treatments-xml/data/03/FB/8C/03FB8C32F81CFFBCFF60F97FFB844A7D.xml
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<document id="0530DE2D99FC270C845709D40654BA41" ID-CLB-Dataset="27714" ID-DOI="10.11646/zootaxa.4543.1.1" ID-GBIF-Dataset="22ef92b8-1d9c-4e9f-b01c-ce1f5d3faa7b" ID-ISSN="1175-5326" ID-Zenodo-Dep="2617607" ID-ZooBank="31ABFA6E-6126-4603-B84F-4BEC7632D1E8" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1553927968043" checkinUser="plazi" docAuthor="Fresneda, Javier, Faille, Arnaud, Fery, Hans &amp; Ribera, Ignacio" docDate="2019" docId="03FB8C32F81CFFBCFF60F97FFB844A7D" docLanguage="en" docName="zootaxa.4543.1.1.pdf" docOrigin="Zootaxa 4543 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Speonemadus comasi Fresneda &amp; Faille &amp; Fery &amp; Ribera 2019, sp. n." docType="treatment" docVersion="9" lastPageNumber="9" masterDocId="FFC2F44AF819FFB4FFF7FF98FFAD4B19" masterDocTitle="A molecular phylogeny of Speonemadus Jeannel, 1922 with description of two new species from Morocco (Coleoptera: Leiodidae: Cholevinae: Anemadini)" masterLastPageNumber="36" masterPageNumber="1" pageNumber="6" updateTime="1698715325246" updateUser="ExternalLinkService">
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<mods:title id="63415F2489318E326EC560B67F539909">A molecular phylogeny of Speonemadus Jeannel, 1922 with description of two new species from Morocco (Coleoptera: Leiodidae: Cholevinae: Anemadini)</mods:title>
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<mods:namePart id="E085870AEBD12F3193AF6D78F076B030">Fresneda, Javier</mods:namePart>
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<mods:namePart id="D74305B5D443393C7DC2828E8D45192B">Faille, Arnaud</mods:namePart>
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<mods:namePart id="7C5A81B8178FEA6D7CEDA8A7E65B916A">Fery, Hans</mods:namePart>
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<mods:namePart id="8C523FD45597C408FCBB76FC5CC75EA8">Ribera, Ignacio</mods:namePart>
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<mods:date id="177D61D870E1A3D906F6D0DDABF74FC8">2019</mods:date>
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<subSubSection id="C3486EAFF81CFFB1FF60F97FFE494C1B" box="[151,484,1767,1794]" pageId="5" pageNumber="6" type="nomenclature">
<paragraph id="8BED3D24F81CFFB1FF60F97FFE494C1B" blockId="5.[151,484,1767,1827]" box="[151,484,1767,1794]" pageId="5" pageNumber="6">
<heading id="D0A58A48F81CFFB1FF60F97FFE494C1B" bold="true" box="[151,484,1767,1794]" fontSize="11" level="1" pageId="5" pageNumber="6" reason="1">
<emphasis id="B926E136F81CFFB1FF60F97FFE494C1B" bold="true" box="[151,484,1767,1794]" pageId="5" pageNumber="6">
<taxonomicName id="4C5246A7F81CFFB1FF60F97FFE364C18" ID-CoL="8Y9NM" authority="Fresneda &amp; Faille &amp; Fery &amp; Ribera, 2019" authorityName="Fresneda &amp; Faille &amp; Fery &amp; Ribera" authorityYear="2019" box="[151,411,1767,1793]" class="Insecta" family="Leiodidae" genus="Speonemadus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="comasi" status="sp. nov.">
<emphasis id="B926E136F81CFFB1FF60F97FFE364C18" bold="true" box="[151,411,1767,1793]" italics="true" pageId="5" pageNumber="6">Speonemadus comasi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2155C4DF81CFFB1FE56F970FE494C1B" box="[417,484,1768,1794]" pageId="5" pageNumber="6" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
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<subSubSection id="C3486EAFF81CFFB1FF60F892FEE14C3A" box="[151,332,1802,1827]" pageId="5" pageNumber="6" type="description">
<paragraph id="8BED3D24F81CFFB1FF60F892FEE14C3A" blockId="5.[151,484,1767,1827]" box="[151,332,1802,1827]" pageId="5" pageNumber="6">
<figureCitation id="136921A1F81CFFB1FF60F892FF734C3A" box="[151,222,1802,1827]" captionStart="FIGURES 12" captionStartId="6.[151,264,1618,1640]" captionTargetBox="[183,1395,224,1571]" captionTargetId="figure@6.[151,1437,193,1597]" captionTargetPageId="6" captionText="FIGURES 12. Habitus of the holotypes of: (1) Speonemadus brusteli sp. n.; (2) S. comasi sp. n." httpUri="https://zenodo.org/record/2617609/files/figure.png" pageId="5" pageNumber="6">Figs 2</figureCitation>
,
<figureCitation id="136921A1F81CFFB1FF1DF893FF574C3A" box="[234,250,1803,1827]" captionStart="FIGURES 34" captionStartId="7.[151,264,1572,1594]" captionTargetBox="[416,1121,437,1489]" captionTargetId="figure@7.[379,1208,409,1550]" captionTargetPageId="7" captionText="FIGURES 34. Protibia of: (3) holotype of Speonemadus comasi sp. n.; (4) S. angusticollis (Rute, Spain)." httpUri="https://zenodo.org/record/2617611/files/figure.png" pageId="5" pageNumber="6">3</figureCitation>
, 12, 27
</paragraph>
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<paragraph id="8BED3D24F81CFFB1FF60F8CAFBF64C72" blockId="5.[151,1115,1874,1899]" box="[151,1115,1874,1899]" pageId="5" pageNumber="6">
<emphasis id="B926E136F81CFFB1FF60F8CAFE824C72" bold="true" box="[151,303,1874,1899]" pageId="5" pageNumber="6">Type locality</emphasis>
:
<materialsCitation id="3B3A3779F81CFFB1FEC9F8CBFBFA4C72" ID-GBIF-Occurrence="2147524156" box="[318,1111,1874,1899]" country="Morocco" elevation="2250" latitude="32.166668" location="Tagelft" longLatPrecision="1212" longitude="-5.95" municipality="Iri Adar Nyighil" pageId="5" pageNumber="6" specimenCount="1">
<collectingCountry id="F3457DB4F81CFFB1FEC9F8CBFE0B4C73" box="[318,422,1875,1898]" name="Morocco" pageId="5" pageNumber="6">Morocco</collectingCountry>
,
<location id="8E8D6BFFF81CFFB1FE46F8CAFDAE4C72" LSID="urn:lsid:plazi:treatment:03FB8C32F81CFFBCFF60F97FFB844A7D:8E8D6BFFF81CFFB1FE46F8CAFDAE4C72" box="[433,515,1874,1899]" country="Morocco" latitude="32.166668" longLatPrecision="1212" longitude="-5.95" municipality="Iri Adar Nyighil" name="Tagelft" pageId="5" pageNumber="6">Tagelft</location>
,
<collectingMunicipality id="6B89A75EF81CFFB1FDF9F8CAFD6A4C72" box="[526,711,1874,1899]" pageId="5" pageNumber="6">Iri Adar Nyighil</collectingMunicipality>
,
<geoCoordinate id="EE665BE3F81CFFB1FD25F8CBFC834C72" box="[722,814,1875,1899]" direction="north" orientation="latitude" pageId="5" pageNumber="6" precision="925" value="32.166668">32°10'N</geoCoordinate>
<geoCoordinate id="EE665BE3F81CFFB1FCC1F8CAFC274C72" box="[822,906,1874,1899]" direction="west" orientation="longitude" pageId="5" pageNumber="6" precision="925" value="-5.95">5°57'W</geoCoordinate>
,
<quantity id="4CAA90C1F81CFFB1FC61F8CAFC464C73" box="[918,1003,1874,1899]" metricMagnitude="3" metricUnit="m" metricValue="2.25" pageId="5" pageNumber="6" unit="m" value="2250.0">
<elevation id="007FDA17F81CFFB1FC61F8CAFC464C73" box="[918,1003,1874,1899]" metricMagnitude="3" metricUnit="m" metricValue="2.25" pageId="5" pageNumber="6" unit="m" value="2250.0">2250 m</elevation>
</quantity>
(
<figureCitation id="136921A1F81CFFB1FC0DF8CAFBE34C72" box="[1018,1102,1874,1899]" captionStart="FIGURE 45" captionStartId="27.[151,250,2008,2030]" captionTargetBox="[151,1436,1021,1985]" captionTargetId="figure@27.[151,1436,1021,1985]" captionTargetPageId="27" captionText="FIGURE 45. Type locality of Speonemadus comasi n. sp., Iri Adar Nyighil, Tagelft, Morocco (photo: J. Comas)." httpUri="https://zenodo.org/record/2617649/files/figure.png" pageId="5" pageNumber="6">Fig. 45</figureCitation>
)
</materialsCitation>
.
</paragraph>
<caption id="DF2D6DACF81FFFB2FF60F9CAFBD94D71" box="[151,1140,1618,1640]" httpUri="https://zenodo.org/record/2617609/files/figure.png" pageId="6" pageNumber="7" startId="6.[151,264,1618,1640]" targetBox="[183,1395,224,1571]" targetPageId="6">
<paragraph id="8BED3D24F81FFFB2FF60F9CAFBD94D71" blockId="6.[151,1140,1618,1640]" box="[151,1140,1618,1640]" pageId="6" pageNumber="7">
<emphasis id="B926E136F81FFFB2FF60F9CAFE994D71" bold="true" box="[151,308,1618,1640]" pageId="6" pageNumber="7">FIGURES 12</emphasis>
. Habitus of the holotypes of:
<emphasis id="B926E136F81FFFB2FD96F9CAFDD34D71" bold="true" box="[609,638,1618,1640]" pageId="6" pageNumber="7">(1)</emphasis>
<taxonomicName id="4C5246A7F81FFFB2FD72F9CAFCCE4D71" authority="Fresneda &amp; Faille &amp; Fery &amp; Ribera, 2019" authorityName="Fresneda &amp; Faille &amp; Fery &amp; Ribera" authorityYear="2019" box="[645,867,1618,1640]" class="Insecta" family="Leiodidae" genus="Speonemadus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="brusteli" status="sp. nov.">
<emphasis id="B926E136F81FFFB2FD72F9CAFCCE4D71" box="[645,867,1618,1640]" italics="true" pageId="6" pageNumber="7">Speonemadus brusteli</emphasis>
</taxonomicName>
<emphasis id="B926E136F81FFFB2FC9DF9CAFC0E4D71" bold="true" box="[874,931,1618,1640]" pageId="6" pageNumber="7">
<taxonomicNameLabel id="A2155C4DF81FFFB2FC9DF9CAFC0E4D71" box="[874,931,1618,1640]" pageId="6" pageNumber="7" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
;
<emphasis id="B926E136F81FFFB2FC46F9CAFC634D71" bold="true" box="[945,974,1618,1640]" pageId="6" pageNumber="7">(2)</emphasis>
<taxonomicName id="4C5246A7F81FFFB2FC23F9CAFB9B4D71" authority="Fresneda &amp; Faille &amp; Fery &amp; Ribera, 2019" authorityName="Fresneda &amp; Faille &amp; Fery &amp; Ribera" authorityYear="2019" box="[980,1078,1618,1640]" class="Insecta" family="Leiodidae" genus="Speonemadus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="comasi" status="sp. nov.">
<emphasis id="B926E136F81FFFB2FC23F9CAFB9B4D71" box="[980,1078,1618,1640]" italics="true" pageId="6" pageNumber="7">S. comasi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2155C4DF81FFFB2FBCBF9CAFBD94D71" box="[1084,1140,1618,1640]" pageId="6" pageNumber="7" rank="species">sp. n.</taxonomicNameLabel>
</paragraph>
</caption>
<paragraph id="8BED3D24F81FFFB2FF30F917FAEA4DD5" blockId="6.[151,1436,1679,2028]" pageId="6" pageNumber="7">
<materialsCitation id="3B3A3779F81FFFB2FF30F917FAEA4DD5" ID-GBIF-Occurrence="2147524153" collectingDate="2005-07-09" collectionCode="MZB" pageId="6" pageNumber="7" specimenCount="1" specimenCount-male="1" typeStatus="holotype">
<emphasis id="B926E136F81FFFB2FF30F917FE904DB1" bold="true" box="[199,317,1679,1704]" pageId="6" pageNumber="7">
<typeStatus id="54E98386F81FFFB2FF30F917FE954DB1" box="[199,312,1679,1704]" pageId="6" pageNumber="7" type="holotype">Holotype</typeStatus>
:
</emphasis>
<specimenCount id="9D54F6ADF81FFFB2FEBEF908FECD4DBE" box="[329,352,1680,1703]" pageId="6" pageNumber="7" type="male"></specimenCount>
, &quot;Iri Adar Nyighil / -Tagelft- Marroc /
<date id="FFEC1BE4F81FFFB2FCCDF917FC174DB1" box="[826,954,1679,1704]" pageId="6" pageNumber="7" value="2005-07-09">
<collectingDate id="EFA8E20CF81FFFB2FCCDF917FC174DB1" box="[826,954,1679,1704]" pageId="6" pageNumber="7" value="2005-07-09">9-VII-2005</collectingDate>
</date>
/ Fadrique- [hw] / Auroux-Comas Leg.&quot; [printed], &quot;
<typeStatus id="54E98386F81FFFB2FEF8F92CFE144DD5" box="[271,441,1716,1740]" pageId="6" pageNumber="7" type="holotype">HOLOTYPUS</typeStatus>
/
<taxonomicName id="4C5246A7F81FFFB2FE39F92BFDC54DD5" authorityName="Jeannel" authorityYear="1922" box="[462,616,1715,1740]" class="Insecta" family="Leiodidae" genus="Speonemadus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="genus">
<emphasis id="B926E136F81FFFB2FE39F92BFDC54DD5" box="[462,616,1715,1740]" italics="true" pageId="6" pageNumber="7">Speonemadus</emphasis>
</taxonomicName>
/
<taxonomicName id="4C5246A7F81FFFB2FD89F92CFD604DD2" authority="Fresneda &amp; Faille &amp; Fery &amp; Ribera, 2019" authorityName="Fresneda &amp; Faille &amp; Fery &amp; Ribera" authorityYear="2019" box="[638,717,1716,1739]" class="Insecta" family="Leiodidae" genus="Speonemadus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="comasi" status="sp. nov.">
<emphasis id="B926E136F81FFFB2FD89F92CFD604DD2" box="[638,717,1716,1739]" italics="true" pageId="6" pageNumber="7">comasi</emphasis>
</taxonomicName>
<emphasis id="B926E136F81FFFB2FD23F92BFCB94DD5" bold="true" box="[724,788,1715,1740]" pageId="6" pageNumber="7">
<taxonomicNameLabel id="A2155C4DF81FFFB2FD23F92BFCB94DD5" box="[724,788,1715,1740]" pageId="6" pageNumber="7" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
/ JF-AF-HF-IR det. 2017&quot; [red label, hw] (
<collectionCode id="ED43A5E1F81FFFB2FB0FF92CFA974DD5" box="[1272,1338,1716,1740]" country="Indonesia" httpUri="http://grbio.org/cool/b9gz-izvg" name="Museum Zoologicum Bogoriense" pageId="6" pageNumber="7" type="Museum">MZB</collectionCode>
).
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C3486EAFF81FFFB3FF30F94FFE0D4D66" lastPageId="7" lastPageNumber="8" pageId="6" pageNumber="7" type="description">
<paragraph id="8BED3D24F81FFFB3FF30F94FFDE64A7C" blockId="6.[151,1436,1679,2028]" lastBlockId="7.[151,1436,151,357]" lastPageId="7" lastPageNumber="8" pageId="6" pageNumber="7">
<emphasis id="B926E136F81FFFB2FF30F94FFDD74DE9" bold="true" box="[199,634,1751,1776]" pageId="6" pageNumber="7">
Description of the
<typeStatus id="54E98386F81FFFB2FE53F94FFDA24DE9" box="[420,527,1751,1776]" pageId="6" pageNumber="7" type="holotype">holotype</typeStatus>
: Habitus
</emphasis>
as in
<figureCitation id="136921A1F81FFFB2FD4AF94FFCAF4DE9" box="[701,770,1751,1776]" captionStart="FIGURES 12" captionStartId="6.[151,264,1618,1640]" captionTargetBox="[183,1395,224,1571]" captionTargetId="figure@6.[151,1437,193,1597]" captionTargetPageId="6" captionText="FIGURES 12. Habitus of the holotypes of: (1) Speonemadus brusteli sp. n.; (2) S. comasi sp. n." httpUri="https://zenodo.org/record/2617609/files/figure.png" pageId="6" pageNumber="7">Fig. 2</figureCitation>
.
<emphasis id="B926E136F81FFFB2FCFAF94FFCE64DE9" bold="true" box="[781,843,1751,1776]" pageId="6" pageNumber="7">Body</emphasis>
length 3.6 mm. Body shape elongate, colour brownreddish, darker on head and pronotal disk.
<emphasis id="B926E136F81FFFB2FD88F963FD124C0D" bold="true" box="[639,703,1787,1812]" pageId="6" pageNumber="7">Head</emphasis>
with eyes well developed, with visible suture between epistome and frons; surface with coarse, dense punctation; antennomeres VIIXI forming a loose club, antennomere X as long as wide.
<emphasis id="B926E136F81FFFB2FECFF8DBFE1C4C45" bold="true" box="[312,433,1859,1884]" pageId="6" pageNumber="7">Pronotum</emphasis>
transverse (maximum width 1.3 mm; maximum length 0.8 mm), sides strongly arched, still more strongly posteriorly; base of pronotum sinuate, as wide as base of elytra; surface with coarse, dense and rough punctation. Elytra very elongate, wider in basal third, regularly tapered towards apex; with parasutural stria; surface with fine and sparse transverse striolae.
<emphasis id="B926E136F81FFFB2FD4CF837FD5E4CD1" bold="true" box="[699,755,1967,1992]" pageId="6" pageNumber="7">Legs</emphasis>
with pentamerous tarsi; protibiae sinuate, more strongly on inner side (
<figureCitation id="136921A1F81FFFB2FEE0F84BFEF14CF5" box="[279,348,2003,2028]" captionStart="FIGURES 34" captionStartId="7.[151,264,1572,1594]" captionTargetBox="[416,1121,437,1489]" captionTargetId="figure@7.[379,1208,409,1550]" captionTargetPageId="7" captionText="FIGURES 34. Protibia of: (3) holotype of Speonemadus comasi sp. n.; (4) S. angusticollis (Rute, Spain)." httpUri="https://zenodo.org/record/2617611/files/figure.png" pageId="6" pageNumber="7">Fig. 3</figureCitation>
); mesotibiae less sinuate, more regularly curved; first three tarsomeres of protarsi and first two tarsomeres of mesotarsi dilated.
<emphasis id="B926E136F81EFFB3FDEDFF0FFD664BA9" bold="true" box="[538,715,151,176]" pageId="7" pageNumber="8">Male genitalia</emphasis>
(Fig. 12). Basal lamina of aedeagus as long as apical part; parameres about as long as median lobe; median lobe in dorsal view with sides slightly sinuate, regularly tapering towards apex; apex with small, rounded extension; basal 2/3 of parameres straight, apical third with external side strongly dilated, narrowing towards apex; apex with triangular expansion on external side, with four fine setae and without conical spine (
<figureCitation id="136921A1F81EFFB3FE50FEBFFDAD4A59" box="[423,512,295,320]" captionStart="FIGURES 1328" captionStartId="12.[151,264,1731,1753]" captionTargetBox="[170,1387,215,1697]" captionTargetId="figure@12.[151,1436,193,1710]" captionTargetPageId="12" captionText="FIGURES 1328. Apex of the parameres of: (13) Speonemadus orchesioides (Ammi-Moussa, Algeria); (14) holotype of S. brusteli sp. n. (Talembote, Morocco); (15) S. gracilis (Hornos, Spain); (16) S. tenuipes (Ifri Semedane, Algeria); (17) S. vandalitiae (Villanúa, Spain); (18) S. pulchellus (Ficuzza, Sicily); (19) holotype of S. algarvensis (Gruta do Vale Telheiro, Portugal); (20) S. angusticollis (Rute, Spain); (21) S. bolivari (Ardales, Spain); (22) S. breuili (Motillas, Spain); (23) S. escalerai (Jumilla, Spain); (24) S. transversostriatus (La Bastida, Spain); (25) S. clathratus (Linarejos, Spain); (26) S. maroccanus (Taza, Morocco); (27) holotype of S. comasi sp. n. (Tagelft, Morocco); (28) Anemadus hajeki (Yunnan, China)." httpUri="https://zenodo.org/record/2617619/files/figure.png" pageId="7" pageNumber="8">Fig. 27</figureCitation>
); endophallus entirely covered with scales and spines, with two dark parallel longitudinal bands scarcely noticeable.
</paragraph>
<caption id="DF2D6DACF81EFFB3FF60F9BCFB7D4D23" box="[151,1232,1572,1594]" httpUri="https://zenodo.org/record/2617611/files/figure.png" pageId="7" pageNumber="8" startId="7.[151,264,1572,1594]" targetBox="[416,1121,437,1489]" targetPageId="7">
<paragraph id="8BED3D24F81EFFB3FF60F9BCFB7D4D23" blockId="7.[151,1232,1572,1594]" box="[151,1232,1572,1594]" pageId="7" pageNumber="8">
<emphasis id="B926E136F81EFFB3FF60F9BCFE974D23" bold="true" box="[151,314,1572,1594]" pageId="7" pageNumber="8">FIGURES 34.</emphasis>
Protibia of:
<emphasis id="B926E136F81EFFB3FE4EF9BCFE7A4D23" bold="true" box="[441,471,1572,1594]" pageId="7" pageNumber="8">(3)</emphasis>
holotype of
<taxonomicName id="4C5246A7F81EFFB3FDA0F9BCFC9C4D23" authority="Fresneda &amp; Faille &amp; Fery &amp; Ribera, 2019" authorityName="Fresneda &amp; Faille &amp; Fery &amp; Ribera" authorityYear="2019" box="[599,817,1572,1594]" class="Insecta" family="Leiodidae" genus="Speonemadus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="comasi" status="sp. nov.">
<emphasis id="B926E136F81EFFB3FDA0F9BCFC9C4D23" box="[599,817,1572,1594]" italics="true" pageId="7" pageNumber="8">Speonemadus comasi</emphasis>
</taxonomicName>
<emphasis id="B926E136F81EFFB3FCC0F9BCFCDD4D23" bold="true" box="[823,880,1572,1594]" pageId="7" pageNumber="8">
<taxonomicNameLabel id="A2155C4DF81EFFB3FCC0F9BCFCDD4D23" box="[823,880,1572,1594]" pageId="7" pageNumber="8" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
;
<emphasis id="B926E136F81EFFB3FC8AF9BCFC364D23" bold="true" box="[893,923,1572,1594]" pageId="7" pageNumber="8">(4)</emphasis>
<taxonomicName id="4C5246A7F81EFFB3FC56F9BCFB914D23" baseAuthorityName="Kraatz" baseAuthorityYear="1870" box="[929,1084,1572,1594]" class="Insecta" family="Leiodidae" genus="Speonemadus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="angusticollis">
<emphasis id="B926E136F81EFFB3FC56F9BCFB914D23" box="[929,1084,1572,1594]" italics="true" pageId="7" pageNumber="8">S. angusticollis</emphasis>
</taxonomicName>
(Rute, Spain).
</paragraph>
</caption>
<paragraph id="8BED3D24F81EFFB3FF30F9FEFE0D4D66" blockId="7.[151,1436,1638,2023]" box="[199,416,1638,1663]" pageId="7" pageNumber="8">
<emphasis id="B926E136F81EFFB3FF30F9FEFE844D66" bold="true" box="[199,297,1638,1663]" pageId="7" pageNumber="8">Females</emphasis>
unknown.
</paragraph>
</subSubSection>
<subSubSection id="C3486EAFF81EFFB3FF30F912FE9D4DF3" pageId="7" pageNumber="8" type="etymology">
<paragraph id="8BED3D24F81EFFB3FF30F912FE9D4DF3" blockId="7.[151,1436,1638,2023]" pageId="7" pageNumber="8">
<emphasis id="B926E136F81EFFB3FF30F912FEE34DBA" bold="true" box="[199,334,1674,1699]" pageId="7" pageNumber="8">Etymology.</emphasis>
The species is dedicated to our friend and colleague Jordi Comas (
<collectingCountry id="F3457DB4F81EFFB3FBB9F912FB3F4DBA" box="[1102,1170,1674,1699]" name="Spain" pageId="7" pageNumber="8">Spain</collectingCountry>
, Barcelona), one of the collectors of the only known specimen of this most interesting species. The specific name is a substantive in the genitive case.
</paragraph>
</subSubSection>
<subSubSection id="C3486EAFF81EFFBCFF30F96EFECF4A59" lastPageId="8" lastPageNumber="9" pageId="7" pageNumber="8" type="discussion">
<paragraph id="8BED3D24F81EFFBCFF30F96EFECF4A59" blockId="7.[151,1436,1638,2023]" lastBlockId="8.[151,1436,151,357]" lastPageId="8" lastPageNumber="9" pageId="7" pageNumber="8">
<emphasis id="B926E136F81EFFB3FF30F96EFE964C16" bold="true" box="[199,315,1782,1807]" pageId="7" pageNumber="8">Remarks.</emphasis>
<taxonomicName id="4C5246A7F81EFFB3FEB5F96EFD9E4C17" authority="Fresneda &amp; Faille &amp; Fery &amp; Ribera, 2019" authorityName="Fresneda &amp; Faille &amp; Fery &amp; Ribera" authorityYear="2019" box="[322,563,1782,1807]" class="Insecta" family="Leiodidae" genus="Speonemadus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="comasi" status="sp. nov.">
<emphasis id="B926E136F81EFFB3FEB5F96EFD9E4C17" box="[322,563,1782,1807]" italics="true" pageId="7" pageNumber="8">Speonemadus comasi</emphasis>
</taxonomicName>
<emphasis id="B926E136F81EFFB3FDCCF96EFDD64C16" bold="true" box="[571,635,1782,1807]" pageId="7" pageNumber="8">
<taxonomicNameLabel id="A2155C4DF81EFFB3FDCCF96EFDD64C16" box="[571,635,1782,1807]" pageId="7" pageNumber="8" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
is a peculiar species, with characters similar to both
<taxonomicName id="4C5246A7F81EFFB3FB25F96EFAC14C16" authorityName="Jeannel" authorityYear="1922" box="[1234,1388,1782,1807]" class="Insecta" family="Leiodidae" genus="Speonemadus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="genus">
<emphasis id="B926E136F81EFFB3FB25F96EFAC14C16" box="[1234,1388,1782,1807]" italics="true" pageId="7" pageNumber="8">Speonemadus</emphasis>
</taxonomicName>
and
<emphasis id="B926E136F81EFFB3FF60F882FEBE4C2A" box="[151,275,1818,1843]" italics="true" pageId="7" pageNumber="8">
<taxonomicName id="4C5246A7F81EFFB3FF60F882FEBD4C2A" authorityName="Reitter" authorityYear="1885" box="[151,272,1818,1843]" class="Insecta" family="Leiodidae" genus="Anemadus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="genus">Anemadus</taxonomicName>
.
</emphasis>
The general slender habitus and the modified protibiae are similar to the species of the
<taxonomicName id="4C5246A7F81EFFB3FB1CF882FAFF4C2A" baseAuthorityName="Uhagon" baseAuthorityYear="1898" box="[1259,1362,1818,1843]" class="Insecta" family="Leiodidae" genus="Speonemadus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="escalerai">
<emphasis id="B926E136F81EFFB3FB1CF882FAFF4C2A" box="[1259,1362,1818,1843]" italics="true" pageId="7" pageNumber="8">escalerai</emphasis>
</taxonomicName>
-group (e.g.
<taxonomicName id="4C5246A7F81EFFB3FF38F8A7FED74C4E" baseAuthorityName="Kraatz" baseAuthorityYear="1870" box="[207,378,1854,1879]" class="Insecta" family="Leiodidae" genus="Speonemadus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="angusticollis">
<emphasis id="B926E136F81EFFB3FF38F8A7FED74C4E" box="[207,378,1854,1879]" italics="true" pageId="7" pageNumber="8">S. angusticollis</emphasis>
</taxonomicName>
,
<figureCitation id="136921A1F81EFFB3FE7FF8A6FE664C4E" box="[392,459,1854,1879]" captionStart="FIGURES 34" captionStartId="7.[151,264,1572,1594]" captionTargetBox="[416,1121,437,1489]" captionTargetId="figure@7.[379,1208,409,1550]" captionTargetPageId="7" captionText="FIGURES 34. Protibia of: (3) holotype of Speonemadus comasi sp. n.; (4) S. angusticollis (Rute, Spain)." httpUri="https://zenodo.org/record/2617611/files/figure.png" pageId="7" pageNumber="8">Fig. 4</figureCitation>
), and the shape of the parameres with their triangular apical expansion is similar to that of
<taxonomicName id="4C5246A7F81EFFB3FF4CF8FBFE9F4C62" authority="Fresneda &amp; Faille &amp; Fery &amp; Ribera, 2019" authorityName="Fresneda &amp; Faille &amp; Fery &amp; Ribera" authorityYear="2019" box="[187,306,1890,1915]" class="Insecta" family="Leiodidae" genus="Speonemadus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="brusteli" status="sp. nov.">
<emphasis id="B926E136F81EFFB3FF4CF8FBFE9F4C62" box="[187,306,1890,1915]" italics="true" pageId="7" pageNumber="8">S. brusteli</emphasis>
</taxonomicName>
<emphasis id="B926E136F81EFFB3FEB7F8FAFE284C62" bold="true" box="[320,389,1890,1915]" pageId="7" pageNumber="8">
<taxonomicNameLabel id="A2155C4DF81EFFB3FEB7F8FAFE284C62" box="[320,389,1890,1915]" pageId="7" pageNumber="8" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
of the
<taxonomicName id="4C5246A7F81EFFB3FE12F8FAFDCE4C62" authorityName="" baseAuthorityName="Heyden" baseAuthorityYear="1870" box="[485,611,1890,1915]" class="Insecta" family="Leiodidae" genus="Speonemadus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="vandalitiae">
<emphasis id="B926E136F81EFFB3FE12F8FAFDCE4C62" box="[485,611,1890,1915]" italics="true" pageId="7" pageNumber="8">vandalitiae</emphasis>
</taxonomicName>
-group. However, the endophallus is similar to that of the species of
<taxonomicName id="4C5246A7F81EFFB3FF60F81EFEA14C86" authorityName="Reitter" authorityYear="1885" box="[151,268,1926,1951]" class="Insecta" family="Leiodidae" genus="Anemadus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="genus">
<emphasis id="B926E136F81EFFB3FF60F81EFEA14C86" box="[151,268,1926,1951]" italics="true" pageId="7" pageNumber="8">Anemadus</emphasis>
</taxonomicName>
, without well defined groups of spines, such as those to the
<taxonomicName id="4C5246A7F81EFFB3FC20F81EFBBA4C86" authorityName="Giachino &amp; Vailati" authorityYear="1993" box="[983,1047,1926,1951]" class="Insecta" family="Leiodidae" genus="Anemadus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="loebli">
<emphasis id="B926E136F81EFFB3FC20F81EFBBA4C86" box="[983,1047,1926,1951]" italics="true" pageId="7" pageNumber="8">loebli</emphasis>
</taxonomicName>
-group (e.g.
<taxonomicName id="4C5246A7F81EFFB3FB5FF81FFE844CDA" authority="Giachino &amp; Vailati, 1993" authorityName="Giachino &amp; Vailati" authorityYear="1993" class="Insecta" family="Leiodidae" genus="Anemadus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="loebli">
<emphasis id="B926E136F81EFFB3FB5FF81FFAA64C86" box="[1192,1291,1926,1951]" italics="true" pageId="7" pageNumber="8">A. loebli</emphasis>
<bibRefCitation id="EFC340D5F81EFFB3FAE2F81EFE844CDA" author="Giachino, P. M. &amp; Vailati, D." pageId="7" pageNumber="8" refId="ref13309" refString="Giachino, P. M. &amp; Vailati, D. (1993) Revisione degli Anemadinae. Monografie di &quot; Natura Bresciana &quot; No. 18. Museo Civico di Scienze Naturali di Brescia, Brescia, 314 pp." type="book" year="1993">Giachino &amp; Vailati, 1993</bibRefCitation>
</taxonomicName>
, from
<collectingCountry id="F3457DB4F81EFFB3FE84F832FE684CDA" box="[371,453,1962,1987]" name="Turkey" pageId="7" pageNumber="8">Turkey</collectingCountry>
) or the
<taxonomicName id="4C5246A7F81EFFB3FDEBF833FD2C4CDB" baseAuthorityName="Kraatz" baseAuthorityYear="1852" box="[540,641,1963,1986]" class="Insecta" family="Leiodidae" genus="Anemadus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="strigosus">
<emphasis id="B926E136F81EFFB3FDEBF833FD2C4CDB" box="[540,641,1963,1986]" italics="true" pageId="7" pageNumber="8">strigosus</emphasis>
</taxonomicName>
-group (e.g.
<taxonomicName id="4C5246A7F81EFFB3FCFBF833FB914CDA" authority="(Kraatz, 1852)" baseAuthorityName="Kraatz" baseAuthorityYear="1852" box="[780,1084,1962,1987]" class="Insecta" family="Leiodidae" genus="Anemadus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="strigosus">
<emphasis id="B926E136F81EFFB3FCFBF833FC3C4CDB" box="[780,913,1963,1986]" italics="true" pageId="7" pageNumber="8">A. strigosus</emphasis>
(Kraatz, 1852)
</taxonomicName>
from
<collectingCountry id="F3457DB4F81EFFB3FB75F832FA9A4CDA" box="[1154,1335,1962,1987]" name="Czech Republic" pageId="7" pageNumber="8">Czech Republic</collectingCountry>
,
<collectingCountry id="F3457DB4F81EFFB3FAB4F832FA354CDA" box="[1347,1432,1962,1987]" name="Austria" pageId="7" pageNumber="8">Austria</collectingCountry>
,
<collectingCountry id="F3457DB4F81EFFB3FF60F856FF664CFE" box="[151,203,1998,2023]" name="Italy" pageId="7" pageNumber="8">Italy</collectingCountry>
,
<collectingCountry id="F3457DB4F81EFFB3FF2FF856FEED4CFE" box="[216,320,1998,2023]" name="Romania" pageId="7" pageNumber="8">Romania</collectingCountry>
,
<collectingCountry id="F3457DB4F81EFFB3FEBAF856FE0F4CFE" box="[333,418,1998,2023]" name="Croatia" pageId="7" pageNumber="8">Croatia</collectingCountry>
,
<collectingCountry id="F3457DB4F81EFFB3FE58F856FD354CFE" box="[431,664,1998,2023]" name="Bosnia and Herzegovina" pageId="7" pageNumber="8">Bosnia-Herzegovina</collectingCountry>
,
<collectingCountry id="F3457DB4F81EFFB3FD52F857FC834CFE" box="[677,814,1999,2023]" name="Serbia" pageId="7" pageNumber="8">Montenegro</collectingCountry>
and
<collectingCountry id="F3457DB4F81EFFB3FC9FF857FC174CFE" box="[872,954,1999,2023]" name="Greece" pageId="7" pageNumber="8">Greece</collectingCountry>
,
<taxonomicName id="4C5246A7F81EFFB3FC30F857FB724CFE" authority="Reitter, 1906" authorityName="Reitter" authorityYear="1906" box="[967,1247,1998,2023]" class="Insecta" family="Leiodidae" genus="Anemadus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="bianchii">
<emphasis id="B926E136F81EFFB3FC30F857FBE94CFE" box="[967,1092,1998,2023]" italics="true" pageId="7" pageNumber="8">A. bianchii</emphasis>
Reitter, 1906
</taxonomicName>
from
<collectingCountry id="F3457DB4F81EFFB3FAD0F856FAF64CFE" box="[1319,1371,1998,2023]" name="Italy" pageId="7" pageNumber="8">Italy</collectingCountry>
or
<taxonomicName id="4C5246A7F81EFFBCFA73F857FE384BA9" authority="Reitter, 1885" authorityName="Reitter" authorityYear="1885" class="Insecta" family="Leiodidae" genus="Anemadus" higherTaxonomySource="GBIF" kingdom="Animalia" lastPageId="8" lastPageNumber="9" order="Coleoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="arcadius">
<emphasis id="B926E136F81EFFBCFA73F857FF554BA9" italics="true" lastPageId="8" lastPageNumber="9" pageId="7" pageNumber="8">A. arcadius</emphasis>
<bibRefCitation id="EFC340D5F811FFBCFEF6FF0FFE384BA9" author="Reitter, E." box="[257,405,151,176]" pageId="8" pageNumber="9" pagination="3 - 122" refId="ref14647" refString="Reitter, E. (1885) Bestimmungs-Tabellen der europaischen Coleopteren, 12 Necrophaga. Ferhandlungen des Naturforschenden Fereines in Brunn, 23 (1884), 3 - 122." type="journal article" year="1885">Reitter, 1885</bibRefCitation>
</taxonomicName>
from
<collectingCountry id="F3457DB4F811FFBCFE29FF00FD9E4BA9" box="[478,563,152,176]" name="Greece" pageId="8" pageNumber="9">Greece</collectingCountry>
). In addition to the morphological characters, the distribution of the species supports its inclusion in
<taxonomicName id="4C5246A7F811FFBCFE51FF24FDED4BCC" authorityName="Jeannel" authorityYear="1922" box="[422,576,188,213]" class="Insecta" family="Leiodidae" genus="Speonemadus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="genus">
<emphasis id="B926E136F811FFBCFE51FF24FDED4BCC" box="[422,576,188,213]" italics="true" pageId="8" pageNumber="9">Speonemadus</emphasis>
</taxonomicName>
, as species of
<taxonomicName id="4C5246A7F811FFBCFD17FF24FCF84BCC" authorityName="Reitter" authorityYear="1885" box="[736,853,188,213]" class="Insecta" family="Leiodidae" genus="Anemadus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="genus">
<emphasis id="B926E136F811FFBCFD17FF24FCF84BCC" box="[736,853,188,213]" italics="true" pageId="8" pageNumber="9">Anemadus</emphasis>
</taxonomicName>
are found from
<collectingCountry id="F3457DB4F811FFBCFBFFFF24FB964BCC" box="[1032,1083,188,213]" name="Italy" pageId="8" pageNumber="9">Italy</collectingCountry>
(including
<collectingRegion id="4996F3C6F811FFBCFB4BFF24FB524BCC" box="[1212,1279,188,213]" country="Italy" name="Sicilia" pageId="8" pageNumber="9">Sicily</collectingRegion>
) and southern
<collectingCountry id="F3457DB4F811FFBCFF60FF78FF4E4BEE" box="[151,227,224,247]" name="France" pageId="8" pageNumber="9">France</collectingCountry>
to the East, reaching
<collectingCountry id="F3457DB4F811FFBCFE20FF47FDB74BE1" box="[471,538,223,248]" name="China" pageId="8" pageNumber="9">China</collectingCountry>
(
<bibRefCitation id="EFC340D5F811FFBCFDDEFF78FD694BE1" author="Perreau, M." box="[553,708,224,248]" pageId="8" pageNumber="9" pagination="133 - 203" refId="ref14426" refString="Perreau, M. (2004) Leiodidae. In: Lobl, I. &amp; Smetana, A. (Eds.), Catalogue of Palaearctic Coleoptera. Fol. 2. Apollo Books, Stenstrup, pp. 133 - 203." type="book chapter" year="2004">Perreau 2004</bibRefCitation>
). In fact, this is one of the westernmost species of
<taxonomicName id="4C5246A7F811FFBCFB0CFF47FA384BE1" authorityName="Jeannel" authorityYear="1922" box="[1275,1429,223,248]" class="Insecta" family="Leiodidae" genus="Speonemadus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="genus">
<emphasis id="B926E136F811FFBCFB0CFF47FA384BE1" box="[1275,1429,223,248]" italics="true" pageId="8" pageNumber="9">Speonemadus</emphasis>
</taxonomicName>
, with other species of the genus distributed further east except for some dubious records of
<taxonomicName id="4C5246A7F811FFBCFB6DFE9DFAB94A05" baseAuthorityName="Peyerimhoff" baseAuthorityYear="1917" box="[1178,1300,261,284]" class="Insecta" family="Leiodidae" genus="Anemadus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="tenuipes">
<emphasis id="B926E136F811FFBCFB6DFE9DFAB94A05" box="[1178,1300,261,284]" italics="true" pageId="8" pageNumber="9">S. tenuipes</emphasis>
</taxonomicName>
in the High Atlas (see below).
</paragraph>
</subSubSection>
<subSubSection id="C3486EAFF811FFBCFF30FED4FB844A7D" box="[199,1065,332,357]" pageId="8" pageNumber="9" type="distribution">
<paragraph id="8BED3D24F811FFBCFF30FED4FB844A7D" blockId="8.[151,1436,151,357]" box="[199,1065,332,357]" pageId="8" pageNumber="9">
<emphasis id="B926E136F811FFBCFF30FED4FEF44A7C" bold="true" box="[199,345,332,357]" pageId="8" pageNumber="9">Distribution</emphasis>
(
<figureCitation id="136921A1F811FFBCFE9FFED4FE114A7D" box="[360,444,332,357]" captionStart="FIGURE 38" captionStartId="21.[151,250,1234,1256]" captionTargetBox="[259,1364,471,1140]" captionTargetId="figure@21.[205,1395,421,1189]" captionTargetPageId="21" captionText="FIGURE 38. Distribution maps of: (38) Speonemadus brusteli sp. n., S. comasi sp. n., S. orchesioides, S. pulchellus, S. tenuipes (empty circles are dubious records of which no material could be studied; the records from continental Italy, given originally as S. orchesoides, are considered S. pulchellus, P.M. Giachino, personal communication 2018; see main text). See Appendix for detailed locality data." httpUri="https://zenodo.org/record/2617639/files/figure.png" pageId="8" pageNumber="9">Fig. 38</figureCitation>
). So far only known from the
<typeStatus id="54E98386F811FFBCFCFAFED5FC904A7C" box="[781,829,333,357]" pageId="8" pageNumber="9">type</typeStatus>
locality in
<collectingCountry id="F3457DB4F811FFBCFC4BFED5FB884A7D" box="[956,1061,333,356]" name="Morocco" pageId="8" pageNumber="9">Morocco</collectingCountry>
.
</paragraph>
</subSubSection>
</treatment>
</document>