250 lines
37 KiB
XML
250 lines
37 KiB
XML
<document id="DEA8179ED34B3DBADB38B701507EF8A1" ID-CLB-Dataset="54981" ID-DOI="10.5252/geodiversitas2022v44a8" ID-GBIF-Dataset="7e93094c-b8b7-4077-9a4e-5b3c0e65f952" ID-ISSN="1638-9395" ID-Zenodo-Dep="6245987" ID-ZooBank="urn:lsid:zoobank.org:pub:E7490671-2900-4DE3-8E2E-D6921D80C6D1" IM.bibliography_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1645530114998" checkinUser="felipe" docAuthor="Stoetzel, Emmanuelle & Pickford, Martin" docDate="2022" docId="DB698781FFE5FFF2FF2FFA56FCE02B67" docLanguage="fr" docName="Geodiversitas.44.8.237-263.pdf" docOrigin="Geodiversitas 44 (8)" docStyle="DocumentStyle:F830B10FF475E64C1F1601E3B32DDC00.4:Geodiversitas.2018-.journal_article" docStyleId="F830B10FF475E64C1F1601E3B32DDC00" docStyleName="Geodiversitas.2018-.journal_article" docStyleVersion="4" docTitle="Crocidura maghrebiana Hutterer 1991" docType="treatment" docVersion="9" lastPageNumber="254" masterDocId="2750FFF9FFF6FFE7FFEDFFE4FFE12D2A" masterDocTitle="Étude d’un assemblage original de microvertébrés du Pléistocène moyen du nord-est de l’Algérie (Ben Kérat, Oued Zenati) et description de deux nouveaux muridés" masterLastPageNumber="263" masterPageNumber="237" pageNumber="254" updateTime="1699316018958" updateUser="ExternalLinkService" zenodo-license-document="CC0-1.0" zenodo-license-figures="CC0-1.0">
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<mods:title id="43DC3F4F33E0BE9BA258A61373BDD3E0">Étude d’un assemblage original de microvertébrés du Pléistocène moyen du nord-est de l’Algérie (Ben Kérat, Oued Zenati) et description de deux nouveaux muridés</mods:title>
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<mods:namePart id="B8AE3AA2C9AA097D6940DB363144563E">Stoetzel, Emmanuelle</mods:namePart>
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<mods:namePart id="5E0B1E9EF472E08A743304FA95543B58">Pickford, Martin</mods:namePart>
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<mods:title id="8EA11A12D9444A214EB872FF6E3FA3F1">Geodiversitas</mods:title>
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<mods:date id="952482D6031B8F59CD243EFA0A68D5C9">2022</mods:date>
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<treatment id="DB698781FFE5FFF2FF2FFA56FCE02B67" ID-DOI="http://doi.org/10.5281/zenodo.6315547" ID-GBIF-Taxon="194518613" ID-Zenodo-Dep="6315547" LSID="urn:lsid:plazi:treatment:DB698781FFE5FFF2FF2FFA56FCE02B67" httpUri="http://treatment.plazi.org/id/DB698781FFE5FFF2FF2FFA56FCE02B67" lastPageId="21" lastPageNumber="254" pageId="19" pageNumber="254">
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<subSubSection id="1BDA651CFFE5FFF4FF2FFA56FD2828E7" box="[194,713,1458,1485]" pageId="19" pageNumber="254" type="nomenclature">
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<paragraph id="537F3697FFE5FFF4FF2FFA56FD2828E7" blockId="19.[194,713,1458,1485]" box="[194,713,1458,1485]" pageId="19" pageNumber="254">
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<treatmentCitation id="D2611086FFE5FFF4FF2FFA56FD2828E7" author="HUTTERER R." box="[194,713,1458,1485]" page="249" pageId="19" pageNumber="254" year="1991">
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<taxonomicName id="94C04D14FFE5FFF4FF2FFA56FD2828E7" authority="Hutterer, 1991: 249" authorityName="Hutterer" authorityPageNumber="249" authorityYear="1991" box="[194,713,1458,1485]" class="Mammalia" family="Soricidae" genus="Crocidura" isUncertain="true" kingdom="Animalia" order="Soricomorpha" pageId="19" pageNumber="254" phylum="Chordata" rank="species" species="maghrebiana">
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<emphasis id="61B4EA85FFE5FFF4FF2FFA56FE0928E7" bold="true" box="[194,488,1458,1485]" italics="true" pageId="19" pageNumber="254">Crocidura cf. maghrebiana</emphasis>
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<bibRefCitation id="37514B66FFE5FFF4FE02FA57FD2828E7" author="HUTTERER R." box="[495,713,1459,1485]" pageId="19" pageNumber="254" pagination="241 - 251" refId="ref19131" refString="HUTTERER R. 1991. - Variation and evolution of the Sicilian shrew: taxonomic conclusions and description of a possibly related species from the Pleistocene of Morocco (Mammalia: Soricidae). Bonner Zoologische Beitrage 42 (3 - 4): 241 - 251." type="journal article" year="1991">Hutterer, 1991: 249</bibRefCitation>
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</taxonomicName>
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</treatmentCitation>
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</paragraph>
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</subSubSection>
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<subSubSection id="1BDA651CFFE5FFF4FF69FA08FD682B2E" box="[132,649,1516,1540]" pageId="19" pageNumber="254" type="reference_group">
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<paragraph id="537F3697FFE5FFF4FF69FA08FD682B2E" blockId="19.[132,649,1516,1540]" box="[132,649,1516,1540]" pageId="19" pageNumber="254">
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<treatmentCitationGroup id="73D011B9FFE5FFF4FF69FA08FD682B2E" box="[132,649,1516,1540]" pageId="19" pageNumber="254">
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<treatmentCitation id="D2611086FFE5FFF4FF69FA08FDDC2B2E" author="GERAADS D." box="[132,573,1516,1540]" page="282" pageId="19" pageNumber="254" year="1993">
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<taxonomicName id="94C04D14FFE5FFF4FF69FA08FDDC2B2E" authority="Geraads, 1993 : 282" authorityName="Geraads" authorityPageNumber="282" authorityYear="1993" box="[132,573,1516,1540]" class="Mammalia" family="Soricidae" genus="Crocidura" isUncertain="true" kingdom="Animalia" order="Soricomorpha" pageId="19" pageNumber="254" phylum="Chordata" rank="species" species="darelbeidae">
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<emphasis id="61B4EA85FFE5FFF4FF69FA08FE912B2E" box="[132,368,1516,1540]" italics="true" pageId="19" pageNumber="254">Crocidura cf. darelbeidae</emphasis>
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<bibRefCitation id="37514B66FFE5FFF4FE9AFA08FDDC2B2E" author="GERAADS D." box="[375,573,1516,1540]" pageId="19" pageNumber="254" pagination="281 - 294" refId="ref18131" refString="GERAADS D. 1993. - Middle Pleistocene Crocidura (Mammalia, Insectivora) from Oulad Hamida I, Morocco, and their phylogenetic relationships. Proceedings of the Koninklijke Nederlandse Akademie van Wetenschappen 96 (3): 281 - 294." type="journal article" year="1993">Geraads, 1993: 282</bibRefCitation>
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</taxonomicName>
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</treatmentCitation>
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,
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<emphasis id="61B4EA85FFE5FFF4FDAAFA08FD682B2E" bold="true" box="[583,649,1516,1540]" pageId="19" pageNumber="254">
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<taxonomicNameLabel id="7A8757FEFFE5FFF4FDAAFA08FD682B2E" box="[583,649,1516,1540]" pageId="19" pageNumber="254">n. syn.</taxonomicNameLabel>
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</emphasis>
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</treatmentCitationGroup>
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</paragraph>
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</subSubSection>
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<subSubSection id="1BDA651CFFE5FFF4FF69F9F8FCE82B92" pageId="19" pageNumber="254" type="materials_examined">
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<paragraph id="537F3697FFE5FFF4FF69F9F8FCE82B92" blockId="19.[131,777,1563,1720]" pageId="19" pageNumber="254">
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<materialsCitation id="E3A83CCAFFE5FFF4FF69F9F8FCE82B92" ID-GBIF-Occurrence="3748611303" pageId="19" pageNumber="254" specimenCount="1">MATÉRIEL. — NR = 15; cinq fragments de mandibules G (trois édentées, une avec m1-m2 en place, une avec m2-m3 en place), un fragment de mandibule D (avec m1-m2 en place), deux fragments de maxillaire G (un avec P4 en place, un avec M1 en place), une unicuspide isolée (u2?), deux m1 G, une m1 D, un fragment de m2 G, une incisive inférieure G et une incisive supérieure D isolées.</materialsCitation>
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</paragraph>
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||
</subSubSection>
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<subSubSection id="1BDA651CFFE5FFF2FF69F916FCE02B67" lastPageId="21" lastPageNumber="256" pageId="19" pageNumber="254" type="description">
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<paragraph id="537F3697FFE5FFF4FF69F916FE012A20" blockId="19.[131,775,1778,2028]" box="[132,480,1778,1804]" pageId="19" pageNumber="254">DESCRIPTION ET COMPARAISONS</paragraph>
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<paragraph id="537F3697FFE5FFF4FF69F8F6FD102AC6" blockId="19.[131,775,1778,2028]" pageId="19" pageNumber="254">
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D’après certains auteurs,
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<taxonomicName id="94C04D14FFE5FFF4FE7CF8F6FD292A06" authority="Geraads, 1993" authorityName="Geraads" authorityYear="1993" box="[401,712,1810,1836]" class="Mammalia" family="Soricidae" genus="Crocidura" kingdom="Animalia" order="Soricomorpha" pageId="19" pageNumber="254" phylum="Chordata" rank="species" species="darelbeidae">
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<emphasis id="61B4EA85FFE5FFF4FE7CF8F6FE422A06" box="[401,419,1810,1836]" italics="true" pageId="19" pageNumber="254">C</emphasis>
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.
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<emphasis id="61B4EA85FFE5FFF4FE5CF8F6FDC72A06" box="[433,550,1810,1836]" italics="true" pageId="19" pageNumber="254">darelbeidae</emphasis>
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<bibRefCitation id="37514B66FFE5FFF4FDC1F8F6FD292A06" author="GERAADS D." box="[556,712,1810,1836]" pageId="19" pageNumber="254" pagination="281 - 294" refId="ref18131" refString="GERAADS D. 1993. - Middle Pleistocene Crocidura (Mammalia, Insectivora) from Oulad Hamida I, Morocco, and their phylogenetic relationships. Proceedings of the Koninklijke Nederlandse Akademie van Wetenschappen 96 (3): 281 - 294." type="journal article" year="1993">Geraads, 1993</bibRefCitation>
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</taxonomicName>
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serait à mettre en synonymie avec
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<taxonomicName id="94C04D14FFE5FFF4FE5BF8D6FCE62A66" authority="Hutterer, 1991" authorityName="Hutterer" authorityYear="1991" box="[438,775,1841,1868]" class="Mammalia" family="Soricidae" genus="Crocidura" kingdom="Animalia" order="Soricomorpha" pageId="19" pageNumber="254" phylum="Chordata" rank="species" species="maghrebiana">
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<emphasis id="61B4EA85FFE5FFF4FE5BF8D6FE292A66" box="[438,456,1842,1868]" italics="true" pageId="19" pageNumber="254">C</emphasis>
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.
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<emphasis id="61B4EA85FFE5FFF4FE3BF8D5FDBC2A61" box="[470,605,1841,1867]" italics="true" pageId="19" pageNumber="254">maghrebiana</emphasis>
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<bibRefCitation id="37514B66FFE5FFF4FD88F8D6FCE62A66" author="HUTTERER R." box="[613,775,1842,1868]" pageId="19" pageNumber="254" pagination="241 - 251" refId="ref19131" refString="HUTTERER R. 1991. - Variation and evolution of the Sicilian shrew: taxonomic conclusions and description of a possibly related species from the Pleistocene of Morocco (Mammalia: Soricidae). Bonner Zoologische Beitrage 42 (3 - 4): 241 - 251." type="journal article" year="1991">Hutterer, 1991</bibRefCitation>
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||
</taxonomicName>
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(
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<bibRefCitation id="37514B66FFE5FFF4FF60F8B5FEF02A46" author="BUTLER P. M." box="[141,273,1873,1900]" pageId="19" pageNumber="254" pagination="121 - 132" refId="ref17710" refString="BUTLER P. M. 1998. - Fossil history of shrews in Africa, in WOJCIK J. M. & WOLSAN M. (eds), Evolution of Shrews. Mammal Research Institute, Polish Academy of Sciences, Bialowieza: 121 - 132." type="book chapter" year="1998">Butler 1998</bibRefCitation>
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||
;
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<bibRefCitation id="37514B66FFE5FFF4FECDF8B5FE252A46" author="VERGILINO M." box="[288,452,1873,1900]" pageId="19" pageNumber="254" refId="ref21319" refString="VERGILINO M. 1999. - Les peuplements de Soricidae (Lypotyphla, Mammalia) du Maghreb depuis le Miocene. Memoire de DEA, Universite de Montpellier, France, 33 p." type="book" year="1999">Vergilino 1999</bibRefCitation>
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||
;
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<bibRefCitation id="37514B66FFE5FFF4FE3AF8B5FD9F2A41" author="GERAADS D." box="[471,638,1873,1899]" pageId="19" pageNumber="254" pagination="105 - 110" refId="ref18435" refString="GERAADS D. 2016 b. - La faune de vertebres du Pleistocene moyen de la Grotte des Rhinoceros, Casablanca, Maroc: 5 - Insectivores, in RAYNAL J. P. & MOHIB A. (eds), Prehistoire de Casablanca. 1 - La Grotte des Rhinoceros (fouilles 1991 et 1996). Villes et Sites Archeologiques du Maroc, Vol. VI, Rabat: 105 - 110." type="book chapter" year="2016">Geraads 2016b</bibRefCitation>
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) et nous les désignons donc sous le terme
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<taxonomicName id="94C04D14FFE5FFF4FE24F896FD902AA1" authorityName="Hutterer" authorityYear="1991" box="[457,625,1905,1932]" class="Mammalia" family="Soricidae" genus="Crocidura" kingdom="Animalia" order="Soricomorpha" pageId="19" pageNumber="254" phylum="Chordata" rank="species" species="maghrebiana">
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<emphasis id="61B4EA85FFE5FFF4FE24F896FD902AA1" box="[457,625,1905,1932]" italics="true" pageId="19" pageNumber="254">C. maghrebiana</emphasis>
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</taxonomicName>
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. Elles présentent une grande taille, proche de l’actuelle
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<taxonomicName id="94C04D14FFE5FFF4FDACF876FD7E2A81" authorityName="Geoffroy Saint-Hilaire" authorityYear="1834" box="[577,671,1938,1964]" class="Mammalia" family="Soricidae" genus="Crocidura" kingdom="Animalia" order="Soricomorpha" pageId="19" pageNumber="254" phylum="Chordata" rank="species" species="viaria">
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<emphasis id="61B4EA85FFE5FFF4FDACF876FDB22A86" box="[577,595,1938,1964]" italics="true" pageId="19" pageNumber="254">C</emphasis>
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.
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<emphasis id="61B4EA85FFE5FFF4FD8CF876FD7E2A81" box="[609,671,1938,1963]" italics="true" pageId="19" pageNumber="254">viaria</emphasis>
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</taxonomicName>
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, bien que certains caractères dentaires et métriques les séparent (
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<bibRefCitation id="37514B66FFE5FFF4FD58F855FED62AC1" author="KOWALSKA B." pageId="19" pageNumber="254" pagination="51 - 90" refId="ref20332" refString="RZEBIK- KOWALSKA B. 1988 b. - Soricidae (Mammalia, Insectivora) from the Plio-Pleistocene and Middle Quaternary of Morocco and Algeria. Folia Quaternaria 57: 51 - 90." type="journal article" year="1988">Rzebik-Kowalska 1988b</bibRefCitation>
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||
;
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<bibRefCitation id="37514B66FFE5FFF4FEABF836FE032AC1" author="HUTTERER R." box="[326,482,2001,2028]" pageId="19" pageNumber="254" pagination="241 - 251" refId="ref19131" refString="HUTTERER R. 1991. - Variation and evolution of the Sicilian shrew: taxonomic conclusions and description of a possibly related species from the Pleistocene of Morocco (Mammalia: Soricidae). Bonner Zoologische Beitrage 42 (3 - 4): 241 - 251." type="journal article" year="1991">Hutterer 1991</bibRefCitation>
|
||
;
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||
<bibRefCitation id="37514B66FFE5FFF4FE19F835FD6B2AC1" author="GERAADS D." box="[500,650,2001,2027]" pageId="19" pageNumber="254" pagination="281 - 294" refId="ref18131" refString="GERAADS D. 1993. - Middle Pleistocene Crocidura (Mammalia, Insectivora) from Oulad Hamida I, Morocco, and their phylogenetic relationships. Proceedings of the Koninklijke Nederlandse Akademie van Wetenschappen 96 (3): 281 - 294." type="journal article" year="1993">Geraads 1993</bibRefCitation>
|
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,
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<bibRefCitation id="37514B66FFE5FFF4FD76F835FD032AC1" author="GERAADS D." box="[667,738,2001,2027]" pageId="19" pageNumber="254" pagination="105 - 110" refId="ref18435" refString="GERAADS D. 2016 b. - La faune de vertebres du Pleistocene moyen de la Grotte des Rhinoceros, Casablanca, Maroc: 5 - Insectivores, in RAYNAL J. P. & MOHIB A. (eds), Prehistoire de Casablanca. 1 - La Grotte des Rhinoceros (fouilles 1991 et 1996). Villes et Sites Archeologiques du Maroc, Vol. VI, Rabat: 105 - 110." type="book chapter" year="2016">2016b</bibRefCitation>
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).
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</paragraph>
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<paragraph id="537F3697FFE5FFF2FCAEFC92FCE02B67" blockId="19.[810,1457,885,2028]" lastBlockId="21.[129,777,1236,1613]" lastPageId="21" lastPageNumber="256" pageId="19" pageNumber="254">
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À Ben Kérat, deux fragments de mandibules, robustes et de grande taille, se rapprochent du groupe
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<emphasis id="61B4EA85FFE5FFF4FB19FC71FC8F2EE4" italics="true" pageId="19" pageNumber="254">
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<taxonomicName id="94C04D14FFE5FFF4FB19FC71FA7C2E85" authorityName="Hutterer" authorityYear="1991" box="[1268,1437,917,943]" class="Mammalia" family="Soricidae" genus="Crocidura" kingdom="Animalia" order="Soricomorpha" pageId="19" pageNumber="254" phylum="Chordata" rank="species" species="maghrebiana">C. maghrebiana</taxonomicName>
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–
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<taxonomicName id="94C04D14FFE5FFF4FCC0FC51FC8F2EE4" authorityName="Geoffroy Saint-Hilaire" authorityYear="1834" box="[813,878,949,974]" class="Mammalia" family="Soricidae" genus="Crocidura" kingdom="Animalia" order="Soricomorpha" pageId="19" pageNumber="254" phylum="Chordata" rank="species" species="viaria">viaria</taxonomicName>
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</emphasis>
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(
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<tableCitation id="1E42032CFFE5FFF4FC6CFC51FC372EE5" box="[897,982,949,975]" captionStart="TABLE" captionStartId="21.[132,143,219,237]" captionTargetPageId="21" captionText="TABLE 7. — Mesures (en mm) prises sur les mandibules et les dents inférieures et supérieures des crocidures de Ben Kérat. *, mesures approximatives (dent abîmée dans sa partie postéro-labiale)." httpUri="http://table.plazi.org/id/07BF661FFFE3FFF2FF69FF3FFE2B2C2E" pageId="19" pageNumber="254" tableUuid="07BF661FFFE3FFF2FF69FF3FFE2B2C2E">Table 7</tableCitation>
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;
|
||
<figureCitation id="CBFB2A12FFE5FFF4FC04FC51FBDB2EE5" box="[1001,1082,949,975]" captionStart="FIG" captionStartId="20.[132,143,1917,1935]" captionTargetBox="[133,1456,205,1881]" captionTargetId="figure-27@20.[699,1268,762,1318]" captionTargetPageId="20" captionText="FIG. 13. — Exemples de restes de Crocidura cf. maghrebiana de Ben Kérat: A, fragment de mandibule gauche en vue linguale et labiale, et condyle en vue postérieure; B, fragment de mandibule droite en vue labiale et linguale, et m1-m2 en vue occlusale; C, fragment de mandibule gauche en vue linguale et labiale, et m2-m3 en place en vue occlusale; D, incisive inférieure gauche en vue labiale et linguale; E, incisive supérieure droite en vue linguale et labiale; F, m1 droite en vue occlusale et labiale; G, fragment de maxillaire gauche avec M1 en place en vue linguale et occlusale; H, fragment de maxillaire gauche avec P4 en place en vue labiale, occlusale et linguale. Barres d'échelle: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.6246056" httpUri="https://zenodo.org/record/6246056/files/figure.png" pageId="19" pageNumber="254">Fig. 13</figureCitation>
|
||
). Le condyle en vue postérieure présente une forme en «L », avec la partie interne de la facette inférieure assez longue et courbée vers le bas, ce qui la rapproche de
|
||
<taxonomicName id="94C04D14FFE5FFF4FC0AFBF1FB722904" authorityName="Hutterer" authorityYear="1991" box="[999,1171,1044,1071]" class="Mammalia" family="Soricidae" genus="Crocidura" kingdom="Animalia" order="Soricomorpha" pageId="19" pageNumber="254" phylum="Chordata" rank="species" species="maghrebiana">
|
||
<emphasis id="61B4EA85FFE5FFF4FC0AFBF1FB722904" box="[999,1171,1044,1071]" italics="true" pageId="19" pageNumber="254">C. maghrebiana</emphasis>
|
||
</taxonomicName>
|
||
, alors que chez
|
||
<taxonomicName id="94C04D14FFE5FFF4FAA3FBF1FA4E2904" authorityName="Geoffroy Saint-Hilaire" authorityYear="1834" box="[1358,1455,1045,1071]" class="Mammalia" family="Soricidae" genus="Crocidura" kingdom="Animalia" order="Soricomorpha" pageId="19" pageNumber="254" phylum="Chordata" rank="species" species="viaria">
|
||
<emphasis id="61B4EA85FFE5FFF4FAA3FBF1FABE2905" box="[1358,1375,1045,1071]" italics="true" pageId="19" pageNumber="254">C</emphasis>
|
||
.
|
||
<emphasis id="61B4EA85FFE5FFF4FA82FBF1FA4E2904" box="[1391,1455,1045,1070]" italics="true" pageId="19" pageNumber="254">viaria</emphasis>
|
||
</taxonomicName>
|
||
la facette inférieure apparaît plus courte et transversale (
|
||
<bibRefCitation id="37514B66FFE5FFF4FCDBFBB1FC382944" author="HUTTERER R." box="[822,985,1108,1135]" pageId="19" pageNumber="254" pagination="521 - 534" refId="ref19093" refString="HUTTERER R. 1986. - The species of Crocidura (Soricidae) in Morocco. Mammalia 50 (4): 521 - 534. https: // doi. org / 10.1515 / mamm. 1986.50.4.521" type="journal article" year="1986">Hutterer 1986</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="37514B66FFE5FFF4FC06FBB0FB672944" author="GERAADS D." box="[1003,1158,1108,1134]" pageId="19" pageNumber="254" pagination="281 - 294" refId="ref18131" refString="GERAADS D. 1993. - Middle Pleistocene Crocidura (Mammalia, Insectivora) from Oulad Hamida I, Morocco, and their phylogenetic relationships. Proceedings of the Koninklijke Nederlandse Akademie van Wetenschappen 96 (3): 281 - 294." type="journal article" year="1993">Geraads 1993</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="37514B66FFE5FFF4FB77FBB0FB032944" author="GERAADS D." box="[1178,1250,1108,1134]" pageId="19" pageNumber="254" pagination="105 - 110" refId="ref18435" refString="GERAADS D. 2016 b. - La faune de vertebres du Pleistocene moyen de la Grotte des Rhinoceros, Casablanca, Maroc: 5 - Insectivores, in RAYNAL J. P. & MOHIB A. (eds), Prehistoire de Casablanca. 1 - La Grotte des Rhinoceros (fouilles 1991 et 1996). Villes et Sites Archeologiques du Maroc, Vol. VI, Rabat: 105 - 110." type="book chapter" year="2016">2016b</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="37514B66FFE5FFF4FB1EFBB0FA7A2944" author="VERGILINO M." box="[1267,1435,1108,1134]" pageId="19" pageNumber="254" refId="ref21319" refString="VERGILINO M. 1999. - Les peuplements de Soricidae (Lypotyphla, Mammalia) du Maghreb depuis le Miocene. Memoire de DEA, Universite de Montpellier, France, 33 p." type="book" year="1999">Vergilino 1999</bibRefCitation>
|
||
). L’incisive inférieure de Ben Kérat se rapproche également de
|
||
<taxonomicName id="94C04D14FFE5FFF4FCC6FB70FC372984" authorityName="Hutterer" authorityYear="1991" box="[811,982,1172,1198]" class="Mammalia" family="Soricidae" genus="Crocidura" kingdom="Animalia" order="Soricomorpha" pageId="19" pageNumber="254" phylum="Chordata" rank="species" species="maghrebiana">
|
||
<emphasis id="61B4EA85FFE5FFF4FCC6FB70FC372984" box="[811,982,1172,1198]" italics="true" pageId="19" pageNumber="254">C. maghrebiana</emphasis>
|
||
</taxonomicName>
|
||
, avec un bord supérieur presque rectiligne, et un sillon lingual positionné plus ventralement que chez l’actuelle
|
||
<taxonomicName id="94C04D14FFE5FFF4FC77FB30FC1F29C7" authorityName="Geoffroy Saint-Hilaire" authorityYear="1834" box="[922,1022,1236,1262]" class="Mammalia" family="Soricidae" genus="Crocidura" kingdom="Animalia" order="Soricomorpha" pageId="19" pageNumber="254" phylum="Chordata" rank="species" species="viaria">
|
||
<emphasis id="61B4EA85FFE5FFF4FC77FB30FC4D29C4" box="[922,940,1236,1262]" italics="true" pageId="19" pageNumber="254">C</emphasis>
|
||
.
|
||
<emphasis id="61B4EA85FFE5FFF4FC50FB30FC1F29C7" box="[957,1022,1236,1261]" italics="true" pageId="19" pageNumber="254">viaria</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<bibRefCitation id="37514B66FFE5FFF4FBFCFB30FB4E29C4" author="GERAADS D." box="[1041,1199,1236,1262]" pageId="19" pageNumber="254" pagination="281 - 294" refId="ref18131" refString="GERAADS D. 1993. - Middle Pleistocene Crocidura (Mammalia, Insectivora) from Oulad Hamida I, Morocco, and their phylogenetic relationships. Proceedings of the Koninklijke Nederlandse Akademie van Wetenschappen 96 (3): 281 - 294." type="journal article" year="1993">Geraads 1993</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="37514B66FFE5FFF4FB2DFB30FAE929C4" author="GERAADS D." box="[1216,1288,1236,1262]" pageId="19" pageNumber="254" pagination="105 - 110" refId="ref18435" refString="GERAADS D. 2016 b. - La faune de vertebres du Pleistocene moyen de la Grotte des Rhinoceros, Casablanca, Maroc: 5 - Insectivores, in RAYNAL J. P. & MOHIB A. (eds), Prehistoire de Casablanca. 1 - La Grotte des Rhinoceros (fouilles 1991 et 1996). Villes et Sites Archeologiques du Maroc, Vol. VI, Rabat: 105 - 110." type="book chapter" year="2016">2016b</bibRefCitation>
|
||
). Les molaires inférieures de Ben Kérat ne présentent pas de caractère particulier: le cingulum labial est épais et plus ou moins onduleux sur toute la longueur des dents, mais le cingulum lingual est incomplet, souvent limité au paraconide; les m1 et m2 présentent un talonide fermé postéro-lingualement, l’hypolophide ne rejoint pas l’entoconide; la m3 montre un talonide réduit dépourvu d’entoconide. L’incisive supérieure est massive, avec un talon épais, un cingulum labial bien marqué, et une forme globale qui se rapproche de
|
||
<taxonomicName id="94C04D14FFE5FFF4FCC0F9F7FC382B06" authorityName="Hutterer" authorityYear="1991" box="[813,985,1554,1581]" class="Mammalia" family="Soricidae" genus="Crocidura" kingdom="Animalia" order="Soricomorpha" pageId="19" pageNumber="254" phylum="Chordata" rank="species" species="maghrebiana">
|
||
<emphasis id="61B4EA85FFE5FFF4FCC0F9F7FC382B06" box="[813,985,1554,1581]" italics="true" pageId="19" pageNumber="254">C. maghrebiana</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<bibRefCitation id="37514B66FFE5FFF4FC06F9F6FB672B07" author="GERAADS D." box="[1003,1158,1554,1581]" pageId="19" pageNumber="254" pagination="281 - 294" refId="ref18131" refString="GERAADS D. 1993. - Middle Pleistocene Crocidura (Mammalia, Insectivora) from Oulad Hamida I, Morocco, and their phylogenetic relationships. Proceedings of the Koninklijke Nederlandse Akademie van Wetenschappen 96 (3): 281 - 294." type="journal article" year="1993">Geraads 1993</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="37514B66FFE5FFF4FB7BF9F6FB3F2B06" author="GERAADS D." box="[1174,1246,1554,1580]" pageId="19" pageNumber="254" pagination="105 - 110" refId="ref18435" refString="GERAADS D. 2016 b. - La faune de vertebres du Pleistocene moyen de la Grotte des Rhinoceros, Casablanca, Maroc: 5 - Insectivores, in RAYNAL J. P. & MOHIB A. (eds), Prehistoire de Casablanca. 1 - La Grotte des Rhinoceros (fouilles 1991 et 1996). Villes et Sites Archeologiques du Maroc, Vol. VI, Rabat: 105 - 110." type="book chapter" year="2016">2016b</bibRefCitation>
|
||
) et la distingue de l’actuelle
|
||
<taxonomicName id="94C04D14FFE5FFF4FC7FF9D7FC0F2B66" authorityName="Geoffroy Saint-Hilaire" authorityYear="1834" box="[914,1006,1587,1613]" class="Mammalia" family="Soricidae" genus="Crocidura" kingdom="Animalia" order="Soricomorpha" pageId="19" pageNumber="254" phylum="Chordata" rank="species" species="viaria">
|
||
<emphasis id="61B4EA85FFE5FFF4FC7FF9D7FC452B67" box="[914,932,1587,1613]" italics="true" pageId="19" pageNumber="254">C</emphasis>
|
||
.
|
||
<emphasis id="61B4EA85FFE5FFF4FC5DF9D7FC0F2B66" box="[944,1006,1587,1612]" italics="true" pageId="19" pageNumber="254">viaria</emphasis>
|
||
</taxonomicName>
|
||
(qui présente notamment une échancrure marquée en avant du talon, cf.
|
||
<bibRefCitation id="37514B66FFE5FFF4FB7DF9B7FAD02B46" author="HUTTERER R." box="[1168,1329,1618,1645]" pageId="19" pageNumber="254" pagination="521 - 534" refId="ref19093" refString="HUTTERER R. 1986. - The species of Crocidura (Soricidae) in Morocco. Mammalia 50 (4): 521 - 534. https: // doi. org / 10.1515 / mamm. 1986.50.4.521" type="journal article" year="1986">Hutterer 1986</bibRefCitation>
|
||
). La P4 de Ben Kérat montre un parastyle bien séparé de la cuspide principale, bien qu’il apparaisse moins développé que sur le spécimen
|
||
<typeStatus id="8C7B8835FFE5FFF4FC51F957FC0D2BE7" box="[956,1004,1715,1741]" pageId="19" pageNumber="254">type</typeStatus>
|
||
de
|
||
<taxonomicName id="94C04D14FFE5FFF4FBF4F956FA692BE7" authority="(Hutterer 1991)" baseAuthorityName="Hutterer" baseAuthorityYear="1991" box="[1049,1416,1714,1741]" class="Mammalia" family="Soricidae" genus="Crocidura" kingdom="Animalia" order="Soricomorpha" pageId="19" pageNumber="254" phylum="Chordata" rank="species" species="maghrebiana">
|
||
<emphasis id="61B4EA85FFE5FFF4FBF4F956FBC82BE6" box="[1049,1065,1714,1740]" italics="true" pageId="19" pageNumber="254">C</emphasis>
|
||
.
|
||
<emphasis id="61B4EA85FFE5FFF4FBD6F956FB262BE6" box="[1083,1223,1714,1740]" italics="true" pageId="19" pageNumber="254">maghrebiana</emphasis>
|
||
(
|
||
<bibRefCitation id="37514B66FFE5FFF4FB37F957FA9E2BE6" author="HUTTERER R." box="[1242,1407,1714,1741]" pageId="19" pageNumber="254" pagination="241 - 251" refId="ref19131" refString="HUTTERER R. 1991. - Variation and evolution of the Sicilian shrew: taxonomic conclusions and description of a possibly related species from the Pleistocene of Morocco (Mammalia: Soricidae). Bonner Zoologische Beitrage 42 (3 - 4): 241 - 251." type="journal article" year="1991">Hutterer 1991</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
; le cingulum labial est épais au niveau du parastyle et s’affine vers la partie postérieure de la dent; la partie linguale de la P4 est de forme carrée. Ces caractères se rapprochent davantage de
|
||
<taxonomicName id="94C04D14FFE5FFF4FC21F8D6FB9B2A61" baseAuthorityName="Hutterer" baseAuthorityYear="1991" box="[972,1146,1841,1868]" class="Mammalia" family="Soricidae" genus="Crocidura" kingdom="Animalia" order="Soricomorpha" pageId="19" pageNumber="254" phylum="Chordata" rank="species" species="maghrebiana">
|
||
<emphasis id="61B4EA85FFE5FFF4FC21F8D6FB9B2A61" box="[972,1146,1841,1868]" italics="true" pageId="19" pageNumber="254">C. maghrebiana</emphasis>
|
||
</taxonomicName>
|
||
que de
|
||
<taxonomicName id="94C04D14FFE5FFF4FB31F8D6FADE2A61" authorityName="Geoffroy Saint-Hilaire" authorityYear="1834" box="[1244,1343,1842,1868]" class="Mammalia" family="Soricidae" genus="Crocidura" kingdom="Animalia" order="Soricomorpha" pageId="19" pageNumber="254" phylum="Chordata" rank="species" species="viaria">
|
||
<emphasis id="61B4EA85FFE5FFF4FB31F8D6FB0D2A66" box="[1244,1260,1842,1868]" italics="true" pageId="19" pageNumber="254">C</emphasis>
|
||
.
|
||
<emphasis id="61B4EA85FFE5FFF4FB13F8D6FADE2A61" box="[1278,1343,1842,1867]" italics="true" pageId="19" pageNumber="254">viaria</emphasis>
|
||
</taxonomicName>
|
||
, chez qui le parastyle est plus rapproché du paracône et le cingulum est plus restreint sur toute la longueur (
|
||
<bibRefCitation id="37514B66FFE5FFF4FAE8F895FA462AA1" author="GERAADS D." box="[1285,1447,1905,1931]" pageId="19" pageNumber="254" pagination="281 - 294" refId="ref18131" refString="GERAADS D. 1993. - Middle Pleistocene Crocidura (Mammalia, Insectivora) from Oulad Hamida I, Morocco, and their phylogenetic relationships. Proceedings of the Koninklijke Nederlandse Akademie van Wetenschappen 96 (3): 281 - 294." type="journal article" year="1993">Geraads 1993</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="37514B66FFE5FFF4FCC0F875FC942A81" author="GERAADS D." box="[813,885,1937,1963]" pageId="19" pageNumber="254" pagination="105 - 110" refId="ref18435" refString="GERAADS D. 2016 b. - La faune de vertebres du Pleistocene moyen de la Grotte des Rhinoceros, Casablanca, Maroc: 5 - Insectivores, in RAYNAL J. P. & MOHIB A. (eds), Prehistoire de Casablanca. 1 - La Grotte des Rhinoceros (fouilles 1991 et 1996). Villes et Sites Archeologiques du Maroc, Vol. VI, Rabat: 105 - 110." type="book chapter" year="2016">2016b</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="37514B66FFE5FFF4FC65F875FBD22A81" author="VERGILINO M." box="[904,1075,1937,1963]" pageId="19" pageNumber="254" refId="ref21319" refString="VERGILINO M. 1999. - Les peuplements de Soricidae (Lypotyphla, Mammalia) du Maghreb depuis le Miocene. Memoire de DEA, Universite de Montpellier, France, 33 p." type="book" year="1999">Vergilino 1999</bibRefCitation>
|
||
). Malheureusement l’absence de U
|
||
<quantity id="94389B72FFE5FFF4FCABF855FC902AE1" box="[838,881,1969,1995]" metricMagnitude="-2" metricUnit="m" metricValue="7.62" pageId="19" pageNumber="254" unit="in" value="3.0">
|
||
3
|
||
<emphasis id="61B4EA85FFE5FFF4FCB7F856FC902AE1" box="[858,881,1970,1995]" italics="true" pageId="19" pageNumber="254">in</emphasis>
|
||
</quantity>
|
||
situ empêche l’observation de la position relative de cette dernière par rapport au parastyle de la P4 en vue labiale. La M1 de Ben Kérat est relativement large avec un hypocône très développé, ce qui là encore rapproche davantage ce spécimen de
|
||
<taxonomicName id="94C04D14FFE3FFF2FE49FAF0FDB12807" box="[420,592,1299,1326]" class="Mammalia" family="Soricidae" genus="Crocidura" kingdom="Animalia" order="Soricomorpha" pageId="21" pageNumber="256" phylum="Chordata" rank="species" species="maghrebiana">
|
||
<emphasis id="61B4EA85FFE3FFF2FE49FAF0FDB12807" box="[420,592,1299,1326]" italics="true" pageId="21" pageNumber="256">C. maghrebiana</emphasis>
|
||
</taxonomicName>
|
||
que de
|
||
<taxonomicName id="94C04D14FFE3FFF2FD4BFAF0FCE62807" authorityName="Geoffroy Saint-Hilaire" authorityYear="1834" box="[678,775,1300,1326]" class="Mammalia" family="Soricidae" genus="Crocidura" kingdom="Animalia" order="Soricomorpha" pageId="21" pageNumber="256" phylum="Chordata" rank="species" species="viaria">
|
||
<emphasis id="61B4EA85FFE3FFF2FD4BFAF0FD592804" box="[678,696,1300,1326]" italics="true" pageId="21" pageNumber="256">C</emphasis>
|
||
.
|
||
<emphasis id="61B4EA85FFE3FFF2FD2AFAF0FCE62807" box="[711,775,1300,1325]" italics="true" pageId="21" pageNumber="256">viaria</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<bibRefCitation id="37514B66FFE3FFF2FF63FAD7FEC62867" author="GERAADS D." box="[142,295,1331,1357]" pageId="21" pageNumber="256" pagination="281 - 294" refId="ref18131" refString="GERAADS D. 1993. - Middle Pleistocene Crocidura (Mammalia, Insectivora) from Oulad Hamida I, Morocco, and their phylogenetic relationships. Proceedings of the Koninklijke Nederlandse Akademie van Wetenschappen 96 (3): 281 - 294." type="journal article" year="1993">Geraads 1993</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="37514B66FFE3FFF2FED8FAD7FE9C2867" author="GERAADS D." box="[309,381,1331,1357]" pageId="21" pageNumber="256" pagination="105 - 110" refId="ref18435" refString="GERAADS D. 2016 b. - La faune de vertebres du Pleistocene moyen de la Grotte des Rhinoceros, Casablanca, Maroc: 5 - Insectivores, in RAYNAL J. P. & MOHIB A. (eds), Prehistoire de Casablanca. 1 - La Grotte des Rhinoceros (fouilles 1991 et 1996). Villes et Sites Archeologiques du Maroc, Vol. VI, Rabat: 105 - 110." type="book chapter" year="2016">2016b</bibRefCitation>
|
||
). Ainsi, bien que le matériel de Ben Kérat soit assez limité et fragmenté, et qu’il paraisse un peu plus gracile que celui de la Grotte des Rhinocéros (
|
||
<collectingCountry id="2BD77607FFE3FFF2FD46FA97FD1828A7" box="[683,761,1395,1421]" name="Morocco" pageId="21" pageNumber="256">Maroc</collectingCountry>
|
||
), une majorité de critères va en faveur d’une attribution de ce matériel à
|
||
<taxonomicName id="94C04D14FFE3FFF2FF18FA57FE7E28E7" box="[245,415,1459,1485]" class="Mammalia" family="Soricidae" genus="Crocidura" kingdom="Animalia" order="Soricomorpha" pageId="21" pageNumber="256" phylum="Chordata" rank="species" species="maghrebiana">
|
||
<emphasis id="61B4EA85FFE3FFF2FF18FA57FE7E28E7" box="[245,415,1459,1485]" italics="true" pageId="21" pageNumber="256">C. maghrebiana</emphasis>
|
||
</taxonomicName>
|
||
. À noter qu’à Jebel Ressas 4, une M3 de crocidure a été attribué à
|
||
<taxonomicName id="94C04D14FFE3FFF2FE03FA37FDAE28C7" authorityName="Geoffroy Saint-Hilaire" authorityYear="1834" box="[494,591,1491,1517]" class="Mammalia" family="Soricidae" genus="Crocidura" kingdom="Animalia" order="Soricomorpha" pageId="21" pageNumber="256" phylum="Chordata" rank="species" species="viaria">
|
||
<emphasis id="61B4EA85FFE3FFF2FE03FA37FDE128C7" box="[494,512,1491,1517]" italics="true" pageId="21" pageNumber="256">C</emphasis>
|
||
.
|
||
<emphasis id="61B4EA85FFE3FFF2FDE2FA30FDAE28C7" box="[527,591,1492,1517]" italics="true" pageId="21" pageNumber="256">viaria</emphasis>
|
||
</taxonomicName>
|
||
uniquement sur la base de sa grande taille (
|
||
<bibRefCitation id="37514B66FFE3FFF2FE42FA17FD502B27" author="MEIN P. & PICKFORD M." box="[431,689,1523,1549]" pageId="21" pageNumber="256" pagination="247 - 253" refId="ref19610" refString="MEIN P. & PICKFORD M. 1992. - Gisements karstiques pleistocenes au Djebel Ressas, Tunisie. Comptes Rendus de l'Academie des Sciences de Paris, Serie II 315: 247 - 253. https: // gallica. bnf. fr / ark: / 12148 / bpt 6 k 6304735 s / f 261. item" type="journal article" year="1992">Mein & Pickford 1992</bibRefCitation>
|
||
; Mein, notes personnelles non publiées), mais sur ce seul critère elle pourrait tout aussi bien appartenir à
|
||
<taxonomicName id="94C04D14FFE3FFF2FDA0F9D7FD1B2B66" box="[589,762,1586,1613]" class="Mammalia" family="Soricidae" genus="Crocidura" kingdom="Animalia" order="Soricomorpha" pageId="21" pageNumber="256" phylum="Chordata" rank="species" species="maghrebiana">
|
||
<emphasis id="61B4EA85FFE3FFF2FDA0F9D7FD1B2B66" box="[589,762,1586,1613]" italics="true" pageId="21" pageNumber="256">C. maghrebiana</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |