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<document ID-DOI="http://dx.doi.org/10.3897/zookeys.1143.94951" ID-Pensoft-Pub="1313-2970-1143-93" ID-Pensoft-UUID="5E978E6177BE5B39B380B105500D7DA5" ID-ZooBank="9156C6A84BF5472FA7010C2F089CE134" ModsDocID="1313-2970-1143-93" checkinTime="1675260478037" checkinUser="pensoft" docAuthor="van Achterberg, Cornelis, Smit, John T. &amp; Ljubomirov, Toshko" docDate="2023" docId="198D3A48865D59F4B6DD6695854AD86B" docLanguage="en" docName="ZooKeys 1143: 93-163" docOrigin="ZooKeys 1143" docPubDate="2023-01-31" docSource="http://dx.doi.org/10.3897/zookeys.1143.94951" docTitle="Eumenes papillarius" docType="treatment" docVersion="1" id="5E978E6177BE5B39B380B105500D7DA5" lastPageNumber="93" masterDocId="5E978E6177BE5B39B380B105500D7DA5" masterDocTitle="Review of the European Eumenes Latreille (Hymenoptera, Vespidae) using morphology and DNA barcodes, with an illustrated key to species" masterLastPageNumber="163" masterPageNumber="93" pageNumber="93" updateTime="1675260478037" updateUser="pensoft">
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<mods:title>Review of the European Eumenes Latreille (Hymenoptera, Vespidae) using morphology and DNA barcodes, with an illustrated key to species</mods:title>
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>van Achterberg, Cornelis</mods:namePart>
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<mods:affiliation>Naturalis Biodiversity Center, P. O. 9517, 2300 RA Leiden, Netherlands</mods:affiliation>
<mods:nameIdentifier type="email">kees@vanachterberg.org</mods:nameIdentifier>
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Smit, John T.</mods:namePart>
<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0002-1568-5183</mods:nameIdentifier>
<mods:affiliation>Naturalis Biodiversity Center, P. O. 9517, 2300 RA Leiden, Netherlands &amp; European Invertebrate Survey - Netherlands, P. O. 9517, 2300 RA Leiden, Netherlands</mods:affiliation>
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Ljubomirov, Toshko</mods:namePart>
<mods:affiliation>Institute of Biodiversity and Ecosystem Research, Bulgarian Academy of Sciences, Tzar Osvoboditel Boulevard 1, 1000 Sofia, Bulgaria</mods:affiliation>
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<mods:title>ZooKeys</mods:title>
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<mods:date>2023</mods:date>
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<mods:number>2023-01-31</mods:number>
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<mods:number>1143</mods:number>
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<treatment LSID="urn:lsid:plazi:treatment:198D3A48865D59F4B6DD6695854AD86B" httpUri="http://treatment.plazi.org/id/198D3A48865D59F4B6DD6695854AD86B" lastPageNumber="93" pageId="0" pageNumber="93">
<subSubSection pageId="0" pageNumber="93" type="nomenclature">
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<taxonomicName ID-CoL="3CDMC" ID-ENA="1507136" LSID="198D3A48-865D-59F4-B6DD-6695854AD86B" authority="(Christ, 1791)" baseAuthorityName="Christ" baseAuthorityYear="1791" class="Insecta" family="Vespidae" genus="Eumenes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Eumenes papillarius" order="Hymenoptera" pageId="0" pageNumber="93" phylum="Arthropoda" rank="species" species="papillarius">Eumenes papillarius (Christ, 1791)</taxonomicName>
</paragraph>
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<subSubSection pageId="0" pageNumber="93" type="description">
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<figureCitation captionStart="Figures 112–121" captionStartId="F16" captionText="Figures 112 - 121. Eumenes papillarius (Christ), France, female 112 metasoma lateral 113 first metasomal tergite dorsal 114 first tergite ventral 115 head and mesosoma dorsal 116 second tergite latero-dorsal 117 head anterior 118 head and pronotum lateral 119 antenna anterior 120 hind tarsal claw 121 hind tibia lateral." figureDoi="10.3897/zookeys.1143.94951.figures112-121" httpUri="https://binary.pensoft.net/fig/803439" pageId="0" pageNumber="93">Figs 112-121</figureCitation>
<figureCitation captionStart="Figures 122–131" captionStartId="F17" captionText="Figures 122 - 131. Eumenes papillarius (Christ), France, male 122 metasoma lateral 123 first metasomal tergite dorsal 124 first tergite ventral 125 head and mesosoma dorsal 126 second tergite dorsal 127 hind tibia lateral 128 head anterior 129 head and mesosoma lateral 130 antenna anterior 131 apical hook of antenna lateral." figureDoi="10.3897/zookeys.1143.94951.figures122-131" httpUri="https://binary.pensoft.net/fig/803440" pageId="0" pageNumber="93">, 122-131</figureCitation>
<figureCitation captionStart="Figures 262–266" captionStartId="F32" captionText="Figures 262 - 266. Eumenes mediterraneus aemilianus Guiglia, holotype, Italy, female 262 metasoma lateral 263 propodeum and first metasomal tergite dorsal 264 head and mesosoma dorsal 265 head anterior 266 head and mesosoma lateral. Photographs: R. Poggi (MSNG)." figureDoi="10.3897/zookeys.1143.94951.figures262-266" httpUri="https://binary.pensoft.net/fig/803455" pageId="0" pageNumber="93">, 262-266</figureCitation>
</paragraph>
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<subSubSection pageId="0" pageNumber="93" type="reference_group">
<paragraph pageId="0" pageNumber="93">
<taxonomicName authorityName="Christ" authorityYear="1791" class="Insecta" family="Vespidae" genus="Sphex" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Sphex papillarius" order="Hymenoptera" pageId="0" pageNumber="93" phylum="Arthropoda" rank="species" species="papillarius">Sphex papillarius</taxonomicName>
Christ, 1791: 325.
