432 lines
56 KiB
XML
432 lines
56 KiB
XML
<document id="45409F1E8E75BEDD49E862B6F7328091" ID-DOI="10.11646/palaeoentomology.6.2.9" ID-ISSN="2624-2834" ID-Zenodo-Dep="7929106" ID-ZooBank="BC5B57E2-679A-4B15-82F7-923825807F5E" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="GgImagineBatch" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1683888728508" checkinUser="plazi" docAuthor="STRELNIKOVA, OLESYA D. & YAN, EVGENY V." docDate="2023" docId="B31E87ADFFA3FFE61AD8FC2FFF7618A9" docLanguage="en" docName="palaeoentomology.6.2.9.pdf" docOrigin="Palaeoentomology 6 (2)" docSource="http://dx.doi.org/10.11646/palaeoentomology.6.2.9" docStyle="DocumentStyle:4EE6FD780FF4521D541D0D0600705D0E.4:Palaeoentomology.2018-.journal_article" docStyleId="4EE6FD780FF4521D541D0D0600705D0E" docStyleName="Palaeoentomology.2018-.journal_article" docStyleVersion="4" docTitle="Notocupes laticella Ponomarenko 1969" docType="treatment" docVersion="2" lastPageNumber="180" masterDocId="4F27FFD5FFA0FFE11A50FF98FFDD191C" masterDocTitle="Redescriptions of the Triassic Notocupes beetles (Archostemata: Ommatidae) from Kyrgyzstan and South Kazakhstan" masterLastPageNumber="190" masterPageNumber="174" pageNumber="177" updateTime="1683889822166" updateUser="ExternalLinkService">
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<mods:title id="96011F1555C160815A93BB4958D91658">Redescriptions of the Triassic Notocupes beetles (Archostemata: Ommatidae) from Kyrgyzstan and South Kazakhstan</mods:title>
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<mods:namePart id="F7BCE27EEF8FAD600DB0F589A9F98B64">STRELNIKOVA, OLESYA D.</mods:namePart>
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<mods:namePart id="D313164A38EA257BBD6ACD2B39AD424B">YAN, EVGENY V.</mods:namePart>
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<mods:title id="D90112520C8A2795A440970CD033280D">Palaeoentomology</mods:title>
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<mods:part id="F67D93C6FD97B2A363B492B0CD2D8422">
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<mods:date id="3EF3E4114D4612C961A55C7E65FC6E27">2023</mods:date>
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<mods:number id="7067F40F3113D96D4E5B91D6E3BCAAEE">2023-04-28</mods:number>
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<treatment id="B31E87ADFFA3FFE61AD8FC2FFF7618A9" LSID="urn:lsid:plazi:treatment:B31E87ADFFA3FFE61AD8FC2FFF7618A9" httpUri="http://treatment.plazi.org/id/B31E87ADFFA3FFE61AD8FC2FFF7618A9" lastPageId="7" lastPageNumber="180" pageId="3" pageNumber="177">
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<subSubSection id="73AD6530FFA3FFE21AD8FC2FFD841ACC" box="[136,601,950,977]" pageId="3" pageNumber="177" type="nomenclature">
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<paragraph id="3B0836BBFFA3FFE21AD8FC2FFD841ACC" blockId="3.[136,601,950,1012]" box="[136,601,950,977]" pageId="3" pageNumber="177">
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<heading id="604081D7FFA3FFE21AD8FC2FFD841ACC" bold="true" box="[136,601,950,977]" fontSize="10" level="2" pageId="3" pageNumber="177" reason="3">
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<taxonomicName id="FCB74D38FFA3FFE21AD8FC2FFD841ACC" authority="Ponomarenko, 1969" authorityName="Ponomarenko" authorityYear="1969" box="[136,601,950,977]" class="Insecta" family="Ommatidae" genus="Notocupes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="177" phylum="Arthropoda" rank="species" species="laticella">
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<emphasis id="09C3EAA9FFA3FFE21AD8FC2FFD841ACC" bold="true" box="[136,601,950,977]" pageId="3" pageNumber="177">
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<emphasis id="09C3EAA9FFA3FFE21AD8FC2FFEB91ACD" bold="true" box="[136,356,950,977]" italics="true" pageId="3" pageNumber="177">Notocupes laticella</emphasis>
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<bibRefCitation id="5F264B4AFFA3FFE21B3BFC2EFD841ACC" author="Ponomarenko, A. G." box="[363,601,950,976]" pageId="3" pageNumber="177" refId="ref10214" refString="Ponomarenko, A. G. (1969) Historical development of Coleoptera - Archostemata. In: Rohdendorf, B. B. (Ed.), Trudy Paleontologicheskogo Instituta, Akademiya Nauk SSSR, 125. Nauka, Moscow, 240 pp. [In Russian]" type="book" year="1969">Ponomarenko, 1969</bibRefCitation>
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</emphasis>
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</taxonomicName>
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</heading>
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</paragraph>
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||
</subSubSection>
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<subSubSection id="73AD6530FFA3FFE21AD8FC42FF211AE8" box="[136,252,986,1012]" pageId="3" pageNumber="177" type="description">
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<paragraph id="3B0836BBFFA3FFE21AD8FC42FF211AE8" blockId="3.[136,601,950,1012]" box="[136,252,986,1012]" pageId="3" pageNumber="177">
|
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(
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||
<figureCitation id="A38C2A3EFFA3FFE21AC0FC42FF2E1AE8" box="[144,243,986,1012]" captionStart-0="FIGURE 1" captionStart-1="FIGURE 2" captionStart-2="FIGURE 3" captionStartId-0="3.[136,235,1925,1949]" captionStartId-1="4.[136,235,1812,1836]" captionStartId-2="5.[136,235,1951,1975]" captionTargetBox-0="[155,1436,1205,1894]" captionTargetBox-1="[273,1314,189,1788]" captionTargetBox-2="[207,1381,189,1923]" captionTargetId-0="figure-396@3.[151,1436,1199,1902]" captionTargetId-1="figure-17@4.[273,1314,189,1788]" captionTargetId-2="figure-27@5.[206,1381,189,1928]" captionTargetPageId-0="3" captionTargetPageId-1="4" captionTargetPageId-2="5" captionText-0="FIGURE 1. Notocupes laticella, holotype (PIN, 2555/1663). A, Photo of part. B, Interpretative line drawing of part. C, Mirrored photo of counterpart wetted with 95% ethanol. D, Interpretative line drawing of mirrored counterpart. Abbreviations here and elsewhere: an.p, antennal pit; cl-lab, clypeolabrum; el, elevation; P1, supraantennal protuberance; pr.cx, procoxae; pr.re, procoxal rests. Pink filling indicates mandibles; light grey—head; dark grey—profemur." captionText-1="FIGURE 2. Notocupes laticella, paratypes (PIN, 2240/71, 74). A, Wetted with 95% ethanol PIN, 2240/74. B, Interpretative line drawing (square here and beyond shows relative size and density of tubercles). С, General scheme of elytra. D, Wetted with 95% ethanol PIN, 2240/71, counterpart. E, Interpretative line drawing. F, Interpretative line drawing of PIN, 2240/71, part. G, Photo of dry PIN, 2240/71, part. Abbreviations here and elsewhere: in.cx.p, intercoxal process; mesan, mesanepisternum; mesep, mesepimeron, mesotr, mesotrochantin; mesovent, mesoventrite; metanep, metanepimeron; metatr, metatrochantin; metavent, metaventrite; mnd, mandible; P2, supraocular protuberance; P3, posteromesal protuberance; s.n.pl, notopleural suture; s.pl.st, pleurosternal suture." captionText-2="FIGURE 3. Notocupes laticella, paratypes. A–C, PIN, 2555/1662. Wetted with 95% ethanol (A), dry (B) and interpretative line drawing (C). D, Photo of dry PIN, 2555/1686. E–G, PIN, 2240/6. Wetted with 95% ethanol (E) and dry (F). G, Interpretative line drawing. Abbreviations: pr.tr, protrochantin; pr.tr-er, protrochanter; pr.tib, protibia." figureDoi-0="http://doi.org/10.5281/zenodo.7929108" figureDoi-1="http://doi.org/10.5281/zenodo.7929110" figureDoi-2="http://doi.org/10.5281/zenodo.7929114" httpUri-0="https://zenodo.org/record/7929108/files/figure.png" httpUri-1="https://zenodo.org/record/7929110/files/figure.png" httpUri-2="https://zenodo.org/record/7929114/files/figure.png" pageId="3" pageNumber="177">Figs 1–3</figureCitation>
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)
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</paragraph>
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</subSubSection>
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<subSubSection id="73AD6530FFA3FFE21AD8FBBFFDE81D27" box="[136,565,1062,1084]" pageId="3" pageNumber="177" type="reference_group">
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<paragraph id="3B0836BBFFA3FFE21AD8FBBFFDE81D27" blockId="3.[136,565,1062,1084]" box="[136,565,1062,1084]" pageId="3" pageNumber="177">
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<treatmentCitationGroup id="1BA71195FFA3FFE21AD8FBBFFDE81D27" box="[136,565,1062,1084]" pageId="3" pageNumber="177">
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<treatmentCitation id="BA1610AAFFA3FFE21AD8FBBFFDEC1D27" author="Ponomarenko, A. G." box="[136,561,1062,1084]" page="90" pageId="3" pageNumber="177" year="1969">
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<taxonomicName id="FCB74D38FFA3FFE21AD8FBBFFDEC1D27" authority="Ponomarenko, 1969: 90" authorityName="Ponomarenko" authorityPageNumber="90" authorityYear="1969" box="[136,561,1062,1084]" class="Insecta" family="Ommatidae" genus="Notocupes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="177" phylum="Arthropoda" rank="species" species="laticella">
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<emphasis id="09C3EAA9FFA3FFE21AD8FBBFFE9F1D27" box="[136,322,1062,1084]" italics="true" pageId="3" pageNumber="177">Notocupes laticella</emphasis>
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<bibRefCitation id="5F264B4AFFA3FFE21B18FBBEFDEC1D27" author="Ponomarenko, A. G." box="[328,561,1062,1084]" pageId="3" pageNumber="177" refId="ref10214" refString="Ponomarenko, A. G. (1969) Historical development of Coleoptera - Archostemata. In: Rohdendorf, B. B. (Ed.), Trudy Paleontologicheskogo Instituta, Akademiya Nauk SSSR, 125. Nauka, Moscow, 240 pp. [In Russian]" type="book" year="1969">Ponomarenko, 1969: 90</bibRefCitation>
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</taxonomicName>
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||
</treatmentCitation>
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.
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||
</treatmentCitationGroup>
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||
</paragraph>
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||
</subSubSection>
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||
<subSubSection id="73AD6530FFA3FFE21AD8FBF2FBCA1865" pageId="3" pageNumber="177" type="materials_examined">
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||
<paragraph id="3B0836BBFFA3FFE21AD8FBF2FBC419C1" blockId="3.[136,777,1130,1157]" lastBlockId="3.[810,1452,158,1156]" pageId="3" pageNumber="177">
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||
<emphasis id="09C3EAA9FFA3FFE21AD8FBF2FF2D1D98" bold="true" box="[136,240,1130,1156]" pageId="3" pageNumber="177">Material</emphasis>
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.
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<materialsCitation id="8BDF3CE6FFA3FFE21B5DFBF2FBC419C1" collectionCode="PIN" pageId="3" pageNumber="177" specimenCode="PIN 2240" specimenCount="1" typeStatus="holotype">
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<typeStatus id="E40C8819FFA3FFE21B5DFBF2FEA81D98" box="[269,373,1130,1156]" pageId="3" pageNumber="177">Holotype</typeStatus>
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||
and
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||
<specimenCount id="2DB1FD32FFA3FFE21B9AFBF2FDA71D99" box="[458,634,1130,1157]" pageId="3" pageNumber="177" type="generic" typeStatus="paratypes">five paratypes</specimenCount>
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, parts and counterparts of almost complete bodies
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<specimenCode id="6B119EC0FFA3FFE21EA4FF07FAB819A4" box="[1268,1381,158,185]" collectionCode="PIN" country="Russia" httpUri="http://grbio.org/cool/kjjc-ziy5" name="Paleontological Institute, Russian Academy of Sciences" pageId="3" pageNumber="177">PIN 2240</specimenCode>
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/6, 71, 74, 2555/1662, 1686.
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</materialsCitation>
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||
</paragraph>
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||
<paragraph id="3B0836BBFFA3FFE2190FFF72FBCA1865" blockId="3.[810,1452,158,1156]" pageId="3" pageNumber="177">
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||
<materialsCitation id="8BDF3CE6FFA3FFE2190FFF72FBCA1865" collectionCode="PIN" country="Kyrgyzstan" county="Dzhayloucho" location="Middle-Upper Triassic" municipality="Batken region" pageId="3" pageNumber="177" specimenCode="PIN 2555" specimenCount="1" stateProvince="Osh Shaary" typeStatus="holotype">
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<emphasis id="09C3EAA9FFA3FFE2190FFF72FC0E1818" bold="true" box="[863,979,234,260]" pageId="3" pageNumber="177">
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<typeStatus id="E40C8819FFA3FFE2190FFF72FC131818" box="[863,974,234,260]" pageId="3" pageNumber="177">Holotype</typeStatus>
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.
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</emphasis>
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<specimenCode id="6B119EC0FFA3FFE2198EFF72FB8D1819" box="[990,1104,234,261]" collectionCode="PIN" country="Russia" httpUri="http://grbio.org/cool/kjjc-ziy5" name="Paleontological Institute, Russian Academy of Sciences" pageId="3" pageNumber="177">PIN 2555</specimenCode>
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/1663, part and counterpart of almost complete body,
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<collectingCounty id="D2694E37FFA3FFE21E7AFE89FB621837" box="[1066,1215,273,299]" pageId="3" pageNumber="177">Dzhayloucho</collectingCounty>
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locality,
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<collectingCountry id="43A0762BFFA3FFE21F74FE89FA7A1837" box="[1316,1447,273,299]" name="Kyrgyzstan" pageId="3" pageNumber="177">Kyrgyzstan</collectingCountry>
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,
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<collectingRegion id="F973F859FFA3FFE2197AFEA0FC11184E" box="[810,972,312,338]" country="Kyrgyzstan" name="Osh Shaary" pageId="3" pageNumber="177">Osh Province</collectingRegion>
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,
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<collectingMunicipality id="DB6CACC1FFA3FFE219B0FEA0FB54184E" box="[992,1161,312,338]" pageId="3" pageNumber="177">Batken region</collectingMunicipality>
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,
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<location id="3E686060FFA3FFE21ECDFEA0FA75184E" LSID="urn:lsid:plazi:treatment:B31E87ADFFA3FFE61AD8FC2FFF7618A9:3E686060FFA3FFE21ECDFEA0FA75184E" box="[1181,1448,312,338]" country="Kyrgyzstan" county="Dzhayloucho" municipality="Batken region" name="Middle-Upper Triassic" pageId="3" pageNumber="177" stateProvince="Osh Shaary">Middle-Upper Triassic</location>
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, Madygen Formation.
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</materialsCitation>
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||
</paragraph>
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||
</subSubSection>
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||
<subSubSection id="73AD6530FFA3FFE2190FFE1DFC561BAD" pageId="3" pageNumber="177" type="diagnosis">
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||
<paragraph id="3B0836BBFFA3FFE2190FFE1DFC561BAD" blockId="3.[810,1452,158,1156]" pageId="3" pageNumber="177">
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<emphasis id="09C3EAA9FFA3FFE2190FFE1DFC0F1883" bold="true" box="[863,978,389,415]" pageId="3" pageNumber="177">Diagnosis</emphasis>
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. Body densely covered with uniform circular tubercles. Antennae distinctly reaching behind base of pronotum; antennomeres 5–10 of subequal length, apical one longer than others. Base of antennae covered by supraantennal protuberance (Р1). Procoxae at pronotum mid-length. Main elytral veins differing only slightly from intercalary ones; no more than 24 cells in one row.
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</paragraph>
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</subSubSection>
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||
<subSubSection id="73AD6530FFA3FFE7190FFD25FE62111B" lastPageId="6" lastPageNumber="180" pageId="3" pageNumber="177" type="description">
|
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<paragraph id="3B0836BBFFA3FFE2190FFD25FC211A52" blockId="3.[810,1452,158,1156]" pageId="3" pageNumber="177">
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||
<emphasis id="09C3EAA9FFA3FFE2190FFD25FC2C1BCB" bold="true" box="[863,1009,701,727]" pageId="3" pageNumber="177">Description.</emphasis>
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Body to head length ratio 5.2–5.7. Cuticular tubercle size 0.02–0.03, space between adjacent tubercles 0.01–0.03; density of cuticular sculpture 380 tubercles per mm².
