plazi/treatments-xml
2
0
Fork 0
Code Issues Pull requests Releases Activity
treatments-xml/data/91/6C/C5/916CC55C9569FFA44009B92EFB07B18D.xml
maintenance 16a7dde1ee added 91
2024-06-21 12:44:05 +02:00

291 lines
37 KiB
XML
Raw Blame History

This file contains ambiguous Unicode characters

This file contains Unicode characters that might be confused with other characters. If you think that this is intentional, you can safely ignore this warning. Use the Escape button to reveal them.

<document id="A3FDF75BF704F689B24BA9C524D29E9E" ID-DOI="10.5281/zenodo.5394764" ID-ISSN="1638-9387" ID-Zenodo-Dep="5394764" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1630323686270" checkinUser="marcus" docAuthor="Lourenço, Wilson R." docDate="2004" docId="916CC55C9569FFA44009B92EFB07B18D" docLanguage="en" docName="Zoosystema.2004.26.3.385-392.pdf" docOrigin="Zoosystema 26 (3)" docStyle="DocumentStyle:2A9C6F9EA5F90D98170D9E0D9CA29375.4:Zoosystema.2000-2008.journal_article.type1" docStyleId="2A9C6F9EA5F90D98170D9E0D9CA29375" docStyleName="Zoosystema.2000-2008.journal_article.type1" docStyleVersion="4" docTitle="Tityobuthus pallidus Lourenço 2004, n. sp." docType="treatment" docVersion="3" lastPageNumber="391" masterDocId="6D55BD24956AFFA24317BA03FF98B358" masterDocTitle="Further considerations regarding Tityobuthus baroni (Pocock, 1890) with the description of a new species from Ste Marie Island, Madagascar (Scorpiones, Buthidae)" masterLastPageNumber="392" masterPageNumber="385" pageNumber="388" updateTime="1699227349855" updateUser="plazi" zenodo-license-document="CC0-1.0" zenodo-license-figures="CC0-1.0" zenodo-license-treatments="UNSPECIFIED">
<mods:mods id="D1A53C575C01FA46DB6D48CE98A273FE" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="3E473C11F3D08F7A9CD188748B372E05">
<mods:title id="CF4CB332B79B64E21151B014B7E13138">Further considerations regarding Tityobuthus baroni (Pocock, 1890) with the description of a new species from Ste Marie Island, Madagascar (Scorpiones, Buthidae)</mods:title>
</mods:titleInfo>
<mods:name id="E66A365D887453ECBF3AD26C2CDAE90D" type="personal">
<mods:role id="D320A19CB34C9595CA27E6B16C861217">
<mods:roleTerm id="6ED5161174CC454389580746C2995C9F">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="7BE3A60E844160501CEF9DECF36A96F8">Lourenço, Wilson R.</mods:namePart>
<mods:affiliation id="660AF8EFBCD2DC30C6ABC24E3FDD6E20">Département Systématique et Évolution, Section Arthropodes (Arachnologie), d’Histoire naturelle, 61 rue Buffon, F- 75231 Paris cedex 05 (France) arachne @ mnhn. fr</mods:affiliation>
<mods:nameIdentifier id="9112414DD4A2004FFD10F529D48E5A5E" type="email">arachne@mnhn.fr</mods:nameIdentifier>
</mods:name>
<mods:typeOfResource id="789BD95FB1A9EC286A0EAF941F3C7D0F">text</mods:typeOfResource>
<mods:relatedItem id="825ABAAE545715254899F319319E0BC4" type="host">
<mods:titleInfo id="8A4DCF71805B934CDEC724904D146455">
<mods:title id="DEA2CD22440E46C724610111EEED0CAB">Zoosystema</mods:title>
</mods:titleInfo>
<mods:part id="9755E13B3F6A3BCDFFFD6958392DB047">
<mods:date id="F3C5FA1C138B3582E711156A6003A4D8">2004</mods:date>
<mods:detail id="AEC6DF87B41BF3BC1792FF6DD8699E62" type="volume">
<mods:number id="07EE9F486C349AE7140E79F1609D02C5">26</mods:number>
</mods:detail>
<mods:detail id="971D0BD74889A053855D731A9F64BBB2" type="issue">
<mods:number id="DD912C1E40EBC0B327485B7703437D49">3</mods:number>
</mods:detail>
<mods:extent id="EDED84B8EB7ED1ADEFCF846BBF2E36FA" unit="page">
<mods:start id="7313143ABC24B29C32EC8F6F3E52CB6F">385</mods:start>
<mods:end id="D3B7B571D3093BFC1EF5B4AE2CF63B80">392</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification id="2F0A5B992540D0208FFD3C9B40992789">journal article</mods:classification>
<mods:identifier id="74DF46015AE31B25E2E185973DC9E99C" type="DOI">10.5281/zenodo.5394764</mods:identifier>
<mods:identifier id="EA2BD99C1630A34022CF53D4566B926D" type="ISSN">1638-9387</mods:identifier>
<mods:identifier id="00CB4870778A05DDA67CF6EBDF49C1EF" type="Zenodo-Dep">5394764</mods:identifier>
</mods:mods>
<treatment id="916CC55C9569FFA44009B92EFB07B18D" LSID="urn:lsid:plazi:treatment:916CC55C9569FFA44009B92EFB07B18D" httpUri="http://treatment.plazi.org/id/916CC55C9569FFA44009B92EFB07B18D" lastPageId="6" lastPageNumber="391" pageId="3" pageNumber="388">
<subSubSection id="51DF27C19569FFA14009B92EFBD8B01E" box="[798,1088,812,839]" pageId="3" pageNumber="388" type="nomenclature">
<paragraph id="197A744A9569FFA14009B92EFBD8B01E" blockId="3.[798,1088,812,871]" box="[798,1088,812,839]" pageId="3" pageNumber="388">
<heading id="4232C3269569FFA14009B92EFBD8B01E" box="[798,1088,812,839]" centered="true" fontSize="11" level="2" pageId="3" pageNumber="388" reason="2">
<taxonomicName id="DEC50FC99569FFA14009B92EFC66B01F" ID-CoL="57978" authority="Lourenço, 2004" authorityName="Lourenço" authorityYear="2004" box="[798,1022,813,839]" class="Arachnida" family="Buthidae" genus="Tityobuthus" kingdom="Animalia" order="Scorpiones" pageId="3" pageNumber="388" phylum="Arthropoda" rank="species" species="pallidus" status="sp. nov.">
<emphasis id="2BB1A8589569FFA14009B92EFC66B01F" bold="true" box="[798,1022,813,839]" italics="true" pageId="3" pageNumber="388">Tityobuthus pallidus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="308215239569FFA14712B92FFBD8B01E" box="[1029,1088,812,838]" pageId="3" pageNumber="388" rank="species">n. sp.</taxonomicNameLabel>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="51DF27C19569FFA14027B94EFBB5B03F" box="[816,1069,845,871]" pageId="3" pageNumber="388" type="description">
<paragraph id="197A744A9569FFA14027B94EFBB5B03F" blockId="3.[798,1088,812,871]" box="[816,1069,845,871]" pageId="3" pageNumber="388">
(
<figureCitation id="81FE68CF9569FFA14020B94EFCE4B03F" box="[823,892,845,871]" captionStart="FIG" captionStartId="3.[113,124,1644,1661]" captionTargetBox="[126,648,256,1541]" captionTargetId="figure-335@3.[102,676,238,1563]" captionTargetPageId="3" captionText="FIG. 2. — Tityobuthus pallidus n. sp., holotype, trichobothrial pattern; A, tibia, dorso-external aspect; B, patella, dorsal aspect; C, femur, dorsal aspect. Scale bar: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5394768" httpUri="https://zenodo.org/record/5394768/files/figure.png" pageId="3" pageNumber="388">Figs 2</figureCitation>
;
<figureCitation id="81FE68CF9569FFA1409FB94EFC32B03F" box="[904,938,845,871]" captionStart="FIG" captionStartId="4.[113,124,1665,1682]" captionTargetBox="[247,1089,239,1622]" captionTargetId="figure-25@4.[156,1166,218,1716]" captionTargetPageId="4" captionText="FIG. 3. — Metasomal segment V and telson, lateral aspect; A, Tityobuthus pallidus n. sp., holotype; B, T. guillaumeti Lourenço, 1985,; C, T. baroni (Pocock, 1890),; D, T. lucileae Lourenço, 1996,. Scale bars: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5394770" httpUri="https://zenodo.org/record/5394770/files/figure.png" pageId="3" pageNumber="388">3A</figureCitation>
;
<figureCitation id="81FE68CF9569FFA140A2B94EFC5DB03F" box="[949,965,845,871]" captionStart="FIG" captionStartId="5.[113,124,948,965]" captionTargetBox="[113,1210,204,992]" captionTargetId="figure-281@5.[117,1206,204,992]" captionTargetPageId="5" captionText="FIG. 4. — Maps of Madagascar and Ste Marie Island, showing the known distribution of Tityobuthus baroni (Pocock, 1890) (1), T. lucileae Lourenço, 1996 (2), T. guillaumeti Lourenço, 1985 (3) and T. pallidus n. sp. (4)." figureDoi="http://doi.org/10.5281/zenodo.5394772" httpUri="https://zenodo.org/record/5394772/files/figure.png" pageId="3" pageNumber="388">4</figureCitation>
;
<tableCitation id="544741F19569FFA140C6B94EFBBAB03F" box="[977,1058,845,871]" captionStart="TABLE" captionStartId="6.[676,687,203,220]" captionText="TABLE 1. — Morphometric values (in mm) of the female holotype" pageId="3" pageNumber="388">Table 1</tableCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="51DF27C19569FFA141B3B996FB94B0A1" pageId="3" pageNumber="388" type="materials_examined">
<paragraph id="197A744A9569FFA141B3B996FB94B0A1" blockId="3.[676,1210,917,1504]" pageId="3" pageNumber="388">
<materialsCitation id="A9AD7E179569FFA141B3B996FB94B0A1" collectingDate="1955-12-27" collectionCode="TYPE, MATERIAL, IRSM" country="Madagascar" latitude="-16.85" location="Ste Marie Island" longLatPrecision="1280" longitude="49.95" municipality="Anbohidena Forest Station" pageId="3" pageNumber="388" specimenCount="1" typeStatus="holotype">
<collectionCode id="7FD4EC8F9569FFA141B3B996FCA3B0F3" box="[676,827,917,940]" pageId="3" pageNumber="388">TYPE MATERIAL</collectionCode>
. —
<typeStatus id="C67ECAE89569FFA1407CB996FC53B0F5" box="[875,971,917,941]" pageId="3" pageNumber="388" type="holotype">Holotype</typeStatus>
:
<collectingCountry id="61D234DA9569FFA140C0B996FBD5B0F5" box="[983,1101,917,941]" name="Madagascar" pageId="3" pageNumber="388">Madagascar</collectingCountry>
,
<location id="1C1A22919569FFA1474FB996FD7AB09F" LSID="urn:lsid:plazi:treatment:916CC55C9569FFA44009B92EFB07B18D:1C1A22919569FFA1474FB996FD7AB09F" country="Madagascar" latitude="-16.85" longLatPrecision="1280" longitude="49.95" municipality="Anbohidena Forest Station" name="Ste Marie Island" pageId="3" pageNumber="388">Ste Marie Island</location>
,
<collectingMunicipality id="F91EEE309569FFA141F9B9ACFC63B09F" box="[750,1019,942,967]" pageId="3" pageNumber="388">Anbohidena Forest Station</collectingMunicipality>
,
<geoCoordinate id="7CF1128D9569FFA14711B9ACFBCEB09F" box="[1030,1110,943,967]" degrees="16" direction="south" minutes="51" orientation="latitude" pageId="3" pageNumber="388" precision="925" value="-16.85">16°51’S</geoCoordinate>
,
<geoCoordinate id="7CF1128D9569FFA14775B9ACFB2DB09F" box="[1122,1205,943,967]" degrees="49" direction="east" minutes="57" orientation="longitude" pageId="3" pageNumber="388" precision="925" value="49.95">49°57’E</geoCoordinate>
,
<date id="6D7B528A9569FFA141B3B9CBFCACB0B8" box="[676,820,968,992]" pageId="3" pageNumber="388" value="1955-12-27">
<collectingDate id="7D3FAB629569FFA141B3B9CBFCACB0B8" box="[676,820,968,992]" pageId="3" pageNumber="388" value="1955-12-27">27.XII.1955</collectingDate>
</date>
,
<collectionCode id="7FD4EC8F9569FFA14055B9CBFC1CB0B8" box="[834,900,968,992]" country="Madagascar" httpUri="http://grbio.org/cool/1qvb-e9wm" name="Institut Recherche Scientifique de Madagascar" pageId="3" pageNumber="388">IRSM</collectionCode>
leg., (Muséum national d’Histoire naturelle, Paris RS-8600).
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="51DF27C19569FFA141B3BE0AFC7EB763" pageId="3" pageNumber="388" type="etymology">
<paragraph id="197A744A9569FFA141B3BE0AFC7EB763" blockId="3.[676,1210,917,1504]" pageId="3" pageNumber="388">ETYMOLOGY. — The specific name makes reference to the pale color of the new species.</paragraph>
</subSubSection>
<subSubSection id="51DF27C19569FFA141B3BE48FC15B6B8" pageId="3" pageNumber="388" type="diagnosis">
<paragraph id="197A744A9569FFA141B3BE48FC15B6B8" blockId="3.[676,1210,917,1504]" pageId="3" pageNumber="388">
DIAGNOSIS. — Moderate to large in size (with respect to the genus) reaching
<quantity id="DE3DD9AF9569FFA140EFBE66FBDDB723" box="[1016,1093,1124,1149]" metricMagnitude="-2" metricUnit="m" metricValue="3.1" pageId="3" pageNumber="388" unit="mm" value="31.0">31 mm</quantity>
in length. General coloration pale yellow throughout both the body and the appendages. Carapace with a moderately pronounced concavity, forming a weak angle. Cheliceral dentition with basal teeth of movable fingers reduced and fused. Pectines with 14-14 teeth; fulcra present. Sternites smooth; sternite V without any bright zone on posterior edge. Telson less elongated than in the other species of the genus, more globular and rounded; aculeus and curved; subaculear tooth moderate and more to rhomboid with two basal granules. Tibial spurs very reduced. Pedipalp fixed and movable fingers with 8/8 almost linear rows of granules. Trichobothrial pattern
<typeStatus id="C67ECAE89569FFA14799BFAEFB21B69D" box="[1166,1209,1453,1477]" pageId="3" pageNumber="388">type</typeStatus>
A-α orthobothriotaxic.
</paragraph>
</subSubSection>
<subSubSection id="51DF27C19569FFA441B3BC0FFB07B18D" lastPageId="6" lastPageNumber="391" pageId="3" pageNumber="388" type="description">
<paragraph id="197A744A9569FFA141B3BC0FFC8BB51F" blockId="3.[676,1209,1548,1703]" pageId="3" pageNumber="388">
<materialsCitation id="A9AD7E179569FFA141B3BC0FFC8BB51F" collectionCode="DESCRIPTION, BASED" pageId="3" pageNumber="388" specimenCount="1" typeStatus="holotype">
<collectionCode id="7FD4EC8F9569FFA141B3BC0FFCE5B57C" box="[676,893,1548,1574]" pageId="3" pageNumber="388">DESCRIPTION BASED</collectionCode>
ON FEMALE
<typeStatus id="C67ECAE89569FFA14714BC12FBEBB57C" box="[1027,1139,1553,1572]" pageId="3" pageNumber="388" type="holotype">HOLOTYPE</typeStatus>
<emphasis id="2BB1A8589569FFA141B3BC2EFC8BB51F" box="[676,787,1581,1607]" italics="true" pageId="3" pageNumber="388">Coloration</emphasis>
</materialsCitation>
</paragraph>
<paragraph id="197A744A9569FFA141B3BC4EFBCBB5FF" blockId="3.[676,1209,1548,1703]" pageId="3" pageNumber="388">Ground color pale yellow throughout the body and appendages; eyes surrounded by black pigment. Cheliceral fingers reddish-yellow.</paragraph>
<caption id="4DBA24C29569FFA14366BC6FFD89B5FF" ID-DOI="http://doi.org/10.5281/zenodo.5394768" ID-Zenodo-Dep="5394768" httpUri="https://zenodo.org/record/5394768/files/figure.png" pageId="3" pageNumber="388" startId="3.[113,124,1644,1661]" targetBox="[126,648,256,1541]" targetPageId="3">
<paragraph id="197A744A9569FFA14366BC6FFD89B5FF" blockId="3.[113,646,1644,1703]" pageId="3" pageNumber="388">
FIG. 2. —
<taxonomicName id="DEC50FC99569FFA143D2BC6FFEF5B525" authority="Lourenço, 2004" authorityName="Lourenço" authorityYear="2004" box="[197,365,1644,1661]" class="Arachnida" family="Buthidae" genus="Tityobuthus" kingdom="Animalia" order="Scorpiones" pageId="3" pageNumber="388" phylum="Arthropoda" rank="species" species="pallidus" status="sp. nov.">
<emphasis id="2BB1A8589569FFA143D2BC6FFEF5B525" box="[197,365,1644,1661]" italics="true" pageId="3" pageNumber="388">Tityobuthus pallidus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="308215239569FFA14264BC6FFE3EB525" box="[371,422,1644,1661]" pageId="3" pageNumber="388" rank="species">n. sp.</taxonomicNameLabel>
, holotype, trichobothrial pattern;
<emphasis id="2BB1A8589569FFA143D7BC82FF55B5CA" bold="true" box="[192,205,1665,1682]" pageId="3" pageNumber="388">A</emphasis>
, tibia, dorso-external aspect;
<emphasis id="2BB1A8589569FFA142FFBC82FE6DB5CA" bold="true" box="[488,501,1665,1682]" pageId="3" pageNumber="388">B</emphasis>
, patella, dorsal aspect;
<emphasis id="2BB1A8589569FFA143A3BC95FF5AB5FF" bold="true" box="[180,194,1686,1703]" pageId="3" pageNumber="388">C</emphasis>
, femur, dorsal aspect. Scale bar: 1 mm.
</paragraph>
</caption>
<caption id="4DBA24C2956EFFA64366BC82FCEFB5FF" ID-DOI="http://doi.org/10.5281/zenodo.5394770" ID-Zenodo-Dep="5394770" httpUri="https://zenodo.org/record/5394770/files/figure.png" pageId="4" pageNumber="389" startId="4.[113,124,1665,1682]" targetBox="[247,1089,239,1622]" targetPageId="4">
<paragraph id="197A744A956EFFA64366BC82FCEFB5FF" blockId="4.[113,1211,1665,1703]" pageId="4" pageNumber="389">
FIG. 3. — Metasomal segment V and telson, lateral aspect;
<emphasis id="2BB1A858956EFFA6417DBC82FDEFB5CA" bold="true" box="[618,631,1665,1682]" pageId="4" pageNumber="389">A</emphasis>
,
<taxonomicName id="DEC50FC9956EFFA64195BC82FCB2B5CA" authority="Lourenço, 2004" authorityName="Lourenço" authorityYear="2004" box="[642,810,1665,1682]" class="Arachnida" family="Buthidae" genus="Tityobuthus" kingdom="Animalia" order="Scorpiones" pageId="4" pageNumber="389" phylum="Arthropoda" rank="species" species="pallidus" status="sp. nov.">
<emphasis id="2BB1A858956EFFA64195BC82FCB2B5CA" box="[642,810,1665,1682]" italics="true" pageId="4" pageNumber="389">Tityobuthus pallidus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="30821523956EFFA64027BC82FCFCB5CA" box="[816,868,1665,1682]" pageId="4" pageNumber="389" rank="species">n. sp.</taxonomicNameLabel>
, holotype;
<emphasis id="2BB1A858956EFFA640C1BC82FC7BB5CA" bold="true" box="[982,995,1665,1682]" pageId="4" pageNumber="389">B</emphasis>
,
<taxonomicName id="DEC50FC9956EFFA640F8BC82FF05B5FF" authority="Lourenco, 1985" authorityName="Lourenco" authorityYear="1985" class="Arachnida" family="Buthidae" genus="Tityobuthus" kingdom="Animalia" order="Scorpiones" pageId="4" pageNumber="389" phylum="Arthropoda" rank="species" species="guillaumeti">
<emphasis id="2BB1A858956EFFA640F8BC82FBC7B5CA" box="[1007,1119,1665,1682]" italics="true" pageId="4" pageNumber="389">T. guillaumeti</emphasis>
Lourenço, 1985
</taxonomicName>
,;
<emphasis id="2BB1A858956EFFA643A8BC95FF55B5FF" bold="true" box="[191,205,1686,1703]" pageId="4" pageNumber="389">C</emphasis>
,
<taxonomicName id="DEC50FC9956EFFA643C0BC95FE3EB5FF" authority="(Pocock, 1890)" baseAuthorityName="Pocock" baseAuthorityYear="1890" box="[215,422,1686,1703]" class="Arachnida" family="Buthidae" genus="Tityobuthus" kingdom="Animalia" order="Scorpiones" pageId="4" pageNumber="389" phylum="Arthropoda" rank="species" species="baroni">
<emphasis id="2BB1A858956EFFA643C0BC95FEB8B5FF" box="[215,288,1686,1703]" italics="true" pageId="4" pageNumber="389">T. baroni</emphasis>
(
<bibRefCitation id="7D5409BB956EFFA6423DBC95FE39B5FF" author="POCOCK R. I." box="[298,417,1686,1703]" pageId="4" pageNumber="389" pagination="114 - 141" refId="ref2982" refString="POCOCK R. I. 1890. - A revision of the genera of scorpions of the family Buthidae, with descriptions of some South-African species. Proceedings of the Zoological Society of London 11: 114 - 141." type="journal article" year="1890">Pocock, 1890</bibRefCitation>
)
</taxonomicName>
,;
<emphasis id="2BB1A858956EFFA642D0BC95FE4DB5FF" bold="true" box="[455,469,1686,1703]" pageId="4" pageNumber="389">D</emphasis>
,
<taxonomicName id="DEC50FC9956EFFA642C8BC95FD25B5FF" authority="Lourenco, 1996" authorityName="Lourenco" authorityYear="1996" box="[479,701,1686,1703]" class="Arachnida" family="Buthidae" genus="Tityobuthus" kingdom="Animalia" order="Scorpiones" pageId="4" pageNumber="389" phylum="Arthropoda" rank="species" species="lucileae">
<emphasis id="2BB1A858956EFFA642C8BC95FDABB5FF" box="[479,563,1686,1703]" italics="true" pageId="4" pageNumber="389">T. lucileae</emphasis>
<bibRefCitation id="7D5409BB956EFFA6412FBC95FD25B5FF" author="LOURENCO W. R." box="[568,701,1686,1703]" pageId="4" pageNumber="389" pagination="1 - 102" refId="ref2888" refString="LOURENCO W. R. 1996. - Scorpions. Faune de Madagascar 87: 1 - 102." type="journal article" year="1996">Lourenço, 1996</bibRefCitation>
</taxonomicName>
,. Scale bars: 1 mm.
</paragraph>
</caption>
<paragraph id="197A744A956FFFA74397BA4CFF6FB307" blockId="5.[128,247,79,95]" box="[128,247,79,95]" pageId="5" pageNumber="390">Lourenço W. R.</paragraph>
<caption id="4DBA24C2956FFFA74366B9B7FCC9B083" ID-DOI="http://doi.org/10.5281/zenodo.5394772" ID-Zenodo-Dep="5394772" httpUri="https://zenodo.org/record/5394772/files/figure.png" pageId="5" pageNumber="390" startId="5.[113,124,948,965]" targetBox="[113,1210,204,992]" targetPageId="5">
<paragraph id="197A744A956FFFA74366B9B7FCC9B083" blockId="5.[113,1210,948,987]" pageId="5" pageNumber="390">
FIG. 4. — Maps of Madagascar and Ste Marie Island, showing the known distribution of
<taxonomicName id="DEC50FC9956FFFA74066B9B7FB01B09D" authority="(Pocock, 1890)" baseAuthorityName="Pocock" baseAuthorityYear="1890" box="[881,1177,948,965]" class="Arachnida" family="Buthidae" genus="Tityobuthus" kingdom="Animalia" order="Scorpiones" pageId="5" pageNumber="390" phylum="Arthropoda" rank="species" species="baroni">
<emphasis id="2BB1A858956FFFA74066B9B7FB88B09D" box="[881,1040,948,965]" italics="true" pageId="5" pageNumber="390">Tityobuthus baroni</emphasis>
(
<bibRefCitation id="7D5409BB956FFFA7470BB9B7FB0CB09D" author="POCOCK R. I." box="[1052,1172,948,965]" pageId="5" pageNumber="390" pagination="114 - 141" refId="ref2982" refString="POCOCK R. I. 1890. - A revision of the genera of scorpions of the family Buthidae, with descriptions of some South-African species. Proceedings of the Zoological Society of London 11: 114 - 141." type="journal article" year="1890">Pocock, 1890</bibRefCitation>
)
</taxonomicName>
(
<emphasis id="2BB1A858956FFFA747B3B9B7FB36B09D" bold="true" box="[1188,1198,948,965]" pageId="5" pageNumber="390">1</emphasis>
),
<taxonomicName id="DEC50FC9956FFFA74366B9C9FED5B083" authority="Lourenco, 1996" authorityName="Lourenco" authorityYear="1996" box="[113,333,970,987]" class="Arachnida" family="Buthidae" genus="Tityobuthus" kingdom="Animalia" order="Scorpiones" pageId="5" pageNumber="390" phylum="Arthropoda" rank="species" species="lucileae">
<emphasis id="2BB1A858956FFFA74366B9C9FF5DB083" box="[113,197,970,987]" italics="true" pageId="5" pageNumber="390">T. lucileae</emphasis>
<bibRefCitation id="7D5409BB956FFFA743DDB9C9FED5B083" author="LOURENCO W. R." box="[202,333,970,987]" pageId="5" pageNumber="390" pagination="1 - 102" refId="ref2888" refString="LOURENCO W. R. 1996. - Scorpions. Faune de Madagascar 87: 1 - 102." type="journal article" year="1996">Lourenço, 1996</bibRefCitation>
</taxonomicName>
(
<emphasis id="2BB1A858956FFFA74241B9C9FEF8B083" bold="true" box="[342,352,970,987]" pageId="5" pageNumber="390">2</emphasis>
),
<taxonomicName id="DEC50FC9956FFFA74267B9C9FDFFB083" authority="Lourenco, 1985" authorityName="Lourenco" authorityYear="1985" box="[368,615,970,987]" class="Arachnida" family="Buthidae" genus="Tityobuthus" kingdom="Animalia" order="Scorpiones" pageId="5" pageNumber="390" phylum="Arthropoda" rank="species" species="guillaumeti">
<emphasis id="2BB1A858956FFFA74267B9C9FE47B083" box="[368,479,970,987]" italics="true" pageId="5" pageNumber="390">T. guillaumeti</emphasis>
Lourenço, 1985
</taxonomicName>
(
<emphasis id="2BB1A858956FFFA74166B9C9FDE3B083" bold="true" box="[625,635,970,987]" pageId="5" pageNumber="390">3</emphasis>
) and
<taxonomicName id="DEC50FC9956FFFA741BDB9C9FD67B083" authority="Lourenço, 2004" authorityName="Lourenço" authorityYear="2004" box="[682,767,970,987]" class="Arachnida" family="Buthidae" genus="Tityobuthus" kingdom="Animalia" order="Scorpiones" pageId="5" pageNumber="390" phylum="Arthropoda" rank="species" species="pallidus" status="sp. nov.">
<emphasis id="2BB1A858956FFFA741BDB9C9FD67B083" box="[682,767,970,987]" italics="true" pageId="5" pageNumber="390">T. pallidus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="30821523956FFFA74014B9C9FCABB083" box="[771,819,970,987]" pageId="5" pageNumber="390" rank="species">n. sp.</taxonomicNameLabel>
(
<emphasis id="2BB1A858956FFFA7402BB9C9FCDEB083" bold="true" box="[828,838,970,987]" pageId="5" pageNumber="390">4</emphasis>
).
</paragraph>
</caption>
<paragraph id="197A744A956FFFA74366BE2EFF73B71F" blockId="5.[113,647,1069,1703]" box="[113,235,1069,1095]" pageId="5" pageNumber="390">
<emphasis id="2BB1A858956FFFA74366BE2EFF73B71F" box="[113,235,1069,1095]" italics="true" pageId="5" pageNumber="390">Morphology</emphasis>
</paragraph>
<paragraph id="197A744A956FFFA74366BE4EFCAEB5FF" blockId="5.[113,647,1069,1703]" lastBlockId="5.[676,1210,1068,1703]" pageId="5" pageNumber="390">
Carapace weakly granular; anterior margin with a moderately pronounced median concavity, forming a weak angle. All carinae weak; furrows moderate to weak. Median ocular tubercle distinctly anterior to the centre of the carapace; median eyes separated by less than one ocular diameter. Three pairs of lateral eyes. Sternum subtriangular to subpentagonal. Mesosoma: tergites weakly granular. Median carina weak on all tergites; other carinae absent. Tergite VII pentacarinate. Venter: genital operculum divided longitudinally, each half being oval-shaped. Pectines: pectinal tooth count 14-14; basal middle lamellae not dilated; fulcra present. Sternites smooth with small, elongate stigmata; VII without carinae. Sternite V without any smooth or bright zone on posterior edge. Metasoma: segments I-III with 10 carinae, moderately crenulate. Segment IV with eight carinae, moderately crenulate. Inter- carinal spaces weakly to moderately granular. Segment V with five carinae, rounded and weakly granular except on the ventral surface which shows thin granulation. Telson smooth, with a globular shape and a long strongly curved aculeus; subaculear tooth moderate and tending to rhomboid in shape with two basal granules. Cheliceral dentition characteristic of the family
<taxonomicName id="DEC50FC9956FFFA741B3BF2EFC9FB61F" authorityName="C. L. Koch" authorityYear="1837" box="[676,775,1325,1351]" class="Arachnida" family="Buthidae" kingdom="Animalia" order="Scorpiones" pageId="5" pageNumber="390" phylum="Arthropoda" rank="family">Buthidae</taxonomicName>
(see
<bibRefCitation id="7D5409BB956FFFA74029BF2EFC4DB61F" author="VACHON M." box="[830,981,1325,1351]" pageId="5" pageNumber="390" pagination="161 - 166" refId="ref3066" refString="VACHON M. 1963. - De l'utilite, en systematique, d'une nomenclature des dents des cheliceres chez les scorpions. Bulletin du Museum national d'Histoire naturelle 2 e ser., 35 (2): 161 - 166." type="journal article" year="1963">Vachon 1963</bibRefCitation>
); basal teeth of movable fingers reduced and fused; ventral surfaces of finger and manus with setae. Pedipalps: femur pentacarinate; patella and chela with some carinae, weakly crenulate; internal face of patella with five spinoid granules; all faces weakly granular; fixed and movable fingers with 8/8 almost linear rows of granules. Trichobothriotaxy; orthobothriotaxy A-α (cf.
<bibRefCitation id="7D5409BB956FFFA740DDBC2EFBFBB51F" author="VACHON M." box="[970,1123,1581,1607]" pageId="5" pageNumber="390" pagination="857 - 958" refId="ref3108" refString="VACHON M. 1974. - Etude des caracteres utilises pour classer les familles et les genres de scorpions (arachnides). 1. La trichobothriotaxie en arachnologie. Sigles trichobothriaux et types de trichobothriotaxie chez les scorpions. Bulletin du Museum national d'Histoire naturelle 3 e ser., n ° 140, Zoologie 104: 857 - 958." type="journal article" year="1974">Vachon 1974</bibRefCitation>
,
<bibRefCitation id="7D5409BB956FFFA74766BC2EFB35B51F" author="VACHON M." box="[1137,1197,1581,1607]" pageId="5" pageNumber="390" pagination="1597 - 1599" refId="ref3170" refString="VACHON M. 1975. - Sur l'utilisation de la trichobothriotaxie du bras des pedipalpes des scorpions (arachnides) dans le classement des genres de la famille des Buthidae Simon. Comptes Rendus de l'Academie des Sciences ser. D, 281: 1597 - 1599." type="journal article" year="1975">1975</bibRefCitation>
). Legs: tarsus with numerous fine median setae ventrally. Pedal spurs moderate and tibial spurs very reduced.
</paragraph>
<paragraph id="197A744A956CFFA44366BACFFE8EB3BC" blockId="6.[113,646,204,743]" box="[113,278,204,230]" pageId="6" pageNumber="391">RELATIONSHIPS</paragraph>
<caption id="4DBA24C2956CFFA441B3BAC8FB21B384" box="[676,1209,203,220]" pageId="6" pageNumber="391" startId="6.[676,687,203,220]">
<paragraph id="197A744A956CFFA441B3BAC8FB21B384" blockId="6.[676,1209,203,725]" box="[676,1209,203,220]" pageId="6" pageNumber="391">TABLE 1. — Morphometric values (in mm) of the female holotype</paragraph>
</caption>
<paragraph id="197A744A956CFFA44366BAEFFE99B1BE" blockId="6.[113,646,204,743]" pageId="6" pageNumber="391">
From its general morphology, and especially in the total lack of pigmentation,
<taxonomicName id="DEC50FC9956CFFA442C4BB0EFF3AB21F" authority="Lourenço, 2004" authorityName="Lourenço" authorityYear="2004" class="Arachnida" family="Buthidae" genus="Tityobuthus" kingdom="Animalia" order="Scorpiones" pageId="6" pageNumber="391" phylum="Arthropoda" rank="species" species="pallidus" status="sp. nov.">
<emphasis id="2BB1A858956CFFA442C4BB0EFF3AB21F" italics="true" pageId="6" pageNumber="391">Tityobuthus pallidus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="30821523956CFFA443BBBB2FFF71B21E" box="[172,233,300,326]" pageId="6" pageNumber="391" rank="species">n. sp.</taxonomicNameLabel>
appears to be most closely related to
<taxonomicName id="DEC50FC9956CFFA44366BB4FFE4DB23F" authority="Lourenco, 1995" authorityName="Lourenco" authorityYear="1995" box="[113,469,332,359]" class="Arachnida" family="Buthidae" genus="Tityobuthus" kingdom="Animalia" order="Scorpiones" pageId="6" pageNumber="391" phylum="Arthropoda" rank="species" species="guillaumeti">
<emphasis id="2BB1A858956CFFA44366BB4FFE8BB23F" box="[113,275,332,359]" italics="true" pageId="6" pageNumber="391">T. guillaumeti</emphasis>
Lourenço, 1995
</taxonomicName>
and
<taxonomicName id="DEC50FC9956CFFA44104BB4FFEBDB2DF" authority="Lourenco, 1996" authorityName="Lourenco" authorityYear="1996" class="Arachnida" family="Buthidae" genus="Tityobuthus" kingdom="Animalia" order="Scorpiones" pageId="6" pageNumber="391" phylum="Arthropoda" rank="species" species="judsoni">
<emphasis id="2BB1A858956CFFA44104BB4FFD1EB23F" box="[531,646,332,359]" italics="true" pageId="6" pageNumber="391">T. judsoni</emphasis>
<bibRefCitation id="7D5409BB956CFFA44366BB6FFEBDB2DF" author="LOURENCO W. R." box="[113,293,364,391]" pageId="6" pageNumber="391" pagination="1 - 102" refId="ref2888" refString="LOURENCO W. R. 1996. - Scorpions. Faune de Madagascar 87: 1 - 102." type="journal article" year="1996">Lourenço, 1996</bibRefCitation>
</taxonomicName>
. The new species can be readily distinguished from the other two by the following features: 1) the presence of fulcra on the pectines of
<taxonomicName id="DEC50FC9956CFFA443E7BBCFFEFBB2BF" authority="Lourenço, 2004" authorityName="Lourenço" authorityYear="2004" box="[240,355,460,487]" class="Arachnida" family="Buthidae" genus="Tityobuthus" kingdom="Animalia" order="Scorpiones" pageId="6" pageNumber="391" phylum="Arthropoda" rank="species" species="pallidus" status="sp. nov.">
<emphasis id="2BB1A858956CFFA443E7BBCFFEFBB2BF" box="[240,355,460,487]" italics="true" pageId="6" pageNumber="391">T. pallidus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="30821523956CFFA4427CBBCFFE32B2BE" box="[363,426,460,486]" pageId="6" pageNumber="391" rank="species">n. sp.</taxonomicNameLabel>
, whereas in
<taxonomicName id="DEC50FC9956CFFA44123BBCFFF5FB15F" authorityName="Lourenco" authorityYear="1995" class="Arachnida" family="Buthidae" genus="Tityobuthus" kingdom="Animalia" order="Scorpiones" pageId="6" pageNumber="391" phylum="Arthropoda" rank="species" species="guillaumeti">
<emphasis id="2BB1A858956CFFA44123BBCFFF5FB15F" italics="true" pageId="6" pageNumber="391">T. guillaumeti</emphasis>
</taxonomicName>
fulcra are absent; 2) a more globular shape of the vesicle in
<taxonomicName id="DEC50FC9956CFFA4427CB80FFE46B17F" authority="Lourenço, 2004" authorityName="Lourenço" authorityYear="2004" box="[363,478,524,551]" class="Arachnida" family="Buthidae" genus="Tityobuthus" kingdom="Animalia" order="Scorpiones" pageId="6" pageNumber="391" phylum="Arthropoda" rank="species" species="pallidus" status="sp. nov.">
<emphasis id="2BB1A858956CFFA4427CB80FFE46B17F" box="[363,478,524,551]" italics="true" pageId="6" pageNumber="391">T. pallidus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="30821523956CFFA442F1B80FFDBBB17E" box="[486,547,524,550]" pageId="6" pageNumber="391" rank="species">n. sp.</taxonomicNameLabel>
(in both
<taxonomicName id="DEC50FC9956CFFA44366B82FFE90B11F" authorityName="Lourenco" authorityYear="1995" box="[113,264,556,583]" class="Arachnida" family="Buthidae" genus="Tityobuthus" kingdom="Animalia" order="Scorpiones" pageId="6" pageNumber="391" phylum="Arthropoda" rank="species" species="guillaumeti">
<emphasis id="2BB1A858956CFFA44366B82FFE90B11F" box="[113,264,556,583]" italics="true" pageId="6" pageNumber="391">T. guillaumeti</emphasis>
</taxonomicName>
[
<figureCitation id="81FE68CF956CFFA44202B82EFEF1B11F" box="[277,361,557,583]" captionStart="FIG" captionStartId="4.[113,124,1665,1682]" captionTargetBox="[247,1089,239,1622]" captionTargetId="figure-25@4.[156,1166,218,1716]" captionTargetPageId="4" captionText="FIG. 3. — Metasomal segment V and telson, lateral aspect; A, Tityobuthus pallidus n. sp., holotype; B, T. guillaumeti Lourenço, 1985,; C, T. baroni (Pocock, 1890),; D, T. lucileae Lourenço, 1996,. Scale bars: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5394770" httpUri="https://zenodo.org/record/5394770/files/figure.png" pageId="6" pageNumber="391">Fig. 3B</figureCitation>
] and
<taxonomicName id="DEC50FC9956CFFA442B1B82FFD89B11F" authorityName="Lourenco" authorityYear="1996" box="[422,529,556,583]" class="Arachnida" family="Buthidae" genus="Tityobuthus" kingdom="Animalia" order="Scorpiones" pageId="6" pageNumber="391" phylum="Arthropoda" rank="species" species="judsoni">
<emphasis id="2BB1A858956CFFA442B1B82FFD89B11F" box="[422,529,556,583]" italics="true" pageId="6" pageNumber="391">T. judsoni</emphasis>
</taxonomicName>
the vesicle is elongated); 3) pectines of
<taxonomicName id="DEC50FC9956CFFA442D6B84FFDA3B13F" authority="Lourenço, 2004" authorityName="Lourenço" authorityYear="2004" box="[449,571,588,615]" class="Arachnida" family="Buthidae" genus="Tityobuthus" kingdom="Animalia" order="Scorpiones" pageId="6" pageNumber="391" phylum="Arthropoda" rank="species" species="pallidus" status="sp. nov.">
<emphasis id="2BB1A858956CFFA442D6B84FFDA3B13F" box="[449,571,588,615]" italics="true" pageId="6" pageNumber="391">T. pallidus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="30821523956CFFA44152B84FFD1EB13E" box="[581,646,588,614]" pageId="6" pageNumber="391" rank="species">n. sp.</taxonomicNameLabel>
have 14-14 teeth whereas those of
<taxonomicName id="DEC50FC9956CFFA44106B86FFD1EB1DF" authorityName="Lourenco" authorityYear="1996" box="[529,646,620,647]" class="Arachnida" family="Buthidae" genus="Tityobuthus" kingdom="Animalia" order="Scorpiones" pageId="6" pageNumber="391" phylum="Arthropoda" rank="species" species="judsoni">
<emphasis id="2BB1A858956CFFA44106B86FFD1EB1DF" box="[529,646,620,647]" italics="true" pageId="6" pageNumber="391">T. judsoni</emphasis>
</taxonomicName>
have 18-19 teeth; and 4) tibial spurs are very reduced in
<taxonomicName id="DEC50FC9956CFFA443FBB8AFFEC6B19F" authority="Lourenço, 2004" authorityName="Lourenço" authorityYear="2004" box="[236,350,684,711]" class="Arachnida" family="Buthidae" genus="Tityobuthus" kingdom="Animalia" order="Scorpiones" pageId="6" pageNumber="391" phylum="Arthropoda" rank="species" species="pallidus" status="sp. nov.">
<emphasis id="2BB1A858956CFFA443FBB8AFFEC6B19F" box="[236,350,684,711]" italics="true" pageId="6" pageNumber="391">T. pallidus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="30821523956CFFA44272B8AFFE39B19E" box="[357,417,684,710]" pageId="6" pageNumber="391" rank="species">n. sp.</taxonomicNameLabel>
but strongly marked in
<taxonomicName id="DEC50FC9956CFFA44399B8CFFF61B1BF" authorityName="Lourenco" authorityYear="1996" box="[142,249,716,743]" class="Arachnida" family="Buthidae" genus="Tityobuthus" kingdom="Animalia" order="Scorpiones" pageId="6" pageNumber="391" phylum="Arthropoda" rank="species" species="judsoni">
<emphasis id="2BB1A858956CFFA44399B8CFFF61B1BF" box="[142,249,716,743]" italics="true" pageId="6" pageNumber="391">T. judsoni</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="197A744A956CFFA441B3BAE3FC0DB3A9" blockId="6.[676,1209,203,725]" box="[676,917,224,241]" pageId="6" pageNumber="391">
of
<taxonomicName id="DEC50FC9956CFFA441ADBAE3FCF8B3A9" authority="Lourenço, 2004" authorityName="Lourenço" authorityYear="2004" box="[698,864,224,241]" class="Arachnida" family="Buthidae" genus="Tityobuthus" kingdom="Animalia" order="Scorpiones" pageId="6" pageNumber="391" phylum="Arthropoda" rank="species" species="pallidus" status="sp. nov.">
<emphasis id="2BB1A858956CFFA441ADBAE3FCF8B3A9" box="[698,864,224,241]" italics="true" pageId="6" pageNumber="391">Tityobuthus pallidus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="30821523956CFFA44072BAE3FC0DB3A9" box="[869,917,224,241]" pageId="6" pageNumber="391" rank="species">n. sp.</taxonomicNameLabel>
</paragraph>
<paragraph id="197A744A956CFFA441B3BB12FB07B27D" blockId="6.[676,1209,203,725]" box="[676,1183,273,293]" pageId="6" pageNumber="391">Total length 31.1</paragraph>
<paragraph id="197A744A956CFFA441B3BB2AFB07B235" blockId="6.[676,1209,203,725]" pageId="6" pageNumber="391">Carapace length 3.5 anterior width 2.4 posterior width 3.8</paragraph>
<paragraph id="197A744A956CFFA441B3BB72FB07B2C5" blockId="6.[676,1209,203,725]" pageId="6" pageNumber="391">Metasomal segment I length 2.3 width 1.8</paragraph>
<paragraph id="197A744A956CFFA441B3BBA2FB07B2BD" blockId="6.[676,1209,203,725]" pageId="6" pageNumber="391">Metasomal segment V length 5.1 width 1.6 depth 1.7</paragraph>
<paragraph id="197A744A956CFFA441B3BBEAFB07B14D" blockId="6.[676,1209,203,725]" pageId="6" pageNumber="391">Vesicle width 1.4 depth 1.4</paragraph>
<paragraph id="197A744A956CFFA441B3B81AFB07B1E5" blockId="6.[676,1209,203,725]" pageId="6" pageNumber="391">Pedipalp femur length 3.6 femur width 1.0 patella length 4.6 patella width 1.5 chela length 6.6 chela width 1.2 chela depth 1.1</paragraph>
<paragraph id="197A744A956CFFA441B3B8C2FB07B18D" blockId="6.[676,1209,203,725]" box="[676,1183,705,725]" pageId="6" pageNumber="391">Movable finger length 4.6</paragraph>
</subSubSection>
</treatment>
</document>
Reference in a new issue View git blame Copy permalink