</paragraph>
<paragraph pageId="0" pageNumber="93">
<taxonomicName baseAuthorityName="Christ" baseAuthorityYear="1791" class="Insecta" family="Vespidae" genus="Eumenes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Eumenes papillarius" order="Hymenoptera" pageId="0" pageNumber="93" phylum="Arthropoda" rank="species" species="papillarius">Eumenes papillarius</taxonomicName>
;
<bibRefCitation author="Tobias, VI" editor="Medvedev, G" journalOrPublisher="Hymenoptera 3 (1)" pageId="0" pageNumber="93" pagination="1 - 584" refId="B82" refString="Tobias, VI, Kurzenko, NV, 1978. Sem. 2. Eumenidae, 152-173. In: Medvedev, G, Ed., Keys to the identification of insects of European USSR. Hymenoptera 3(1): 1 - 584" title="Sem. 2. Eumenidae, 152 - 173." volumeTitle="Keys to the identification of insects of European USSR." year="1978">Tobias and Kurzenko 1978</bibRefCitation>
: 160;
<bibRefCitation author="Hensen, R" journalOrPublisher="Jeugdbondsuitgeverij, ' s Graveland" pageId="0" pageNumber="93" refId="B57" refString="Hensen, R, 1985. De plooivleugelwespen. Jeugdbondsuitgeverij, 's Graveland" title="De plooivleugelwespen." year="1985">Hensen 1985</bibRefCitation>
: 45;
<bibRefCitation author="Castro, L" journalOrPublisher="Catalogus de la entomofauna aragonesa" pageId="0" pageNumber="93" pagination="3 - 8" refId="B18" refString="Castro, L, 1997. Familia Vespidae: Subfamilia Eumeninae. Catalogus de la entomofauna aragonesa 16: 3 - 8" title="Familia Vespidae: Subfamilia Eumeninae." volume="16" year="1997">Castro 1997</bibRefCitation>
: 4;
<bibRefCitation author="Schmid-Egger, C" journalOrPublisher="Bembix" pageId="0" pageNumber="93" pagination="13 - 28" refId="B78" refString="Schmid-Egger, C, Schmidt, K, 2002. Kritisches Verzeichnis der deutschen Arten der Faltenwespen (Hymenoptera, Vespidae). Bembix 16: 13 - 28" title="Kritisches Verzeichnis der deutschen Arten der Faltenwespen (Hymenoptera, Vespidae)." volume="16" year="2002">Schmid-Egger and Schmidt 2002</bibRefCitation>
: 18;
<bibRefCitation author="Schmid-Egger, C" journalOrPublisher="Ampulex" pageId="0" pageNumber="93" refId="B75" refString="Schmid-Egger, C, 2004. Bestimmungsschluessel fuer die deutschen Arten der solitaeren Faltenwespen (Hymenoptera: Eumeninae). Deutscher Jugendbund fuer Naturbeobachtung Hamburg, 54-102. 3. ueberarbeitete Auflage. Hamburg." title="Bestimmungsschluessel fuer die deutschen Arten der solitaeren Faltenwespen (Hymenoptera: Eumeninae). Deutscher Jugendbund fuer Naturbeobachtung Hamburg, 54 - 102. 3. ueberarbeitete Auflage. Hamburg." year="2004">Schmid-Egger 2004</bibRefCitation>
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<bibRefCitation author="Schmid-Egger, C" journalOrPublisher="Ampulex" pageId="0" pageNumber="93" pagination="5 - 40" refId="B76" refString="Schmid-Egger, C, 2010. Rote Liste der Wespen Deutschlands. Hymenoptera Aculeata: Grabwespen (Ampulicidae, Crabronidae, Sphecidae), Wegwespen (Pompilidae), Goldwespen (Chrysididae), Faltenwespen (Vespidae), Spinnenameisen (Mutillidae), Dolchwespen (Scoliidae), Rollwespen (Tiphiidae) und Keulhornwespen (Sapygidae). Ampulex 1: 5 - 40" title="Rote Liste der Wespen Deutschlands. Hymenoptera Aculeata: Grabwespen (Ampulicidae, Crabronidae, Sphecidae), Wegwespen (Pompilidae), Goldwespen (Chrysididae), Faltenwespen (Vespidae), Spinnenameisen (Mutillidae), Dolchwespen (Scoliidae), Rollwespen (Tiphiidae) und Keulhornwespen (Sapygidae)." volume="1" year="2010">2010</bibRefCitation>
: 23;
<bibRefCitation author="Smit, J" editor="Peeters, TMJ" journalOrPublisher="Nederlandse Fauna" pageId="0" pageNumber="93" pagination="1 - 507" refId="B80" refString="Smit, J, Peeters, TMJ, Lefeber, V, 2004. Vespidae - plooivleugelwespen, 321-344. In: Peeters, TMJ, van Achterberg, C, Heitmans, WRB, Klein, WF, Lefeber, V, van Loon, AJ, Mabelis, AA, Nieuwenhuijsen, H, Reemer, M, de Rond, J, Smit, J, Velthuis, HHW, Eds., De wespen en mieren van Nederland (Hymenoptera: Aculeata). Nederlandse Fauna 6: 1 - 507" title="Vespidae - plooivleugelwespen, 321 - 344." volume="6" volumeTitle="De wespen en mieren van Nederland (Hymenoptera: Aculeata)." year="2004">Smit et al. 2004</bibRefCitation>
: 327;
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: 41-44;
<bibRefCitation author="Castro, L" journalOrPublisher="Boletin de la SEA" pageId="0" pageNumber="93" pagination="259 - 278" refId="B19" refString="Castro, L, Sanza, F, 2009. Aportacion al conocimiento de los Vespidae (Hymenoptera) de Sierra Nevada (Espana), con algunos comentarios taxonomicos. Boletin de la SEA 45: 259 - 278" title="Aportacion al conocimiento de los Vespidae (Hymenoptera) de Sierra Nevada (Espana), con algunos comentarios taxonomicos." volume="45" year="2009">Castro and Sanza 2009</bibRefCitation>
: 266;
<bibRefCitation author="Abenius, J" editor="Douwes, P" journalOrPublisher="Hymenoptera: Formicidae - Vespidae. Nationalnyckeln till Sveriges flora och fauna. ArtDatabanken, SLU, Uppsala" pageId="0" pageNumber="93" pagination="213 - 275" refId="B1" refString="Abenius, J, 2012. Vespidae, Eumeninae - solitaere getingar. In: Douwes, P, Abenius, J, Cederberg, B, Wahlstedt, U, Eds., Steklar: Myror-getingar. Hymenoptera: Formicidae - Vespidae. Nationalnyckeln till Sveriges flora och fauna. ArtDatabanken, SLU, Uppsala: 213 - 275" title="Vespidae, Eumeninae - solitaere getingar." volumeTitle="Steklar: Myror-getingar." year="2012">Abenius 2012</bibRefCitation>
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<bibRefCitation author="Arens, W" journalOrPublisher="Linzer Biologische Beitrage" pageId="0" pageNumber="93" pagination="481 - 522" refId="B5" refString="Arens, W, 2012. Die solitaeren Faltenwespen der Peloponnes (Hymenoptera: Vespidae: Raphiglossinae, Eumeninae) 1. Teil. Linzer Biologische Beitrage 44 (1): 481 - 522" title="Die solitaeren Faltenwespen der Peloponnes (Hymenoptera: Vespidae: Raphiglossinae, Eumeninae) 1. Teil." volume="44" year="2012">Arens 2012</bibRefCitation>
: 488;
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</paragraph>
<paragraph pageId="0" pageNumber="93">
<taxonomicName authority="papillarius" class="Insecta" family="Vespidae" genus="Eumenes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Eumenes (Eumenes) papillarius subsp. papillarius" order="Hymenoptera" pageId="0" pageNumber="93" phylum="Arthropoda" rank="subSpecies" species="papillarius" subGenus="Eumenes" subSpecies="papillarius">Eumenes (Eumenes) papillarius papillarius</taxonomicName>
;
<bibRefCitation author="van der Vecht, J" journalOrPublisher="Hymenopterorum catalogus nova editio" pageId="0" pageNumber="93" pagination="1 - 200" refId="B84" refString="van der Vecht, J, Fischer, FCJ, 1972. Palaearctic Eumenidae. Hymenopterorum catalogus nova editio 8: 1 - 200" title="Palaearctic Eumenidae." volume="8" year="1972">van der Vecht and Fischer 1972</bibRefCitation>
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<bibRefCitation author="Verges Serra, F" journalOrPublisher="Graellsia" pageId="0" pageNumber="93" pagination="137 - 152" refId="B85" refString="Verges Serra, F, 1985. Estudio monografico de los Eumenes de Espana (Hymenoptera, Eumenidae). Graellsia 41: 137 - 152" title="Estudio monografico de los Eumenes de Espana (Hymenoptera, Eumenidae)." volume="41" year="1985">
<normalizedToken originalValue="Vergés">Verges</normalizedToken>
Serra 1985
</bibRefCitation>
: 143;
<bibRefCitation author="Castro, L" journalOrPublisher="Catalogus de la entomofauna aragonesa" pageId="0" pageNumber="93" pagination="3 - 8" refId="B18" refString="Castro, L, 1997. Familia Vespidae: Subfamilia Eumeninae. Catalogus de la entomofauna aragonesa 16: 3 - 8" title="Familia Vespidae: Subfamilia Eumeninae." volume="16" year="1997">Castro 1997</bibRefCitation>
: 4;
<bibRefCitation author="Sanza, F" journalOrPublisher="Ampulex" pageId="0" pageNumber="93" refId="B74" refString="Sanza, F, 1997. Revision de los eumenidos de la Peninsula Iberica (Hymenoptera: Eumenidae) (tesis doctoral, inedita). Unpublished thesis, Dept de Biologia Animal, Ecologia, Parasitologia, Edafologia y Quimica Agricola de la Universidad de Salamanca, Salamanca, i-vii + 1-541." title="Revision de los eumenidos de la Peninsula Iberica (Hymenoptera: Eumenidae) (tesis doctoral, inedita). Unpublished thesis, Dept de Biologia Animal, Ecologia, Parasitologia, Edafologia y Quimica Agricola de la Universidad de Salamanca, Salamanca, i-vii + 1 - 541." year="1997">Sanza 1997</bibRefCitation>
: 451;
<bibRefCitation author="Gereys, B" journalOrPublisher="Faune de France" pageId="0" pageNumber="93" pagination="1 - 330" refId="B40" refString="Gereys, B, 2016. Vespidae solitaire de France metropolitaine (Hymenoptera: Eumeninae, Masarinae). Faune de France 98: 1 - 330" title="Vespidae solitaire de France metropolitaine (Hymenoptera: Eumeninae, Masarinae)." volume="98" year="2016">Gereys 2016</bibRefCitation>
: 132;
<bibRefCitation DOI="https://doi.org/10.11646/zootaxa.5137.1.1" author="Dal Pos, D" journalOrPublisher="Zootaxa" pageId="0" pageNumber="93" pagination="1 - 111" refId="B21" refString="Dal Pos, D, Carpenter, JM, Uliana, M, 2022. The Giordani Soika collection of Eumeninae at the Natural History Museum of Venice Giancarlo Ligabue: Catalogue of species and type specimens (Insecta, Hymenoptera, Vespidae). Zootaxa 5137 (1): 1 - 111, DOI: https://doi.org/10.11646/zootaxa.5137.1.1" title="The Giordani Soika collection of Eumeninae at the Natural History Museum of Venice Giancarlo Ligabue: Catalogue of species and type specimens (Insecta, Hymenoptera, Vespidae)." url="https://doi.org/10.11646/zootaxa.5137.1.1" volume="5137" year="2022">Dal Pos et al. 2022</bibRefCitation>
: 16.
</paragraph>
<paragraph pageId="0" pageNumber="93">
<taxonomicName authority="papillarius" class="Insecta" family="Vespidae" genus="Eumenes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Eumenes papillarius subsp. papillarius" order="Hymenoptera" pageId="0" pageNumber="93" phylum="Arthropoda" rank="subSpecies" species="papillarius" subSpecies="papillarius">Eumenes papillarius papillarius</taxonomicName>
;
<bibRefCitation author="Gusenleitner, J" journalOrPublisher="Bollettino della Societa Entomologica Italiana" pageId="0" pageNumber="93" refId="B51" refString="Gusenleitner, J, 1972. Uebersicht ueber die derzeit bekannten westpalaearktischen Arten der Gattung Eumenes Latr. 1802 (Hym. Vespoidea). Bollettino del Museo civico di storia naturale di Venezia XXII-XXIII: 67-117." title="Uebersicht ueber die derzeit bekannten westpalaearktischen Arten der Gattung Eumenes Latr. 1802 (Hym. Vespoidea). Bollettino del Museo civico di storia naturale di Venezia XXII-XXIII: 67 - 117." year="1972">Gusenleitner 1972</bibRefCitation>
: 87-88,
<bibRefCitation author="Gusenleitner, J" journalOrPublisher="Linzer Biologische Beitrage" pageId="0" pageNumber="93" pagination="5 - 107" refId="B53" refString="Gusenleitner, J, 2013. Die Gattungen der Eumeninae im Nahen Osten, in Nordafrika und in Arabien (Hymenoptera: Vespidae: Eumeninae). Linzer Biologische Beitrage 45 (1): 5 - 107" title="Die Gattungen der Eumeninae im Nahen Osten, in Nordafrika und in Arabien (Hymenoptera: Vespidae: Eumeninae)." volume="45" year="2013">2013</bibRefCitation>
: 28;
<bibRefCitation author="Borsato, W" journalOrPublisher="Bollettino del Museo Civico di Storia Naturale di Venezia" pageId="0" pageNumber="93" pagination="127 - 150" refId="B14" refString="Borsato, W, Turrisi, GF, 2004. Contributo alla conoscenza degli Eumenidae di Sicilia (Hymenoptera, Vespoidea). Bollettino del Museo Civico di Storia Naturale di Venezia 55: 127 - 150" title="Contributo alla conoscenza degli Eumenidae di Sicilia (Hymenoptera, Vespoidea)." volume="55" year="2004">Borsato and Turrisi 2004</bibRefCitation>
: 144;
<bibRefCitation author="Gogala, A" journalOrPublisher="Scopolia" pageId="0" pageNumber="93" pagination="1 - 79" refId="B48" refString="Gogala, A, 2022. Ose druzine Vespidae v Sloveniji (Hymenoptera: Vespoidea: Vespidae) / Vespid wasps of Slovenia (Hymenoptera: Vespoidea: Vespidae). Scopolia 102: 1 - 79" title="Ose druzine Vespidae v Sloveniji (Hymenoptera: Vespoidea: Vespidae) / Vespid wasps of Slovenia (Hymenoptera: Vespoidea: Vespidae)." volume="102" year="2022">Gogala 2022</bibRefCitation>
: 26-28.
</paragraph>
<paragraph pageId="0" pageNumber="93">
<taxonomicName baseAuthorityName="Christ" baseAuthorityYear="1791" class="Insecta" family="Vespidae" genus="Eumenes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Eumenes (Eumenes) papillarius" order="Hymenoptera" pageId="0" pageNumber="93" phylum="Arthropoda" rank="species" species="papillarius" subGenus="Eumenes">Eumenes (Eumenes) papillarius</taxonomicName>
;
<bibRefCitation author="Fateryga, AV" editor="Belokobylskij, SA" journalOrPublisher="Symphyta and Apocrita: Aculeata" pageId="0" pageNumber="93" pagination="1 - 476" refId="B29" refString="Fateryga, AV, 2017. Subfamily Eumeninae - Potter wasps, 178-192. In: Belokobylskij, SA, Lelej, AS, Eds., Annotated catalogue of the Hymenoptera of Russia. Symphyta and Apocrita: Aculeata 1: 1 - 476" title="Subfamily Eumeninae - Potter wasps, 178 - 192." volume="1" volumeTitle="Annotated catalogue of the Hymenoptera of Russia." year="2017">Fateryga 2017</bibRefCitation>
: 182,
<bibRefCitation author="Fateryga, AV" journalOrPublisher="Symphyta and Apocrita: Aculeata" pageId="0" pageNumber="93" refId="B30" refString="Fateryga, AV, 2018. Wasps of the family Vespidae (Hymenoptera) of the Crimean Peninsula Entomofauna 39: 193-233." title="Wasps of the family Vespidae (Hymenoptera) of the Crimean Peninsula Entomofauna 39: 193 - 233." year="2018">2018</bibRefCitation>
: 207.
