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<document id="0F70D1F218C8E57E57B7D3AD23D9D1EF" ID-CLB-Dataset="29346" ID-DOI="10.11646/zootaxa.4446.2.7" ID-GBIF-Dataset="475a54d9-e98e-4be3-8f56-0af7063cf66a" ID-ISSN="1175-5326" ID-Zenodo-Dep="1444173" ID-ZooBank="78204F54-9DF7-4807-84A4-6954F4197AA0" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1538626174513" checkinUser="plazi" docAuthor="Stec, Daniel, Kristensen, Reinhardt Møbjerg &amp; Michalczyk, Łukasz" docDate="2018" docId="196C5B1C784FFF8BFF5DFC0EB3F771E4" docLanguage="en" docName="zootaxa.4446.2.7.pdf" docOrigin="Zootaxa 4446 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Macrobiotus papei Stec &amp; Kristensen &amp; Michalczyk 2018, sp. nov." docType="treatment" docVersion="7" lastPageNumber="287" masterDocId="E5552364784BFF85FFCAFFB1B03C732C" masterDocTitle="Integrative taxonomy identifies Macrobiotus papei, a new tardigrade species of the Macrobiotus hufelandi complex (Eutardigrada: Macrobiotidae) from the Udzungwa Mountains National Park (Tanzania)" masterLastPageNumber="291" masterPageNumber="273" pageNumber="277" updateTime="1698562572642" updateUser="ExternalLinkService">
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<mods:title id="2F78A9CFD969A0056DDD3DE55B604BC1">Integrative taxonomy identifies Macrobiotus papei, a new tardigrade species of the Macrobiotus hufelandi complex (Eutardigrada: Macrobiotidae) from the Udzungwa Mountains National Park (Tanzania)</mods:title>
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<mods:namePart id="4E9014EA5236FF7F785ABD7CE30A4E8C">Stec, Daniel</mods:namePart>
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<mods:namePart id="332FE47AE68DFE3B2E2FD0DBFD589957">Kristensen, Reinhardt Møbjerg</mods:namePart>
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<mods:namePart id="DE881CF2699B94F05F10939E259978C8">Michalczyk, Łukasz</mods:namePart>
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<mods:title id="9B9D08A7728BABE5A815522B7CA57FFF">Zootaxa</mods:title>
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<mods:date id="C33DD3A94E7A41C1A540BF93D5F05831">2018</mods:date>
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<mods:number id="87A488EE7402B5DBD8C58AEE9DF3A0AB">2018-07-17</mods:number>
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<mods:number id="76530E83000ED425E7CE20AC91CEB388">4446</mods:number>
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<mods:classification id="99F2B56C6444EAEF24A16585A3F6F549">journal article</mods:classification>
<mods:identifier id="A29D9BB0CBA0A9C8632399436D5FAA36" type="CLB-Dataset">29346</mods:identifier>
<mods:identifier id="F4B27207BEF8AF3B8F72F7A513E802A8" type="DOI">10.11646/zootaxa.4446.2.7</mods:identifier>
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<treatment id="196C5B1C784FFF8BFF5DFC0EB3F771E4" ID-DOI="http://doi.org/10.5281/zenodo.5963664" ID-GBIF-Taxon="148456414" ID-Zenodo-Dep="5963664" LSID="urn:lsid:plazi:treatment:196C5B1C784FFF8BFF5DFC0EB3F771E4" httpUri="http://treatment.plazi.org/id/196C5B1C784FFF8BFF5DFC0EB3F771E4" lastPageId="14" lastPageNumber="287" pageId="4" pageNumber="277">
<subSubSection id="D9DFB981784FFF81FF5DFC0EB18A70D0" pageId="4" pageNumber="277" type="nomenclature">
<paragraph id="917AEA0A784FFF81FF5DFC0EB1F370F4" blockId="4.[151,463,959,1021]" box="[151,463,959,984]" pageId="4" pageNumber="277">
<heading id="CA325D66784FFF81FF5DFC0EB1F370F4" bold="true" box="[151,463,959,984]" fontSize="10" level="2" pageId="4" pageNumber="277" reason="2">
<emphasis id="A3B13618784FFF81FF5DFC0EB1F370F4" bold="true" box="[151,463,959,984]" pageId="4" pageNumber="277">
<taxonomicName id="56C59189784FFF81FF5DFC0EB15270F4" ID-CoL="ef3a897c-b881-4fb3-b015-088b349f57b2" authority="Stec &amp; Kristensen &amp; Michalczyk, 2018" authorityName="Stec &amp; Kristensen &amp; Michalczyk" authorityYear="2018" box="[151,366,959,984]" class="Eutardigrada" family="Macrobiotidae" genus="Macrobiotus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Parachela" pageId="4" pageNumber="277" phylum="Tardigrada" rank="species" species="papei" status="sp. nov.">
<emphasis id="A3B13618784FFF81FF5DFC0EB15270F4" bold="true" box="[151,366,959,984]" italics="true" pageId="4" pageNumber="277">Macrobiotus papei</emphasis>
</taxonomicName>
<taxonomicNameLabel id="B8828B63784FFF81FEBFFC0EB1F370F4" box="[373,463,959,984]" pageId="4" pageNumber="277" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="917AEA0A784FFF81FF5DFC55B18A70D0" blockId="4.[151,463,959,1021]" box="[151,438,996,1021]" pageId="4" pageNumber="277">
(
<tableCitation id="DC47DFB1784FFF81FF54FC55B12070D0" box="[158,284,996,1021]" captionStart-0="TABLE 3" captionStart-1="TABLE 4" captionStartId-0="13.[151,239,152,175]" captionStartId-1="14.[151,239,760,783]" captionText-0="TABLE 3. Measurements [in µm] and the pt values [in %] of selected morphological structures of individuals of Macrobiotus papei sp. nov. mounted in Hoyer’s medium (N–number of specimens/structures measured, RANGE refers to the smallest and the largest structure among all measured specimens; SD–standard deviation)." captionText-1="TABLE 4. Measurements [in µm] of selected morphological structures of the eggs of Macrobiotus papei sp. nov. mounted in Hoyer’s medium (N–number of eggs/structures measured, RANGE refers to the smallest and the largest structure among all measured specimens; SD–standard deviation)." pageId="4" pageNumber="277">Tables 3–4</tableCitation>
,
<figureCitation id="09FEF68F784FFF81FEE2FC55B19170D0" box="[296,429,996,1021]" captionStart-0="FIGURE 1" captionStart-1="FIGURE 2" captionStart-2="FIGURE 3" captionStart-3="FIGURE 4" captionStart-4="FIGURE 5" captionStart-5="FIGURE 6" captionStart-6="FIGURE 7" captionStartId-0="6.[151,250,1900,1922]" captionStartId-1="7.[151,250,1822,1844]" captionStartId-2="8.[151,250,1797,1819]" captionStartId-3="9.[151,250,1762,1784]" captionStartId-4="10.[151,250,1717,1739]" captionStartId-5="11.[151,250,1801,1823]" captionStartId-6="12.[151,250,1924,1946]" captionTargetBox-0="[163,1427,405,1878]" captionTargetBox-1="[151,1436,203,1787]" captionTargetBox-2="[151,1436,213,1756]" captionTargetBox-3="[191,1396,185,1745]" captionTargetBox-4="[151,1436,603,1706]" captionTargetBox-5="[184,1408,263,1785]" captionTargetBox-6="[151,1436,213,1875]" captionTargetId-0="figure@6.[163,1427,405,1878]" captionTargetId-1="figure@7.[151,1436,203,1787]" captionTargetId-2="figure@8.[151,1436,213,1756]" captionTargetId-3="figure@9.[191,1396,185,1745]" captionTargetId-4="figure@10.[151,1436,603,1706]" captionTargetId-5="figure@11.[184,1408,263,1785]" captionTargetId-6="figure@12.[151,1436,213,1875]" captionTargetPageId-0="6" captionTargetPageId-1="7" captionTargetPageId-2="8" captionTargetPageId-3="9" captionTargetPageId-4="10" captionTargetPageId-5="11" captionTargetPageId-6="12" captionText-0="FIGURE 1. Macrobiotus papei sp. nov.—habitus: A—dorso-ventral projection (holotype, Hoyer’s medium, PCM); B–C— cuticular pores seen in PCM on the dorsal (B) and ventral (C) part of the body (paratype); D–E—cuticular pores seen in SEM on the dorsal (D) and ventral (E) part of the body (paratype). Scale bars in µm." captionText-1="FIGURE 2. Macrobiotus papei sp. nov.—cuticular structures on legs: A–B—external granulation on leg II seen in PCM (A) and SEM (B) (paratypes); C–D—a cuticular bulge resembling a pulvinus, faint cuticular fold on the internal surface of leg II with visible muscle attachment seen in PCM (C) and SEM (D) (paratypes); E—two patches of granulation on leg IV (PCM, paratype); F—distal patch of granulation on leg IV (SEM, paratype). Filled, flat arrowheads indicate the cuticular bulge, empty, flat arrowheads indicate the faint cuticular fold under the claws, arrows indicate muscle attachments, whereas filled and empty indented arrowheads indicate proximal and distal patch of granulation on the leg IV respectively. Scale bars in µm." captionText-2="FIGURE 3. Macrobiotus papei sp. nov.—buccal apparatus and the oral cavity armature (OCA) seen in PCM (all paratypes): A—dorso-ventral projection of the entire buccal apparatus; B–C oral cavity armature detected in the majority of specimens, i.e. OCA of the patagonicus type (i.e. the second and third band of teeth visible), dorsal (B) and ventral (C) view, respectively; D–E oral cavity armature detected in small specimens, i.e. OCA of the maculatus type (i.e. only the third band of teeth visible), dorsal (D) and ventral (E) view, respectively; F–G—placoid morphology. The arrow indicates pores surrounding mouth opening, flat empty arrowheads indicate the second band of teeth in the oral cavity, filled indented arrowheads indicate the third band of teeth in the oral cavity whereas filled arrowheads indicate faint constrictions in macroplacoids. Scale bars in µm." captionText-3="FIGURE 4. Macrobiotus papei sp. nov.—mouth opening and the oral cavity armature seen in SEM (paratype): A– mouth opening with six peribuccal sensory lobes and ten lamellae surrounded by large pores; B–C—the oral cavity armature of a single paratype seen in SEM from different angles. Arrows indicate pores in the irregular ring of pores surrounding the peribuccal lobes, filled arrowheads indicate teeth of the first band, empty arrowheads indicate teeth of the second band, the peaks in the dorsal ridge of the third band that correspond with teeth of the third band, in species with better developed oral cavity armatures are marked with “L d ” and “M d ” (the dorso-median tooth comprises several small peaks visible as thickenings in PCM), whereas the teeth of the ventral portion are marked with “M v ” (medio-ventral tooth) and “L v ” (lateral teeth). Scale bars in µm." captionText-4="FIGURE 5. Macrobiotus papei sp. nov.—claws (paratypes): A–B—claws II and IV seen in PCM, with smooth and slightly dentate lunules, respectively; C–D—claws I and IV seen in SEM, with smooth and slightly dentate lunules, respectively. Figs A and B assembled from several photos. Scale bars in µm." captionText-5="FIGURE 6. Macrobiotus papei sp. nov.—egg seen in PCM: A—midsection; B—surface; C–D—magnification on surface; E—magnification on midsection; F—surface of terminal discs of egg processes. Filled arrowheads indicate thin flexible filaments at the edges of terminal discs on the midsections. A, B and D show the details of a single egg whereas other photos are taken from more than one egg. Scale bars in µm." captionText-6="FIGURE 7. Macrobiotus papei sp. nov.—egg chorion morphology seen in SEM: A—entire egg with clearly visible flexible filaments on the egg processes; B–C—processes with filaments of variable length and the surface between processes; D— detail of the egg surface; E–F—details of the terminal disc and its flexible filaments. Scale bars in µm." httpUri-0="https://zenodo.org/record/1444175/files/figure.png" httpUri-1="https://zenodo.org/record/1444177/files/figure.png" httpUri-2="https://zenodo.org/record/1444179/files/figure.png" httpUri-3="https://zenodo.org/record/1444181/files/figure.png" httpUri-4="https://zenodo.org/record/1444183/files/figure.png" httpUri-5="https://zenodo.org/record/1444185/files/figure.png" httpUri-6="https://zenodo.org/record/1444187/files/figure.png" pageId="4" pageNumber="277">Figures 1–7</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="D9DFB981784FFF81FF5DFB9DB4A777A1" pageId="4" pageNumber="277" type="materials_examined">
<paragraph id="917AEA0A784FFF81FF5DFB9DB4A777A1" blockId="4.[151,1437,1068,2029]" pageId="4" pageNumber="277">
<emphasis id="A3B13618784FFF81FF5DFB9DB1BE7769" bold="true" box="[151,386,1068,1093]" pageId="4" pageNumber="277">Material examined:</emphasis>
131 animals (including
<specimenCount id="87C32183784FFF81FD5EFB9DB3107769" box="[660,812,1068,1093]" pageId="4" pageNumber="277" type="generic">10 specimens</specimenCount>
in simplex stage), and
<specimenCount id="87C32183784FFF81FBFBFB9DB4B07769" box="[1073,1164,1068,1093]" pageId="4" pageNumber="277" type="egg">81 eggs</specimenCount>
. Specimens mounted on microscope slides in Hoyer’s medium (49 animals +
<specimenCount id="87C32183784FFF81FCCAFBFEB3617744" box="[768,861,1103,1128]" pageId="4" pageNumber="277" type="egg">66 eggs</specimenCount>
), fixed on SEM stubs (20+15), and processed for
<collectionCode id="F7D472CF784FFF81FF5DFBC5B0E877A0" box="[151,212,1140,1164]" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15536" name="Department of Natural Resources, Environment, The Arts and Sport" pageId="4" pageNumber="277">DNA</collectionCode>
sequencing (6+0), aceto-orcein staining (50+0), and single individual culturing (6+0).
</paragraph>
</subSubSection>
<subSubSection id="D9DFB981784FFF8BFF0DFB26B21B7234" lastPageId="14" lastPageNumber="287" pageId="4" pageNumber="277" type="description">
<paragraph id="917AEA0A784FFF81FF0DFB26B2B575A8" blockId="4.[151,1437,1068,2029]" pageId="4" pageNumber="277">
<emphasis id="A3B13618784FFF81FF0DFB26B20A779C" bold="true" box="[199,566,1175,1200]" pageId="4" pageNumber="277">Description of the new species</emphasis>
. Animals (measurements and statistics in
<tableCitation id="DC47DFB1784FFF81FBD6FB26B44F779C" box="[1052,1139,1175,1200]" captionStart="TABLE 3" captionStartId="13.[151,239,152,175]" captionText="TABLE 3. Measurements [in µm] and the pt values [in %] of selected morphological structures of individuals of Macrobiotus papei sp. nov. mounted in Hoyer’s medium (N–number of specimens/structures measured, RANGE refers to the smallest and the largest structure among all measured specimens; SD–standard deviation)." pageId="4" pageNumber="277">Table 3</tableCitation>
). Body white in juveniles and slightly yellowish in adults, after fixation in Hoyer’s medium transparent (
<figureCitation id="09FEF68F784FFF81FBEBFB0DB44177F8" box="[1057,1149,1212,1237]" captionStart="FIGURE 1" captionStartId="6.[151,250,1900,1922]" captionTargetBox="[163,1427,405,1878]" captionTargetId="figure@6.[163,1427,405,1878]" captionTargetPageId="6" captionText="FIGURE 1. Macrobiotus papei sp. nov.—habitus: A—dorso-ventral projection (holotype, Hoyer’s medium, PCM); B–C— cuticular pores seen in PCM on the dorsal (B) and ventral (C) part of the body (paratype); D–E—cuticular pores seen in SEM on the dorsal (D) and ventral (E) part of the body (paratype). Scale bars in µm." httpUri="https://zenodo.org/record/1444175/files/figure.png" pageId="4" pageNumber="277">Fig. 1A</figureCitation>
). Eyes present in all live animals (dissolved in 80% of specimens mounted in Hoyer’s medium). Small round and oval pores (0.6–1.1 µm in diameter), visible under PCM and SEM, scattered randomly over the entire body cuticle (
<figureCitation id="09FEF68F784FFF81FBB4FAB5B4C97630" box="[1150,1269,1284,1309]" captionStart="FIGURE 1" captionStartId="6.[151,250,1900,1922]" captionTargetBox="[163,1427,405,1878]" captionTargetId="figure@6.[163,1427,405,1878]" captionTargetPageId="6" captionText="FIGURE 1. Macrobiotus papei sp. nov.—habitus: A—dorso-ventral projection (holotype, Hoyer’s medium, PCM); B–C— cuticular pores seen in PCM on the dorsal (B) and ventral (C) part of the body (paratype); D–E—cuticular pores seen in SEM on the dorsal (D) and ventral (E) part of the body (paratype). Scale bars in µm." httpUri="https://zenodo.org/record/1444175/files/figure.png" pageId="4" pageNumber="277">Fig. 1B–E</figureCitation>
), including the external and internal surface of all legs (
<figureCitation id="09FEF68F784FFF81FD93FA96B2EE766C" box="[601,722,1319,1344]" captionStart="FIGURE 2" captionStartId="7.[151,250,1822,1844]" captionTargetBox="[151,1436,203,1787]" captionTargetId="figure@7.[151,1436,203,1787]" captionTargetPageId="7" captionText="FIGURE 2. Macrobiotus papei sp. nov.—cuticular structures on legs: A–B—external granulation on leg II seen in PCM (A) and SEM (B) (paratypes); C–D—a cuticular bulge resembling a pulvinus, faint cuticular fold on the internal surface of leg II with visible muscle attachment seen in PCM (C) and SEM (D) (paratypes); E—two patches of granulation on leg IV (PCM, paratype); F—distal patch of granulation on leg IV (SEM, paratype). Filled, flat arrowheads indicate the cuticular bulge, empty, flat arrowheads indicate the faint cuticular fold under the claws, arrows indicate muscle attachments, whereas filled and empty indented arrowheads indicate proximal and distal patch of granulation on the leg IV respectively. Scale bars in µm." httpUri="https://zenodo.org/record/1444177/files/figure.png" pageId="4" pageNumber="277">Fig. 2B–E</figureCitation>
). Two cuticular granulation patches, one on the external and the other on the internal surface, are present on legs I–III (
<figureCitation id="09FEF68F784FFF81FD3FFAFDB34E7648" box="[757,882,1356,1381]" captionStart="FIGURE 2" captionStartId="7.[151,250,1822,1844]" captionTargetBox="[151,1436,203,1787]" captionTargetId="figure@7.[151,1436,203,1787]" captionTargetPageId="7" captionText="FIGURE 2. Macrobiotus papei sp. nov.—cuticular structures on legs: A–B—external granulation on leg II seen in PCM (A) and SEM (B) (paratypes); C–D—a cuticular bulge resembling a pulvinus, faint cuticular fold on the internal surface of leg II with visible muscle attachment seen in PCM (C) and SEM (D) (paratypes); E—two patches of granulation on leg IV (PCM, paratype); F—distal patch of granulation on leg IV (SEM, paratype). Filled, flat arrowheads indicate the cuticular bulge, empty, flat arrowheads indicate the faint cuticular fold under the claws, arrows indicate muscle attachments, whereas filled and empty indented arrowheads indicate proximal and distal patch of granulation on the leg IV respectively. Scale bars in µm." httpUri="https://zenodo.org/record/1444177/files/figure.png" pageId="4" pageNumber="277">Fig. 2A–D</figureCitation>
). Granulation on the external surface (
<figureCitation id="09FEF68F784FFF81FAD6FAFDB5A87648" box="[1308,1428,1356,1381]" captionStart="FIGURE 2" captionStartId="7.[151,250,1822,1844]" captionTargetBox="[151,1436,203,1787]" captionTargetId="figure@7.[151,1436,203,1787]" captionTargetPageId="7" captionText="FIGURE 2. Macrobiotus papei sp. nov.—cuticular structures on legs: A–B—external granulation on leg II seen in PCM (A) and SEM (B) (paratypes); C–D—a cuticular bulge resembling a pulvinus, faint cuticular fold on the internal surface of leg II with visible muscle attachment seen in PCM (C) and SEM (D) (paratypes); E—two patches of granulation on leg IV (PCM, paratype); F—distal patch of granulation on leg IV (SEM, paratype). Filled, flat arrowheads indicate the cuticular bulge, empty, flat arrowheads indicate the faint cuticular fold under the claws, arrows indicate muscle attachments, whereas filled and empty indented arrowheads indicate proximal and distal patch of granulation on the leg IV respectively. Scale bars in µm." httpUri="https://zenodo.org/record/1444177/files/figure.png" pageId="4" pageNumber="277">Fig. 2A–B</figureCitation>
) is bigger and more distinct than the on the internal surface (
<figureCitation id="09FEF68F784FFF81FCAAFADEB3DC76A4" box="[864,992,1391,1416]" captionStart="FIGURE 2" captionStartId="7.[151,250,1822,1844]" captionTargetBox="[151,1436,203,1787]" captionTargetId="figure@7.[151,1436,203,1787]" captionTargetPageId="7" captionText="FIGURE 2. Macrobiotus papei sp. nov.—cuticular structures on legs: A–B—external granulation on leg II seen in PCM (A) and SEM (B) (paratypes); C–D—a cuticular bulge resembling a pulvinus, faint cuticular fold on the internal surface of leg II with visible muscle attachment seen in PCM (C) and SEM (D) (paratypes); E—two patches of granulation on leg IV (PCM, paratype); F—distal patch of granulation on leg IV (SEM, paratype). Filled, flat arrowheads indicate the cuticular bulge, empty, flat arrowheads indicate the faint cuticular fold under the claws, arrows indicate muscle attachments, whereas filled and empty indented arrowheads indicate proximal and distal patch of granulation on the leg IV respectively. Scale bars in µm." httpUri="https://zenodo.org/record/1444177/files/figure.png" pageId="4" pageNumber="277">Fig. 2C–D</figureCitation>
). A cuticular bulge/fold, resembling a pulvinus, is present on the internal surface of legs I–III (
<figureCitation id="09FEF68F784FFF81FCC3FA25B3407680" box="[777,892,1428,1453]" captionStart="FIGURE 2" captionStartId="7.[151,250,1822,1844]" captionTargetBox="[151,1436,203,1787]" captionTargetId="figure@7.[151,1436,203,1787]" captionTargetPageId="7" captionText="FIGURE 2. Macrobiotus papei sp. nov.—cuticular structures on legs: A–B—external granulation on leg II seen in PCM (A) and SEM (B) (paratypes); C–D—a cuticular bulge resembling a pulvinus, faint cuticular fold on the internal surface of leg II with visible muscle attachment seen in PCM (C) and SEM (D) (paratypes); E—two patches of granulation on leg IV (PCM, paratype); F—distal patch of granulation on leg IV (SEM, paratype). Filled, flat arrowheads indicate the cuticular bulge, empty, flat arrowheads indicate the faint cuticular fold under the claws, arrows indicate muscle attachments, whereas filled and empty indented arrowheads indicate proximal and distal patch of granulation on the leg IV respectively. Scale bars in µm." httpUri="https://zenodo.org/record/1444177/files/figure.png" pageId="4" pageNumber="277">Fig 2C–D</figureCitation>
, filled arrowhead), whereas a faint cuticular fold is present just above the claws (
<figureCitation id="09FEF68F784FFF81FDC8FA06B24176FC" box="[514,637,1463,1488]" captionStart="FIGURE 2" captionStartId="7.[151,250,1822,1844]" captionTargetBox="[151,1436,203,1787]" captionTargetId="figure@7.[151,1436,203,1787]" captionTargetPageId="7" captionText="FIGURE 2. Macrobiotus papei sp. nov.—cuticular structures on legs: A–B—external granulation on leg II seen in PCM (A) and SEM (B) (paratypes); C–D—a cuticular bulge resembling a pulvinus, faint cuticular fold on the internal surface of leg II with visible muscle attachment seen in PCM (C) and SEM (D) (paratypes); E—two patches of granulation on leg IV (PCM, paratype); F—distal patch of granulation on leg IV (SEM, paratype). Filled, flat arrowheads indicate the cuticular bulge, empty, flat arrowheads indicate the faint cuticular fold under the claws, arrows indicate muscle attachments, whereas filled and empty indented arrowheads indicate proximal and distal patch of granulation on the leg IV respectively. Scale bars in µm." httpUri="https://zenodo.org/record/1444177/files/figure.png" pageId="4" pageNumber="277">Fig. 2C–D</figureCitation>
, flat empty arrowhead). Both structures are visible only if the legs are fully extended and well oriented on the slide (particularly in the case of the cuticular fold above the claws). Cuticular granulation on legs IV is always clearly visible and consists of two granulation patches: the distal patch with densely distributed granules situated just above the claws and the proximal patch being wider with more sparsely distributed granules located just above the first patch (
<figureCitation id="09FEF68F784FFF81FC92F9F6B3F1754C" box="[856,973,1607,1632]" captionStart="FIGURE 2" captionStartId="7.[151,250,1822,1844]" captionTargetBox="[151,1436,203,1787]" captionTargetId="figure@7.[151,1436,203,1787]" captionTargetPageId="7" captionText="FIGURE 2. Macrobiotus papei sp. nov.—cuticular structures on legs: A–B—external granulation on leg II seen in PCM (A) and SEM (B) (paratypes); C–D—a cuticular bulge resembling a pulvinus, faint cuticular fold on the internal surface of leg II with visible muscle attachment seen in PCM (C) and SEM (D) (paratypes); E—two patches of granulation on leg IV (PCM, paratype); F—distal patch of granulation on leg IV (SEM, paratype). Filled, flat arrowheads indicate the cuticular bulge, empty, flat arrowheads indicate the faint cuticular fold under the claws, arrows indicate muscle attachments, whereas filled and empty indented arrowheads indicate proximal and distal patch of granulation on the leg IV respectively. Scale bars in µm." httpUri="https://zenodo.org/record/1444177/files/figure.png" pageId="4" pageNumber="277">Fig. 2E–F</figureCitation>
). Sometimes muscle attachments are also visible under claws I–III (
<figureCitation id="09FEF68F784FFF81FE73F9DDB20E75A8" box="[441,562,1644,1669]" captionStart="FIGURE 2" captionStartId="7.[151,250,1822,1844]" captionTargetBox="[151,1436,203,1787]" captionTargetId="figure@7.[151,1436,203,1787]" captionTargetPageId="7" captionText="FIGURE 2. Macrobiotus papei sp. nov.—cuticular structures on legs: A–B—external granulation on leg II seen in PCM (A) and SEM (B) (paratypes); C–D—a cuticular bulge resembling a pulvinus, faint cuticular fold on the internal surface of leg II with visible muscle attachment seen in PCM (C) and SEM (D) (paratypes); E—two patches of granulation on leg IV (PCM, paratype); F—distal patch of granulation on leg IV (SEM, paratype). Filled, flat arrowheads indicate the cuticular bulge, empty, flat arrowheads indicate the faint cuticular fold under the claws, arrows indicate muscle attachments, whereas filled and empty indented arrowheads indicate proximal and distal patch of granulation on the leg IV respectively. Scale bars in µm." httpUri="https://zenodo.org/record/1444177/files/figure.png" pageId="4" pageNumber="277">Fig. 2C–D</figureCitation>
, arrow)
</paragraph>
<paragraph id="917AEA0A784FFF80FF0DF93EB18C7184" blockId="4.[151,1437,1068,2029]" lastBlockId="5.[151,1437,151,1725]" lastPageId="5" lastPageNumber="278" pageId="4" pageNumber="277">
Mouth antero-ventral. Bucco-pharyngeal apparatus of the
<taxonomicName id="56C59189784FFF81FC94F93EB3D07584" authorityName="Stec &amp; Kristensen &amp; Michalczyk" authorityYear="2018" box="[862,1004,1679,1704]" class="Eutardigrada" family="Macrobiotidae" genus="Macrobiotus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Parachela" pageId="4" pageNumber="277" phylum="Tardigrada" rank="genus">
<emphasis id="A3B13618784FFF81FC94F93EB3D07584" box="[862,1004,1679,1704]" italics="true" pageId="4" pageNumber="277">Macrobiotus</emphasis>
</taxonomicName>
<typeStatus id="4E7E54A8784FFF81FC3FF920B4147585" box="[1013,1064,1681,1705]" pageId="4" pageNumber="277">type</typeStatus>
, with the ventral lamina and ten small peribuccal lamellae followed by six buccal sensory lobes (
<figureCitation id="09FEF68F784FFF81FCB0F905B3EB75E0" box="[890,983,1716,1741]" captionStart="FIGURE 3" captionStartId="8.[151,250,1797,1819]" captionTargetBox="[151,1436,213,1756]" captionTargetId="figure@8.[151,1436,213,1756]" captionTargetPageId="8" captionText="FIGURE 3. Macrobiotus papei sp. nov.—buccal apparatus and the oral cavity armature (OCA) seen in PCM (all paratypes): A—dorso-ventral projection of the entire buccal apparatus; B–C oral cavity armature detected in the majority of specimens, i.e. OCA of the patagonicus type (i.e. the second and third band of teeth visible), dorsal (B) and ventral (C) view, respectively; D–E oral cavity armature detected in small specimens, i.e. OCA of the maculatus type (i.e. only the third band of teeth visible), dorsal (D) and ventral (E) view, respectively; F–G—placoid morphology. The arrow indicates pores surrounding mouth opening, flat empty arrowheads indicate the second band of teeth in the oral cavity, filled indented arrowheads indicate the third band of teeth in the oral cavity whereas filled arrowheads indicate faint constrictions in macroplacoids. Scale bars in µm." httpUri="https://zenodo.org/record/1444179/files/figure.png" pageId="4" pageNumber="277">Figs 3A</figureCitation>
and
<figureCitation id="09FEF68F784FFF81FBDBF905B46575E0" box="[1041,1113,1716,1740]" captionStart="FIGURE 4" captionStartId="9.[151,250,1762,1784]" captionTargetBox="[191,1396,185,1745]" captionTargetId="figure@9.[191,1396,185,1745]" captionTargetPageId="9" captionText="FIGURE 4. Macrobiotus papei sp. nov.—mouth opening and the oral cavity armature seen in SEM (paratype): A– mouth opening with six peribuccal sensory lobes and ten lamellae surrounded by large pores; B–C—the oral cavity armature of a single paratype seen in SEM from different angles. Arrows indicate pores in the irregular ring of pores surrounding the peribuccal lobes, filled arrowheads indicate teeth of the first band, empty arrowheads indicate teeth of the second band, the peaks in the dorsal ridge of the third band that correspond with teeth of the third band, in species with better developed oral cavity armatures are marked with “L d ” and “M d ” (the dorso-median tooth comprises several small peaks visible as thickenings in PCM), whereas the teeth of the ventral portion are marked with “M v ” (medio-ventral tooth) and “L v ” (lateral teeth). Scale bars in µm." httpUri="https://zenodo.org/record/1444181/files/figure.png" pageId="4" pageNumber="277">4A–C</figureCitation>
). An irregular ring of pores, clearly visible using SEM and occasionally under PCM, is present around the mouth opening, immediately behind the peribuccal sensory lobes (
<figureCitation id="09FEF68F784FFF81FE2CF94DB2037438" box="[486,575,1788,1813]" captionStart="FIGURE 3" captionStartId="8.[151,250,1797,1819]" captionTargetBox="[151,1436,213,1756]" captionTargetId="figure@8.[151,1436,213,1756]" captionTargetPageId="8" captionText="FIGURE 3. Macrobiotus papei sp. nov.—buccal apparatus and the oral cavity armature (OCA) seen in PCM (all paratypes): A—dorso-ventral projection of the entire buccal apparatus; B–C oral cavity armature detected in the majority of specimens, i.e. OCA of the patagonicus type (i.e. the second and third band of teeth visible), dorsal (B) and ventral (C) view, respectively; D–E oral cavity armature detected in small specimens, i.e. OCA of the maculatus type (i.e. only the third band of teeth visible), dorsal (D) and ventral (E) view, respectively; F–G—placoid morphology. The arrow indicates pores surrounding mouth opening, flat empty arrowheads indicate the second band of teeth in the oral cavity, filled indented arrowheads indicate the third band of teeth in the oral cavity whereas filled arrowheads indicate faint constrictions in macroplacoids. Scale bars in µm." httpUri="https://zenodo.org/record/1444179/files/figure.png" pageId="4" pageNumber="277">Figs 3B</figureCitation>
and
<figureCitation id="09FEF68F784FFF81FDB2F94DB2A17438" box="[632,669,1788,1812]" captionStart="FIGURE 4" captionStartId="9.[151,250,1762,1784]" captionTargetBox="[191,1396,185,1745]" captionTargetId="figure@9.[191,1396,185,1745]" captionTargetPageId="9" captionText="FIGURE 4. Macrobiotus papei sp. nov.—mouth opening and the oral cavity armature seen in SEM (paratype): A– mouth opening with six peribuccal sensory lobes and ten lamellae surrounded by large pores; B–C—the oral cavity armature of a single paratype seen in SEM from different angles. Arrows indicate pores in the irregular ring of pores surrounding the peribuccal lobes, filled arrowheads indicate teeth of the first band, empty arrowheads indicate teeth of the second band, the peaks in the dorsal ridge of the third band that correspond with teeth of the third band, in species with better developed oral cavity armatures are marked with “L d ” and “M d ” (the dorso-median tooth comprises several small peaks visible as thickenings in PCM), whereas the teeth of the ventral portion are marked with “M v ” (medio-ventral tooth) and “L v ” (lateral teeth). Scale bars in µm." httpUri="https://zenodo.org/record/1444181/files/figure.png" pageId="4" pageNumber="277">4A</figureCitation>
, arrow). Under PCM, the oral cavity armature is of the
<emphasis id="A3B13618784FFF81FAD9F94CB5A07438" box="[1299,1436,1789,1812]" italics="true" pageId="4" pageNumber="277">patagonicus</emphasis>
<typeStatus id="4E7E54A8784FFF81FF5DF890B0FB7415" box="[151,199,1825,1849]" pageId="4" pageNumber="277">type</typeStatus>
(only the second and third band of teeth visible under light microscopy (LM) in the majority of specimens (
<figureCitation id="09FEF68F784FFF81FF55F8F5B1297470" box="[159,277,1860,1885]" captionStart="FIGURE 3" captionStartId="8.[151,250,1797,1819]" captionTargetBox="[151,1436,213,1756]" captionTargetId="figure@8.[151,1436,213,1756]" captionTargetPageId="8" captionText="FIGURE 3. Macrobiotus papei sp. nov.—buccal apparatus and the oral cavity armature (OCA) seen in PCM (all paratypes): A—dorso-ventral projection of the entire buccal apparatus; B–C oral cavity armature detected in the majority of specimens, i.e. OCA of the patagonicus type (i.e. the second and third band of teeth visible), dorsal (B) and ventral (C) view, respectively; D–E oral cavity armature detected in small specimens, i.e. OCA of the maculatus type (i.e. only the third band of teeth visible), dorsal (D) and ventral (E) view, respectively; F–G—placoid morphology. The arrow indicates pores surrounding mouth opening, flat empty arrowheads indicate the second band of teeth in the oral cavity, filled indented arrowheads indicate the third band of teeth in the oral cavity whereas filled arrowheads indicate faint constrictions in macroplacoids. Scale bars in µm." httpUri="https://zenodo.org/record/1444179/files/figure.png" pageId="4" pageNumber="277">Fig. 3B–C</figureCitation>
) but in smaller individuals the oral cavity armature is of the
<emphasis id="A3B13618784FFF81FC7FF8F5B4147471" box="[949,1064,1860,1885]" italics="true" pageId="4" pageNumber="277">maculatus</emphasis>
<typeStatus id="4E7E54A8784FFF81FBE5F8F4B4637471" box="[1071,1119,1861,1885]" pageId="4" pageNumber="277">type</typeStatus>
(only the third band of teeth visible under LM) (
<figureCitation id="09FEF68F784FFF81FEB0F8D6B1CB74AC" box="[378,503,1895,1920]" captionStart="FIGURE 3" captionStartId="8.[151,250,1797,1819]" captionTargetBox="[151,1436,213,1756]" captionTargetId="figure@8.[151,1436,213,1756]" captionTargetPageId="8" captionText="FIGURE 3. Macrobiotus papei sp. nov.—buccal apparatus and the oral cavity armature (OCA) seen in PCM (all paratypes): A—dorso-ventral projection of the entire buccal apparatus; B–C oral cavity armature detected in the majority of specimens, i.e. OCA of the patagonicus type (i.e. the second and third band of teeth visible), dorsal (B) and ventral (C) view, respectively; D–E oral cavity armature detected in small specimens, i.e. OCA of the maculatus type (i.e. only the third band of teeth visible), dorsal (D) and ventral (E) view, respectively; F–G—placoid morphology. The arrow indicates pores surrounding mouth opening, flat empty arrowheads indicate the second band of teeth in the oral cavity, filled indented arrowheads indicate the third band of teeth in the oral cavity whereas filled arrowheads indicate faint constrictions in macroplacoids. Scale bars in µm." httpUri="https://zenodo.org/record/1444179/files/figure.png" pageId="4" pageNumber="277">Fig. 3D–E</figureCitation>
). However, under SEM the oral cavity comprises three bands of teeth,
<emphasis id="A3B13618784FFF81FAFAF8D8B56F74AC" box="[1328,1363,1897,1920]" italics="true" pageId="4" pageNumber="277">i.e.</emphasis>
under PCM only the third band or only the second and the third band of teeth are visible (
<figureCitation id="09FEF68F784FFF81FB84F83DB4FB7488" box="[1102,1223,1932,1957]" captionStart="FIGURE 3" captionStartId="8.[151,250,1797,1819]" captionTargetBox="[151,1436,213,1756]" captionTargetId="figure@8.[151,1436,213,1756]" captionTargetPageId="8" captionText="FIGURE 3. Macrobiotus papei sp. nov.—buccal apparatus and the oral cavity armature (OCA) seen in PCM (all paratypes): A—dorso-ventral projection of the entire buccal apparatus; B–C oral cavity armature detected in the majority of specimens, i.e. OCA of the patagonicus type (i.e. the second and third band of teeth visible), dorsal (B) and ventral (C) view, respectively; D–E oral cavity armature detected in small specimens, i.e. OCA of the maculatus type (i.e. only the third band of teeth visible), dorsal (D) and ventral (E) view, respectively; F–G—placoid morphology. The arrow indicates pores surrounding mouth opening, flat empty arrowheads indicate the second band of teeth in the oral cavity, filled indented arrowheads indicate the third band of teeth in the oral cavity whereas filled arrowheads indicate faint constrictions in macroplacoids. Scale bars in µm." httpUri="https://zenodo.org/record/1444179/files/figure.png" pageId="4" pageNumber="277">Fig. 3B–E</figureCitation>
), whereas all three bands are always detectable in SEM (
<figureCitation id="09FEF68F784FFF81FD81F801B2F474E4" box="[587,712,1968,1993]" captionStart="FIGURE 4" captionStartId="9.[151,250,1762,1784]" captionTargetBox="[191,1396,185,1745]" captionTargetId="figure@9.[191,1396,185,1745]" captionTargetPageId="9" captionText="FIGURE 4. Macrobiotus papei sp. nov.—mouth opening and the oral cavity armature seen in SEM (paratype): A– mouth opening with six peribuccal sensory lobes and ten lamellae surrounded by large pores; B–C—the oral cavity armature of a single paratype seen in SEM from different angles. Arrows indicate pores in the irregular ring of pores surrounding the peribuccal lobes, filled arrowheads indicate teeth of the first band, empty arrowheads indicate teeth of the second band, the peaks in the dorsal ridge of the third band that correspond with teeth of the third band, in species with better developed oral cavity armatures are marked with “L d ” and “M d ” (the dorso-median tooth comprises several small peaks visible as thickenings in PCM), whereas the teeth of the ventral portion are marked with “M v ” (medio-ventral tooth) and “L v ” (lateral teeth). Scale bars in µm." httpUri="https://zenodo.org/record/1444181/files/figure.png" pageId="4" pageNumber="277">Fig. 4B–C</figureCitation>
). The first band of teeth consists of numerous extremely small cones arranged in one to two rows situated anteriorly in the oral cavity, just behind the base of the peribuccal lamellae (
<figureCitation id="09FEF68F784EFF80FECDFF26B1BE739C" box="[263,386,151,176]" captionStart="FIGURE 4" captionStartId="9.[151,250,1762,1784]" captionTargetBox="[191,1396,185,1745]" captionTargetId="figure@9.[191,1396,185,1745]" captionTargetPageId="9" captionText="FIGURE 4. Macrobiotus papei sp. nov.—mouth opening and the oral cavity armature seen in SEM (paratype): A– mouth opening with six peribuccal sensory lobes and ten lamellae surrounded by large pores; B–C—the oral cavity armature of a single paratype seen in SEM from different angles. Arrows indicate pores in the irregular ring of pores surrounding the peribuccal lobes, filled arrowheads indicate teeth of the first band, empty arrowheads indicate teeth of the second band, the peaks in the dorsal ridge of the third band that correspond with teeth of the third band, in species with better developed oral cavity armatures are marked with “L d ” and “M d ” (the dorso-median tooth comprises several small peaks visible as thickenings in PCM), whereas the teeth of the ventral portion are marked with “M v ” (medio-ventral tooth) and “L v ” (lateral teeth). Scale bars in µm." httpUri="https://zenodo.org/record/1444181/files/figure.png" pageId="5" pageNumber="278">Fig. 4B–C</figureCitation>
, flat filled arrowhead). The second band of teeth are situated between the ring fold and the third band of teeth and comprises 4–5 rows of small cones, slightly bigger than those of the first band (
<figureCitation id="09FEF68F784EFF80FAD5FF0DB5AB73F8" box="[1311,1431,188,213]" captionStart="FIGURE 4" captionStartId="9.[151,250,1762,1784]" captionTargetBox="[191,1396,185,1745]" captionTargetId="figure@9.[191,1396,185,1745]" captionTargetPageId="9" captionText="FIGURE 4. Macrobiotus papei sp. nov.—mouth opening and the oral cavity armature seen in SEM (paratype): A– mouth opening with six peribuccal sensory lobes and ten lamellae surrounded by large pores; B–C—the oral cavity armature of a single paratype seen in SEM from different angles. Arrows indicate pores in the irregular ring of pores surrounding the peribuccal lobes, filled arrowheads indicate teeth of the first band, empty arrowheads indicate teeth of the second band, the peaks in the dorsal ridge of the third band that correspond with teeth of the third band, in species with better developed oral cavity armatures are marked with “L d ” and “M d ” (the dorso-median tooth comprises several small peaks visible as thickenings in PCM), whereas the teeth of the ventral portion are marked with “M v ” (medio-ventral tooth) and “L v ” (lateral teeth). Scale bars in µm." httpUri="https://zenodo.org/record/1444181/files/figure.png" pageId="5" pageNumber="278">Fig. 4B–C</figureCitation>
, empty flat arrowhead). Under PCM, only the larger teeth of second band are visible in larger specimens and often appear as very weakly developed (
<figureCitation id="09FEF68F784EFF80FDD3FEB5B2AD7230" box="[537,657,260,285]" captionStart="FIGURE 3" captionStartId="8.[151,250,1797,1819]" captionTargetBox="[151,1436,213,1756]" captionTargetId="figure@8.[151,1436,213,1756]" captionTargetPageId="8" captionText="FIGURE 3. Macrobiotus papei sp. nov.—buccal apparatus and the oral cavity armature (OCA) seen in PCM (all paratypes): A—dorso-ventral projection of the entire buccal apparatus; B–C oral cavity armature detected in the majority of specimens, i.e. OCA of the patagonicus type (i.e. the second and third band of teeth visible), dorsal (B) and ventral (C) view, respectively; D–E oral cavity armature detected in small specimens, i.e. OCA of the maculatus type (i.e. only the third band of teeth visible), dorsal (D) and ventral (E) view, respectively; F–G—placoid morphology. The arrow indicates pores surrounding mouth opening, flat empty arrowheads indicate the second band of teeth in the oral cavity, filled indented arrowheads indicate the third band of teeth in the oral cavity whereas filled arrowheads indicate faint constrictions in macroplacoids. Scale bars in µm." httpUri="https://zenodo.org/record/1444179/files/figure.png" pageId="5" pageNumber="278">Fig. 3B–C</figureCitation>
, empty flat arrowhead). The teeth of the third band are located within the posterior portion of the oral cavity, between the second band of teeth and the buccal tube opening (
<figureCitation id="09FEF68F784EFF80FADCFE96B5AC726C" box="[1302,1424,295,320]" captionStart="FIGURE 4" captionStartId="9.[151,250,1762,1784]" captionTargetBox="[191,1396,185,1745]" captionTargetId="figure@9.[191,1396,185,1745]" captionTargetPageId="9" captionText="FIGURE 4. Macrobiotus papei sp. nov.—mouth opening and the oral cavity armature seen in SEM (paratype): A– mouth opening with six peribuccal sensory lobes and ten lamellae surrounded by large pores; B–C—the oral cavity armature of a single paratype seen in SEM from different angles. Arrows indicate pores in the irregular ring of pores surrounding the peribuccal lobes, filled arrowheads indicate teeth of the first band, empty arrowheads indicate teeth of the second band, the peaks in the dorsal ridge of the third band that correspond with teeth of the third band, in species with better developed oral cavity armatures are marked with “L d ” and “M d ” (the dorso-median tooth comprises several small peaks visible as thickenings in PCM), whereas the teeth of the ventral portion are marked with “M v ” (medio-ventral tooth) and “L v ” (lateral teeth). Scale bars in µm." httpUri="https://zenodo.org/record/1444181/files/figure.png" pageId="5" pageNumber="278">Fig. 4B–C</figureCitation>
). The third band of teeth is discontinuous and divided into a dorsal and a ventral portion. Under PCM, the dorsal teeth form a single transverse ridge with obvious thickenings, whereas the ventral teeth appear as two separate lateral transverse ridges between which a roundish median tooth is visible (
<figureCitation id="09FEF68F784EFF80FC2FFE25B4617280" box="[997,1117,404,429]" captionStart="FIGURE 3" captionStartId="8.[151,250,1797,1819]" captionTargetBox="[151,1436,213,1756]" captionTargetId="figure@8.[151,1436,213,1756]" captionTargetPageId="8" captionText="FIGURE 3. Macrobiotus papei sp. nov.—buccal apparatus and the oral cavity armature (OCA) seen in PCM (all paratypes): A—dorso-ventral projection of the entire buccal apparatus; B–C oral cavity armature detected in the majority of specimens, i.e. OCA of the patagonicus type (i.e. the second and third band of teeth visible), dorsal (B) and ventral (C) view, respectively; D–E oral cavity armature detected in small specimens, i.e. OCA of the maculatus type (i.e. only the third band of teeth visible), dorsal (D) and ventral (E) view, respectively; F–G—placoid morphology. The arrow indicates pores surrounding mouth opening, flat empty arrowheads indicate the second band of teeth in the oral cavity, filled indented arrowheads indicate the third band of teeth in the oral cavity whereas filled arrowheads indicate faint constrictions in macroplacoids. Scale bars in µm." httpUri="https://zenodo.org/record/1444179/files/figure.png" pageId="5" pageNumber="278">Fig. 3B–E</figureCitation>
). The medio-ventral tooth is occasionally divided into two smaller rounded teeth. Under SEM, the dorsal teeth form a single ridge with two larger lateral peaks and several smaller median peaks and indentations (
<figureCitation id="09FEF68F784EFF80FC0EFE6DB42072D8" box="[964,1052,476,501]" captionStart="FIGURE 4" captionStartId="9.[151,250,1762,1784]" captionTargetBox="[191,1396,185,1745]" captionTargetId="figure@9.[191,1396,185,1745]" captionTargetPageId="9" captionText="FIGURE 4. Macrobiotus papei sp. nov.—mouth opening and the oral cavity armature seen in SEM (paratype): A– mouth opening with six peribuccal sensory lobes and ten lamellae surrounded by large pores; B–C—the oral cavity armature of a single paratype seen in SEM from different angles. Arrows indicate pores in the irregular ring of pores surrounding the peribuccal lobes, filled arrowheads indicate teeth of the first band, empty arrowheads indicate teeth of the second band, the peaks in the dorsal ridge of the third band that correspond with teeth of the third band, in species with better developed oral cavity armatures are marked with “L d ” and “M d ” (the dorso-median tooth comprises several small peaks visible as thickenings in PCM), whereas the teeth of the ventral portion are marked with “M v ” (medio-ventral tooth) and “L v ” (lateral teeth). Scale bars in µm." httpUri="https://zenodo.org/record/1444181/files/figure.png" pageId="5" pageNumber="278">Fig. 4B</figureCitation>
) which correspond to thickenings visible under LM (
<figureCitation id="09FEF68F784EFF80FEBDFE4EB1EF7134" box="[375,467,511,536]" captionStart="FIGURE 3" captionStartId="8.[151,250,1797,1819]" captionTargetBox="[151,1436,213,1756]" captionTargetId="figure@8.[151,1436,213,1756]" captionTargetPageId="8" captionText="FIGURE 3. Macrobiotus papei sp. nov.—buccal apparatus and the oral cavity armature (OCA) seen in PCM (all paratypes): A—dorso-ventral projection of the entire buccal apparatus; B–C oral cavity armature detected in the majority of specimens, i.e. OCA of the patagonicus type (i.e. the second and third band of teeth visible), dorsal (B) and ventral (C) view, respectively; D–E oral cavity armature detected in small specimens, i.e. OCA of the maculatus type (i.e. only the third band of teeth visible), dorsal (D) and ventral (E) view, respectively; F–G—placoid morphology. The arrow indicates pores surrounding mouth opening, flat empty arrowheads indicate the second band of teeth in the oral cavity, filled indented arrowheads indicate the third band of teeth in the oral cavity whereas filled arrowheads indicate faint constrictions in macroplacoids. Scale bars in µm." httpUri="https://zenodo.org/record/1444179/files/figure.png" pageId="5" pageNumber="278">Fig. 3B</figureCitation>
). The ventral teeth are separated into one median and two lateral teeth (
<figureCitation id="09FEF68F784EFF80FAF8FE4EB5AC7134" box="[1330,1424,511,536]" captionStart="FIGURE 4" captionStartId="9.[151,250,1762,1784]" captionTargetBox="[191,1396,185,1745]" captionTargetId="figure@9.[191,1396,185,1745]" captionTargetPageId="9" captionText="FIGURE 4. Macrobiotus papei sp. nov.—mouth opening and the oral cavity armature seen in SEM (paratype): A– mouth opening with six peribuccal sensory lobes and ten lamellae surrounded by large pores; B–C—the oral cavity armature of a single paratype seen in SEM from different angles. Arrows indicate pores in the irregular ring of pores surrounding the peribuccal lobes, filled arrowheads indicate teeth of the first band, empty arrowheads indicate teeth of the second band, the peaks in the dorsal ridge of the third band that correspond with teeth of the third band, in species with better developed oral cavity armatures are marked with “L d ” and “M d ” (the dorso-median tooth comprises several small peaks visible as thickenings in PCM), whereas the teeth of the ventral portion are marked with “M v ” (medio-ventral tooth) and “L v ” (lateral teeth). Scale bars in µm." httpUri="https://zenodo.org/record/1444181/files/figure.png" pageId="5" pageNumber="278">Fig. 4C</figureCitation>
). Pharyngeal bulb spherical, with triangular apophyses, two rod-shaped macroplacoids and a triangular small microplacoid (
<figureCitation id="09FEF68F784EFF80FEF0FDF6B1A8714C" box="[314,404,583,608]" captionStart="FIGURE 3" captionStartId="8.[151,250,1797,1819]" captionTargetBox="[151,1436,213,1756]" captionTargetId="figure@8.[151,1436,213,1756]" captionTargetPageId="8" captionText="FIGURE 3. Macrobiotus papei sp. nov.—buccal apparatus and the oral cavity armature (OCA) seen in PCM (all paratypes): A—dorso-ventral projection of the entire buccal apparatus; B–C oral cavity armature detected in the majority of specimens, i.e. OCA of the patagonicus type (i.e. the second and third band of teeth visible), dorsal (B) and ventral (C) view, respectively; D–E oral cavity armature detected in small specimens, i.e. OCA of the maculatus type (i.e. only the third band of teeth visible), dorsal (D) and ventral (E) view, respectively; F–G—placoid morphology. The arrow indicates pores surrounding mouth opening, flat empty arrowheads indicate the second band of teeth in the oral cavity, filled indented arrowheads indicate the third band of teeth in the oral cavity whereas filled arrowheads indicate faint constrictions in macroplacoids. Scale bars in µm." httpUri="https://zenodo.org/record/1444179/files/figure.png" pageId="5" pageNumber="278">Fig. 3A</figureCitation>
). The macroplacoid length sequence 2&lt;1. The first and the second macroplacoid are smooth in the majority of specimens, only occasionally with very weak central and subterminal constrictions, respectively (
<figureCitation id="09FEF68F784EFF80FF55FD3EB12E7184" box="[159,274,655,680]" captionStart="FIGURE 3" captionStartId="8.[151,250,1797,1819]" captionTargetBox="[151,1436,213,1756]" captionTargetId="figure@8.[151,1436,213,1756]" captionTargetPageId="8" captionText="FIGURE 3. Macrobiotus papei sp. nov.—buccal apparatus and the oral cavity armature (OCA) seen in PCM (all paratypes): A—dorso-ventral projection of the entire buccal apparatus; B–C oral cavity armature detected in the majority of specimens, i.e. OCA of the patagonicus type (i.e. the second and third band of teeth visible), dorsal (B) and ventral (C) view, respectively; D–E oral cavity armature detected in small specimens, i.e. OCA of the maculatus type (i.e. only the third band of teeth visible), dorsal (D) and ventral (E) view, respectively; F–G—placoid morphology. The arrow indicates pores surrounding mouth opening, flat empty arrowheads indicate the second band of teeth in the oral cavity, filled indented arrowheads indicate the third band of teeth in the oral cavity whereas filled arrowheads indicate faint constrictions in macroplacoids. Scale bars in µm." httpUri="https://zenodo.org/record/1444179/files/figure.png" pageId="5" pageNumber="278">Fig. 3F–G</figureCitation>
, arrowheads).
</paragraph>
<paragraph id="917AEA0A784EFF80FF0DFD05B4DD7039" blockId="5.[151,1437,151,1725]" pageId="5" pageNumber="278">
Claws long and slender, of the
<taxonomicName id="56C59189784EFF80FDEFFD05B2B371E1" authorityName=", C. A. S. Schultze" authorityYear="1834" box="[549,655,692,717]" class="Eutardigrada" family="Macrobiotidae" genus="Macrobiotus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Parachela" pageId="5" pageNumber="278" phylum="Tardigrada" rank="species" species="hufelandi">
<emphasis id="A3B13618784EFF80FDEFFD05B2B371E1" box="[549,655,692,717]" italics="true" pageId="5" pageNumber="278">hufelandi</emphasis>
</taxonomicName>
<typeStatus id="4E7E54A8784EFF80FD5CFD04B2FA71E1" box="[662,710,693,717]" pageId="5" pageNumber="278">type</typeStatus>
(
<figureCitation id="09FEF68F784EFF80FD1CFD05B36971E1" box="[726,853,692,717]" captionStart="FIGURE 5" captionStartId="10.[151,250,1717,1739]" captionTargetBox="[151,1436,603,1706]" captionTargetId="figure@10.[151,1436,603,1706]" captionTargetPageId="10" captionText="FIGURE 5. Macrobiotus papei sp. nov.—claws (paratypes): A–B—claws II and IV seen in PCM, with smooth and slightly dentate lunules, respectively; C–D—claws I and IV seen in SEM, with smooth and slightly dentate lunules, respectively. Figs A and B assembled from several photos. Scale bars in µm." httpUri="https://zenodo.org/record/1444183/files/figure.png" pageId="5" pageNumber="278">Fig. 5A–D</figureCitation>
). Primary branches with distinct accessory points, a long common tract, and with an evident stalk connecting the claw to the lunula (
<figureCitation id="09FEF68F784EFF80FBD6FD66B4A571DC" box="[1052,1177,727,752]" captionStart="FIGURE 5" captionStartId="10.[151,250,1717,1739]" captionTargetBox="[151,1436,603,1706]" captionTargetId="figure@10.[151,1436,603,1706]" captionTargetPageId="10" captionText="FIGURE 5. Macrobiotus papei sp. nov.—claws (paratypes): A–B—claws II and IV seen in PCM, with smooth and slightly dentate lunules, respectively; C–D—claws I and IV seen in SEM, with smooth and slightly dentate lunules, respectively. Figs A and B assembled from several photos. Scale bars in µm." httpUri="https://zenodo.org/record/1444183/files/figure.png" pageId="5" pageNumber="278">Fig. 5A–D</figureCitation>
). Lunulae I–III smooth (
<figureCitation id="09FEF68F784EFF80FF55FD4DB1257038" box="[159,281,764,789]" captionStart="FIGURE 5" captionStartId="10.[151,250,1717,1739]" captionTargetBox="[151,1436,603,1706]" captionTargetId="figure@10.[151,1436,603,1706]" captionTargetPageId="10" captionText="FIGURE 5. Macrobiotus papei sp. nov.—claws (paratypes): A–B—claws II and IV seen in PCM, with smooth and slightly dentate lunules, respectively; C–D—claws I and IV seen in SEM, with smooth and slightly dentate lunules, respectively. Figs A and B assembled from several photos. Scale bars in µm." httpUri="https://zenodo.org/record/1444183/files/figure.png" pageId="5" pageNumber="278">Fig. 5A, C</figureCitation>
), whereas lunulae IV sparsely dentate (
<figureCitation id="09FEF68F784EFF80FD1BFD4DB3707038" box="[721,844,764,789]" captionStart="FIGURE 5" captionStartId="10.[151,250,1717,1739]" captionTargetBox="[151,1436,603,1706]" captionTargetId="figure@10.[151,1436,603,1706]" captionTargetPageId="10" captionText="FIGURE 5. Macrobiotus papei sp. nov.—claws (paratypes): A–B—claws II and IV seen in PCM, with smooth and slightly dentate lunules, respectively; C–D—claws I and IV seen in SEM, with smooth and slightly dentate lunules, respectively. Figs A and B assembled from several photos. Scale bars in µm." httpUri="https://zenodo.org/record/1444183/files/figure.png" pageId="5" pageNumber="278">Fig. 5B, D</figureCitation>
). Cuticular bars under claws absent.
</paragraph>
<paragraph id="917AEA0A784EFF80FF0DFC91B4897621" blockId="5.[151,1437,151,1725]" pageId="5" pageNumber="278">
Eggs (measurements and statistics in
<tableCitation id="DC47DFB1784EFF80FDBAFCAEB2FB7014" box="[624,711,799,824]" captionStart="TABLE 4" captionStartId="14.[151,239,760,783]" captionText="TABLE 4. Measurements [in µm] of selected morphological structures of the eggs of Macrobiotus papei sp. nov. mounted in Hoyer’s medium (N–number of eggs/structures measured, RANGE refers to the smallest and the largest structure among all measured specimens; SD–standard deviation)." pageId="5" pageNumber="278">Table 4</tableCitation>
). Laid freely, white/light yellow, spherical or slightly oval (
<figureCitation id="09FEF68F784EFF80FAA0FCAEB0E57071" captionStart="FIGURE 6" captionStartId="11.[151,250,1801,1823]" captionTargetBox="[184,1408,263,1785]" captionTargetId="figure@11.[184,1408,263,1785]" captionTargetPageId="11" captionText="FIGURE 6. Macrobiotus papei sp. nov.—egg seen in PCM: A—midsection; B—surface; C–D—magnification on surface; E—magnification on midsection; F—surface of terminal discs of egg processes. Filled arrowheads indicate thin flexible filaments at the edges of terminal discs on the midsections. A, B and D show the details of a single egg whereas other photos are taken from more than one egg. Scale bars in µm." httpUri="https://zenodo.org/record/1444185/files/figure.png" pageId="5" pageNumber="278">Figs 6A–B</figureCitation>
and
<figureCitation id="09FEF68F784EFF80FEDEFCF5B1057070" box="[276,313,836,860]" captionStart="FIGURE 7" captionStartId="12.[151,250,1924,1946]" captionTargetBox="[151,1436,213,1875]" captionTargetId="figure@12.[151,1436,213,1875]" captionTargetPageId="12" captionText="FIGURE 7. Macrobiotus papei sp. nov.—egg chorion morphology seen in SEM: A—entire egg with clearly visible flexible filaments on the egg processes; B–C—processes with filaments of variable length and the surface between processes; D— detail of the egg surface; E–F—details of the terminal disc and its flexible filaments. Scale bars in µm." httpUri="https://zenodo.org/record/1444187/files/figure.png" pageId="5" pageNumber="278">7A</figureCitation>
). The surface between processes of an intermediate
<typeStatus id="4E7E54A8784EFF80FC5AFCF4B3FC7071" box="[912,960,837,861]" pageId="5" pageNumber="278">type</typeStatus>
between the
<taxonomicName id="56C59189784EFF80FB90FCF5B4F87071" authorityName=", C. A. S. Schultze" authorityYear="1834" box="[1114,1220,836,861]" class="Eutardigrada" family="Macrobiotidae" genus="Macrobiotus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Parachela" pageId="5" pageNumber="278" phylum="Tardigrada" rank="species" species="hufelandi">
<emphasis id="A3B13618784EFF80FB90FCF5B4F87071" box="[1114,1220,836,861]" italics="true" pageId="5" pageNumber="278">hufelandi</emphasis>
</taxonomicName>
and the
<emphasis id="A3B13618784EFF80FAE3FCF5B5A07071" box="[1321,1436,836,861]" italics="true" pageId="5" pageNumber="278">maculatus</emphasis>
<typeStatus id="4E7E54A8784EFF80FF5DFCD9B0F670AC" box="[151,202,872,896]" pageId="5" pageNumber="278">type</typeStatus>
,
<emphasis id="A3B13618784EFF80FF1EFCD8B0CB70AC" box="[212,247,873,896]" italics="true" pageId="5" pageNumber="278">i.e.</emphasis>
chorion surface covered numerous small pores that form a reticulum with thick walls (
<figureCitation id="09FEF68F784EFF80FB71FCD6B50B70AC" box="[1211,1335,871,896]" captionStart="FIGURE 6" captionStartId="11.[151,250,1801,1823]" captionTargetBox="[184,1408,263,1785]" captionTargetId="figure@11.[184,1408,263,1785]" captionTargetPageId="11" captionText="FIGURE 6. Macrobiotus papei sp. nov.—egg seen in PCM: A—midsection; B—surface; C–D—magnification on surface; E—magnification on midsection; F—surface of terminal discs of egg processes. Filled arrowheads indicate thin flexible filaments at the edges of terminal discs on the midsections. A, B and D show the details of a single egg whereas other photos are taken from more than one egg. Scale bars in µm." httpUri="https://zenodo.org/record/1444185/files/figure.png" pageId="5" pageNumber="278">Figs 6B–D</figureCitation>
and
<figureCitation id="09FEF68F784EFF80FAA1FCD9B0967088" captionStart="FIGURE 7" captionStartId="12.[151,250,1924,1946]" captionTargetBox="[151,1436,213,1875]" captionTargetId="figure@12.[151,1436,213,1875]" captionTargetPageId="12" captionText="FIGURE 7. Macrobiotus papei sp. nov.—egg chorion morphology seen in SEM: A—entire egg with clearly visible flexible filaments on the egg processes; B–C—processes with filaments of variable length and the surface between processes; D— detail of the egg surface; E–F—details of the terminal disc and its flexible filaments. Scale bars in µm." httpUri="https://zenodo.org/record/1444187/files/figure.png" pageId="5" pageNumber="278">7A– F</figureCitation>
). Pores are similar in size and uniformly scattered over the entire surface, i.e. there no peribasal rings of elongated pores/bigger mesh around the processes. There are several rows of pores between processes and the mesh bars and nodes are often wider than the pore diameter (
<figureCitation id="09FEF68F784EFF80FC89FC65B38270C1" box="[835,958,980,1005]" captionStart="FIGURE 6" captionStartId="11.[151,250,1801,1823]" captionTargetBox="[184,1408,263,1785]" captionTargetId="figure@11.[184,1408,263,1785]" captionTargetPageId="11" captionText="FIGURE 6. Macrobiotus papei sp. nov.—egg seen in PCM: A—midsection; B—surface; C–D—magnification on surface; E—magnification on midsection; F—surface of terminal discs of egg processes. Filled arrowheads indicate thin flexible filaments at the edges of terminal discs on the midsections. A, B and D show the details of a single egg whereas other photos are taken from more than one egg. Scale bars in µm." httpUri="https://zenodo.org/record/1444185/files/figure.png" pageId="5" pageNumber="278">Figs 6C–D</figureCitation>
and
<figureCitation id="09FEF68F784EFF80FC3FFC65B40B70C0" box="[1013,1079,980,1004]" captionStart="FIGURE 7" captionStartId="12.[151,250,1924,1946]" captionTargetBox="[151,1436,213,1875]" captionTargetId="figure@12.[151,1436,213,1875]" captionTargetPageId="12" captionText="FIGURE 7. Macrobiotus papei sp. nov.—egg chorion morphology seen in SEM: A—entire egg with clearly visible flexible filaments on the egg processes; B–C—processes with filaments of variable length and the surface between processes; D— detail of the egg surface; E–F—details of the terminal disc and its flexible filaments. Scale bars in µm." httpUri="https://zenodo.org/record/1444187/files/figure.png" pageId="5" pageNumber="278">7B–F</figureCitation>
). The pores in the reticulum are circular or slightly oval (0.3–0.6 µm in diameter) and under SEM almost all pores are empty inside (
<figureCitation id="09FEF68F784EFF80FADCFC46B5AC773C" box="[1302,1424,1015,1040]" captionStart="FIGURE 7" captionStartId="12.[151,250,1924,1946]" captionTargetBox="[151,1436,213,1875]" captionTargetId="figure@12.[151,1436,213,1875]" captionTargetPageId="12" captionText="FIGURE 7. Macrobiotus papei sp. nov.—egg chorion morphology seen in SEM: A—entire egg with clearly visible flexible filaments on the egg processes; B–C—processes with filaments of variable length and the surface between processes; D— detail of the egg surface; E–F—details of the terminal disc and its flexible filaments. Scale bars in µm." httpUri="https://zenodo.org/record/1444187/files/figure.png" pageId="5" pageNumber="278">Fig. 7C–E</figureCitation>
). Processes are in the shape of inverted goblets with slightly concave conical trunks and well-defined terminal discs (
<figureCitation id="09FEF68F784EFF80FF55FB8EB1297774" box="[159,277,1087,1112]" captionStart="FIGURE 6" captionStartId="11.[151,250,1801,1823]" captionTargetBox="[184,1408,263,1785]" captionTargetId="figure@11.[184,1408,263,1785]" captionTargetPageId="11" captionText="FIGURE 6. Macrobiotus papei sp. nov.—egg seen in PCM: A—midsection; B—surface; C–D—magnification on surface; E—magnification on midsection; F—surface of terminal discs of egg processes. Filled arrowheads indicate thin flexible filaments at the edges of terminal discs on the midsections. A, B and D show the details of a single egg whereas other photos are taken from more than one egg. Scale bars in µm." httpUri="https://zenodo.org/record/1444185/files/figure.png" pageId="5" pageNumber="278">Figs 6E–F</figureCitation>
and
<figureCitation id="09FEF68F784EFF80FE86FBF1B1AC7774" box="[332,400,1088,1112]" captionStart="FIGURE 7" captionStartId="12.[151,250,1924,1946]" captionTargetBox="[151,1436,213,1875]" captionTargetId="figure@12.[151,1436,213,1875]" captionTargetPageId="12" captionText="FIGURE 7. Macrobiotus papei sp. nov.—egg chorion morphology seen in SEM: A—entire egg with clearly visible flexible filaments on the egg processes; B–C—processes with filaments of variable length and the surface between processes; D— detail of the egg surface; E–F—details of the terminal disc and its flexible filaments. Scale bars in µm." httpUri="https://zenodo.org/record/1444187/files/figure.png" pageId="5" pageNumber="278">7A–F</figureCitation>
). Terminal discs are cog-shaped, with a concave central area and with 10–15 small irregular teeth (
<figureCitation id="09FEF68F784EFF80FF14FBD5B1697751" box="[222,341,1124,1149]" captionStart="FIGURE 6" captionStartId="11.[151,250,1801,1823]" captionTargetBox="[184,1408,263,1785]" captionTargetId="figure@11.[184,1408,263,1785]" captionTargetPageId="11" captionText="FIGURE 6. Macrobiotus papei sp. nov.—egg seen in PCM: A—midsection; B—surface; C–D—magnification on surface; E—magnification on midsection; F—surface of terminal discs of egg processes. Filled arrowheads indicate thin flexible filaments at the edges of terminal discs on the midsections. A, B and D show the details of a single egg whereas other photos are taken from more than one egg. Scale bars in µm." httpUri="https://zenodo.org/record/1444185/files/figure.png" pageId="5" pageNumber="278">Figs 6E–F</figureCitation>
and
<figureCitation id="09FEF68F784EFF80FE45FBD5B1ED7750" box="[399,465,1124,1148]" captionStart="FIGURE 7" captionStartId="12.[151,250,1924,1946]" captionTargetBox="[151,1436,213,1875]" captionTargetId="figure@12.[151,1436,213,1875]" captionTargetPageId="12" captionText="FIGURE 7. Macrobiotus papei sp. nov.—egg chorion morphology seen in SEM: A—entire egg with clearly visible flexible filaments on the egg processes; B–C—processes with filaments of variable length and the surface between processes; D— detail of the egg surface; E–F—details of the terminal disc and its flexible filaments. Scale bars in µm." httpUri="https://zenodo.org/record/1444187/files/figure.png" pageId="5" pageNumber="278">7B–F</figureCitation>
). Almost all teeth on the terminal disc are elongated into thin flexible filaments, less than 0.3 µm in diameter and 2–7 µm in length (
<figureCitation id="09FEF68F784EFF80FD61FB36B31D778C" box="[683,801,1159,1184]" captionStart="FIGURE 6" captionStartId="11.[151,250,1801,1823]" captionTargetBox="[184,1408,263,1785]" captionTargetId="figure@11.[184,1408,263,1785]" captionTargetPageId="11" captionText="FIGURE 6. Macrobiotus papei sp. nov.—egg seen in PCM: A—midsection; B—surface; C–D—magnification on surface; E—magnification on midsection; F—surface of terminal discs of egg processes. Filled arrowheads indicate thin flexible filaments at the edges of terminal discs on the midsections. A, B and D show the details of a single egg whereas other photos are taken from more than one egg. Scale bars in µm." httpUri="https://zenodo.org/record/1444185/files/figure.png" pageId="5" pageNumber="278">Figs 6E–F</figureCitation>
, filled arrowhead and 7B–F). The filaments are hair-like under PCM (
<figureCitation id="09FEF68F784EFF80FEE7FB1DB19E77E9" box="[301,418,1196,1221]" captionStart="FIGURE 6" captionStartId="11.[151,250,1801,1823]" captionTargetBox="[184,1408,263,1785]" captionTargetId="figure@11.[184,1408,263,1785]" captionTargetPageId="11" captionText="FIGURE 6. Macrobiotus papei sp. nov.—egg seen in PCM: A—midsection; B—surface; C–D—magnification on surface; E—magnification on midsection; F—surface of terminal discs of egg processes. Filled arrowheads indicate thin flexible filaments at the edges of terminal discs on the midsections. A, B and D show the details of a single egg whereas other photos are taken from more than one egg. Scale bars in µm." httpUri="https://zenodo.org/record/1444185/files/figure.png" pageId="5" pageNumber="278">Fig. 6E–F</figureCitation>
) and under SEM are completely smooth as is the surface of terminal discs (
<figureCitation id="09FEF68F784EFF80FADFFB1DB5AC77E8" box="[1301,1424,1196,1221]" captionStart="FIGURE 7" captionStartId="12.[151,250,1924,1946]" captionTargetBox="[151,1436,213,1875]" captionTargetId="figure@12.[151,1436,213,1875]" captionTargetPageId="12" captionText="FIGURE 7. Macrobiotus papei sp. nov.—egg chorion morphology seen in SEM: A—entire egg with clearly visible flexible filaments on the egg processes; B–C—processes with filaments of variable length and the surface between processes; D— detail of the egg surface; E–F—details of the terminal disc and its flexible filaments. Scale bars in µm." httpUri="https://zenodo.org/record/1444187/files/figure.png" pageId="5" pageNumber="278">Fig. 7D–F</figureCitation>
). Under PCM, the filaments are clearly visible, however the filaments are very thin and could be overlooked and/or misinterpreted as debris attached to the eggs, if close attention is not paid during examination.
</paragraph>
<paragraph id="917AEA0A784EFF80FF0DFAA6B1127559" blockId="5.[151,1437,151,1725]" pageId="5" pageNumber="278">
<materialsCitation id="21ADE057784EFF80FF0DFAA6B1127559" ID-GBIF-Occurrence="1914122839" collectionCode="PCM" county="Only" location="Furthermore" municipality="Moreover" pageId="5" pageNumber="278" specimenCount="50" specimenCount-adult="50" stateProvince="Nelson" typeStatus="holotype">
<emphasis id="A3B13618784EFF80FF0DFAA6B186761C" bold="true" box="[199,442,1303,1328]" pageId="5" pageNumber="278">Reproductive mode.</emphasis>
Staining
<specimenCount id="87C32183784EFF80FDE3FAA6B2AD761C" box="[553,657,1303,1328]" pageId="5" pageNumber="278" type="adult">50 adults</specimenCount>
of the new species with aceto-orcein did not reveal any individuals with a testicle filled with spermatozoa (males), an ovotestis with spermatozoa and oocytes (hermaphrodites), or with spermatozoa in a spermatheca (inseminated females).
<collectingCounty id="781B9286784EFF80FCE7FAEEB3597654" box="[813,869,1375,1400]" pageId="5" pageNumber="278">Only</collectingCounty>
females with oocytes in the ovary or specimens of an undetermined sex were observed.
<collectingMunicipality id="711E7070784EFF80FD98FA34B2FE76B0" box="[594,706,1413,1436]" pageId="5" pageNumber="278">Moreover</collectingMunicipality>
, both
<collectionCode id="F7D472CF784EFF80FCDBFA35B37076B0" box="[785,844,1412,1436]" country="Poland" httpUri="http://grbio.org/cool/f3v1-41fg" name="Polish Collection of Microorganisms" pageId="5" pageNumber="278">PCM</collectionCode>
and SEM observations showed no morphological secondary sexual dimorphism such as gibbosities on legs IV (
<emphasis id="A3B13618784EFF80FC9FFA18B34176EC" box="[853,893,1449,1472]" italics="true" pageId="5" pageNumber="278">e.g.</emphasis>
<bibRefCitation id="F55497FB784EFF80FC4FFA16B49B76EC" box="[901,1191,1447,1472]" pageId="5" pageNumber="278" refId="ref8666" refString="Rebecchi, L. &amp; Nelson, D. R. (1998) Evaluation of a secondary sex character in eutardigrades. Invertebrate Biology, 117, 194 - 198." type="journal article">
Rebecchi &amp;
<collectingRegion id="530124E8784EFF80FBDEFA16B45F76EC" box="[1044,1123,1447,1472]" country="New Zealand" name="Nelson" pageId="5" pageNumber="278">Nelson</collectingRegion>
1998
</bibRefCitation>
).
<location id="941ABCD1784EFF80FB76FA16B57776EC" LSID="urn:lsid:plazi:treatment:196C5B1C784FFF8BFF5DFC0EB3F771E4:941ABCD1784EFF80FB76FA16B57776EC" box="[1212,1355,1447,1472]" county="Only" municipality="Moreover" name="Furthermore" pageId="5" pageNumber="278" stateProvince="Nelson">Furthermore</location>
, to test whether females are capable of parthenogenetic reproduction, we isolated eggs and observed the full cycle of hatchlings maturing to adults, laying the next generation of eggs still in isolation, and the emergence of the second generation, also from individually isolated eggs (
<emphasis id="A3B13618784EFF80FD1AF9A4B2CE7500" box="[720,754,1557,1580]" italics="true" pageId="5" pageNumber="278">i.e.</emphasis>
there was no possibility of fertilisation).
<location id="941ABCD1784EFF80FB1CF9A5B52D7501" LSID="urn:lsid:plazi:treatment:196C5B1C784FFF8BFF5DFC0EB3F771E4:941ABCD1784EFF80FB1CF9A5B52D7501" box="[1238,1297,1556,1581]" county="Only" municipality="Moreover" name="Thus" pageId="5" pageNumber="278" stateProvince="Nelson">Thus</location>
, all lines of evidence show the
<typeStatus id="4E7E54A8784EFF80FEB7F988B191757D" box="[381,429,1593,1617]" pageId="5" pageNumber="278">type</typeStatus>
population of
<taxonomicName id="56C59189784EFF80FDA8F988B2F6757C" authority="Stec &amp; Kristensen &amp; Michalczyk, 2018" authorityName="Stec &amp; Kristensen &amp; Michalczyk" authorityYear="2018" box="[610,714,1593,1616]" class="Eutardigrada" family="Macrobiotidae" genus="Macrobiotus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Parachela" pageId="5" pageNumber="278" phylum="Tardigrada" rank="species" species="papei" status="sp. nov.">
<emphasis id="A3B13618784EFF80FDA8F988B2F6757C" box="[610,714,1593,1616]" italics="true" pageId="5" pageNumber="278">M. papei</emphasis>
</taxonomicName>
<taxonomicNameLabel id="B8828B63784EFF80FD12F986B30B757C" box="[728,823,1591,1616]" pageId="5" pageNumber="278" rank="species">
<emphasis id="A3B13618784EFF80FD12F986B30B757C" bold="true" box="[728,823,1591,1616]" pageId="5" pageNumber="278">sp. nov.</emphasis>
</taxonomicNameLabel>
comprises only females capable of parthenogenetic reproduction.
</materialsCitation>
</paragraph>
<paragraph id="917AEA0A784EFF80FF0DF9CEB4CB7591" blockId="5.[151,1437,151,1725]" pageId="5" pageNumber="278">
<emphasis id="A3B13618784EFF80FF0DF9CEB1B575B4" bold="true" box="[199,393,1663,1688]" pageId="5" pageNumber="278">DNA sequences.</emphasis>
We obtained very good quality sequences for all four molecular markers from all analysed specimens (paragenophores). DNA sequences of all markers were represented by single haplotypes:
</paragraph>
<paragraph id="917AEA0A784EFF80FF5DF957B35E75D3" blockId="5.[151,866,1766,1791]" box="[151,866,1766,1791]" pageId="5" pageNumber="278">
The
<emphasis id="A3B13618784EFF80FF03F957B17975D3" bold="true" box="[201,325,1766,1791]" pageId="5" pageNumber="278">18S rRNA</emphasis>
sequence (GenBank:
<accessionNumber id="8E9677E9784EFF80FDF6F957B2FD75D3" box="[572,705,1766,1791]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/MH063881" pageId="5" pageNumber="278">MH063881</accessionNumber>
), 905 bp long:
</paragraph>
<paragraph id="917AEA0A784EFF83FF5DF89AB21B7271" blockId="5.[151,1431,1835,2030]" box="[151,1431,154,2030]" lastBlockId="6.[151,1431,154,349]" lastPageId="6" lastPageNumber="279" pageId="5" pageNumber="278">TAGATCGTAATTTTACACGGATAACTGTGGTAATTCTAGAGCTAATACGTGCAACCAGCTCGTTCCCTTGTGGAGCGAGC GCAGTTATTAGAACAAGACCAATCCGGCCTTCGGGTCGGTACAATTGGTGACTCTGAATAACCGAAGCGGAGCGCATGGT CTCGTACCGGCGCCAGATCTTTCAAGTGTCTGACTTATCAGCTTGTTGTTAGGTTATGTTCCTAACAAGGCTTCAACGGG TAACGGGGTATCAGGGTCCGATACCGGAGAGGGAGCCTGAGAAACGGCTACCACATCCAAGGAAGGCAGCAGGCGCGCAA ATTACCCACTCCTAGCACAGGGAGGTAGTGACGAAAAATAACGATGCGAGGGCTAATAGCTTCTCGTAATCGGAATGGGT ACACTTTAAATCCTTTAACGAGGATCTATTGGAGGGCAAGTCTGGTGCCAGCAGCCGCGGTAATTCCAGCTCCAATAGCG TATATTAAAGTTGCTGCGGTTAAAAGCTCGTAGTTGGATCTGGGCTTCTGAATGGATGGTTCACTTTACGGTGTAACTGT TCGTTTGGTGCCACAAGCCGGCCATGTCTTGCATGCCCTTTACTGGGTGTGCATGGCGACCGGAACGTTTACTTTGAAAA AATTAGAGTGCTCAAAGCAGGCGTATGGCCTTGCATAATGGTGCATGGAATAATGGAATAGGACCTCGGTTCTATTTTGT TGGTTTTCGGAACTCGAGGTAATGATTAAGAGGAACAGACGGGGGCATTCGTATTGCGGCGTTAGAGGTGAAATTCTTGG ATCGTCGCAAGACGAACTACTGCGAAAGCATTTGCCAAGAATGTTTTCATTAATCAAGAACGAAAGTTAGAGGTTCGAAG GCGATCAGATACCGCCCTAGTTCTA</paragraph>
<caption id="C5BABA82784DFF83FF5DF8DDB38F74EC" httpUri="https://zenodo.org/record/1444175/files/figure.png" pageId="6" pageNumber="279" startId="6.[151,250,1900,1922]" targetBox="[163,1427,405,1878]" targetPageId="6">
<paragraph id="917AEA0A784DFF83FF5DF8DDB38F74EC" blockId="6.[151,1436,1900,1984]" pageId="6" pageNumber="279">
<emphasis id="A3B13618784DFF83FF5DF8DDB29874AE" bold="true" box="[151,676,1900,1923]" pageId="6" pageNumber="279">
FIGURE 1.
<taxonomicName id="56C59189784DFF83FED6F8DDB1DC74AE" authority="Stec &amp; Kristensen &amp; Michalczyk, 2018" authorityName="Stec &amp; Kristensen &amp; Michalczyk" authorityYear="2018" box="[284,480,1900,1923]" class="Eutardigrada" family="Macrobiotidae" genus="Macrobiotus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Parachela" pageId="6" pageNumber="279" phylum="Tardigrada" rank="species" species="papei" status="sp. nov.">
<emphasis id="A3B13618784DFF83FED6F8DDB1DC74AE" bold="true" box="[284,480,1900,1923]" italics="true" pageId="6" pageNumber="279">Macrobiotus papei</emphasis>
</taxonomicName>
<taxonomicNameLabel id="B8828B63784DFF83FE22F8DDB20574AE" box="[488,569,1900,1922]" pageId="6" pageNumber="279" rank="species">sp. nov.</taxonomicNameLabel>
—habitus
</emphasis>
:
<emphasis id="A3B13618784DFF83FD79F8DDB2F974AE" bold="true" box="[691,709,1900,1922]" pageId="6" pageNumber="279">A</emphasis>
—dorso-ventral projection (holotype, Hoyer’s medium, PCM);
<emphasis id="A3B13618784DFF83FA99F8DDB5BE74AE" bold="true" box="[1363,1410,1900,1922]" pageId="6" pageNumber="279">B–C</emphasis>
— cuticular pores seen in PCM on the dorsal (B) and ventral (C) part of the body (paratype);
<emphasis id="A3B13618784DFF83FBF9F83AB45F748D" bold="true" box="[1075,1123,1931,1953]" pageId="6" pageNumber="279">D–E</emphasis>
—cuticular pores seen in SEM on the dorsal (D) and ventral (E) part of the body (paratype). Scale bars in µm.
</paragraph>
</caption>
<caption id="C5BABA82784CFF82FF5DF8AFB51E74E1" httpUri="https://zenodo.org/record/1444177/files/figure.png" pageId="7" pageNumber="280" startId="7.[151,250,1822,1844]" targetBox="[151,1436,203,1787]" targetPageId="7">
<paragraph id="917AEA0A784CFF82FF5DF8AFB51E74E1" blockId="7.[151,1436,1821,1997]" pageId="7" pageNumber="280">
<emphasis id="A3B13618784CFF82FF5DF8AFB3907418" bold="true" box="[151,940,1821,1844]" pageId="7" pageNumber="280">
FIGURE 2.
<taxonomicName id="56C59189784CFF82FED2F8ACB1E6741F" authority="Stec &amp; Kristensen &amp; Michalczyk, 2018" authorityName="Stec &amp; Kristensen &amp; Michalczyk" authorityYear="2018" box="[280,474,1821,1844]" class="Eutardigrada" family="Macrobiotidae" genus="Macrobiotus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Parachela" pageId="7" pageNumber="280" phylum="Tardigrada" rank="species" species="papei" status="sp. nov.">
<emphasis id="A3B13618784CFF82FED2F8ACB1E6741F" bold="true" box="[280,474,1821,1844]" italics="true" pageId="7" pageNumber="280">Macrobiotus papei</emphasis>
</taxonomicName>
<taxonomicNameLabel id="B8828B63784CFF82FE2AF8AFB2137418" box="[480,559,1822,1844]" pageId="7" pageNumber="280" rank="species">sp. nov.</taxonomicNameLabel>
—cuticular structures on legs: A–B
</emphasis>
—external granulation on leg II seen in PCM (A) and SEM (B) (paratypes);
<emphasis id="A3B13618784CFF82FE62F88DB1CF747E" bold="true" box="[424,499,1852,1874]" pageId="7" pageNumber="280">C–D—</emphasis>
a cuticular bulge resembling a pulvinus, faint cuticular fold on the internal surface of leg II with visible muscle attachment seen in PCM (C) and SEM (D) (paratypes);
<emphasis id="A3B13618784CFF82FC63F8EAB386745D" bold="true" box="[937,954,1883,1905]" pageId="7" pageNumber="280">E</emphasis>
—two patches of granulation on leg IV (PCM, paratype);
<emphasis id="A3B13618784CFF82FEC9F8CBB11074BC" bold="true" box="[259,300,1914,1936]" pageId="7" pageNumber="280">F—</emphasis>
distal patch of granulation on leg IV (SEM, paratype). Filled, flat arrowheads indicate the cuticular bulge, empty, flat arrowheads indicate the faint cuticular fold under the claws, arrows indicate muscle attachments, whereas filled and empty indented arrowheads indicate proximal and distal patch of granulation on the leg IV respectively. Scale bars in µm.
</paragraph>
</caption>
<caption id="C5BABA827843FF8DFF5DF8B4B08374DE" httpUri="https://zenodo.org/record/1444179/files/figure.png" pageId="8" pageNumber="281" startId="8.[151,250,1797,1819]" targetBox="[151,1436,213,1756]" targetPageId="8">
<paragraph id="917AEA0A7843FF8DFF5DF8B4B08374DE" blockId="8.[151,1436,1797,2034]" pageId="8" pageNumber="281">
<emphasis id="A3B136187843FF8DFF5DF8B4B1107416" bold="true" pageId="8" pageNumber="281">
FIGURE 3.
<taxonomicName id="56C591897843FF8DFEE9F8B4B1D67437" authority="Stec &amp; Kristensen &amp; Michalczyk, 2018" authorityName="Stec &amp; Kristensen &amp; Michalczyk" authorityYear="2018" box="[291,490,1797,1820]" class="Eutardigrada" family="Macrobiotidae" genus="Macrobiotus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Parachela" pageId="8" pageNumber="281" phylum="Tardigrada" rank="species" species="papei" status="sp. nov.">
<emphasis id="A3B136187843FF8DFEE9F8B4B1D67437" bold="true" box="[291,490,1797,1820]" italics="true" pageId="8" pageNumber="281">Macrobiotus papei</emphasis>
</taxonomicName>
<taxonomicNameLabel id="B8828B637843FF8DFE3CF8B4B2777437" box="[502,587,1797,1819]" pageId="8" pageNumber="281" rank="species">sp. nov.</taxonomicNameLabel>
—buccal apparatus and the oral cavity armature (OCA) seen in PCM (all paratypes): A
</emphasis>
—dorso-ventral projection of the entire buccal apparatus;
<emphasis id="A3B136187843FF8DFCB5F895B3927416" bold="true" box="[895,942,1828,1850]" pageId="8" pageNumber="281">B–C</emphasis>
oral cavity armature detected in the majority of specimens, i.e. OCA of the
<emphasis id="A3B136187843FF8DFE79F8F5B2137475" box="[435,559,1860,1881]" italics="true" pageId="8" pageNumber="281">patagonicus</emphasis>
type (
<emphasis id="A3B136187843FF8DFDB8F8F5B2AC7475" box="[626,656,1860,1881]" italics="true" pageId="8" pageNumber="281">i.e.</emphasis>
the second and third band of teeth visible), dorsal (B) and ventral (C) view, respectively;
<emphasis id="A3B136187843FF8DFED4F8D0B173745B" bold="true" box="[286,335,1889,1911]" pageId="8" pageNumber="281">D–E</emphasis>
oral cavity armature detected in small specimens, i.e. OCA of the
<emphasis id="A3B136187843FF8DFC38F8D0B466745B" box="[1010,1114,1889,1911]" italics="true" pageId="8" pageNumber="281">maculatus</emphasis>
type (
<emphasis id="A3B136187843FF8DFB51F8D3B485745B" box="[1179,1209,1890,1911]" italics="true" pageId="8" pageNumber="281">i.e.</emphasis>
only the third band of teeth visible), dorsal (D) and ventral (E) view, respectively;
<emphasis id="A3B136187843FF8DFD3FF831B30274BA" bold="true" box="[757,830,1920,1942]" pageId="8" pageNumber="281">F–G—</emphasis>
placoid morphology. The arrow indicates pores surrounding mouth opening, flat empty arrowheads indicate the second band of teeth in the oral cavity, filled indented arrowheads indicate the third band of teeth in the oral cavity whereas filled arrowheads indicate faint constrictions in macroplacoids. Scale bars in µm.
</paragraph>
</caption>
<caption id="C5BABA827842FF8CFF5DF953B13674C5" httpUri="https://zenodo.org/record/1444181/files/figure.png" pageId="9" pageNumber="282" startId="9.[151,250,1762,1784]" targetBox="[191,1396,185,1745]" targetPageId="9">
<paragraph id="917AEA0A7842FF8CFF5DF953B13674C5" blockId="9.[151,1436,1762,2025]" pageId="9" pageNumber="282">
<emphasis id="A3B136187842FF8CFF5DF953B57475D4" bold="true" box="[151,1352,1762,1785]" pageId="9" pageNumber="282">
FIGURE 4.
<taxonomicName id="56C591897842FF8CFED2F953B1E675DB" authority="Stec &amp; Kristensen &amp; Michalczyk, 2018" authorityName="Stec &amp; Kristensen &amp; Michalczyk" authorityYear="2018" box="[280,474,1762,1785]" class="Eutardigrada" family="Macrobiotidae" genus="Macrobiotus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Parachela" pageId="9" pageNumber="282" phylum="Tardigrada" rank="species" species="papei" status="sp. nov.">
<emphasis id="A3B136187842FF8CFED2F953B1E675DB" bold="true" box="[280,474,1762,1785]" italics="true" pageId="9" pageNumber="282">Macrobiotus papei</emphasis>
</taxonomicName>
<taxonomicNameLabel id="B8828B637842FF8CFE2AF953B21375D4" box="[480,559,1762,1784]" pageId="9" pageNumber="282" rank="species">sp. nov.</taxonomicNameLabel>
—mouth opening and the oral cavity armature seen in SEM (paratype): A
</emphasis>
– mouth opening with six peribuccal sensory lobes and ten lamellae surrounded by large pores;
<emphasis id="A3B136187842FF8CFBE2F8B5B46B7436" bold="true" box="[1064,1111,1796,1818]" pageId="9" pageNumber="282">B–C</emphasis>
—the oral cavity armature of a single paratype seen in SEM from different angles. Arrows indicate pores in the irregular ring of pores surrounding the peribuccal lobes, filled arrowheads indicate teeth of the first band, empty arrowheads indicate teeth of the second band, the peaks in the dorsal ridge of the third band that correspond with teeth of the third band, in species with better developed oral cavity armatures are marked with “L d” and “M d” (the dorso-median tooth comprises several small peaks visible as thickenings in PCM), whereas the teeth of the ventral portion are marked with “M v” (medio-ventral tooth) and “L v” (lateral teeth). Scale bars in µm.
</paragraph>
</caption>
<paragraph id="917AEA0A7841FF8FFF5DFF26B35E739C" blockId="10.[151,866,151,176]" box="[151,866,151,176]" pageId="10" pageNumber="283">
The
<emphasis id="A3B136187841FF8FFF03FF26B179739C" bold="true" box="[201,325,151,176]" pageId="10" pageNumber="283">28S rRNA</emphasis>
sequence (GenBank:
<accessionNumber id="8E9677E97841FF8FFDF6FF26B2FD739C" box="[572,705,151,176]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/MH063880" pageId="10" pageNumber="283">MH063880</accessionNumber>
), 787 bp long:
</paragraph>
<paragraph id="917AEA0A7841FF8FFF5DFF53B4FB711C" blockId="10.[151,1431,226,560]" box="[151,1431,226,560]" pageId="10" pageNumber="283">TACTAAGCGGAGGAAAAGAAACCAACGGGGATGCCGACAGTAACTGCGAGTGAAATCGGCCAAGCCCAGCGCCGAATCCT GTTGCTGGTAACGGTGGTAGGAACTGTGGCGTGAAGAACGTCCTTACCGGTACGGTTTGCGTGCGTAAGTTCTCCTGAGT GAGGCTCCATTCCAAGGAGGGTGCAAGACCCGTATCGCGTGCAACCGGTATCGGTGTAAGATGTTCGGAGAGTCGCCTTG TTTGTGAGTACAAGGTGAAGTCGGTGGTAAACTCCATCGAAGGCTAAATATGACCACGAGTCCGATAGCGAACAAGTACC GTGAGGGAAAATTGAAAAGCACTTTGAAGAGAGAGCGAAACAGTGCGTGAAACCGCTCAGAGGCAAGCAAATGGGGCCTC GAAGGCAAGGCAGCGAATTCAGCTGGTGGTCTGCGTGGCTGGCCGGTTAAGTGATCTTAACGACTCTTGCCGGTTATGTC TAGCGTAGGTGCCAGTGCACTTTCGTTGCTTGTACGCCACCGCCGTTGAGTGGGCATCCGTCGGGTTGGTAACGCGAAGC CTTACGCCTTCACGGGCGTAGGTGCTTGCAGCCGACTTTGTACGCGTTTGCACTTCAACCGGTCATGTTTGCATGTGCCA GCATTTGCGTTGGATTGGCTCGCTCTGCCGTTTGTCGTGAGATGACGAGCTTGCTCGGCTCTTCGGCATCTATGGTAGAC TCGTGTCGGTTTTCAACGTGGGCACATTGTTAATTCGGTGGCGAGTAGATGGCTGCCCATTTAACCC</paragraph>
<caption id="C5BABA827841FF8FFF5DF904B2F27424" httpUri="https://zenodo.org/record/1444183/files/figure.png" pageId="10" pageNumber="283" startId="10.[151,250,1717,1739]" targetBox="[151,1436,603,1706]" targetPageId="10">
<paragraph id="917AEA0A7841FF8FFF5DF904B2F27424" blockId="10.[151,1436,1716,1800]" pageId="10" pageNumber="283">
<emphasis id="A3B136187841FF8FFF5DF904B37175E7" bold="true" box="[151,845,1716,1739]" pageId="10" pageNumber="283">
FIGURE 5.
<taxonomicName id="56C591897841FF8FFED0F905B1E175E6" authority="Stec &amp; Kristensen &amp; Michalczyk, 2018" authorityName="Stec &amp; Kristensen &amp; Michalczyk" authorityYear="2018" box="[282,477,1716,1739]" class="Eutardigrada" family="Macrobiotidae" genus="Macrobiotus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Parachela" pageId="10" pageNumber="283" phylum="Tardigrada" rank="species" species="papei" status="sp. nov.">
<emphasis id="A3B136187841FF8FFED0F905B1E175E6" bold="true" box="[282,477,1716,1739]" italics="true" pageId="10" pageNumber="283">Macrobiotus papei</emphasis>
</taxonomicName>
<taxonomicNameLabel id="B8828B637841FF8FFE2FF904B20A75E7" box="[485,566,1717,1739]" pageId="10" pageNumber="283" rank="species">sp. nov.</taxonomicNameLabel>
—claws (paratypes): A–B
</emphasis>
—claws II and IV seen in PCM, with smooth and slightly dentate lunules, respectively;
<emphasis id="A3B136187841FF8FFE08F965B1CF75C6" bold="true" box="[450,499,1748,1770]" pageId="10" pageNumber="283">C–D</emphasis>
—claws I and IV seen in SEM, with smooth and slightly dentate lunules, respectively. Figs A and B assembled from several photos. Scale bars in µm.
</paragraph>
</caption>
<paragraph id="917AEA0A7841FF8FFF5DF884B3177462" blockId="10.[151,811,1845,1870]" box="[151,811,1845,1870]" pageId="10" pageNumber="283">
The
<emphasis id="A3B136187841FF8FFF03F884B1327462" bold="true" box="[201,270,1845,1870]" pageId="10" pageNumber="283">ITS-2</emphasis>
sequence (GenBank:
<accessionNumber id="8E9677E97841FF8FFDCEF884B2B67462" box="[516,650,1845,1870]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/MH063921" pageId="10" pageNumber="283">MH063921</accessionNumber>
), 397 bp long:
</paragraph>
<paragraph id="917AEA0A7841FF8FFF5DF8CEB5AB74F5" blockId="10.[151,1431,1919,2009]" box="[151,1431,1919,2009]" pageId="10" pageNumber="283">TTTGTGAACGTTAATTCTTCGAACGCACATTGCGGCTTCGGGTTAACTGAAGCCATGCCTGGTTGAGGGTCAGTTGAAGA AAAAAATCGTAATCGCGCATTGATTACGGAGTGTCTGGTTAATGGCTCGTCCGTTTCCAGATGAAGTATAGACCAGATGT GTGCGCTCATTTGACCGGTGCAAGCAACGCTTTGCCGAGTTGGAGCATTCGGCTTTCTTAGCCGTGCGCCGCAGTTGCAC</paragraph>
<paragraph id="917AEA0A7840FF8EFF5DFF2BB55B73FE" blockId="11.[151,1431,154,210]" box="[151,1431,154,210]" pageId="11" pageNumber="284">GATGGCTAAGTTGGCTACCAACAATGGCGAAGTAAGACCGGTTCAGAGGTGCGCAACGCAATAGGCACATCTGTGTACCA AAAAGAACGTACGGTCGCGGTGTTTCGACCGATGCGGACTTAACTCATTCTTTTGACCTCAGCTCAGACAAGATTAC</paragraph>
<caption id="C5BABA827840FF8EFF5DF8B8B2957457" httpUri="https://zenodo.org/record/1444185/files/figure.png" pageId="11" pageNumber="284" startId="11.[151,250,1801,1823]" targetBox="[184,1408,263,1785]" targetPageId="11">
<paragraph id="917AEA0A7840FF8EFF5DF8B8B2957457" blockId="11.[151,1436,1800,1915]" pageId="11" pageNumber="284">
<emphasis id="A3B136187840FF8EFF5DF8B8B30D7433" bold="true" box="[151,817,1800,1823]" pageId="11" pageNumber="284">
FIGURE 6.
<taxonomicName id="56C591897840FF8EFED6F8B9B1DD7432" authority="Stec &amp; Kristensen &amp; Michalczyk, 2018" authorityName="Stec &amp; Kristensen &amp; Michalczyk" authorityYear="2018" box="[284,481,1800,1823]" class="Eutardigrada" family="Macrobiotidae" genus="Macrobiotus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Parachela" pageId="11" pageNumber="284" phylum="Tardigrada" rank="species" species="papei" status="sp. nov.">
<emphasis id="A3B136187840FF8EFED6F8B9B1DD7432" bold="true" box="[284,481,1800,1823]" italics="true" pageId="11" pageNumber="284">Macrobiotus papei</emphasis>
</taxonomicName>
<taxonomicNameLabel id="B8828B637840FF8EFE23F8B8B2067433" box="[489,570,1801,1823]" pageId="11" pageNumber="284" rank="species">sp. nov.</taxonomicNameLabel>
—egg seen in PCM: A
</emphasis>
—midsection;
<emphasis id="A3B136187840FF8EFC03F8B8B3E67433" bold="true" box="[969,986,1801,1823]" pageId="11" pageNumber="284">B</emphasis>
—surface;
<emphasis id="A3B136187840FF8EFB81F8B8B4AA7433" bold="true" box="[1099,1174,1801,1823]" pageId="11" pageNumber="284">C–D—</emphasis>
magnification on surface;
<emphasis id="A3B136187840FF8EFF5DF899B0FD7412" bold="true" box="[151,193,1832,1854]" pageId="11" pageNumber="284">E—</emphasis>
magnification on midsection;
<emphasis id="A3B136187840FF8EFE34F899B2317412" bold="true" box="[510,525,1832,1854]" pageId="11" pageNumber="284">F</emphasis>
—surface of terminal discs of egg processes. Filled arrowheads indicate thin flexible filaments at the edges of terminal discs on the midsections. A, B and D show the details of a single egg whereas other photos are taken from more than one egg. Scale bars in µm.
</paragraph>
</caption>
<caption id="C5BABA827847FF89FF5DF835B49474F4" httpUri="https://zenodo.org/record/1444187/files/figure.png" pageId="12" pageNumber="285" startId="12.[151,250,1924,1946]" targetBox="[151,1436,213,1875]" targetPageId="12">
<paragraph id="917AEA0A7847FF89FF5DF835B49474F4" blockId="12.[151,1436,1924,2008]" pageId="12" pageNumber="285">
<emphasis id="A3B136187847FF89FF5DF835B43E74B6" bold="true" box="[151,1026,1924,1947]" pageId="12" pageNumber="285">
FIGURE 7.
<taxonomicName id="56C591897847FF89FED3F835B1E074B6" authority="Stec &amp; Kristensen &amp; Michalczyk, 2018" authorityName="Stec &amp; Kristensen &amp; Michalczyk" authorityYear="2018" box="[281,476,1924,1947]" class="Eutardigrada" family="Macrobiotidae" genus="Macrobiotus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Parachela" pageId="12" pageNumber="285" phylum="Tardigrada" rank="species" species="papei" status="sp. nov.">
<emphasis id="A3B136187847FF89FED3F835B1E074B6" bold="true" box="[281,476,1924,1947]" italics="true" pageId="12" pageNumber="285">Macrobiotus papei</emphasis>
</taxonomicName>
<taxonomicNameLabel id="B8828B637847FF89FE28F835B20E74B6" box="[482,562,1924,1946]" pageId="12" pageNumber="285" rank="species">sp. nov.</taxonomicNameLabel>
—egg chorion morphology seen in SEM: A
</emphasis>
—entire egg with clearly visible flexible filaments on the egg processes;
<emphasis id="A3B136187847FF89FE2DF812B22A7495" bold="true" box="[487,534,1955,1977]" pageId="12" pageNumber="285">B–C</emphasis>
—processes with filaments of variable length and the surface between processes;
<emphasis id="A3B136187847FF89FABAF812B5BE7495" bold="true" box="[1392,1410,1955,1977]" pageId="12" pageNumber="285">D</emphasis>
— detail of the egg surface;
<emphasis id="A3B136187847FF89FE5DF873B1F874F4" bold="true" box="[407,452,1986,2008]" pageId="12" pageNumber="285">E–F</emphasis>
—details of the terminal disc and its flexible filaments. Scale bars in µm.
</paragraph>
</caption>
<caption id="C5BABA827846FF88FF5DFF29B4A773C3" pageId="13" pageNumber="286">
<paragraph id="917AEA0A7846FF88FF5DFF29B4A773C3" blockId="13.[151,1435,152,239]" pageId="13" pageNumber="286">
<emphasis id="A3B136187846FF88FF5DFF29B1327383" bold="true" box="[151,270,152,175]" pageId="13" pageNumber="286">TABLE 3.</emphasis>
Measurements [in µm] and the
<emphasis id="A3B136187846FF88FD4BFF29B2AA7383" box="[641,662,152,175]" italics="true" pageId="13" pageNumber="286">pt</emphasis>
values [in %] of selected morphological structures of individuals of
<taxonomicName id="56C591897846FF88FF5DFF06B15C73E3" authority="Stec &amp; Kristensen &amp; Michalczyk, 2018" authorityName="Stec &amp; Kristensen &amp; Michalczyk" authorityYear="2018" box="[151,352,183,207]" class="Eutardigrada" family="Macrobiotidae" genus="Macrobiotus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Parachela" pageId="13" pageNumber="286" phylum="Tardigrada" rank="species" species="papei" status="sp. nov.">
<emphasis id="A3B136187846FF88FF5DFF06B15C73E3" box="[151,352,183,207]" italics="true" pageId="13" pageNumber="286">Macrobiotus papei</emphasis>
</taxonomicName>
<taxonomicNameLabel id="B8828B637846FF88FEA2FF09B18273E3" box="[360,446,184,207]" pageId="13" pageNumber="286" rank="species">
<emphasis id="A3B136187846FF88FEA2FF09B18273E3" bold="true" box="[360,446,184,207]" pageId="13" pageNumber="286">sp. nov.</emphasis>
</taxonomicNameLabel>
mounted in Hoyer’s medium (N–number of specimens/structures measured, RANGE refers to the smallest and the largest structure among all measured specimens; SD–standard deviation).
</paragraph>
</caption>
<paragraph id="917AEA0A7846FF88FF55FEBBB5B875B2" pageId="13" pageNumber="286">
<table id="E3C518AA7846007AFF55FEBBB5B975B2" box="[159,1413,266,1694]" gridcols="10" gridrows="35" pageId="13" pageNumber="286">
<tr id="2FF5E8487846007AFF55FEBBB5B9720C" box="[159,1413,266,288]" gridrow="0" pageId="13" pageNumber="286">
<th id="6C2481347846007AFF55FEBBB1F7720C" box="[159,459,266,288]" gridcol="0" gridrow="0" pageId="13" pageNumber="286">CHARACTER</th>
<th id="6C2481347846007AFDC6FEBBB21B720C" box="[524,551,266,288]" gridcol="1" gridrow="0" pageId="13" pageNumber="286">N</th>
<th id="6C2481347846007AFD92FEBBB36F720C" box="[600,851,266,288]" colspan="2" colspanRight="1" gridcol="2" gridrow="0" pageId="13" pageNumber="286">RANGE</th>
<th id="6C2481347846007AFC5DFEBBB411720C" box="[919,1069,266,288]" colspan="2" colspanRight="1" gridcol="4" gridrow="0" pageId="13" pageNumber="286">MEAN</th>
<th id="6C2481347846007AFB95FEBBB4EB720C" box="[1119,1239,266,288]" colspan="2" colspanRight="1" gridcol="6" gridrow="0" pageId="13" pageNumber="286">SD</th>
<th id="6C2481347846007AFAC8FEBBB5B9720C" box="[1282,1413,266,288]" colspan="2" colspanRight="1" gridcol="8" gridrow="0" pageId="13" pageNumber="286">Holotype</th>
</tr>
<tr id="2FF5E8487846007AFF55FE84B5B97266" box="[159,1413,309,330]" gridrow="1" pageId="13" pageNumber="286" rowspan-0="1" rowspan-1="1">
<td id="6C2481347846007AFD92FE84B2827266" box="[600,702,309,330]" gridcol="2" gridrow="1" pageId="13" pageNumber="286">µm</td>
<td id="6C2481347846007AFD27FE84B36F7266" box="[749,851,309,330]" gridcol="3" gridrow="1" pageId="13" pageNumber="286">
<emphasis id="A3B136187846FF88FCDFFE84B3157266" box="[789,809,309,330]" italics="true" pageId="13" pageNumber="286">pt</emphasis>
</td>
<td id="6C2481347846007AFC5DFE84B3FF7266" box="[919,963,309,330]" gridcol="4" gridrow="1" pageId="13" pageNumber="286">µm</td>
<td id="6C2481347846007AFC34FE84B4117266" box="[1022,1069,309,330]" gridcol="5" gridrow="1" pageId="13" pageNumber="286">
<emphasis id="A3B136187846FF88FBC7FE84B41C7266" box="[1037,1056,309,330]" italics="true" pageId="13" pageNumber="286">pt</emphasis>
</td>
<td id="6C2481347846007AFB95FE84B4BD7266" box="[1119,1153,309,330]" gridcol="6" gridrow="1" pageId="13" pageNumber="286">µm</td>
<td id="6C2481347846007AFB7BFE84B4EB7266" box="[1201,1239,309,330]" gridcol="7" gridrow="1" pageId="13" pageNumber="286">
<emphasis id="A3B136187846FF88FB70FE84B4F27266" box="[1210,1230,309,330]" italics="true" pageId="13" pageNumber="286">pt</emphasis>
</td>
<td id="6C2481347846007AFAC8FE84B5127266" box="[1282,1326,309,330]" gridcol="8" gridrow="1" pageId="13" pageNumber="286">µm</td>
<td id="6C2481347846007AFA9CFE84B5B97266" box="[1366,1413,309,330]" gridcol="9" gridrow="1" pageId="13" pageNumber="286">
<emphasis id="A3B136187846FF88FAAFFE84B5447266" box="[1381,1400,309,330]" italics="true" pageId="13" pageNumber="286">pt</emphasis>
</td>
</tr>
<tr id="2FF5E8487846007AFF55FEECB5B9725F" box="[159,1413,349,371]" gridrow="2" pageId="13" pageNumber="286">
<th id="6C2481347846007AFF55FEECB1F7725F" box="[159,459,349,371]" gridcol="0" gridrow="2" pageId="13" pageNumber="286">Body length</th>
<td id="6C2481347846007AFDC6FEECB21B725F" box="[524,551,349,371]" gridcol="1" gridrow="2" pageId="13" pageNumber="286">30</td>
<td id="6C2481347846007AFD92FEECB282725F" box="[600,702,349,371]" gridcol="2" gridrow="2" pageId="13" pageNumber="286">245–591</td>
<td id="6C2481347846007AFD27FEECB36F725F" box="[749,851,349,371]" gridcol="3" gridrow="2" pageId="13" pageNumber="286">
<emphasis id="A3B136187846FF88FD27FEECB36E725F" box="[749,850,349,371]" italics="true" pageId="13" pageNumber="286">881–1313</emphasis>
</td>
<td id="6C2481347846007AFC5DFEECB3FF725F" box="[919,963,349,371]" gridcol="4" gridrow="2" pageId="13" pageNumber="286">450</td>
<td id="6C2481347846007AFC34FEECB411725F" box="[1022,1069,349,371]" gridcol="5" gridrow="2" pageId="13" pageNumber="286">
<emphasis id="A3B136187846FF88FC34FEECB411725F" box="[1022,1069,349,371]" italics="true" pageId="13" pageNumber="286">1102</emphasis>
</td>
<td id="6C2481347846007AFB95FEECB4BD725F" box="[1119,1153,349,371]" gridcol="6" gridrow="2" pageId="13" pageNumber="286">83</td>
<td id="6C2481347846007AFB7BFEECB4EB725F" box="[1201,1239,349,371]" gridcol="7" gridrow="2" pageId="13" pageNumber="286">
<emphasis id="A3B136187846FF88FB7BFEECB4EB725F" box="[1201,1239,349,371]" italics="true" pageId="13" pageNumber="286">106</emphasis>
</td>
<td id="6C2481347846007AFAC8FEECB512725F" box="[1282,1326,349,371]" gridcol="8" gridrow="2" pageId="13" pageNumber="286">526</td>
<td id="6C2481347846007AFA9CFEECB5B9725F" box="[1366,1413,349,371]" gridcol="9" gridrow="2" pageId="13" pageNumber="286">
<emphasis id="A3B136187846FF88FA9CFEECB5B9725F" box="[1366,1413,349,371]" italics="true" pageId="13" pageNumber="286">1174</emphasis>
</td>
</tr>
<tr id="2FF5E8487846007AFF55FE37B5B972B0" box="[159,1413,390,412]" gridrow="3" pageId="13" pageNumber="286">
<th id="6C2481347846007AFF55FE37B5B972B0" box="[159,1413,390,412]" colspan="10" colspanRight="9" gridcol="0" gridrow="3" pageId="13" pageNumber="286">Buccal apparatus</th>
</tr>
<tr id="2FF5E8487846007AFF55FE01B5B972EA" box="[159,1413,432,454]" gridrow="4" pageId="13" pageNumber="286" rowspan-3="1">
<th id="6C2481347846007AFF55FE01B1F772EA" box="[159,459,432,454]" gridcol="0" gridrow="4" pageId="13" pageNumber="286">Buccal tube length</th>
<td id="6C2481347846007AFDC6FE01B21B72EA" box="[524,551,432,454]" gridcol="1" gridrow="4" pageId="13" pageNumber="286">30</td>
<td id="6C2481347846007AFD92FE01B28272EA" box="[600,702,432,454]" gridcol="2" gridrow="4" pageId="13" pageNumber="286">27.8–50.1</td>
<td id="6C2481347846007AFC5DFE01B3FF72EA" box="[919,963,432,454]" gridcol="4" gridrow="4" pageId="13" pageNumber="286">40.6</td>
<td id="6C2481347846007AFC34FE01B41172EA" box="[1022,1069,432,454]" gridcol="5" gridrow="4" pageId="13" pageNumber="286">
<emphasis id="A3B136187846FF88FBDBFE00B42272EA" box="[1041,1054,433,454]" italics="true" pageId="13" pageNumber="286">–</emphasis>
</td>
<td id="6C2481347846007AFB95FE01B4BD72EA" box="[1119,1153,432,454]" gridcol="6" gridrow="4" pageId="13" pageNumber="286">5.9</td>
<td id="6C2481347846007AFB7BFE01B4EB72EA" box="[1201,1239,432,454]" gridcol="7" gridrow="4" pageId="13" pageNumber="286">
<emphasis id="A3B136187846FF88FB74FE00B4F772EA" box="[1214,1227,433,454]" italics="true" pageId="13" pageNumber="286">–</emphasis>
</td>
<td id="6C2481347846007AFAC8FE01B51272EA" box="[1282,1326,432,454]" gridcol="8" gridrow="4" pageId="13" pageNumber="286">44.8</td>
<td id="6C2481347846007AFA9CFE01B5B972EA" box="[1366,1413,432,454]" gridcol="9" gridrow="4" pageId="13" pageNumber="286">
<emphasis id="A3B136187846FF88FAA2FE00B54972EA" box="[1384,1397,433,454]" italics="true" pageId="13" pageNumber="286">–</emphasis>
</td>
</tr>
<tr id="2FF5E8487846007AFF55FE68B5B972C3" box="[159,1413,473,495]" gridrow="5" pageId="13" pageNumber="286">
<th id="6C2481347846007AFF55FE68B1F772C3" box="[159,459,473,495]" gridcol="0" gridrow="5" pageId="13" pageNumber="286">Stylet support insertion point</th>
<td id="6C2481347846007AFDC6FE68B21B72C3" box="[524,551,473,495]" gridcol="1" gridrow="5" pageId="13" pageNumber="286">30</td>
<td id="6C2481347846007AFD92FE68B28272C3" box="[600,702,473,495]" gridcol="2" gridrow="5" pageId="13" pageNumber="286">20.5–37.5</td>
<td id="6C2481347846007AFD27FE68B36F72C3" box="[749,851,473,495]" gridcol="3" gridrow="5" pageId="13" pageNumber="286">
<emphasis id="A3B136187846FF88FD27FE68B36E72C3" box="[749,850,473,495]" italics="true" pageId="13" pageNumber="286">72.4–76.4</emphasis>
</td>
<td id="6C2481347846007AFC5DFE68B3FF72C3" box="[919,963,473,495]" gridcol="4" gridrow="5" pageId="13" pageNumber="286">30.6</td>
<td id="6C2481347846007AFC34FE68B41172C3" box="[1022,1069,473,495]" gridcol="5" gridrow="5" pageId="13" pageNumber="286">
<emphasis id="A3B136187846FF88FBCBFE68B41172C3" box="[1025,1069,473,495]" italics="true" pageId="13" pageNumber="286">75.1</emphasis>
</td>
<td id="6C2481347846007AFB95FE68B4BD72C3" box="[1119,1153,473,495]" gridcol="6" gridrow="5" pageId="13" pageNumber="286">4.6</td>
<td id="6C2481347846007AFB7BFE68B4EB72C3" box="[1201,1239,473,495]" gridcol="7" gridrow="5" pageId="13" pageNumber="286">
<emphasis id="A3B136187846FF88FB7EFE68B4EF72C3" box="[1204,1235,473,495]" italics="true" pageId="13" pageNumber="286">0.9</emphasis>
</td>
<td id="6C2481347846007AFAC8FE68B51272C3" box="[1282,1326,473,495]" gridcol="8" gridrow="5" pageId="13" pageNumber="286">34.0</td>
<td id="6C2481347846007AFA9CFE68B5B972C3" box="[1366,1413,473,495]" gridcol="9" gridrow="5" pageId="13" pageNumber="286">
<emphasis id="A3B136187846FF88FA92FE68B5B872C3" box="[1368,1412,473,495]" italics="true" pageId="13" pageNumber="286">75.9</emphasis>
</td>
</tr>
<tr id="2FF5E8487846007AFF55FDB3B5B97134" box="[159,1413,514,536]" gridrow="6" pageId="13" pageNumber="286">
<th id="6C2481347846007AFF55FDB3B1F77134" box="[159,459,514,536]" gridcol="0" gridrow="6" pageId="13" pageNumber="286">Buccal tube external width</th>
<td id="6C2481347846007AFDC6FDB3B21B7134" box="[524,551,514,536]" gridcol="1" gridrow="6" pageId="13" pageNumber="286">30</td>
<td id="6C2481347846007AFD92FDB3B2827134" box="[600,702,514,536]" gridcol="2" gridrow="6" pageId="13" pageNumber="286">3.4–7.2</td>
<td id="6C2481347846007AFD27FDB3B36F7134" box="[749,851,514,536]" gridcol="3" gridrow="6" pageId="13" pageNumber="286">
<emphasis id="A3B136187846FF88FD27FDB3B36E7134" box="[749,850,514,536]" italics="true" pageId="13" pageNumber="286">12.0–14.5</emphasis>
</td>
<td id="6C2481347846007AFC5DFDB3B3FF7134" box="[919,963,514,536]" gridcol="4" gridrow="6" pageId="13" pageNumber="286">5.5</td>
<td id="6C2481347846007AFC34FDB3B4117134" box="[1022,1069,514,536]" gridcol="5" gridrow="6" pageId="13" pageNumber="286">
<emphasis id="A3B136187846FF88FBCBFDB3B4117134" box="[1025,1069,514,536]" italics="true" pageId="13" pageNumber="286">13.4</emphasis>
</td>
<td id="6C2481347846007AFB95FDB3B4BD7134" box="[1119,1153,514,536]" gridcol="6" gridrow="6" pageId="13" pageNumber="286">1.0</td>
<td id="6C2481347846007AFB7BFDB3B4EB7134" box="[1201,1239,514,536]" gridcol="7" gridrow="6" pageId="13" pageNumber="286">
<emphasis id="A3B136187846FF88FB7EFDB3B4EF7134" box="[1204,1235,514,536]" italics="true" pageId="13" pageNumber="286">0.7</emphasis>
</td>
<td id="6C2481347846007AFAC8FDB3B5127134" box="[1282,1326,514,536]" gridcol="8" gridrow="6" pageId="13" pageNumber="286">6.4</td>
<td id="6C2481347846007AFA9CFDB3B5B97134" box="[1366,1413,514,536]" gridcol="9" gridrow="6" pageId="13" pageNumber="286">
<emphasis id="A3B136187846FF88FA92FDB3B5B87134" box="[1368,1412,514,536]" italics="true" pageId="13" pageNumber="286">14.3</emphasis>
</td>
</tr>
<tr id="2FF5E8487846007AFF55FD9DB5B9716E" box="[159,1413,556,578]" gridrow="7" pageId="13" pageNumber="286">
<th id="6C2481347846007AFF55FD9DB1F7716E" box="[159,459,556,578]" gridcol="0" gridrow="7" pageId="13" pageNumber="286">Buccal tube internal width</th>
<td id="6C2481347846007AFDC6FD9DB21B716E" box="[524,551,556,578]" gridcol="1" gridrow="7" pageId="13" pageNumber="286">30</td>
<td id="6C2481347846007AFD92FD9DB282716E" box="[600,702,556,578]" gridcol="2" gridrow="7" pageId="13" pageNumber="286">2.1–5.4</td>
<td id="6C2481347846007AFD27FD9DB36F716E" box="[749,851,556,578]" gridcol="3" gridrow="7" pageId="13" pageNumber="286">
<emphasis id="A3B136187846FF88FD3EFD9DB376716E" box="[756,842,556,578]" italics="true" pageId="13" pageNumber="286">7.6–11.2</emphasis>
</td>
<td id="6C2481347846007AFC5DFD9DB3FF716E" box="[919,963,556,578]" gridcol="4" gridrow="7" pageId="13" pageNumber="286">4.0</td>
<td id="6C2481347846007AFC34FD9DB411716E" box="[1022,1069,556,578]" gridcol="5" gridrow="7" pageId="13" pageNumber="286">
<emphasis id="A3B136187846FF88FBCDFD9DB41B716E" box="[1031,1063,556,578]" italics="true" pageId="13" pageNumber="286">9.6</emphasis>
</td>
<td id="6C2481347846007AFB95FD9DB4BD716E" box="[1119,1153,556,578]" gridcol="6" gridrow="7" pageId="13" pageNumber="286">0.8</td>
<td id="6C2481347846007AFB7BFD9DB4EB716E" box="[1201,1239,556,578]" gridcol="7" gridrow="7" pageId="13" pageNumber="286">
<emphasis id="A3B136187846FF88FB7EFD9DB4EF716E" box="[1204,1235,556,578]" italics="true" pageId="13" pageNumber="286">0.9</emphasis>
</td>
<td id="6C2481347846007AFAC8FD9DB512716E" box="[1282,1326,556,578]" gridcol="8" gridrow="7" pageId="13" pageNumber="286">5.0</td>
<td id="6C2481347846007AFA9CFD9DB5B9716E" box="[1366,1413,556,578]" gridcol="9" gridrow="7" pageId="13" pageNumber="286">
<emphasis id="A3B136187846FF88FA93FD9DB5BE716E" box="[1369,1410,556,578]" italics="true" pageId="13" pageNumber="286">11.2</emphasis>
</td>
</tr>
<tr id="2FF5E8487846007AFF55FDE4B5B97147" box="[159,1413,597,619]" gridrow="8" pageId="13" pageNumber="286">
<th id="6C2481347846007AFF55FDE4B1F77147" box="[159,459,597,619]" gridcol="0" gridrow="8" pageId="13" pageNumber="286">Ventral lamina length</th>
<td id="6C2481347846007AFDC6FDE4B21B7147" box="[524,551,597,619]" gridcol="1" gridrow="8" pageId="13" pageNumber="286">25</td>
<td id="6C2481347846007AFD92FDE4B2827147" box="[600,702,597,619]" gridcol="2" gridrow="8" pageId="13" pageNumber="286">16.3–29.6</td>
<td id="6C2481347846007AFD27FDE4B36F7147" box="[749,851,597,619]" gridcol="3" gridrow="8" pageId="13" pageNumber="286">
<emphasis id="A3B136187846FF88FD27FDE4B36F7147" box="[749,851,597,619]" italics="true" pageId="13" pageNumber="286">56.1–63.7</emphasis>
</td>
<td id="6C2481347846007AFC5DFDE4B3FF7147" box="[919,963,597,619]" gridcol="4" gridrow="8" pageId="13" pageNumber="286">24.7</td>
<td id="6C2481347846007AFC34FDE4B4117147" box="[1022,1069,597,619]" gridcol="5" gridrow="8" pageId="13" pageNumber="286">
<emphasis id="A3B136187846FF88FBCBFDE4B4117147" box="[1025,1069,597,619]" italics="true" pageId="13" pageNumber="286">59.5</emphasis>
</td>
<td id="6C2481347846007AFB95FDE4B4BD7147" box="[1119,1153,597,619]" gridcol="6" gridrow="8" pageId="13" pageNumber="286">3.3</td>
<td id="6C2481347846007AFB7BFDE4B4EB7147" box="[1201,1239,597,619]" gridcol="7" gridrow="8" pageId="13" pageNumber="286">
<emphasis id="A3B136187846FF88FB7EFDE4B4EF7147" box="[1204,1235,597,619]" italics="true" pageId="13" pageNumber="286">1.9</emphasis>
</td>
<td id="6C2481347846007AFAC8FDE4B5127147" box="[1282,1326,597,619]" gridcol="8" gridrow="8" pageId="13" pageNumber="286">26.7</td>
<td id="6C2481347846007AFA9CFDE4B5B97147" box="[1366,1413,597,619]" gridcol="9" gridrow="8" pageId="13" pageNumber="286">
<emphasis id="A3B136187846FF88FA92FDE4B5B87147" box="[1368,1412,597,619]" italics="true" pageId="13" pageNumber="286">59.6</emphasis>
</td>
</tr>
<tr id="2FF5E8487846007AFF55FDCFB5B971B8" box="[159,1413,638,660]" gridrow="9" pageId="13" pageNumber="286">
<th id="6C2481347846007AFF55FDCFB5B971B8" box="[159,1413,638,660]" colspan="10" colspanRight="9" gridcol="0" gridrow="9" pageId="13" pageNumber="286">Placoid lengths</th>
</tr>
<tr id="2FF5E8487846007AFF55FD19B5B97192" box="[159,1413,680,702]" gridrow="10" pageId="13" pageNumber="286">
<th id="6C2481347846007AFF55FD19B1F77192" box="[159,459,680,702]" gridcol="0" gridrow="10" pageId="13" pageNumber="286">Macroplacoid 1</th>
<td id="6C2481347846007AFDC6FD19B21B7192" box="[524,551,680,702]" gridcol="1" gridrow="10" pageId="13" pageNumber="286">30</td>
<td id="6C2481347846007AFD92FD19B2827192" box="[600,702,680,702]" gridcol="2" gridrow="10" pageId="13" pageNumber="286">5.1–13.3</td>
<td id="6C2481347846007AFD27FD19B36F7192" box="[749,851,680,702]" gridcol="3" gridrow="10" pageId="13" pageNumber="286">
<emphasis id="A3B136187846FF88FD27FD19B36F7192" box="[749,851,680,702]" italics="true" pageId="13" pageNumber="286">18.0–27.5</emphasis>
</td>
<td id="6C2481347846007AFC5DFD19B3FF7192" box="[919,963,680,702]" gridcol="4" gridrow="10" pageId="13" pageNumber="286">10.0</td>
<td id="6C2481347846007AFC34FD19B4117192" box="[1022,1069,680,702]" gridcol="5" gridrow="10" pageId="13" pageNumber="286">
<emphasis id="A3B136187846FF88FBCBFD19B4117192" box="[1025,1069,680,702]" italics="true" pageId="13" pageNumber="286">24.5</emphasis>
</td>
<td id="6C2481347846007AFB95FD19B4BD7192" box="[1119,1153,680,702]" gridcol="6" gridrow="10" pageId="13" pageNumber="286">2.0</td>
<td id="6C2481347846007AFB7BFD19B4EB7192" box="[1201,1239,680,702]" gridcol="7" gridrow="10" pageId="13" pageNumber="286">
<emphasis id="A3B136187846FF88FB7EFD19B4EF7192" box="[1204,1235,680,702]" italics="true" pageId="13" pageNumber="286">2.1</emphasis>
</td>
<td id="6C2481347846007AFAC8FD19B5127192" box="[1282,1326,680,702]" gridcol="8" gridrow="10" pageId="13" pageNumber="286">11.5</td>
<td id="6C2481347846007AFA9CFD19B5B97192" box="[1366,1413,680,702]" gridcol="9" gridrow="10" pageId="13" pageNumber="286">
<emphasis id="A3B136187846FF88FA92FD19B5B87192" box="[1368,1412,680,702]" italics="true" pageId="13" pageNumber="286">25.7</emphasis>
</td>
</tr>
<tr id="2FF5E8487846007AFF55FD60B5B971CB" box="[159,1413,721,743]" gridrow="11" pageId="13" pageNumber="286">
<th id="6C2481347846007AFF55FD60B1F771CB" box="[159,459,721,743]" gridcol="0" gridrow="11" pageId="13" pageNumber="286">Macroplacoid 2</th>
<td id="6C2481347846007AFDC6FD60B21B71CB" box="[524,551,721,743]" gridcol="1" gridrow="11" pageId="13" pageNumber="286">30</td>
<td id="6C2481347846007AFD92FD60B28271CB" box="[600,702,721,743]" gridcol="2" gridrow="11" pageId="13" pageNumber="286">2.9–7.9</td>
<td id="6C2481347846007AFD27FD60B36F71CB" box="[749,851,721,743]" gridcol="3" gridrow="11" pageId="13" pageNumber="286">
<emphasis id="A3B136187846FF88FD27FD60B36E71CB" box="[749,850,721,743]" italics="true" pageId="13" pageNumber="286">10.3–16.2</emphasis>
</td>
<td id="6C2481347846007AFC5DFD60B3FF71CB" box="[919,963,721,743]" gridcol="4" gridrow="11" pageId="13" pageNumber="286">5.9</td>
<td id="6C2481347846007AFC34FD60B41171CB" box="[1022,1069,721,743]" gridcol="5" gridrow="11" pageId="13" pageNumber="286">
<emphasis id="A3B136187846FF88FBCBFD60B41171CB" box="[1025,1069,721,743]" italics="true" pageId="13" pageNumber="286">14.4</emphasis>
</td>
<td id="6C2481347846007AFB95FD60B4BD71CB" box="[1119,1153,721,743]" gridcol="6" gridrow="11" pageId="13" pageNumber="286">1.3</td>
<td id="6C2481347846007AFB7BFD60B4EB71CB" box="[1201,1239,721,743]" gridcol="7" gridrow="11" pageId="13" pageNumber="286">
<emphasis id="A3B136187846FF88FB7EFD60B4EF71CB" box="[1204,1235,721,743]" italics="true" pageId="13" pageNumber="286">1.5</emphasis>
</td>
<td id="6C2481347846007AFAC8FD60B51271CB" box="[1282,1326,721,743]" gridcol="8" gridrow="11" pageId="13" pageNumber="286">6.4</td>
<td id="6C2481347846007AFA9CFD60B5B971CB" box="[1366,1413,721,743]" gridcol="9" gridrow="11" pageId="13" pageNumber="286">
<emphasis id="A3B136187846FF88FA92FD60B5B871CB" box="[1368,1412,721,743]" italics="true" pageId="13" pageNumber="286">14.3</emphasis>
</td>
</tr>
<tr id="2FF5E8487846007AFF55FD4BB5B9703C" box="[159,1413,762,784]" gridrow="12" pageId="13" pageNumber="286">
<th id="6C2481347846007AFF55FD4BB1F7703C" box="[159,459,762,784]" gridcol="0" gridrow="12" pageId="13" pageNumber="286">Microplacoid</th>
<td id="6C2481347846007AFDC6FD4BB21B703C" box="[524,551,762,784]" gridcol="1" gridrow="12" pageId="13" pageNumber="286">30</td>
<td id="6C2481347846007AFD92FD4BB282703C" box="[600,702,762,784]" gridcol="2" gridrow="12" pageId="13" pageNumber="286">1.7–4.8</td>
<td id="6C2481347846007AFD27FD4BB36F703C" box="[749,851,762,784]" gridcol="3" gridrow="12" pageId="13" pageNumber="286">
<emphasis id="A3B136187846FF88FD39FD4BB370703C" box="[755,844,762,784]" italics="true" pageId="13" pageNumber="286">6.0–10.5</emphasis>
</td>
<td id="6C2481347846007AFC5DFD4BB3FF703C" box="[919,963,762,784]" gridcol="4" gridrow="12" pageId="13" pageNumber="286">3.2</td>
<td id="6C2481347846007AFC34FD4BB411703C" box="[1022,1069,762,784]" gridcol="5" gridrow="12" pageId="13" pageNumber="286">
<emphasis id="A3B136187846FF88FBCDFD4AB41B703C" box="[1031,1063,763,784]" italics="true" pageId="13" pageNumber="286">7.7</emphasis>
</td>
<td id="6C2481347846007AFB95FD4BB4BD703C" box="[1119,1153,762,784]" gridcol="6" gridrow="12" pageId="13" pageNumber="286">0.7</td>
<td id="6C2481347846007AFB7BFD4BB4EB703C" box="[1201,1239,762,784]" gridcol="7" gridrow="12" pageId="13" pageNumber="286">
<emphasis id="A3B136187846FF88FB7EFD4BB4EF703C" box="[1204,1235,762,784]" italics="true" pageId="13" pageNumber="286">1.0</emphasis>
</td>
<td id="6C2481347846007AFAC8FD4BB512703C" box="[1282,1326,762,784]" gridcol="8" gridrow="12" pageId="13" pageNumber="286">3.8</td>
<td id="6C2481347846007AFA9CFD4BB5B9703C" box="[1366,1413,762,784]" gridcol="9" gridrow="12" pageId="13" pageNumber="286">
<emphasis id="A3B136187846FF88FA95FD4BB543703C" box="[1375,1407,762,784]" italics="true" pageId="13" pageNumber="286">8.5</emphasis>
</td>
</tr>
<tr id="2FF5E8487846007AFF55FC95B5B97016" box="[159,1413,804,826]" gridrow="13" pageId="13" pageNumber="286">
<th id="6C2481347846007AFF55FC95B1F77016" box="[159,459,804,826]" gridcol="0" gridrow="13" pageId="13" pageNumber="286">Macroplacoid row</th>
<td id="6C2481347846007AFDC6FC95B21B7016" box="[524,551,804,826]" gridcol="1" gridrow="13" pageId="13" pageNumber="286">30</td>
<td id="6C2481347846007AFD92FC95B2827016" box="[600,702,804,826]" gridcol="2" gridrow="13" pageId="13" pageNumber="286">9.3–22.3</td>
<td id="6C2481347846007AFD27FC95B36F7016" box="[749,851,804,826]" gridcol="3" gridrow="13" pageId="13" pageNumber="286">
<emphasis id="A3B136187846FF88FD27FC95B36E7016" box="[749,850,804,826]" italics="true" pageId="13" pageNumber="286">32.9–52.9</emphasis>
</td>
<td id="6C2481347846007AFC5DFC95B3FF7016" box="[919,963,804,826]" gridcol="4" gridrow="13" pageId="13" pageNumber="286">17.4</td>
<td id="6C2481347846007AFC34FC95B4117016" box="[1022,1069,804,826]" gridcol="5" gridrow="13" pageId="13" pageNumber="286">
<emphasis id="A3B136187846FF88FBCBFC95B4117016" box="[1025,1069,804,826]" italics="true" pageId="13" pageNumber="286">42.6</emphasis>
</td>
<td id="6C2481347846007AFB95FC95B4BD7016" box="[1119,1153,804,826]" gridcol="6" gridrow="13" pageId="13" pageNumber="286">3.4</td>
<td id="6C2481347846007AFB7BFC95B4EB7016" box="[1201,1239,804,826]" gridcol="7" gridrow="13" pageId="13" pageNumber="286">
<emphasis id="A3B136187846FF88FB7EFC95B4EF7016" box="[1204,1235,804,826]" italics="true" pageId="13" pageNumber="286">3.9</emphasis>
</td>
<td id="6C2481347846007AFAC8FC95B5127016" box="[1282,1326,804,826]" gridcol="8" gridrow="13" pageId="13" pageNumber="286">19.3</td>
<td id="6C2481347846007AFA9CFC95B5B97016" box="[1366,1413,804,826]" gridcol="9" gridrow="13" pageId="13" pageNumber="286">
<emphasis id="A3B136187846FF88FA92FC95B5B87016" box="[1368,1412,804,826]" italics="true" pageId="13" pageNumber="286">43.1</emphasis>
</td>
</tr>
<tr id="2FF5E8487846007AFF55FCFCB5B9704F" box="[159,1413,845,867]" gridrow="14" pageId="13" pageNumber="286">
<th id="6C2481347846007AFF55FCFCB1F7704F" box="[159,459,845,867]" gridcol="0" gridrow="14" pageId="13" pageNumber="286">Placoid row</th>
<td id="6C2481347846007AFDC6FCFCB21B704F" box="[524,551,845,867]" gridcol="1" gridrow="14" pageId="13" pageNumber="286">30</td>
<td id="6C2481347846007AFD92FCFCB282704F" box="[600,702,845,867]" gridcol="2" gridrow="14" pageId="13" pageNumber="286">11.2–26.3</td>
<td id="6C2481347846007AFD27FCFCB36F704F" box="[749,851,845,867]" gridcol="3" gridrow="14" pageId="13" pageNumber="286">
<emphasis id="A3B136187846FF88FD27FCFCB36E704F" box="[749,850,845,867]" italics="true" pageId="13" pageNumber="286">40.3–58.7</emphasis>
</td>
<td id="6C2481347846007AFC5DFCFCB3FF704F" box="[919,963,845,867]" gridcol="4" gridrow="14" pageId="13" pageNumber="286">21.0</td>
<td id="6C2481347846007AFC34FCFCB411704F" box="[1022,1069,845,867]" gridcol="5" gridrow="14" pageId="13" pageNumber="286">
<emphasis id="A3B136187846FF88FBCBFCFCB411704F" box="[1025,1069,845,867]" italics="true" pageId="13" pageNumber="286">51.4</emphasis>
</td>
<td id="6C2481347846007AFB95FCFCB4BD704F" box="[1119,1153,845,867]" gridcol="6" gridrow="14" pageId="13" pageNumber="286">4.0</td>
<td id="6C2481347846007AFB7BFCFCB4EB704F" box="[1201,1239,845,867]" gridcol="7" gridrow="14" pageId="13" pageNumber="286">
<emphasis id="A3B136187846FF88FB7EFCFCB4EF704F" box="[1204,1235,845,867]" italics="true" pageId="13" pageNumber="286">4.1</emphasis>
</td>
<td id="6C2481347846007AFAC8FCFCB512704F" box="[1282,1326,845,867]" gridcol="8" gridrow="14" pageId="13" pageNumber="286">23.8</td>
<td id="6C2481347846007AFA9CFCFCB5B9704F" box="[1366,1413,845,867]" gridcol="9" gridrow="14" pageId="13" pageNumber="286">
<emphasis id="A3B136187846FF88FA92FCFCB5B8704F" box="[1368,1412,845,867]" italics="true" pageId="13" pageNumber="286">53.1</emphasis>
</td>
</tr>
<tr id="2FF5E8487846007AFF55FCC7B5B970A0" box="[159,1413,886,908]" gridrow="15" pageId="13" pageNumber="286">
<th id="6C2481347846007AFF55FCC7B5B970A0" box="[159,1413,886,908]" colspan="10" colspanRight="9" gridcol="0" gridrow="15" pageId="13" pageNumber="286">Claw 1 lengths</th>
</tr>
<tr id="2FF5E8487846007AFF55FC11B5B9709A" box="[159,1413,928,950]" gridrow="16" pageId="13" pageNumber="286">
<th id="6C2481347846007AFF55FC11B1F7709A" box="[159,459,928,950]" gridcol="0" gridrow="16" pageId="13" pageNumber="286">External primary branch</th>
<td id="6C2481347846007AFDC6FC11B21B709A" box="[524,551,928,950]" gridcol="1" gridrow="16" pageId="13" pageNumber="286">29</td>
<td id="6C2481347846007AFD92FC11B282709A" box="[600,702,928,950]" gridcol="2" gridrow="16" pageId="13" pageNumber="286">7.5–16.2</td>
<td id="6C2481347846007AFD27FC11B36F709A" box="[749,851,928,950]" gridcol="3" gridrow="16" pageId="13" pageNumber="286">
<emphasis id="A3B136187846FF88FD27FC11B36E709A" box="[749,850,928,950]" italics="true" pageId="13" pageNumber="286">27.0–36.5</emphasis>
</td>
<td id="6C2481347846007AFC5DFC11B3FF709A" box="[919,963,928,950]" gridcol="4" gridrow="16" pageId="13" pageNumber="286">13.1</td>
<td id="6C2481347846007AFC34FC11B411709A" box="[1022,1069,928,950]" gridcol="5" gridrow="16" pageId="13" pageNumber="286">
<emphasis id="A3B136187846FF88FBCBFC11B411709A" box="[1025,1069,928,950]" italics="true" pageId="13" pageNumber="286">32.3</emphasis>
</td>
<td id="6C2481347846007AFB95FC11B4BD709A" box="[1119,1153,928,950]" gridcol="6" gridrow="16" pageId="13" pageNumber="286">2.2</td>
<td id="6C2481347846007AFB7BFC11B4EB709A" box="[1201,1239,928,950]" gridcol="7" gridrow="16" pageId="13" pageNumber="286">
<emphasis id="A3B136187846FF88FB7EFC11B4EF709A" box="[1204,1235,928,950]" italics="true" pageId="13" pageNumber="286">2.3</emphasis>
</td>
<td id="6C2481347846007AFAC8FC11B512709A" box="[1282,1326,928,950]" gridcol="8" gridrow="16" pageId="13" pageNumber="286">15.0</td>
<td id="6C2481347846007AFA9CFC11B5B9709A" box="[1366,1413,928,950]" gridcol="9" gridrow="16" pageId="13" pageNumber="286">
<emphasis id="A3B136187846FF88FA92FC11B5B8709A" box="[1368,1412,928,950]" italics="true" pageId="13" pageNumber="286">33.5</emphasis>
</td>
</tr>
<tr id="2FF5E8487846007AFF55FC78B5B970F3" box="[159,1413,969,991]" gridrow="17" pageId="13" pageNumber="286">
<th id="6C2481347846007AFF55FC78B1F770F3" box="[159,459,969,991]" gridcol="0" gridrow="17" pageId="13" pageNumber="286">External secondary branch</th>
<td id="6C2481347846007AFDC6FC78B21B70F3" box="[524,551,969,991]" gridcol="1" gridrow="17" pageId="13" pageNumber="286">22</td>
<td id="6C2481347846007AFD92FC78B28270F3" box="[600,702,969,991]" gridcol="2" gridrow="17" pageId="13" pageNumber="286">4.8–13.0</td>
<td id="6C2481347846007AFD27FC78B36F70F3" box="[749,851,969,991]" gridcol="3" gridrow="17" pageId="13" pageNumber="286">
<emphasis id="A3B136187846FF88FD27FC78B36F70F3" box="[749,851,969,991]" italics="true" pageId="13" pageNumber="286">17.3–29.1</emphasis>
</td>
<td id="6C2481347846007AFC5DFC78B3FF70F3" box="[919,963,969,991]" gridcol="4" gridrow="17" pageId="13" pageNumber="286">10.3</td>
<td id="6C2481347846007AFC34FC78B41170F3" box="[1022,1069,969,991]" gridcol="5" gridrow="17" pageId="13" pageNumber="286">
<emphasis id="A3B136187846FF88FBCBFC78B41170F3" box="[1025,1069,969,991]" italics="true" pageId="13" pageNumber="286">24.8</emphasis>
</td>
<td id="6C2481347846007AFB95FC78B4BD70F3" box="[1119,1153,969,991]" gridcol="6" gridrow="17" pageId="13" pageNumber="286">2.1</td>
<td id="6C2481347846007AFB7BFC78B4EB70F3" box="[1201,1239,969,991]" gridcol="7" gridrow="17" pageId="13" pageNumber="286">
<emphasis id="A3B136187846FF88FB7EFC78B4EF70F3" box="[1204,1235,969,991]" italics="true" pageId="13" pageNumber="286">3.1</emphasis>
</td>
<td id="6C2481347846007AFAC8FC78B51270F3" box="[1282,1326,969,991]" gridcol="8" gridrow="17" pageId="13" pageNumber="286">12.1</td>
<td id="6C2481347846007AFA9CFC78B5B970F3" box="[1366,1413,969,991]" gridcol="9" gridrow="17" pageId="13" pageNumber="286">
<emphasis id="A3B136187846FF88FA92FC78B5B870F3" box="[1368,1412,969,991]" italics="true" pageId="13" pageNumber="286">27.0</emphasis>
</td>
</tr>
<tr id="2FF5E8487846007AFF55FC43B5B97724" box="[159,1413,1010,1032]" gridrow="18" pageId="13" pageNumber="286">
<th id="6C2481347846007AFF55FC43B1F77724" box="[159,459,1010,1032]" gridcol="0" gridrow="18" pageId="13" pageNumber="286">Internal primary branch</th>
<td id="6C2481347846007AFDC6FC43B21B7724" box="[524,551,1010,1032]" gridcol="1" gridrow="18" pageId="13" pageNumber="286">30</td>
<td id="6C2481347846007AFD92FC43B2827724" box="[600,702,1010,1032]" gridcol="2" gridrow="18" pageId="13" pageNumber="286">7.3–15.8</td>
<td id="6C2481347846007AFD27FC43B36F7724" box="[749,851,1010,1032]" gridcol="3" gridrow="18" pageId="13" pageNumber="286">
<emphasis id="A3B136187846FF88FD27FC43B36E7724" box="[749,850,1010,1032]" italics="true" pageId="13" pageNumber="286">26.3–34.1</emphasis>
</td>
<td id="6C2481347846007AFC5DFC43B3FF7724" box="[919,963,1010,1032]" gridcol="4" gridrow="18" pageId="13" pageNumber="286">12.3</td>
<td id="6C2481347846007AFC34FC43B4117724" box="[1022,1069,1010,1032]" gridcol="5" gridrow="18" pageId="13" pageNumber="286">
<emphasis id="A3B136187846FF88FBCBFC43B4117724" box="[1025,1069,1010,1032]" italics="true" pageId="13" pageNumber="286">30.3</emphasis>
</td>
<td id="6C2481347846007AFB95FC43B4BD7724" box="[1119,1153,1010,1032]" gridcol="6" gridrow="18" pageId="13" pageNumber="286">2.1</td>
<td id="6C2481347846007AFB7BFC43B4EB7724" box="[1201,1239,1010,1032]" gridcol="7" gridrow="18" pageId="13" pageNumber="286">
<emphasis id="A3B136187846FF88FB7EFC43B4EF7724" box="[1204,1235,1010,1032]" italics="true" pageId="13" pageNumber="286">2.1</emphasis>
</td>
<td id="6C2481347846007AFAC8FC43B5127724" box="[1282,1326,1010,1032]" gridcol="8" gridrow="18" pageId="13" pageNumber="286">13.7</td>
<td id="6C2481347846007AFA9CFC43B5B97724" box="[1366,1413,1010,1032]" gridcol="9" gridrow="18" pageId="13" pageNumber="286">
<emphasis id="A3B136187846FF88FA92FC43B5B87724" box="[1368,1412,1010,1032]" italics="true" pageId="13" pageNumber="286">30.6</emphasis>
</td>
</tr>
<tr id="2FF5E8487846007AFF55FBADB5B9771E" box="[159,1413,1052,1074]" gridrow="19" pageId="13" pageNumber="286">
<th id="6C2481347846007AFF55FBADB1F7771E" box="[159,459,1052,1074]" gridcol="0" gridrow="19" pageId="13" pageNumber="286">Internal secondary branch</th>
<td id="6C2481347846007AFDC6FBADB21B771E" box="[524,551,1052,1074]" gridcol="1" gridrow="19" pageId="13" pageNumber="286">24</td>
<td id="6C2481347846007AFD92FBADB282771E" box="[600,702,1052,1074]" gridcol="2" gridrow="19" pageId="13" pageNumber="286">4.8–12.7</td>
<td id="6C2481347846007AFD27FBADB36F771E" box="[749,851,1052,1074]" gridcol="3" gridrow="19" pageId="13" pageNumber="286">
<emphasis id="A3B136187846FF88FD27FBADB36F771E" box="[749,851,1052,1074]" italics="true" pageId="13" pageNumber="286">17.3–27.1</emphasis>
</td>
<td id="6C2481347846007AFC5DFBADB3FF771E" box="[919,963,1052,1074]" gridcol="4" gridrow="19" pageId="13" pageNumber="286">9.4</td>
<td id="6C2481347846007AFC34FBADB411771E" box="[1022,1069,1052,1074]" gridcol="5" gridrow="19" pageId="13" pageNumber="286">
<emphasis id="A3B136187846FF88FBCBFBADB411771E" box="[1025,1069,1052,1074]" italics="true" pageId="13" pageNumber="286">22.7</emphasis>
</td>
<td id="6C2481347846007AFB95FBADB4BD771E" box="[1119,1153,1052,1074]" gridcol="6" gridrow="19" pageId="13" pageNumber="286">2.1</td>
<td id="6C2481347846007AFB7BFBADB4EB771E" box="[1201,1239,1052,1074]" gridcol="7" gridrow="19" pageId="13" pageNumber="286">
<emphasis id="A3B136187846FF88FB7EFBADB4EF771E" box="[1204,1235,1052,1074]" italics="true" pageId="13" pageNumber="286">2.6</emphasis>
</td>
<td id="6C2481347846007AFAC8FBADB512771E" box="[1282,1326,1052,1074]" gridcol="8" gridrow="19" pageId="13" pageNumber="286">10.6</td>
<td id="6C2481347846007AFA9CFBADB5B9771E" box="[1366,1413,1052,1074]" gridcol="9" gridrow="19" pageId="13" pageNumber="286">
<emphasis id="A3B136187846FF88FA92FBADB5B8771E" box="[1368,1412,1052,1074]" italics="true" pageId="13" pageNumber="286">23.7</emphasis>
</td>
</tr>
<tr id="2FF5E8487846007AFF55FBF4B5B97777" box="[159,1413,1093,1115]" gridrow="20" pageId="13" pageNumber="286">
<th id="6C2481347846007AFF55FBF4B5B97777" box="[159,1413,1093,1115]" colspan="10" colspanRight="9" gridcol="0" gridrow="20" pageId="13" pageNumber="286">Claw 2 lengths</th>
</tr>
<tr id="2FF5E8487846007AFF55FBDFB5B977A8" box="[159,1413,1134,1156]" gridrow="21" pageId="13" pageNumber="286">
<th id="6C2481347846007AFF55FBDFB1F777A8" box="[159,459,1134,1156]" gridcol="0" gridrow="21" pageId="13" pageNumber="286">External primary branch</th>
<td id="6C2481347846007AFDC6FBDFB21B77A8" box="[524,551,1134,1156]" gridcol="1" gridrow="21" pageId="13" pageNumber="286">27</td>
<td id="6C2481347846007AFD92FBDFB28277A8" box="[600,702,1134,1156]" gridcol="2" gridrow="21" pageId="13" pageNumber="286">8.6–17.2</td>
<td id="6C2481347846007AFD27FBDFB36F77A8" box="[749,851,1134,1156]" gridcol="3" gridrow="21" pageId="13" pageNumber="286">
<emphasis id="A3B136187846FF88FD27FBDFB36E77A8" box="[749,850,1134,1156]" italics="true" pageId="13" pageNumber="286">30.4–38.9</emphasis>
</td>
<td id="6C2481347846007AFC5DFBDFB3FF77A8" box="[919,963,1134,1156]" gridcol="4" gridrow="21" pageId="13" pageNumber="286">14.3</td>
<td id="6C2481347846007AFC34FBDFB41177A8" box="[1022,1069,1134,1156]" gridcol="5" gridrow="21" pageId="13" pageNumber="286">
<emphasis id="A3B136187846FF88FBCBFBDFB41177A8" box="[1025,1069,1134,1156]" italics="true" pageId="13" pageNumber="286">34.9</emphasis>
</td>
<td id="6C2481347846007AFB95FBDFB4BD77A8" box="[1119,1153,1134,1156]" gridcol="6" gridrow="21" pageId="13" pageNumber="286">2.3</td>
<td id="6C2481347846007AFB7BFBDFB4EB77A8" box="[1201,1239,1134,1156]" gridcol="7" gridrow="21" pageId="13" pageNumber="286">
<emphasis id="A3B136187846FF88FB7EFBDFB4EF77A8" box="[1204,1235,1134,1156]" italics="true" pageId="13" pageNumber="286">2.3</emphasis>
</td>
<td id="6C2481347846007AFAC8FBDFB51277A8" box="[1282,1326,1134,1156]" gridcol="8" gridrow="21" pageId="13" pageNumber="286">15.9</td>
<td id="6C2481347846007AFA9CFBDFB5B977A8" box="[1366,1413,1134,1156]" gridcol="9" gridrow="21" pageId="13" pageNumber="286">
<emphasis id="A3B136187846FF88FA92FBDFB5B877A8" box="[1368,1412,1134,1156]" italics="true" pageId="13" pageNumber="286">35.5</emphasis>
</td>
</tr>
<tr id="2FF5E8487846007AFF55FB29B5B97782" box="[159,1413,1176,1198]" gridrow="22" pageId="13" pageNumber="286">
<th id="6C2481347846007AFF55FB29B1F77782" box="[159,459,1176,1198]" gridcol="0" gridrow="22" pageId="13" pageNumber="286">External secondary branch</th>
<td id="6C2481347846007AFDC6FB29B21B7782" box="[524,551,1176,1198]" gridcol="1" gridrow="22" pageId="13" pageNumber="286">23</td>
<td id="6C2481347846007AFD92FB29B2827782" box="[600,702,1176,1198]" gridcol="2" gridrow="22" pageId="13" pageNumber="286">5.7–13.4</td>
<td id="6C2481347846007AFD27FB29B36F7782" box="[749,851,1176,1198]" gridcol="3" gridrow="22" pageId="13" pageNumber="286">
<emphasis id="A3B136187846FF88FD27FB29B36E7782" box="[749,850,1176,1198]" italics="true" pageId="13" pageNumber="286">20.3–30.7</emphasis>
</td>
<td id="6C2481347846007AFC5DFB29B3FF7782" box="[919,963,1176,1198]" gridcol="4" gridrow="22" pageId="13" pageNumber="286">10.6</td>
<td id="6C2481347846007AFC34FB29B4117782" box="[1022,1069,1176,1198]" gridcol="5" gridrow="22" pageId="13" pageNumber="286">
<emphasis id="A3B136187846FF88FBCBFB29B4117782" box="[1025,1069,1176,1198]" italics="true" pageId="13" pageNumber="286">26.0</emphasis>
</td>
<td id="6C2481347846007AFB95FB29B4BD7782" box="[1119,1153,1176,1198]" gridcol="6" gridrow="22" pageId="13" pageNumber="286">2.1</td>
<td id="6C2481347846007AFB7BFB29B4EB7782" box="[1201,1239,1176,1198]" gridcol="7" gridrow="22" pageId="13" pageNumber="286">
<emphasis id="A3B136187846FF88FB7EFB29B4EF7782" box="[1204,1235,1176,1198]" italics="true" pageId="13" pageNumber="286">2.9</emphasis>
</td>
<td id="6C2481347846007AFAC8FB29B5127782" box="[1282,1326,1176,1198]" gridcol="8" gridrow="22" pageId="13" pageNumber="286">13.0</td>
<td id="6C2481347846007AFA9CFB29B5B97782" box="[1366,1413,1176,1198]" gridcol="9" gridrow="22" pageId="13" pageNumber="286">
<emphasis id="A3B136187846FF88FA92FB29B5B87782" box="[1368,1412,1176,1198]" italics="true" pageId="13" pageNumber="286">29.0</emphasis>
</td>
</tr>
<tr id="2FF5E8487846007AFF55FB70B5B977FB" box="[159,1413,1217,1239]" gridrow="23" pageId="13" pageNumber="286">
<th id="6C2481347846007AFF55FB70B1F777FB" box="[159,459,1217,1239]" gridcol="0" gridrow="23" pageId="13" pageNumber="286">Internal primary branch</th>
<td id="6C2481347846007AFDC6FB70B21B77FB" box="[524,551,1217,1239]" gridcol="1" gridrow="23" pageId="13" pageNumber="286">29</td>
<td id="6C2481347846007AFD92FB70B28277FB" box="[600,702,1217,1239]" gridcol="2" gridrow="23" pageId="13" pageNumber="286">8.1–14.9</td>
<td id="6C2481347846007AFD27FB70B36F77FB" box="[749,851,1217,1239]" gridcol="3" gridrow="23" pageId="13" pageNumber="286">
<emphasis id="A3B136187846FF88FD27FB70B36E77FB" box="[749,850,1217,1239]" italics="true" pageId="13" pageNumber="286">28.6–36.2</emphasis>
</td>
<td id="6C2481347846007AFC5DFB70B3FF77FB" box="[919,963,1217,1239]" gridcol="4" gridrow="23" pageId="13" pageNumber="286">13.0</td>
<td id="6C2481347846007AFC34FB70B41177FB" box="[1022,1069,1217,1239]" gridcol="5" gridrow="23" pageId="13" pageNumber="286">
<emphasis id="A3B136187846FF88FBCBFB70B41177FB" box="[1025,1069,1217,1239]" italics="true" pageId="13" pageNumber="286">31.9</emphasis>
</td>
<td id="6C2481347846007AFB95FB70B4BD77FB" box="[1119,1153,1217,1239]" gridcol="6" gridrow="23" pageId="13" pageNumber="286">2.0</td>
<td id="6C2481347846007AFB7BFB70B4EB77FB" box="[1201,1239,1217,1239]" gridcol="7" gridrow="23" pageId="13" pageNumber="286">
<emphasis id="A3B136187846FF88FB7EFB70B4EF77FB" box="[1204,1235,1217,1239]" italics="true" pageId="13" pageNumber="286">2.1</emphasis>
</td>
<td id="6C2481347846007AFAC8FB70B51277FB" box="[1282,1326,1217,1239]" gridcol="8" gridrow="23" pageId="13" pageNumber="286">14.5</td>
<td id="6C2481347846007AFA9CFB70B5B977FB" box="[1366,1413,1217,1239]" gridcol="9" gridrow="23" pageId="13" pageNumber="286">
<emphasis id="A3B136187846FF88FA92FB70B5B877FB" box="[1368,1412,1217,1239]" italics="true" pageId="13" pageNumber="286">32.4</emphasis>
</td>
</tr>
<tr id="2FF5E8487846007AFF55FB5BB5B9762C" box="[159,1413,1258,1280]" gridrow="24" pageId="13" pageNumber="286">
<th id="6C2481347846007AFF55FB5BB1F7762C" box="[159,459,1258,1280]" gridcol="0" gridrow="24" pageId="13" pageNumber="286">Internal secondary branch</th>
<td id="6C2481347846007AFDC6FB5BB21B762C" box="[524,551,1258,1280]" gridcol="1" gridrow="24" pageId="13" pageNumber="286">23</td>
<td id="6C2481347846007AFD92FB5BB282762C" box="[600,702,1258,1280]" gridcol="2" gridrow="24" pageId="13" pageNumber="286">5.4–11.7</td>
<td id="6C2481347846007AFD27FB5BB36F762C" box="[749,851,1258,1280]" gridcol="3" gridrow="24" pageId="13" pageNumber="286">
<emphasis id="A3B136187846FF88FD27FB5BB36E762C" box="[749,850,1258,1280]" italics="true" pageId="13" pageNumber="286">19.2–28.8</emphasis>
</td>
<td id="6C2481347846007AFC5DFB5BB3FF762C" box="[919,963,1258,1280]" gridcol="4" gridrow="24" pageId="13" pageNumber="286">9.6</td>
<td id="6C2481347846007AFC34FB5BB411762C" box="[1022,1069,1258,1280]" gridcol="5" gridrow="24" pageId="13" pageNumber="286">
<emphasis id="A3B136187846FF88FBCBFB5BB411762C" box="[1025,1069,1258,1280]" italics="true" pageId="13" pageNumber="286">23.7</emphasis>
</td>
<td id="6C2481347846007AFB95FB5BB4BD762C" box="[1119,1153,1258,1280]" gridcol="6" gridrow="24" pageId="13" pageNumber="286">1.9</td>
<td id="6C2481347846007AFB7BFB5BB4EB762C" box="[1201,1239,1258,1280]" gridcol="7" gridrow="24" pageId="13" pageNumber="286">
<emphasis id="A3B136187846FF88FB7EFB5BB4EF762C" box="[1204,1235,1258,1280]" italics="true" pageId="13" pageNumber="286">2.4</emphasis>
</td>
<td id="6C2481347846007AFAC8FB5BB512762C" box="[1282,1326,1258,1280]" gridcol="8" gridrow="24" pageId="13" pageNumber="286">11.0</td>
<td id="6C2481347846007AFA9CFB5BB5B9762C" box="[1366,1413,1258,1280]" gridcol="9" gridrow="24" pageId="13" pageNumber="286">
<emphasis id="A3B136187846FF88FA92FB5BB5B8762C" box="[1368,1412,1258,1280]" italics="true" pageId="13" pageNumber="286">24.6</emphasis>
</td>
</tr>
<tr id="2FF5E8487846007AFF55FAA5B5B97606" box="[159,1413,1300,1322]" gridrow="25" pageId="13" pageNumber="286">
<th id="6C2481347846007AFF55FAA5B5B97606" box="[159,1413,1300,1322]" colspan="10" colspanRight="9" gridcol="0" gridrow="25" pageId="13" pageNumber="286">Claw 3 lengths</th>
</tr>
<tr id="2FF5E8487846007AFF55FA8CB5B9767F" box="[159,1413,1341,1363]" gridrow="26" pageId="13" pageNumber="286">
<th id="6C2481347846007AFF55FA8CB1F7767F" box="[159,459,1341,1363]" gridcol="0" gridrow="26" pageId="13" pageNumber="286">External primary branch</th>
<td id="6C2481347846007AFDC6FA8CB21B767F" box="[524,551,1341,1363]" gridcol="1" gridrow="26" pageId="13" pageNumber="286">27</td>
<td id="6C2481347846007AFD92FA8CB282767F" box="[600,702,1341,1363]" gridcol="2" gridrow="26" pageId="13" pageNumber="286">8.4–16.7</td>
<td id="6C2481347846007AFD27FA8CB36F767F" box="[749,851,1341,1363]" gridcol="3" gridrow="26" pageId="13" pageNumber="286">
<emphasis id="A3B136187846FF88FD27FA8CB36E767F" box="[749,850,1341,1363]" italics="true" pageId="13" pageNumber="286">29.7–39.1</emphasis>
</td>
<td id="6C2481347846007AFC5DFA8CB3FF767F" box="[919,963,1341,1363]" gridcol="4" gridrow="26" pageId="13" pageNumber="286">14.4</td>
<td id="6C2481347846007AFC34FA8CB411767F" box="[1022,1069,1341,1363]" gridcol="5" gridrow="26" pageId="13" pageNumber="286">
<emphasis id="A3B136187846FF88FBCBFA8CB411767F" box="[1025,1069,1341,1363]" italics="true" pageId="13" pageNumber="286">34.8</emphasis>
</td>
<td id="6C2481347846007AFB95FA8CB4BD767F" box="[1119,1153,1341,1363]" gridcol="6" gridrow="26" pageId="13" pageNumber="286">2.2</td>
<td id="6C2481347846007AFB7BFA8CB4EB767F" box="[1201,1239,1341,1363]" gridcol="7" gridrow="26" pageId="13" pageNumber="286">
<emphasis id="A3B136187846FF88FB7EFA8CB4EF767F" box="[1204,1235,1341,1363]" italics="true" pageId="13" pageNumber="286">2.3</emphasis>
</td>
<td id="6C2481347846007AFAC8FA8CB512767F" box="[1282,1326,1341,1363]" gridcol="8" gridrow="26" pageId="13" pageNumber="286">16.0</td>
<td id="6C2481347846007AFA9CFA8CB5B9767F" box="[1366,1413,1341,1363]" gridcol="9" gridrow="26" pageId="13" pageNumber="286">
<emphasis id="A3B136187846FF88FA92FA8CB5B8767F" box="[1368,1412,1341,1363]" italics="true" pageId="13" pageNumber="286">35.7</emphasis>
</td>
</tr>
<tr id="2FF5E8487846007AFF55FAD7B5B97650" box="[159,1413,1382,1404]" gridrow="27" pageId="13" pageNumber="286">
<th id="6C2481347846007AFF55FAD7B1F77650" box="[159,459,1382,1404]" gridcol="0" gridrow="27" pageId="13" pageNumber="286">External secondary branch</th>
<td id="6C2481347846007AFDC6FAD7B21B7650" box="[524,551,1382,1404]" gridcol="1" gridrow="27" pageId="13" pageNumber="286">19</td>
<td id="6C2481347846007AFD92FAD7B2827650" box="[600,702,1382,1404]" gridcol="2" gridrow="27" pageId="13" pageNumber="286">6.0–12.9</td>
<td id="6C2481347846007AFD27FAD7B36F7650" box="[749,851,1382,1404]" gridcol="3" gridrow="27" pageId="13" pageNumber="286">
<emphasis id="A3B136187846FF88FD27FAD7B36E7650" box="[749,850,1382,1404]" italics="true" pageId="13" pageNumber="286">21.2–33.3</emphasis>
</td>
<td id="6C2481347846007AFC5DFAD7B3FF7650" box="[919,963,1382,1404]" gridcol="4" gridrow="27" pageId="13" pageNumber="286">11.2</td>
<td id="6C2481347846007AFC34FAD7B4117650" box="[1022,1069,1382,1404]" gridcol="5" gridrow="27" pageId="13" pageNumber="286">
<emphasis id="A3B136187846FF88FBCBFAD7B4117650" box="[1025,1069,1382,1404]" italics="true" pageId="13" pageNumber="286">26.5</emphasis>
</td>
<td id="6C2481347846007AFB95FAD7B4BD7650" box="[1119,1153,1382,1404]" gridcol="6" gridrow="27" pageId="13" pageNumber="286">1.5</td>
<td id="6C2481347846007AFB7BFAD7B4EB7650" box="[1201,1239,1382,1404]" gridcol="7" gridrow="27" pageId="13" pageNumber="286">
<emphasis id="A3B136187846FF88FB7EFAD7B4EF7650" box="[1204,1235,1382,1404]" italics="true" pageId="13" pageNumber="286">3.0</emphasis>
</td>
<td id="6C2481347846007AFAC8FAD7B5127650" box="[1282,1326,1382,1404]" gridcol="8" gridrow="27" pageId="13" pageNumber="286">11.6</td>
<td id="6C2481347846007AFA9CFAD7B5B97650" box="[1366,1413,1382,1404]" gridcol="9" gridrow="27" pageId="13" pageNumber="286">
<emphasis id="A3B136187846FF88FA92FAD7B5B87650" box="[1368,1412,1382,1404]" italics="true" pageId="13" pageNumber="286">25.9</emphasis>
</td>
</tr>
<tr id="2FF5E8487846007AFF55FA21B5B9768A" box="[159,1413,1424,1446]" gridrow="28" pageId="13" pageNumber="286">
<th id="6C2481347846007AFF55FA21B1F7768A" box="[159,459,1424,1446]" gridcol="0" gridrow="28" pageId="13" pageNumber="286">Internal primary branch</th>
<td id="6C2481347846007AFDC6FA21B21B768A" box="[524,551,1424,1446]" gridcol="1" gridrow="28" pageId="13" pageNumber="286">28</td>
<td id="6C2481347846007AFD92FA21B282768A" box="[600,702,1424,1446]" gridcol="2" gridrow="28" pageId="13" pageNumber="286">7.5–17.0</td>
<td id="6C2481347846007AFD27FA21B36F768A" box="[749,851,1424,1446]" gridcol="3" gridrow="28" pageId="13" pageNumber="286">
<emphasis id="A3B136187846FF88FD27FA21B36F768A" box="[749,851,1424,1446]" italics="true" pageId="13" pageNumber="286">26.5–37.2</emphasis>
</td>
<td id="6C2481347846007AFC5DFA21B3FF768A" box="[919,963,1424,1446]" gridcol="4" gridrow="28" pageId="13" pageNumber="286">13.4</td>
<td id="6C2481347846007AFC34FA21B411768A" box="[1022,1069,1424,1446]" gridcol="5" gridrow="28" pageId="13" pageNumber="286">
<emphasis id="A3B136187846FF88FBCBFA21B411768A" box="[1025,1069,1424,1446]" italics="true" pageId="13" pageNumber="286">32.4</emphasis>
</td>
<td id="6C2481347846007AFB95FA21B4BD768A" box="[1119,1153,1424,1446]" gridcol="6" gridrow="28" pageId="13" pageNumber="286">2.3</td>
<td id="6C2481347846007AFB7BFA21B4EB768A" box="[1201,1239,1424,1446]" gridcol="7" gridrow="28" pageId="13" pageNumber="286">
<emphasis id="A3B136187846FF88FB7EFA21B4EF768A" box="[1204,1235,1424,1446]" italics="true" pageId="13" pageNumber="286">2.2</emphasis>
</td>
<td id="6C2481347846007AFAC8FA21B512768A" box="[1282,1326,1424,1446]" gridcol="8" gridrow="28" pageId="13" pageNumber="286">14.9</td>
<td id="6C2481347846007AFA9CFA21B5B9768A" box="[1366,1413,1424,1446]" gridcol="9" gridrow="28" pageId="13" pageNumber="286">
<emphasis id="A3B136187846FF88FA92FA21B5B8768A" box="[1368,1412,1424,1446]" italics="true" pageId="13" pageNumber="286">33.3</emphasis>
</td>
</tr>
<tr id="2FF5E8487846007AFF55FA08B5B976E3" box="[159,1413,1465,1487]" gridrow="29" pageId="13" pageNumber="286">
<th id="6C2481347846007AFF55FA08B1F776E3" box="[159,459,1465,1487]" gridcol="0" gridrow="29" pageId="13" pageNumber="286">Internal secondary branch</th>
<td id="6C2481347846007AFDC6FA08B21B76E3" box="[524,551,1465,1487]" gridcol="1" gridrow="29" pageId="13" pageNumber="286">21</td>
<td id="6C2481347846007AFD92FA08B28276E3" box="[600,702,1465,1487]" gridcol="2" gridrow="29" pageId="13" pageNumber="286">5.6–14.9</td>
<td id="6C2481347846007AFD27FA08B36F76E3" box="[749,851,1465,1487]" gridcol="3" gridrow="29" pageId="13" pageNumber="286">
<emphasis id="A3B136187846FF88FD27FA08B36E76E3" box="[749,850,1465,1487]" italics="true" pageId="13" pageNumber="286">20.1–29.9</emphasis>
</td>
<td id="6C2481347846007AFC5DFA08B3FF76E3" box="[919,963,1465,1487]" gridcol="4" gridrow="29" pageId="13" pageNumber="286">9.9</td>
<td id="6C2481347846007AFC34FA08B41176E3" box="[1022,1069,1465,1487]" gridcol="5" gridrow="29" pageId="13" pageNumber="286">
<emphasis id="A3B136187846FF88FBCBFA08B41176E3" box="[1025,1069,1465,1487]" italics="true" pageId="13" pageNumber="286">24.3</emphasis>
</td>
<td id="6C2481347846007AFB95FA08B4BD76E3" box="[1119,1153,1465,1487]" gridcol="6" gridrow="29" pageId="13" pageNumber="286">2.2</td>
<td id="6C2481347846007AFB7BFA08B4EB76E3" box="[1201,1239,1465,1487]" gridcol="7" gridrow="29" pageId="13" pageNumber="286">
<emphasis id="A3B136187846FF88FB7EFA08B4EF76E3" box="[1204,1235,1465,1487]" italics="true" pageId="13" pageNumber="286">2.6</emphasis>
</td>
<td id="6C2481347846007AFAC8FA08B51276E3" box="[1282,1326,1465,1487]" gridcol="8" gridrow="29" pageId="13" pageNumber="286">11.8</td>
<td id="6C2481347846007AFA9CFA08B5B976E3" box="[1366,1413,1465,1487]" gridcol="9" gridrow="29" pageId="13" pageNumber="286">
<emphasis id="A3B136187846FF88FA92FA08B5B876E3" box="[1368,1412,1465,1487]" italics="true" pageId="13" pageNumber="286">26.3</emphasis>
</td>
</tr>
<tr id="2FF5E8487846007AFF55FA53B5B976D4" box="[159,1413,1506,1528]" gridrow="30" pageId="13" pageNumber="286">
<th id="6C2481347846007AFF55FA53B5B976D4" box="[159,1413,1506,1528]" colspan="10" colspanRight="9" gridcol="0" gridrow="30" pageId="13" pageNumber="286">Claw 4 lengths</th>
</tr>
<tr id="2FF5E8487846007AFF55F9BDB5B9750E" box="[159,1413,1548,1570]" gridrow="31" pageId="13" pageNumber="286">
<th id="6C2481347846007AFF55F9BDB1F7750E" box="[159,459,1548,1570]" gridcol="0" gridrow="31" pageId="13" pageNumber="286">Anterior primary branch</th>
<td id="6C2481347846007AFDC6F9BDB21B750E" box="[524,551,1548,1570]" gridcol="1" gridrow="31" pageId="13" pageNumber="286">29</td>
<td id="6C2481347846007AFD92F9BDB282750E" box="[600,702,1548,1570]" gridcol="2" gridrow="31" pageId="13" pageNumber="286">8.3–18.5</td>
<td id="6C2481347846007AFD27F9BDB36F750E" box="[749,851,1548,1570]" gridcol="3" gridrow="31" pageId="13" pageNumber="286">
<emphasis id="A3B136187846FF88FD27F9BDB36E750E" box="[749,850,1548,1570]" italics="true" pageId="13" pageNumber="286">29.5–45.9</emphasis>
</td>
<td id="6C2481347846007AFC5DF9BDB3FF750E" box="[919,963,1548,1570]" gridcol="4" gridrow="31" pageId="13" pageNumber="286">15.6</td>
<td id="6C2481347846007AFC34F9BDB411750E" box="[1022,1069,1548,1570]" gridcol="5" gridrow="31" pageId="13" pageNumber="286">
<emphasis id="A3B136187846FF88FBCBF9BDB411750E" box="[1025,1069,1548,1570]" italics="true" pageId="13" pageNumber="286">38.4</emphasis>
</td>
<td id="6C2481347846007AFB95F9BDB4BD750E" box="[1119,1153,1548,1570]" gridcol="6" gridrow="31" pageId="13" pageNumber="286">3.0</td>
<td id="6C2481347846007AFB7BF9BDB4EB750E" box="[1201,1239,1548,1570]" gridcol="7" gridrow="31" pageId="13" pageNumber="286">
<emphasis id="A3B136187846FF88FB7EF9BDB4EF750E" box="[1204,1235,1548,1570]" italics="true" pageId="13" pageNumber="286">3.9</emphasis>
</td>
<td id="6C2481347846007AFAC8F9BDB512750E" box="[1282,1326,1548,1570]" gridcol="8" gridrow="31" pageId="13" pageNumber="286">18.5</td>
<td id="6C2481347846007AFA9CF9BDB5B9750E" box="[1366,1413,1548,1570]" gridcol="9" gridrow="31" pageId="13" pageNumber="286">
<emphasis id="A3B136187846FF88FA92F9BDB5B8750E" box="[1368,1412,1548,1570]" italics="true" pageId="13" pageNumber="286">41.3</emphasis>
</td>
</tr>
<tr id="2FF5E8487846007AFF55F984B5B97567" box="[159,1413,1589,1611]" gridrow="32" pageId="13" pageNumber="286">
<th id="6C2481347846007AFF55F984B1F77567" box="[159,459,1589,1611]" gridcol="0" gridrow="32" pageId="13" pageNumber="286">Anterior secondary branch</th>
<td id="6C2481347846007AFDC6F984B21B7567" box="[524,551,1589,1611]" gridcol="1" gridrow="32" pageId="13" pageNumber="286">26</td>
<td id="6C2481347846007AFD92F984B2827567" box="[600,702,1589,1611]" gridcol="2" gridrow="32" pageId="13" pageNumber="286">5.2–13.3</td>
<td id="6C2481347846007AFD27F984B36F7567" box="[749,851,1589,1611]" gridcol="3" gridrow="32" pageId="13" pageNumber="286">
<emphasis id="A3B136187846FF88FD27F984B36E7567" box="[749,850,1589,1611]" italics="true" pageId="13" pageNumber="286">18.5–32.4</emphasis>
</td>
<td id="6C2481347846007AFC5DF984B3FF7567" box="[919,963,1589,1611]" gridcol="4" gridrow="32" pageId="13" pageNumber="286">10.9</td>
<td id="6C2481347846007AFC34F984B4117567" box="[1022,1069,1589,1611]" gridcol="5" gridrow="32" pageId="13" pageNumber="286">
<emphasis id="A3B136187846FF88FBCBF984B4117567" box="[1025,1069,1589,1611]" italics="true" pageId="13" pageNumber="286">26.7</emphasis>
</td>
<td id="6C2481347846007AFB95F984B4BD7567" box="[1119,1153,1589,1611]" gridcol="6" gridrow="32" pageId="13" pageNumber="286">2.3</td>
<td id="6C2481347846007AFB7BF984B4EB7567" box="[1201,1239,1589,1611]" gridcol="7" gridrow="32" pageId="13" pageNumber="286">
<emphasis id="A3B136187846FF88FB7EF984B4EF7567" box="[1204,1235,1589,1611]" italics="true" pageId="13" pageNumber="286">3.0</emphasis>
</td>
<td id="6C2481347846007AFAC8F984B5127567" box="[1282,1326,1589,1611]" gridcol="8" gridrow="32" pageId="13" pageNumber="286">12.5</td>
<td id="6C2481347846007AFA9CF984B5B97567" box="[1366,1413,1589,1611]" gridcol="9" gridrow="32" pageId="13" pageNumber="286">
<emphasis id="A3B136187846FF88FA92F984B5B87567" box="[1368,1412,1589,1611]" italics="true" pageId="13" pageNumber="286">27.9</emphasis>
</td>
</tr>
<tr id="2FF5E8487846007AFF55F9EFB5B97558" box="[159,1413,1630,1652]" gridrow="33" pageId="13" pageNumber="286">
<th id="6C2481347846007AFF55F9EFB1F77558" box="[159,459,1630,1652]" gridcol="0" gridrow="33" pageId="13" pageNumber="286">Posterior primary branch</th>
<td id="6C2481347846007AFDC6F9EFB21B7558" box="[524,551,1630,1652]" gridcol="1" gridrow="33" pageId="13" pageNumber="286">28</td>
<td id="6C2481347846007AFD92F9EFB2827558" box="[600,702,1630,1652]" gridcol="2" gridrow="33" pageId="13" pageNumber="286">8.9–20.7</td>
<td id="6C2481347846007AFD27F9EFB36F7558" box="[749,851,1630,1652]" gridcol="3" gridrow="33" pageId="13" pageNumber="286">
<emphasis id="A3B136187846FF88FD27F9EFB36E7558" box="[749,850,1630,1652]" italics="true" pageId="13" pageNumber="286">32.0–46.5</emphasis>
</td>
<td id="6C2481347846007AFC5DF9EFB3FF7558" box="[919,963,1630,1652]" gridcol="4" gridrow="33" pageId="13" pageNumber="286">17.3</td>
<td id="6C2481347846007AFC34F9EFB4117558" box="[1022,1069,1630,1652]" gridcol="5" gridrow="33" pageId="13" pageNumber="286">
<emphasis id="A3B136187846FF88FBCBF9EFB4117558" box="[1025,1069,1630,1652]" italics="true" pageId="13" pageNumber="286">42.2</emphasis>
</td>
<td id="6C2481347846007AFB95F9EFB4BD7558" box="[1119,1153,1630,1652]" gridcol="6" gridrow="33" pageId="13" pageNumber="286">2.9</td>
<td id="6C2481347846007AFB7BF9EFB4EB7558" box="[1201,1239,1630,1652]" gridcol="7" gridrow="33" pageId="13" pageNumber="286">
<emphasis id="A3B136187846FF88FB7EF9EFB4EF7558" box="[1204,1235,1630,1652]" italics="true" pageId="13" pageNumber="286">3.4</emphasis>
</td>
<td id="6C2481347846007AFAC8F9EFB5127558" box="[1282,1326,1630,1652]" gridcol="8" gridrow="33" pageId="13" pageNumber="286">20.7</td>
<td id="6C2481347846007AFA9CF9EFB5B97558" box="[1366,1413,1630,1652]" gridcol="9" gridrow="33" pageId="13" pageNumber="286">
<emphasis id="A3B136187846FF88FA92F9EFB5B87558" box="[1368,1412,1630,1652]" italics="true" pageId="13" pageNumber="286">46.2</emphasis>
</td>
</tr>
<tr id="2FF5E8487846007AFF55F939B5B975B2" box="[159,1413,1672,1694]" gridrow="34" pageId="13" pageNumber="286">
<th id="6C2481347846007AFF55F939B1F775B2" box="[159,459,1672,1694]" gridcol="0" gridrow="34" pageId="13" pageNumber="286">Posterior secondary branch</th>
<td id="6C2481347846007AFDC6F939B21B75B2" box="[524,551,1672,1694]" gridcol="1" gridrow="34" pageId="13" pageNumber="286">19</td>
<td id="6C2481347846007AFD92F939B28275B2" box="[600,702,1672,1694]" gridcol="2" gridrow="34" pageId="13" pageNumber="286">10.1–14.8</td>
<td id="6C2481347846007AFD27F939B36F75B2" box="[749,851,1672,1694]" gridcol="3" gridrow="34" pageId="13" pageNumber="286">
<emphasis id="A3B136187846FF88FD27F939B36E75B2" box="[749,850,1672,1694]" italics="true" pageId="13" pageNumber="286">26.3–34.1</emphasis>
</td>
<td id="6C2481347846007AFC5DF939B3FF75B2" box="[919,963,1672,1694]" gridcol="4" gridrow="34" pageId="13" pageNumber="286">13.1</td>
<td id="6C2481347846007AFC34F939B41175B2" box="[1022,1069,1672,1694]" gridcol="5" gridrow="34" pageId="13" pageNumber="286">
<emphasis id="A3B136187846FF88FBCBF939B41175B2" box="[1025,1069,1672,1694]" italics="true" pageId="13" pageNumber="286">31.0</emphasis>
</td>
<td id="6C2481347846007AFB95F939B4BD75B2" box="[1119,1153,1672,1694]" gridcol="6" gridrow="34" pageId="13" pageNumber="286">1.2</td>
<td id="6C2481347846007AFB7BF939B4EB75B2" box="[1201,1239,1672,1694]" gridcol="7" gridrow="34" pageId="13" pageNumber="286">
<emphasis id="A3B136187846FF88FB7EF939B4EF75B2" box="[1204,1235,1672,1694]" italics="true" pageId="13" pageNumber="286">2.0</emphasis>
</td>
<td id="6C2481347846007AFAC8F939B51275B2" box="[1282,1326,1672,1694]" gridcol="8" gridrow="34" pageId="13" pageNumber="286">13.9</td>
<td id="6C2481347846007AFA9CF939B5B975B2" box="[1366,1413,1672,1694]" gridcol="9" gridrow="34" pageId="13" pageNumber="286">31.0</td>
</tr>
</table>
</paragraph>
<paragraph id="917AEA0A7846FF88FF5DF96FB32775DB" blockId="13.[151,795,1758,1783]" box="[151,795,1758,1783]" pageId="13" pageNumber="286">
The
<emphasis id="A3B136187846FF88FF03F96FB0C275DB" bold="true" box="[201,254,1758,1783]" pageId="13" pageNumber="286">COI</emphasis>
sequence (GenBank:
<accessionNumber id="8E9677E97846FF88FE3FF96FB24775DB" box="[501,635,1758,1783]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/MH057763" pageId="13" pageNumber="286">MH057763</accessionNumber>
), 745 bp long:
</paragraph>
<paragraph id="917AEA0A7846FF8BFF5DF898B21B7234" blockId="13.[151,1431,1833,2027]" box="[151,1431,154,2027]" lastBlockId="14.[151,1431,154,280]" lastPageId="14" lastPageNumber="287" pageId="13" pageNumber="286">ACTGCCTCTGTGGGAACGTCCTTAAGATTTCTAATTCGGAGGGAATTAAGCCAACCAGGGCTTCTTCTCTCCGACGAACA AATATATAATGTGATTGTAACAAGCCACGCATTTATTATAATTTTTTTTTTCGTGATACCAATTTTAATCGGGGGATTCG GGAACTGACTGGTCCCCCTTATAATTAGAGCACCAGACATAGCTTTTCCACGAATAAATAATTTAAGCTTCTGGATACTG CCCCCCTCTTTTTTCCTAATTACTTTAAGCTCGATAACTGAACAGGGGGCCGGAACCGGATGAACTGTTTACCCACCCCT TTCTCATTTTTTTGCTCACAGGGGCCCTAGAGTAGATCTTACAATTTTTTCACTTCATGTGGCGGGAATTTCATCAATTT TAGGGGCTATTAATTTCATTTCTACAATTCTAAATATGCGAGTACCCCACCTAACCTTAGAAAAAATACCTCTCTTTGTG TGATCCGTTTTTTTAACAGCTATTTTATTACTTCTGGCCCTCCCAGTATTGGCAGGAGGTATTACTATGCTATTACTGGA CCGAAACTTTAATACTTCTTTTTTTGACCCTGCGGGGGGGGGAGACCCTATTTTATACCAACACCTATTTTGGTTTTTTG GCCACCCCGAAGTATATATTTTAATCTTACCAGGCTTTGGAATTATCTCACAAATCGTTATTCACTTCAGAGGAAAATCT CTCACATTCGGACATTTAGGAATAA</paragraph>
</subSubSection>
<subSubSection id="D9DFB9817845FF8BFF0DFEF6B44A72A8" pageId="14" pageNumber="287" type="materials_examined">
<paragraph id="917AEA0A7845FF8BFF0DFEF6B44A72A8" blockId="14.[151,1437,327,712]" pageId="14" pageNumber="287">
<materialsCitation id="21ADE0577845FF8BFF0DFEF6B44A72A8" ID-GBIF-Occurrence="1914122841" collectingDate="2016-08-16" country="Tanzania" county="Udzungwa Mts." elevation="2050" latitude="-7.8236113" location="Mwanihana Peak" longLatPrecision="21" longitude="36.825554" municipality="Mts. National Park" pageId="14" pageNumber="287" specimenCount="1" stateProvince="Morogoro Region" typeStatus="holotype">
<emphasis id="A3B136187845FF8BFF0DFEF6B151724C" bold="true" box="[199,365,327,352]" pageId="14" pageNumber="287">
<typeStatus id="4E7E54A87845FF8BFF0DFEF6B13E724C" box="[199,258,327,352]" pageId="14" pageNumber="287">Type</typeStatus>
locality:
</emphasis>
<geoCoordinate id="F4F18CCD7845FF8BFEB2FEF6B1D6724C" box="[376,490,327,352]" direction="south" orientation="latitude" pageId="14" pageNumber="287" precision="15" value="-7.8236113">7°49'25''S</geoCoordinate>
,
<geoCoordinate id="F4F18CCD7845FF8BFE30FEF6B241724C" box="[506,637,327,352]" direction="east" orientation="longitude" pageId="14" pageNumber="287" precision="15" value="36.825554">36°49'32''E</geoCoordinate>
;
<elevation id="1AE80D397845FF8BFD46FEF6B32F724C" box="[652,787,327,352]" metricMagnitude="3" metricUnit="m" metricValue="2.05" pageId="14" pageNumber="287" unit="m" value="2050.0">
<quantity id="563D47EF7845FF8BFD46FEF6B2D97273" box="[652,741,327,352]" metricMagnitude="3" metricUnit="m" metricValue="2.05" pageId="14" pageNumber="287" unit="m" value="2050.0">2050 m</quantity>
asl
</elevation>
:
<collectingCountry id="E9D2AA9A7845FF8BFCE8FEF6B3B6724C" box="[802,906,327,352]" name="Tanzania" pageId="14" pageNumber="287">Tanzania</collectingCountry>
,
<collectingRegion id="530124E87845FF8BFC53FEF9B456724C" box="[921,1130,327,352]" country="Tanzania" name="Morogoro" pageId="14" pageNumber="287">Morogoro Region</collectingRegion>
,
<collectingCounty id="781B92867845FF8BFBB3FEF6B50D7273" box="[1145,1329,327,352]" pageId="14" pageNumber="287">
Udzungwa
<collectingMunicipality id="711E70707845FF8BFB34FEF9B50D7273" box="[1278,1329,328,351]" pageId="14" pageNumber="287">Mts.</collectingMunicipality>
</collectingCounty>
National Park,
<location id="941ABCD17845FF8BFF12FEDDB1A172A9" LSID="urn:lsid:plazi:treatment:196C5B1C784FFF8BFF5DFC0EB3F771E4:941ABCD17845FF8BFF12FEDDB1A172A9" box="[216,413,364,389]" country="Tanzania" county="Udzungwa Mts." latitude="-7.8236113" longLatPrecision="21" longitude="36.825554" municipality="Mts. National Park" name="Mwanihana Peak" pageId="14" pageNumber="287" stateProvince="Morogoro Region">Mwanihana Peak</location>
; lichen on branches of a bush; coll.
<collectingDate id="F53F35227845FF8BFCE5FEDDB39172A9" box="[815,941,364,389]" pageId="14" pageNumber="287" value="2016-08-16">
<date id="E57BCCCA7845FF8BFCE5FEDDB39172A9" box="[815,941,364,389]" pageId="14" pageNumber="287" value="2016-08-16">16.08.2016</date>
</collectingDate>
by Thomas Pape.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="D9DFB9817845FF8BFF0DFE3EB34672DC" pageId="14" pageNumber="287" type="etymology">
<paragraph id="917AEA0A7845FF8BFF0DFE3EB34672DC" blockId="14.[151,1437,327,712]" pageId="14" pageNumber="287">
<emphasis id="A3B136187845FF8BFF0DFE3EB16D7284" bold="true" box="[199,337,399,424]" pageId="14" pageNumber="287">Etymology:</emphasis>
We take great pleasure in dedicating this new species to the colleague and friend of the first and the second authors, an established taxonomist of Diptera, Thomas Pape, Head of Section of Biosystematics of the Natural History Museum of
<collectingCountry id="E9D2AA9A7845FF8BFE1EFE66B20372DC" box="[468,575,471,496]" name="Denmark" pageId="14" pageNumber="287">Denmark</collectingCountry>
, University of Copenhagen.
</paragraph>
</subSubSection>
<subSubSection id="D9DFB9817845FF8BFF0DFE4DB3F771E4" pageId="14" pageNumber="287" type="materials_examined">
<paragraph id="917AEA0A7845FF8BFF0DFE4DB3F771E4" blockId="14.[151,1437,327,712]" pageId="14" pageNumber="287">
<materialsCitation id="21ADE0577845FF8BFF0DFE4DB3FB71E4" ID-GBIF-Occurrence="1914122840" collectorName="Zoological Museum &amp; Universitetsparken &amp; Copenhagen O" country="Poland" county="University of Copenhagen" location="Institute of Zoology" municipality="Natural History Museum of Denmark" pageId="14" pageNumber="287" specimenCount="114" specimenCount-egg="66" typeStatus="holotype">
<emphasis id="A3B136187845FF8BFF0DFE4DB19C7139" bold="true" box="[199,416,508,533]" pageId="14" pageNumber="287">
<typeStatus id="4E7E54A87845FF8BFF0DFE4DB13E7139" box="[199,258,508,533]" pageId="14" pageNumber="287">Type</typeStatus>
depositories:
</emphasis>
<typeStatus id="4E7E54A87845FF8BFE6DFE4DB2337139" box="[423,527,508,533]" pageId="14" pageNumber="287" type="holotype">Holotype</typeStatus>
(slide TZ.027.01 with
<specimenCount id="87C321837845FF8BFCC5FE4DB3AD7138" box="[783,913,508,532]" pageId="14" pageNumber="287" type="generic" typeStatus="paratypes">2 paratypes</specimenCount>
) and
<specimenCount id="87C321837845FF8BFC07FE4DB4677139" box="[973,1115,508,533]" pageId="14" pageNumber="287" type="generic" typeStatus="paratypes">34 paratypes</specimenCount>
(slides: TZ.027.*, where the asterisk can be substituted by any of the following numbers 02–06, 0 8, 22) and
<specimenCount id="87C321837845FF8BFBD5FDAEB44A7114" box="[1055,1142,543,568]" pageId="14" pageNumber="287" type="egg">56 eggs</specimenCount>
(slides: TZ.027.*: 10–16, 18, 19, 21) are deposited at the
<location id="941ABCD17845FF8BFDF1FDF5B3137171" LSID="urn:lsid:plazi:treatment:196C5B1C784FFF8BFF5DFC0EB3F771E4:941ABCD17845FF8BFDF1FDF5B3137171" box="[571,815,580,605]" country="Poland" county="University of Copenhagen" municipality="Natural History Museum of Denmark" name="Institute of Zoology" pageId="14" pageNumber="287">Institute of Zoology</location>
and Biomedical Research, Jagiellonian University, Gronostajowa 9, 30-387, Kraków,
<collectingCountry id="E9D2AA9A7845FF8BFDFCFDD6B2B871AC" box="[566,644,615,640]" name="Poland" pageId="14" pageNumber="287">Poland</collectingCountry>
and
<specimenCount id="87C321837845FF8BFD0DFDD9B36071AC" box="[711,860,616,640]" pageId="14" pageNumber="287" type="generic" typeStatus="paratypes">12 paratypes</specimenCount>
(slides: TZ.027.*: 0 7, 09) and
<specimenCount id="87C321837845FF8BFB2BFDD9B50171AC" box="[1249,1341,616,640]" pageId="14" pageNumber="287" type="egg">10 eggs</specimenCount>
(slides: TZ.027.*: 17, 20) are deposited in the
<collectorName id="3C308FDC7845FF8BFD9EFD3DB3077188" box="[596,827,652,677]" pageId="14" pageNumber="287">Zoological Museum</collectorName>
,
<collectingMunicipality id="711E70707845FF8BFC82FD3DB4C47189" box="[840,1272,652,677]" pageId="14" pageNumber="287">Natural History Museum of Denmark</collectingMunicipality>
,
<collectingCounty id="781B92867845FF8BFACCFD3DB11A71E4" pageId="14" pageNumber="287">University of Copenhagen</collectingCounty>
,
<collectorName id="3C308FDC7845FF8BFEFBFD1EB23971E4" box="[305,517,687,712]" pageId="14" pageNumber="287">Universitetsparken</collectorName>
15, DK-2100
<collectorName id="3C308FDC7845FF8BFD6CFD1EB37371E4" box="[678,847,687,712]" pageId="14" pageNumber="287">Copenhagen Ø</collectorName>
,
<collectingCountry id="E9D2AA9A7845FF8BFC91FD1EB3FB71E4" box="[859,967,687,712]" name="Denmark" pageId="14" pageNumber="287">Denmark</collectingCountry>
</materialsCitation>
.
</paragraph>
</subSubSection>
</treatment>
</document>
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