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<document id="EAB65D9784642D7385772509E22D4808" ID-CLB-Dataset="22256" ID-DOI="10.11646/zootaxa.4772.3.7" ID-GBIF-Dataset="8b277405-a91c-4d0c-b77e-8b1d4820c7cd" ID-ISSN="1175-5326" ID-Zenodo-Dep="3819764" ID-ZooBank="8DE2F40F-0931-4002-97C4-5603E363B7E8" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1589175825495" checkinUser="plazi" docAuthor="Arce-Pérez, Roberto &amp; Morón, Miguel Ángel" docDate="2020" docId="6C5796057B70FFCAFF0939109C2F2E56" docLanguage="en" docName="zootaxa.4772.3.7.pdf" docOrigin="Zootaxa 4772 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Macrodactylus zunilensis Bates 1887" docType="treatment" docVersion="7" lastPageNumber="582" masterDocId="906EEE7D7B7EFFC5FF9E3F1F990C2A65" masterDocTitle="Review of the species of Macrodactylus Dejean (Coleoptera: Scarabaeidae Melolonthinae) from the Central American Nucleus" masterLastPageNumber="584" masterPageNumber="567" pageNumber="581" updateTime="1698839127964" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="45D77531BAA7052187AFEC19092CC2BB">Review of the species of Macrodactylus Dejean (Coleoptera: Scarabaeidae Melolonthinae) from the Central American Nucleus</mods:title>
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<mods:namePart id="B224E6510229782D3185EBF1DBE5DF8E">Arce-Pérez, Roberto</mods:namePart>
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<mods:namePart id="4652BBB872275FE785A0C87D3B58A607">Morón, Miguel Ángel</mods:namePart>
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<mods:title id="306FDC41B5667543C18BD90162319F8A">Zootaxa</mods:title>
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<mods:date id="155E0F01D4C22868BEFA40E9EA0F5841">2020</mods:date>
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<mods:number id="663DFD38E318E87F79A7AD8B22510718">2020-05-11</mods:number>
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<mods:number id="CFB9A5FE18F7CA8F55389CB6622F361D">4772</mods:number>
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<treatment id="6C5796057B70FFCAFF0939109C2F2E56" ID-DOI="http://doi.org/10.5281/zenodo.3843995" ID-GBIF-Taxon="164035936" ID-Zenodo-Dep="3843995" LSID="urn:lsid:plazi:treatment:6C5796057B70FFCAFF0939109C2F2E56" httpUri="http://treatment.plazi.org/id/6C5796057B70FFCAFF0939109C2F2E56" lastPageId="15" lastPageNumber="582" pageId="14" pageNumber="581">
<subSubSection id="ACE474987B70FFCBFF09391098632C28" pageId="14" pageNumber="581" type="nomenclature">
<paragraph id="E44127137B70FFCBFF0939109B6C2C4C" blockId="14.[151,608,1551,1613]" box="[151,608,1551,1578]" pageId="14" pageNumber="581">
<heading id="BF09907F7B70FFCBFF0939109B6C2C4C" bold="true" box="[151,608,1551,1578]" fontSize="11" level="1" pageId="14" pageNumber="581" reason="1">
<taxonomicName id="23FE5C907B70FFCBFF0939109B6C2C4C" authority="Bates, 1887" authorityName="Bates" authorityYear="1887" box="[151,608,1551,1578]" class="Insecta" family="Melolonthidae" genus="Macrodactylus" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="581" phylum="Arthropoda" rank="species" species="zunilensis">
<emphasis id="D68AFB017B70FFCBFF0939109B6C2C4C" bold="true" box="[151,608,1551,1578]" pageId="14" pageNumber="581">
<emphasis id="D68AFB017B70FFCBFF09391098C72C4F" bold="true" box="[151,459,1551,1578]" italics="true" pageId="14" pageNumber="581">Macrodactylus zunilensis</emphasis>
<bibRefCitation id="806F5AE27B70FFCBFE4C39109B6C2C4C" author="Bates, H. W." box="[466,608,1551,1577]" pageId="14" pageNumber="581" pagination="25 - 160" refId="ref10965" refString="Bates, H. W. (1887) Pectinicornia and Lamellicornia. In: Salvin, O. &amp; Godman, F. (Eds.), Biologia Centrali-Americana, Insecta, Coleoptera, volume 2, part 2. Taylor and Francis, London, pp. 25 - 160. Available from: www. biodiversitylibrary. org / item / 135502 # page / 10 / mode / 1 up (accessed 27 February 2020)" type="journal article" year="1887">Bates, 1887</bibRefCitation>
</emphasis>
</taxonomicName>
</heading>
</paragraph>
<paragraph id="E44127137B70FFCBFF09392C98632C28" blockId="14.[151,608,1551,1613]" box="[151,367,1586,1613]" pageId="14" pageNumber="581">Figs. 56, 8183, 87</paragraph>
</subSubSection>
<subSubSection id="ACE474987B70FFCBFF0939659BBF2CF3" box="[151,691,1658,1686]" pageId="14" pageNumber="581" type="materials_examined">
<paragraph id="E44127137B70FFCBFF0939659BBF2CF3" blockId="14.[151,1436,1658,2046]" box="[151,691,1658,1686]" pageId="14" pageNumber="581">
<emphasis id="D68AFB017B70FFCBFF093965988D2CF0" bold="true" box="[151,385,1658,1685]" pageId="14" pageNumber="581">Material examined.</emphasis>
<specimenCount id="F2F8EC9A7B70FFCBFE1939649B2F2CF0" box="[391,547,1659,1685]" pageId="14" pageNumber="581" type="generic">96 specimens</specimenCount>
:
<specimenCount id="F2F8EC9A7B70FFCBFDB039649B642CF3" box="[558,616,1659,1686]" pageId="14" pageNumber="581" type="male">48 ♂</specimenCount>
,
<specimenCount id="F2F8EC9A7B70FFCBFDEA39649BA12CF3" box="[628,685,1659,1686]" pageId="14" pageNumber="581" type="female">48 ♀</specimenCount>
.
</paragraph>
</subSubSection>
<subSubSection id="ACE474987B70FFCAFF5939819C2F2E56" lastPageId="15" lastPageNumber="582" pageId="14" pageNumber="581" type="diagnosis">
<paragraph id="E44127137B70FFCBFF5939819D502D08" blockId="14.[151,1436,1658,2046]" pageId="14" pageNumber="581">
<emphasis id="D68AFB017B70FFCBFF593981984E2CDC" bold="true" box="[199,322,1694,1721]" pageId="14" pageNumber="581">Diagnosis.</emphasis>
Body length 8.510.0 mm; integument reddish yellow; dorsally with dense vestiture of scale-like, reddish yellow, slightly flattened setae, most dense in sutural region of elytra; pygidium with scattered setae; femora reddish yellow; tibiae reddish yellow with the apices black; tarsi black with rings of white setae (Fig. 56); parameres stout, oval, glabrous, completely fused dorsobasally, forming an oval in distal view, apices lanceolate, the dorsobasal region with an extra laminar projection internally, but in lateral view, it is evident that it is part of the parameres (Figs. 8182); female with vestiture and color similar to male; genital plates (Fig. 83).
</paragraph>
<paragraph id="E44127137B70FFCBFF59386898D82DD0" blockId="14.[151,1436,1658,2046]" pageId="14" pageNumber="581">
<emphasis id="D68AFB017B70FFCBFF593868988B2DF4" bold="true" box="[199,391,1910,1937]" pageId="14" pageNumber="581">Natural history.</emphasis>
This species inhabits rain forests ranging from
<quantity id="23068AF67B70FFCBFC3438689D632DF7" box="[938,1135,1911,1938]" metricMagnitude="0" metricUnit="m" metricValue="2.0" metricValueMax="3.0" metricValueMin="1.0" pageId="14" pageNumber="581" unit="m" value="2.0" valueMax="3.0" valueMin="1.0">1,000 to 3,000 m</quantity>
(additional information in
<bibRefCitation id="806F5AE27B70FFCBFF09388498C42DD0" author="Arce-Perez, R. &amp; Moron, M. A." box="[151,456,1947,1973]" pageId="14" pageNumber="581" refId="ref10871" refString="Arce-Perez, R. &amp; Moron, M. A. (2011) Los Macrodactylus (Coleoptera: Melolonthidae) de America Central. Editorial Academica Espanola, Saarbrucken, 275 pp." type="book" year="2011">Arce-Pérez &amp; Morón 2011</bibRefCitation>
).
</paragraph>
<paragraph id="E44127137B70FFCBFF5938A19A972D98" blockId="14.[151,1436,1658,2046]" pageId="14" pageNumber="581">
<emphasis id="D68AFB017B70FFCBFF5938A19B0F2DBC" bold="true" box="[199,515,1982,2009]" pageId="14" pageNumber="581">Geographical distribution</emphasis>
.
<materialsCitation id="54962D4E7B70FFCBFD8B38A09BEA2DBF" ID-GBIF-Occurrence="2611565328" box="[533,742,1983,2010]" country="Mexico" location="Chiapas" pageId="14" pageNumber="581" specimenCount="1" stateProvince="Chiapas">
<collectingCountry id="9CE967837B70FFCBFD8B38A09B672DBC" box="[533,619,1983,2009]" name="Mexico" pageId="14" pageNumber="581">Mexico</collectingCountry>
(
<collectingRegion id="263AE9F17B70FFCBFDE038A09BD22DBF" box="[638,734,1983,2010]" country="Mexico" name="Chiapas" pageId="14" pageNumber="581">Chiapas</collectingRegion>
)
</materialsCitation>
,
<materialsCitation id="54962D4E7B70FFCBFD6838A09BAC2D98" ID-GBIF-Occurrence="2611565320" country="Guatemala" location="Chimaltenango" pageId="14" pageNumber="581" specimenCount="1" stateProvince="Chimaltenango">
<collectingCountry id="9CE967837B70FFCBFD6838A09A632DBC" box="[758,879,1983,2009]" name="Guatemala" pageId="14" pageNumber="581">Guatemala</collectingCountry>
(
<collectingRegion id="263AE9F17B70FFCBFC1C38A09D3E2DBC" box="[898,1074,1983,2009]" country="Guatemala" name="Chimaltenango" pageId="14" pageNumber="581">Chímaltenango</collectingRegion>
,
<collectingRegion id="263AE9F17B70FFCBFBDC38A09DA12DBC" box="[1090,1197,1983,2009]" country="Guatemala" name="Escuintla" pageId="14" pageNumber="581">Escuintla</collectingRegion>
,
<collectingCountry id="9CE967837B70FFCBFB2238A09C342DBC" box="[1212,1336,1983,2009]" name="Guatemala" pageId="14" pageNumber="581">Guatemala</collectingCountry>
,
<collectingRegion id="263AE9F17B70FFCBFAD638A099F32D9B" country="Guatemala" name="Huehuetenango" pageId="14" pageNumber="581">Huehuetenango</collectingRegion>
,
<collectingRegion id="263AE9F17B70FFCBFE9438FC98B12D98" box="[266,445,2019,2045]" country="Guatemala" name="Quetzaltenango" pageId="14" pageNumber="581">Quetzaltenango</collectingRegion>
,
<collectingRegion id="263AE9F17B70FFCBFE5638FC9B692D98" box="[456,613,2019,2045]" country="Guatemala" name="Sacatepequez" pageId="14" pageNumber="581">Sacatepéquez</collectingRegion>
), and
</materialsCitation>
<collectingCountry id="9CE967837B70FFCBFD3938FC9A272D98" box="[679,811,2019,2045]" name="El Salvador" pageId="14" pageNumber="581">El Salvador</collectingCountry>
(Fig. 87).
</paragraph>
<paragraph id="E44127137B71FFCAFF093CE99C2F2E56" blockId="15.[151,1436,1014,1075]" pageId="15" pageNumber="582">
<emphasis id="D68AFB017B71FFCAFF093CE999F22E6A" bold="true" box="[151,254,1014,1039]" pageId="15" pageNumber="582">Figure 87</emphasis>
. Distribution of
<taxonomicName id="23FE5C907B71FFCAFE393CE99B322E6B" authorityName="Dejean" authorityYear="1821" box="[423,574,1014,1038]" class="Insecta" family="Melolonthidae" genus="Macrodactylus" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="582" phylum="Arthropoda" rank="genus">
<emphasis id="D68AFB017B71FFCAFE393CE99B322E6B" box="[423,574,1014,1038]" italics="true" pageId="15" pageNumber="582">Macrodactylus</emphasis>
</taxonomicName>
in the Central American Nucleus:
<taxonomicName id="23FE5C907B71FFCAFC003CE89D1C2E6B" authorityName="Chevrolat" authorityYear="1834" box="[926,1040,1014,1038]" class="Insecta" family="Melolonthidae" genus="Macrodactylus" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="582" phylum="Arthropoda" rank="species" species="lineatus">
<emphasis id="D68AFB017B71FFCAFC003CE89D1C2E6B" box="[926,1040,1014,1038]" italics="true" pageId="15" pageNumber="582">M. lineatus</emphasis>
</taxonomicName>
(red circle),
<taxonomicName id="23FE5C907B71FFCAFB0D3CE89C042E6B" authorityName="Bates" authorityYear="1887" box="[1171,1288,1015,1038]" class="Insecta" family="Melolonthidae" genus="Macrodactylus" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="582" phylum="Arthropoda" rank="species" species="nigripes">
<emphasis id="D68AFB017B71FFCAFB0D3CE89C042E6B" box="[1171,1288,1015,1038]" italics="true" pageId="15" pageNumber="582">M. nigripes</emphasis>
</taxonomicName>
(green circle),
<taxonomicName id="23FE5C907B71FFCAFF093B0498152E57" authorityName="Bates" authorityYear="1887" box="[151,281,1050,1074]" class="Insecta" family="Melolonthidae" genus="Macrodactylus" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="582" phylum="Arthropoda" rank="species" species="rufescens">
<emphasis id="D68AFB017B71FFCAFF093B0498152E57" box="[151,281,1050,1074]" italics="true" pageId="15" pageNumber="582">M. rufescens</emphasis>
</taxonomicName>
(blue circle),
<taxonomicName id="23FE5C907B71FFCAFE393B049B4E2E57" authorityName="Bates" authorityYear="1887" box="[423,578,1050,1074]" class="Insecta" family="Melolonthidae" genus="Macrodactylus" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="582" phylum="Arthropoda" rank="species" species="sericeicollis">
<emphasis id="D68AFB017B71FFCAFE393B049B4E2E57" box="[423,578,1050,1074]" italics="true" pageId="15" pageNumber="582">M. sericeicollis</emphasis>
</taxonomicName>
(yellow circle),
<taxonomicName id="23FE5C907B71FFCAFD743B049A7C2E57" authorityName="Arce-Perez y Moron" authorityYear="2010" box="[746,880,1051,1074]" class="Insecta" family="Melolonthidae" genus="Macrodactylus" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="582" phylum="Arthropoda" rank="species" species="zaragozai">
<emphasis id="D68AFB017B71FFCAFD743B049A7C2E57" box="[746,880,1051,1074]" italics="true" pageId="15" pageNumber="582">M. zaragozai</emphasis>
</taxonomicName>
(pink circle),
<taxonomicName id="23FE5C907B71FFCAFC613B049D892E57" box="[1023,1157,1050,1074]" class="Insecta" family="Melolonthidae" genus="Macrodactylus" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="582" phylum="Arthropoda" rank="species" species="zunilensis">
<emphasis id="D68AFB017B71FFCAFC613B049D892E57" box="[1023,1157,1050,1074]" italics="true" pageId="15" pageNumber="582">M. zunilensis</emphasis>
</taxonomicName>
(purple circle).
</paragraph>
</subSubSection>
</treatment>
</document>