasis id="61B4EA85FFE6FFF7FC64FD53FC772FE3" bold="true" box="[905,918,695,713]" pageId="16" pageNumber="251">A</emphasis>
, m1 droite;
<emphasis id="61B4EA85FFE6FFF7FC13FD53FBEA2FE3" bold="true" box="[1022,1035,695,713]" pageId="16" pageNumber="251">B</emphasis>
, m2 gauche;
<emphasis id="61B4EA85FFE6FFF7FB6FFD52FB712FE2" bold="true" box="[1154,1168,694,712]" pageId="16" pageNumber="251">C</emphasis>
, deux m3 droites;
<emphasis id="61B4EA85FFE6FFF7FADCFD53FADE2FE3" bold="true" box="[1329,1343,695,713]" pageId="16" pageNumber="251">D</emphasis>
, m3 gauche;
<emphasis id="61B4EA85FFE6FFF7FCC0FD29FCD82FF5" bold="true" box="[813,825,717,735]" pageId="16" pageNumber="251">E</emphasis>
, M2 droite. Barre d'échelle: 0,5 mm.
</paragraph>
</caption>
<paragraph id="537F3697FFE6FFF7FCAEFB50FB0A2AC1" blockId="16.[811,1457,789,2028]" pageId="16" pageNumber="251">
Au moins deux m1 de Ben Kérat (
<figureCitation id="CBFB2A12FFE6FFF7FB46FB50FB1729E4" box="[1195,1270,1204,1230]" captionStart="FIG" captionStartId="17.[132,143,1985,2003]" captionTargetBox="[132,1455,994,1973]" captionTargetId="figure-18@17.[698,1267,981,1526]" captionTargetPageId="17" captionText="FIG. 11. — Exemples de molaires d’Ellobius de Ben Kérat en vue latérale (A) et occlusale (B-K): A et B, trois m1 gauches; C, m1 droite; D, deux m2 gauches; E, m2 droite; F, deux m3 gauches; G, deux m3 droites; H, M1 gauche; I, M1 droite; J, M3 gauche; K, M3 droite. Barre d'échelle: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.6246047" httpUri="https://zenodo.org/record/6246047/files/figure.png" pageId="16" pageNumber="251">Fig. 11</figureCitation>
) présentent plusieurs indentations sur leur lobe antérieur (la plupart des dents sont abimées sur leur partie antérieure), un caractère qui est habituellement plutôt associé aux formes anciennes (comme
<taxonomicName id="94C04D14FFE6FFF7FCC1FAD0FC492867" authorityName="Jaeger" authorityYear="1988" box="[812,936,1331,1358]" class="Mammalia" family="Cricetidae" genus="Ellobius" kingdom="Animalia" order="Rodentia" pageId="16" pageNumber="251" phylum="Chordata" rank="species" species="africanus">
<emphasis id="61B4EA85FFE6FFF7FCC1FAD0FCDD2867" box="[812,828,1332,1357]" italics="true" pageId="16" pageNumber="251">E</emphasis>
.
<emphasis id="61B4EA85FFE6FFF7FCA7FAD7FC492867" box="[842,936,1331,1357]" italics="true" pageId="16" pageNumber="251">africanus</emphasis>
</taxonomicName>
), bien qu’il se retrouve aussi parfois chez
<taxonomicName id="94C04D14FFE6FFF7FA88FAD0FC1F2844" authority="(Jaeger 1988)" baseAuthorityName="Jaeger" baseAuthorityYear="1988" class="Mammalia" family="Cricetidae" genus="Ellobius" kingdom="Animalia" order="Rodentia" pageId="16" pageNumber="251" phylum="Chordata" rank="species" species="barbarus">
<emphasis id="61B4EA85FFE6FFF7FA88FAD0FA942867" box="[1381,1397,1332,1357]" italics="true" pageId="16" pageNumber="251">E</emphasis>
.
