treatments-xml/data/CC/79/87/CC7987FCFF8E5C4C52C732D9FC4B6769.xml

<document ID-DOI="10.11646/phytotaxa.583.3.7" ID-ISSN="1179-3163" ID-Zenodo-Dep="7621526" checkinTime="1675871161665" checkinUser="plazi" docAuthor="Sunojkumar, P." docDate="2023" docId="CC7987FCFF8E5C4C52C732D9FC4B6769" docLanguage="en" docName="phytotaxa.583.3.7.pdf" docOrigin="Phytotaxa 583 (3)" docSource="http://dx.doi.org/10.11646/phytotaxa.583.3.7" docStyle="DocumentStyle:96748F8F1B6C902996E134952A3A36B9.5:Phytotaxa.2014-.journal_article" docStyleId="96748F8F1B6C902996E134952A3A36B9" docStyleName="Phytotaxa.2014-.journal_article" docStyleVersion="5" docTitle="Hedyotis recurvata Jiji, Jeomol, Manudev &amp; Sunojk. A. A 2023, sp. nov." docType="treatment" docVersion="3" lastPageNumber="289" masterDocId="3040FF84FF8C5C48524F322DFFFD664D" masterDocTitle="Hedyotis recurvata (Rubiaceae): a new species from southern Western Ghats of India" masterLastPageNumber="291" masterPageNumber="285" pageNumber="287" updateTime="1676043973764" updateUser="valdenar" zenodo-license-figures="UNSPECIFIED" zenodo-license-treatments="UNSPECIFIED">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>Hedyotis recurvata (Rubiaceae): a new species from southern Western Ghats of India</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Sunojkumar, P.</mods:namePart>
<mods:nameIdentifier type="ORCID">0000-0003-0453-7684</mods:nameIdentifier>
<mods:affiliation>Department of Botany, University of Calicut, Calicut University PO, Malappuram District, Kerala- 673 635, India &amp; drsunoj @ gmail. com; https: // orcid. org / 0000 - 0003 - 0453 - 7684</mods:affiliation>
<mods:nameIdentifier type="email">drsunoj@gmail.com</mods:nameIdentifier>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>Phytotaxa</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2023</mods:date>
<mods:detail type="pubDate">
<mods:number>2023-02-07</mods:number>
</mods:detail>
<mods:detail type="volume">
<mods:number>583</mods:number>
</mods:detail>
<mods:detail type="issue">
<mods:number>3</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>285</mods:start>
<mods:end>291</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:location>
<mods:url>http://dx.doi.org/10.11646/phytotaxa.583.3.7</mods:url>
</mods:location>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">10.11646/phytotaxa.583.3.7</mods:identifier>
<mods:identifier type="ISSN">1179-3163</mods:identifier>
<mods:identifier type="Zenodo-Dep">7621526</mods:identifier>
</mods:mods>
<treatment LSID="urn:lsid:plazi:treatment:CC7987FCFF8E5C4C52C732D9FC4B6769" httpUri="http://treatment.plazi.org/id/CC7987FCFF8E5C4C52C732D9FC4B6769" lastPageId="4" lastPageNumber="289" pageId="2" pageNumber="287">
<subSubSection box="[136,976,244,271]" pageId="2" pageNumber="287" type="nomenclature">
<paragraph blockId="2.[136,976,244,271]" box="[136,976,244,271]" pageId="2" pageNumber="287">
<heading box="[136,976,244,271]" fontSize="11" level="2" pageId="2" pageNumber="287" reason="3">
<taxonomicName authority="Jiji, Jeomol, Manudev &amp; Sunojk." authorityName="Jiji, Jeomol, Manudev &amp; Sunojk. A. A" authorityYear="2023" box="[136,733,244,271]" class="Magnoliopsida" family="Rubiaceae" genus="Hedyotis" kingdom="Plantae" order="Gentianales" pageId="2" pageNumber="287" phylum="Tracheophyta" rank="species" species="recurvata" status="sp. nov.">
<emphasis bold="true" box="[136,354,244,271]" italics="true" pageId="2" pageNumber="287">Hedyotis recurvata</emphasis>
Jiji, Jeomol, Manudev &amp; Sunojk.
</taxonomicName>
<emphasis box="[740,825,245,270]" italics="true" pageId="2" pageNumber="287">
<taxonomicNameLabel box="[740,825,245,270]" pageId="2" pageNumber="287" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(
<figureCitation box="[839,916,244,270]" captionStart="FIGURE 1" captionStartId="1.[136,229,1910,1932]" captionTargetBox="[305,1283,544,1883]" captionTargetId="figure-136@1.[305,1283,544,1883]" captionTargetPageId="1" captionText="FIGURE 1. Hedyotis recurvata Jiji, Jeomol, Manudev &amp; Sunojk. A. A flowering twig; B &amp; C. Inflorescence; D. &amp; E. Stipule; F. Bract; G. Bracteole; H. Flower bud; I. Flower; J. Calyx; K. Calyx with gynoecium; L. Corolla split open showing stamens; M. Capsules; N. Seeds. (All from P. Jiji, K.K. Jeomol &amp; K.M. Manudev 153887)." figureDoi="http://doi.org/10.5281/zenodo.7621530" httpUri="https://zenodo.org/record/7621530/files/figure.png" pageId="2" pageNumber="287">Figs. 1</figureCitation>
&amp; 
<figureCitation box="[952,967,244,270]" captionStart="FIGURE 2" captionStartId="2.[136,229,1926,1948]" captionTargetBox="[292,1288,559,1884]" captionTargetId="figure-145@2.[280,1308,547,1901]" captionTargetPageId="2" captionText="FIGURE 2. Hedyotis recurvata Jiji, Jeomol, Manudev &amp; Sunojk. A. A flowering twig; B &amp; C. Stipule; D. Inflorescence; E. Flower; F. Corolla split open showing stamens; G. Calyx; H. Calyx with gynoecium; I. Capsules. (All from P. Jiji, K.K. Jeomol &amp; K.M. Manudev 153887)." figureDoi="http://doi.org/10.5281/zenodo.7621536" httpUri="https://zenodo.org/record/7621536/files/figure.png" pageId="2" pageNumber="287">2</figureCitation>
)
</heading>
</paragraph>
</subSubSection>
<subSubSection pageId="2" pageNumber="287" type="description">
<paragraph blockId="2.[136,1452,316,523]" pageId="2" pageNumber="287">
<taxonomicName authorityName="Jiji, Jeomol, Manudev &amp; Sunojk. A. A" authorityYear="2023" box="[136,355,316,342]" class="Magnoliopsida" family="Rubiaceae" genus="Hedyotis" kingdom="Plantae" order="Gentianales" pageId="2" pageNumber="287" phylum="Tracheophyta" rank="species" species="recurvata">
<emphasis box="[136,355,316,342]" italics="true" pageId="2" pageNumber="287">Hedyotis recurvata</emphasis>
</taxonomicName>
is similar to 
<taxonomicName authorityName="Hooker" authorityYear="1880" box="[519,667,316,342]" class="Magnoliopsida" family="Rubiaceae" genus="Hedyotis" kingdom="Plantae" order="Gentianales" pageId="2" pageNumber="287" phylum="Tracheophyta" rank="species" species="beddomei">
<emphasis box="[519,667,316,342]" italics="true" pageId="2" pageNumber="287">H. beddomei</emphasis>
</taxonomicName>
but can be distinguished by its stem trichotomously branched (
<emphasis box="[1421,1444,317,342]" italics="true" pageId="2" pageNumber="287">vs</emphasis>
. dichotomously branched), hairy to glabrescent (
<emphasis box="[686,709,353,378]" italics="true" pageId="2" pageNumber="287">vs</emphasis>
. glabrous); stipules recurved (
<emphasis box="[1062,1085,353,378]" italics="true" pageId="2" pageNumber="287">vs</emphasis>
. adpressed), pectinate with 2–4 distant subulate segments (
<emphasis box="[444,467,389,414]" italics="true" pageId="2" pageNumber="287">vs</emphasis>
. triangular), fully tomentose (
<emphasis box="[811,834,389,414]" italics="true" pageId="2" pageNumber="287">vs</emphasis>
. pubescent); petioles pubescent (
<emphasis box="[1214,1237,389,414]" italics="true" pageId="2" pageNumber="287">vs</emphasis>
. glabrous); lamina glabrous above and pubescent on nerves below (
<emphasis box="[670,693,425,450]" italics="true" pageId="2" pageNumber="287">vs</emphasis>
. both sides glabrous), margins ciliate (
<emphasis box="[1120,1151,425,450]" italics="true" pageId="2" pageNumber="287">vs.</emphasis>
not ciliate); calyx glabrous with ciliate margins (
<emphasis box="[385,408,461,486]" italics="true" pageId="2" pageNumber="287">vs</emphasis>
. fully pubescent); corolla glabrous outside (
<emphasis box="[926,949,461,486]" italics="true" pageId="2" pageNumber="287">vs</emphasis>
. pubescent outside) and capsules glabrous, ellipsoid (
<emphasis box="[249,272,497,522]" italics="true" pageId="2" pageNumber="287">vs</emphasis>
. hispid, oblanceolate).
</paragraph>
</subSubSection>
<caption ID-DOI="http://doi.org/10.5281/zenodo.7621536" ID-Zenodo-Dep="7621536" httpUri="https://zenodo.org/record/7621536/files/figure.png" pageId="2" pageNumber="287" startId="2.[136,229,1926,1948]" targetBox="[292,1288,559,1884]" targetPageId="2">
<paragraph blockId="2.[136,1451,1926,2020]" pageId="2" pageNumber="287">
<emphasis bold="true" box="[136,254,1926,1948]" pageId="2" pageNumber="287">FIGURE 2.</emphasis>
<taxonomicName authority="Jiji, Jeomol, Manudev &amp; Sunojk. A. A" authorityName="Jiji, Jeomol, Manudev &amp; Sunojk. A. A" authorityYear="2023" box="[261,825,1926,1948]" class="Magnoliopsida" family="Rubiaceae" genus="Hedyotis" kingdom="Plantae" order="Gentianales" pageId="2" pageNumber="287" phylum="Tracheophyta" rank="species" species="recurvata">
<emphasis box="[261,445,1926,1948]" italics="true" pageId="2" pageNumber="287">Hedyotis recurvata</emphasis>
Jiji, Jeomol, Manudev &amp; Sunojk. 
<emphasis bold="true" box="[780,797,1926,1947]" pageId="2" pageNumber="287">A</emphasis>
. A
</taxonomicName>
flowering twig; 
<emphasis bold="true" box="[988,1004,1926,1947]" pageId="2" pageNumber="287">B</emphasis>
&amp; 
<emphasis bold="true" box="[1036,1053,1926,1948]" pageId="2" pageNumber="287">C</emphasis>
. Stipule; 
<emphasis bold="true" box="[1148,1165,1926,1947]" pageId="2" pageNumber="287">D</emphasis>
. Inflorescence; 
<emphasis bold="true" box="[1320,1336,1926,1947]" pageId="2" pageNumber="287">E</emphasis>
. Flower; 
<emphasis bold="true" box="[1431,1446,1926,1947]" pageId="2" pageNumber="287">F</emphasis>
. Corolla split open showing stamens; 
<emphasis bold="true" box="[496,515,1962,1984]" pageId="2" pageNumber="287">G</emphasis>
. Calyx; 
<emphasis bold="true" box="[598,617,1962,1983]" pageId="2" pageNumber="287">H</emphasis>
. Calyx with gynoecium; 
<emphasis bold="true" box="[863,872,1962,1983]" pageId="2" pageNumber="287">I</emphasis>
. Capsules. (All from 
<emphasis box="[1084,1099,1963,1984]" italics="true" pageId="2" pageNumber="287">P</emphasis>
. 
<emphasis box="[1111,1451,1962,1984]" italics="true" pageId="2" pageNumber="287">Jiji, K.K. Jeomol &amp; K.M. Manudev</emphasis>
153887).
</paragraph>
</caption>
<caption ID-DOI="http://doi.org/10.5281/zenodo.7621542" ID-Zenodo-Dep="7621542" httpUri="https://zenodo.org/record/7621542/files/figure.png" pageId="3" pageNumber="288" startId="3.[136,229,1123,1145]" targetBox="[151,1436,190,1099]" targetPageId="3">
<paragraph blockId="3.[136,1451,1123,1181]" pageId="3" pageNumber="288">
<emphasis bold="true" box="[136,256,1123,1145]" pageId="3" pageNumber="288">FIGURE 3.</emphasis>
Map showing the distribution of 
<taxonomicName authority="Jiji, Jeomol, Manudev &amp; Sunojk." authorityName="Jiji, Jeomol, Manudev &amp; Sunojk. A. A" authorityYear="2023" box="[595,1120,1123,1145]" class="Magnoliopsida" family="Rubiaceae" genus="Hedyotis" kingdom="Plantae" order="Gentianales" pageId="3" pageNumber="288" phylum="Tracheophyta" rank="species" species="recurvata">
<emphasis box="[595,781,1123,1145]" italics="true" pageId="3" pageNumber="288">Hedyotis recurvata</emphasis>
Jiji, Jeomol, Manudev &amp; Sunojk.