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</paragraph>
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<paragraph id="3B0836BBFFA3FFE7190FFCC1FE1E1BE9" blockId="3.[810,1452,158,1156]" lastBlockId="6.[136,778,158,2056]" lastPageId="6" lastPageNumber="180" pageId="3" pageNumber="177">
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||
<emphasis id="09C3EAA9FFA3FFE2190FFCC1FC411A6F" box="[863,924,857,883]" italics="true" pageId="3" pageNumber="177">Head</emphasis>
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length definitely exceeds width, its basal 1/3 covered by pronotum. Eyes lie close to mandibular bases or distance between them not exceeding eye diameter; head length behind eyes 1.6–1.7× longer than eye diameter. Head not narrow posteriorly, only slightly so in front of eyes. Mandibles large, protruding forward, more or less straight, with only apices curved inwards, with three apical denticles with a vertical arrangement of teeth (
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<figureCitation id="A38C2A3EFFA6FFE71AA4FF06FEE719A4" box="[244,314,158,184]" captionStart="FIGURE 1" captionStartId="3.[136,235,1925,1949]" captionTargetBox="[155,1436,1205,1894]" captionTargetId="figure-396@3.[151,1436,1199,1902]" captionTargetPageId="3" captionText="FIGURE 1. Notocupes laticella, holotype (PIN, 2555/1663). A, Photo of part. B, Interpretative line drawing of part. C, Mirrored photo of counterpart wetted with 95% ethanol. D, Interpretative line drawing of mirrored counterpart. Abbreviations here and elsewhere: an.p, antennal pit; cl-lab, clypeolabrum; el, elevation; P1, supraantennal protuberance; pr.cx, procoxae; pr.re, procoxal rests. Pink filling indicates mandibles; light grey—head; dark grey—profemur." figureDoi="http://doi.org/10.5281/zenodo.7929108" httpUri="https://zenodo.org/record/7929108/files/figure.png" pageId="6" pageNumber="180">Fig. 1</figureCitation>
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, pink filling). Clypeolabrum protruding forward, have elongate-oval shape (
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<figureCitation id="A38C2A3EFFA6FFE71810FF5CFD0F19C3" box="[576,722,196,223]" captionStart="FIGURE 1" captionStartId="3.[136,235,1925,1949]" captionTargetBox="[155,1436,1205,1894]" captionTargetId="figure-396@3.[151,1436,1199,1902]" captionTargetPageId="3" captionText="FIGURE 1. Notocupes laticella, holotype (PIN, 2555/1663). A, Photo of part. B, Interpretative line drawing of part. C, Mirrored photo of counterpart wetted with 95% ethanol. D, Interpretative line drawing of mirrored counterpart. Abbreviations here and elsewhere: an.p, antennal pit; cl-lab, clypeolabrum; el, elevation; P1, supraantennal protuberance; pr.cx, procoxae; pr.re, procoxal rests. Pink filling indicates mandibles; light grey—head; dark grey—profemur." figureDoi="http://doi.org/10.5281/zenodo.7929108" httpUri="https://zenodo.org/record/7929108/files/figure.png" pageId="6" pageNumber="180">Figs 1B, D</figureCitation>
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,
|
||
<figureCitation id="A38C2A3EFFA6FFE718B7FF5CFCD819C2" box="[743,773,196,222]" captionStart="FIGURE 2" captionStartId="4.[136,235,1812,1836]" captionTargetBox="[273,1314,189,1788]" captionTargetId="figure-17@4.[273,1314,189,1788]" captionTargetPageId="4" captionText="FIGURE 2. Notocupes laticella, paratypes (PIN, 2240/71, 74). A, Wetted with 95% ethanol PIN, 2240/74. B, Interpretative line drawing (square here and beyond shows relative size and density of tubercles). С, General scheme of elytra. D, Wetted with 95% ethanol PIN, 2240/71, counterpart. E, Interpretative line drawing. F, Interpretative line drawing of PIN, 2240/71, part. G, Photo of dry PIN, 2240/71, part. Abbreviations here and elsewhere: in.cx.p, intercoxal process; mesan, mesanepisternum; mesep, mesepimeron, mesotr, mesotrochantin; mesovent, mesoventrite; metanep, metanepimeron; metatr, metatrochantin; metavent, metaventrite; mnd, mandible; P2, supraocular protuberance; P3, posteromesal protuberance; s.n.pl, notopleural suture; s.pl.st, pleurosternal suture." figureDoi="http://doi.org/10.5281/zenodo.7929110" httpUri="https://zenodo.org/record/7929110/files/figure.png" pageId="6" pageNumber="180">2F</figureCitation>
|
||
, cl-lab). Eyes large, slightly protruding laterally, with longitudinal diameter is 0.5, transverse diameter 0.2– 0.3, head width (excluding eyes) to eye diameter 4–4.3. Antennae filiform, attached immediately in front of eyes, antennomeres 3–6 are slightly widened anteriorly, others transverse, parallel-sided.
|
||
<taxonomicName id="FCB74D38FFA6FFE71BE1FE32FDDD18D8" authorityName="Norman" authorityYear="1875" box="[433,512,426,452]" genus="Scapus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="6" pageNumber="184" phylum="Cnidaria" rank="genus">Scapus</taxonomicName>
|
||
enlarged, 1.5× as wide as long, 3× size of 2
|
||
<superScript id="CCC29BF3FFA6FFE71B23FE57FE5E18C1" attach="left" box="[371,387,463,477]" fontSize="6" pageId="6" pageNumber="180">nd</superScript>
|
||
antennomere, slightly longer than 3
|
||
<superScript id="CCC29BF3FFA6FFE71AC6FE6EFF791B18" attach="none" box="[150,164,502,516]" fontSize="6" pageId="6" pageNumber="180">rd</superScript>
|
||
, equal in size with 4
|
||
<superScript id="CCC29BF3FFA6FFE71BD5FE6EFE4F1B18" attach="none" box="[389,402,502,516]" fontSize="6" pageId="6" pageNumber="180">th</superScript>
|
||
, other antennal segments of equal size except that apical one slightly longer. Head with three pairs of protuberances (supraantennal protuberance, Р1; supraocular protuberance, Р2; posteromesal protuberance, Р3). Furrow visible in central part of the head dividing right and left protuberances; areas on sides of furrow forming ridges around eyes.
|
||
</paragraph>
|
||
<caption id="6FC86633FFA3FFE21AD8F81DFC751116" ID-DOI="http://doi.org/10.5281/zenodo.7929108" ID-Zenodo-Dep="7929108" httpUri="https://zenodo.org/record/7929108/files/figure.png" pageId="3" pageNumber="177" startId="3.[136,235,1925,1949]" targetBox="[155,1436,1205,1894]" targetPageId="3" targetType="figure">
|
||
<paragraph id="3B0836BBFFA3FFE21AD8F81DFC751116" blockId="3.[136,1452,1925,2058]" pageId="3" pageNumber="177">
|
||
<emphasis id="09C3EAA9FFA3FFE21AD8F81DFED91E82" bold="true" box="[136,260,1925,1950]" pageId="3" pageNumber="177">FIGURE 1.</emphasis>
|
||
<taxonomicName id="FCB74D38FFA3FFE21B5BF81EFE0D1E82" authorityName="Ponomarenko" authorityYear="1969" box="[267,464,1926,1950]" class="Insecta" family="Ommatidae" genus="Notocupes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="177" phylum="Arthropoda" rank="species" species="laticella">
|
||
<emphasis id="09C3EAA9FFA3FFE21B5BF81EFE0D1E82" box="[267,464,1926,1950]" italics="true" pageId="3" pageNumber="177">Notocupes laticella</emphasis>
|
||
</taxonomicName>
|
||
, holotype (PIN, 2555/1663).
|
||
<emphasis id="09C3EAA9FFA3FFE218ACF81DFCD31E81" bold="true" box="[764,782,1925,1949]" pageId="3" pageNumber="177">A</emphasis>
|
||
, Photo of part.
|
||
<emphasis id="09C3EAA9FFA3FFE219FDF81EFC631E82" bold="true" box="[941,958,1926,1950]" pageId="3" pageNumber="177">B</emphasis>
|
||
, Interpretative line drawing of part.
|
||
<emphasis id="09C3EAA9FFA3FFE21F61F81DFA9E1E81" bold="true" box="[1329,1347,1925,1949]" pageId="3" pageNumber="177">C</emphasis>
|
||
, Mirrored photo of counterpart wetted with 95% ethanol.