<emphasis id="61B4EA85FFE6FFF7FA6EFAD7FC842847" italics="true" pageId="16" pageNumber="251">barbarus</emphasis>
(
<bibRefCitation id="37514B66FFE6FFF7FC98FAB0FC122847" author="JAEGER J. J." box="[885,1011,1363,1390]" pageId="16" pageNumber="251" pagination="3 - 50" refId="ref19224" refString="JAEGER J. J. 1988. - Origine et evolution du genre Ellobius (Mammalia, Rodentia) en Afrique nord-occidentale. Folia Quaternaria 57: 3 - 50." type="journal article" year="1988">Jaeger 1988</bibRefCitation>
)
</taxonomicName>
. De plus, l’indice d’hypsodontie observé dans le matériel de Ben Kérat est assez modéré (moy 0,83; min0,73; max 1,02), se rapprochant davantage de
<taxonomicName id="94C04D14FFE6FFF7FADBFA70FA512887" authorityName="Jaeger" authorityYear="1988" box="[1334,1456,1427,1454]" class="Mammalia" family="Cricetidae" genus="Ellobius" kingdom="Animalia" order="Rodentia" pageId="16" pageNumber="251" phylum="Chordata" rank="species" species="africanus">
<emphasis id="61B4EA85FFE6FFF7FADBFA70FAA72887" box="[1334,1350,1428,1453]" italics="true" pageId="16" pageNumber="251">E</emphasis>
.
<emphasis id="61B4EA85FFE6FFF7FAB9FA77FA512887" box="[1364,1456,1427,1453]" italics="true" pageId="16" pageNumber="251">africanus</emphasis>
</taxonomicName>
(0,92 à Tighennif) ou
<taxonomicName id="94C04D14FFE6FFF7FBFEFA50FB7528E7" authorityName="Jaeger" authorityYear="1988" box="[1043,1172,1459,1485]" class="Mammalia" family="Cricetidae" genus="Ellobius" kingdom="Animalia" order="Rodentia" pageId="16" pageNumber="251" phylum="Chordata" rank="species" species="atlanticus">
<emphasis id="61B4EA85FFE6FFF7FBFEFA50FBC228E7" box="[1043,1059,1460,1485]" italics="true" pageId="16" pageNumber="251">E</emphasis>
.
<emphasis id="61B4EA85FFE6FFF7FBDCFA57FB7528E7" box="[1073,1172,1459,1485]" italics="true" pageId="16" pageNumber="251">atlanticus</emphasis>
</taxonomicName>
(1,03 à la carrière Thomas) que de
<taxonomicName id="94C04D14FFE6FFF7FC9BFA30FC0F28C7" baseAuthorityName="Jaeger" baseAuthorityYear="1988" box="[886,1006,1491,1517]" class="Mammalia" family="Cricetidae" genus="Ellobius" kingdom="Animalia" order="Rodentia" pageId="16" pageNumber="251" phylum="Chordata" rank="species" species="barbarus">
<emphasis id="61B4EA85FFE6FFF7FC9BFA30FC6728C7" box="[886,902,1492,1517]" italics="true" pageId="16" pageNumber="251">E</emphasis>
.