</taxonomicName>
[drawn using the software QGIS 3.24.1-Tisler (
<bibRefCitation author="QGIS DEVELOPMENT TEAM" box="[272,589,1159,1181]" pageId="3" pageNumber="288" refId="ref3603" refString="QGIS DEVELOPMENT TEAM (2022) QGIS Geographic Information System. QGIS 3.24.1 - Tisler. Open Source Geospatial Foundation Project. [http: // qgis. osgeo. org]" type="url" year="2022">QGIS Development Team, 2022</bibRefCitation>
)].
</paragraph>
</caption>
<subSubSection pageId="3" pageNumber="288" type="materials_examined">
<paragraph blockId="3.[136,1452,1227,2082]" pageId="3" pageNumber="288">
<emphasis bold="true" box="[189,248,1228,1254]" pageId="3" pageNumber="288">Type</emphasis>
:— 
<materialsCitation collectingDate="2021-11-02" collectionCode="K" collectorName="P. Jiji &amp; Jeomol &amp; M. Manudev" country="India" elevation="2197" latitude="11.292084" location="Cauliflower Shola" longLatPrecision="1" longitude="76.57953" municipality="Avalanche" pageId="3" pageNumber="288" specimenCount="1" stateProvince="Tamil Nadu" typeStatus="holotype">
<collectingCountry box="[282,367,1228,1254]" name="India" pageId="3" pageNumber="288">INDIA</collectingCountry>
. 
<collectingRegion box="[375,509,1227,1253]" country="India" name="Tamil Nadu" pageId="3" pageNumber="288">Tamil Nadu</collectingRegion>
, 
<collectingMunicipality box="[517,638,1227,1253]" pageId="3" pageNumber="288">Avalanche</collectingMunicipality>
, on the way to 
<location LSID="urn:lsid:plazi:treatment:CC7987FCFF8E5C4C52C732D9FC4B6769:410F6031FF8F5C4B516F36E6FC1362A8" box="[800,1006,1227,1253]" country="India" latitude="11.292084" longLatPrecision="1" longitude="76.57953" municipality="Avalanche" name="Cauliflower Shola" pageId="3" pageNumber="288" stateProvince="Tamil Nadu">Cauliflower Shola</location>
, ± 
<quantity box="[1030,1116,1228,1254]" metricMagnitude="3" metricUnit="m" metricValue="2.197" pageId="3" pageNumber="288" unit="m" value="2197.0">
<elevation box="[1030,1116,1228,1254]" metricMagnitude="3" metricUnit="m" metricValue="2.197" pageId="3" pageNumber="288" unit="m" value="2197.0">2197 m</elevation>
</quantity>
, 
<geoCoordinate box="[1127,1284,1227,1254]" degrees="11" direction="north" minutes="17" orientation="latitude" pageId="3" pageNumber="288" precision="1" seconds="31.5" value="11.292084">11°17’31.5”N</geoCoordinate>
<geoCoordinate box="[1289,1447,1227,1253]" degrees="76" direction="east" minutes="34" orientation="longitude" pageId="3" pageNumber="288" precision="1" seconds="46.3" value="76.57953">76°34’46.3”E</geoCoordinate>
, 
<date box="[136,355,1263,1290]" pageId="3" pageNumber="288" value="2021-11-02">
<collectingDate box="[136,355,1263,1290]" pageId="3" pageNumber="288" value="2021-11-02">02 November 2021</collectingDate>
</date>
, 
<collectorName box="[366,435,1264,1290]" pageId="3" pageNumber="288">
<emphasis box="[366,383,1264,1289]" italics="true" pageId="3" pageNumber="288">P</emphasis>
. 
<emphasis box="[397,435,1264,1289]" italics="true" pageId="3" pageNumber="288">Jiji</emphasis>
</collectorName>
, 
<collectionCode box="[447,469,1264,1289]" country="United Kingdom" lsid="urn:lsid:biocol.org:col:15867" name="Royal Botanic Gardens" pageId="3" pageNumber="288" type="Herbarium">K</collectionCode>
. 
<collectionCode box="[473,495,1264,1289]" country="United Kingdom" lsid="urn:lsid:biocol.org:col:15867" name="Royal Botanic Gardens" pageId="3" pageNumber="288" type="Herbarium">K</collectionCode>
. 
<collectorName box="[505,586,1263,1289]" pageId="3" pageNumber="288">Jeomol</collectorName>
&amp; 
<collectionCode box="[622,644,1264,1289]" country="United Kingdom" lsid="urn:lsid:biocol.org:col:15867" name="Royal Botanic Gardens" pageId="3" pageNumber="288" type="Herbarium">K</collectionCode>
. 
<collectorName box="[647,789,1263,1289]" pageId="3" pageNumber="288">M. Manudev</collectorName>
153887 (
<typeStatus box="[895,995,1263,1289]" pageId="3" pageNumber="288">holotype</typeStatus>
CALI!; isotype MH!, DEV!)
</materialsCitation>
.
</paragraph>
<paragraph blockId="3.[136,1452,1227,2082]" pageId="3" pageNumber="288">
Perennial, shrubs, up to 
<quantity box="[472,544,1299,1326]" metricMagnitude="-1" metricUnit="m" metricValue="6.0" pageId="3" pageNumber="288" unit="cm" value="60.0">60 cm</quantity>
tall. 
<emphasis box="[607,672,1300,1326]" italics="true" pageId="3" pageNumber="288">Stems</emphasis>
erect, trichotomously branched, woody at base; 
<emphasis box="[1237,1355,1299,1325]" italics="true" pageId="3" pageNumber="288">internodes</emphasis>
<quantity box="[1365,1451,1300,1326]" metricMagnitude="-2" metricUnit="m" metricValue="3.0" metricValueMax="4.0" metricValueMin="2.0" pageId="3" pageNumber="288" unit="cm" value="3.0" valueMax="4.0" valueMin="2.0">2–4 cm</quantity>
long, 4-angular, winged, hairy to glabrescent, whitish when dry. 
<emphasis box="[856,945,1335,1361]" italics="true" pageId="3" pageNumber="288">Stipules</emphasis>
recurved, pectinate with 2–4 distant subulate segments, 4–10 × 
<quantity box="[343,480,1371,1398]" metricMagnitude="-3" metricUnit="m" metricValue="2.5" metricValueMax="3.5" metricValueMin="1.5" pageId="3" pageNumber="288" unit="mm" value="2.5" valueMax="3.5" valueMin="1.5">1.5–3.5 mm</quantity>
, tomentose, yellowish green with segments black at apex. 
<emphasis box="[1143,1221,1372,1397]" italics="true" pageId="3" pageNumber="288">Leaves</emphasis>
compactly arranged to the apex, opposite; 
<emphasis box="[385,472,1407,1433]" italics="true" pageId="3" pageNumber="288">petioles</emphasis>
<quantity box="[480,574,1408,1434]" metricMagnitude="-3" metricUnit="m" metricValue="2.5" metricValueMax="4.0" metricValueMin="1.0" pageId="3" pageNumber="288" unit="mm" value="2.5" valueMax="4.0" valueMin="1.0">1–4 mm</quantity>
long, canaliculate, pubescent, green; 
<emphasis box="[998,1076,1407,1433]" italics="true" pageId="3" pageNumber="288">lamina</emphasis>
elliptic-ovate, 2.5–5.5 × 
<quantity box="[1360,1447,1408,1434]" metricMagnitude="-2" metricUnit="m" metricValue="1.5" metricValueMax="2.0" metricValueMin="1.0" pageId="3" pageNumber="288" unit="cm" value="1.5" valueMax="2.0" valueMin="1.0">1–2 cm</quantity>
, base cuneate or round, margins entire, ciliate, apex acuminate, glabrous above, pubescent on nerves below, green with black margins, nerves 
<quantity box="[445,516,1479,1505]" metricMagnitude="-1" metricUnit="m" metricValue="1.143" metricValueMax="1.524" metricValueMin="0.762" pageId="3" pageNumber="288" unit="in" value="4.5" valueMax="6.0" valueMin="3.0">3–6 in</quantity>
pairs, prominent below, convergent. 
<emphasis box="[933,1008,1480,1506]" italics="true" pageId="3" pageNumber="288">Cymes</emphasis>
umbellate, terminal and axillary, 5–30- flowered; 
<emphasis box="[254,366,1515,1541]" italics="true" pageId="3" pageNumber="288">peduncles</emphasis>
4-angular, 
<quantity box="[501,587,1516,1542]" metricMagnitude="-2" metricUnit="m" metricValue="2.0" metricValueMax="3.0" metricValueMin="1.0" pageId="3" pageNumber="288" unit="cm" value="2.0" valueMax="3.0" valueMin="1.0">1–3 cm</quantity>
long, pubescent, green; 
<emphasis box="[876,946,1515,1541]" italics="true" pageId="3" pageNumber="288">bracts</emphasis>
leafy, elliptic-ovate, 5–10 × 
<quantity box="[1288,1386,1515,1542]" metricMagnitude="-3" metricUnit="m" metricValue="4.5" metricValueMax="6.0" metricValueMin="3.0" pageId="3" pageNumber="288" unit="mm" value="4.5" valueMax="6.0" valueMin="3.0">3–6 mm</quantity>
, base cuneate, margins entire, apex acuminate, nerves 
<quantity box="[689,762,1551,1578]" metricMagnitude="-2" metricUnit="m" metricValue="8.89" metricValueMax="10.16" metricValueMin="7.62" pageId="3" pageNumber="288" unit="in" value="3.5" valueMax="4.0" valueMin="3.0">3–4 in</quantity>
pairs, hairy as lamina, green with margins black; 
<emphasis box="[1335,1452,1551,1577]" italics="true" pageId="3" pageNumber="288">bracteoles</emphasis>
ovate to lanceolate, 4–5 × 0.5–1.0 mm, ciliate, green. 
<emphasis box="[737,829,1587,1613]" italics="true" pageId="3" pageNumber="288">Flowers</emphasis>
compactly arranged, 
<quantity box="[1071,1164,1588,1614]" metricMagnitude="-3" metricUnit="m" metricValue="8.5" metricValueMax="9.0" metricValueMin="8.0" pageId="3" pageNumber="288" unit="mm" value="8.5" valueMax="9.0" valueMin="8.0">8–9 mm</quantity>
long; 
<emphasis box="[1235,1327,1587,1613]" italics="true" pageId="3" pageNumber="288">pedicels</emphasis>
terete, 
<quantity metricMagnitude="-3" metricUnit="m" metricValue="1.5" metricValueMax="2.0" metricValueMin="1.0" pageId="3" pageNumber="288" unit="mm" value="1.5" valueMax="2.0" valueMin="1.0">1–2 mm</quantity>
, glabrous, pale green. 
<taxonomicName box="[441,506,1623,1649]" class="Demospongiae" family="Phloeodictyidae" genus="Calyx" kingdom="Animalia" order="Haplosclerida" pageId="3" pageNumber="288" phylum="Porifera" rank="genus">
<emphasis box="[441,506,1623,1649]" italics="true" pageId="3" pageNumber="288">Calyx</emphasis>
</taxonomicName>
4-lobed, infundibular, 
<quantity box="[770,867,1624,1650]" metricMagnitude="-3" metricUnit="m" metricValue="3.5" metricValueMax="4.0" metricValueMin="3.0" pageId="3" pageNumber="288" unit="mm" value="3.5" valueMax="4.0" valueMin="3.0">3–4 mm</quantity>
, purple with lobes green; 
<emphasis box="[1166,1225,1623,1649]" italics="true" pageId="3" pageNumber="288">lobes</emphasis>
triangular, 
<quantity box="[1357,1452,1624,1650]" metricMagnitude="-3" metricUnit="m" metricValue="2.5" metricValueMax="3.0" metricValueMin="2.0" pageId="3" pageNumber="288" unit="mm" value="2.5" valueMax="3.0" valueMin="2.0">2–3 mm</quantity>
long, valvate, apex acute, margins ciliate. 
<emphasis box="[602,688,1659,1685]" italics="true" pageId="3" pageNumber="288">Corolla</emphasis>
4-lobed, rotate; 
<emphasis box="[871,930,1659,1685]" italics="true" pageId="3" pageNumber="288">tubes</emphasis>
<quantity box="[936,1028,1659,1686]" metricMagnitude="-3" metricUnit="m" metricValue="7.0" metricValueMax="8.0" metricValueMin="6.0" pageId="3" pageNumber="288" unit="mm" value="7.0" valueMax="8.0" valueMin="6.0">6–8 mm</quantity>
long, glabrous outside, woolly inside, denser at the throat; lobes ovate with beaked apex, 2.0–2.5, recurved, margins ciliate, glabrous towards apex, purple. 
<emphasis box="[136,229,1732,1758]" italics="true" pageId="3" pageNumber="288">Stamens</emphasis>
4, epipetalous, exserted; 
<emphasis box="[508,611,1731,1757]" italics="true" pageId="3" pageNumber="288">filaments</emphasis>
terete, 
<quantity box="[691,783,1731,1758]" metricMagnitude="-3" metricUnit="m" metricValue="4.5" metricValueMax="5.0" metricValueMin="4.0" pageId="3" pageNumber="288" unit="mm" value="4.5" valueMax="5.0" valueMin="4.0">4–5 mm</quantity>
long, glabrous, white to pale blue; 
<emphasis box="[1169,1253,1731,1757]" italics="true" pageId="3" pageNumber="288">anthers</emphasis>
ellipsoid, 1.0–1.5 × 
<quantity box="[159,297,1767,1794]" metricMagnitude="-4" metricUnit="m" metricValue="4.0" metricValueMax="5.0" metricValueMin="3.0" pageId="3" pageNumber="288" unit="mm" value="0.4" valueMax="0.5" valueMin="0.3">0.3–0.5 mm</quantity>
, dorsifixed, dark purple. 
<emphasis box="[580,650,1768,1794]" italics="true" pageId="3" pageNumber="288">Ovary</emphasis>
2.0–2.5 × 1.5–2.0 mm, glabrous, pale purple or green; 
<emphasis box="[1274,1325,1767,1793]" italics="true" pageId="3" pageNumber="288">style</emphasis>
terete, 
<quantity metricMagnitude="-3" metricUnit="m" metricValue="4.5" metricValueMax="5.0" metricValueMin="4.0" pageId="3" pageNumber="288" unit="mm" value="4.5" valueMax="5.0" valueMin="4.0">4–5 mm</quantity>
long, glabrous; 
<emphasis box="[359,434,1804,1829]" italics="true" pageId="3" pageNumber="288">stigma</emphasis>
bilobed; 
<emphasis box="[537,596,1803,1829]" italics="true" pageId="3" pageNumber="288">lobes</emphasis>
1.5–2.0 mm long, pappillose, pink. 
<emphasis box="[995,1098,1803,1829]" italics="true" pageId="3" pageNumber="288">Capsules</emphasis>
ovoid to ellipsoid, 2.0– 
<quantity box="[1361,1447,1803,1830]" metricMagnitude="-3" metricUnit="m" metricValue="2.5" pageId="3" pageNumber="288" unit="mm" value="2.5">2.5 mm</quantity>
, glabrous. 
<emphasis box="[246,310,1839,1865]" italics="true" pageId="3" pageNumber="288">Seeds</emphasis>
narrowly winged with a raised hilar ridge 0.5–1.0× 0.4– 
<quantity box="[950,1038,1839,1866]" metricMagnitude="-4" metricUnit="m" metricValue="6.0" pageId="3" pageNumber="288" unit="mm" value="0.6">0.6 mm</quantity>
, glabrous, black.
</paragraph>
<paragraph blockId="3.[136,1452,1227,2082]" box="[189,738,1875,1902]" pageId="3" pageNumber="288">
<emphasis bold="true" box="[189,440,1875,1901]" pageId="3" pageNumber="288">Flowering &amp; fruiting</emphasis>
:—September to February.
</paragraph>
</subSubSection>
<subSubSection pageId="3" pageNumber="288" type="description">
<paragraph blockId="3.[136,1452,1227,2082]" pageId="3" pageNumber="288">
<emphasis bold="true" box="[189,281,1911,1937]" pageId="3" pageNumber="288">Habitat</emphasis>
:—Occurring at an elevations of ± 
<quantity box="[666,752,1912,1938]" metricMagnitude="3" metricUnit="m" metricValue="2.197" pageId="3" pageNumber="288" unit="m" value="2197.0">2197 m</quantity>
above sea level, growing among grasses along the margins of evergreen shola forests.
</paragraph>
</subSubSection>
<subSubSection pageId="3" pageNumber="288" type="distribution">
<paragraph blockId="3.[136,1452,1227,2082]" pageId="3" pageNumber="288">
<emphasis bold="true" box="[189,335,1983,2009]" pageId="3" pageNumber="288">Distribution</emphasis>
:— 
<collectingCountry box="[369,453,1984,2010]" name="India" pageId="3" pageNumber="288">INDIA</collectingCountry>
, hitherto known only from the 
<typeStatus box="[815,863,1984,2010]" pageId="3" pageNumber="288">type</typeStatus>
locality, Avalanche in Nilgiris district, 
<collectingRegion box="[1316,1452,1983,2009]" country="India" name="Tamil Nadu" pageId="3" pageNumber="288">Tamil Nadu</collectingRegion>
(
<figureCitation box="[144,213,2019,2046]" captionStart="FIGURE 3" captionStartId="3.[136,229,1123,1145]" captionTargetBox="[151,1436,190,1099]" captionTargetId="figure-19@3.[151,1436,190,1099]" captionTargetPageId="3" captionText="FIGURE 3. Map showing the distribution of Hedyotis recurvata Jiji, Jeomol, Manudev &amp; Sunojk. [drawn using the software QGIS 3.24.1-Tisler (QGIS Development Team, 2022)]." figureDoi="http://doi.org/10.5281/zenodo.7621542" httpUri="https://zenodo.org/record/7621542/files/figure.png" pageId="3" pageNumber="288">Fig. 3</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection lastPageId="4" lastPageNumber="289" pageId="3" pageNumber="288" type="etymology">
<paragraph blockId="3.[136,1452,1227,2082]" box="[189,1152,2055,2082]" pageId="3" pageNumber="288">
<emphasis bold="true" box="[189,318,2056,2082]" pageId="3" pageNumber="288">Etymology</emphasis>
:—The specific epithet “ 
<taxonomicName authorityName="Jiji, Jeomol, Manudev &amp; Sunojk. A. A" authorityYear="2023" box="[594,702,2056,2081]" class="Magnoliopsida" family="Rubiaceae" genus="Hedyotis" kingdom="Plantae" order="Gentianales" pageId="3" pageNumber="288" phylum="Tracheophyta" rank="species" species="recurvata">
<emphasis box="[594,702,2056,2081]" italics="true" pageId="3" pageNumber="288">recurvata</emphasis>
</taxonomicName>
” refers to its strongly recurved stipules.
