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<document id="87AB9B01F9375B78BB2FC20D0E0016E5" ID-DOI="10.11646/palaeoentomology.6.2.9" ID-ISSN="2624-2834" ID-ZooBank="BC5B57E2-679A-4B15-82F7-923825807F5E" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="GgImagineBatch" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1683888728508" checkinUser="plazi" docAuthor="STRELNIKOVA, OLESYA D. &amp; YAN, EVGENY V." docDate="2023" docId="B31E87ADFFACFFEF1AD8FA4BFD741CCC" docLanguage="en" docName="palaeoentomology.6.2.9.pdf" docOrigin="Palaeoentomology 6 (2)" docSource="http://dx.doi.org/10.11646/palaeoentomology.6.2.9" docStyle="DocumentStyle:4EE6FD780FF4521D541D0D0600705D0E.4:Palaeoentomology.2018-.journal_article" docStyleId="4EE6FD780FF4521D541D0D0600705D0E" docStyleName="Palaeoentomology.2018-.journal_article" docStyleVersion="4" docTitle="Notocupes oxypygus Ponomarenko 1969" docType="treatment" docVersion="2" lastPageNumber="188" masterDocId="4F27FFD5FFA0FFE11A50FF98FFDD191C" masterDocTitle="Redescriptions of the Triassic Notocupes beetles (Archostemata: Ommatidae) from Kyrgyzstan and South Kazakhstan" masterLastPageNumber="190" masterPageNumber="174" pageNumber="186" updateTime="1683889138702" updateUser="ExternalLinkService">
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<mods:title id="8E7930BF871486878C11D87C3017C95A">Redescriptions of the Triassic Notocupes beetles (Archostemata: Ommatidae) from Kyrgyzstan and South Kazakhstan</mods:title>
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<mods:namePart id="B1FEC77C4D7965C3D472CDA1FCE3AC37">STRELNIKOVA, OLESYA D.</mods:namePart>
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<subSubSection id="73AD6530FFACFFED1AD8FA4BFDB51CF0" box="[136,616,1490,1517]" pageId="12" pageNumber="186" type="nomenclature">
<paragraph id="3B0836BBFFACFFED1AD8FA4BFDB51CF0" blockId="12.[136,616,1490,1552]" box="[136,616,1490,1517]" pageId="12" pageNumber="186">
<heading id="604081D7FFACFFED1AD8FA4BFDB51CF0" bold="true" box="[136,616,1490,1517]" fontSize="10" level="2" pageId="12" pageNumber="186" reason="3">
<taxonomicName id="FCB74D38FFACFFED1AD8FA4BFDB51CF0" authority="Ponomarenko, 1969" authorityName="Ponomarenko" authorityYear="1969" box="[136,616,1490,1517]" class="Insecta" family="Ommatidae" genus="Notocupes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="186" phylum="Arthropoda" rank="species" species="oxypygus">
<emphasis id="09C3EAA9FFACFFED1AD8FA4BFDB51CF0" bold="true" box="[136,616,1490,1517]" pageId="12" pageNumber="186">
<emphasis id="09C3EAA9FFACFFED1AD8FA4BFEAE1CF1" bold="true" box="[136,371,1491,1517]" italics="true" pageId="12" pageNumber="186">Notocupes oxypygus</emphasis>
<bibRefCitation id="5F264B4AFFACFFED1B2BFA4AFDB51CF0" author="Ponomarenko, A. G." box="[379,616,1490,1516]" pageId="12" pageNumber="186" refId="ref10214" refString="Ponomarenko, A. G. (1969) Historical development of Coleoptera - Archostemata. In: Rohdendorf, B. B. (Ed.), Trudy Paleontologicheskogo Instituta, Akademiya Nauk SSSR, 125. Nauka, Moscow, 240 pp. [In Russian]" type="book" year="1969">Ponomarenko, 1969</bibRefCitation>
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="73AD6530FFACFFED1AD8FA6EFEC51F0C" box="[136,280,1526,1552]" pageId="12" pageNumber="186" type="description">
<paragraph id="3B0836BBFFACFFED1AD8FA6EFEC51F0C" blockId="12.[136,616,1490,1552]" box="[136,280,1526,1552]" pageId="12" pageNumber="186">
(