</paragraph>
<paragraph pageId="0" pageNumber="93">
<taxonomicName authorityName="de Saussure" authorityYear="1852" class="Insecta" family="Vespidae" genus="Eumenes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Eumenes bipunctis" order="Hymenoptera" pageId="0" pageNumber="93" phylum="Arthropoda" rank="species" species="bipunctis">Eumenes bipunctis</taxonomicName>
de Saussure, 1852: 33;
<bibRefCitation author="van der Vecht, J" journalOrPublisher="Hymenopterorum catalogus nova editio" pageId="0" pageNumber="93" pagination="1 - 200" refId="B84" refString="van der Vecht, J, Fischer, FCJ, 1972. Palaearctic Eumenidae. Hymenopterorum catalogus nova editio 8: 1 - 200" title="Palaearctic Eumenidae." volume="8" year="1972">van der Vecht and Fischer 1972</bibRefCitation>
: 134 (as unidentified species; literature before 1972);
<bibRefCitation author="Gusenleitner, J" journalOrPublisher="Bollettino della Societa Entomologica Italiana" pageId="0" pageNumber="93" refId="B51" refString="Gusenleitner, J, 1972. Uebersicht ueber die derzeit bekannten westpalaearktischen Arten der Gattung Eumenes Latr. 1802 (Hym. Vespoidea). Bollettino del Museo civico di storia naturale di Venezia XXII-XXIII: 67-117." title="Uebersicht ueber die derzeit bekannten westpalaearktischen Arten der Gattung Eumenes Latr. 1802 (Hym. Vespoidea). Bollettino del Museo civico di storia naturale di Venezia XXII-XXIII: 67 - 117." year="1972">Gusenleitner 1972</bibRefCitation>
: 87 (as synonym of
<taxonomicName family="Vespidae" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="E. papillarius" order="Hymenoptera" pageId="0" pageNumber="93" rank="species" species="papillarius">E. papillarius</taxonomicName>
);
<bibRefCitation author="Gereys, B" journalOrPublisher="Faune de France" pageId="0" pageNumber="93" pagination="1 - 330" refId="B40" refString="Gereys, B, 2016. Vespidae solitaire de France metropolitaine (Hymenoptera: Eumeninae, Masarinae). Faune de France 98: 1 - 330" title="Vespidae solitaire de France metropolitaine (Hymenoptera: Eumeninae, Masarinae)." volume="98" year="2016">Gereys 2016</bibRefCitation>
: 132 (id.);
<bibRefCitation author="Fateryga, AV" editor="Belokobylskij, SA" journalOrPublisher="Symphyta and Apocrita: Aculeata" pageId="0" pageNumber="93" pagination="1 - 476" refId="B29" refString="Fateryga, AV, 2017. Subfamily Eumeninae - Potter wasps, 178-192. In: Belokobylskij, SA, Lelej, AS, Eds., Annotated catalogue of the Hymenoptera of Russia. Symphyta and Apocrita: Aculeata 1: 1 - 476" title="Subfamily Eumeninae - Potter wasps, 178 - 192." volume="1" volumeTitle="Annotated catalogue of the Hymenoptera of Russia." year="2017">Fateryga 2017</bibRefCitation>
: 182 (id.).
</paragraph>
<paragraph pageId="0" pageNumber="93">
<taxonomicName authorityName="Andre" authorityYear="1884" class="Insecta" family="Vespidae" genus="Eumenes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Eumenes bimaculatus" order="Hymenoptera" pageId="0" pageNumber="93" phylum="Arthropoda" rank="species" species="bimaculatus">Eumenes bimaculatus</taxonomicName>
<normalizedToken originalValue="André">Andre</normalizedToken>
, 1884: 645;
<bibRefCitation author="van der Vecht, J" journalOrPublisher="Hymenopterorum catalogus nova editio" pageId="0" pageNumber="93" pagination="1 - 200" refId="B84" refString="van der Vecht, J, Fischer, FCJ, 1972. Palaearctic Eumenidae. Hymenopterorum catalogus nova editio 8: 1 - 200" title="Palaearctic Eumenidae." volume="8" year="1972">van der Vecht and Fischer 1972</bibRefCitation>
: 130 (as synonym of
<taxonomicName family="Vespidae" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="E. papillarius" order="Hymenoptera" pageId="0" pageNumber="93" rank="species" species="papillarius">E. papillarius</taxonomicName>
; literature before 1972):
<bibRefCitation author="Gusenleitner, J" journalOrPublisher="Bollettino della Societa Entomologica Italiana" pageId="0" pageNumber="93" refId="B51" refString="Gusenleitner, J, 1972. Uebersicht ueber die derzeit bekannten westpalaearktischen Arten der Gattung Eumenes Latr. 1802 (Hym. Vespoidea). Bollettino del Museo civico di storia naturale di Venezia XXII-XXIII: 67-117." title="Uebersicht ueber die derzeit bekannten westpalaearktischen Arten der Gattung Eumenes Latr. 1802 (Hym. Vespoidea). Bollettino del Museo civico di storia naturale di Venezia XXII-XXIII: 67 - 117." year="1972">Gusenleitner 1972</bibRefCitation>
: 87 (as synonym of
<taxonomicName family="Vespidae" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="E. papillarius" order="Hymenoptera" pageId="0" pageNumber="93" rank="species" species="papillarius">E. papillarius</taxonomicName>
);
<bibRefCitation author="Gereys, B" journalOrPublisher="Faune de France" pageId="0" pageNumber="93" pagination="1 - 330" refId="B40" refString="Gereys, B, 2016. Vespidae solitaire de France metropolitaine (Hymenoptera: Eumeninae, Masarinae). Faune de France 98: 1 - 330" title="Vespidae solitaire de France metropolitaine (Hymenoptera: Eumeninae, Masarinae)." volume="98" year="2016">Gereys 2016</bibRefCitation>
: 132 (id.);
<bibRefCitation author="Fateryga, AV" editor="Belokobylskij, SA" journalOrPublisher="Symphyta and Apocrita: Aculeata" pageId="0" pageNumber="93" pagination="1 - 476" refId="B29" refString="Fateryga, AV, 2017. Subfamily Eumeninae - Potter wasps, 178-192. In: Belokobylskij, SA, Lelej, AS, Eds., Annotated catalogue of the Hymenoptera of Russia. Symphyta and Apocrita: Aculeata 1: 1 - 476" title="Subfamily Eumeninae - Potter wasps, 178 - 192." volume="1" volumeTitle="Annotated catalogue of the Hymenoptera of Russia." year="2017">Fateryga 2017</bibRefCitation>
: 182 (id.).
</paragraph>
<paragraph pageId="0" pageNumber="93">
<taxonomicName class="Insecta" family="Vespidae" genus="Eumenes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Eumenes papillarius subsp. var. var. baltica" order="Hymenoptera" pageId="0" pageNumber="93" phylum="Arthropoda" rank="variety" species="papillarius" subSpecies="var." variety="baltica">Eumenes papillarius var. baltica</taxonomicName>
<normalizedToken originalValue="Blüthgen">Bluethgen</normalizedToken>
, 1938: 485;
<bibRefCitation author="Fateryga, AV" editor="Belokobylskij, SA" journalOrPublisher="Symphyta and Apocrita: Aculeata" pageId="0" pageNumber="93" pagination="1 - 476" refId="B29" refString="Fateryga, AV, 2017. Subfamily Eumeninae - Potter wasps, 178-192. In: Belokobylskij, SA, Lelej, AS, Eds., Annotated catalogue of the Hymenoptera of Russia. Symphyta and Apocrita: Aculeata 1: 1 - 476" title="Subfamily Eumeninae - Potter wasps, 178 - 192." volume="1" volumeTitle="Annotated catalogue of the Hymenoptera of Russia." year="2017">Fateryga 2017</bibRefCitation>
: 182 (as synonym of
<taxonomicName family="Vespidae" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="E. papillarius" order="Hymenoptera" pageId="0" pageNumber="93" rank="species" species="papillarius">E. papillarius</taxonomicName>
).
</paragraph>
<paragraph pageId="0" pageNumber="93">
<taxonomicName authorityName="Bluthgen" authorityYear="1938" class="Insecta" family="Vespidae" genus="Eumenes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Eumenes (Eumenes) papillarius subsp. balticus" order="Hymenoptera" pageId="0" pageNumber="93" phylum="Arthropoda" rank="subSpecies" species="papillarius" subGenus="Eumenes" subSpecies="balticus">Eumenes (Eumenes) papillarius balticus</taxonomicName>
;
<bibRefCitation author="van der Vecht, J" journalOrPublisher="Hymenopterorum catalogus nova editio" pageId="0" pageNumber="93" pagination="1 - 200" refId="B84" refString="van der Vecht, J, Fischer, FCJ, 1972. Palaearctic Eumenidae. Hymenopterorum catalogus nova editio 8: 1 - 200" title="Palaearctic Eumenidae." volume="8" year="1972">van der Vecht and Fischer 1972</bibRefCitation>
: 130 (literature before 1972);
<bibRefCitation DOI="https://doi.org/10.11646/zootaxa.5137.1.1" author="Dal Pos, D" journalOrPublisher="Zootaxa" pageId="0" pageNumber="93" pagination="1 - 111" refId="B21" refString="Dal Pos, D, Carpenter, JM, Uliana, M, 2022. The Giordani Soika collection of Eumeninae at the Natural History Museum of Venice Giancarlo Ligabue: Catalogue of species and type specimens (Insecta, Hymenoptera, Vespidae). Zootaxa 5137 (1): 1 - 111, DOI: https://doi.org/10.11646/zootaxa.5137.1.1" title="The Giordani Soika collection of Eumeninae at the Natural History Museum of Venice Giancarlo Ligabue: Catalogue of species and type specimens (Insecta, Hymenoptera, Vespidae)." url="https://doi.org/10.11646/zootaxa.5137.1.1" volume="5137" year="2022">Dal Pos et al. 2022</bibRefCitation>
: 16.
</paragraph>
<paragraph pageId="0" pageNumber="93">
<taxonomicName authorityName="Bluthgen" authorityYear="1938" class="Insecta" family="Vespidae" genus="Eumenes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Eumenes papillarius subsp. balticus" order="Hymenoptera" pageId="0" pageNumber="93" phylum="Arthropoda" rank="subSpecies" species="papillarius" subSpecies="balticus">Eumenes papillarius balticus</taxonomicName>
;
<bibRefCitation author="Gusenleitner, J" journalOrPublisher="Bollettino della Societa Entomologica Italiana" pageId="0" pageNumber="93" refId="B51" refString="Gusenleitner, J, 1972. Uebersicht ueber die derzeit bekannten westpalaearktischen Arten der Gattung Eumenes Latr. 1802 (Hym. Vespoidea). Bollettino del Museo civico di storia naturale di Venezia XXII-XXIII: 67-117." title="Uebersicht ueber die derzeit bekannten westpalaearktischen Arten der Gattung Eumenes Latr. 1802 (Hym. Vespoidea). Bollettino del Museo civico di storia naturale di Venezia XXII-XXIII: 67 - 117." year="1972">Gusenleitner 1972</bibRefCitation>
: 89-90,
<bibRefCitation author="Gusenleitner, J" journalOrPublisher="Linzer Biologische Beitrage" pageId="0" pageNumber="93" pagination="561 - 584" refId="B52" refString="Gusenleitner, J, 1999. Bestimmungstabellen mittel- und suedeuropaeischer Eumeniden (Vespoidea, Hymenoptera) Teil 11. Die Gattungen Discoelius Latreille 1809, Eumenes Latreille 1802, Katamenes Meade-Waldo 1910, Delta Saussure 1855, Ischnogasteroides Magretti 1884 und Pareumenes Saussure 1855. Linzer Biologische Beitrage 31 (2): 561 - 584" title="Bestimmungstabellen mittel- und suedeuropaeischer Eumeniden (Vespoidea, Hymenoptera) Teil 11. Die Gattungen Discoelius Latreille 1809, Eumenes Latreille 1802, Katamenes Meade-Waldo 1910, Delta Saussure 1855, Ischnogasteroides Magretti 1884 und Pareumenes Saussure 1855." volume="31" year="1999">1999</bibRefCitation>
: 572.
</paragraph>
<paragraph pageId="0" pageNumber="93">
<taxonomicName class="Insecta" family="Vespidae" genus="Eumenes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Eumenes mediterraneus subsp. aemilianus" order="Hymenoptera" pageId="0" pageNumber="93" phylum="Arthropoda" rank="subSpecies" species="mediterraneus" subSpecies="aemilianus">Eumenes mediterraneus aemilianus</taxonomicName>
Guiglia, 1951: 28. Syn. nov.
</paragraph>
<paragraph pageId="0" pageNumber="93">
<taxonomicName class="Insecta" family="Vespidae" genus="Eumenes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Eumenes (Eumenes) mediterraneus subsp. aemilianus" order="Hymenoptera" pageId="0" pageNumber="93" phylum="Arthropoda" rank="subSpecies" species="mediterraneus" subGenus="Eumenes" subSpecies="aemilianus">Eumenes (Eumenes) mediterraneus aemilianus</taxonomicName>
;
<bibRefCitation author="van der Vecht, J" journalOrPublisher="Hymenopterorum catalogus nova editio" pageId="0" pageNumber="93" pagination="1 - 200" refId="B84" refString="van der Vecht, J, Fischer, FCJ, 1972. Palaearctic Eumenidae. Hymenopterorum catalogus nova editio 8: 1 - 200" title="Palaearctic Eumenidae." volume="8" year="1972">van der Vecht and Fischer 1972</bibRefCitation>
: 125 (literature before 1972).
</paragraph>
<paragraph pageId="0" pageNumber="93">
<taxonomicName authorityName="Guiglia" authorityYear="1951" class="Insecta" family="Vespidae" genus="Eumenes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Eumenes aemilianus" order="Hymenoptera" pageId="0" pageNumber="93" phylum="Arthropoda" rank="species" species="aemilianus">Eumenes aemilianus</taxonomicName>
;
<bibRefCitation author="Gusenleitner, J" journalOrPublisher="Bollettino della Societa Entomologica Italiana" pageId="0" pageNumber="93" refId="B51" refString="Gusenleitner, J, 1972. Uebersicht ueber die derzeit bekannten westpalaearktischen Arten der Gattung Eumenes Latr. 1802 (Hym. Vespoidea). Bollettino del Museo civico di storia naturale di Venezia XXII-XXIII: 67-117." title="Uebersicht ueber die derzeit bekannten westpalaearktischen Arten der Gattung Eumenes Latr. 1802 (Hym. Vespoidea). Bollettino del Museo civico di storia naturale di Venezia XXII-XXIII: 67 - 117." year="1972">Gusenleitner 1972</bibRefCitation>
: 95-96,
<bibRefCitation author="Gusenleitner, J" journalOrPublisher="Linzer Biologische Beitrage" pageId="0" pageNumber="93" pagination="561 - 584" refId="B52" refString="Gusenleitner, J, 1999. Bestimmungstabellen mittel- und suedeuropaeischer Eumeniden (Vespoidea, Hymenoptera) Teil 11. Die Gattungen Discoelius Latreille 1809, Eumenes Latreille 1802, Katamenes Meade-Waldo 1910, Delta Saussure 1855, Ischnogasteroides Magretti 1884 und Pareumenes Saussure 1855. Linzer Biologische Beitrage 31 (2): 561 - 584" title="Bestimmungstabellen mittel- und suedeuropaeischer Eumeniden (Vespoidea, Hymenoptera) Teil 11. Die Gattungen Discoelius Latreille 1809, Eumenes Latreille 1802, Katamenes Meade-Waldo 1910, Delta Saussure 1855, Ischnogasteroides Magretti 1884 und Pareumenes Saussure 1855." volume="31" year="1999">1999</bibRefCitation>
: 569;
<bibRefCitation author="Borsato, W" journalOrPublisher="Bollettino del Museo Civico di Storia Naturale di Venezia" pageId="0" pageNumber="93" pagination="123 - 147" refId="B13" refString="Borsato, W, 2006. Contributo alla conoscenza degli Eumenidae di Sardegna e note sulla fauna corsa (Hymenoptera, Vespoidea). Bollettino del Museo Civico di Storia Naturale di Venezia 57: 123 - 147" title="Contributo alla conoscenza degli Eumenidae di Sardegna e note sulla fauna corsa (Hymenoptera, Vespoidea)." volume="57" year="2006">Borsato 2006</bibRefCitation>
: 140.
</paragraph>
<paragraph pageId="0" pageNumber="93">
<taxonomicName authorityName="Bluthgen" authorityYear="1956" class="Insecta" family="Vespidae" genus="Eumenes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Eumenes papillarius subsp. monticola" order="Hymenoptera" pageId="0" pageNumber="93" phylum="Arthropoda" rank="subSpecies" species="papillarius" subSpecies="monticola">Eumenes papillarius monticola</taxonomicName>
<normalizedToken originalValue="Blüthgen">Bluethgen</normalizedToken>
, 1956: 2;
<bibRefCitation author="van der Vecht, J" journalOrPublisher="Hymenopterorum catalogus nova editio" pageId="0" pageNumber="93" pagination="1 - 200" refId="B84" refString="van der Vecht, J, Fischer, FCJ, 1972. Palaearctic Eumenidae. Hymenopterorum catalogus nova editio 8: 1 - 200" title="Palaearctic Eumenidae." volume="8" year="1972">van der Vecht and Fischer 1972</bibRefCitation>
: 130;
<bibRefCitation author="Castro, L" journalOrPublisher="Catalogus de la entomofauna aragonesa" pageId="0" pageNumber="93" pagination="3 - 8" refId="B18" refString="Castro, L, 1997. Familia Vespidae: Subfamilia Eumeninae. Catalogus de la entomofauna aragonesa 16: 3 - 8" title="Familia Vespidae: Subfamilia Eumeninae." volume="16" year="1997">Castro 1997</bibRefCitation>
: 4 (as synonym of
<taxonomicName family="Vespidae" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="E. papillarius" order="Hymenoptera" pageId="0" pageNumber="93" rank="species" species="papillarius">E. papillarius</taxonomicName>
);
<bibRefCitation author="Gereys, B" journalOrPublisher="Faune de France" pageId="0" pageNumber="93" pagination="1 - 330" refId="B40" refString="Gereys, B, 2016. Vespidae solitaire de France metropolitaine (Hymenoptera: Eumeninae, Masarinae). Faune de France 98: 1 - 330" title="Vespidae solitaire de France metropolitaine (Hymenoptera: Eumeninae, Masarinae)." volume="98" year="2016">Gereys 2016</bibRefCitation>
: 132 (id.);
<bibRefCitation author="Fateryga, AV" editor="Belokobylskij, SA" journalOrPublisher="Symphyta and Apocrita: Aculeata" pageId="0" pageNumber="93" pagination="1 - 476" refId="B29" refString="Fateryga, AV, 2017. Subfamily Eumeninae - Potter wasps, 178-192. In: Belokobylskij, SA, Lelej, AS, Eds., Annotated catalogue of the Hymenoptera of Russia. Symphyta and Apocrita: Aculeata 1: 1 - 476" title="Subfamily Eumeninae - Potter wasps, 178 - 192." volume="1" volumeTitle="Annotated catalogue of the Hymenoptera of Russia." year="2017">Fateryga 2017</bibRefCitation>
: 182 (id.).
</paragraph>
<paragraph pageId="0" pageNumber="93">
<taxonomicName authorityName="Gusenleitner" authorityYear="1972" class="Insecta" family="Vespidae" genus="Eumenes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Eumenes papillarius subsp. rubricornis" order="Hymenoptera" pageId="0" pageNumber="93" phylum="Arthropoda" rank="subSpecies" species="papillarius" subSpecies="rubricornis">Eumenes papillarius rubricornis</taxonomicName>
Giordani Soika (in Gusenleitner), 1972: 90;
<bibRefCitation author="Gusenleitner, J" journalOrPublisher="Linzer Biologische Beitrage" pageId="0" pageNumber="93" pagination="561 - 584" refId="B52" refString="Gusenleitner, J, 1999. Bestimmungstabellen mittel- und suedeuropaeischer Eumeniden (Vespoidea, Hymenoptera) Teil 11. Die Gattungen Discoelius Latreille 1809, Eumenes Latreille 1802, Katamenes Meade-Waldo 1910, Delta Saussure 1855, Ischnogasteroides Magretti 1884 und Pareumenes Saussure 1855. Linzer Biologische Beitrage 31 (2): 561 - 584" title="Bestimmungstabellen mittel- und suedeuropaeischer Eumeniden (Vespoidea, Hymenoptera) Teil 11. Die Gattungen Discoelius Latreille 1809, Eumenes Latreille 1802, Katamenes Meade-Waldo 1910, Delta Saussure 1855, Ischnogasteroides Magretti 1884 und Pareumenes Saussure 1855." volume="31" year="1999">Gusenleitner 1999</bibRefCitation>
: 572,
<bibRefCitation author="Gusenleitner, J" journalOrPublisher="Linzer Biologische Beitrage" pageId="0" pageNumber="93" pagination="5 - 107" refId="B53" refString="Gusenleitner, J, 2013. Die Gattungen der Eumeninae im Nahen Osten, in Nordafrika und in Arabien (Hymenoptera: Vespidae: Eumeninae). Linzer Biologische Beitrage 45 (1): 5 - 107" title="Die Gattungen der Eumeninae im Nahen Osten, in Nordafrika und in Arabien (Hymenoptera: Vespidae: Eumeninae)." volume="45" year="2013">2013</bibRefCitation>
: 28;
<bibRefCitation author="Fateryga, AV" editor="Belokobylskij, SA" journalOrPublisher="Symphyta and Apocrita: Aculeata" pageId="0" pageNumber="93" pagination="1 - 476" refId="B29" refString="Fateryga, AV, 2017. Subfamily Eumeninae - Potter wasps, 178-192. In: Belokobylskij, SA, Lelej, AS, Eds., Annotated catalogue of the Hymenoptera of Russia. Symphyta and Apocrita: Aculeata 1: 1 - 476" title="Subfamily Eumeninae - Potter wasps, 178 - 192." volume="1" volumeTitle="Annotated catalogue of the Hymenoptera of Russia." year="2017">Fateryga 2017</bibRefCitation>
: 182 (as synonym of
<taxonomicName family="Vespidae" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="E. papillarius" order="Hymenoptera" pageId="0" pageNumber="93" rank="species" species="papillarius">E. papillarius</taxonomicName>
).
</paragraph>
<paragraph pageId="0" pageNumber="93">
<taxonomicName authorityName="Gusenleitner" authorityYear="1972" class="Insecta" family="Vespidae" genus="Eumenes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Eumenes (Eumenes) papillarius subsp. rubricornis" order="Hymenoptera" pageId="0" pageNumber="93" phylum="Arthropoda" rank="subSpecies" species="papillarius" subGenus="Eumenes" subSpecies="rubricornis">Eumenes (Eumenes) papillarius rubricornis</taxonomicName>
;
<bibRefCitation DOI="https://doi.org/10.11646/zootaxa.5137.1.1" author="Dal Pos, D" journalOrPublisher="Zootaxa" pageId="0" pageNumber="93" pagination="1 - 111" refId="B21" refString="Dal Pos, D, Carpenter, JM, Uliana, M, 2022. The Giordani Soika collection of Eumeninae at the Natural History Museum of Venice Giancarlo Ligabue: Catalogue of species and type specimens (Insecta, Hymenoptera, Vespidae). Zootaxa 5137 (1): 1 - 111, DOI: https://doi.org/10.11646/zootaxa.5137.1.1" title="The Giordani Soika collection of Eumeninae at the Natural History Museum of Venice Giancarlo Ligabue: Catalogue of species and type specimens (Insecta, Hymenoptera, Vespidae)." url="https://doi.org/10.11646/zootaxa.5137.1.1" volume="5137" year="2022">Dal Pos et al. 2022</bibRefCitation>
: 16.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="93" type="notes">
<paragraph pageId="0" pageNumber="93">Notes.</paragraph>
<paragraph pageId="0" pageNumber="93">
Large specimens (fore wing length about 10 mm) have frequently a pair of yellow patches on the mesoscutum antero-laterally; the patches vary from minute to large. The photographs of the female holotype kindly supplied by Roberto Poggi (MSNG) show that
<taxonomicName family="Vespidae" kingdom="Animalia" lsidName="E. aemilianus" order="Hymenoptera" pageId="0" pageNumber="93" rank="species" species="aemilianus">
<emphasis italics="true" pageId="0" pageNumber="93">E. aemilianus</emphasis>
</taxonomicName>
Guiglia, 1951 is a junior synonym of
<taxonomicName family="Vespidae" kingdom="Animalia" lsidName="E. papillarius" order="Hymenoptera" pageId="0" pageNumber="93" rank="species" species="papillarius">
<emphasis italics="true" pageId="0" pageNumber="93">E. papillarius</emphasis>
</taxonomicName>
(Christ, 1791) (syn. nov.) because of the entirely dark brown labrum (Fig.
<figureCitation captionStart="Figures 262–266" captionStartId="F32" captionText="Figures 262 - 266. Eumenes mediterraneus aemilianus Guiglia, holotype, Italy, female 262 metasoma lateral 263 propodeum and first metasomal tergite dorsal 264 head and mesosoma dorsal 265 head anterior 266 head and mesosoma lateral. Photographs: R. Poggi (MSNG)." figureDoi="10.3897/zookeys.1143.94951.figures262-266" httpUri="https://binary.pensoft.net/fig/803455" pageId="0" pageNumber="93">265</figureCitation>
), the shallow apical emargination of the clypeus (Fig.
<figureCitation captionStart="Figures 262–266" captionStartId="F32" captionText="Figures 262 - 266. Eumenes mediterraneus aemilianus Guiglia, holotype, Italy, female 262 metasoma lateral 263 propodeum and first metasomal tergite dorsal 264 head and mesosoma dorsal 265 head anterior 266 head and mesosoma lateral. Photographs: R. Poggi (MSNG)." figureDoi="10.3897/zookeys.1143.94951.figures262-266" httpUri="https://binary.pensoft.net/fig/803455" pageId="0" pageNumber="93">265</figureCitation>
), the pair of large yellow patches on the mesoscutum (Fig.
<figureCitation captionStart="Figures 262–266" captionStartId="F32" captionText="Figures 262 - 266. Eumenes mediterraneus aemilianus Guiglia, holotype, Italy, female 262 metasoma lateral 263 propodeum and first metasomal tergite dorsal 264 head and mesosoma dorsal 265 head anterior 266 head and mesosoma lateral. Photographs: R. Poggi (MSNG)." figureDoi="10.3897/zookeys.1143.94951.figures262-266" httpUri="https://binary.pensoft.net/fig/803455" pageId="0" pageNumber="93">264</figureCitation>
) and the medium-sized setosity of the second tergite (mentioned in the original description). The holotype female (deposited in MSNG) is a comparatively yellowish specimen with slender first tergite, it has the clypeus narrowly black apically, the mesoscutum with a pair of large and curved yellow lateral patches, the apex of the antenna black and the apical rim of the second tergite pale yellowish (Figs
<figureCitation captionStart="Figures 262–266" captionStartId="F32" captionText="Figures 262 - 266. Eumenes mediterraneus aemilianus Guiglia, holotype, Italy, female 262 metasoma lateral 263 propodeum and first metasomal tergite dorsal 264 head and mesosoma dorsal 265 head anterior 266 head and mesosoma lateral. Photographs: R. Poggi (MSNG)." figureDoi="10.3897/zookeys.1143.94951.figures262-266" httpUri="https://binary.pensoft.net/fig/803455" pageId="0" pageNumber="93">262-266</figureCitation>
). The presence or absence of a pair of yellow mesoscutal patches is a variable (and at least partly size related) character in
<taxonomicName family="Vespidae" kingdom="Animalia" lsidName="E. papillarius" order="Hymenoptera" pageId="0" pageNumber="93" rank="species" species="papillarius">
<emphasis italics="true" pageId="0" pageNumber="93">E. papillarius</emphasis>
</taxonomicName>
and does not indicate that it concerns a separate species (see also comments of
<bibRefCitation author="Gusenleitner, J" journalOrPublisher="Bollettino della Societa Entomologica Italiana" pageId="0" pageNumber="93" refId="B51" refString="Gusenleitner, J, 1972. Uebersicht ueber die derzeit bekannten westpalaearktischen Arten der Gattung Eumenes Latr. 1802 (Hym. Vespoidea). Bollettino del Museo civico di storia naturale di Venezia XXII-XXIII: 67-117." title="Uebersicht ueber die derzeit bekannten westpalaearktischen Arten der Gattung Eumenes Latr. 1802 (Hym. Vespoidea). Bollettino del Museo civico di storia naturale di Venezia XXII-XXIII: 67 - 117." year="1972">Gusenleitner (1972)</bibRefCitation>
under
<taxonomicName family="Vespidae" kingdom="Animalia" lsidName="E. papillarius" order="Hymenoptera" pageId="0" pageNumber="93" rank="species" species="papillarius">
<emphasis italics="true" pageId="0" pageNumber="93">E. papillarius</emphasis>
</taxonomicName>
). Aberrant specimens from the Iberian Peninsula have medium-sized to long setae on the second sternite and the apex of the hind tibia is yellow. The apical lamella of the second tergite varies from pale yellowish to dark brown.
</paragraph>
<caption doi="10.3897/zookeys.1143.94951.figures112-121" httpUri="https://binary.pensoft.net/fig/803439" pageId="0" pageNumber="93" start="Figures 112–121" startId="F16">
<paragraph pageId="0" pageNumber="93">
<emphasis bold="true" pageId="0" pageNumber="93">Figures 112-121.</emphasis>
<taxonomicName baseAuthorityName="Christ" baseAuthorityYear="1791" class="Insecta" family="Vespidae" genus="Eumenes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Eumenes papillarius" order="Hymenoptera" pageId="0" pageNumber="93" phylum="Arthropoda" rank="species" species="papillarius">
<emphasis italics="true" pageId="0" pageNumber="93">Eumenes papillarius</emphasis>
</taxonomicName>
(Christ), France, female
<emphasis bold="true" pageId="0" pageNumber="93">112</emphasis>
metasoma lateral
<emphasis bold="true" pageId="0" pageNumber="93">113</emphasis>
first metasomal tergite dorsal
<emphasis bold="true" pageId="0" pageNumber="93">114</emphasis>
first tergite ventral
<emphasis bold="true" pageId="0" pageNumber="93">115</emphasis>
head and mesosoma dorsal
<emphasis bold="true" pageId="0" pageNumber="93">116</emphasis>
second tergite latero-dorsal
<emphasis bold="true" pageId="0" pageNumber="93">117</emphasis>
head anterior
<emphasis bold="true" pageId="0" pageNumber="93">118</emphasis>
head and pronotum lateral
<emphasis bold="true" pageId="0" pageNumber="93">119</emphasis>
antenna anterior
<emphasis bold="true" pageId="0" pageNumber="93">120</emphasis>
hind tarsal claw
<emphasis bold="true" pageId="0" pageNumber="93">121</emphasis>
hind tibia lateral.
</paragraph>
</caption>
<caption doi="10.3897/zookeys.1143.94951.figures122-131" httpUri="https://binary.pensoft.net/fig/803440" pageId="0" pageNumber="93" start="Figures 122–131" startId="F17">
<paragraph pageId="0" pageNumber="93">
<emphasis bold="true" pageId="0" pageNumber="93">Figures 122-131.</emphasis>
<taxonomicName baseAuthorityName="Christ" baseAuthorityYear="1791" class="Insecta" family="Vespidae" genus="Eumenes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Eumenes papillarius" order="Hymenoptera" pageId="0" pageNumber="93" phylum="Arthropoda" rank="species" species="papillarius">
<emphasis italics="true" pageId="0" pageNumber="93">Eumenes papillarius</emphasis>
</taxonomicName>
(Christ), France, male
<emphasis bold="true" pageId="0" pageNumber="93">122</emphasis>
metasoma lateral
<emphasis bold="true" pageId="0" pageNumber="93">123</emphasis>
first metasomal tergite dorsal
<emphasis bold="true" pageId="0" pageNumber="93">124</emphasis>
first tergite ventral
<emphasis bold="true" pageId="0" pageNumber="93">125</emphasis>
head and mesosoma dorsal
<emphasis bold="true" pageId="0" pageNumber="93">126</emphasis>
second tergite dorsal
<emphasis bold="true" pageId="0" pageNumber="93">127</emphasis>
hind tibia lateral
<emphasis bold="true" pageId="0" pageNumber="93">128</emphasis>
head anterior
<emphasis bold="true" pageId="0" pageNumber="93">129</emphasis>
head and mesosoma lateral
<emphasis bold="true" pageId="0" pageNumber="93">130</emphasis>
antenna anterior
<emphasis bold="true" pageId="0" pageNumber="93">131</emphasis>
apical hook of antenna lateral.
</paragraph>
</caption>
</subSubSection>
<subSubSection pageId="0" pageNumber="93" type="distribution">
<paragraph pageId="0" pageNumber="93">Distribution.</paragraph>
<paragraph pageId="0" pageNumber="93">
Widespread in most of Europe, but considered absent from UK (only known as vagrant in England;
<bibRefCitation author="Archer, ME" journalOrPublisher="Species des hymenopteres d'Europe &amp; d'Algerie" pageId="0" pageNumber="93" refId="B4" refString="Archer, ME, 2014. The vespoid wasps (Tiphiidae, Mutillidae, Sapygidae, Scoliidae and Vespidae) of the British Isles. Handbooks for the identification of British Insects 6(6): [i-vi +] 1-82." title="The vespoid wasps (Tiphiidae, Mutillidae, Sapygidae, Scoliidae and Vespidae) of the British Isles. Handbooks for the identification of British Insects 6 (6): [i-vi +] 1 - 82." year="2014">Archer 2014</bibRefCitation>
), Norway, Sweden and Finland. Found up to 1550 m altitude in Switzerland (
<bibRefCitation author="Neumeyer, R" journalOrPublisher="Fauna Helvetica" pageId="0" pageNumber="93" pagination="1 - 381" refId="B69" refString="Neumeyer, R, 2019. Vespidae. Fauna Helvetica 31: 1 - 381" title="Vespidae." volume="31" year="2019">Neumeyer 2019</bibRefCitation>
), but in S Greece rather rare and restricted to lowland and submontane habitats (
<bibRefCitation author="Arens, W" journalOrPublisher="Linzer Biologische Beitrage" pageId="0" pageNumber="93" pagination="481 - 522" refId="B5" refString="Arens, W, 2012. Die solitaeren Faltenwespen der Peloponnes (Hymenoptera: Vespidae: Raphiglossinae, Eumeninae) 1. Teil. Linzer Biologische Beitrage 44 (1): 481 - 522" title="Die solitaeren Faltenwespen der Peloponnes (Hymenoptera: Vespidae: Raphiglossinae, Eumeninae) 1. Teil." volume="44" year="2012">Arens 2012</bibRefCitation>
). This species is often found near human settlements and using small crevices of buildings (e.g., under roof tiles) to construct small groups of clay nests.
</paragraph>
</subSubSection>
</treatment>
</document>
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