treatments-xml/data/19/BC/4D/19BC4D641F675A5B9B4F22489B3563A5.xml

<document id="E11A4B9CA3F1FF24EA7097F84AA15F8A" ID-DOI="10.3897/zookeys.1209.125897" ID-publisher-id="125897" URI-arpha="6ED9C06E-4DBB-5A79-A98B-FE427E12ED58" URI-zoobank="http://zoobank.org/CEE78803-61EB-40CC-8D63-46142E6383A" XM.bibliography_approvedBy="admin" XM.materialsCitations_approvedBy="admin" XM.treatmentCitations_approvedBy="admin" article-type="research-article" checkinTime="1723117912200" checkinUser="pensoft" docAuthor="Harden, Curt W. &amp; Caterino, Michael S." docDate="2024" docId="19BC4D641F675A5B9B4F22489B3563A5" docLanguage="en" docName="ZooKeys 1209: 69-197" docOrigin="ZooKeys 1209" docSource="https://zookeys.pensoft.net/article/125897/download/xml/" docStyle="DocumentStyle:PensoftTaxPub.0000.journal_article.generic" docStyleName="PensoftTaxPub.0000.journal_article.generic" docTitle="Anillinus simplex Harden &amp; Caterino, 2024, sp. nov." docType="treatment" docUuid="CA863666-9208-48EC-8AB0-B2E006B1F0E9" docUuidSource="ZooBank" docVersion="3" dtd-version="3.0" lastPageNumber="197" masterDocId="6ED9C06E4DBB5A79A98BFE427E12ED58" masterDocTitle="Systematics and biogeography of Appalachian Anillini, and a taxonomic review of the species of South Carolina (Coleoptera, Carabidae, Trechinae, Anillini)" masterLastPageNumber="197" masterPageNumber="69" pageNumber="69" updateTime="1723146111653" updateUser="admin">
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<mods:title id="495ED5D934042F63E8EEEDB5A8AAFE33">Systematics and biogeography of Appalachian Anillini, and a taxonomic review of the species of South Carolina (Coleoptera, Carabidae, Trechinae, Anillini)</mods:title>
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<mods:namePart id="12EA21D5AE8B9FC7C5C22B9F32A35C4B">Harden, Curt W.</mods:namePart>
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<mods:affiliation id="C8E39AE794EF5519AA5A0BA826D0607E">Department of Plant and Environmental Sciences, Clemson University, 277 Poole Agricultural Center, Clemson, South Carolina, 29634, USA</mods:affiliation>
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<mods:namePart id="8E82C6C6DBA726BCE169CEF4C68482C2">Caterino, Michael S.</mods:namePart>
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<mods:affiliation id="116A477442319C422A7E4AC25415B528">Department of Plant and Environmental Sciences, Clemson University, 277 Poole Agricultural Center, Clemson, South Carolina, 29634, USA</mods:affiliation>
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<subSubSection id="5DAE6EB156960EE38D512EFC4E621AC0" type="nomenclature">
<paragraph id="AD24F080CB8997D170770272EB80D910">
<taxonomicName id="7AE81FD5E24FBE539F02312DA58EDB3D" authorityName="Harden &amp; Caterino" authorityYear="2024" class="Insecta" family="Carabidae" genus="Anillinus" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="simplex" status="sp. nov.">
<emphasis id="B316A58B4955BBA823123E6DAFB4D087" italics="true">Anillinus simplex</emphasis>
</taxonomicName>
<taxonomicNameLabel id="8FE3A86D5A12FE22D7A5260ADD0B008B" rank="species">sp. nov.</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection id="2B35248D19188310D50278C0415A5527" type="description">
<paragraph id="F59CD4B4027B6F76C05879E74772F9DD">
<figureCitation id="D13EA933F299A02E7770F89932ACCAE9" captionStart="Figure 21" captionStartId="F21" captionText="Figure 21. Spermathecae of Anillini species A, B Anillinus chandleri C Anillinus castaneus D Anillinus murrayae E Anillinus simplex F Anillinus dentatus G Anillinus sp. “ South Carolina, Coon Branch ” H Anillinus micamicus I Anillinus mica J Anillinus choestoea K Anillinus seneca L Anillinus montrex M Anillinus arenicollis N Anillinus jancae O Anillinus cf. nantahala P Anillinus cherokee Q Anillinus loweae R Anillinus merritti S Serranillus dunavani T Serranillus jeanneli. Scale bar: 0.1 mm." figureDoi="10.3897/zookeys.1209.125897.figure21" httpUri="https://binary.pensoft.net/fig/1110360">Figs 21 E</figureCitation>
, 
<figureCitation id="87390ED2FB0FD7A836307A5CD420DF87" captionStart="Figure 24" captionStartId="F24" captionText="Figure 24. Male ring sclerite, ventral aspect. A Anillinus chandleri B Anillinus castaneus C Anillinus sp. “ South Carolina, Chestnut Ridge ” D Anillinus murrayae E Anillinus simplex F Anillinus dentatus G Anillinus mica H Anillinus micamicus I Anillinus choestoea J Anillinus seneca K Anillinus arenicollis L Anillinus montrex M Anillinus jancae N Anillinus loweae O Anillinus cherokee P Anillinus merritti. Scale bar: 0.5 mm." figureDoi="10.3897/zookeys.1209.125897.figure24" httpUri="https://binary.pensoft.net/fig/1110363">24 E</figureCitation>
, 
<figureCitation id="133754517677660F9AAD67580A91A8C5" captionStart="Figure 29" captionStartId="F29" captionText="Figure 29. Distribution map of valentinei - group species in South Carolina and adjacent states. Data are from Harden (2024)." figureDoi="10.3897/zookeys.1209.125897.figure29" httpUri="https://binary.pensoft.net/fig/1110368">29</figureCitation>
, 
<figureCitation id="3B64A928688E36DC8196926D4C03A037" captionStart="Figure 30" captionStartId="F30" captionText="Figure 30. Aedeagi of valentinei group Anillinus species A – C Anillinus castaneus sp. nov. D – F Anillinus cornelli G – I Anillinus simplex sp. nov. J – N Anillinus murrayae. Median lobes C, F, I in right dorsolateral aspect, median lobe K in right lateral aspect. Left A, D, G, J and right B, E, H, L – N parameres in left and right lateral aspects, respectively. Scale bar: 0.1 mm." figureDoi="10.3897/zookeys.1209.125897.figure30" httpUri="https://binary.pensoft.net/fig/1110369">30 G – I</figureCitation>
, 
<figureCitation id="C5B13F87E7A8FF18F8F1678EE5D4DF73" captionStart="Figure 33" captionStartId="F33" captionText="Figure 33. Anillinus simplex sp. nov. A dorsal habitus, abdomen removed for DNA extraction B last abdominal ventrite, ventral aspect C head, left lateral aspect (black arrow = frontoclypeal horn). Scale bars: 1 mm (A); 0.1 mm (B)." figureDoi="10.3897/zookeys.1209.125897.figure33" httpUri="https://binary.pensoft.net/fig/1110372">33 A – C</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="SECID0EUNCM" type="type material">
<paragraph id="49BA7813FC77737717833C806BA259B4">
<heading id="A598A942A1E2529472B5572D08121572" reason="title">Type material.</heading>
</paragraph>
<paragraph id="5146650A0F4A91E74D94D5D2FE2C965E">
<materialsCitation id="0DFEEB1E5CCB665C7D50A7CA7AF21847" collectedFrom="Under rock near stream" collectingDate="2020-09-30" collectionCode="USNM" collectorName="C. W. Harden" country="United States of America" county="York Co." latitude="35.1307" location="Kings Mountain S. P." longLatPrecision="6" longitude="-81.3649" municipality="Original" specimenCode="CUAC 000185896" specimenCount="1" specimenCount-male="1" stateProvince="SC" typeStatus="holotype">
<emphasis id="A1243A11D9D1C71B17B4A9EA46120D78" bold="true" italics="true">
<typeStatus id="CE7E1B2E41B8C15E7DC317FE6E72F987" type="holotype">Holotype</typeStatus>
<specimenCount id="69BB9688EFB667F8EBF6AE4C11AC1650" type="male">male</specimenCount>
</emphasis>
: (
<named-content id="CA1014AEF65DB058D37A731DE5A25805" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/smithsonian-institution-national-museum-natural-history-0" xlink_title="United States National Museum of Natural History">
<emphasis id="4C065134ED6918BDA387B7674A41EE25" bold="true">
<collectionCode id="4A775EE32D8DB91AEBFD82EEF7A5C5E8" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" type="Museum">USNM</collectionCode>
</emphasis>
</named-content>
), point mounted, with abdominal ventrites glued to point and genitalia in Euparal on microslide pinned beneath specimen. Original label: “ 
<collectingCountry id="301846A537ECE6B9143CE9BE96D750B9" name="United States of America">USA</collectingCountry>
: 
<collectingRegion id="743933A328D3B54C6BD83184FF69AC2C">SC</collectingRegion>
, 
<collectingCounty id="4048BBF7F339822FEE39D763AA74406F">York Co.</collectingCounty>
<location id="D5AF67B02013412F8519E32EFDC2D215" LSID="urn:lsid:plazi:treatment:19BC4D641F675A5B9B4F22489B3563A5:D5AF67B02013412F8519E32EFDC2D215" country="United States of America" county="York Co." latitude="35.1307" longLatPrecision="6" longitude="-81.3649" municipality="Original" name="Kings Mountain S. P." stateProvince="SC">Kings Mountain S. P.</location>
<named-content content-type="dwc:verbatimCoordinates" id="NCID0ENOCM" specific-use="{&quot;type&quot;:&quot;Point&quot;,&quot;coordinates&quot;:[-81.364900,35.130700]}">
<geoCoordinate id="6E1345EE70AAAAF54F439E38D01B81D1" degrees="35.1307" direction="north" orientation="latitude" precision="5" value="35.1307">35.1307</geoCoordinate>
, 
<geoCoordinate id="5C591E71BF8C6E754CDB4676096B51AD" degrees="81.3649" direction="west" orientation="longitude" precision="5" value="-81.3649">- 81.3649</geoCoordinate>
</named-content>
. 26. September. 2020. 
<collectorName id="2F8C73F5FF0A93FF9EC30D0F561B5583">C. W. Harden</collectorName>
. 
<collectedFrom id="995AB9FB28844195536B0FBED04EA0F0">Under rock near stream</collectedFrom>
. ” “ [QR Code] CLEMSON-ENT 
<specimenCode id="CB58D20DFE857CABDC7CD7BC425ED979" collectionCode="CUAC" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:33168" lsid="urn:lsid:biocol.org:col:33168" name="Clemson University Arthropod Collection" type="University or college">CUAC 000185896</specimenCode>
” “ Harden DNA Voucher CWH- 247 Anill. ‘ kingsmtnsp 1 ’ M Ext. 
<date id="0B487A5099928153CE4AA0B75FCD4D0F" value="2020-09-30">
<collectingDate id="0AD3D407C764D30DD80DC460C49EA2AB" value="2020-09-30">30 / September / 2020</collectingDate>
</date>
[green-bordered label]. ” “ HOLOTYPE 
<taxonomicName id="FFA6479D814D023919220857ED0774CB" authority="Harden &amp; Caterino" authorityName="Harden &amp; Caterino" authorityYear="2024" class="Insecta" family="Carabidae" genus="Anillinus" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="simplex">
<originalDetermination id="7AB018CA581D56916315C9D377F2B28D">
<emphasis id="E0A227443C0FB3C01217A902D9A14787" italics="true">Anillinus simplex</emphasis>
Harden &amp; Caterino
</originalDetermination>
</taxonomicName>
[orange cardstock] ”
</materialsCitation>
</paragraph>
<paragraph id="D6A2F17B7626844BEE0588C2AC0F6A8D">
GenBank: 
<ext-link id="0C26BAE83F52DB9B2EC1EEC0F894D531" ext-link-type="gen" xlink_href="OR853342" xlink_type="simple">OR 853342</ext-link>
, 
<ext-link id="6A4ECF6F03E64A2FC1E579A1248272AA" ext-link-type="gen" xlink_href="OR839331" xlink_type="simple">OR 839331</ext-link>
, 
<ext-link id="B74F0610EAE40D5D611A1269BDBC7264" ext-link-type="gen" xlink_href="OR839570" xlink_type="simple">OR 839570</ext-link>
, 
<ext-link id="555277911089947A21539BA1FF3AD257" ext-link-type="gen" xlink_href="OR837888" xlink_type="simple">OR 837888</ext-link>
, 
<ext-link id="3E75CB71C9975D38CEC35D8968B7FC84" ext-link-type="gen" xlink_href="OR838032" xlink_type="simple">OR 838032</ext-link>
, 
<ext-link id="F228D49A621A705CFAEC4EFDE591779E" ext-link-type="gen" xlink_href="OR838222" xlink_type="simple">OR 838222</ext-link>
.