</paragraph>
<paragraph blockId="4.[136,1451,158,293]" pageId="4" pageNumber="289">
<emphasis bold="true" box="[189,430,158,184]" pageId="4" pageNumber="289">Conservation status</emphasis>
:— 
<materialsCitation country="India" county="Data Deficient" location="Avalanche in Nilgiris" municipality="According" pageId="4" pageNumber="289" specimenCount="2" stateProvince="Tamil Nadu">
Until now, 
<taxonomicName authorityName="Jiji, Jeomol, Manudev &amp; Sunojk. A. A" authorityYear="2023" box="[598,744,159,184]" class="Magnoliopsida" family="Rubiaceae" genus="Hedyotis" kingdom="Plantae" order="Gentianales" pageId="4" pageNumber="289" phylum="Tracheophyta" rank="species" species="recurvata">
<emphasis box="[598,744,159,184]" italics="true" pageId="4" pageNumber="289">H. recurvata</emphasis>
</taxonomicName>
is known only from the 
<typeStatus box="[1041,1089,159,185]" pageId="4" pageNumber="289">type</typeStatus>
locality, 
<location LSID="urn:lsid:plazi:treatment:CC7987FCFF8E5C4C52C732D9FC4B6769:410F6031FF885C4C56E132B3FA5A66F5" box="[1198,1447,158,184]" country="India" county="Data Deficient" municipality="According" name="Avalanche in Nilgiris" pageId="4" pageNumber="289" stateProvince="Tamil Nadu">Avalanche in Nilgiris</location>
, 
<collectingRegion box="[136,272,194,220]" country="India" name="Tamil Nadu" pageId="4" pageNumber="289">Tamil Nadu</collectingRegion>
. The population size at the type locality is very low. 
<location LSID="urn:lsid:plazi:treatment:CC7987FCFF8E5C4C52C732D9FC4B6769:410F6031FF885C4C512C32EEFC366690" box="[867,971,195,221]" country="India" county="Data Deficient" municipality="According" name="However" pageId="4" pageNumber="289" stateProvince="Tamil Nadu">However</location>
, thorough exploration in adjoining regions of 
<location LSID="urn:lsid:plazi:treatment:CC7987FCFF8E5C4C52C732D9FC4B6769:410F6031FF885C4C52E732CBFF03674D" box="[168,254,230,256]" country="India" county="Data Deficient" municipality="According" name="Nilgiris" pageId="4" pageNumber="289" stateProvince="Tamil Nadu">Nilgiris</location>
is needed to assess the distribution status of this species in 
<location LSID="urn:lsid:plazi:treatment:CC7987FCFF8E5C4C52C732D9FC4B6769:410F6031FF885C4C51E232CBFC13674D" box="[941,1006,230,256]" country="India" county="Data Deficient" municipality="According" name="South" pageId="4" pageNumber="289" stateProvince="Tamil Nadu">South</location>
<collectingCountry box="[1015,1076,230,256]" name="India" pageId="4" pageNumber="289">India</collectingCountry>
. 
<collectingMunicipality box="[1087,1205,230,256]" pageId="4" pageNumber="289">According</collectingMunicipality>
to IUCN criteria, the species falls under the category 
<collectingCounty box="[495,658,266,293]" pageId="4" pageNumber="289">Data Deficient</collectingCounty>
(DD) (IUCN SPS, 2022)
</materialsCitation>
.
</paragraph>
</subSubSection>
</treatment>
</document>