<figureCitation id="A38C2A3EFFACFFED1AC0FA6EFECD1F0C" box="[144,272,1526,1552]" captionStart-0="FIGURE 10" captionStart-1="FIGURE 11" captionStart-2="FIGURE 12" captionStartId-0="13.[136,235,1078,1102]" captionStartId-1="13.[136,235,1993,2017]" captionStartId-2="14.[136,235,1153,1177]" captionTargetBox-0="[183,1404,191,1054]" captionTargetBox-1="[184,1404,1177,1970]" captionTargetBox-2="[141,774,183,1130]" captionTargetId-0="figure-27@13.[183,1404,191,1054]" captionTargetId-1="figure-57@13.[183,1404,1176,1970]" captionTargetId-2="figure-654@14.[139,774,183,1130]" captionTargetPageId-0="13" captionTargetPageId-1="13" captionTargetPageId-2="14" captionText-0="FIGURE 10. Notocupes oxypygus, holotype (PIN, 1240/6, part). A, Dry photo. B, Interpretative line drawing, dorsal aspect. C, Interpretative line drawing, ventral aspect." captionText-1="FIGURE 11. Notocupes oxypygus, holotype (PIN, 1240/6, counterpart). A, Wetted with 95% ethanol. B, Dry photo.C, Interpretative line drawing." captionText-2="FIGURE 12. Notocupes oxypygus, paratype (PIN, 1361/128, part). A, Dry photo. B, Interpretative line drawing." pageId="12" pageNumber="186">Figs 1012</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="73AD6530FFACFFED1AD8F9DAFDE21F4B" box="[136,575,1602,1624]" pageId="12" pageNumber="186" type="reference_group">
<paragraph id="3B0836BBFFACFFED1AD8F9DAFDE21F4B" blockId="12.[136,575,1602,1624]" box="[136,575,1602,1624]" pageId="12" pageNumber="186">
<treatmentCitationGroup id="1BA71195FFACFFED1AD8F9DAFDE21F4B" box="[136,575,1602,1624]" pageId="12" pageNumber="186">
<treatmentCitation id="BA1610AAFFACFFED1AD8F9DAFDE11F4B" author="Ponomarenko, A. G." box="[136,572,1602,1624]" page="91" pageId="12" pageNumber="186" year="1969">
<taxonomicName id="FCB74D38FFACFFED1AD8F9DAFDE11F4B" authority="Ponomarenko, 1969: 91" authorityName="Ponomarenko" authorityPageNumber="91" authorityYear="1969" box="[136,572,1602,1624]" class="Insecta" family="Ommatidae" genus="Notocupes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="186" phylum="Arthropoda" rank="species" species="oxypygus">
<emphasis id="09C3EAA9FFACFFED1AD8F9DAFE911F4B" box="[136,332,1602,1623]" italics="true" pageId="12" pageNumber="186">Notocupes oxypygus</emphasis>
<bibRefCitation id="5F264B4AFFACFFED1B03F9DAFDE11F4B" author="Ponomarenko, A. G." box="[339,572,1602,1624]" pageId="12" pageNumber="186" refId="ref10214" refString="Ponomarenko, A. G. (1969) Historical development of Coleoptera - Archostemata. In: Rohdendorf, B. B. (Ed.), Trudy Paleontologicheskogo Instituta, Akademiya Nauk SSSR, 125. Nauka, Moscow, 240 pp. [In Russian]" type="book" year="1969">Ponomarenko, 1969: 91</bibRefCitation>
</taxonomicName>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="73AD6530FFACFFED1AD8F91EFF3A1E48" pageId="12" pageNumber="186" type="materials_examined">
<paragraph id="3B0836BBFFACFFED1AD8F91EFE591FD8" blockId="12.[136,778,1670,2092]" pageId="12" pageNumber="186">
<emphasis id="09C3EAA9FFACFFED1AD8F91EFF2A1FBC" bold="true" box="[136,247,1670,1696]" pageId="12" pageNumber="186">Material.</emphasis>
<materialsCitation id="8BDF3CE6FFACFFED1AAEF91EFE5D1FD8" collectionCode="PIN" pageId="12" pageNumber="186" specimenCode="PIN 1361" specimenCount="2" typeStatus="holotype">
<typeStatus id="E40C8819FFACFFED1AAEF91EFEBB1FBC" box="[254,358,1670,1696]" pageId="12" pageNumber="186">Holotype</typeStatus>
and
<typeStatus id="E40C8819FFACFFED1BCCF91FFE211FBD" box="[412,508,1671,1697]" pageId="12" pageNumber="186">paratype</typeStatus>
<specimenCode id="6B119EC0FFACFFED1853F91FFDAD1FBC" box="[515,624,1670,1697]" collectionCode="PIN" country="Russia" httpUri="http://grbio.org/cool/kjjc-ziy5" name="Paleontological Institute, Russian Academy of Sciences" pageId="12" pageNumber="186">PIN 1361</specimenCode>
/128, part and counterpart of elytron
</materialsCitation>
.
</paragraph>
<paragraph id="3B0836BBFFACFFED1AEDF956FF3A1E48" blockId="12.[136,778,1670,2092]" pageId="12" pageNumber="186">
<materialsCitation id="8BDF3CE6FFACFFED1AEDF956FF3A1E48" collectionCode="PIN" county="Ketmen'" location="Ketmen' Ridge" municipality="Uygur District" pageId="12" pageNumber="186" specimenCode="PIN 1240" specimenCount="1" stateProvince="Ongtustik Qazaqstan Oblysy" typeStatus="holotype">
<emphasis id="09C3EAA9FFACFFED1AEDF956FEEC1FF4" bold="true" box="[189,305,1742,1768]" pageId="12" pageNumber="186">
<typeStatus id="E40C8819FFACFFED1AEDF956FEF11FF4" box="[189,300,1742,1768]" pageId="12" pageNumber="186">Holotype</typeStatus>
.
</emphasis>
<specimenCode id="6B119EC0FFACFFED1B6AF957FE761FF4" box="[314,427,1742,1769]" collectionCode="PIN" country="Russia" httpUri="http://grbio.org/cool/kjjc-ziy5" name="Paleontological Institute, Russian Academy of Sciences" pageId="12" pageNumber="186">PIN 1240</specimenCode>
/6, body part without the head, pronotum and most legs,
<collectingCounty id="D2694E37FFACFFED1BCEF96AFE231E10" box="[414,510,1778,1804]" pageId="12" pageNumber="186">Ketmen</collectingCounty>
locality,
<collectingMunicipality id="DB6CACC1FFACFFED1832F96BFCDB1E10" box="[610,774,1778,1805]" pageId="12" pageNumber="186">Uygur District</collectingMunicipality>
,
<location id="3E686060FFACFFED1AD8F88EFEE61E2C" LSID="urn:lsid:plazi:treatment:B31E87ADFFACFFEF1AD8FA4BFD741CCC:3E686060FFACFFED1AD8F88EFEE61E2C" box="[136,315,1814,1840]" county="Ketmen'" municipality="Uygur District" name="Ketmen' Ridge" pageId="12" pageNumber="186" stateProvince="Ongtustik Qazaqstan Oblysy">Ketmen Ridge</location>
,
<location id="3E686060FFACFFED1B1CF88EFE161E2C" LSID="urn:lsid:plazi:treatment:B31E87ADFFACFFEF1AD8FA4BFD741CCC:3E686060FFACFFED1B1CF88EFE161E2C" box="[332,459,1814,1840]" county="Ketmen'" municipality="Uygur District" name="Kyzyl-Tam" pageId="12" pageNumber="186" stateProvince="Ongtustik Qazaqstan Oblysy">Kyzyl-Tam</location>
spring,
<collectingRegion id="F973F859FFACFFED1862F88EFCDB1E2C" box="[562,774,1814,1840]" country="Kazakhstan" name="Ongtustik Qazaqstan Oblysy" pageId="12" pageNumber="186">South Kazakhstan</collectingRegion>
, Triassic.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="73AD6530FFACFFED1AEDF8C6FD041114" pageId="12" pageNumber="186" type="diagnosis">
<paragraph id="3B0836BBFFACFFED1AEDF8C6FD041114" blockId="12.[136,778,1670,2092]" pageId="12" pageNumber="186">
<emphasis id="09C3EAA9FFACFFED1AEDF8C6FEE51E64" bold="true" box="[189,312,1886,1912]" pageId="12" pageNumber="186">Diagnosis.</emphasis>
Elytron with up to 32 cells in a row; elytral cells closer to its distal half round, cells on epipleuron larger than elsewhere on elytron; base of field IV widened and with three rows of cells instead of two. Рaracoxal suture merged with posterior margin of metaventrite.
</paragraph>
</subSubSection>
<subSubSection id="73AD6530FFACFFED1AEDF78AFC6B1C9C" pageId="12" pageNumber="186" type="description">
<paragraph id="3B0836BBFFACFFED1AEDF78AFAE119C3" blockId="12.[136,778,1670,2092]" lastBlockId="12.[810,1452,158,2093]" pageId="12" pageNumber="186">
<emphasis id="09C3EAA9FFACFFED1AEDF78AFE921130" bold="true" box="[189,335,2066,2092]" pageId="12" pageNumber="186">Description.</emphasis>
Cuticular tubercle size is approximately
<quantity id="FC4F9B5EFFACFFED197AFF06FC5019A5" box="[810,909,158,185]" metricMagnitude="-5" metricUnit="m" metricValue="3.0" pageId="12" pageNumber="186" unit="mm" value="0.03">0.03 mm</quantity>
, space between adjacent tubercles is 0.02; density of cuticular sculpture is 550 tubercles per mm².
</paragraph>
<paragraph id="3B0836BBFFACFFED190FFF7EFC3B1AB0" blockId="12.[810,1452,158,2093]" pageId="12" pageNumber="186">
<emphasis id="09C3EAA9FFACFFED190FFF7EFC6F181C" box="[863,946,230,256]" italics="true" pageId="12" pageNumber="186">Elytron</emphasis>
with apex not exceeding that of abdomen. Vein 2 straight, not following curved epipleuron, merging with vein 1 before elytron apex, space between vein convergence and elytron apex accommodating up to seven cells. Vein 3 straight, not following curved epipleuron. Veins 4 and 5 merge three cells before elytron apex.Elytron maximum width at its midlength. Epipleuron width ratio at elytron base and near apex 1.5. Elytron maximum width ratio to that of epipleuron 8.4. Humerus indistinct. Elytral apex rounded, not protruding posteriorly. Field I with single row of cells, other fields all with double rows; cells in field I 29; field II 25, field III 30, field IV 32, field V 23, field VI 27, VII 02. Maximum number of cells in row to elytral length ratio 4.3. Maculae present, varying in number from 45 at elytron base and apex to 78 at midlength. Bigger epipleural cells have more maculae embedded to cell edges. Cells in fields III and IV with 34 maculae near elytron base. Cells occupying more than half distance between veins, but clearly not reaching borders of veins.
</paragraph>
<paragraph id="3B0836BBFFACFFED190FFC2FFBEF1D04" blockId="12.[810,1452,158,2093]" pageId="12" pageNumber="186">
<emphasis id="09C3EAA9FFACFFED190FFC2FFC331ACC" box="[863,1006,951,976]" italics="true" pageId="12" pageNumber="186">Metaventrite</emphasis>
width at posterior margin to its anterior margin ratio 1.9. Рaracoxal suture merging with posterior margin of metaventrite.
</paragraph>
<paragraph id="3B0836BBFFACFFED190FFBBAFB091DD0" blockId="12.[810,1452,158,2093]" pageId="12" pageNumber="186">
<emphasis id="09C3EAA9FFACFFED190FFBBAFC141D20" box="[863,969,1058,1084]" italics="true" pageId="12" pageNumber="186">Abdomen</emphasis>
with weakly defined relief, with transverse impressions on anterior and posterior margins of sternites IIIV. Abdomen is not narrowing at the base, narrowing posteriorly at level of sternite IV, abdominal apex acute. Sternite length ratio is 1.6:1:1.1:1:2.4.
</paragraph>
<paragraph id="3B0836BBFFACFFED190FFB4FFA441DED" blockId="12.[810,1452,158,2093]" box="[863,1433,1238,1265]" pageId="12" pageNumber="186">
<emphasis id="09C3EAA9FFACFFED190FFB4FFC001DEC" box="[863,989,1239,1264]" italics="true" pageId="12" pageNumber="186">Mesocoxae</emphasis>
narrowly separated, weakly transverse.
</paragraph>
<paragraph id="3B0836BBFFACFFED190FFB63FC6B1C9C" blockId="12.[810,1452,158,2093]" pageId="12" pageNumber="186">
Measurements (in mm).
<typeStatus id="E40C8819FFACFFED1E23FB62FB061C08" box="[1139,1243,1274,1300]" pageId="12" pageNumber="186">Holotype</typeStatus>
PIN 1240/6: body length estimated 1013, width 3.8, elytral length 7.27.3, elytral width 1.9;
<typeStatus id="E40C8819FFACFFED19A5FADBFB881C41" box="[1013,1109,1347,1373]" pageId="12" pageNumber="186">paratype</typeStatus>
PIN 1361/128: elytron length 7.4, width 2.
</paragraph>
</subSubSection>
<subSubSection id="73AD6530FFACFFEF190FFA12FD741CCC" lastPageId="14" lastPageNumber="188" pageId="12" pageNumber="186" type="discussion">
<paragraph id="3B0836BBFFACFFEF190FFA12FE8A1C20" blockId="12.[810,1452,158,2093]" lastBlockId="14.[136,778,1277,1488]" lastPageId="14" lastPageNumber="188" pageId="12" pageNumber="186">
<emphasis id="09C3EAA9FFACFFED190FFA12FC0E1CB8" bold="true" box="[863,979,1418,1444]" pageId="12" pageNumber="186">Remarks.</emphasis>
Additional rows of cells at the elytron base are also known for
<taxonomicName id="FCB74D38FFACFFED1E15FA37FC7F1CF0" authority="Strelnikova &amp; Yan, 2021" authorityName="Strelnikova &amp; Yan" authorityYear="2021" class="Insecta" family="Ommatidae" genus="Zygadenia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="186" phylum="Arthropoda" rank="species" species="alexrasnitsyni">
<emphasis id="09C3EAA9FFACFFED1E15FA37FADF1CD4" box="[1093,1282,1454,1480]" italics="true" pageId="12" pageNumber="186">Z. alexrasnitsyni</emphasis>
<bibRefCitation id="5F264B4AFFACFFED1F5BFA36FC7F1CF0" author="Strelnikova, O. D. &amp; Yan, E. V." pageId="12" pageNumber="186" pagination="508 - 523" refId="ref10747" refString="Strelnikova, O. D. &amp; Yan, E. V. (2021) Redescriptions of beetles of the Notocupes generic complex (Coleoptera: Archostemata: Ommatidae) from the Lower Cretaceous of Buryatia. Palaeoentomology, 4 (5), 508 - 523. https: // doi. org / 10.11646 / palaeoentomology. 4.5.15" type="journal article" year="2021">Strelnikova &amp; Yan, 2021</bibRefCitation>
</taxonomicName>
. Cell rows with more than 30 cells found in
<taxonomicName id="FCB74D38FFACFFED197AFA6FFC6C1F0C" authorityName="Ponomarenko" authorityYear="1969" box="[810,945,1527,1553]" class="Insecta" family="Ommatidae" genus="Notocupes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="186" phylum="Arthropoda" rank="species" species="rostratus">
<emphasis id="09C3EAA9FFACFFED197AFA6FFCE01F0C" box="[810,829,1527,1552]" italics="true" pageId="12" pageNumber="186">N</emphasis>
.
<emphasis id="09C3EAA9FFACFFED191DFA6FFC6C1F0C" box="[845,945,1527,1552]" italics="true" pageId="12" pageNumber="186">rostratus</emphasis>
</taxonomicName>
,
<taxonomicName id="FCB74D38FFACFFED1992FA6FFAF41F0D" authority="Ponomarenko, 1985" authorityName="Ponomarenko" authorityYear="1985" box="[962,1321,1526,1553]" class="Insecta" family="Ommatidae" genus="Notocupes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="186" phylum="Arthropoda" rank="species" species="patulus">
<emphasis id="09C3EAA9FFACFFED1992FA6FFC081F0C" box="[962,981,1527,1552]" italics="true" pageId="12" pageNumber="186">N</emphasis>
.
<emphasis id="09C3EAA9FFACFFED19B6FA6EFBE51F0C" box="[998,1080,1526,1552]" italics="true" pageId="12" pageNumber="186">patulus</emphasis>
<bibRefCitation id="5F264B4AFFACFFED1E12FA6EFAF41F0D" author="Ponomarenko, A. G." box="[1090,1321,1526,1553]" pageId="12" pageNumber="186" pagination="47 - 87" refId="ref10264" refString="Ponomarenko, A. G. (1985) Coleoptera from the Jurassic of Siberia and Western Mongolia. In: Rasnitsyn, A. P. (Ed.), Jurassic insects of Siberia and Mongolia. Trudy Paleontologicheskogo Instituta, Akademiya Nauk SSSR, 211, 47 - 87. [In Russian]" type="journal article" year="1985">Ponomarenko, 1985</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="FCB74D38FFACFFED1F67FA6FFBDC1F28" authority="(Tan &amp; Ren, 2006)" authorityName="Tan &amp; Ren" authorityYear="2006" baseAuthorityName="Tan &amp; Ren" baseAuthorityYear="2006" class="Insecta" family="Ommatidae" genus="Ovatocupes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="186" phylum="Arthropoda" rank="species" species="alienus">
<emphasis id="09C3EAA9FFACFFED1F67FA6FFA761F0C" box="[1335,1451,1526,1552]" italics="true" pageId="12" pageNumber="186">N. alienus</emphasis>
(Tan &amp;
<bibRefCitation id="5F264B4AFFACFFED19D1F983FC241F28" author="Ren, D. &amp; Tan, J." box="[897,1017,1562,1589]" pageId="12" pageNumber="186" pagination="457 - 463" refId="ref10525" refString="Ren, D. &amp; Tan, J. (2006) A new cupedid genus (Coleoptera: Archostemata: Cupedidae) from Jehol biota of Western Liaoning, China. Annales Zoologici, 56 (3), 457 - 463." type="journal article" year="2006">Ren, 2006</bibRefCitation>
)
</taxonomicName>
,
<taxonomicName id="FCB74D38FFACFFED1E5CF983FAA11F29" authority="Jarzembowski, 2015" authorityName="Jarzembowski" authorityYear="2015" box="[1036,1404,1562,1589]" class="Insecta" family="Ommatidae" genus="Zygadenia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="186" phylum="Arthropoda" rank="species" species="simpsoni">
<emphasis id="09C3EAA9FFACFFED1E5CF983FBC11F28" box="[1036,1052,1563,1588]" italics="true" pageId="12" pageNumber="186">Z</emphasis>
.
<emphasis id="09C3EAA9FFACFFED1E7AF983FB531F28" box="[1066,1166,1563,1588]" italics="true" pageId="12" pageNumber="186">simpsoni</emphasis>
<bibRefCitation id="5F264B4AFFACFFED1EC5F982FAA11F29" author="Jarzembowski, E. A. &amp; Wang, B. &amp; Zhang, H. C. &amp; Fang, Y." box="[1173,1404,1562,1589]" pageId="12" pageNumber="186" pagination="431 - 439" refId="ref9420" refString="Jarzembowski, E. A., Wang, B., Zhang, H. C. &amp; Fang, Y. (2015) Boring beetles are not necessarily dull: New notocupedins (Insecta: Coleoptera) from the Mesozoic of Eurasia and East Gondwana. Cretaceous Research, 52, 431 - 439. https: // doi. org / 10.1016 / j. cretres. 2014.03.006" type="journal article" year="2015">Jarzembowski, 2015</bibRefCitation>
</taxonomicName>
and
<taxonomicName id="FCB74D38FFACFFED197AF9A7FAD91F44" authority="Strelnikova &amp; Yan, 2021" authorityName="Strelnikova &amp; Yan" authorityYear="2021" box="[810,1284,1598,1625]" class="Insecta" family="Ommatidae" genus="Zygadenia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="186" phylum="Arthropoda" rank="species" species="alexrasnitsyni">
<emphasis id="09C3EAA9FFACFFED197AF9A7FCE71F44" box="[810,826,1599,1624]" italics="true" pageId="12" pageNumber="186">Z</emphasis>
.
<emphasis id="09C3EAA9FFACFFED1917F9A6FC391F44" box="[839,996,1598,1624]" italics="true" pageId="12" pageNumber="186">alexrasnitsyni</emphasis>
<bibRefCitation id="5F264B4AFFACFFED19BBF9A6FAD91F44" author="Strelnikova, O. D. &amp; Yan, E. V." box="[1003,1284,1598,1625]" pageId="12" pageNumber="186" pagination="508 - 523" refId="ref10747" refString="Strelnikova, O. D. &amp; Yan, E. V. (2021) Redescriptions of beetles of the Notocupes generic complex (Coleoptera: Archostemata: Ommatidae) from the Lower Cretaceous of Buryatia. Palaeoentomology, 4 (5), 508 - 523. https: // doi. org / 10.11646 / palaeoentomology. 4.5.15" type="journal article" year="2021">Strelnikova &amp; Yan, 2021</bibRefCitation>
</taxonomicName>
. The species is similar to
<taxonomicName id="FCB74D38FFACFFED19C5F9FBFBCA1F60" authorityName="Ponomarenko" authorityYear="1969" box="[917,1047,1635,1661]" class="Insecta" family="Ommatidae" genus="Notocupes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="186" phylum="Arthropoda" rank="species" species="rostratus">
<emphasis id="09C3EAA9FFACFFED19C5F9FBFC751F60" box="[917,936,1635,1660]" italics="true" pageId="12" pageNumber="186">N</emphasis>
.
<emphasis id="09C3EAA9FFACFFED19E3F9FBFBCA1F60" box="[947,1047,1635,1660]" italics="true" pageId="12" pageNumber="186">rostratus</emphasis>
</taxonomicName>
in strongly acute abdominal apex but differs in median fossa on abdominal apex and humerus indistinct. It differs from
<taxonomicName id="FCB74D38FFACFFED1E01F933FB1B1FD8" authorityName="Ponomarenko" authorityYear="1985" box="[1105,1222,1706,1733]" class="Insecta" family="Ommatidae" genus="Notocupes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="186" phylum="Arthropoda" rank="species" species="patulus">
<emphasis id="09C3EAA9FFACFFED1E01F933FBB91FD8" box="[1105,1124,1707,1732]" italics="true" pageId="12" pageNumber="186">N</emphasis>
.
<emphasis id="09C3EAA9FFACFFED1E24F932FB1B1FD8" box="[1140,1222,1706,1732]" italics="true" pageId="12" pageNumber="186">patulus</emphasis>
</taxonomicName>
in paracoxal suture merging with posterior margin of metaventrite; from
<taxonomicName id="FCB74D38FFACFFED1FC2F957FCA61E10" authorityName="Tan &amp; Ren" authorityYear="2006" baseAuthorityName="Tan &amp; Ren" baseAuthorityYear="2006" class="Insecta" family="Ommatidae" genus="Ovatocupes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="186" phylum="Arthropoda" rank="species" species="alienus">
<emphasis id="09C3EAA9FFACFFED1FC2F957FA781FF4" box="[1426,1445,1743,1768]" italics="true" pageId="12" pageNumber="186">N</emphasis>
.
<emphasis id="09C3EAA9FFACFFED197AF96AFCA61E10" box="[810,891,1778,1804]" italics="true" pageId="12" pageNumber="186">alienus</emphasis>
</taxonomicName>
in epipleuron without cells; from
<taxonomicName id="FCB74D38FFACFFED1F58F96BFA511E10" authorityName="Jarzembowski" authorityYear="2015" box="[1288,1420,1779,1805]" class="Insecta" family="Ommatidae" genus="Zygadenia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="186" phylum="Arthropoda" rank="species" species="simpsoni">
<emphasis id="09C3EAA9FFACFFED1F58F96BFAC51E10" box="[1288,1304,1779,1804]" italics="true" pageId="12" pageNumber="186">Z</emphasis>
.
<emphasis id="09C3EAA9FFACFFED1F78F96BFA511E10" box="[1320,1420,1779,1804]" italics="true" pageId="12" pageNumber="186">simpsoni</emphasis>
</taxonomicName>
in smaller size and elytron without colour pattern; from
<taxonomicName id="FCB74D38FFACFFED1FC5F88FFC1F1E48" class="Insecta" family="Ommatidae" genus="Zygadenia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="186" phylum="Arthropoda" rank="species" species="alexrasnitsini">
<emphasis id="09C3EAA9FFACFFED1FC5F88FFA781E2C" box="[1429,1445,1815,1840]" italics="true" pageId="12" pageNumber="186">Z</emphasis>
.
<emphasis id="09C3EAA9FFACFFED197AF8A2FC1F1E48" box="[810,962,1850,1876]" italics="true" pageId="12" pageNumber="186">alexrasnitsini</emphasis>
</taxonomicName>
in smaller size, wider base of epipleuron and smaller VII field with only 2 cells.
<taxonomicName id="FCB74D38FFACFFED1E86F8C7FA801E64" authorityName="Ponomarenko" authorityYear="1969" box="[1238,1373,1887,1913]" class="Insecta" family="Ommatidae" genus="Notocupes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="186" phylum="Arthropoda" rank="species" species="oxypygus">
<emphasis id="09C3EAA9FFACFFED1E86F8C7FB341E64" box="[1238,1257,1887,1912]" italics="true" pageId="12" pageNumber="186">N</emphasis>
.
<emphasis id="09C3EAA9FFACFFED1EA5F8C7FA801E64" box="[1269,1373,1887,1912]" italics="true" pageId="12" pageNumber="186">oxypygus</emphasis>
</taxonomicName>
differs from other Triassic species in paracoxal suture merging with the posterior margin of the metaventrite, but similar to them in having dense, uniform cuticular tubercles. Differs from all other species of the genus in the relief of abdomen, in which depressions are visible not only along the anterior margin of the sternite, but also along the posterior ones.
</paragraph>
<caption id="6FC86633FFADFFEC1AD8FBAEFDEB1D6D" pageId="13" pageNumber="187" startId="13.[136,235,1078,1102]" targetBox="[183,1404,191,1054]" targetPageId="13" targetType="figure">
<paragraph id="3B0836BBFFADFFEC1AD8FBAEFDEB1D6D" blockId="13.[136,1451,1078,1138]" pageId="13" pageNumber="187">
<emphasis id="09C3EAA9FFADFFEC1AD8FBAEFECF1D52" bold="true" box="[136,274,1078,1102]" pageId="13" pageNumber="187">FIGURE 10.</emphasis>
<taxonomicName id="FCB74D38FFADFFEC1B4AFBAFFE371D52" authorityName="Ponomarenko" authorityYear="1969" box="[282,490,1079,1102]" class="Insecta" family="Ommatidae" genus="Notocupes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="187" phylum="Arthropoda" rank="species" species="oxypygus">
<emphasis id="09C3EAA9FFADFFEC1B4AFBAFFE371D52" box="[282,490,1079,1102]" italics="true" pageId="13" pageNumber="187">Notocupes oxypygus</emphasis>
</taxonomicName>
, holotype (PIN, 1240/6, part).
<emphasis id="09C3EAA9FFADFFEC197BFBAEFCE01D52" bold="true" box="[811,829,1078,1102]" pageId="13" pageNumber="187">A</emphasis>
, Dry photo.