|
||
<emphasis id="09C3EAA9FFA3FFE2182AF832FD511EDE" bold="true" box="[634,652,1962,1986]" pageId="3" pageNumber="177">D</emphasis>
|
||
, Interpretative line drawing of mirrored counterpart. Abbreviations here and elsewhere: an.p, antennal pit; cl-lab, clypeolabrum; el, elevation; P1, supraantennal protuberance; pr.cx, procoxae; pr.re, procoxal rests. Pink filling indicates mandibles; light grey—head; dark grey—profemur.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="6FC86633FFA4FFE51AD8F88CFE851118" ID-DOI="http://doi.org/10.5281/zenodo.7929110" ID-Zenodo-Dep="7929110" httpUri="https://zenodo.org/record/7929110/files/figure.png" pageId="4" pageNumber="178" startId="4.[136,235,1812,1836]" targetBox="[273,1314,189,1788]" targetPageId="4" targetType="figure">
|
||
<paragraph id="3B0836BBFFA4FFE51AD8F88CFE851118" blockId="4.[136,1452,1812,2052]" pageId="4" pageNumber="178">
|
||
<emphasis id="09C3EAA9FFA4FFE51AD8F88CFED51E30" bold="true" box="[136,264,1812,1836]" pageId="4" pageNumber="178">FIGURE 2.</emphasis>
|
||
<taxonomicName id="FCB74D38FFA4FFE51B42F88DFE071E30" authorityName="Ponomarenko" authorityYear="1969" box="[274,474,1812,1836]" class="Insecta" family="Ommatidae" genus="Notocupes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="178" phylum="Arthropoda" rank="species" species="laticella">
|
||
<emphasis id="09C3EAA9FFA4FFE51B42F88DFE071E30" box="[274,474,1812,1836]" italics="true" pageId="4" pageNumber="178">Notocupes laticella</emphasis>
|
||
</taxonomicName>
|
||
, paratypes (PIN, 2240/71, 74).
|
||
<emphasis id="09C3EAA9FFA4FFE5197DF88CFCE21E30" bold="true" box="[813,831,1812,1836]" pageId="4" pageNumber="178">A</emphasis>
|
||
, Wetted with 95% ethanol PIN, 2240/74.
|
||
<emphasis id="09C3EAA9FFA4FFE51F52F88CFACE1E30" bold="true" box="[1282,1299,1812,1836]" pageId="4" pageNumber="178">B</emphasis>
|
||
, Interpretative line drawing (square here and beyond shows relative size and density of tubercles).
|
||
<emphasis id="09C3EAA9FFA4FFE519B3F8A0FC281E4C" bold="true" box="[995,1013,1848,1872]" pageId="4" pageNumber="178">С</emphasis>
|
||
, General scheme of elytra.
|
||
<emphasis id="09C3EAA9FFA4FFE51F41F8A0FAFE1E4C" bold="true" box="[1297,1315,1848,1872]" pageId="4" pageNumber="178">D</emphasis>
|
||
, Wetted with 95% ethanol PIN, 2240/71, counterpart.
|
||
<emphasis id="09C3EAA9FFA4FFE5187DF8C4FDE31E68" bold="true" box="[557,574,1884,1908]" pageId="4" pageNumber="178">E</emphasis>
|
||
, Interpretative line drawing.
|
||
<emphasis id="09C3EAA9FFA4FFE5193DF8C4FCA11E68" bold="true" box="[877,892,1884,1908]" pageId="4" pageNumber="178">F</emphasis>
|
||
, Interpretative line drawing of PIN, 2240/71, part.
|
||
<emphasis id="09C3EAA9FFA4FFE51FC1F8C4FA781E68" bold="true" box="[1425,1445,1884,1908]" pageId="4" pageNumber="178">G</emphasis>
|
||
, Photo of dry PIN, 2240/71, part. Abbreviations here and elsewhere: in.cx.p, intercoxal process; mesan, mesanepisternum; mesep, mesepimeron, mesotr, mesotrochantin; mesovent, mesoventrite; metanep, metanepimeron; metatr, metatrochantin; metavent, metaventrite; mnd, mandible; P2, supraocular protuberance; P3, posteromesal protuberance; s.n.pl, notopleural suture; s.pl.st, pleurosternal suture.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="6FC86633FFA5FFE41AD8F807FC101EE3" ID-DOI="http://doi.org/10.5281/zenodo.7929114" ID-Zenodo-Dep="7929114" httpUri="https://zenodo.org/record/7929114/files/figure.png" pageId="5" pageNumber="179" startId="5.[136,235,1951,1975]" targetBox="[207,1381,189,1923]" targetPageId="5" targetType="figure">
|
||
<paragraph id="3B0836BBFFA5FFE41AD8F807FC101EE3" blockId="5.[136,1451,1951,2047]" pageId="5" pageNumber="179">
|
||
<emphasis id="09C3EAA9FFA5FFE41AD8F807FED81EAB" bold="true" box="[136,261,1951,1975]" pageId="5" pageNumber="179">FIGURE 3.</emphasis>
|
||
<taxonomicName id="FCB74D38FFA5FFE41B5CF838FE0F1EAB" authorityName="Ponomarenko" authorityYear="1969" box="[268,466,1951,1975]" class="Insecta" family="Ommatidae" genus="Notocupes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="179" phylum="Arthropoda" rank="species" species="laticella">
|
||
<emphasis id="09C3EAA9FFA5FFE41B5CF838FE0F1EAB" box="[268,466,1951,1975]" italics="true" pageId="5" pageNumber="179">Notocupes laticella</emphasis>
|
||
</taxonomicName>
|
||
, paratypes.
|
||
<emphasis id="09C3EAA9FFA5FFE4181EF807FDA21EAB" bold="true" box="[590,639,1951,1975]" pageId="5" pageNumber="179">A–C</emphasis>
|
||
, PIN, 2555/1662. Wetted with 95% ethanol (
|
||
<emphasis id="09C3EAA9FFA5FFE41E00F807FBBF1EAB" bold="true" box="[1104,1122,1951,1975]" pageId="5" pageNumber="179">A</emphasis>
|
||
), dry (
|
||
<emphasis id="09C3EAA9FFA5FFE41EFAF807FB661EAB" bold="true" box="[1194,1211,1951,1975]" pageId="5" pageNumber="179">B</emphasis>
|
||
) and interpretative line drawing (
|
||
<emphasis id="09C3EAA9FFA5FFE41ABAF85BFF211EC7" bold="true" box="[234,252,1987,2011]" pageId="5" pageNumber="179">C</emphasis>
|
||
).
|
||
<emphasis id="09C3EAA9FFA5FFE41B41F85BFEFE1EC7" bold="true" box="[273,291,1987,2011]" pageId="5" pageNumber="179">D</emphasis>
|
||
, Photo of dry PIN, 2555/1686.
|
||
<emphasis id="09C3EAA9FFA5FFE41833F85BFD491EC7" bold="true" box="[611,660,1987,2011]" pageId="5" pageNumber="179">E–G</emphasis>
|
||
, PIN, 2240/6. Wetted with 95% ethanol (
|
||
<emphasis id="09C3EAA9FFA5FFE41E67F85BFB951EC7" bold="true" box="[1079,1096,1987,2011]" pageId="5" pageNumber="179">E</emphasis>
|
||
) and dry (
|
||
<emphasis id="09C3EAA9FFA5FFE41EE2F85BFB1C1EC7" bold="true" box="[1202,1217,1987,2011]" pageId="5" pageNumber="179">F</emphasis>
|
||
).
|
||
<emphasis id="09C3EAA9FFA5FFE41E86F85BFB371EC7" bold="true" box="[1238,1258,1987,2011]" pageId="5" pageNumber="179">G</emphasis>
|
||
, Interpretative line drawing. Abbreviations: pr.tr, protrochantin; pr.tr-er, protrochanter; pr.tib, protibia.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="3B0836BBFFA6FFE7197AFF07FBC019A9" blockId="6.[810,1053,159,181]" box="[810,1053,159,181]" pageId="6" pageNumber="180">
|
||
<tableNote id="C6513735FFA6FFE7197AFF07FBC019A9" box="[810,1053,159,181]" pageId="6" pageNumber="180" targetBox="[817,1431,195,494]" targetPageId="6">Measurements (in mm).</tableNote>
|
||
</paragraph>
|
||
<paragraph id="3B0836BBFFA6FFE71E12FF5BFAAB18F1" pageId="6" pageNumber="180">
|
||
<table id="49B7C41BFFA6001E1961FF5BFA4A18F2" box="[817,1431,195,494]" colsContinueIn="8.[817,1423,1245,1589]" gridcols="5" gridrows="4" pageId="6" pageNumber="180">
|
||
<tr id="858734F9FFA6001E1961FF5BFA4A19EF" box="[817,1431,195,243]" gridrow="0" pageId="6" pageNumber="180" rowspan-0="1">
|
||
<th id="C6565D85FFA6001E1E12FF5BFBAA19EF" box="[1090,1143,195,243]" gridcol="1" gridrow="0" pageId="6" pageNumber="180">body length</th>
|
||
<th id="C6565D85FFA6001E1ECFFF5BFB0D19EF" box="[1183,1232,195,243]" gridcol="2" gridrow="0" pageId="6" pageNumber="180">body width</th>
|
||
<th id="C6565D85FFA6001E1EABFF5BFAEA19EF" box="[1275,1335,195,243]" gridcol="3" gridrow="0" pageId="6" pageNumber="180">elytron length</th>
|
||
<th id="C6565D85FFA6001E1F0BFF5BFA4A19EF" box="[1371,1431,195,243]" gridcol="4" gridrow="0" pageId="6" pageNumber="180">elytron width</th>
|
||
</tr>
|
||
<tr id="858734F9FFA6001E1961FE88FA4A1838" box="[817,1431,272,292]" gridrow="1" pageId="6" pageNumber="180">
|
||
<th id="C6565D85FFA6001E1961FE88FBD41838" box="[817,1033,272,292]" gridcol="0" gridrow="1" pageId="6" pageNumber="180">Holotype PIN 2555/1663</th>
|
||
<td id="C6565D85FFA6001E1E12FE88FBAA1838" box="[1090,1143,272,292]" gridcol="1" gridrow="1" pageId="6" pageNumber="180">11</td>
|
||
<td id="C6565D85FFA6001E1ECFFE88FB0D1838" box="[1183,1232,272,292]" gridcol="2" gridrow="1" pageId="6" pageNumber="180">4.4</td>
|
||
<td id="C6565D85FFA6001E1EABFE88FAEA1838" box="[1275,1335,272,292]" gridcol="3" gridrow="1" pageId="6" pageNumber="180">7.7</td>
|
||
<td id="C6565D85FFA6001E1F0BFE88FA4A1838" box="[1371,1431,272,292]" gridcol="4" gridrow="1" pageId="6" pageNumber="180">2.2</td>
|
||
</tr>
|
||
<tr id="858734F9FFA6001E1961FEDCFA4A1861" box="[817,1431,324,381]" gridrow="2" pageId="6" pageNumber="180">
|
||
<th id="C6565D85FFA6001E1961FEDCFBD41861" box="[817,1033,324,381]" gridcol="0" gridrow="2" pageId="6" pageNumber="180">Paratype PIN 2240/6 Paratype PIN 2240/71</th>
|
||
<td id="C6565D85FFA6001E1E12FEDCFBAA1861" box="[1090,1143,324,381]" gridcol="1" gridrow="2" pageId="6" pageNumber="180">13.4 12.9</td>
|
||
<td id="C6565D85FFA6001E1ECFFEDCFB0D1861" box="[1183,1232,324,381]" gridcol="2" gridrow="2" pageId="6" pageNumber="180">4.4 3.7</td>
|
||
<td id="C6565D85FFA6001E1EABFEDCFAEA1861" box="[1275,1335,324,381]" gridcol="3" gridrow="2" pageId="6" pageNumber="180">9.8 9.3</td>
|
||
<td id="C6565D85FFA6001E1F0BFEDCFA4A1861" box="[1371,1431,324,381]" gridcol="4" gridrow="2" pageId="6" pageNumber="180">1.7 1.9</td>
|
||
</tr>
|
||
<tr id="858734F9FFA6001E1961FE17FA4A18F2" box="[817,1431,399,494]" gridrow="3" pageId="6" pageNumber="180">
|
||
<th id="C6565D85FFA6001E1961FE17FBD418F2" box="[817,1033,399,494]" gridcol="0" gridrow="3" pageId="6" pageNumber="180">Paratype PIN 2240/74 Paratype PIN 2555/1662 Paratype PIN 2555/1686</th>
|
||
<td id="C6565D85FFA6001E1E12FE17FBAA18F2" box="[1090,1143,399,494]" gridcol="1" gridrow="3" pageId="6" pageNumber="180">14.6* 10.7 11.1</td>
|
||
<td id="C6565D85FFA6001E1ECFFE17FB0D18F2" box="[1183,1232,399,494]" gridcol="2" gridrow="3" pageId="6" pageNumber="180">- - 3.6</td>
|
||
<td id="C6565D85FFA6001E1EABFE17FAEA18F2" box="[1275,1335,399,494]" gridcol="3" gridrow="3" pageId="6" pageNumber="180">- - 8.7</td>
|
||
<td id="C6565D85FFA6001E1F0BFE17FA4A18F2" box="[1371,1431,399,494]" gridcol="4" gridrow="3" pageId="6" pageNumber="180">- - 1.8</td>
|
||
</tr>
|
||
</table>
|
||
</paragraph>
|
||
<paragraph id="3B0836BBFFA6FFE7197AFE67FBB81B08" blockId="6.[810,1125,510,532]" box="[810,1125,510,532]" pageId="6" pageNumber="180">
|
||
<tableNote id="C6513735FFA6FFE7197AFE67FBB81B08" box="[810,1125,510,532]" pageId="6" pageNumber="180" targetBox="[817,1431,195,494]" targetPageId="6">Note: * incomplete: as preserved</tableNote>
|
||
</paragraph>
|
||
<paragraph id="3B0836BBFFA6FFE71AEDFC9AFDE01AC6" blockId="6.[136,778,158,2056]" pageId="6" pageNumber="180">
|
||
<emphasis id="09C3EAA9FFA6FFE71AEDFC9AFEF11A07" box="[189,300,770,795]" italics="true" pageId="6" pageNumber="180">Pronotum</emphasis>
|
||
transverse, almost not narrowing anteriorly. Pronotum 1.6× as wide as head; pronotal anterior margin not concave; anterior angles slightly protruding anteriorly. Posterior angles and posterior margin rounded; sides of pronotum straight. Dorsal aspect with pair of elongate-oval or bean-shaped medial elevations.
|
||
</paragraph>
|
||
<paragraph id="3B0836BBFFA6FFE71AEDFC7FFD511D50" blockId="6.[136,778,158,2056]" pageId="6" pageNumber="180">
|
||
<emphasis id="09C3EAA9FFA6FFE71AEDFC7FFE9D1D1C" box="[189,320,999,1024]" italics="true" pageId="6" pageNumber="180">Prosternum</emphasis>
|
||
. Notopleural suture straight, its apical third curved outward. Pleurosternal suture straight. Propleuron of equal width all along its length.
|
||
</paragraph>
|
||
<paragraph id="3B0836BBFFA6FFE71AEDFBC1FD3C1CEC" blockId="6.[136,778,158,2056]" pageId="6" pageNumber="180">
|
||
<emphasis id="09C3EAA9FFA6FFE71AEDFBC1FED61D6F" box="[189,267,1113,1139]" italics="true" pageId="6" pageNumber="180">Elytral</emphasis>
|
||
length to width ratio 3.4–3.8; elytron to pronotum width ratio 1.7. Humerus indistinct; basal 1/3 of epipleural margin straight; epipleuron to elytron width ratio at elytron mid-length 13–18. Main veins almost indistinguishable from intercalary ones. Veins 4 and 5 merge together before reaching elytral apex. Elytral cells large, rounded, occupying almost entire distance between the veins. Field I with full row of cells, cells in field II 23, field III 24, field IV 22–23, field V 18, field VI
|
||
<quantity id="FC4F9B5EFFA6FFE718F6FA12FCD71CB8" box="[678,778,1418,1444]" metricMagnitude="-1" metricUnit="m" metricValue="4.953" metricValueMax="5.334" metricValueMin="4.572" pageId="6" pageNumber="180" unit="in" value="19.5" valueMax="21.0" valueMin="18.0">18–21 in</quantity>
|
||
one row. Maculae present, cells near elytral base with 5–6 of them, cells at elytral mid-length with 7–8 maculae.
|
||
</paragraph>
|
||
<paragraph id="3B0836BBFFA6FFE71AEDFA65FD661F0B" blockId="6.[136,778,158,2056]" box="[189,699,1533,1559]" pageId="6" pageNumber="180">
|
||
<emphasis id="09C3EAA9FFA6FFE71AEDFA65FE921F0A" box="[189,335,1533,1558]" italics="true" pageId="6" pageNumber="180">Mesoventrite</emphasis>
|
||
slightly longer than mesocoxae.
|
||
</paragraph>
|
||
<paragraph id="3B0836BBFFA6FFE71AEDF9BCFDEE1FB3" blockId="6.[136,778,158,2056]" pageId="6" pageNumber="180">
|
||
<emphasis id="09C3EAA9FFA6FFE71AEDF9BCFE911F21" box="[189,332,1572,1597]" italics="true" pageId="6" pageNumber="180">Metaventrite</emphasis>
|
||
as long as wide, anterior margin to posterior margin width ratio 1.9; longitudinal suture distinct, paracoxal suture approximate to but not merging with posterior margin of metaventrite.
|
||
</paragraph>
|
||
<paragraph id="3B0836BBFFA6FFE71AEDF923FE62111B" blockId="6.[136,778,158,2056]" pageId="6" pageNumber="180">
|
||
<emphasis id="09C3EAA9FFA6FFE71AEDF923FEFA1FC9" box="[189,295,1723,1749]" italics="true" pageId="6" pageNumber="180">Abdomen</emphasis>
|
||
not narrowing forward, narrowing posteriorly at level of sternite III. Sternites I–IV with depression along their anterior margins. Sternites length ratio is 1.7–2:1:1:1:1.6–1.7.
|
||
<emphasis id="09C3EAA9FFA6FFE71B83F8B7FDE01E54" box="[467,573,1839,1864]" italics="true" pageId="6" pageNumber="180">Procoxae</emphasis>
|
||
adjacent, weakly transverse, large, making 1/3 of overall pronotal length; prosternal intercoxal process not fully separating procoxae (
|
||
<figureCitation id="A38C2A3EFFA6FFE71AC0F839FF391EA7" box="[144,228,1953,1979]" captionStart="FIGURE 2" captionStartId="4.[136,235,1812,1836]" captionTargetBox="[273,1314,189,1788]" captionTargetId="figure-17@4.[273,1314,189,1788]" captionTargetPageId="4" captionText="FIGURE 2. Notocupes laticella, paratypes (PIN, 2240/71, 74). A, Wetted with 95% ethanol PIN, 2240/74. B, Interpretative line drawing (square here and beyond shows relative size and density of tubercles). С, General scheme of elytra. D, Wetted with 95% ethanol PIN, 2240/71, counterpart. E, Interpretative line drawing. F, Interpretative line drawing of PIN, 2240/71, part. G, Photo of dry PIN, 2240/71, part. Abbreviations here and elsewhere: in.cx.p, intercoxal process; mesan, mesanepisternum; mesep, mesepimeron, mesotr, mesotrochantin; mesovent, mesoventrite; metanep, metanepimeron; metatr, metatrochantin; metavent, metaventrite; mnd, mandible; P2, supraocular protuberance; P3, posteromesal protuberance; s.n.pl, notopleural suture; s.pl.st, pleurosternal suture." figureDoi="http://doi.org/10.5281/zenodo.7929110" httpUri="https://zenodo.org/record/7929110/files/figure.png" pageId="6" pageNumber="180">Fig. 2B</figureCitation>
|
||
, in.cx.p). First two tarsomeres of fore legs of same length, other 3 shorter. First visible abdominal sternite to metacoxae length ratio 2–3.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="73AD6530FFA6FFE6190FFDDBFF7618A9" lastPageId="7" lastPageNumber="181" pageId="6" pageNumber="180" type="discussion">
|
||
<paragraph id="3B0836BBFFA6FFE7190FFDDBFBEA1C2C" blockId="6.[810,1452,579,2051]" pageId="6" pageNumber="180">
|
||
<emphasis id="09C3EAA9FFA6FFE7190FFDDBFC0E1B41" bold="true" box="[863,979,579,605]" pageId="6" pageNumber="180">Remarks.</emphasis>
|
||
Similar to
|
||
<taxonomicName id="FCB74D38FFA6FFE71E25FDDCFB381B41" authorityName="Ponomarenko" authorityYear="1969" box="[1141,1253,580,605]" class="Insecta" family="Ommatidae" genus="Notocupes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="180" phylum="Arthropoda" rank="species" species="tenuis">
|
||
<emphasis id="09C3EAA9FFA6FFE71E25FDDCFB381B41" box="[1141,1253,580,605]" italics="true" pageId="6" pageNumber="180">N. tenuis</emphasis>
|
||
</taxonomicName>
|
||
in head relief, procoxae shifted closer to pronotal mid-length and main veins almost indistinguishable from intercalary ones, but differs in bigger elytral cells and pronotum not narrowing in anterior half. Similar to
|
||
<taxonomicName id="FCB74D38FFA6FFE71E32FD4CFB151BF1" authorityName="Ponomarenko" authorityYear="1969" box="[1122,1224,724,750]" class="Insecta" family="Ommatidae" genus="Notocupes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="180" phylum="Arthropoda" rank="species" species="tenuis">
|
||
<emphasis id="09C3EAA9FFA6FFE71E32FD4CFBA81BF1" box="[1122,1141,724,749]" italics="true" pageId="6" pageNumber="180">N</emphasis>
|
||
.
|
||
<emphasis id="09C3EAA9FFA6FFE71ED5FD4CFB151BF1" box="[1157,1224,724,749]" italics="true" pageId="6" pageNumber="180">tenuis</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="FCB74D38FFA6FFE71F54FD4CFA511BF1" authorityName="Ponomarenko" authorityYear="1969" box="[1284,1420,724,750]" class="Insecta" family="Ommatidae" genus="Notocupes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="180" phylum="Arthropoda" rank="species" species="rostratus">
|
||
<emphasis id="09C3EAA9FFA6FFE71F54FD4CFACA1BF1" box="[1284,1303,724,749]" italics="true" pageId="6" pageNumber="180">N</emphasis>
|
||
.
|
||
<emphasis id="09C3EAA9FFA6FFE71F78FD4CFA511BF1" box="[1320,1420,724,749]" italics="true" pageId="6" pageNumber="180">rostratus</emphasis>
|
||
</taxonomicName>
|
||
in antennal bases covered by supraantennal protuberance. Similar to
|
||
<taxonomicName id="FCB74D38FFA6FFE719FDFC84FBCA1A29" authorityName="Ponomarenko" authorityYear="1969" box="[941,1047,796,822]" class="Insecta" family="Ommatidae" genus="Notocupes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="180" phylum="Arthropoda" rank="species" species="tenuis">
|
||
<emphasis id="09C3EAA9FFA6FFE719FDFC84FC1D1A29" box="[941,960,796,821]" italics="true" pageId="6" pageNumber="180">N</emphasis>
|
||
.
|
||
<emphasis id="09C3EAA9FFA6FFE71984FC84FBCA1A29" box="[980,1047,796,821]" italics="true" pageId="6" pageNumber="180">tenuis</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="FCB74D38FFA6FFE71E7BFC84FB6B1A29" authorityName="Ponomarenko" authorityYear="1969" box="[1067,1206,796,822]" class="Insecta" family="Ommatidae" genus="Notocupes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="180" phylum="Arthropoda" rank="species" species="rostratus">
|
||
<emphasis id="09C3EAA9FFA6FFE71E7BFC84FBE31A29" box="[1067,1086,796,821]" italics="true" pageId="6" pageNumber="180">N</emphasis>
|
||
.
|
||
<emphasis id="09C3EAA9FFA6FFE71E02FC84FB6B1A29" box="[1106,1206,796,821]" italics="true" pageId="6" pageNumber="180">rostratus</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="FCB74D38FFA6FFE71EA9FC84FA551A29" authorityName="Ponomarenko" authorityYear="1969" box="[1273,1416,796,822]" class="Insecta" family="Ommatidae" genus="Notocupes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="180" phylum="Arthropoda" rank="species" species="oxypygus">
|
||
<emphasis id="09C3EAA9FFA6FFE71EA9FC84FAD11A29" box="[1273,1292,796,821]" italics="true" pageId="6" pageNumber="180">N</emphasis>
|
||
.
|
||
<emphasis id="09C3EAA9FFA6FFE71F70FC84FA551A29" box="[1312,1416,796,821]" italics="true" pageId="6" pageNumber="180">oxypygus</emphasis>
|
||
</taxonomicName>
|
||
in having dense, uniform cuticular tubercles. In having large elytral cells similar to
|
||
<taxonomicName id="FCB74D38FFA6FFE71E13FCFDFA7A1A63" authority="(Giebel, 1856)" baseAuthorityName="Giebel" baseAuthorityYear="1856" box="[1091,1447,868,895]" class="Insecta" family="Ommatidae" genus="Zygadenia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="180" phylum="Arthropoda" rank="species" species="tuberculata">
|
||
<emphasis id="09C3EAA9FFA6FFE71E13FCFDFB8B1A62" box="[1091,1110,869,894]" italics="true" pageId="6" pageNumber="180">N</emphasis>
|
||
.
|
||
<emphasis id="09C3EAA9FFA6FFE71E3BFCFCFB361A62" box="[1131,1259,868,894]" italics="true" pageId="6" pageNumber="180">tuberculata</emphasis>
|
||
(
|
||
<bibRefCitation id="5F264B4AFFA6FFE71F51FCFCFA421A63" author="Giebel, C. G. A." box="[1281,1439,868,895]" pageId="6" pageNumber="180" refId="ref9196" refString="Giebel, C. G. A. (1856) Fauna der Vorwelt mit steter Berucksichtigung der lebenden Thiere, Bd. 2, Abth. I. Die Insecten und Spinnen der Vorwelt, mit steter Berucksichtigung der lebenden Insecten und Spinnen, monographisch dargestellt. Brockhaus, Leipzig, xviii + 512 pp." type="book" year="1856">Giebel, 1856</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="FCB74D38FFA6FFE7197AFC11FB6A1ABE" authority="Ponomarenko, 1968" authorityName="Ponomarenko" authorityYear="1968" box="[810,1207,904,931]" class="Insecta" family="Ommatidae" genus="Notocupes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="180" phylum="Arthropoda" rank="species" species="foersteri">
|
||
<emphasis id="09C3EAA9FFA6FFE7197AFC11FCE01ABE" box="[810,829,905,930]" italics="true" pageId="6" pageNumber="180">N</emphasis>
|
||
.
|
||
<emphasis id="09C3EAA9FFA6FFE7191AFC10FC741ABE" box="[842,937,904,930]" italics="true" pageId="6" pageNumber="180">foersteri</emphasis>
|
||
<bibRefCitation id="5F264B4AFFA6FFE71991FC10FB6A1ABE" author="Ponomarenko, A. G." box="[961,1207,904,930]" pageId="6" pageNumber="180" pagination="118 - 138" refId="ref10163" refString="Ponomarenko, A. G. (1968) Archostemata beetles from the Jurassic of the Kara-Tau. In: Rohdendorf, B. B. (Ed.), Jurassic insects of the Kara-Tau. Nauka, Moscow, pp. 118 - 138. [In Russian]" type="book chapter" year="1968">Ponomarenko, 1968</bibRefCitation>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="FCB74D38FFA6FFE71E83FC11FBCC1ADA" authority="Ponomarenko, 1968" authorityName="Ponomarenko" authorityYear="1968" class="Insecta" family="Ommatidae" genus="Notocupes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="180" phylum="Arthropoda" rank="species" species="nigrimonticola">
|
||
<emphasis id="09C3EAA9FFA6FFE71E83FC11FB3B1ABE" box="[1235,1254,905,930]" italics="true" pageId="6" pageNumber="180">N</emphasis>
|
||
.
|
||
<emphasis id="09C3EAA9FFA6FFE71F55FC10FA761ABE" box="[1285,1451,904,930]" italics="true" pageId="6" pageNumber="180">nigrimonticola</emphasis>
|
||
<bibRefCitation id="5F264B4AFFA6FFE7197AFC34FBCC1ADA" author="Ponomarenko, A. G." box="[810,1041,940,966]" pageId="6" pageNumber="180" pagination="118 - 138" refId="ref10163" refString="Ponomarenko, A. G. (1968) Archostemata beetles from the Jurassic of the Kara-Tau. In: Rohdendorf, B. B. (Ed.), Jurassic insects of the Kara-Tau. Nauka, Moscow, pp. 118 - 138. [In Russian]" type="book chapter" year="1968">Ponomarenko, 1968</bibRefCitation>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="FCB74D38FFA6FFE71E02FC35FA7A1ADA" authority="Tan et al., 2006" authorityName="Tan" authorityYear="2006" box="[1106,1447,940,967]" class="Insecta" family="Ommatidae" genus="Notocupes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="180" phylum="Arthropoda" rank="species" species="protensa">
|
||
<emphasis id="09C3EAA9FFA6FFE71E02FC35FBB81ADA" box="[1106,1125,941,966]" italics="true" pageId="6" pageNumber="180">N</emphasis>
|
||
.
|
||
<emphasis id="09C3EAA9FFA6FFE71E28FC35FB041ADA" box="[1144,1241,941,966]" italics="true" pageId="6" pageNumber="180">protensa</emphasis>
|
||
Tan
|
||
<emphasis id="09C3EAA9FFA6FFE71F4BFC35FA8F1ADA" box="[1307,1362,940,966]" italics="true" pageId="6" pageNumber="180">et al</emphasis>
|
||
., 2006
|
||
</taxonomicName>
|
||
, but differs in main veins almost indistinguishable from intercalary ones. It differs from
|
||
<taxonomicName id="FCB74D38FFA6FFE71ED8FC6DFAFB1D12" box="[1160,1318,1012,1039]" class="Insecta" family="Ommatidae" genus="Notocupes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="180" phylum="Arthropoda" rank="species" species="tuberculate">
|
||
<emphasis id="09C3EAA9FFA6FFE71ED8FC6DFB461D12" box="[1160,1179,1013,1038]" italics="true" pageId="6" pageNumber="180">N</emphasis>
|
||
.
|
||
<emphasis id="09C3EAA9FFA6FFE71EF7FC6CFAFB1D12" box="[1191,1318,1012,1038]" italics="true" pageId="6" pageNumber="180">tuberculate</emphasis>
|
||
</taxonomicName>
|
||
in narrower epipleuron and smaller size, from
|
||
<taxonomicName id="FCB74D38FFA6FFE71EEFFB81FA571D2F" authorityName="Ponomarenko" authorityYear="1968" box="[1215,1418,1049,1075]" class="Insecta" family="Ommatidae" genus="Notocupes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="180" phylum="Arthropoda" rank="species" species="nigrimonticola">
|
||
<emphasis id="09C3EAA9FFA6FFE71EEFFB81FB0F1D2E" box="[1215,1234,1049,1074]" italics="true" pageId="6" pageNumber="180">N</emphasis>
|
||
.
|
||
<emphasis id="09C3EAA9FFA6FFE71EB4FB81FA571D2F" box="[1252,1418,1049,1075]" italics="true" pageId="6" pageNumber="180">nigrimonticola</emphasis>
|
||
</taxonomicName>
|
||
in uniform cuticular tubercles, not serrated lateral margin of pronotum, narrower and straighter epipleuron with row of cells, rounded elytral cells and smaller size; from
|
||
<taxonomicName id="FCB74D38FFA6FFE7197AFB32FC6C1DDF" box="[810,945,1194,1219]" class="Insecta" family="Ommatidae" genus="Notocupes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="180" phylum="Arthropoda" rank="species" species="protensa">
|
||
<emphasis id="09C3EAA9FFA6FFE7197AFB32FC6C1DDF" box="[810,945,1194,1219]" italics="true" pageId="6" pageNumber="180">N. protensa</emphasis>
|
||
</taxonomicName>
|
||
in eyes close to mandibular bases, lateral margin of pronotum not serrated, epipleuron narrower and straighter, and last sternite more tapering and sharper and with more maculae.
|
||
</paragraph>
|
||
<paragraph id="3B0836BBFFA6FFE7190FFAA2FC351CF8" blockId="6.[810,1452,579,2051]" pageId="6" pageNumber="180">Differs from the congeners in having antennae clearly reaching behind pronotal posterior margin and pronotum almost rectangular and not narrowing anteriorly. The long antenna is unusual for ommatids which usually have antennae shorter.</paragraph>
|
||
<paragraph id="3B0836BBFFA6FFE7190FFA77FC5D1EA6" blockId="6.[810,1452,579,2051]" pageId="6" pageNumber="180">
|
||
In having procoxae located closer to the center of pronotum instead of its posterior margin, make
|
||
<taxonomicName id="FCB74D38FFA6FFE7197AF9A0FC7C1F4D" authorityName="Ponomarenko" authorityYear="1969" box="[810,929,1591,1617]" class="Insecta" family="Ommatidae" genus="Notocupes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="180" phylum="Arthropoda" rank="species" species="laticella">
|
||
<emphasis id="09C3EAA9FFA6FFE7197AF9A0FC7C1F4D" box="[810,929,1591,1617]" italics="true" pageId="6" pageNumber="180">N. laticella</emphasis>
|
||
</taxonomicName>
|
||
similar to representatives of
|
||
<taxonomicName id="FCB74D38FFA6FFE71E8CF9AFFA751F4D" box="[1244,1448,1591,1617]" class="Insecta" family="Tshekardocoleidae" kingdom="Animalia" order="Protocoleoptera" pageId="6" pageNumber="180" phylum="Arthropoda" rank="family">Tshekardocoleidae</taxonomicName>
|
||
,
|
||
<taxonomicName id="FCB74D38FFA6FFE7197AF9C3FC3D1F69" authorityName="Rohdendorf" authorityYear="1961" box="[810,992,1627,1653]" class="Insecta" family="Rhombocoleidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="180" phylum="Arthropoda" rank="family">Rhombocoleidae</taxonomicName>
|
||
and
|
||
<taxonomicName id="FCB74D38FFA6FFE71E42F9C3FB141F69" box="[1042,1225,1627,1653]" class="Insecta" family="Permocupedidae" kingdom="Animalia" order="Protocoleoptera" pageId="6" pageNumber="180" phylum="Arthropoda" rank="family">Permocupedidae</taxonomicName>
|
||
. However, elongated head, mandible three-toothed with a vertical arrangement of teeth, prothoracic intercoxal process incomplete and not reaching behind the coxae, double elytral cell rows with almost longitudinally-oriented veins and the veins 3 and 4 fused before reaching elytral apex suggest placement in
|
||
<taxonomicName id="FCB74D38FFA6FFE7197AF8ACFC6D1E52" authorityName="Sharp & Muir" authorityYear="1912" box="[810,944,1844,1870]" class="Insecta" family="Ommatidae" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="180" phylum="Arthropoda" rank="family">Ommatidae</taxonomicName>
|
||
. It seems that only this species of
|
||
<taxonomicName id="FCB74D38FFA6FFE71F65F8ADFA761E52" box="[1333,1451,1845,1870]" class="Insecta" family="Ommatidae" genus="Notocupes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="180" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="09C3EAA9FFA6FFE71F65F8ADFA761E52" box="[1333,1451,1845,1870]" italics="true" pageId="6" pageNumber="180">Notocupes</emphasis>
|
||
</taxonomicName>
|
||
has the procoxae lying closest to the procoxal center: Even
|
||
<taxonomicName id="FCB74D38FFA6FFE7197AF8E5FC4F1E8A" authorityName="Ponomarenko" authorityYear="1969" box="[810,914,1917,1943]" class="Insecta" family="Ommatidae" genus="Notocupes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="180" phylum="Arthropoda" rank="species" species="tenuis">
|
||
<emphasis id="09C3EAA9FFA6FFE7197AF8E5FCE01E8A" box="[810,829,1917,1942]" italics="true" pageId="6" pageNumber="180">N</emphasis>
|
||
.
|
||
<emphasis id="09C3EAA9FFA6FFE7191FF8E5FC4F1E8A" box="[847,914,1917,1942]" italics="true" pageId="6" pageNumber="180">tenuis</emphasis>
|
||
</taxonomicName>
|
||
has the procoxae slightly shifted to posterior margin.
|
||
</paragraph>
|
||
<paragraph id="3B0836BBFFA6FFE6190FF85DFF7618A9" blockId="6.[810,1452,579,2051]" lastBlockId="7.[136,778,158,437]" lastPageId="7" lastPageNumber="181" pageId="6" pageNumber="180">
|
||
The pronotal intercoxal process was originally mentioned in regard of material from
|
||
<collectingCountry id="43A0762BFFA6FFE71F4CF871FA82111F" box="[1308,1375,2025,2051]" name="China" pageId="6" pageNumber="180">China</collectingCountry>
|
||
(
|
||
<emphasis id="09C3EAA9FFA6FFE71F2CF872FA55111F" box="[1404,1416,2026,2051]" italics="true" pageId="6" pageNumber="180">e</emphasis>
|
||
.
|
||
<emphasis id="09C3EAA9FFA6FFE71FDFF872FA40111F" box="[1423,1437,2026,2051]" italics="true" pageId="6" pageNumber="180">g</emphasis>
|
||
.,
|
||
<taxonomicName id="FCB74D38FFA7FFE61AD8FF07FDC219A4" authority="(Tan & Ren, 2006))" authorityName="Tan & Ren" authorityYear="2006" baseAuthorityName="Tan & Ren" baseAuthorityYear="2006" box="[136,543,158,185]" class="Insecta" family="Ommatidae" genus="Ovatocupes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="181" phylum="Arthropoda" rank="species" species="alienus">
|
||
<emphasis id="09C3EAA9FFA7FFE61AD8FF07FF4619A4" box="[136,155,159,184]" italics="true" pageId="7" pageNumber="181">N</emphasis>
|
||
.
|
||
<emphasis id="09C3EAA9FFA7FFE61AE6FF06FEDA19A4" box="[182,263,158,184]" italics="true" pageId="7" pageNumber="181">alienus</emphasis>
|
||
(Tan &
|
||
<bibRefCitation id="5F264B4AFFA7FFE61BDCFF07FDD019A4" author="Ren, D. & Tan, J." box="[396,525,158,185]" pageId="7" pageNumber="181" pagination="457 - 463" refId="ref10525" refString="Ren, D. & Tan, J. (2006) A new cupedid genus (Coleoptera: Archostemata: Cupedidae) from Jehol biota of Western Liaoning, China. Annales Zoologici, 56 (3), 457 - 463." type="journal article" year="2006">Ren, 2006</bibRefCitation>
|
||
))
|
||
</taxonomicName>
|
||
and
|
||
<collectingCountry id="43A0762BFFA7FFE61820FF06FCD419A5" box="[624,777,158,185]" name="South Korea" pageId="7" pageNumber="181">South Korea</collectingCountry>
|
||
(
|
||
<taxonomicName id="FCB74D38FFA7FFE61AC1FF5BFE3D19C0" authority="Lee et al., 2022" authorityName="Lee" authorityYear="2022" box="[145,480,194,221]" class="Insecta" family="Ommatidae" genus="Notocupes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="181" phylum="Arthropoda" rank="species" species="premeris">
|
||
<emphasis id="09C3EAA9FFA7FFE61AC1FF5BFF7919C0" box="[145,164,195,220]" italics="true" pageId="7" pageNumber="181">N</emphasis>
|
||
.
|
||
<emphasis id="09C3EAA9FFA7FFE61AE5FF5BFEC519C0" box="[181,280,195,220]" italics="true" pageId="7" pageNumber="181">premeris</emphasis>
|
||
<bibRefCitation id="5F264B4AFFA7FFE61B72FF5BFE3D19C0" author="Lee, S. B. & Nam, G. S. & Li, Y. D." box="[290,480,194,221]" pageId="7" pageNumber="181" refId="ref9899" refString="Lee, S. B., Nam, G. S. & Li, Y. D. (2022) A new species of Notocupes (Coleoptera: Archostemata) from the Lower Cretaceous (Albian) Jinju Formation in South Korea. Cretaceous Research, 105357. https: // doi. org / 10.1016 / j. cretres. 2022.105357" type="book" year="2022">
|
||
Lee
|
||
<emphasis id="09C3EAA9FFA7FFE61B07FF5BFE5619C0" box="[343,395,194,220]" italics="true" pageId="7" pageNumber="181">et al</emphasis>
|
||
., 2022
|
||
</bibRefCitation>
|
||
</taxonomicName>
|
||
). Among
|
||
<taxonomicName id="FCB74D38FFA7FFE61803FF5BFD1419C0" box="[595,713,195,220]" class="Insecta" family="Ommatidae" genus="Notocupes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="181" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="09C3EAA9FFA7FFE61803FF5BFD1419C0" box="[595,713,195,220]" italics="true" pageId="7" pageNumber="181">Notocupes</emphasis>
|
||
</taxonomicName>
|
||
from the PIN RAS collection, this is the only specimen that enables observation of this character. The aforementioned species were depicted with intercoxal process dividing the procoxae, but our observations suggest contiguous procoxae underneath a short, triangular process (
|
||
<figureCitation id="A38C2A3EFFA7FFE618FCFEEFFD33188D" box="[684,750,375,401]" captionStart="FIGURE 2" captionStartId="4.[136,235,1812,1836]" captionTargetBox="[273,1314,189,1788]" captionTargetId="figure-17@4.[273,1314,189,1788]" captionTargetPageId="4" captionText="FIGURE 2. Notocupes laticella, paratypes (PIN, 2240/71, 74). A, Wetted with 95% ethanol PIN, 2240/74. B, Interpretative line drawing (square here and beyond shows relative size and density of tubercles). С, General scheme of elytra. D, Wetted with 95% ethanol PIN, 2240/71, counterpart. E, Interpretative line drawing. F, Interpretative line drawing of PIN, 2240/71, part. G, Photo of dry PIN, 2240/71, part. Abbreviations here and elsewhere: in.cx.p, intercoxal process; mesan, mesanepisternum; mesep, mesepimeron, mesotr, mesotrochantin; mesovent, mesoventrite; metanep, metanepimeron; metatr, metatrochantin; metavent, metaventrite; mnd, mandible; P2, supraocular protuberance; P3, posteromesal protuberance; s.n.pl, notopleural suture; s.pl.st, pleurosternal suture." figureDoi="http://doi.org/10.5281/zenodo.7929110" httpUri="https://zenodo.org/record/7929110/files/figure.png" pageId="7" pageNumber="181">Fig. 2</figureCitation>
|
||
А, B).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |