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<mods:title id="137E8AA62831B43F22B6F62477D45DE0">Revision and Phylogeny of the Genus Phausis (Coleoptera: Lampyridae) with the Description of Three New Species</mods:title>
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<mods:namePart id="62FBE755D2238E1BE56151B3F4153B8F">Branham, Marc A.</mods:namePart>
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<subSubSection id="C36F18AEFFBEFFD57DB255C856C9F901" box="[151,648,1672,1699]" pageId="16" pageNumber="17" type="nomenclature">
<paragraph id="8BCA4B25FFBEFFD57DB255C856C9F901" blockId="16.[151,648,1672,1735]" box="[151,648,1672,1699]" pageId="16" pageNumber="17">
<heading id="D082FC49FFBEFFD57DB255C856C9F901" bold="true" box="[151,648,1672,1699]" fontSize="11" level="1" pageId="16" pageNumber="17" reason="1">
<emphasis id="B9019737FFBEFFD57DB255C856C9F901" bold="true" box="[151,648,1672,1699]" pageId="16" pageNumber="17">
<taxonomicName id="4C7530A6FFBEFFD57DB255C8558AF900" ID-CoL="4FXJG" authority="Fender" authorityName="Fender" authorityYear="1962" box="[151,459,1672,1699]" class="Insecta" family="Lampyridae" genus="Phausis" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="dorothae">
<emphasis id="B9019737FFBEFFD57DB255C8552AF900" bold="true" box="[151,363,1672,1699]" italics="true" pageId="16" pageNumber="17">Phausis dorothae</emphasis>
Fender
</taxonomicName>
in Hatch, 1962
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C36F18AEFFBEFFD57DB255EC55BFF964" box="[151,510,1708,1735]" pageId="16" pageNumber="17" type="description">
<paragraph id="8BCA4B25FFBEFFD57DB255EC55BFF964" blockId="16.[151,648,1672,1735]" box="[151,510,1708,1735]" pageId="16" pageNumber="17">
(
<figureCitation id="134E57A0FFBEFFD57DBA55EC54A9F964" box="[159,232,1708,1735]" captionStart="FIGURE 7" captionStartId="18.[152,255,1491,1515]" captionTargetBox="[582,1006,561,1468]" captionTargetId="figure-170@18.[582,1006,561,1468]" captionTargetPageId="18" captionText="FIGURE 7. Detail of North American distribution of Phausis dorothae." figureDoi="http://doi.org/10.5281/zenodo.11373261" httpUri="https://zenodo.org/record/11373261/files/figure.png" pageId="16" pageNumber="17">Figs 7</figureCitation>
;
<figureCitation id="134E57A0FFBEFFD57DD055ED5567F964" box="[245,294,1709,1735]" captionStart="FIGURE 17" captionStartId="42.[152,255,1550,1574]" captionTargetBox="[169,1418,181,1526]" captionTargetId="figure-28@42.[169,1418,181,1526]" captionTargetPageId="42" captionText="FIGURE 17. Dorsal habitus of male Phausis (AF). (A) Phausis branhami sp. nov.; (B). Phausis californica; (C) Phausis dorothae; (D) Phausis inaccensa; (E) Phausis luminosa; (F) Phausis marina." figureDoi="http://doi.org/10.5281/zenodo.11348305" httpUri="https://zenodo.org/record/11348305/files/figure.png" pageId="16" pageNumber="17">17C</figureCitation>
;
<figureCitation id="134E57A0FFBEFFD57C1755ED553CF964" box="[306,381,1709,1735]" captionStart="FIGURE 22" captionStartId="46.[152,255,1881,1905]" captionTargetBox="[170,1418,309,1869]" captionTargetId="figure-84@46.[170,1418,309,1869]" captionTargetPageId="46" captionText="FIGURE 22. Phausis aedeagi (AJj) (from left to right: dorsal, lateral, ventral view). (AC) Phausis branhami sp. nov.; (DF) Phausis californica; (GI) Phausis dorothae; (JL) Phausis inaccensa; (MO) Phausis luminosa; (PR) Phausis marina; (SU) Phausis milleri sp. nov.; (VX) Phausis nigra; (YAa) Phausis reticulata; (BbDd) Phausis rhombica; (EeGg) Phausis riversi; (HhJj) Phausis skelleyi." figureDoi="http://doi.org/10.5281/zenodo.11348323" httpUri="https://zenodo.org/record/11348323/files/figure.png" pageId="16" pageNumber="17">22GI</figureCitation>
;
<figureCitation id="134E57A0FFBEFFD57CAD55ED55FBF964" box="[392,442,1709,1735]" captionStart="FIGURE 23" captionStartId="47.[152,255,1365,1389]" captionTargetBox="[346,1242,181,1341]" captionTargetId="figure-17@47.[346,1242,181,1341]" captionTargetPageId="47" captionText="FIGURE 23. Sensilla of Phausis aedeagi (AD). (A) Phausis luminosa (medial lobe, ventral view); (B) Phausis dorothae (paramere, dorsal view); (C) Phausis riversi (paramere, dorsal view); (D) Phausis marina (paramere, lateral view). Not pictured is Phausis californica which also has sensilla on the apical tip of the parameres (holotype, dorsal view)." figureDoi="http://doi.org/10.5281/zenodo.11348325" httpUri="https://zenodo.org/record/11348325/files/figure.png" pageId="16" pageNumber="17">23B</figureCitation>
;
<figureCitation id="134E57A0FFBEFFD57CE355ED55B7F964" box="[454,502,1709,1735]" captionStart="FIGURE 24" captionStartId="48.[152,255,1943,1967]" captionTargetBox="[234,1354,181,1919]" captionTargetId="figure-28@48.[234,1354,181,1919]" captionTargetPageId="48" captionText="FIGURE 24. Type labels of Phausis (AL). (A) Phausis branhami sp. nov.; (B). Phausis californica; (C) Phausis dorothae; (D) Phausis inaccensa; (E) Phausis luminosa; (F) Phausis marina; (G) Phausis milleri sp. nov.; (H) Phausis nigra; (I) Phausis reticulata; (J) Phausis rhombica; (K) Phausis riversi; (L) Phausis skelleyi." figureDoi="http://doi.org/10.5281/zenodo.11348327" httpUri="https://zenodo.org/record/11348327/files/figure.png" pageId="16" pageNumber="17">24C</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C36F18AEFFBEFFD57DB255AD55B5F8A6" box="[151,500,1773,1797]" pageId="16" pageNumber="17" type="reference_group">
<paragraph id="8BCA4B25FFBEFFD57DB255AD55B5F8A6" blockId="16.[151,500,1773,1797]" box="[151,500,1773,1797]" pageId="16" pageNumber="17">
<treatmentCitationGroup id="AB656C0BFFBEFFD57DB255AD55B5F8A6" box="[151,500,1773,1797]" pageId="16" pageNumber="17">
<treatmentCitation id="0AD46D34FFBEFFD57DB255AD55B5F8A6" author="Fender, K. M." box="[151,500,1773,1797]" page="42" pageId="16" pageNumber="17" year="1962">
<taxonomicName id="4C7530A6FFBEFFD57DB255AD55B5F8A6" ID-CoL="4FXJG" authority="Fender 1962: 42" authorityName="Fender" authorityPageNumber="42" authorityYear="1962" box="[151,500,1773,1797]" class="Insecta" family="Lampyridae" genus="Phausis" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="dorothae">
<emphasis id="B9019737FFBEFFD57DB255AD5508F8A6" box="[151,329,1773,1797]" italics="true" pageId="16" pageNumber="17">Phausis dorothae</emphasis>
<bibRefCitation id="EFE436D4FFBEFFD57C6A55AD55B5F8A6" author="Fender, K. M." box="[335,500,1773,1797]" pageId="16" pageNumber="17" refId="ref39267" refString="Fender, K. M. (1962) Lampyridae. In: Hatch, M. H. (Ed.), The Beetles of the Pacific Northwest. Vol. 16. University of Washington Press, Seattle, Washington, pp. 41 - 43." type="book" year="1962">Fender 1962: 42</bibRefCitation>
</taxonomicName>
</treatmentCitation>
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C36F18AEFFBEFFD47DB2546F558FF9AA" lastPageId="17" lastPageNumber="18" pageId="16" pageNumber="17" type="diagnosis">
<paragraph id="8BCA4B25FFBEFFD57DB2546F57BCF817" blockId="16.[151,1437,1839,2043]" pageId="16" pageNumber="17">
Diagnosis.
<emphasis id="B9019737FFBEFFD57C33546F5512F8EA" bold="true" box="[278,339,1839,1865]" pageId="16" pageNumber="17">Male</emphasis>
.
<taxonomicName id="4C7530A6FFBEFFD57C47546F5667F8EA" authorityName="Fender" authorityYear="1962" box="[354,550,1839,1865]" class="Insecta" family="Lampyridae" genus="Phausis" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="dorothae">
<emphasis id="B9019737FFBEFFD57C47546F5667F8EA" box="[354,550,1839,1865]" italics="true" pageId="16" pageNumber="17">Phausis dorothae</emphasis>
</taxonomicName>
differs from its congeners by the following characters: short, simple, plumose setae covering base of mandibles; anterior vitreous areas (pronotal spots) are absent on the pronotum (
<figureCitation id="134E57A0FFBEFFD57802541251CEF8CF" box="[1319,1423,1874,1900]" captionStart="FIGURE 17" captionStartId="42.[152,255,1550,1574]" captionTargetBox="[169,1418,181,1526]" captionTargetId="figure-28@42.[169,1418,181,1526]" captionTargetPageId="42" captionText="FIGURE 17. Dorsal habitus of male Phausis (AF). (A) Phausis branhami sp. nov.; (B). Phausis californica; (C) Phausis dorothae; (D) Phausis inaccensa; (E) Phausis luminosa; (F) Phausis marina." figureDoi="http://doi.org/10.5281/zenodo.11348305" httpUri="https://zenodo.org/record/11348305/files/figure.png" pageId="16" pageNumber="17">Fig. 17C</figureCitation>
); central pronotal disc absent; vertex of head not concave as in other species of
<taxonomicName id="4C7530A6FFBEFFD57ECF54355002F82C" authorityName="LeConte" authorityYear="1852" box="[1002,1091,1909,1935]" class="Insecta" family="Lampyridae" genus="Phausis" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="genus">
<emphasis id="B9019737FFBEFFD57ECF54355002F82C" box="[1002,1091,1909,1935]" italics="true" pageId="16" pageNumber="17">Phausis</emphasis>
</taxonomicName>
. Labial palps long; Metathorax without posterior apex notched; medial line absent; surface sparsely punctate.
</paragraph>
<paragraph id="8BCA4B25FFBEFFD47DE254FC54A7FEE2" blockId="16.[151,1437,1839,2043]" lastBlockId="17.[151,1437,151,2013]" lastPageId="17" lastPageNumber="18" pageId="16" pageNumber="17">
Redescription.
<emphasis id="B9019737FFBEFFD57C4A54FD55EDF874" bold="true" box="[367,428,1981,2007]" pageId="16" pageNumber="17">Male</emphasis>
(
<figureCitation id="134E57A0FFBEFFD57C9F54FC566BF874" box="[442,554,1980,2007]" captionStart="FIGURE 17" captionStartId="42.[152,255,1550,1574]" captionTargetBox="[169,1418,181,1526]" captionTargetId="figure-28@42.[169,1418,181,1526]" captionTargetPageId="42" captionText="FIGURE 17. Dorsal habitus of male Phausis (AF). (A) Phausis branhami sp. nov.; (B). Phausis californica; (C) Phausis dorothae; (D) Phausis inaccensa; (E) Phausis luminosa; (F) Phausis marina." figureDoi="http://doi.org/10.5281/zenodo.11348305" httpUri="https://zenodo.org/record/11348305/files/figure.png" pageId="16" pageNumber="17">Figs. 17C</figureCitation>
;
<figureCitation id="134E57A0FFBEFFD57F1154FD563EF874" box="[564,639,1981,2007]" captionStart="FIGURE 22" captionStartId="46.[152,255,1881,1905]" captionTargetBox="[170,1418,309,1869]" captionTargetId="figure-84@46.[170,1418,309,1869]" captionTargetPageId="46" captionText="FIGURE 22. Phausis aedeagi (AJj) (from left to right: dorsal, lateral, ventral view). (AC) Phausis branhami sp. nov.; (DF) Phausis californica; (GI) Phausis dorothae; (JL) Phausis inaccensa; (MO) Phausis luminosa; (PR) Phausis marina; (SU) Phausis milleri sp. nov.; (VX) Phausis nigra; (YAa) Phausis reticulata; (BbDd) Phausis rhombica; (EeGg) Phausis riversi; (HhJj) Phausis skelleyi." figureDoi="http://doi.org/10.5281/zenodo.11348323" httpUri="https://zenodo.org/record/11348323/files/figure.png" pageId="16" pageNumber="17">22GI</figureCitation>
;
<figureCitation id="134E57A0FFBEFFD57FAC54FD56FDF874" box="[649,700,1981,2007]" captionStart="FIGURE 23" captionStartId="47.[152,255,1365,1389]" captionTargetBox="[346,1242,181,1341]" captionTargetId="figure-17@47.[346,1242,181,1341]" captionTargetPageId="47" captionText="FIGURE 23. Sensilla of Phausis aedeagi (AD). (A) Phausis luminosa (medial lobe, ventral view); (B) Phausis dorothae (paramere, dorsal view); (C) Phausis riversi (paramere, dorsal view); (D) Phausis marina (paramere, lateral view). Not pictured is Phausis californica which also has sensilla on the apical tip of the parameres (holotype, dorsal view)." figureDoi="http://doi.org/10.5281/zenodo.11348325" httpUri="https://zenodo.org/record/11348325/files/figure.png" pageId="16" pageNumber="17">23B</figureCitation>
). Length 5.56.0 mm. Width 2.0
<quantity id="4C8DE6C0FFBEFFD5791554FC50C6F874" box="[1072,1159,1980,2007]" metricMagnitude="-3" metricUnit="m" metricValue="2.5" pageId="16" pageNumber="17" unit="mm" value="2.5">2.5 mm</quantity>
.
<emphasis id="B9019737FFBEFFD579B554FC5097F875" box="[1168,1238,1980,2006]" italics="true" pageId="16" pageNumber="17">Color.</emphasis>
Body dark brown to chestnut brown; eyes black; mouthparts flavus to brown; antennae brown; pronotum generally ochre-yellow with a dark brown central spot(brown central spot sometimes absent); scutellum pale brown to flavus; thorax overall brunneous; legs brownish-yellow to brunneous depending on specimen examined; lateral margins of elytra paler than elytral disc; photic organs absent on abdominal segments VI VII and VIII; abdominal segment VI flavus along lateral and posterior margin, brunneous medially; pubescence cinereous. Anterior vitreous areas (pronotal spots) absent.
</paragraph>
<paragraph id="8BCA4B25FFBFFFD47DE2520B56B1FDBA" blockId="17.[151,1437,151,2013]" pageId="17" pageNumber="18">
<emphasis id="B9019737FFBFFFD47DE2520B5545FEC6" box="[199,260,331,357]" italics="true" pageId="17" pageNumber="18">Head</emphasis>
small and transverse (frontal view); vertex not concave, with fine close punctures dorsally. Eyes small and separated, not partially obstructing labium postventrally. Mandibles short (apices of mandibles not crossing frontally), robust, tapering abruptly, beginning at base; incisor region elongate (greater than half of total length of mandible); outer margin convex versus straight as in other taxa (
<taxonomicName id="4C7530A6FFBFFFD47EB452F757BFFE73" authorityName="Fender" authorityYear="1966" box="[913,1022,439,464]" class="Insecta" family="Lampyridae" genus="Phausis" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="marina">
<emphasis id="B9019737FFBFFFD47EB452F757BFFE73" box="[913,1022,439,464]" italics="true" pageId="17" pageNumber="18">P. marina</emphasis>
</taxonomicName>
,
<emphasis id="B9019737FFBFFFD4792852F750A4FE72" box="[1037,1253,439,465]" italics="true" pageId="17" pageNumber="18">
<taxonomicName id="4C7530A6FFBFFFD4792852F75029FE73" authorityName="Hopping, Det. Ralph Hopping" authorityYear="1937" box="[1037,1128,439,464]" class="Insecta" family="Lampyridae" genus="Phausis" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="nigra">P. nigra</taxonomicName>
,
<taxonomicName id="4C7530A6FFBFFFD4795052F750A4FE72" box="[1141,1253,439,465]" class="Insecta" family="Lampyridae" genus="Phausis" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="skelleyi">P. skelleyi</taxonomicName>
</emphasis>
, and
<taxonomicName id="4C7530A6FFBFFFD4780152F751CAFE72" authorityName="Hodson &amp; Branham" authorityYear="2024" box="[1316,1419,439,465]" class="Insecta" family="Lampyridae" genus="Phausis" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="milleri">
<emphasis id="B9019737FFBFFFD4780152F751CAFE72" box="[1316,1419,439,465]" italics="true" pageId="17" pageNumber="18">P. milleri</emphasis>
</taxonomicName>
); basal angle approximately less than 90 degrees dorsally. Maxillary palps with three palpomeres, last palpomere securiform; Labial palps long, with three palpomeres.
</paragraph>
<paragraph id="8BCA4B25FFBFFFD47DE251635156FD6E" blockId="17.[151,1437,151,2013]" pageId="17" pageNumber="18">
<emphasis id="B9019737FFBFFFD47DE251635570FD9E" box="[199,305,547,573]" italics="true" pageId="17" pageNumber="18">Antennae</emphasis>
filiform, long, extending to middle of metathorax (
<figureCitation id="134E57A0FFBFFFD47EB7516357BCFD9E" box="[914,1021,547,573]" captionStart="FIGURE 17" captionStartId="42.[152,255,1550,1574]" captionTargetBox="[169,1418,181,1526]" captionTargetId="figure-28@42.[169,1418,181,1526]" captionTargetPageId="42" captionText="FIGURE 17. Dorsal habitus of male Phausis (AF). (A) Phausis branhami sp. nov.; (B). Phausis californica; (C) Phausis dorothae; (D) Phausis inaccensa; (E) Phausis luminosa; (F) Phausis marina." figureDoi="http://doi.org/10.5281/zenodo.11348305" httpUri="https://zenodo.org/record/11348305/files/figure.png" pageId="17" pageNumber="18">Fig. 17C</figureCitation>
). Scape as long as pedicel. Pedicel approximately 1/2 the length of antennomere 3. Antennomere 3 as long as antennomere 4. Antennomeres 5 to 10 shorter than 3. Antennomere 11 slightly longer, rounded at apex. Flagellomeres feebly compressed, but not as compressed as in
<taxonomicName id="4C7530A6FFBFFFD47CA151CF55A9FD0A" authorityName="Laporte" authorityYear="1833" box="[388,488,655,681]" class="Insecta" family="Lampyridae" genus="Lucidota" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="genus">
<emphasis id="B9019737FFBFFFD47CA151CF55A9FD0A" box="[388,488,655,681]" italics="true" pageId="17" pageNumber="18">Lucidota</emphasis>
</taxonomicName>
or
<taxonomicName id="4C7530A6FFBFFFD47F3451CF563DFD0A" box="[529,636,655,681]" class="Insecta" family="Lampyridae" genus="Ellychnia" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="genus">
<emphasis id="B9019737FFBFFFD47F3451CF563DFD0A" box="[529,636,655,681]" italics="true" pageId="17" pageNumber="18">Ellychnia</emphasis>
</taxonomicName>
. When viewed laterally, flagellomeres are wider than in ventral view. Sensorium present on the distal end of antennomere 11, semi-transparent, small, and truncate-subulate.
</paragraph>
<paragraph id="8BCA4B25FFBFFFD47DE251975552FC22" blockId="17.[151,1437,151,2013]" pageId="17" pageNumber="18">
<emphasis id="B9019737FFBFFFD47DE251975577FD53" box="[199,310,727,752]" italics="true" pageId="17" pageNumber="18">Pronotum</emphasis>
subquadrate (almost as long as wide), subelliptical, and coarsely punctate (
<figureCitation id="134E57A0FFBFFFD479A6519750ABFD52" box="[1155,1258,727,753]" captionStart="FIGURE 17" captionStartId="42.[152,255,1550,1574]" captionTargetBox="[169,1418,181,1526]" captionTargetId="figure-28@42.[169,1418,181,1526]" captionTargetPageId="42" captionText="FIGURE 17. Dorsal habitus of male Phausis (AF). (A) Phausis branhami sp. nov.; (B). Phausis californica; (C) Phausis dorothae; (D) Phausis inaccensa; (E) Phausis luminosa; (F) Phausis marina." figureDoi="http://doi.org/10.5281/zenodo.11348305" httpUri="https://zenodo.org/record/11348305/files/figure.png" pageId="17" pageNumber="18">Fig. 17C</figureCitation>
); median carina absent; all margins strongly reflexed, except less so along anterior and posterior margins, becoming explanate toward the hind angles; basal margin undulate; hind angles rounded. Central pronotal disc present; quadrate; shiny and finely punctate medially, becoming dull and coarsely punctate basally. Anterior vitreous areas (spots) absent (
<figureCitation id="134E57A0FFBFFFD47DBA50275547FC22" box="[159,262,871,897]" captionStart="FIGURE 17" captionStartId="42.[152,255,1550,1574]" captionTargetBox="[169,1418,181,1526]" captionTargetId="figure-28@42.[169,1418,181,1526]" captionTargetPageId="42" captionText="FIGURE 17. Dorsal habitus of male Phausis (AF). (A) Phausis branhami sp. nov.; (B). Phausis californica; (C) Phausis dorothae; (D) Phausis inaccensa; (E) Phausis luminosa; (F) Phausis marina." figureDoi="http://doi.org/10.5281/zenodo.11348305" httpUri="https://zenodo.org/record/11348305/files/figure.png" pageId="17" pageNumber="18">Fig. 17C</figureCitation>
).
</paragraph>
<paragraph id="8BCA4B25FFBFFFD47DE250CB5172FC06" blockId="17.[151,1437,151,2013]" box="[199,1331,907,933]" pageId="17" pageNumber="18">
<taxonomicName id="4C7530A6FFBFFFD47DE250CB5574FC06" box="[199,309,907,933]" class="Trilobita" family="Styginidae" genus="Scutellum" kingdom="Animalia" order="Corynexochida" pageId="17" pageNumber="18" phylum="Arthropoda" rank="genus">
<emphasis id="B9019737FFBFFFD47DE250CB5574FC06" box="[199,309,907,933]" italics="true" pageId="17" pageNumber="18">Scutellum</emphasis>
</taxonomicName>
spatulate, sides curving sinuate apex truncate (
<figureCitation id="134E57A0FFBFFFD47E6050CB57EDFC06" box="[837,940,907,933]" captionStart="FIGURE 17" captionStartId="42.[152,255,1550,1574]" captionTargetBox="[169,1418,181,1526]" captionTargetId="figure-28@42.[169,1418,181,1526]" captionTargetPageId="42" captionText="FIGURE 17. Dorsal habitus of male Phausis (AF). (A) Phausis branhami sp. nov.; (B). Phausis californica; (C) Phausis dorothae; (D) Phausis inaccensa; (E) Phausis luminosa; (F) Phausis marina." figureDoi="http://doi.org/10.5281/zenodo.11348305" httpUri="https://zenodo.org/record/11348305/files/figure.png" pageId="17" pageNumber="18">Fig. 17C</figureCitation>
); surface dull and finely reticulate.
</paragraph>
<paragraph id="8BCA4B25FFBFFFD47DE250EF51C0FBB2" blockId="17.[151,1437,151,2013]" pageId="17" pageNumber="18">
<emphasis id="B9019737FFBFFFD47DE250EF554CFC6A" box="[199,269,943,969]" italics="true" pageId="17" pageNumber="18">Elytra</emphasis>
well developed; moderately punctate to reticulate, punctate apically, becoming more finely reticulate basally (
<figureCitation id="134E57A0FFBFFFD47DD05093551DFC4E" box="[245,348,979,1005]" captionStart="FIGURE 17" captionStartId="42.[152,255,1550,1574]" captionTargetBox="[169,1418,181,1526]" captionTargetId="figure-28@42.[169,1418,181,1526]" captionTargetPageId="42" captionText="FIGURE 17. Dorsal habitus of male Phausis (AF). (A) Phausis branhami sp. nov.; (B). Phausis californica; (C) Phausis dorothae; (D) Phausis inaccensa; (E) Phausis luminosa; (F) Phausis marina." figureDoi="http://doi.org/10.5281/zenodo.11348305" httpUri="https://zenodo.org/record/11348305/files/figure.png" pageId="17" pageNumber="18">Fig. 17C</figureCitation>
). Elytral costae absent on each elytron. Longitudinal elevated lines weakly present and transverse lines present, transverse lines somewhat irregular in the interstitial spaces giving elytra a reticulated appearance.
</paragraph>
<paragraph id="8BCA4B25FFBFFFD47DE2575B50ECFB02" blockId="17.[151,1437,151,2013]" pageId="17" pageNumber="18">
<emphasis id="B9019737FFBFFFD47DE2575B5552FB97" box="[199,275,1051,1076]" italics="true" pageId="17" pageNumber="18">Venter.</emphasis>
Thorax elongate, narrowly rhombus. Metathorax without posterior apex notched; medial line absent; surface finely sparsely punctate. Tergal apicolateral angles strongly lobed. Abdominal segments VI and VII without photic organs. Photic organ absent on abdominal segment VIII. Abdominal segment VII elongate (longer than wide), apical margin emarginated. Abdominal segment VIII elongate, ovate, rounded at apex.
</paragraph>
<paragraph id="8BCA4B25FFBFFFD47DE257EB558FF9AA" blockId="17.[151,1437,151,2013]" pageId="17" pageNumber="18">
<emphasis id="B9019737FFBFFFD47DE257EB5538FB66" box="[199,377,1195,1221]" italics="true" pageId="17" pageNumber="18">Male Genitalia.</emphasis>
Basal piece (dorsal view) wider than long, quadrate, segments approximate (almost touching), and moderately sclerotized (light brown) (
<figureCitation id="134E57A0FFBFFFD47F55578F5698FB4A" box="[624,729,1231,1257]" captionStart="FIGURE 22" captionStartId="46.[152,255,1881,1905]" captionTargetBox="[170,1418,309,1869]" captionTargetId="figure-84@46.[170,1418,309,1869]" captionTargetPageId="46" captionText="FIGURE 22. Phausis aedeagi (AJj) (from left to right: dorsal, lateral, ventral view). (AC) Phausis branhami sp. nov.; (DF) Phausis californica; (GI) Phausis dorothae; (JL) Phausis inaccensa; (MO) Phausis luminosa; (PR) Phausis marina; (SU) Phausis milleri sp. nov.; (VX) Phausis nigra; (YAa) Phausis reticulata; (BbDd) Phausis rhombica; (EeGg) Phausis riversi; (HhJj) Phausis skelleyi." figureDoi="http://doi.org/10.5281/zenodo.11348323" httpUri="https://zenodo.org/record/11348323/files/figure.png" pageId="17" pageNumber="18">Fig. 22G</figureCitation>
). Median lobe (dorsal view) does not exceed tips of parameres, elongate and acutely rounded (less than 45 degrees) at apex; extending forward ventrally with respect to parameres (
<figureCitation id="134E57A0FFBFFFD47DBA5657554AFA92" box="[159,267,1303,1329]" captionStart="FIGURE 22" captionStartId="46.[152,255,1881,1905]" captionTargetBox="[170,1418,309,1869]" captionTargetId="figure-84@46.[170,1418,309,1869]" captionTargetPageId="46" captionText="FIGURE 22. Phausis aedeagi (AJj) (from left to right: dorsal, lateral, ventral view). (AC) Phausis branhami sp. nov.; (DF) Phausis californica; (GI) Phausis dorothae; (JL) Phausis inaccensa; (MO) Phausis luminosa; (PR) Phausis marina; (SU) Phausis milleri sp. nov.; (VX) Phausis nigra; (YAa) Phausis reticulata; (BbDd) Phausis rhombica; (EeGg) Phausis riversi; (HhJj) Phausis skelleyi." figureDoi="http://doi.org/10.5281/zenodo.11348323" httpUri="https://zenodo.org/record/11348323/files/figure.png" pageId="17" pageNumber="18">Fig. 22H</figureCitation>
); sensilla absent on apex; dorsal inner-basal margins truncate; dorsal opening present, triangular with acuminate apices; longitudinal dorsal carina absent; lateral line absent (lateral view); ventral opening absent. Parameres (dorsal view;
<figureCitation id="134E57A0FFBFFFD47C9E561F5664FADA" box="[443,549,1375,1401]" captionStart="FIGURE 22" captionStartId="46.[152,255,1881,1905]" captionTargetBox="[170,1418,309,1869]" captionTargetId="figure-84@46.[170,1418,309,1869]" captionTargetPageId="46" captionText="FIGURE 22. Phausis aedeagi (AJj) (from left to right: dorsal, lateral, ventral view). (AC) Phausis branhami sp. nov.; (DF) Phausis californica; (GI) Phausis dorothae; (JL) Phausis inaccensa; (MO) Phausis luminosa; (PR) Phausis marina; (SU) Phausis milleri sp. nov.; (VX) Phausis nigra; (YAa) Phausis reticulata; (BbDd) Phausis rhombica; (EeGg) Phausis riversi; (HhJj) Phausis skelleyi." figureDoi="http://doi.org/10.5281/zenodo.11348323" httpUri="https://zenodo.org/record/11348323/files/figure.png" pageId="17" pageNumber="18">Fig. 22G</figureCitation>
) elongate, slender, sub-parallel; inner lateral margins concave; outer lateral margins broadly convex; apex roundly acute (lateral view;
<figureCitation id="134E57A0FFBFFFD47E1056C357DCFA3E" box="[821,925,1411,1437]" captionStart="FIGURE 22" captionStartId="46.[152,255,1881,1905]" captionTargetBox="[170,1418,309,1869]" captionTargetId="figure-84@46.[170,1418,309,1869]" captionTargetPageId="46" captionText="FIGURE 22. Phausis aedeagi (AJj) (from left to right: dorsal, lateral, ventral view). (AC) Phausis branhami sp. nov.; (DF) Phausis californica; (GI) Phausis dorothae; (JL) Phausis inaccensa; (MO) Phausis luminosa; (PR) Phausis marina; (SU) Phausis milleri sp. nov.; (VX) Phausis nigra; (YAa) Phausis reticulata; (BbDd) Phausis rhombica; (EeGg) Phausis riversi; (HhJj) Phausis skelleyi." figureDoi="http://doi.org/10.5281/zenodo.11348323" httpUri="https://zenodo.org/record/11348323/files/figure.png" pageId="17" pageNumber="18">Fig. 22H</figureCitation>
)); proximal flanges (extensions of parameres that curve or extend outward from inner margin) absent (dorsal view). Sensilla present, sparse to abundant along proximal apical margins (dorsal view). Number of sensilla vary from 1 to greater than 10, if absent then a pore can be seen (
<figureCitation id="134E57A0FFBFFFD47DDF56AF55C5F9AA" box="[250,388,1519,1545]" captionStart="FIGURE 22" captionStartId="46.[152,255,1881,1905]" captionTargetBox="[170,1418,309,1869]" captionTargetId="figure-84@46.[170,1418,309,1869]" captionTargetPageId="46" captionText="FIGURE 22. Phausis aedeagi (AJj) (from left to right: dorsal, lateral, ventral view). (AC) Phausis branhami sp. nov.; (DF) Phausis californica; (GI) Phausis dorothae; (JL) Phausis inaccensa; (MO) Phausis luminosa; (PR) Phausis marina; (SU) Phausis milleri sp. nov.; (VX) Phausis nigra; (YAa) Phausis reticulata; (BbDd) Phausis rhombica; (EeGg) Phausis riversi; (HhJj) Phausis skelleyi." figureDoi="http://doi.org/10.5281/zenodo.11348323" httpUri="https://zenodo.org/record/11348323/files/figure.png" pageId="17" pageNumber="18">Figs. 22GI</figureCitation>
;
<figureCitation id="134E57A0FFBFFFD47CAA56AF5583F9AA" box="[399,450,1519,1545]" captionStart="FIGURE 23" captionStartId="47.[152,255,1365,1389]" captionTargetBox="[346,1242,181,1341]" captionTargetId="figure-17@47.[346,1242,181,1341]" captionTargetPageId="47" captionText="FIGURE 23. Sensilla of Phausis aedeagi (AD). (A) Phausis luminosa (medial lobe, ventral view); (B) Phausis dorothae (paramere, dorsal view); (C) Phausis riversi (paramere, dorsal view); (D) Phausis marina (paramere, lateral view). Not pictured is Phausis californica which also has sensilla on the apical tip of the parameres (holotype, dorsal view)." figureDoi="http://doi.org/10.5281/zenodo.11348325" httpUri="https://zenodo.org/record/11348325/files/figure.png" pageId="17" pageNumber="18">23B</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C36F18AEFFBFFFD77DE2555355E5FDBA" lastPageId="18" lastPageNumber="19" pageId="17" pageNumber="18" type="description">
<paragraph id="8BCA4B25FFBFFFD47DE2555355E3F98E" blockId="17.[151,1437,151,2013]" box="[199,418,1555,1581]" pageId="17" pageNumber="18">
<emphasis id="B9019737FFBFFFD47DE255535564F98E" bold="true" box="[199,293,1555,1581]" pageId="17" pageNumber="18">Female.</emphasis>
Unknown.
</paragraph>
<paragraph id="8BCA4B25FFBFFFD47DE2557755D5F9F2" blockId="17.[151,1437,151,2013]" box="[199,404,1591,1617]" pageId="17" pageNumber="18">
<emphasis id="B9019737FFBFFFD47DE255775556F9F2" bold="true" box="[199,279,1591,1617]" pageId="17" pageNumber="18">Larva.</emphasis>
Unknown.
</paragraph>
<paragraph id="8BCA4B25FFBFFFD47DE2551B55C4F9D6" blockId="17.[151,1437,151,2013]" box="[199,389,1627,1653]" pageId="17" pageNumber="18">
<emphasis id="B9019737FFBFFFD47DE2551B5549F9D6" bold="true" box="[199,264,1627,1653]" pageId="17" pageNumber="18">Eggs.</emphasis>
Unknown.
</paragraph>
<paragraph id="8BCA4B25FFBFFFD47DE2553F5791F93A" blockId="17.[151,1437,151,2013]" box="[199,976,1663,1689]" pageId="17" pageNumber="18">
Etymology. Named after K.M. Fenders wife Dorothy (
<bibRefCitation id="EFE436D4FFBFFFD47E14553F5785F93A" author="Fender, K. M." box="[817,964,1663,1689]" pageId="17" pageNumber="18" refId="ref39267" refString="Fender, K. M. (1962) Lampyridae. In: Hatch, M. H. (Ed.), The Beetles of the Pacific Northwest. Vol. 16. University of Washington Press, Seattle, Washington, pp. 41 - 43." type="book" year="1962">Fender 1962</bibRefCitation>
).
</paragraph>
<paragraph id="8BCA4B25FFBFFFD47DE255E35046F8EE" blockId="17.[151,1437,151,2013]" pageId="17" pageNumber="18">
<materialsCitation id="3B1D4178FFBFFFD47DE255E3561BF8A6" collectingDate="1937-06-18" collectionCode="VI, CAS" collectorName="Specimens Examined. &amp; Co." country="United States of America" location="Cal. Mill Valley" pageId="17" pageNumber="18" specimenCount="1" specimenCount-male="1" stateProvince="Oregon" typeStatus="holotype">
<collectorName id="26802EF3FFBFFFD47DE255E355FFF91E" box="[199,446,1699,1725]" pageId="17" pageNumber="18">Specimens Examined.</collectorName>
<emphasis id="B9019737FFBFFFD47CE755E3561CF91E" box="[450,605,1699,1725]" italics="true" pageId="17" pageNumber="18">Type Material</emphasis>
.
<typeStatus id="54CEF587FFBFFFD47F4C55E35690F91E" box="[617,721,1699,1725]" pageId="17" pageNumber="18" type="holotype">Holotype</typeStatus>
(
<figureCitation id="134E57A0FFBFFFD47FFB55E3570DF91E" box="[734,844,1699,1725]" captionStart="FIGURE 17" captionStartId="42.[152,255,1550,1574]" captionTargetBox="[169,1418,181,1526]" captionTargetId="figure-28@42.[169,1418,181,1526]" captionTargetPageId="42" captionText="FIGURE 17. Dorsal habitus of male Phausis (AF). (A) Phausis branhami sp. nov.; (B). Phausis californica; (C) Phausis dorothae; (D) Phausis inaccensa; (E) Phausis luminosa; (F) Phausis marina." figureDoi="http://doi.org/10.5281/zenodo.11348305" httpUri="https://zenodo.org/record/11348305/files/figure.png" pageId="17" pageNumber="18">Figs. 17C</figureCitation>
;
<figureCitation id="134E57A0FFBFFFD47E7055E357C9F91E" box="[853,904,1699,1725]" captionStart="FIGURE 24" captionStartId="48.[152,255,1943,1967]" captionTargetBox="[234,1354,181,1919]" captionTargetId="figure-28@48.[234,1354,181,1919]" captionTargetPageId="48" captionText="FIGURE 24. Type labels of Phausis (AL). (A) Phausis branhami sp. nov.; (B). Phausis californica; (C) Phausis dorothae; (D) Phausis inaccensa; (E) Phausis luminosa; (F) Phausis marina; (G) Phausis milleri sp. nov.; (H) Phausis nigra; (I) Phausis reticulata; (J) Phausis rhombica; (K) Phausis riversi; (L) Phausis skelleyi." figureDoi="http://doi.org/10.5281/zenodo.11348327" httpUri="https://zenodo.org/record/11348327/files/figure.png" pageId="17" pageNumber="18">24C</figureCitation>
).
<emphasis id="B9019737FFBFFFD47EBC55E25024F91F" bold="true" box="[921,1125,1698,1724]" pageId="17" pageNumber="18">
<collectingCountry id="F3620BB5FFBFFFD47EBC55E25794F91F" box="[921,981,1698,1724]" name="United States of America" pageId="17" pageNumber="18">USA</collectingCountry>
:
<collectingRegion id="49B185C7FFBFFFD47EFA55E25021F91F" box="[991,1120,1698,1724]" country="United States of America" name="Oregon" pageId="17" pageNumber="18">OREGON</collectingRegion>
:
</emphasis>
“Reuben Cr., Ore,
<collectionCode id="ED64D3E0FFBFFFD4781755E35113F91E" box="[1330,1362,1699,1725]" country="Norway" name="Mykotektet, National Veterinary Institute" pageId="17" pageNumber="18">VI</collectionCode>
.18.37, HOLOTYPE
<taxonomicName id="4C7530A6FFBFFFD47C1355875615F942" authority="Fender" authorityName="Fender" authorityYear="1962" box="[310,596,1735,1761]" class="Insecta" family="Lampyridae" genus="Phausis" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="dorothae">Phausis dorothae Fender</taxonomicName>
, Type No 9789,” [more detailed locality information: JOSEPHINE
<collectorName id="26802EF3FFBFFFD4784B558751D9F942" box="[1390,1432,1735,1761]" pageId="17" pageNumber="18">Co.</collectorName>
, Reuben Creek,
<date id="FFCB6DE5FFBFFFD47C6655AB5586F8A6" box="[323,455,1771,1797]" pageId="17" pageNumber="18" value="1937-06-18">
<collectingDate id="EF8F940DFFBFFFD47C6655AB5586F8A6" box="[323,455,1771,1797]" pageId="17" pageNumber="18" value="1937-06-18">18.VI.1937</collectingDate>
</date>
],
<specimenCount id="9D7380ACFFBFFFD47CFF55AB5642F8A6" box="[474,515,1771,1797]" count="1" pageId="17" pageNumber="18" type="male">1 ♂</specimenCount>
(
<collectionCode id="ED64D3E0FFBFFFD47F3755AB560FF8A6" box="[530,590,1771,1797]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" lsid="urn:lsid:biocol.org:col:34700" name="California Academy of Sciences" pageId="17" pageNumber="18" type="Museum">CAS</collectionCode>
).
</materialsCitation>
<materialsCitation id="3B1D4178FFBFFFD47F4555AB555BF8EE" collectingDate="1937-06-18" collectionCode="VI, CAS" collectorName="Specimens Examined. &amp; Co." country="United States of America" location="Cal. Mill Valley" pageId="17" pageNumber="18" specimenCount="1" specimenCount-male="1" stateProvince="Oregon" typeStatus="holotype">
<typeStatus id="54CEF587FFBFFFD47F4555AB5690F8A6" box="[608,721,1771,1797]" pageId="17" pageNumber="18" type="paratype">Paratypes</typeStatus>
.
<emphasis id="B9019737FFBFFFD47FFF55AA57D9F8A7" bold="true" box="[730,920,1770,1796]" pageId="17" pageNumber="18">
<collectingRegion id="49B185C7FFBFFFD47FFF55AA57D2F8A7" box="[730,915,1770,1796]" country="United States of America" name="California" pageId="17" pageNumber="18">CALIFORNIA</collectingRegion>
:
</emphasis>
“MARION Co.,
<location id="8EAA1DFEFFBFFFD4797D55AB514DF8A6" LSID="urn:lsid:plazi:treatment:03DCFA33FFBEFFD77DB255C855E5FDBA:8EAA1DFEFFBFFFD4797D55AB514DF8A6" box="[1112,1292,1771,1797]" country="United States of America" name="Cal. Mill Valley" pageId="17" pageNumber="18" stateProvince="Oregon">Cal. Mill Valley</location>
,
<date id="FFCB6DE5FFBFFFD4783355AB5119F8A6" box="[1302,1368,1771,1797]" pageId="17" pageNumber="18" value="1949-06-29">
29.
<collectionCode id="ED64D3E0FFBFFFD4781255AB5119F8A6" box="[1335,1368,1771,1797]" country="Norway" name="Mykotektet, National Veterinary Institute" pageId="17" pageNumber="18">VI</collectionCode>
</date>
.1949,
<collectorName id="26802EF3FFBFFFD47DB2544F55C6F88A" box="[151,391,1807,1833]" pageId="17" pageNumber="18">H.B. Leech Collector</collectorName>
,
<collectorName id="26802EF3FFBFFFD47CB7544F56A7F88A" box="[402,742,1807,1833]" pageId="17" pageNumber="18">Kenneth M. Fender Collection</collectorName>
1988 and 1992, Gifts to the
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,”
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(
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same except Upland, 529 Apr.,
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,”
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(
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)
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.
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<emphasis id="B9019737FFBFFFD47DE25417553BF8D2" box="[199,378,1879,1905]" italics="true" pageId="17" pageNumber="18">Other material.</emphasis>
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<emphasis id="B9019737FFBFFFD47CA1541656CFF8D3" bold="true" box="[388,654,1878,1904]" pageId="17" pageNumber="18">
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:
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:
</emphasis>
EL DORADO
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,
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E.
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,
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.
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18.1989, screening fume,
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and D.
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,
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(
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); MONTEREY
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, Andrew Molera
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,
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24.
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. 27.
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.2006,
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,
<collectorName id="26802EF3FFBFFFD47E7D54DF57D1F81A" box="[856,912,1951,1977]" pageId="17" pageNumber="18">Coll.</collectorName>
A.
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.
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<collectorName id="26802EF3FFBFFFD47EF554DF504CF81A" box="[976,1037,1951,1977]" pageId="17" pageNumber="18">Cline</collectorName>
and M.
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,
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(
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)
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.
</paragraph>
<paragraph id="8BCA4B25FFBFFFD47DE2548351D6F87E" blockId="17.[151,1437,151,2013]" box="[199,1431,1987,2013]" pageId="17" pageNumber="18">
Distribution. Found primarily within northern
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and parts of
<collectingRegion id="49B185C7FFBFFFD47EE654835059F87E" box="[963,1048,1987,2013]" country="United States of America" name="Oregon" pageId="17" pageNumber="18">Oregon</collectingRegion>
,
<collectingCountry id="F3620BB5FFBFFFD4790654835176F87E" box="[1059,1335,1987,2013]" name="United States of America" pageId="17" pageNumber="18">United States of America</collectingCountry>
(
<figureCitation id="134E57A0FFBFFFD47863548351CBF87E" box="[1350,1418,1987,2013]" captionStart="FIGURE 7" captionStartId="18.[152,255,1491,1515]" captionTargetBox="[582,1006,561,1468]" captionTargetId="figure-170@18.[582,1006,561,1468]" captionTargetPageId="18" captionText="FIGURE 7. Detail of North American distribution of Phausis dorothae." figureDoi="http://doi.org/10.5281/zenodo.11373261" httpUri="https://zenodo.org/record/11373261/files/figure.png" pageId="17" pageNumber="18">Fig. 7</figureCitation>
).
</paragraph>
<paragraph id="8BCA4B25FFBCFFD77DE253D756C7FF12" blockId="18.[151,1437,151,537]" box="[199,646,151,177]" pageId="18" pageNumber="19">Phenology. Occurs in late April to June.</paragraph>
<paragraph id="8BCA4B25FFBCFFD77DE253FB559FFEE2" blockId="18.[151,1437,151,537]" pageId="18" pageNumber="19">
Remarks. Some characters described above are not congruent with Fenders description of the species (
<bibRefCitation id="EFE436D4FFBCFFD7786953FB5492FF5A" author="Fender, K. M." pageId="18" pageNumber="19" pagination="83 - 95" refId="ref39317" refString="Fender, K. M. (1966) The genus Phausis in America North of Mexico (Coleoptera-Lampyridae). Northwest Science, 40 (3), 83 - 95." type="journal article" year="1966">Fender 1966</bibRefCitation>
). The characters are as follows: vertex head is not shallowly longitudinally concave; not all specimens examined have flavus palpi, legs, and last four abdominal segments, some are brunneous with abdominal segment VI lateral and posterior margins flavus.
</paragraph>
<paragraph id="8BCA4B25FFBCFFD77DE2520B573BFE56" blockId="18.[151,1437,151,537]" pageId="18" pageNumber="19">
Variation. Identified. In some specimens, the mouthparts and legs are either flavus or brunneous. The abdominal segments on the
<typeStatus id="54CEF587FFBCFFD77C7D522F55FBFE2A" box="[344,442,367,393]" pageId="18" pageNumber="19" type="holotype">holotype</typeStatus>
are paler along the lateral margins, but this is not present in the
<typeStatus id="54CEF587FFBCFFD77981522F5152FE2A" box="[1188,1299,367,393]" pageId="18" pageNumber="19" type="paratype">paratypes</typeStatus>
. Abdominal segment VI in some specimens appearing entirely yellow, except for the presence of an anterior-medially small dark brown spot. Tergal apicolateral angles on Abdominal segment VI entirely yellow to brown. The pronotum is entirely yellow in some specimens. This was also noted by Fender (1961).
</paragraph>
<paragraph id="8BCA4B25FFBCFFD77DE252BF55E5FDBA" blockId="18.[151,1437,151,537]" box="[199,420,511,537]" pageId="18" pageNumber="19">Biology. Unknown.</paragraph>
</subSubSection>
</treatment>
</document>