<emphasis id="09C3EAA9FFADFFEC1992FBAEFC0E1D52" bold="true" box="[962,979,1078,1102]" pageId="13" pageNumber="187">B</emphasis>
, Interpretative line drawing, dorsal aspect.
<emphasis id="09C3EAA9FFADFFEC1FC3FBAEFA781D52" bold="true" box="[1427,1445,1078,1102]" pageId="13" pageNumber="187">C</emphasis>
, Interpretative line drawing, ventral aspect.
</paragraph>
</caption>
<caption id="6FC86633FFADFFEC1AD8F851FED21119" pageId="13" pageNumber="187" startId="13.[136,235,1993,2017]" targetBox="[184,1404,1177,1970]" targetPageId="13" targetType="figure">
<paragraph id="3B0836BBFFADFFEC1AD8F851FED21119" blockId="13.[136,1451,1993,2053]" pageId="13" pageNumber="187">
<emphasis id="09C3EAA9FFADFFEC1AD8F851FED01EFD" bold="true" box="[136,269,1993,2017]" pageId="13" pageNumber="187">FIGURE 11.</emphasis>
<taxonomicName id="FCB74D38FFADFFEC1B41F852FE021EFD" authorityName="Ponomarenko" authorityYear="1969" box="[273,479,1994,2017]" class="Insecta" family="Ommatidae" genus="Notocupes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="187" phylum="Arthropoda" rank="species" species="oxypygus">
<emphasis id="09C3EAA9FFADFFEC1B41F852FE021EFD" box="[273,479,1994,2017]" italics="true" pageId="13" pageNumber="187">Notocupes oxypygus</emphasis>
</taxonomicName>
, holotype (PIN, 1240/6, counterpart).
<emphasis id="09C3EAA9FFADFFEC190AF851FCB11EFD" bold="true" box="[858,876,1993,2017]" pageId="13" pageNumber="187">A</emphasis>
, Wetted with 95% ethanol.
<emphasis id="09C3EAA9FFADFFEC1E2BF852FB511EFE" bold="true" box="[1147,1164,1994,2018]" pageId="13" pageNumber="187">B</emphasis>
, Dry photo.
<emphasis id="09C3EAA9FFADFFEC1F56F851FAC51EFD" bold="true" box="[1286,1304,1993,2017]" pageId="13" pageNumber="187">C</emphasis>
, Interpretative line drawing.
</paragraph>
</caption>
<caption id="6FC86633FFAEFFEF1AD8FB19FD561DA1" pageId="14" pageNumber="188" startId="14.[136,235,1153,1177]" targetBox="[141,774,183,1130]" targetPageId="14" targetType="figure">
<paragraph id="3B0836BBFFAEFFEF1AD8FB19FD561DA1" blockId="14.[136,778,1153,1214]" pageId="14" pageNumber="188">
<emphasis id="09C3EAA9FFAEFFEF1AD8FB19FEC91D85" bold="true" box="[136,276,1153,1177]" pageId="14" pageNumber="188">FIGURE 12.</emphasis>
<taxonomicName id="FCB74D38FFAEFFEF1B4EFB1AFE2C1D85" authorityName="Ponomarenko" authorityYear="1969" box="[286,497,1154,1177]" class="Insecta" family="Ommatidae" genus="Notocupes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="188" phylum="Arthropoda" rank="species" species="oxypygus">
<emphasis id="09C3EAA9FFAEFFEF1B4EFB1AFE2C1D85" box="[286,497,1154,1177]" italics="true" pageId="14" pageNumber="188">Notocupes oxypygus</emphasis>
</taxonomicName>
, paratype (PIN, 1361/128, part).
<emphasis id="09C3EAA9FFAEFFEF1A95FB3DFF0A1DA1" bold="true" box="[197,215,1189,1213]" pageId="14" pageNumber="188">A</emphasis>
, Dry photo.
<emphasis id="09C3EAA9FFAEFFEF1B08FB3EFEB41DA2" bold="true" box="[344,361,1190,1214]" pageId="14" pageNumber="188">B</emphasis>
, Interpretative line drawing.
</paragraph>
</caption>
<paragraph id="3B0836BBFFAEFFEF1AEDFADFFD741CCC" blockId="14.[136,778,1277,1488]" pageId="14" pageNumber="188">
Portion of the right elytron is most likely deformed on the
<typeStatus id="E40C8819FFAEFFEF1A80FAF3FEE81C99" box="[208,309,1387,1413]" pageId="14" pageNumber="188">holotype</typeStatus>
, due to experiencing tectonic deformation, thus the additional “line” visible between the sutural margin and vein 5 is most likely a bending zone.
</paragraph>
</subSubSection>
</treatment>
</document>