<emphasis id="61B4EA85FFE6FFF7FC7EFA37FC0F28C7" box="[915,1006,1491,1517]" italics="true" pageId="16" pageNumber="251">barbarus</emphasis>
</taxonomicName>
(1,14 à Aïn Mefta) (
<bibRefCitation id="37514B66FFE6FFF7FB2AFA37FAA528C7" author="JAEGER J. J." box="[1223,1348,1491,1517]" pageId="16" pageNumber="251" pagination="3 - 50" refId="ref19224" refString="JAEGER J. J. 1988. - Origine et evolution du genre Ellobius (Mammalia, Rodentia) en Afrique nord-occidentale. Folia Quaternaria 57: 3 - 50." type="journal article" year="1988">Jaeger 1988</bibRefCitation>
;
<tableCitation id="1E42032CFFE6FFF7FAA3FA37FA7E28C7" box="[1358,1439,1491,1517]" captionStart="TABLE" captionStartId="21.[132,143,219,237]" captionTargetPageId="21" captionText="TABLE 7. — Mesures (en mm) prises sur les mandibules et les dents inférieures et supérieures des crocidures de Ben Kérat. *, mesures approximatives (dent abîmée dans sa partie postéro-labiale)." httpUri="http://table.plazi.org/id/07BF661FFFE3FFF2FF69FF3FFE2B2C2E" pageId="16" pageNumber="251" tableUuid="07BF661FFFE3FFF2FF69FF3FFE2B2C2E">Table 7</tableCitation>
). Il faut cependant garder à l’esprit que la valeur de l’indice d’hypsodontie peut légèrement varier avec l’âge individuel et l’usure dentaire (
<bibRefCitation id="37514B66FFE6FFF7FBEAF9D6FB402B67" author="CHALINE J." box="[1031,1185,1586,1613]" pageId="16" pageNumber="251" refId="ref17758" refString="CHALINE J. 1972. - Les rongeurs du Pleistocene moyen et superieur de France (systematique - biostratigraphie - paleoclimatologie). Cahiers de Paleontologie, Editions du Centre national de la Recherche scientifique, Paris, 410 p." type="book" year="1972">Chaline 1972</bibRefCitation>
). Les spécimens de Ben Kérat présentent de plus une taille particulièrement importante, se situant dans la variabilité haute de
<taxonomicName id="94C04D14FFE6FFF7FB19F997FA8C2BA6" baseAuthorityName="Jaeger" baseAuthorityYear="1988" box="[1268,1389,1650,1677]" class="Mammalia" family="Cricetidae" genus="Ellobius" kingdom="Animalia" order="Rodentia" pageId="16" pageNumber="251" phylum="Chordata" rank="species" species="barbarus">
<emphasis id="61B4EA85FFE6FFF7FB19F997FAE52BA6" box="[1268,1284,1651,1676]" italics="true" pageId="16" pageNumber="251">E</emphasis>
.
<emphasis id="61B4EA85FFE6FFF7FAFFF996FA8C2BA6" box="[1298,1389,1650,1676]" italics="true" pageId="16" pageNumber="251">barbarus</emphasis>
</taxonomicName>
, voire au-delà (
<tableCitation id="1E42032CFFE6FFF7FC61F976FC3C2B87" box="[908,989,1682,1709]" captionStart="TABLE" captionStartId="19.[385,395,219,237]" captionTargetPageId="19" captionText="TABLE 5. — Mesures de longueur et largeur (en mm) des molaires inférieures et supérieures des El-" httpUri="http://table.plazi.org/id/07BF661FFFE5FFF4FE6CFF3FFDCA2C29" pageId="16" pageNumber="251" tableUuid="07BF661FFFE5FFF4FE6CFF3FFDCA2C29">Table 5</tableCitation>
;
<figureCitation id="CBFB2A12FFE6FFF7FC00F976FBDB2B86" box="[1005,1082,1682,1708]" captionStart="FIG" captionStartId="18.[132,143,1603,1621]" captionTargetBox="[178,1405,218,1563]" captionTargetId="graphics-142@18.[264,1122,266,1491]" captionTargetPageId="18" captionText="FIG. 12. — Replacement des premières molaires supérieures (A) et inférieures (B) d’Ellobius de Ben Kérat au sein de la variabilité de taille d’autres Ellobius fossiles du Maghreb (Jaeger 1988;Geraads 1994, 2016a).Abréviations: Actuel, spécimens actuels d’Iran; BR2, Bulla Regia 2; IDV, Irhoud Derbala Virage; IL3, Irhoud Lanz 3; Mef, Aïn Mefta; OH1, Oulad Hamida 1; SA1, Sidi Abderrahmane 1; Salé, Salé; Tadj, Tadjera; Th, carrière Thomas; Tig, Tighennif. Barres d’erreur: écart-type." figureDoi="http://doi.org/10.5281/zenodo.6246051" httpUri="https://zenodo.org/record/6246051/files/figure.png" pageId="16" pageNumber="251">Fig. 12</figureCitation>
), les m2 et m3 sont peu réduites, et les molaires inférieures sont relativement larges (
<tableCitation id="1E42032CFFE6FFF7FAA2F956FA7E2BE6" box="[1359,1439,1714,1740]" captionStart="TABLE" captionStartId="19.[132,143,516,534]" captionTargetPageId="19" captionText="TABLE 6. — Proportion relative des molaires (% par rapport à la longueur de la M1 ou m1), rapport de la largeur sur la longueur de la m1 (l/L m1) et indice d’hypsodontie (Ih max) des Ellobius fossiles du Maghreb incluant le matériel de Ben Kérat (données de référence issues de Jaeger 1988)." httpUri="http://table.plazi.org/id/07BF661FFFE5FFF4FF69FDE0FB1D2F07" pageId="16" pageNumber="251" tableUuid="07BF661FFFE5FFF4FF69FDE0FB1D2F07">Table 6</tableCitation>
). Ainsi, les restes de Ben Kérat se rapprochent davantage de
<taxonomicName id="94C04D14FFE6FFF7FCC3F917FC472A26" baseAuthorityName="Jaeger" baseAuthorityYear="1988" box="[814,934,1778,1804]" class="Mammalia" family="Cricetidae" genus="Ellobius" kingdom="Animalia" order="Rodentia" pageId="16" pageNumber="251" phylum="Chordata" rank="species" species="barbarus">
<emphasis id="61B4EA85FFE6FFF7FCC3F917FCDD2A26" box="[814,828,1779,1804]" italics="true" pageId="16" pageNumber="251">E</emphasis>
.
<emphasis id="61B4EA85FFE6FFF7FCA7F916FC472A26" box="[842,934,1778,1804]" italics="true" pageId="16" pageNumber="251">barbarus</emphasis>
</taxonomicName>
, malgré un indice d’hypsodontie plus modéré et une taille un peu plus importante. Devant la faible quantité de matériel et son mauvais état de préservation, nous préférons rester prudents et limiter l’attribution à
<taxonomicName id="94C04D14FFE6FFF7FAE3F8B6FA492A41" baseAuthorityName="Jaeger" baseAuthorityYear="1988" box="[1294,1448,1873,1900]" class="Mammalia" family="Cricetidae" genus="Ellobius" isUncertain="true" kingdom="Animalia" order="Rodentia" pageId="16" pageNumber="251" phylum="Chordata" rank="species" species="barbarus">
<emphasis id="61B4EA85FFE6FFF7FAE3F8B6FAFF2A41" box="[1294,1310,1874,1899]" italics="true" pageId="16" pageNumber="251">E</emphasis>
. cf.
<emphasis id="61B4EA85FFE6FFF7FAA1F8B5FA492A41" box="[1356,1448,1873,1899]" italics="true" pageId="16" pageNumber="251">barbarus</emphasis>
</taxonomicName>
. Les différences observées pourraient aller dans le sens de l’hypothèse de «différentiations géographiques» liées à des conditions climatiques locales particulières déjà évoquée par
<bibRefCitation id="37514B66FFE6FFF7FCC1F836FC5F2AC6" author="JAEGER J. J." box="[812,958,2001,2028]" pageId="16" pageNumber="251" pagination="3 - 50" refId="ref19224" refString="JAEGER J. J. 1988. - Origine et evolution du genre Ellobius (Mammalia, Rodentia) en Afrique nord-occidentale. Folia Quaternaria 57: 3 - 50." type="journal article" year="1988">Jaeger (1988)</bibRefCitation>
pour
<taxonomicName id="94C04D14FFE6FFF7FC12F836FBBA2AC1" authorityName="Jaeger" authorityYear="1988" box="[1023,1115,2002,2028]" class="Mammalia" family="Cricetidae" genus="Ellobius" kingdom="Animalia" order="Rodentia" pageId="16" pageNumber="251" phylum="Chordata" rank="species" species="zimae">
<emphasis id="61B4EA85FFE6FFF7FC12F836FBEE2AC1" box="[1023,1039,2002,2027]" italics="true" pageId="16" pageNumber="251">E</emphasis>
.
<emphasis id="61B4EA85FFE6FFF7FBF0F836FBBA2AC1" box="[1053,1115,2002,2027]" italics="true" pageId="16" pageNumber="251">zimae</emphasis>
</taxonomicName>
par exemple.
</paragraph>
</subSubSection>
</treatment>
</document>
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