</paragraph>
<paragraph id="465139A704B1717800A6579A8C5CDDEF">
<emphasis id="85DFDC4F7F8FDD8691234C0F82228C62" bold="true" italics="true">
<typeStatus id="68CFE45BCFEA60CFD075A3E8302AD37C" type="paratype">Paratypes</typeStatus>
</emphasis>
(
<emphasis id="04D9200F90B2C6E999D184FF72545E64" italics="true">n</emphasis>
= 46, 
<abbrev id="ABBRID0EGQCM" xlink_title="Augusto DeGiovanni personal collection, Bologna, Italy">
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</abbrev>
, 
<named-content id="361405F9A72169F56B031BEEE6E9FF78" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/carnegie-museum-natural-history" xlink_title="Carnegie Museum of Natural History">
<collectionCode id="F4E4B363AB7C155EFC2C929E8309E1F8" country="USA" httpUri="http://grbio.org/cool/daw7-8h44" name="The Cleveland Museum of Natural History" type="Museum">CMNH</collectionCode>
</named-content>
, 
<named-content id="63C8A412A02FBC9F735F4572983F43CE" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/canadian-national-collection-insects-arachnids-and-nematodes" xlink_title="Canadian National Collection of Insects">
<collectionCode id="6D8F32BE6FF4F481D41E5145079C5D94" country="Canada" httpUri="http://grbio.org/cool/y2kv-9w3k" name="Canadian National Collection of Insects, Arachnids, and Nematodes" type="">CNC</collectionCode>
</named-content>
, 
<named-content id="C0ADEDF4ABD244AC8889D2F2EACC1E2D" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/clemson-university-arthropod-collection" xlink_title="Clemson University Arthropod Collection">
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</named-content>
, 
<named-content id="8174DB3DDF09EAAC92180E0431E2B4F8" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/florida-state-collection-arthropods-museum-entomology" xlink_title="Florida State Collection of Arthropods">
<collectionCode id="977864F141B447F849E14ECF32DA1B65" country="USA" httpUri="http://grbio.org/cool/jekt-x82m" name="Florida State Collection of Arthropods, The Museum of Entomology" type="Museum">FSCA</collectionCode>
</named-content>
, 
<named-content id="FE10DE7E508FDEF416B2002093D8B2D3" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/louisiana-state-arthropod-museum" xlink_title="Louisiana State Arthropod Museum">
<collectionCode id="AE45F531EA54E3D5AEF65D8327E0B4A9" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:33726" lsid="urn:lsid:biocol.org:col:33726" name="Louisiana State Arthropod Museum" type="">LSAM</collectionCode>
</named-content>
, 
<named-content id="AF140619C7B59E34C5C07690445BC45D" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/north-carolina-state-university-insect-museum" xlink_title="North Carolina State University">
<collectionCode id="F78738FCB8DF3A6F4C11D25827721C3F" country="USA" httpUri="http://grbio.org/cool/ij62-iybb" name="North Carolina State University Insect Museum" type="Museum">NCSU</collectionCode>
</named-content>
, 
<named-content id="B63E8AAF20E6760A6517475F0F237E48" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/natural-history-museum-london" xlink_title="The Natural History Museum, London">
<collectionCode id="4557E3E6A4063FB470E0CA7024F7B8FB" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34665" lsid="urn:lsid:biocol.org:col:34665" name="Natural History Museum, London" type="Museum">NHMUK</collectionCode>
</named-content>
, 
<named-content id="06E023E4C2B512067C8A18B95C707070" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/oregon-state-arthropod-collection" xlink_title="Oregon State Arthropod Collection">
<collectionCode id="B8AE5CC19B3511F30486729AEDA41A07" country="USA" httpUri="http://grbio.org/cool/vn0n-jrfa" name="Oregon State Arthropod Collection" type="University or college">OSAC</collectionCode>
</named-content>
, 
<named-content id="89DADAEBAB748BE507DC55E12509ABB1" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/smithsonian-institution-national-museum-natural-history-0" xlink_title="United States National Museum of Natural History">
<collectionCode id="2B361DE4EBF021F6A02A0AEA29357A76" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" type="Museum">USNM</collectionCode>
</named-content>
, 
<named-content id="ABF56BBBE2BDABD1289FD40A48F46B7B" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/virginia-museum-natural-history" xlink_title="Virginia Museum of Natural History">
<collectionCode id="7B79706B0A6663272DF739450ECBE6D7" country="USA" httpUri="http://grbio.org/cool/zxpe-c0as" name="Virginia Museum of Natural History" type="">VMNH</collectionCode>
</named-content>
). 
<materialsCitation id="2EFD8C33B542C995505523D08394B550" collectedFrom="Under rock near stream" collectingDate="2020-09-30" collectionCode="CUAC" collectorName="York Co. Kings Mountain S. P. &amp; C. W. Harden. Under" country="United States of America" county="York Co." latitude="35.1307" location="Kings Mountain State Park" longLatPrecision="6" longitude="-81.3649" municipality="Kings Mountain State Park" specimenCode="CUAC 000170024" specimenCount="1" specimenCount-male="1" stateProvince="South Carolina" typeStatus="paratype">
<emphasis id="E5D82B47D03446D6B0AA6DEBB83669AF" bold="true">
<collectingCountry id="DA9F04D45E5A40AA947C0E4B9825A2DC" name="United States of America">USA</collectingCountry>
</emphasis>
• 
<emphasis id="2CA89D4EA11CEF3665AE75CA65C1B1A7" bold="true">
<collectingRegion id="CD9D750C8E88D9D02C295064187C7BF2" country="United States of America" name="South Carolina">South Carolina</collectingRegion>
</emphasis>
• 
<emphasis id="79C2CE3FA5C13CE57EB2320605FEDFC0" bold="true">
<collectingCounty id="641DE4FF96701B13ACFED9C18A9EEAAD">York Co.</collectingCounty>
</emphasis>
• 
<collectingMunicipality id="DA0A05A7AD15952E531AB66640E01EAF" country="United States of America" county="York Co." name="Kings Mountain State Park" stateProvince="South Carolina">Kings Mountain State Park</collectingMunicipality>
; • 
<specimenCount id="EEACB34C6A19FED6C875243C24144871" count="1" type="male">1 ♂</specimenCount>
; same data as holotype; 
<specimenCode id="F893743B919E50F207FEE4CBACE35CD5" collectionCode="CUAC" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:33168" lsid="urn:lsid:biocol.org:col:33168" name="Clemson University Arthropod Collection" type="University or college">CUAC 000170024</specimenCode>
, CWH- 248; 
<named-content id="1C86CA79C6B5DBC73B27F7F9EE6CAD38" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/clemson-university-arthropod-collection" xlink_title="Clemson University Arthropod Collection">
<collectionCode id="4E201C2B7D61F5F00C213320D52FECFC" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:33168" lsid="urn:lsid:biocol.org:col:33168" name="Clemson University Arthropod Collection" type="University or college">CUAC</collectionCode>
</named-content>
</materialsCitation>
; • 
<materialsCitation id="E21932546C5B657F3DA376F4EAC15FFC" collectedFrom="Under rock near stream" collectingDate="2020-09-30" collectionCode="CUAC" collectorName="York Co. Kings Mountain S. P. &amp; C. W. Harden. Under" country="United States of America" county="York Co." latitude="35.1307" location="Kings Mountain S. P." longLatPrecision="6" longitude="-81.3649" municipality="Original" specimenCode="CUAC 000170025" specimenCount="1" specimenCount-female="1" stateProvince="South Carolina" typeStatus="paratype">
<specimenCount id="FAD770A6079FBADB2101212369930FAA" count="1" type="female">1 ♀</specimenCount>
; same data as holotype; 
<specimenCode id="4E1641A8FD18FBE4630316C3606545E3" collectionCode="CUAC" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:33168" lsid="urn:lsid:biocol.org:col:33168" name="Clemson University Arthropod Collection" type="University or college">CUAC 000170025</specimenCode>
, CWH- 249; 
<named-content id="0A554A64566E14F3648C3D22E37FE8C2" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/clemson-university-arthropod-collection" xlink_title="Clemson University Arthropod Collection">
<collectionCode id="8AEF0D74000FB656545D158AECF10CFD" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:33168" lsid="urn:lsid:biocol.org:col:33168" name="Clemson University Arthropod Collection" type="University or college">CUAC</collectionCode>
</named-content>
</materialsCitation>
; • 
<materialsCitation id="364DD0E6410520768A067D44F18C6297" collectedFrom="Under rock near stream" collectingDate="2020-09-30" collectionCode="CNC, CUAC" collectorName="York Co. Kings Mountain S. P. &amp; C. W. Harden. Under" country="United States of America" county="York Co." latitude="35.1307" location="Kings Mountain S. P." longLatPrecision="6" longitude="-81.3649" municipality="Original" specimenCode="CUAC 000170026, CUAC 000170035" specimenCount="10" specimenCount-female="2" specimenCount-male="8" stateProvince="South Carolina" typeStatus="paratype">
<specimenCount id="4F8E5FDFDB87600CE696E3F3ED651D9F" count="8" type="male">8 ♂</specimenCount>
, 
<specimenCount id="C77D4E9793A00D65C9E9287AA4684B7B" count="2" type="female">2 ♀</specimenCount>
; same data as holotype; 
<specimenCode id="BC800E593ECD18E0E00D681D9037470E" collectionCode="CUAC" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:33168" lsid="urn:lsid:biocol.org:col:33168" name="Clemson University Arthropod Collection" type="University or college">CUAC 000170026</specimenCode>
to 
<specimenCode id="B67E78D0F0D8C991D7B54802F556337E" collectionCode="CUAC" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:33168" lsid="urn:lsid:biocol.org:col:33168" name="Clemson University Arthropod Collection" type="University or college">CUAC 000170035</specimenCode>
; 
<named-content id="535F5404E736273D7C7B4321BACDFFEC" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/canadian-national-collection-insects-arachnids-and-nematodes" xlink_title="Canadian National Collection of Insects">
<collectionCode id="11564CDA3B4D460D9537D793B6B705A2" country="Canada" httpUri="http://grbio.org/cool/y2kv-9w3k" name="Canadian National Collection of Insects, Arachnids, and Nematodes" type="">CNC</collectionCode>
</named-content>
, 
<named-content id="D7060205241CC86AAE22AC138D2E0033" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/clemson-university-arthropod-collection" xlink_title="Clemson University Arthropod Collection">
<collectionCode id="C97F1F36D400DD58CAA4D3B4C52BF660" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:33168" lsid="urn:lsid:biocol.org:col:33168" name="Clemson University Arthropod Collection" type="University or college">CUAC</collectionCode>
</named-content>
</materialsCitation>
; • 
<materialsCitation id="E465526A3031B460C6F4AE7205CD837F" collectedFrom="Under rock near stream" collectingDate="2020-09-30" collectionCode="ADGc" collectorName="York Co. Kings Mountain S. P. &amp; C. W. Harden. Under" country="United States of America" county="York Co." latitude="35.1307" location="Kings Mountain S. P." longLatPrecision="6" longitude="-81.3649" municipality="Original" specimenCount="3" specimenCount-male="3" stateProvince="South Carolina" typeStatus="paratype">
<specimenCount id="DF74E203ADB8E8668C94B2B7E7CC265C" count="3" type="male">3 ♂</specimenCount>
; same data as holotype; 
<abbrev id="ABBRID0EISCM" xlink_title="Augusto DeGiovanni personal collection, Bologna, Italy">
<collectionCode id="0A39E37C14842D41405284D4B7E35447">ADGc</collectionCode>
</abbrev>
</materialsCitation>
; • 
<materialsCitation id="9191599B7EB7171D5E0A84D624ACA7F3" collectedFrom="deep litter and topsoil, oak, maple, sourwood, pine" collectingDate="2020-09-26" collectingMethod="Berlese" collectionCode="CMNH, CNC, FSCA, LSAM, OSAC, USNM, VMNH" collectorName="C. W. Harden" country="United States of America" county="York Co." latitude="35.13198" longLatPrecision="1" longitude="-81.36608" specimenCode="CUAC 000170036, CUAC 000170059" specimenCount="24" specimenCount-female="12" specimenCount-male="12" stateProvince="South Carolina" typeStatus="paratype">
<specimenCount id="F6458C208BE4D0A9FE7E8E6D0390904F" count="12" type="male">12 ♂</specimenCount>
, 
<specimenCount id="7759CE597E74E2E5FEF17B45D30AF5F8" count="12" type="female">12 ♀</specimenCount>
; 
<named-content content-type="dwc:verbatimCoordinates" id="NCID0EPSCM" specific-use="{&quot;type&quot;:&quot;Point&quot;,&quot;coordinates&quot;:[-81.366080,35.131980]}">
<geoCoordinate id="2A83E33813DA1E51EC098ED93EB13C66" degrees="35.13198" direction="north" orientation="latitude" precision="1" value="35.13198">35.13198</geoCoordinate>
, 
<geoCoordinate id="8D765693A4D9CCA01F06FA3CBEAD7E1C" degrees="81.36608" direction="west" orientation="longitude" precision="1" value="-81.36608">- 81.36608</geoCoordinate>
</named-content>
; 
<date id="2D6AE541751653C0F59605027DA7F08E" value="2020-09-26">
<collectingDate id="61BB7854D94F83E4EC35AA256A608BC7" value="2020-09-26">26 Sep. 2020</collectingDate>
</date>
; 
<collectorName id="E7323E6747B543FF0056DA611446302E">C. W. Harden</collectorName>
leg.; 
<collectingMethod id="0FDD3BBDD0C8B1E9AEF9BA58A4D9D14B">Berlese</collectingMethod>
, 
<collectedFrom id="282A5CDA5A3AFAEEEA7FD3653D7A92DE">deep litter and topsoil, oak, maple, sourwood, pine</collectedFrom>
; 
<specimenCode id="509B6DF80C430BBEDABA9C95760DC059" collectionCode="CUAC" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:33168" lsid="urn:lsid:biocol.org:col:33168" name="Clemson University Arthropod Collection" type="University or college">CUAC 000170036</specimenCode>
to 
<specimenCode id="2D241EFCE514F46932FDCA704EC5B8FE" collectionCode="CUAC" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:33168" lsid="urn:lsid:biocol.org:col:33168" name="Clemson University Arthropod Collection" type="University or college">CUAC 000170059</specimenCode>
; 
<named-content id="A50DDF1159C36621B3E0062C3F693C59" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/carnegie-museum-natural-history" xlink_title="Carnegie Museum of Natural History">
<collectionCode id="3F4FB09C935DA2C49137906E93F4328D" country="USA" httpUri="http://grbio.org/cool/daw7-8h44" name="The Cleveland Museum of Natural History" type="Museum">CMNH</collectionCode>
</named-content>
, 
<named-content id="40663272C83376FE869A01D4358B6DA8" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/canadian-national-collection-insects-arachnids-and-nematodes" xlink_title="Canadian National Collection of Insects">
<collectionCode id="9EDF7C55B8FE865CAE352C6C0FCEFD31" country="Canada" httpUri="http://grbio.org/cool/y2kv-9w3k" name="Canadian National Collection of Insects, Arachnids, and Nematodes" type="">CNC</collectionCode>
</named-content>
, 
<named-content id="4F5F82A70289C18EB295E4C1BBF01B93" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/florida-state-collection-arthropods-museum-entomology" xlink_title="Florida State Collection of Arthropods">
<collectionCode id="09661621D39506E02FC5BB6DE41DC7FA" country="USA" httpUri="http://grbio.org/cool/jekt-x82m" name="Florida State Collection of Arthropods, The Museum of Entomology" type="Museum">FSCA</collectionCode>
</named-content>
, 
<named-content id="7F17197A4BA44C75225ED6C139AA8B55" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/louisiana-state-arthropod-museum" xlink_title="Louisiana State Arthropod Museum">
<collectionCode id="60AD089091BB54DA1FB06A12D8E80319" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:33726" lsid="urn:lsid:biocol.org:col:33726" name="Louisiana State Arthropod Museum" type="">LSAM</collectionCode>
</named-content>
, 
<named-content id="A6E6064A1D883B21FE5A29F05E95AEFF" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/oregon-state-arthropod-collection" xlink_title="Oregon State Arthropod Collection">
<collectionCode id="F263CD447D7CD13EEB32267597666893" country="USA" httpUri="http://grbio.org/cool/vn0n-jrfa" name="Oregon State Arthropod Collection" type="University or college">OSAC</collectionCode>
</named-content>
, 
<named-content id="07E5795034AC358D4A85967262B1D249" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/smithsonian-institution-national-museum-natural-history-0" xlink_title="United States National Museum of Natural History">
<collectionCode id="1923AA243B257DB2299CCDACF41A67D6" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" type="Museum">USNM</collectionCode>
</named-content>
, 
<named-content id="575FD9D39F139FFCC32AD2439F90B2B0" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/virginia-museum-natural-history" xlink_title="Virginia Museum of Natural History">
<collectionCode id="3779B2F3BA4CEE505ED6E67ED9474A67" country="USA" httpUri="http://grbio.org/cool/zxpe-c0as" name="Virginia Museum of Natural History" type="">VMNH</collectionCode>
</named-content>
</materialsCitation>
; • 
<materialsCitation id="63FCA0741A0DA200288D046E2D676733" collectedFrom="litter near stream, mesic hardwoods" collectingDate="2020-09-26" collectingMethod="Berlese" collectionCode="VMNH" collectorName="C. W. Harden" country="United States of America" county="York Co." latitude="35.13062" longLatPrecision="1" longitude="-81.36439" specimenCode="CUAC 00017060" specimenCount="1" specimenCount-female="1" stateProvince="South Carolina" typeStatus="paratype">
<specimenCount id="7226237FC217A31FBAF0184D75DDEB08" count="1" type="female">1 ♀</specimenCount>
; 
<named-content content-type="dwc:verbatimCoordinates" id="NCID0ETTCM" specific-use="{&quot;type&quot;:&quot;Point&quot;,&quot;coordinates&quot;:[-81.364390,35.130620]}">
<geoCoordinate id="4B5C3370D45D54476364D04FD845E3DF" degrees="35.13062" direction="north" orientation="latitude" precision="1" value="35.13062">35.13062</geoCoordinate>
, 
<geoCoordinate id="8C46DDA0A2EDF2DE204EB4346947367E" degrees="81.36439" direction="west" orientation="longitude" precision="1" value="-81.36439">- 81.36439</geoCoordinate>
</named-content>
; 
<date id="4D1338D008874E048692022C27EA548D" value="2020-09-26">
<collectingDate id="BEBC5F9652D4ED9170579058E7ABB568" value="2020-09-26">26 Sep. 2020</collectingDate>
</date>
; 
<collectorName id="48B558CAC0E1E6FB3CF2EEBECD8AAF3D">C. W. Harden</collectorName>
leg.; 
<collectingMethod id="C669B5F8C9328D72545D33A1D3978031">Berlese</collectingMethod>
, 
<collectedFrom id="2CCD226846C44D1830E7001897006B27">litter near stream, mesic hardwoods</collectedFrom>
; 
<specimenCode id="A5ED9B98CA7A466182D535379FB7E7B3" collectionCode="CUAC" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:33168" lsid="urn:lsid:biocol.org:col:33168" name="Clemson University Arthropod Collection" type="University or college">CUAC 00017060</specimenCode>
; 
<named-content id="9DBC926F00EDD3218258FFAF79C2FB94" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/virginia-museum-natural-history" xlink_title="Virginia Museum of Natural History">
<collectionCode id="70D7C1BD052E0EC9E9E21894E3B4F291" country="USA" httpUri="http://grbio.org/cool/zxpe-c0as" name="Virginia Museum of Natural History" type="">VMNH</collectionCode>
</named-content>
</materialsCitation>
; • 
<materialsCitation id="97E25E41ABA5D5FDA6EB061CFD74AF9E" collectedFrom="under embedded rock" collectingDate="2021-12-23" collectionCode="NCSU" collectorName="C. W. Harden" country="United States of America" county="York Co." latitude="35.13018" longLatPrecision="1" longitude="-81.36205" specimenCode="CUAC 000167147, CUAC 000167149" specimenCount="3" specimenCount-female="1" specimenCount-male="2" stateProvince="South Carolina" typeStatus="paratype">
<specimenCount id="A1E7E86342B7D1B87A5BC637F82649CE" count="2" type="male">2 ♂</specimenCount>
, 
<specimenCount id="DE586F7F768C7513B63A47E9CBC055B1" count="1" type="female">1 ♀</specimenCount>
; 
<named-content content-type="dwc:verbatimCoordinates" id="NCID0E6TCM" specific-use="{&quot;type&quot;:&quot;Point&quot;,&quot;coordinates&quot;:[-81.362050,35.130180]}">
<geoCoordinate id="9E79DC484EE59676CC15067228E2F4C3" degrees="35.13018" direction="north" orientation="latitude" precision="1" value="35.13018">35.13018</geoCoordinate>
, 
<geoCoordinate id="2DD7491090385BB34D92E4118F22329E" degrees="81.36205" direction="west" orientation="longitude" precision="1" value="-81.36205">- 81.36205</geoCoordinate>
</named-content>
; 
<date id="B1E1FE74FC9DFA5FB593D7E124AB6DB3" value="2021-12-23">
<collectingDate id="B935D92E7E003F6C0543047800C6A644" value="2021-12-23">23 Dec. 2021</collectingDate>
</date>
; 
<collectorName id="696EF71AB6F634A373CF4761E554C635">C. W. Harden</collectorName>
leg.; 
<collectedFrom id="C8DA285581518FD41DC212456049E2E2">under embedded rock</collectedFrom>
; 
<specimenCode id="6D3BBB3000F61A1ED8B83185D60ECD16" collectionCode="CUAC" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:33168" lsid="urn:lsid:biocol.org:col:33168" name="Clemson University Arthropod Collection" type="University or college">CUAC 000167147</specimenCode>
to 
<specimenCode id="CB7669B7F88D0B072525E396937483DA" collectionCode="CUAC" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:33168" lsid="urn:lsid:biocol.org:col:33168" name="Clemson University Arthropod Collection" type="University or college">CUAC 000167149</specimenCode>
; 
<named-content id="4A8A2F3B673E9E84F0459AE8E1552C55" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/north-carolina-state-university-insect-museum" xlink_title="North Carolina State University">
<collectionCode id="101B765571AD5C1F0FD540E108C94192" country="USA" httpUri="http://grbio.org/cool/ij62-iybb" name="North Carolina State University Insect Museum" type="Museum">NCSU</collectionCode>
</named-content>
</materialsCitation>
; • 
<materialsCitation id="D7A6A3918DECFFE88F5FB1DCB662B9B9" collectedFrom="soil near seepage" collectingDate="2021-12-23" collectingMethod="Berlese" collectionCode="NHMUK" collectorName="C. W. Harden" country="United States of America" county="York Co." latitude="35.1301" longLatPrecision="6" longitude="-81.3637" specimenCode="CUAC 000170061, CUAC 000170063" specimenCount="3" specimenCount-female="2" specimenCount-male="1" stateProvince="South Carolina" typeStatus="paratype">
<specimenCount id="E82F8FDAD083E2207A70295BEFB8DD0A" count="1" type="male">1 ♂</specimenCount>
, 
<specimenCount id="4071AB716F8F40C2D04F824FCF5AB133" count="2" type="female">2 ♀</specimenCount>
; 
<named-content content-type="dwc:verbatimCoordinates" id="NCID0ELUCM" specific-use="{&quot;type&quot;:&quot;Point&quot;,&quot;coordinates&quot;:[-81.363700,35.130100]}">
<geoCoordinate id="DF7CA5F251106FC519EF098D16F4636F" degrees="35.1301" direction="north" orientation="latitude" precision="5" value="35.1301">35.1301</geoCoordinate>
, 
<geoCoordinate id="5DB93B5EF0DEA3C9861B0A5C631B4500" degrees="81.3637" direction="west" orientation="longitude" precision="5" value="-81.3637">- 81.3637</geoCoordinate>
</named-content>
; 
<date id="CC730B8FFAD452365AAC727329598450" value="2021-12-23">
<collectingDate id="4F8208CD51FC041AE09F248A3D145AE0" value="2021-12-23">23 Dec. 2021</collectingDate>
</date>
; 
<collectorName id="C8504800CB881184E0675C6A2D365E72">C. W. Harden</collectorName>
leg.; 
<collectingMethod id="CF6634073FB041A71B1FE41BD9D36CE8">Berlese</collectingMethod>
, 
<collectedFrom id="7D3C737984235D23FFAC48AA85C689D2">soil near seepage</collectedFrom>
; 
<specimenCode id="68C312BFC6B377370B34C8389DB07694" collectionCode="CUAC" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:33168" lsid="urn:lsid:biocol.org:col:33168" name="Clemson University Arthropod Collection" type="University or college">CUAC 000170061</specimenCode>
to 
<specimenCode id="ABF940DF8C34239B51DF1A32E5FD3588" collectionCode="CUAC" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:33168" lsid="urn:lsid:biocol.org:col:33168" name="Clemson University Arthropod Collection" type="University or college">CUAC 000170063</specimenCode>
; 
<named-content id="7679E72D11D23CFA7DFC036561B21546" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/natural-history-museum-london" xlink_title="The Natural History Museum, London">
<collectionCode id="6CEFF0B581767A24D062C68FD793866B" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34665" lsid="urn:lsid:biocol.org:col:34665" name="Natural History Museum, London" type="Museum">NHMUK</collectionCode>
</named-content>
</materialsCitation>
.
</paragraph>
<paragraph id="D5E5E54CEA4767497838D82A5FE60EBC">
GenBank accession numbers for 
<typeStatus id="73F03A679D9D251F819DC9996AD029FC" type="paratype">paratypes</typeStatus>
: 
<ext-link id="83295EC40D3C0CFDCDE8C74435C1EB0D" ext-link-type="gen" xlink_href="OR839571" xlink_type="simple">OR 839571</ext-link>
, 
<ext-link id="6076E64DA7452414F4D1A52538B512CC" ext-link-type="gen" xlink_href="OR853339" xlink_type="simple">OR 853339</ext-link>
, 
<ext-link id="43965C1A41BEDA9520E1444DAE237E32" ext-link-type="gen" xlink_href="OR839328" xlink_type="simple">OR 839328</ext-link>
, 
<ext-link id="C0C8317A601837F0472A3980058A2952" ext-link-type="gen" xlink_href="OR839572" xlink_type="simple">OR 839572</ext-link>
, 
<ext-link id="00B0DBDF232B29736CB1DEFB3A6CB460" ext-link-type="gen" xlink_href="OR837889" xlink_type="simple">OR 837889</ext-link>
, 
<ext-link id="2DC36D07582D269091B32EDA3E09EBA7" ext-link-type="gen" xlink_href="OR838033" xlink_type="simple">OR 838033</ext-link>
, 
<ext-link id="4DB3679DB5DF37E8D1CCEF1EB9CB0F7F" ext-link-type="gen" xlink_href="OR838223" xlink_type="simple">OR 838223</ext-link>
.
</paragraph>
</subSubSection>
<subSubSection id="SECID0EZVCM" type="material">
<paragraph id="BB6A6BACD9DBFAB1F853CB2392FAC4D8">
<heading id="11D97B0D2F5826EF9C8E703642E5EB87" reason="title">Other material</heading>
</paragraph>
<paragraph id="425A02DD6BBAD849E85FBBA1CE4ECAF9">
<emphasis id="CE7F85331262A73F2AA9D627F7EC3EB3" bold="true">
(
<emphasis id="4BA995E4744E43A3055556DB6CF10A6A" bold="true" italics="true">n</emphasis>
= 30, 
<named-content id="66880E7283DF48F7C557C1925D008466" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/clemson-university-arthropod-collection" xlink_title="Clemson University Arthropod Collection">
<collectionCode id="9101DB12087675FFFE740DE6A17DAE0B" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:33168" lsid="urn:lsid:biocol.org:col:33168" name="Clemson University Arthropod Collection" type="University or college">CUAC</collectionCode>
</named-content>
, 
<abbrev id="ABBRID0EHWCM" xlink_title="Curt Harden personal collection, Central, SC, USA">
<collectionCode id="702EDAF23C89BACF39E538ADE5321B09">CWHc</collectionCode>
</abbrev>
, 
<named-content id="1870BCE7A1AC310BCAD3C0319592E583" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/north-carolina-state-university-insect-museum" xlink_title="North Carolina State University">
<collectionCode id="4F7D178E41AE88C9A28495DEEC7AA22A" country="USA" httpUri="http://grbio.org/cool/ij62-iybb" name="North Carolina State University Insect Museum" type="Museum">NCSU</collectionCode>
</named-content>
). 
<materialsCitation id="6A77EB5718F279AA6EAA147A7328534F" collectedFrom="sifted duff / litter" collectingDate="2023-04-29" collectionCode="NCSU" collectorName="C. W. Harden &amp; Berlese" country="United States of America" county="Gaston Co." latitude="35.2417" location="near Linwood parking" longLatPrecision="6" longitude="-81.2717" municipality="Crowders Mountain State Park" specimenCount="8" specimenCount-female="5" specimenCount-male="3" stateProvince="North Carolina">
<collectingCountry id="C7DF059F91E7A4CCB0F7FC713855E9D3" name="United States of America">USA</collectingCountry>
• 
<emphasis id="E256AE83C0987B5F38AC55E5569696F2" bold="true">
<collectingRegion id="D483A94B7DB0FF4B8A87B042013CF227" country="United States of America" name="North Carolina">North Carolina</collectingRegion>
</emphasis>
• 
<specimenCount id="7295C72AFC8FE7213169B650BA84146E" count="3" type="male">3 ♂</specimenCount>
, 
<specimenCount id="FBDC25415376FF49713CFFADCC574ED1" count="5" type="female">5 ♀</specimenCount>
; 
<emphasis id="44A49D37B03B96866A7A95E942EBC6E9" bold="true">
<collectingCounty id="743C5DC66B649397A35342D3DF0F56C3">Gaston Co.</collectingCounty>
</emphasis>
; 
<collectingMunicipality id="573EA83C009B3D66BB05E610004E5C08">Crowders Mountain State Park</collectingMunicipality>
, 
<location id="0B16329BD750A66930221F83C2887854" LSID="urn:lsid:plazi:treatment:19BC4D641F675A5B9B4F22489B3563A5:0B16329BD750A66930221F83C2887854" country="United States of America" county="Gaston Co." latitude="35.2417" longLatPrecision="6" longitude="-81.2717" municipality="Crowders Mountain State Park" name="near Linwood parking" stateProvince="North Carolina">near Linwood parking</location>
; 
<named-content content-type="dwc:verbatimCoordinates" id="NCID0EXWCM" specific-use="{&quot;type&quot;:&quot;Point&quot;,&quot;coordinates&quot;:[-81.271700,35.241700]}">
<geoCoordinate id="9C3112505D94831BF3811EC37C3BFC27" degrees="35.2417" direction="north" orientation="latitude" precision="5" value="35.2417">35.2417</geoCoordinate>
, 
<geoCoordinate id="D7AEEB4E1D5EE6103EC4215427DAB74D" degrees="81.2717" direction="west" orientation="longitude" precision="5" value="-81.2717">- 81.2717</geoCoordinate>
</named-content>
; 
<date id="A9DC1CFC3CC10EF695179BB681631067" value="2023-04-29">
<collectingDate id="2052742A6FADF86DF8A0543AFC74136B" value="2023-04-29">29 Apr. 2023</collectingDate>
</date>
; 
<collectorName id="DC6B518E38029851059CA83BE9F63CF7">C. W. Harden</collectorName>
leg.; 
<collectorName id="3FDAABCF537A6A24CC79D48F3E11A612">Berlese</collectorName>
, 
<collectedFrom id="67AFD172D7C9E59902FEFB6858A275EB">sifted duff / litter</collectedFrom>
; 
<named-content id="F998ED6ABD0066AAA38C854129A4450F" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/north-carolina-state-university-insect-museum" xlink_title="North Carolina State University">
<collectionCode id="2BFFD230A7078ABE2B9E68639EE75607" country="USA" httpUri="http://grbio.org/cool/ij62-iybb" name="North Carolina State University Insect Museum" type="Museum">NCSU</collectionCode>
</named-content>
</materialsCitation>
</emphasis>
; • 
<materialsCitation id="DE714D6349299960DBF1E38ED5A3C478" collectedFrom="under embedded rock" collectingDate="2023-04-29" collectionCode="CUAC" collectorName="C. W. Harden &amp; Berlese" country="United States of America" county="Gaston Co." latitude="35.2417" location="Linwood" longLatPrecision="6" longitude="-81.2717" municipality="Crowders Mountain State Park" specimenCode="CUAC 000182312, CUAC 000182313" specimenCount="2" specimenCount-male="2" stateProvince="North Carolina">
<specimenCount id="812B8ABB64C424C9740395B48CF5F7A2" count="2" type="male">2 ♂</specimenCount>
; same data as previous; 
<collectedFrom id="3F3502CD5A92FD6E83D9C72AF643A086">under embedded rock</collectedFrom>
; 
<specimenCode id="F7C0E9D429DC09C33E353278C6767663" collectionCode="CUAC" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:33168" lsid="urn:lsid:biocol.org:col:33168" name="Clemson University Arthropod Collection" type="University or college">CUAC 000182312</specimenCode>
and 
<specimenCode id="C00F71FAB38B19325C7411B05260CD30" collectionCode="CUAC" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:33168" lsid="urn:lsid:biocol.org:col:33168" name="Clemson University Arthropod Collection" type="University or college">CUAC 000182313</specimenCode>
, CWH- 510 and CWH- 511; 
<named-content id="58D6E378A6BD8908D81A4F4C1CB035C9" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/clemson-university-arthropod-collection" xlink_title="Clemson University Arthropod Collection">
<collectionCode id="2699C47EEF8A0FEBD480C1684C99D495" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:33168" lsid="urn:lsid:biocol.org:col:33168" name="Clemson University Arthropod Collection" type="University or college">CUAC</collectionCode>
</named-content>
</materialsCitation>
; • 
<materialsCitation id="AA9E397233BFAD0B28A635BD3C533987" collectedFrom="under embedded rock" collectingDate="2023-04-29" collectionCode="CUAC" collectorName="C. W. Harden" country="United States of America" county="Gaston Co." latitude="35.1977" location="Ridgeline Trail at Road 1104" longLatPrecision="6" longitude="-81.3152" municipality="Crowders Mountain State Park" specimenCode="CUAC 000182314" specimenCount="1" specimenCount-male="1" stateProvince="North Carolina">
<specimenCount id="FED17C72425F19DCF949C3500232517B" count="1" type="male">1 ♂</specimenCount>
; 
<emphasis id="423CA010F09FA474B28859D48E907466" bold="true">
<collectingCounty id="7D46A39BA158BBD1335F648A832EA40B">Gaston Co.</collectingCounty>
</emphasis>
; 
<collectingMunicipality id="B449C73C3BD80931C1C79A1EFD8F5512">Crowders Mountain State Park</collectingMunicipality>
; 
<location id="3420D4E7580BF429078C8C8B8E589DBD" LSID="urn:lsid:plazi:treatment:19BC4D641F675A5B9B4F22489B3563A5:3420D4E7580BF429078C8C8B8E589DBD" country="United States of America" county="Gaston Co." latitude="35.1977" longLatPrecision="6" longitude="-81.3152" municipality="Crowders Mountain State Park" name="Ridgeline Trail at Road 1104" stateProvince="North Carolina">Ridgeline Trail at Road 1104</location>
; 
<named-content content-type="dwc:verbatimCoordinates" id="NCID0EJXCM" specific-use="{&quot;type&quot;:&quot;Point&quot;,&quot;coordinates&quot;:[-81.315200,35.197700]}">
<geoCoordinate id="17E8CEBD30F4DCF7ED500F972003A911" degrees="35.1977" direction="north" orientation="latitude" precision="5" value="35.1977">35.1977</geoCoordinate>
, 
<geoCoordinate id="B3E3E9A45DFD4DE965AFF55EF88E9247" degrees="81.3152" direction="west" orientation="longitude" precision="5" value="-81.3152">- 81.3152</geoCoordinate>
</named-content>
; 
<date id="E0E48F0D8329C7C25A51F4DE25006874" value="2023-04-29">
<collectingDate id="09EB234BFD8E0857172F5AC2EF0543F3" value="2023-04-29">29 Apr. 2023</collectingDate>
</date>
; 
<collectorName id="592A843BA026FC9F5D5F4EA29D780D27">C. W. Harden</collectorName>
leg.; 
<collectedFrom id="8775D7A9F08DEF3A89115294B35C826E">under embedded rock</collectedFrom>
; 
<specimenCode id="34B5AF8C92B796457C6D9DAD2F83C9A1" collectionCode="CUAC" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:33168" lsid="urn:lsid:biocol.org:col:33168" name="Clemson University Arthropod Collection" type="University or college">CUAC 000182314</specimenCode>
, CWH- 512; 
<named-content id="3510F79478B7A8E0A9960641490BEE65" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/clemson-university-arthropod-collection" xlink_title="Clemson University Arthropod Collection">
<collectionCode id="CA192DC994F1C3A647FBD2660187F25A" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:33168" lsid="urn:lsid:biocol.org:col:33168" name="Clemson University Arthropod Collection" type="University or college">CUAC</collectionCode>
</named-content>
</materialsCitation>
; • 
<materialsCitation id="C6B23AC254CEC4F4304D4AD91775CC26" collectedFrom="under embedded rock" collectingDate="2023-04-29" collectionCode="CWHc" collectorName="C. W. Harden" country="United States of America" county="Gaston Co." latitude="35.1977" location="Ridgeline Trail at Road 1104" longLatPrecision="6" longitude="-81.3152" municipality="Crowders Mountain State Park" specimenCount="3" specimenCount-male="3" stateProvince="North Carolina">
<specimenCount id="AF2091228DB7475C3E101C9519A4CF32" count="3" type="male">3 ♂</specimenCount>
; 
<emphasis id="EEFFD079D6DF8143D2B104233F0DACC6" bold="true">
<collectingCounty id="DE00BD02A696BFFE4157196204C7E027">Gaston Co.</collectingCounty>
</emphasis>
; 
<collectingMunicipality id="139196B151B92FA0A377C5C6D73329C9">Crowders Mountain State Park</collectingMunicipality>
; 
<location id="5BF080B38C591353017BC75DBCDF393F" LSID="urn:lsid:plazi:treatment:19BC4D641F675A5B9B4F22489B3563A5:5BF080B38C591353017BC75DBCDF393F" country="United States of America" county="Gaston Co." latitude="35.1977" longLatPrecision="6" longitude="-81.3152" municipality="Crowders Mountain State Park" name="Ridgeline Trail at Road 1104" stateProvince="North Carolina">Ridgeline Trail at Road 1104</location>
; 
<named-content content-type="dwc:verbatimCoordinates" id="NCID0EXXCM" specific-use="{&quot;type&quot;:&quot;Point&quot;,&quot;coordinates&quot;:[-81.315200,35.197700]}">
<geoCoordinate id="611F7A3B8A1C9DFC65B746EAF609E927" degrees="35.1977" direction="north" orientation="latitude" precision="5" value="35.1977">35.1977</geoCoordinate>
, 
<geoCoordinate id="C6014D9A82DEEE7D2436190349666D3B" degrees="81.3152" direction="west" orientation="longitude" precision="5" value="-81.3152">- 81.3152</geoCoordinate>
</named-content>
; 
<date id="B5D279767CB10A051427EEED7E47E855" value="2023-04-29">
<collectingDate id="7A12065A0D795601CAFA28D763FD5610" value="2023-04-29">29 Apr. 2023</collectingDate>
</date>
; 
<collectorName id="D95C4DF3829C7CB15F0E401BE7421E16">C. W. Harden</collectorName>
leg.; 
<collectedFrom id="0FFB7A8DCC145D80D8936DDA186910CA">under embedded rock</collectedFrom>
; 
<abbrev id="ABBRID0E3XCM" xlink_title="Curt Harden personal collection, Central, SC, USA">
<collectionCode id="509ACD4A6F3E41CAF3A1EFF5E4DBDE54">CWHc</collectionCode>
</abbrev>
</materialsCitation>
; • 
<materialsCitation id="DF6EBD59354B9010D5EEB18CD3765C69" collectedFrom="under rock" collectingDate="2023-04-29" collectionCode="CWHc" collectorName="C. W. Harden" country="United States of America" county="Gaston Co." latitude="35.2011" location="near Pinnacle, open piney ridge" longLatPrecision="6" longitude="-81.3148" municipality="Crowders Mountain State Park" specimenCount="1" specimenCount-female="1" stateProvince="North Carolina">
<specimenCount id="B7CF3432694461ABCE4325F721C4498C" count="1" type="female">1 ♀</specimenCount>
; 
<emphasis id="6809FDB41CFAD0CA52B6269541C514E0" bold="true">
<collectingCounty id="D1395AD73DAFCC9A50B0BFBCE0F2FBDF">Gaston Co.</collectingCounty>
</emphasis>
; 
<collectingMunicipality id="759D056894EC91A5DC0E0794C32A841E">Crowders Mountain State Park</collectingMunicipality>
, 
<location id="89383EE72DDD0A857E87C8F1789E06B4" LSID="urn:lsid:plazi:treatment:19BC4D641F675A5B9B4F22489B3563A5:89383EE72DDD0A857E87C8F1789E06B4" country="United States of America" county="Gaston Co." latitude="35.2011" longLatPrecision="6" longitude="-81.3148" municipality="Crowders Mountain State Park" name="near Pinnacle, open piney ridge" stateProvince="North Carolina">near Pinnacle, open piney ridge</location>
; 
<named-content content-type="dwc:verbatimCoordinates" id="NCID0EFYCM" specific-use="{&quot;type&quot;:&quot;Point&quot;,&quot;coordinates&quot;:[-81.314800,35.201100]}">
<geoCoordinate id="94F9C025A6D6034720A270AB6422C2DE" degrees="35.2011" direction="north" orientation="latitude" precision="5" value="35.2011">35.2011</geoCoordinate>
, 
<geoCoordinate id="C05F1BF22ABF67FF6B1B4655D1DFC8A2" degrees="81.3148" direction="west" orientation="longitude" precision="5" value="-81.3148">- 81.3148</geoCoordinate>
</named-content>
; 
<date id="0C6BFBC4B695CCD08B65AFD950A76FD7" value="2023-04-29">
<collectingDate id="F3F38E7E2F0863707E8AE23F5B2B9F51" value="2023-04-29">29 Apr. 2023</collectingDate>
</date>
; 
<collectorName id="EBDDC503B9204920367FA5D89B42EC6D">C. W. Harden</collectorName>
leg.; 
<collectedFrom id="B8A5C497156AB8686D7FB6360170724C">under rock</collectedFrom>
; 
<abbrev id="ABBRID0EKYCM" xlink_title="Curt Harden personal collection, Central, SC, USA">
<collectionCode id="5AE0A9A015931DE49E0996264B7A557E">CWHc</collectionCode>
</abbrev>
</materialsCitation>
; • 
<materialsCitation id="47E542DD7957261B96D07770821D4479" collectedFrom="under rock" collectingDate="2023-04-29" collectionCode="CWHc" collectorName="C. W. Harden" country="United States of America" county="Gaston Co." latitude="35.1993" location="near Pinnacle, recently washed gully" longLatPrecision="6" longitude="-81.3152" municipality="Crowders Mountain State Park" specimenCount="1" specimenCount-male="1" stateProvince="North Carolina">
<specimenCount id="F79EA40A7A214241EC17DB21F663E40F" count="1" type="male">1 ♂</specimenCount>
; 
<emphasis id="EFEA8A76324336DFF82E0BDA76B4F9DB" bold="true">
<collectingCounty id="A1D04343E95E8815EE983E06F79A6484">Gaston Co.</collectingCounty>
</emphasis>
; 
<collectingMunicipality id="102B018192569C668FBC1A9DC2DF8FAB">Crowders Mountain State Park</collectingMunicipality>
, 
<location id="0BBD573198001B907F8F1575B4BA65BC" LSID="urn:lsid:plazi:treatment:19BC4D641F675A5B9B4F22489B3563A5:0BBD573198001B907F8F1575B4BA65BC" country="United States of America" county="Gaston Co." latitude="35.1993" longLatPrecision="6" longitude="-81.3152" municipality="Crowders Mountain State Park" name="near Pinnacle, recently washed gully" stateProvince="North Carolina">near Pinnacle, recently washed gully</location>
; 
<named-content content-type="dwc:verbatimCoordinates" id="NCID0ETYCM" specific-use="{&quot;type&quot;:&quot;Point&quot;,&quot;coordinates&quot;:[-81.315200,35.199300]}">
<geoCoordinate id="0865BB6DC97A3C527435633AF016C063" degrees="35.1993" direction="north" orientation="latitude" precision="5" value="35.1993">35.1993</geoCoordinate>
, 
<geoCoordinate id="4544C78E61BB0A9E05DDC2EC59A4219B" degrees="81.3152" direction="west" orientation="longitude" precision="5" value="-81.3152">- 81.3152</geoCoordinate>
</named-content>
; 
<date id="C64A80A7F164FF3176E853C644D197E9" value="2023-04-29">
<collectingDate id="65C7D2BBD9096FAD50DDAA32C0D9721D" value="2023-04-29">29 Apr. 2023</collectingDate>
</date>
; 
<collectorName id="42FFCAD89E4435A5EB54A8DAECD401E6">C. W. Harden</collectorName>
leg.; 
<collectedFrom id="DD73274E8D0C304CAD461312144C54E8">under rock</collectedFrom>
; 
<abbrev id="ABBRID0EYYCM" xlink_title="Curt Harden personal collection, Central, SC, USA">
<collectionCode id="10A146AF3A664A99B1853A78F41D4D36">CWHc</collectionCode>
</abbrev>
</materialsCitation>
; • 
<materialsCitation id="98FC43824976335E4B7B96C8119D5F04" collectedFrom="under rock" collectingDate="2023-04-29" collectionCode="CWHc" collectorName="C. W. Harden" country="United States of America" county="Gaston Co." latitude="35.2031" location="near Pinnacle, recently washed gully" longLatPrecision="6" longitude="-81.3144" municipality="Crowders Mountain State Park" specimenCount="3" specimenCount-male="3" stateProvince="North Carolina">
<specimenCount id="3DCDAEC0983BA6C7284D07CE5938C366" count="3" type="male">3 ♂</specimenCount>
; 
<emphasis id="FE0B325ACACFA2B6074B711ADE38B061" bold="true">
<collectingCounty id="6A4648DEB816653F3943EC78688D4871">Gaston Co.</collectingCounty>
</emphasis>
; 
<collectingMunicipality id="1F57D45B6BD63F677400AF026F8BF496">Crowders Mountain State Park</collectingMunicipality>
, 
<location id="27A0E8166F6DF830F0ED6005FEAF88F6" LSID="urn:lsid:plazi:treatment:19BC4D641F675A5B9B4F22489B3563A5:27A0E8166F6DF830F0ED6005FEAF88F6" country="United States of America" county="Gaston Co." latitude="35.2031" longLatPrecision="6" longitude="-81.3144" municipality="Crowders Mountain State Park" name="near Pinnacle, recently washed gully" stateProvince="North Carolina">near Pinnacle, recently washed gully</location>
; 
<named-content content-type="dwc:verbatimCoordinates" id="NCID0EBZCM" specific-use="{&quot;type&quot;:&quot;Point&quot;,&quot;coordinates&quot;:[-81.314400,35.203100]}">
<geoCoordinate id="634C80522D305F269F1064FB82567689" degrees="35.2031" direction="north" orientation="latitude" precision="5" value="35.2031">35.2031</geoCoordinate>
, 
<geoCoordinate id="F8348A5AEA5701C8AE3F390A23686E5E" degrees="81.3144" direction="west" orientation="longitude" precision="5" value="-81.3144">- 81.3144</geoCoordinate>
</named-content>
; 
<date id="42D8ADE71203175A00C3BA4659A6FF4D" value="2023-04-29">
<collectingDate id="DB2DE5BBF6852D859223AA672385DF38" value="2023-04-29">29 Apr. 2023</collectingDate>
</date>
; 
<collectorName id="D7DF79E0E591586E5BEE2ABE8C8B896F">C. W. Harden</collectorName>
leg.; 
<collectedFrom id="F7185A26C53E73E92B8BDD6213B12EEE">under rock</collectedFrom>
; 
<abbrev id="ABBRID0EGZCM" xlink_title="Curt Harden personal collection, Central, SC, USA">
<collectionCode id="F8CE95A945C049F74B4DD1469240127E">CWHc</collectionCode>
</abbrev>
</materialsCitation>
; • 
<materialsCitation id="F2A6D585FCBED6A226EFB3675A93F31D" collectedFrom="deep litter and topsoil, oak, maple, sourwood, pine" collectingDate="2020-09-26" collectingMethod="Berlese" collectionCode="CWHc" collectorName="C. W. Harden" country="United States of America" county="York Co." latitude="35.13198" location="Kings Mountain State Park" longLatPrecision="1" longitude="-81.36608" specimenCount="10" specimenCount-female="3" specimenCount-male="7" stateProvince="South Carolina">
<emphasis id="7E5EBFDF11CBC7E16AAC703F3D9B7C6C" bold="true">
<collectingRegion id="EB71402630B72936B6DAF1459E38519B" country="United States of America" name="South Carolina">South Carolina</collectingRegion>
</emphasis>
• 
<emphasis id="0121E8D63638EB77FC28D78F71BA7A30" bold="true">
<collectingCounty id="5CD2C0BF39B99C04E8BCDFDE8DC1C214">York Co.</collectingCounty>
</emphasis>
• 
<specimenCount id="779C9C797F75FB264266FD91D21F16D6" count="7" type="male">7 ♂</specimenCount>
, 
<specimenCount id="A6E8E43F59A3CF2C892156C3D3AF9854" count="3" type="female">3 ♀</specimenCount>
; 
<location id="4C9D94C2495C6C85E11B0F512A521FC3" LSID="urn:lsid:plazi:treatment:19BC4D641F675A5B9B4F22489B3563A5:4C9D94C2495C6C85E11B0F512A521FC3" country="United States of America" county="York Co." latitude="35.13198" longLatPrecision="1" longitude="-81.36608" name="Kings Mountain State Park" stateProvince="South Carolina">Kings Mountain State Park</location>
; 
<named-content content-type="dwc:verbatimCoordinates" id="NCID0ERZCM" specific-use="{&quot;type&quot;:&quot;Point&quot;,&quot;coordinates&quot;:[-81.366080,35.131980]}">
<geoCoordinate id="062C5B75593B3ED895629C407CD5CC06" degrees="35.13198" direction="north" orientation="latitude" precision="1" value="35.13198">35.13198</geoCoordinate>
, 
<geoCoordinate id="4B1838DEE545ECB36902EEE1B3E103A9" degrees="81.36608" direction="west" orientation="longitude" precision="1" value="-81.36608">- 81.36608</geoCoordinate>
</named-content>
; 
<date id="193703AA11E393E55CB57F9EB20EC6EA" value="2020-09-26">
<collectingDate id="54F0F0D6A778B3CDF7DAD2F14125B6B4" value="2020-09-26">26 Sep. 2020</collectingDate>
</date>
; 
<collectorName id="D7C68F6A79D915977A0E0C00DD5EBAA2">C. W. Harden</collectorName>
leg.; 
<collectingMethod id="0716360DD8C734880DD3EA0DAB7E9D53">Berlese</collectingMethod>
, 
<collectedFrom id="629142415A509B59A3B175F04D74D175">deep litter and topsoil, oak, maple, sourwood, pine</collectedFrom>
; 
<abbrev id="ABBRID0EWZCM" xlink_title="Curt Harden personal collection, Central, SC, USA">
<collectionCode id="41E4BFF175DCEB937AC7BDC0FED525BC">CWHc</collectionCode>
</abbrev>
</materialsCitation>
; • 
<materialsCitation id="AE496A45E77D60BADBF024AB9475215E" collectedFrom="in leaf litter" collectingDate="1998-02-09" collectionCode="CUAC" collectorName="J. C. Ciegler" country="United States of America" latitude="35.116" location="Bethany" longLatPrecision="71" longitude="-81.305" specimenCount="1" specimenCount-female="1" stateProvince="South Carolina">
<specimenCount id="56DCA0590FCF09B7720FA3B0B55589DE" count="1" type="female">1 ♀</specimenCount>
; 
<location id="AEA9D0964124E8C4117C8E321D489EDD" LSID="urn:lsid:plazi:treatment:19BC4D641F675A5B9B4F22489B3563A5:AEA9D0964124E8C4117C8E321D489EDD" country="United States of America" latitude="35.116" longLatPrecision="71" longitude="-81.305" name="Bethany" stateProvince="South Carolina">Bethany</location>
; 
<named-content content-type="dwc:verbatimCoordinates" id="NCID0E4ZCM" specific-use="{&quot;type&quot;:&quot;Point&quot;,&quot;coordinates&quot;:[-81.305000,35.116000]}">
<geoCoordinate id="A458023B3077895FF7DB6638398F3B7B" degrees="35.116" direction="north" orientation="latitude" precision="55" value="35.116">35.116</geoCoordinate>
, 
<geoCoordinate id="16755CB15F4ED697731B8F646D8C14C2" degrees="81.305" direction="west" orientation="longitude" precision="55" value="-81.305">- 81.305</geoCoordinate>
</named-content>
; 
<date id="A0A06FE2EAB96850490EFCB1E5C38E52" value="1998-02-09">
<collectingDate id="14F7661F54A5BF63ABE93A16585E1B58" value="1998-02-09">9 Feb. 1998</collectingDate>
</date>
; 
<collectorName id="397B98DC313A8FAAFF29F0AAA387FAF5">J. C. Ciegler</collectorName>
leg.; 
<collectedFrom id="5C7C166CE42EC603E48A391EC852A705">in leaf litter</collectedFrom>
; 
<named-content id="78B624B4BDED060C69727FFEB9D45C85" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/clemson-university-arthropod-collection" xlink_title="Clemson University Arthropod Collection">
<collectionCode id="006D6DEEDCF34687EB200F1744BC8F52" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:33168" lsid="urn:lsid:biocol.org:col:33168" name="Clemson University Arthropod Collection" type="University or college">CUAC</collectionCode>
</named-content>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="SECID0EG1CM" type="diagnosis">
<paragraph id="C22D7788BF77997DD364BA98F56DF1BA">
<heading id="4AD741BC79D840E321842A171DAEB73C" reason="title">Diagnosis.</heading>
</paragraph>
<paragraph id="6BEEA90E0610760342EDA184F4614501">
Males of 
<taxonomicName id="7CC0005BC8228904A774785AA3810A19" authorityName="Harden &amp; Caterino" authorityYear="2024" class="Insecta" family="Carabidae" genus="Anillinus" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="simplex">
<emphasis id="23B152B1712A0C4A80CBAF2A1BC2E596" italics="true">A. simplex</emphasis>
</taxonomicName>
are the only members of the 
<taxonomicName id="772DD51E35CBD815B5CC737D96F0F62D" authorityName="Jeannel" authorityYear="1963" class="Insecta" family="Carabidae" genus="Troglanillus" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="valentinei">
<emphasis id="629BEE9F30C29AAAB4F6D7D8A2B54482" italics="true">valentinei</emphasis>
</taxonomicName>
group known to lack ventral adhesive setae on protarsomere 2, but this character is difficult to confirm without strong magnification. From most anillines occurring in 
<collectingRegion id="C7696638BAB22A42CA6B06F77E24BBAD" country="United States of America" name="South Carolina">South Carolina</collectingRegion>
, members of 
<taxonomicName id="D13F80254013AAD489032E36DC6E1C42" authorityName="Harden &amp; Caterino" authorityYear="2024" class="Insecta" family="Carabidae" genus="Anillinus" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="simplex">
<emphasis id="32EFC5592B1D1585D844FFA314C37019" italics="true">A. simplex</emphasis>
</taxonomicName>
can be recognized by their small size (
<abbrev id="ABBRID0EM2CM" xlink_title="Apparent body length, measured from anterior edge of clypeus to apex of elytra">ABL</abbrev>
&lt;
<quantity id="58742017A70DE38BDA5CF36FC06CE7DD" metricMagnitude="-3" metricUnit="m" metricValue="1.65" unit="mm" value="1.65">1.65 mm</quantity>
) and effaced microsculpture on the sides of the vertex. The median lobe of 
<taxonomicName id="C0F1987B4CC14D5CE8D2014984F294B7" authorityName="Harden &amp; Caterino" authorityYear="2024" class="Insecta" family="Carabidae" genus="Anillinus" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="simplex">
<emphasis id="42B3F63464F93345B45673CFD3C90BCC" italics="true">A. simplex</emphasis>
</taxonomicName>
is distinctive, with a short, thick well-sclerotized curved flagellum in the internal sac and the absence of other well-sclerotized structures (Fig. 
<figureCitation id="79EE66BC4F16EEAF39DBE03536767EA1" captionStart="Figure 30" captionStartId="F30" captionText="Figure 30. Aedeagi of valentinei group Anillinus species A – C Anillinus castaneus sp. nov. D – F Anillinus cornelli G – I Anillinus simplex sp. nov. J – N Anillinus murrayae. Median lobes C, F, I in right dorsolateral aspect, median lobe K in right lateral aspect. Left A, D, G, J and right B, E, H, L – N parameres in left and right lateral aspects, respectively. Scale bar: 0.1 mm." figureDoi="10.3897/zookeys.1209.125897.figure30" httpUri="https://binary.pensoft.net/fig/1110369">30 I</figureCitation>
). Some individuals of 
<taxonomicName id="5D205F3000D6E08EBC370F2E47B7CC15" authorityName="Sokolov &amp; Carlton" authorityYear="2004" class="Insecta" family="Carabidae" genus="Anillinus" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="murrayae">
<emphasis id="327E8729A920171F429A157B4A19A4EE" italics="true">A. murrayae</emphasis>
</taxonomicName>
from southern Pickens Co. overlap in size with members of 
<taxonomicName id="35A08B1A5247A474B406FFD38BDD476C" authorityName="Harden &amp; Caterino" authorityYear="2024" class="Insecta" family="Carabidae" genus="Anillinus" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="simplex">
<emphasis id="7FBDE1AD854933FDD1CC8D482FB38F89" italics="true">A. simplex</emphasis>
</taxonomicName>
but are easily distinguished by the different male and female genitalia (Figs 
<figureCitation id="45A0D4D5154A8491A3BF9AE91DA4ABCF" captionStart="Figure 21" captionStartId="F21" captionText="Figure 21. Spermathecae of Anillini species A, B Anillinus chandleri C Anillinus castaneus D Anillinus murrayae E Anillinus simplex F Anillinus dentatus G Anillinus sp. “ South Carolina, Coon Branch ” H Anillinus micamicus I Anillinus mica J Anillinus choestoea K Anillinus seneca L Anillinus montrex M Anillinus arenicollis N Anillinus jancae O Anillinus cf. nantahala P Anillinus cherokee Q Anillinus loweae R Anillinus merritti S Serranillus dunavani T Serranillus jeanneli. Scale bar: 0.1 mm." figureDoi="10.3897/zookeys.1209.125897.figure21" httpUri="https://binary.pensoft.net/fig/1110360">21 D</figureCitation>
, 
<figureCitation id="23DA543530DD264C4EF5286EAB66149C" captionStart="Figure 30" captionStartId="F30" captionText="Figure 30. Aedeagi of valentinei group Anillinus species A – C Anillinus castaneus sp. nov. D – F Anillinus cornelli G – I Anillinus simplex sp. nov. J – N Anillinus murrayae. Median lobes C, F, I in right dorsolateral aspect, median lobe K in right lateral aspect. Left A, D, G, J and right B, E, H, L – N parameres in left and right lateral aspects, respectively. Scale bar: 0.1 mm." figureDoi="10.3897/zookeys.1209.125897.figure30" httpUri="https://binary.pensoft.net/fig/1110369">30 J – N</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="SECID0E43CM" type="description">
<paragraph id="CA3735FD840E14D18A444A227BF1DBDA">
<heading id="F9B45E3F56815FCE2965F83CEBE73D59" reason="title">Description.</heading>
</paragraph>
<paragraph id="D4C822318363CDABE5F47E03C017371E">
<emphasis id="E3AC9650FD2CDFC2DDE6CA26913D9344" bold="true" italics="true">Habitus</emphasis>
Body small (
<abbrev id="ABBRID0EI4CM" xlink_title="Apparent body length, measured from anterior edge of clypeus to apex of elytra">ABL</abbrev>
= 
<quantity id="7C98352FB2B10E2F2EA8520DD4DFF75D" metricMagnitude="-3" metricUnit="m" metricValue="1.545" metricValueMax="1.64" metricValueMin="1.45" unit="mm" value="1.545" valueMax="1.64" valueMin="1.45">1.45–1.64 mm</quantity>
), moderately convex, slightly ovoid, robust (
<abbrev id="ABBRID0EM4CM" xlink_title="Maximum elytral width">EW</abbrev>
/ 
<abbrev id="ABBRID0EQ4CM" xlink_title="Apparent body length, measured from anterior edge of clypeus to apex of elytra">ABL</abbrev>
= 0.35–0.38) (Fig. 
<figureCitation id="1595A8B79441634F4AAA7C952E67AF3B" captionStart="Figure 33" captionStartId="F33" captionText="Figure 33. Anillinus simplex sp. nov. A dorsal habitus, abdomen removed for DNA extraction B last abdominal ventrite, ventral aspect C head, left lateral aspect (black arrow = frontoclypeal horn). Scale bars: 1 mm (A); 0.1 mm (B)." figureDoi="10.3897/zookeys.1209.125897.figure33" httpUri="https://binary.pensoft.net/fig/1110372">33 A</figureCitation>
). 
<emphasis id="163741F9ADB23FA2FAA031825FB23FAB" bold="true" italics="true">Integument</emphasis>
Dorsal microsculpture of forebody variable. In specimens from the type locality, microsculpture is effaced from most of the pronotum, present only along extreme outer margins, and weakly impressed there, and absent from large portions on both sides of the vertex posteriorly. In specimens from Crowders Mountain, the pronotum is entirely covered in strong, easily visible isodiametric microsculpture, and the smooth patches on the vertex are smaller. 
<emphasis id="437F936919AAD70290B091F4ED50F005" bold="true" italics="true">Head</emphasis>
<abbrev id="ABBRID0E54CM" xlink_title="Maximum head width">HW</abbrev>
/ 
<abbrev id="ABBRID0EC5CM" xlink_title="Maximum pronotal width">PW</abbrev>
= 0.71–0.77. Three pairs of supraorbital setae present, outer posterior pair shorter than other two. Frontoclypeal horn well-developed (Fig. 
<figureCitation id="96EADB8E0DF05890F0044CA5923354D6" captionStart="Figure 33" captionStartId="F33" captionText="Figure 33. Anillinus simplex sp. nov. A dorsal habitus, abdomen removed for DNA extraction B last abdominal ventrite, ventral aspect C head, left lateral aspect (black arrow = frontoclypeal horn). Scale bars: 1 mm (A); 0.1 mm (B)." figureDoi="10.3897/zookeys.1209.125897.figure33" httpUri="https://binary.pensoft.net/fig/1110372">33 C</figureCitation>
). 
<emphasis id="72EE50923D0DD6009810414A0BA5C889" bold="true" italics="true">Pronotum</emphasis>
Relatively short (
<abbrev id="ABBRID0EN5CM" xlink_title="Pronotal length, measured along midline">PL</abbrev>
/ 
<abbrev id="ABBRID0ER5CM" xlink_title="Apparent body length, measured from anterior edge of clypeus to apex of elytra">ABL</abbrev>
= 0.22–0.24) and broad (
<abbrev id="ABBRID0EV5CM" xlink_title="Maximum pronotal width">PW</abbrev>
/ 
<abbrev id="ABBRID0EZ5CM" xlink_title="Maximum elytral width">EW</abbrev>
= 0.80–0.85), sides evenly convergent or slightly sinuate before slightly narrowed posterior angles (
<abbrev id="ABBRID0E45CM" xlink_title="Pronotal basal width, measured at posterior angles and not including external denticles">PbW</abbrev>
/ 
<abbrev id="ABBRID0EB6CM" xlink_title="Maximum pronotal width">PW</abbrev>
= 0.73–0.78). 
<emphasis id="01691BAF97CB2E97A39132E1453C34EA" bold="true" italics="true">Elytra</emphasis>
Slightly ovoid, widest approximately middle, length variable (
<abbrev id="ABBRID0EI6CM" xlink_title="Elytral length, measured from posterior edge of scutellum to apex of elytra">EL</abbrev>
/ 
<abbrev id="ABBRID0EM6CM" xlink_title="Apparent body length, measured from anterior edge of clypeus to apex of elytra">ABL</abbrev>
= 0.51–0.57). Umbilicate punctures relatively large and conspicuous. Striae weakly impressed, difficult to trace. 
<emphasis id="7DDF484BDC3D43F24ED6F02545C0DCBC" bold="true" italics="true">Legs</emphasis>
Profemora of males unmodified; protarsomere 1 of males expanded and with inner margin spinose, protarsomere 2 unmodified and apparently lacking ventral adhesive setae in most specimens (a single adhesive seta observed in 
<specimenCount id="CDC269319890F1AD4DCDCFE17721ECEF" count="1" type="male">one male</specimenCount>
). Mesofemora of males unmodified. Metafemora of males with patch of coarse papillate microsculpture medially on posterior face. 
<emphasis id="A70F53717D7D01577D19954C4AEEC525" bold="true" italics="true">Abdominal ventrites</emphasis>
Unmodified in either sex (Fig. 
<figureCitation id="5D274D044DDDB411FD587F5E5E2A586D" captionStart="Figure 33" captionStartId="F33" captionText="Figure 33. Anillinus simplex sp. nov. A dorsal habitus, abdomen removed for DNA extraction B last abdominal ventrite, ventral aspect C head, left lateral aspect (black arrow = frontoclypeal horn). Scale bars: 1 mm (A); 0.1 mm (B)." figureDoi="10.3897/zookeys.1209.125897.figure33" httpUri="https://binary.pensoft.net/fig/1110372">33 B</figureCitation>
). 
<emphasis id="048E0FB382FA186622471C82CB2A3118" bold="true" italics="true">Male genitalia</emphasis>
Ring sclerite (Fig. 
<figureCitation id="3F695532E12FFBFB7EBB27FA87419120" captionStart="Figure 24" captionStartId="F24" captionText="Figure 24. Male ring sclerite, ventral aspect. A Anillinus chandleri B Anillinus castaneus C Anillinus sp. “ South Carolina, Chestnut Ridge ” D Anillinus murrayae E Anillinus simplex F Anillinus dentatus G Anillinus mica H Anillinus micamicus I Anillinus choestoea J Anillinus seneca K Anillinus arenicollis L Anillinus montrex M Anillinus jancae N Anillinus loweae O Anillinus cherokee P Anillinus merritti. Scale bar: 0.5 mm." figureDoi="10.3897/zookeys.1209.125897.figure24" httpUri="https://binary.pensoft.net/fig/1110363">24 E</figureCitation>
) moderately small (
<abbrev id="ABBRID0EBADM" xlink_title="Ring sclerite length">RL</abbrev>
/ 
<abbrev id="ABBRID0EFADM" xlink_title="Apparent body length, measured from anterior edge of clypeus to apex of elytra">ABL</abbrev>
= 0.28) and asymmetrical, strongly narrowed anteriorly. Median lobe (Fig. 
<figureCitation id="FD3BB01BC7A8EED57C6850307F404963" captionStart="Figure 30" captionStartId="F30" captionText="Figure 30. Aedeagi of valentinei group Anillinus species A – C Anillinus castaneus sp. nov. D – F Anillinus cornelli G – I Anillinus simplex sp. nov. J – N Anillinus murrayae. Median lobes C, F, I in right dorsolateral aspect, median lobe K in right lateral aspect. Left A, D, G, J and right B, E, H, L – N parameres in left and right lateral aspects, respectively. Scale bar: 0.1 mm." figureDoi="10.3897/zookeys.1209.125897.figure30" httpUri="https://binary.pensoft.net/fig/1110369">30 I</figureCitation>
) evenly curved, slightly twisted dorsally from level of basal lobes, gradually expanded in width from base to apex. Ventral margin slightly expanded, asetose. Apex small and evenly rounded. Left side of base with semicircular excavation visible in left posterolateral aspect, with carina extending distally from the excavation to approximately middle. Internal sac with well-sclerotized flagellum short and stout, evenly curved, with one short basal extension. Ventral surface of internal sac with field of weakly sclerotized teeth. Sides of internal sac near ostium appearing grooved with curved parallel lines. Ostial plate on left side, small and weakly sclerotized. Right paramere narrow, with four long apical setae (Fig. 
<figureCitation id="93308B8B2BFA81002FDDA9820FA333D5" captionStart="Figure 30" captionStartId="F30" captionText="Figure 30. Aedeagi of valentinei group Anillinus species A – C Anillinus castaneus sp. nov. D – F Anillinus cornelli G – I Anillinus simplex sp. nov. J – N Anillinus murrayae. Median lobes C, F, I in right dorsolateral aspect, median lobe K in right lateral aspect. Left A, D, G, J and right B, E, H, L – N parameres in left and right lateral aspects, respectively. Scale bar: 0.1 mm." figureDoi="10.3897/zookeys.1209.125897.figure30" httpUri="https://binary.pensoft.net/fig/1110369">30 H</figureCitation>
). Left paramere conchoidal, with four ventral subapical pores, the apical two bearing short setae (Fig. 
<figureCitation id="CCFDB180AAEFBD4945650523F7D83314" captionStart="Figure 30" captionStartId="F30" captionText="Figure 30. Aedeagi of valentinei group Anillinus species A – C Anillinus castaneus sp. nov. D – F Anillinus cornelli G – I Anillinus simplex sp. nov. J – N Anillinus murrayae. Median lobes C, F, I in right dorsolateral aspect, median lobe K in right lateral aspect. Left A, D, G, J and right B, E, H, L – N parameres in left and right lateral aspects, respectively. Scale bar: 0.1 mm." figureDoi="10.3897/zookeys.1209.125897.figure30" httpUri="https://binary.pensoft.net/fig/1110369">30 G</figureCitation>
). 
<emphasis id="485153A18BEA5BA18A3C63ED9BAFB868" bold="true" italics="true">Female genitalia</emphasis>
Spermatheca long, gradually and slightly enlarged distally, stem slightly sinuate before basal curve, which is rotated perpendicular from rest of stem (Fig. 
<figureCitation id="E88200C901323C6E01FCE9189CA9C786" captionStart="Figure 21" captionStartId="F21" captionText="Figure 21. Spermathecae of Anillini species A, B Anillinus chandleri C Anillinus castaneus D Anillinus murrayae E Anillinus simplex F Anillinus dentatus G Anillinus sp. “ South Carolina, Coon Branch ” H Anillinus micamicus I Anillinus mica J Anillinus choestoea K Anillinus seneca L Anillinus montrex M Anillinus arenicollis N Anillinus jancae O Anillinus cf. nantahala P Anillinus cherokee Q Anillinus loweae R Anillinus merritti S Serranillus dunavani T Serranillus jeanneli. Scale bar: 0.1 mm." figureDoi="10.3897/zookeys.1209.125897.figure21" httpUri="https://binary.pensoft.net/fig/1110360">21 E</figureCitation>
). Spermathecal duct short and not coiled.
</paragraph>
<caption id="5FC00CEF892AE8FACFFED3CA9B6A0A3D" ID-DOI="10.3897/zookeys.1209.125897.figure33" ID-arpha="C1B92850-E3E5-5732-BF90-7324547FA2C3" httpUri="https://binary.pensoft.net/fig/1110372" startId="F33">
<paragraph id="720E1A5B56FE516303916252565FE866">
<label id="43865CE45138A9C1FC894A2B74022245">Figure 33.</label>
</paragraph>
<paragraph id="CFE696A0F6AEA010DA1BCE8A092A6749">
<taxonomicName id="94022BB2E99E58ECF08EC68487EFC4C8" authorityName="Harden &amp; Caterino" authorityYear="2024" class="Insecta" family="Carabidae" genus="Anillinus" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="simplex" status="sp. nov.">
<emphasis id="3E668BD07F6F7FCD9275493C868E5F05" italics="true">Anillinus simplex</emphasis>
</taxonomicName>
<taxonomicNameLabel id="717832DC5C5FC1EDFE7AEE8D62D50069" rank="species">sp. nov.</taxonomicNameLabel>
<emphasis id="B9ADFDF409A3EC8FA0518FA2A0BD322E" bold="true">A</emphasis>
dorsal habitus, abdomen removed for DNA extraction 
<emphasis id="AA4ED4815EE1EEB4BBB82D41C8076674" bold="true">B</emphasis>
last abdominal ventrite, ventral aspect 
<emphasis id="B48AB56332677022376A1E22887B301F" bold="true">C</emphasis>
head, left lateral aspect (black arrow = frontoclypeal horn). Scale bars: 1 mm (
<emphasis id="EE094A5525F28DB3CC80FF8C1C54FADE" bold="true">A</emphasis>
); 0.1 mm (
<emphasis id="C602D68A9CFB1CD8FFAF130BAB874E5A" bold="true">B</emphasis>
).
</paragraph>
</caption>
</subSubSection>
<subSubSection id="SECID0E5BDM" type="distribution">
<paragraph id="CCF8ACF0FE0D1B24FA8B1724DB6C9245">
<heading id="F9B6064AB274F1B1F60F1A96898C9D97" reason="title">Distribution.</heading>
</paragraph>
<paragraph id="13DF2B1EFB1C691D2F7C841F3B2A2685">
Known only from Kings Mountain, a short linear 
<taxonomicName id="307B8D665F4F068976F7A1949BFAF2B2" authorityName="Harden &amp; Caterino" authorityYear="2024" class="Insecta" family="Carabidae" genus="Serranillus" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="monadnock">
<emphasis id="EE933D3DA60A0A0F08CA19D16D9787E5" italics="true">monadnock</emphasis>
</taxonomicName>
spanning from York County, 
<collectingRegion id="7B37ABE71D80888178AB31C57F2B2157" country="United States of America" name="South Carolina">South Carolina</collectingRegion>
to Gaston County, 
<collectingRegion id="DA7EC6A793EA0EC6778F6B185B34E3C1" country="United States of America" name="North Carolina">North Carolina</collectingRegion>
(Fig. 
<figureCitation id="BF7E84E3E2500A55BBE4D5BB96FEABF9" captionStart="Figure 29" captionStartId="F29" captionText="Figure 29. Distribution map of valentinei - group species in South Carolina and adjacent states. Data are from Harden (2024)." figureDoi="10.3897/zookeys.1209.125897.figure29" httpUri="https://binary.pensoft.net/fig/1110368">29</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="SECID0ERCDM" type="Sympatry">
<paragraph id="4577E4427C502EB105154CC64E7B0052">
<heading id="DFD564EA647DD3E92920F3C43539D318" reason="title">Sympatry.</heading>
</paragraph>
<paragraph id="5D31CE6B3B20722157E46311F90481F2">
This species has been collected with 
<taxonomicName id="2DDB2A9CBA61FFDB03678E44E0E1231F" authorityName="Harden &amp; Caterino" authorityYear="2024" class="Insecta" family="Carabidae" genus="Anillinus" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="montrex">
<emphasis id="C84C713B1DF3FD35966B71052DEC45FB" italics="true">A. montrex</emphasis>
</taxonomicName>
and 
<taxonomicName id="4D2B1E3094856EC0971406A85D571B3A" authorityName="Sokolov &amp; Carlton" authorityYear="2004" class="Insecta" family="Carabidae" genus="Anillinus" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="cornelli">
<emphasis id="0A7B28F3EBF5563BC420A109819FEF7D" italics="true">A. cornelli</emphasis>
</taxonomicName>
underneath embedded rocks. 
<taxonomicName id="21212CE04843D4B0818249645C715F21" authorityName="Harden &amp; Caterino" authorityYear="2024" class="Insecta" family="Carabidae" genus="Serranillus" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="monadnock" status="sp. nov.">
<emphasis id="9B2FA927D4DD364052A0FE5AFDD6892F" italics="true">Serranillus monadnock</emphasis>
</taxonomicName>
<taxonomicNameLabel id="FBAB6967C3F8783663C513C726225256" rank="species">sp. nov.</taxonomicNameLabel>
also occurs at Kings Mountain State Park.
</paragraph>
</subSubSection>
<subSubSection id="SECID0EYDDM" type="natural history">
<paragraph id="CF12C4357DF6AAF78C1A670AFF71DCCA">
<heading id="F5AAA01A6F6D6B6558DF388363457CA9" reason="title">Natural history.</heading>
</paragraph>
<paragraph id="332C0D30ADE206268E077D9C8C41F417">Specimens have been collected in February, April, September, and December, from underneath embedded rocks on sandy clay rich soil near an ephemeral stream and through Berlese extraction of sifted litter and soil.</paragraph>
</subSubSection>
<subSubSection id="SECID0E4DDM" type="Species status justification">
<paragraph id="E7D3FC7EABA63FA4AFC2BA13421C3117">
<heading id="7665E20BB23DE93733835A2B83DE252E" reason="title">Species status justification.</heading>
</paragraph>
<paragraph id="20CDA3620D3484198B8FD003426CAF77">
The male and female genitalic characters are unique within the genus, and the DNA sequence data indicate 
<taxonomicName id="BA5427F653DB68B7DEAE67F81E257FC6" authorityName="Harden &amp; Caterino" authorityYear="2024" class="Insecta" family="Carabidae" genus="Anillinus" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="simplex">
<emphasis id="7382FB88DBD513B090AF5BB7EEA22EF8" italics="true">A. simplex</emphasis>
</taxonomicName>
is most closely related to 
<taxonomicName id="EBD79DDF7FC561F03B689DC792983C31" authorityName="Sokolov &amp; Carlton" authorityYear="2004" class="Insecta" family="Carabidae" genus="Anillinus" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="cornelli">
<emphasis id="0CFA2886D2A913C4200C26FE304864C2" italics="true">A. cornelli</emphasis>
</taxonomicName>
, 
<taxonomicName id="1886E3366379FC1FE6030A85FCBDFCE8" authorityName="Harden &amp; Caterino" authorityYear="2024" class="Insecta" family="Carabidae" genus="Anillinus" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="castaneus">
<emphasis id="831F8855699BA25E0D9C4FE0616FF55A" italics="true">A. castaneus</emphasis>
</taxonomicName>
, and “ 
<collectingRegion id="D5B0D81F761B06A0B7FF2FF503F109CA" country="United States of America" name="South Carolina">South Carolina</collectingRegion>
, Chestnut Ridge ”, all of which differ markedly in male genitalic characters, especially by possessing several sclerotized spines in the internal sac.
</paragraph>
</subSubSection>
<subSubSection id="SECID0EEFDM" type="Derivation of species name">
<paragraph id="8A45BCE678D92ABF41A27F9F6DBF83B1">
<heading id="ED22D37422EF89B94223E98DFE76B8A4" reason="title">Derivation of species name.</heading>
</paragraph>
<paragraph id="442C5C92223050827DD0449972ECAC69">
A noun in apposition, from the Latin, meaning “ simple ”, in reference to the structure of the median lobe of the aedeagus, which lacks the complex sclerotized structures in the internal sac that are present in the other 
<collectingRegion id="351E8E07C29B42823214680F9A16AAD6" country="United States of America" name="South Carolina">South Carolina</collectingRegion>
species in the 
<taxonomicName id="2362E7CF97273739C0267998A2161C5C" authorityName="Jeannel" authorityYear="1963" class="Insecta" family="Carabidae" genus="Troglanillus" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="valentinei">
<emphasis id="669298849A5085E67C84079B4AFD3BC1" italics="true">valentinei</emphasis>
</taxonomicName>
group, and the lack of modifications to male protarsomere 2.
</paragraph>
<paragraph id="47D7063684BB01F8018D4170E195AD08">
<title id="F288FFB90DF130A46B4307667C6A6B70">
<taxonomicName id="BCA2624EB24346A2AB6D943BD48E63B9" authorityName="Jeannel" authorityYear="1963" class="Insecta" family="Carabidae" genus="Anillinus" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="elongatus">
‘ 
<emphasis id="DA4E2CBFF6D5230BD6794F1BE9E769D7" italics="true">elongatus</emphasis>
</taxonomicName>
group’
</title>
</paragraph>
<paragraph id="8575246BA1B1907BE7DD0A194D677FF0">
The five described species of the 
<taxonomicName id="CA137A84E988150A77E6A473D4414B09" authorityName="Jeannel" authorityYear="1963" class="Insecta" family="Carabidae" genus="Anillinus" kingdom="Animalia" order="Coleoptera" phylum="Arthropoda" rank="species" species="elongatus">
<emphasis id="3D05389363743958A50ED358E8AFF484" italics="true">elongatus</emphasis>
</taxonomicName>
group were revised by 
<bibRefCitation id="DFAD50D09602FFF13E32D96B4697E3F9" author="Harden CW &amp; Caterino MS" firstAuthor="Harden" journalOrPublisher="Insecta Mundi" pagination="1-31" refId="B40" refString="Harden CW, Caterino MS (2024) Revision of the elongatus and pecki species groups of Anillinus Casey, 1918 (Coleoptera, Carabidae, Trechinae, Anillini), with four new species and notes on Anillinus turneri Jeannel, 1963. Insecta Mundi 1057: 1–31." title="Revision of the elongatus and pecki species groups of Anillinus Casey, 1918 (Coleoptera, Carabidae, Trechinae, Anillini), with four new species and notes on Anillinus turneri Jeannel, 1963." volume="1057" year="2024">Harden and Caterino (2024)</bibRefCitation>
and are discussed and illustrated in more detail in that paper.
</paragraph>
</subSubSection>
</treatment>
</document>