242 lines
18 KiB
XML
242 lines
18 KiB
XML
<document ID-DOI="http://dx.doi.org/10.3897/jhr.94.94263" ID-Pensoft-Pub="1314-2607-94-13" ID-Pensoft-UUID="ADCFB8021287566FB2D7E8A8711D5CAE" ID-ZooBank="6CB807239A47403FABEC9AF8AE7F417F" ModsDocID="1314-2607-94-13" checkinTime="1671601504499" checkinUser="pensoft" docAuthor="Burks, Roger, Mitroiu, Mircea-Dan, Fusu, Lucian, Heraty, John M., Jansta, Petr, Heydon, Steve, Papilloud, Natalie Dale-Skey, Peters, Ralph S., Tselikh, Ekaterina V., Woolley, James B., van Noort, Simon, Baur, Hannes, Cruaud, Astrid, Darling, Christopher, Haas, Michael, Hanson, Paul, Krogmann, Lars & Rasplus, Jean-Yves" docDate="2022" docId="1997E87CD30C5DF29278851C0CB900AC" docLanguage="en" docName="JourHymenoptRes 94: 13-88" docOrigin="Journal of Hymenoptera Research 94" docPubDate="2022-12-20" docSource="http://dx.doi.org/10.3897/jhr.94.94263" docTitle="Perniphora Ruschka 1923" docType="treatment" docVersion="1" id="ADCFB8021287566FB2D7E8A8711D5CAE" lastPageNumber="13" masterDocId="ADCFB8021287566FB2D7E8A8711D5CAE" masterDocTitle="From hell's heart I stab at thee! A determined approach towards a monophyletic Pteromalidae and reclassification of Chalcidoidea (Hymenoptera)" masterLastPageNumber="88" masterPageNumber="13" pageNumber="13" updateTime="1671601504499" updateUser="pensoft">
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<mods:titleInfo>
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<mods:title>From hell's heart I stab at thee! A determined approach towards a monophyletic Pteromalidae and reclassification of Chalcidoidea (Hymenoptera)</mods:title>
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<mods:namePart>Burks, Roger</mods:namePart>
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<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0003-3032-7939</mods:nameIdentifier>
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<mods:affiliation>Department of Entomology, University of California Riverside, Riverside, CA, USA</mods:affiliation>
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<mods:nameIdentifier type="email">burks.roger@gmail.com</mods:nameIdentifier>
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Mitroiu, Mircea-Dan</mods:namePart>
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<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0003-1368-7721</mods:nameIdentifier>
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<mods:affiliation>Faculty of Biology, Alexandru Ioan Cuza University, Iasi, Romania</mods:affiliation>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Fusu, Lucian</mods:namePart>
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<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0003-0819-026X</mods:nameIdentifier>
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<mods:affiliation>Faculty of Biology, Alexandru Ioan Cuza University, Iasi, Romania</mods:affiliation>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Heraty, John M.</mods:namePart>
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<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0002-9246-5651</mods:nameIdentifier>
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<mods:affiliation>Department of Entomology, University of California Riverside, Riverside, CA, USA</mods:affiliation>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Jansta, Petr</mods:namePart>
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<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0001-6409-3603</mods:nameIdentifier>
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<mods:affiliation>Department of Zoology, Faculty of Science, Charles University, Prague, Czech Republic & Department of Entomology, State Museum of Natural History, Stuttgart, Germany</mods:affiliation>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Heydon, Steve</mods:namePart>
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<mods:affiliation>Bohart Museum of Entomology, University of California, Davis, CA, 95616, USA</mods:affiliation>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Papilloud, Natalie Dale-Skey</mods:namePart>
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<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0001-7582-0386</mods:nameIdentifier>
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<mods:affiliation>Insects Division, Natural History Museum, London, UK</mods:affiliation>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Peters, Ralph S.</mods:namePart>
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<mods:affiliation>Zoologisches Forschungsmuseum Alexander Koenig, Leibniz Institute for the Analysis of Biodiversity Change, Bonn, Germany</mods:affiliation>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Tselikh, Ekaterina V.</mods:namePart>
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<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0002-9184-043X</mods:nameIdentifier>
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<mods:affiliation>Zoological Institute, Russian Academy of Sciences, St. Petersburg, Russia</mods:affiliation>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Woolley, James B.</mods:namePart>
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<mods:affiliation>Department of Entomology, Texas A & M University, College Station, TX, USA</mods:affiliation>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>van Noort, Simon</mods:namePart>
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<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0001-6930-9741</mods:nameIdentifier>
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<mods:affiliation>Research and Exhibitions Department, South African Museum, Iziko Museums of South Africa, PO Box 61, Cape Town 8000 South Africa & Department of Biological Sciences, University of Cape Town, Private Bag, Rondebosch, 7701, South Africa</mods:affiliation>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Baur, Hannes</mods:namePart>
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<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0003-1360-3487</mods:nameIdentifier>
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<mods:affiliation>Department of Invertebrates, Natural History Museum Bern, Bern, Switzerland & Institute of Ecology and Evolution, University of Bern, Bern, Switzerland</mods:affiliation>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Cruaud, Astrid</mods:namePart>
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<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0001-8932-4199</mods:nameIdentifier>
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<mods:affiliation>CBGP, INRAE, CIRAD, IRD, Montpellier SupAgro, University of Montpellier, Montpellier, France</mods:affiliation>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Darling, Christopher</mods:namePart>
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<mods:affiliation>Department of Natural History, Royal Ontario Museum, Toronto, ON, M 5 S 2 C 6, Canada & Department of Ecology and Evolutionary Biology, University of Toronto, Toronto, ON, M 5 S 1 A 1, Canada</mods:affiliation>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Haas, Michael</mods:namePart>
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<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0001-6869-6698</mods:nameIdentifier>
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<mods:affiliation>Department of Entomology, State Museum of Natural History, Stuttgart, Germany</mods:affiliation>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Hanson, Paul</mods:namePart>
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<mods:affiliation>Escuela de Biologia, Universidad de Costa Rica, San Pedro de Montes de Oca, San Jose 11501 - 2060, Costa Rica</mods:affiliation>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Krogmann, Lars</mods:namePart>
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<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0002-3724-1735</mods:nameIdentifier>
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<mods:affiliation>Department of Entomology, State Museum of Natural History, Stuttgart, Germany & Institute of Biology, Biological Systematics (190 n) University of Hohenheim, Stuttgart, Germany</mods:affiliation>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Rasplus, Jean-Yves</mods:namePart>
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<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0001-8614-6665</mods:nameIdentifier>
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<mods:affiliation>CBGP, INRAE, CIRAD, IRD, Montpellier SupAgro, University of Montpellier, Montpellier, France</mods:affiliation>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:titleInfo>
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<mods:title>Journal of Hymenoptera Research</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2022</mods:date>
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<mods:detail type="pubDate">
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<mods:number>2022-12-20</mods:number>
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</mods:detail>
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<mods:detail type="volume">
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<mods:number>94</mods:number>
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||
</mods:detail>
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<mods:extent unit="page">
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<mods:start>13</mods:start>
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<mods:end>88</mods:end>
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||
</mods:extent>
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||
</mods:part>
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</mods:relatedItem>
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<mods:location>
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<mods:url>http://dx.doi.org/10.3897/jhr.94.94263</mods:url>
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</mods:location>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">http://dx.doi.org/10.3897/jhr.94.94263</mods:identifier>
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<mods:identifier type="Pensoft-Pub">1314-2607-94-13</mods:identifier>
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<mods:identifier type="ZooBank">6CB807239A47403FABEC9AF8AE7F417F</mods:identifier>
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<treatment LSID="urn:lsid:plazi:treatment:1997E87CD30C5DF29278851C0CB900AC" httpUri="http://treatment.plazi.org/id/1997E87CD30C5DF29278851C0CB900AC" lastPageNumber="13" pageId="0" pageNumber="13">
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<subSubSection pageId="0" pageNumber="13" type="nomenclature">
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<paragraph pageId="0" pageNumber="13">
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||
<taxonomicName LSID="1997E87C-D30C-5DF2-9278-851C0CB900AC" authority="new placement" authorityName="Ruschka" authorityYear="1923" family="Agaonidae" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="" order="Hymenoptera" pageId="0" pageNumber="13" rank="subFamily" subFamily="Sycophaginae">Sycophaginae new placement</taxonomicName>
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</paragraph>
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||
</subSubSection>
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<subSubSection pageId="0" pageNumber="13" type="reference_group">
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<paragraph pageId="0" pageNumber="13">
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||
<taxonomicName family="Sycophagoidae" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="" order="Hymenoptera" pageId="0" pageNumber="13" rank="family">Sycophagoidae</taxonomicName>
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Walker, 1875. Type genus:
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<taxonomicName class="Insecta" family="Agaonidae" genus="Sycophaga" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Sycophaga" order="Hymenoptera" pageId="0" pageNumber="13" phylum="Arthropoda" rank="genus">Sycophaga</taxonomicName>
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Westwood, 1840. Treated as
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<taxonomicName authorityName="Walker" authorityYear="1875" family="Agaonidae" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="" order="Hymenoptera" pageId="0" pageNumber="13" rank="subFamily" subFamily="Sycophaginae">Sycophaginae</taxonomicName>
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by
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<bibRefCitation DOI="https://doi.org/10.5962/bhl.title.10341" author="Ashmead, WH" journalOrPublisher="Proceedings of the Entomological Society of Washington" pageId="0" pageNumber="13" refId="B3" refString="Ashmead, WH, 1904. Classification of the chalcid flies, or the superfamily Chalcidoidea, with descriptions of new species in the Carnegie Museum, collected in South America by Herbert H. Smith. Memoirs of the Carnegie Museum 1: i-xi, 225-551. [pls xxxi-xxxix] https://doi.org/10.5962/bhl.title.10341" title="Classification of the chalcid flies, or the superfamily Chalcidoidea, with descriptions of new species in the Carnegie Museum, collected in South America by Herbert H. Smith. Memoirs of the Carnegie Museum 1: i-xi, 225 - 551. [pls xxxi-xxxix]" url="https://doi.org/10.5962/bhl.title.10341" year="1904">Ashmead (1904)</bibRefCitation>
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.
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</paragraph>
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||
</subSubSection>
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||
<subSubSection pageId="0" pageNumber="13" type="diagnosis">
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<paragraph pageId="0" pageNumber="13">Diagnosis.</paragraph>
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<paragraph pageId="0" pageNumber="13">
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Antennal variable in flagellomere count. Mandibles not falcate. Scapula anteriorly exposed by narrow pronotum. Axilla not strongly advanced. Axillula enlarged and convex (Fig.
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<figureCitation captionStart="Figures 103–108" captionStartId="F18" captionText="Figures 103 - 108. 103 Apocrypta caudata (Girault) (Pteromalidae, Otitesellini): mesosoma dorsal view 104 Philotrypesis caricae L. (Otitesellini): propodeum 105 Seres wardi van Noort (Otitesellini): fore tibia 106 Grandiana sp. (Otitesellini): male dorsal view 107 Pseudidarnes minerva Girault (Pteromalidae, Sycophaginae): metasoma dorsal view 108 Sycophaga sp. (Pteromalidae, Sycophaginae): male lateral view." figureDoi="10.3897/jhr.94.94263.figures103-108" httpUri="https://binary.pensoft.net/fig/781081" pageId="0" pageNumber="13">107</figureCitation>
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). Petiole simple, transverse. Males usually apterous (except in a few genera), residing inside figs (Fig.
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<figureCitation captionStart="Figures 103–108" captionStartId="F18" captionText="Figures 103 - 108. 103 Apocrypta caudata (Girault) (Pteromalidae, Otitesellini): mesosoma dorsal view 104 Philotrypesis caricae L. (Otitesellini): propodeum 105 Seres wardi van Noort (Otitesellini): fore tibia 106 Grandiana sp. (Otitesellini): male dorsal view 107 Pseudidarnes minerva Girault (Pteromalidae, Sycophaginae): metasoma dorsal view 108 Sycophaga sp. (Pteromalidae, Sycophaginae): male lateral view." figureDoi="10.3897/jhr.94.94263.figures103-108" httpUri="https://binary.pensoft.net/fig/781081" pageId="0" pageNumber="13">108</figureCitation>
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).
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</paragraph>
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</subSubSection>
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||
<subSubSection pageId="0" pageNumber="13" type="discussion">
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<paragraph pageId="0" pageNumber="13">Discussion.</paragraph>
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<paragraph pageId="0" pageNumber="13">
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The taxonomic placement of the subfamily
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<taxonomicName authorityName="Walker" authorityYear="1875" family="Agaonidae" kingdom="Animalia" lsidName="" order="Hymenoptera" pageId="0" pageNumber="13" rank="subFamily" subFamily="Sycophaginae">Sycophaginae</taxonomicName>
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has long been controversial.
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<taxonomicName authorityName="Walker" authorityYear="1875" family="Agaonidae" kingdom="Animalia" lsidName="" order="Hymenoptera" pageId="0" pageNumber="13" rank="subFamily" subFamily="Sycophaginae">Sycophaginae</taxonomicName>
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were previously classified in
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<taxonomicName authorityName="Walker" authorityYear="1833" class="Insecta" family="Torymidae" kingdom="Animalia" lsidName="" order="Hymenoptera" pageId="0" pageNumber="13" phylum="Arthropoda" rank="family">Torymidae</taxonomicName>
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(Wiebes, 1967) and
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<taxonomicName class="Insecta" family="Agaonidae" kingdom="Animalia" lsidName="" order="Hymenoptera" pageId="0" pageNumber="13" phylum="Arthropoda" rank="family">Agaonidae</taxonomicName>
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||
(
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||
<bibRefCitation author="Boucek, Z" journalOrPublisher="CAB International, Wallingford, UK" pageId="0" pageNumber="13" refId="B6" refString="Boucek, Z, 1988. Australasian Chalcidoidea (Hymenoptera). A biosystematic revision of genera of fourteen families, with a reclassification of species. CAB International, Wallingford, UK" title="Australasian Chalcidoidea (Hymenoptera). A biosystematic revision of genera of fourteen families, with a reclassification of species." year="1988">
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||
<normalizedToken originalValue="Bouček">Boucek</normalizedToken>
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||
1988
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</bibRefCitation>
|
||
;
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||
<bibRefCitation DOI="https://doi.org/10.1111/cla.12006" author="Heraty, JM" journalOrPublisher="Cladistics" pageId="0" pageNumber="13" pagination="466 - 542" refId="B33" refString="Heraty, JM, Burks, RA, Cruaud, A, Gibson, GA, Liljeblad, J, Munro, JB, Rasplus, J-Y, Delvare, G, Jansta, P, Gumovsky, A, Huber, JT, Woolley, JB, Krogmann, L, Heydon, SL, Polaszek, A, Schmidt, S, Darling, DC, Gates, M, Mottern, J, Murray, E, Dal Molin, A, Triapitsyn, SV, Baur, H, Pinto, JD, Van Noort, S, George, JN, Yoder, MJ, 2013. A phylogenetic analysis of the megadiverse Chalcidoidea (Hymenoptera). Cladistics 29: 466 - 542, DOI: https://doi.org/10.1111/cla.12006" title="A phylogenetic analysis of the megadiverse Chalcidoidea (Hymenoptera)." url="https://doi.org/10.1111/cla.12006" volume="29" year="2013">Heraty et al. 2013</bibRefCitation>
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||
) as the family shares at least few features with these families. Different strategies to reduce biases in our phylogenetic inference (Cruaud et al., submitted) stabilized the position of
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||
<taxonomicName authorityName="Walker" authorityYear="1875" family="Agaonidae" kingdom="Animalia" lsidName="" order="Hymenoptera" pageId="0" pageNumber="13" rank="subFamily" subFamily="Sycophaginae">Sycophaginae</taxonomicName>
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as sister to all other
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<taxonomicName authorityName="Dalman" authorityYear="1820" family="Pteromalidae" kingdom="Animalia" lsidName="" order="Hymenoptera" pageId="0" pageNumber="13" rank="family">Pteromalidae</taxonomicName>
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. This position is corroborated by several features shared with other members of the family and a few others shared with
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<taxonomicName family="Pteromalidae" kingdom="Animalia" lsidName="" order="Hymenoptera" pageId="0" pageNumber="13" rank="subFamily" subFamily="Colotrechninae">Colotrechninae</taxonomicName>
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(large and convex axillula), the next lineage in the
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<taxonomicName authorityName="Dalman" authorityYear="1820" family="Pteromalidae" kingdom="Animalia" lsidName="" order="Hymenoptera" pageId="0" pageNumber="13" rank="family">Pteromalidae</taxonomicName>
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topology.
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<taxonomicName authorityName="Walker" authorityYear="1875" family="Agaonidae" kingdom="Animalia" lsidName="" order="Hymenoptera" pageId="0" pageNumber="13" rank="subFamily" subFamily="Sycophaginae">Sycophaginae</taxonomicName>
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and
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<taxonomicName family="Pteromalidae" kingdom="Animalia" lsidName="" order="Hymenoptera" pageId="0" pageNumber="13" rank="subFamily" subFamily="Colotrechninae">Colotrechninae</taxonomicName>
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also share a gall-associated biology, indeed most
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<taxonomicName authorityName="Walker" authorityYear="1875" family="Agaonidae" kingdom="Animalia" lsidName="" order="Hymenoptera" pageId="0" pageNumber="13" rank="subFamily" subFamily="Sycophaginae">Sycophaginae</taxonomicName>
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are either gall-makers or parasitoids of gall-makers within figs. We therefore propose to include
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<taxonomicName authorityName="Walker" authorityYear="1875" family="Agaonidae" kingdom="Animalia" lsidName="" order="Hymenoptera" pageId="0" pageNumber="13" rank="subFamily" subFamily="Sycophaginae">Sycophaginae</taxonomicName>
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in
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<taxonomicName authorityName="Dalman" authorityYear="1820" family="Pteromalidae" kingdom="Animalia" lsidName="" order="Hymenoptera" pageId="0" pageNumber="13" rank="family">Pteromalidae</taxonomicName>
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new placement.
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</paragraph>
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||
</subSubSection>
|
||
</treatment>
|
||
</document> |