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<document id="010FD28CBA0E13890D13C6847E874090" ID-DOI="10.26879/832" ID-ISSN="1094-8074" ID-Zenodo-Dep="10961470" ID-ZooBank="152941A8-052F-4134-9E98-ABC4CAC8F01B" IM.bibliography_approvedBy="carolina" IM.illustrations_approvedBy="carolina" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="carolina" IM.tables_approvedBy="carolina" IM.taxonomicNames_approvedBy="carolina" IM.treatments_approvedBy="carolina" checkinTime="1712855045557" checkinUser="felipe" docAuthor="Alekseev, Vitalii I. &amp; Tomaszewska, Wioletta" docDate="2018" docId="03986E15FFD24B0F0E854B26FD50F914" docLanguage="en" docName="PalaeontolElectron.21.1.6A.pdf" docOrigin="Palaeontologia Electronica (6 A) 21 (1)" docSource="http://dx.doi.org/10.26879/832" docStyle="DocumentStyle:322FC57CC99045555090E8A84CFB029A.3:PalaeontolElectron.2018-.journal_article" docStyleId="322FC57CC99045555090E8A84CFB029A" docStyleName="PalaeontolElectron.2018-.journal_article" docStyleVersion="3" docTitle="Symbiotes borussiaeorientalis Alekseev &amp; Tomaszewska 2018, sp. nov." docType="treatment" docUuid="DB91C913-B619-447D-930A-E816A976F371" docUuidSource="ZooBank" docVersion="3" lastPageNumber="12" masterDocId="FFA1166DFFDB4B040D174A35FFF2FFD6" masterDocTitle="New handsome fungus beetles (Coleoptera: Coccinelloidea: Anamorphidae, Endomychidae) from European amber of the Upper Eocene" masterLastPageNumber="23" masterPageNumber="1" pageNumber="10" updateTime="1713983817784" updateUser="carolina" zenodo-license-document="CC-BY-NC-SA-4.0">
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<mods:title id="971CC4409CD401E2BBDAAB08FF4FF34B">New handsome fungus beetles (Coleoptera: Coccinelloidea: Anamorphidae, Endomychidae) from European amber of the Upper Eocene</mods:title>
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<mods:namePart id="4153F7228E0773E556B08CFF1915E35E">Alekseev, Vitalii I.</mods:namePart>
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<mods:title id="914F6757C0123BE98F30C52ECD2CA4C2">Palaeontologia Electronica</mods:title>
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<mods:date id="FCD8DAA3F37E45EC82E51268095F5C81">2018</mods:date>
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<mods:title id="DEC4A9AF3B0CAB286D22B9F95D0248DF">6 A</mods:title>
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<subSubSection id="C32B8C88FFD24B0D0E854B26FAA5FEFA" box="[914,1367,275,301]" pageId="9" pageNumber="10" type="nomenclature">
<paragraph id="8B8EDF03FFD24B0D0E854B26FAA5FEFA" blockId="9.[896,1384,243,332]" box="[914,1367,275,301]" pageId="9" pageNumber="10">
<heading id="D0C6686FFFD24B0D0E854B26FAA5FEFA" box="[914,1367,275,301]" centered="true" fontSize="10" level="2" pageId="9" pageNumber="10" reason="7">
<taxonomicName id="4C31A480FFD24B0D0E854B26FB06FEFA" authority="Alekseev &amp; Tomaszewska, 2018" authorityName="Alekseev &amp; Tomaszewska" authorityYear="2018" box="[914,1268,275,301]" class="Insecta" family="Anamorphidae" genus="Symbiotes" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="borussiaeorientalis" status="sp. nov.">
<emphasis id="B9450311FFD24B0D0E854B26FB06FEFA" box="[914,1268,275,301]" italics="true" pageId="9" pageNumber="10">Symbiotes borussiaeorientalis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A276BE6AFFD24B0D09EC4B26FAA5FEFA" box="[1275,1367,275,300]" pageId="9" pageNumber="10" rank="species">sp. nov.</taxonomicNameLabel>
</heading>
</paragraph>
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<subSubSection id="C32B8C88FFD24B0D093B4B06FA72FEA4" pageId="9" pageNumber="10" type="description">
<paragraph id="8B8EDF03FFD24B0D093B4B06FB4EFE9A" blockId="9.[896,1384,243,332]" box="[1068,1212,307,332]" pageId="9" pageNumber="10">
<figureCitation id="130AC386FFD24B0D093B4B06FB4EFE9A" box="[1068,1212,307,332]" captionStart="FIGURE 4" captionStartId="10.[198,289,1808,1831]" captionTargetBox="[251,1383,166,1774]" captionTargetId="figure-13@10.[251,1383,166,1774]" captionTargetPageId="10" captionText="FIGURE 4. Anamorphid and Endomychid from Baltic amber. 1-3, Symbiotes borussiaeorientalis sp. nov. (No. 1799-2 [CCHH], holotype), habitus in dorso-lateral (1) and ventro-lateral (2) views, and a reconstruction of the habitus in dorsal view (3). 4, Laima andreei gen. et sp. nov., a reconstruction of the habitus in dorsal view." figureDoi="http://doi.org/10.5281/zenodo.10961479" httpUri="https://zenodo.org/record/10961479/files/figure.png" pageId="9" pageNumber="10">Figure 4.1-3</figureCitation>
</paragraph>
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<emphasis id="B9450311FFD24B0D0E5F4B6BFA72FEA4" bold="true" box="[840,1408,350,370]" pageId="9" pageNumber="10">
zoobank.org/
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</emphasis>
</paragraph>
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<subSubSection id="C32B8C88FFD24B0D0E5F4BB6FB64FD2A" pageId="9" pageNumber="10" type="materials_examined">
<paragraph id="8B8EDF03FFD24B0D0E5F4BB6FB7DFDAA" blockId="9.[840,1441,387,1916]" pageId="9" pageNumber="10">
<emphasis id="B9450311FFD24B0D0E5F4BB6FBF4FE4A" bold="true" box="[840,1030,387,412]" pageId="9" pageNumber="10">Type material.</emphasis>
<materialsCitation id="3B59D55EFFD24B0D090D4BB6FB7DFDAA" collectionCode="CCHH" pageId="9" pageNumber="10" specimenCount="1" typeStatus="holotype">
<typeStatus id="548A61A1FFD24B0D090D4BB6FB71FE4A" box="[1050,1155,387,412]" pageId="9" pageNumber="10" type="holotype">Holotype</typeStatus>
No. 1799-2 [
<collectionCode id="ED2047C6FFD24B0D08544BB6FA65FE4B" box="[1347,1431,387,413]" pageId="9" pageNumber="10">CCHH</collectionCode>
], female (terminal parts of genitalia are visible; apices of elytra without small tubercles and other modifications). The beetle inclusion is preserved in a polished piece of transparent amber with an orange shade and the amber piece is embedded in polyester resin (total measurements are 11 x 9 x
<quantity id="4CC972E6FFD24B0D08864876FC88FDAA" metricMagnitude="-3" metricUnit="m" metricValue="4.0" pageId="9" pageNumber="10" unit="mm" value="4.0">4 mm</quantity>
). Syninclusions absent.
</materialsCitation>
</paragraph>
<paragraph id="8B8EDF03FFD24B0D0E5F48B6FB14FD4A" blockId="9.[840,1441,387,1916]" box="[840,1254,643,668]" pageId="9" pageNumber="10">
<emphasis id="B9450311FFD24B0D0E5F48B6FC2FFD4A" bold="true" box="[840,989,643,668]" pageId="9" pageNumber="10">
<typeStatus id="548A61A1FFD24B0D0E5F48B6FC74FD4A" box="[840,902,643,668]" pageId="9" pageNumber="10">Type</typeStatus>
strata.
</emphasis>
Baltic amber. Eocene.
</paragraph>
<paragraph id="8B8EDF03FFD24B0D0E5F4896FB64FD2A" blockId="9.[840,1441,387,1916]" pageId="9" pageNumber="10">
<materialsCitation id="3B59D55EFFD24B0D0E5F4896FB60FD2A" country="Russia" location="Yantarny" municipality="Sambian" pageId="9" pageNumber="10" specimenCount="1" stateProvince="Kaliningrad" typeStatus="holotype">
<emphasis id="B9450311FFD24B0D0E5F4896FC08FD6A" bold="true" box="[840,1018,675,700]" pageId="9" pageNumber="10">
<typeStatus id="548A61A1FFD24B0D0E5F4896FC76FD6A" box="[840,900,675,700]" pageId="9" pageNumber="10">Type</typeStatus>
locality.
</emphasis>
<collectingCountry id="F3269F93FFD24B0D09194896FB96FD6A" box="[1038,1124,675,700]" name="Russia" pageId="9" pageNumber="10">Russia</collectingCountry>
,
<collectingRegion id="49F511E1FFD24B0D096C4896FA91FD6A" box="[1147,1379,675,700]" country="Russia" name="Kaliningrad" pageId="9" pageNumber="10">Kaliningrad region</collectingRegion>
, the
<collectingMunicipality id="6BEA4579FFD24B0D0E5F48F6FC43FD0B" box="[840,945,707,733]" pageId="9" pageNumber="10">Sambian</collectingMunicipality>
(Samland) peninsula,
<location id="8EEE89D8FFD24B0D09CA48F6FAB7FD0A" LSID="urn:lsid:plazi:treatment:03986E15FFD24B0F0E854B26FD50F914:8EEE89D8FFD24B0D09CA48F6FAB7FD0A" box="[1245,1349,707,732]" country="Russia" municipality="Sambian" name="Yantarny" pageId="9" pageNumber="10" stateProvince="Kaliningrad">Yantarny</location>
settlement (formerly Palmnicken)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C32B8C88FFD24B0D0E5F4936FC40FCAA" pageId="9" pageNumber="10" type="etymology">
<paragraph id="8B8EDF03FFD24B0D0E5F4936FC40FCAA" blockId="9.[840,1441,387,1916]" pageId="9" pageNumber="10">
<emphasis id="B9450311FFD24B0D0E5F4936FC24FCCA" bold="true" box="[840,982,771,796]" pageId="9" pageNumber="10">Etymology.</emphasis>
The specific epithet refers to the region of the specimens origin and is formed after the Medieval Latin
<emphasis id="B9450311FFD24B0D092B4976FADCFC8A" box="[1084,1326,835,860]" italics="true" pageId="9" pageNumber="10">
<taxonomicName id="4C31A480FFD24B0D09554976FAD4FC8A" box="[1090,1318,835,860]" class="Insecta" family="Anamorphidae" genus="Borussia" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="orientalis">Borussia orientalis</taxonomicName>
</emphasis>
(Eastern Prussia).
</paragraph>
</subSubSection>
<subSubSection id="C32B8C88FFD24B0D0E5F49B6FA64FBAA" pageId="9" pageNumber="10" type="discussion">
<paragraph id="8B8EDF03FFD24B0D0E5F49B6FA64FBAA" blockId="9.[840,1441,387,1916]" pageId="9" pageNumber="10">
<emphasis id="B9450311FFD24B0D0E5F49B6FC32FC4A" bold="true" box="[840,960,899,924]" pageId="9" pageNumber="10">Remarks.</emphasis>
The newly described species from Baltic amber was assigned to the genus
<taxonomicName id="4C31A480FFD24B0D08334996FA52FC6B" authorityName="Redtenbacher" authorityYear="1849" box="[1316,1440,931,957]" class="Insecta" family="Anamorphidae" genus="Symbiotes" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="genus">
<emphasis id="B9450311FFD24B0D08334996FA52FC6B" box="[1316,1440,931,957]" italics="true" pageId="9" pageNumber="10">Symbiotes</emphasis>
</taxonomicName>
based of following combination of morphological characters: body form subparallel; antenna composed of 11 antennomeres; procoxal cavities open externally; pronotum with posterior margin truncate and lateral margins raised; pronotal carinae not reaching posterior margin; and tarsi 4-segmented.
</paragraph>
</subSubSection>
<subSubSection id="C32B8C88FFD24B0D0E5F4EB6FBD8F96A" pageId="9" pageNumber="10" type="diagnosis">
<paragraph id="8B8EDF03FFD24B0D0E5F4EB6FBD8F96A" blockId="9.[840,1441,387,1916]" pageId="9" pageNumber="10">
<emphasis id="B9450311FFD24B0D0E5F4EB6FC3CFB4A" bold="true" box="[840,974,1155,1180]" pageId="9" pageNumber="10">Diagnosis.</emphasis>
<taxonomicName id="4C31A480FFD24B0D0EC94EB6FABAFB4A" authorityName="Alekseev &amp; Tomaszewska" authorityYear="2018" box="[990,1352,1155,1181]" class="Insecta" family="Anamorphidae" genus="Symbiotes" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="borussiaeorientalis">
<emphasis id="B9450311FFD24B0D0EC94EB6FABAFB4A" box="[990,1352,1155,1181]" italics="true" pageId="9" pageNumber="10">Symbiotes borussiaeorientalis</emphasis>
</taxonomicName>
differs from all known congeners in having coarsely and irregularly punctured elytra with distinct subsutural striae only, and scarce, poorly visible dorsal pubescence. Newly described species is
<quantity id="4CC972E6FFD24B0D09EA4F36FAA8FACA" box="[1277,1370,1283,1308]" metricMagnitude="-3" metricUnit="m" metricValue="1.6" pageId="9" pageNumber="10" unit="mm" value="1.6">1.6 mm</quantity>
long, thereby smaller than most extant representatives of
<taxonomicName id="4C31A480FFD24B0D0E704F76FC11FA8B" authorityName="Redtenbacher" authorityYear="1849" box="[871,995,1347,1373]" class="Insecta" family="Anamorphidae" genus="Symbiotes" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="genus">
<emphasis id="B9450311FFD24B0D0E704F76FC11FA8B" box="[871,995,1347,1373]" italics="true" pageId="9" pageNumber="10">Symbiotes</emphasis>
</taxonomicName>
(
<quantity id="4CC972E6FFD24B0D0EE44F76FB7BFA8A" box="[1011,1161,1347,1372]" metricMagnitude="-3" metricUnit="m" metricValue="2.05" metricValueMax="2.4" metricValueMin="1.7" pageId="9" pageNumber="10" unit="mm" value="2.05" valueMax="2.4" valueMin="1.7">1.72.4 mm</quantity>
), except
<taxonomicName id="4C31A480FFD24B0D09E34F76FA6AFA8A" baseAuthorityName="Lucas" baseAuthorityYear="1846" box="[1268,1432,1347,1372]" class="Insecta" family="Anamorphidae" genus="Symbiotes" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="gibberosus">
<emphasis id="B9450311FFD24B0D09E34F76FA6AFA8A" box="[1268,1432,1347,1372]" italics="true" pageId="9" pageNumber="10">S. gibberosus</emphasis>
</taxonomicName>
, which is
<quantity id="4CC972E6FFD24B0D0EA44F56FBB0FAAA" box="[947,1090,1379,1404]" metricMagnitude="-3" metricUnit="m" metricValue="1.65" metricValueMax="1.8" metricValueMin="1.5" pageId="9" pageNumber="10" unit="mm" value="1.65" valueMax="1.8" valueMin="1.5">1.51.8 mm</quantity>
long. Additionally, new fossil species can be distinguished from
<taxonomicName id="4C31A480FFD24B0D09FC4FB6FABAFA4A" authorityName="Blatchley" authorityYear="1910" box="[1259,1352,1411,1436]" class="Insecta" family="Anamorphidae" genus="Symbiotes" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="duryi">
<emphasis id="B9450311FFD24B0D09FC4FB6FABAFA4A" box="[1259,1352,1411,1436]" italics="true" pageId="9" pageNumber="10">S. duryi</emphasis>
</taxonomicName>
and
<taxonomicName id="4C31A480FFD24B0D08904FB6FC38FA6A" baseAuthorityName="Lucas" baseAuthorityYear="1846" class="Insecta" family="Anamorphidae" genus="Symbiotes" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="gibberosus">
<emphasis id="B9450311FFD24B0D08904FB6FC38FA6A" italics="true" pageId="9" pageNumber="10">S. gibberosus</emphasis>
</taxonomicName>
by having smooth (not denticulate) lateral pronotal margins; from
<taxonomicName id="4C31A480FFD24B0D09BB4FF6FAFFFA0A" authorityName="Redtenbacher" authorityYear="1849" box="[1196,1293,1475,1500]" class="Insecta" family="Anamorphidae" genus="Symbiotes" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="latus">
<emphasis id="B9450311FFD24B0D09BB4FF6FAFFFA0A" box="[1196,1293,1475,1500]" italics="true" pageId="9" pageNumber="10">S. latus</emphasis>
</taxonomicName>
by its less transverse pronotum (2.5 times wider than long in
<taxonomicName id="4C31A480FFD24B0D0E5F4C36FC50F9CA" authorityName="Redtenbacher" authorityYear="1849" box="[840,930,1539,1564]" class="Insecta" family="Anamorphidae" genus="Symbiotes" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="latus">
<emphasis id="B9450311FFD24B0D0E5F4C36FC50F9CA" box="[840,930,1539,1564]" italics="true" pageId="9" pageNumber="10">S. latus</emphasis>
</taxonomicName>
), wider elytra, and dark color; from
<taxonomicName id="4C31A480FFD24B0D08464C36FC55F9EA" baseAuthorityName="Lucas" baseAuthorityYear="1846" class="Insecta" family="Anamorphidae" genus="Symbiotes" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="gibberosus">
<emphasis id="B9450311FFD24B0D08464C36FC55F9EA" italics="true" pageId="9" pageNumber="10">S. gibberosus</emphasis>
</taxonomicName>
by less convex body, and absence of pits at base of lateral sulci; from
<emphasis id="B9450311FFD24B0D09B14C76FA52F98A" box="[1190,1440,1603,1628]" italics="true" pageId="9" pageNumber="10">
<taxonomicName id="4C31A480FFD24B0D09B14C76FAF5F98A" authorityName="Blatchley" authorityYear="1910" box="[1190,1287,1603,1628]" class="Insecta" family="Anamorphidae" genus="Symbiotes" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="duryi">S. duryi</taxonomicName>
,
<taxonomicName id="4C31A480FFD24B0D08024C76FA69F98A" authorityName="Reitter" authorityYear="1881" box="[1301,1435,1603,1628]" class="Insecta" family="Anamorphidae" genus="Symbiotes" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="armatus">S. armatus</taxonomicName>
,
</emphasis>
and
<taxonomicName id="4C31A480FFD24B0D0E6B4C56FC26F9AA" authorityName="Redtenbacher" authorityYear="1849" box="[892,980,1635,1660]" class="Insecta" family="Anamorphidae" genus="Symbiotes" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="latus">
<emphasis id="B9450311FFD24B0D0E6B4C56FC26F9AA" box="[892,980,1635,1660]" italics="true" pageId="9" pageNumber="10">S. latus</emphasis>
</taxonomicName>
by presence of subsutural stria without distinct punctation; and from
<taxonomicName id="4C31A480FFD24B0D09B34CB6FAD5F94A" authorityName="Reitter" authorityYear="1881" box="[1188,1319,1667,1692]" class="Insecta" family="Anamorphidae" genus="Symbiotes" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="armatus">
<emphasis id="B9450311FFD24B0D09B34CB6FAD5F94A" box="[1188,1319,1667,1692]" italics="true" pageId="9" pageNumber="10">S. armatus</emphasis>
</taxonomicName>
by widely rounded scutellum.
</paragraph>
</subSubSection>
<subSubSection id="C32B8C88FFD24B0F0E5F4CF6FD50F914" lastPageId="11" lastPageNumber="12" pageId="9" pageNumber="10" type="description">
<paragraph id="8B8EDF03FFD24B0D0E5F4CF6FBF0F8EA" blockId="9.[840,1441,387,1916]" pageId="9" pageNumber="10">
<emphasis id="B9450311FFD24B0D0E5F4CF6FC10F90A" bold="true" box="[840,994,1731,1756]" pageId="9" pageNumber="10">Description.</emphasis>
Body uniformly dark brown, elongate oval, moderately convex, shiny, scarcely pubescent. Total body length
<quantity id="4CC972E6FFD24B0D097A4D36FB3DF8CA" box="[1133,1231,1795,1820]" metricMagnitude="-3" metricUnit="m" metricValue="1.6" pageId="9" pageNumber="10" unit="mm" value="1.6">1.6 mm</quantity>
, width (at widest point)
<quantity id="4CC972E6FFD24B0D0E854D16FC0FF8EA" box="[914,1021,1827,1852]" metricMagnitude="-4" metricUnit="m" metricValue="7.5" pageId="9" pageNumber="10" unit="mm" value="0.75">0.75 mm</quantity>
.
</paragraph>
<paragraph id="8B8EDF03FFD24B0F0E954D76FD5CFDF4" blockId="9.[840,1441,387,1916]" lastBlockId="11.[192,793,169,1730]" lastPageId="11" lastPageNumber="12" pageId="9" pageNumber="10">Head deeply retracted in pronotum, transverse; with fine and very sparse punctation. Eyes large, prominent. Fronto-clypeal suture distinct, straight. Antennae comparatively long, extending to base of elytra; composed of 11 antennomeres, with loose club composed of three antennomeres. Antennomere length proportions follow formula: 2.5: 2.0: 1.0: 1.0: 1.0: 1.0: 1.0: 1.0: 1.5: 2.5: 3.5. Antennomeres 38 subequal in length and form, short and cylindrical; antennomere 9 triangular; antennomere 10 as long as wide; terminal antennomere subcylindrical, rounded apically, almost 2.0 times as long as wide. Terminal maxillary palpomere elongate, subcylindrical, tapering.</paragraph>
<caption id="DF4E8F8BFFD14B0E0DD14D25FB67F8B4" ID-DOI="http://doi.org/10.5281/zenodo.10961479" ID-Zenodo-Dep="10961479" httpUri="https://zenodo.org/record/10961479/files/figure.png" pageId="10" pageNumber="11" startId="10.[198,289,1808,1831]" targetBox="[251,1383,166,1774]" targetPageId="10" targetType="figure">
<paragraph id="8B8EDF03FFD14B0E0DD14D25FB67F8B4" blockId="10.[198,1430,1808,1890]" pageId="10" pageNumber="11">
<emphasis id="B9450311FFD14B0E0DD14D25FECEF8FE" bold="true" box="[198,316,1808,1832]" pageId="10" pageNumber="11">FIGURE 4.</emphasis>
Anamorphid and Endomychid from Baltic amber.
<emphasis id="B9450311FFD14B0E0E5D4D24FC86F8FE" bold="true" box="[842,884,1809,1832]" pageId="10" pageNumber="11">1-3,</emphasis>
<taxonomicName id="4C31A480FFD14B0E0E6D4D25FB4AF8FE" authority="Alekseev &amp; Tomaszewska, 2018" authorityName="Alekseev &amp; Tomaszewska" authorityYear="2018" box="[890,1208,1808,1832]" class="Insecta" family="Anamorphidae" genus="Symbiotes" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="borussiaeorientalis" status="sp. nov.">
<emphasis id="B9450311FFD14B0E0E6D4D25FB4AF8FE" box="[890,1208,1808,1832]" italics="true" pageId="10" pageNumber="11">Symbiotes borussiaeorientalis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A276BE6AFFD14B0E09A84D24FAE0F8FE" box="[1215,1298,1809,1832]" pageId="10" pageNumber="11" rank="species">sp. nov.</taxonomicNameLabel>
(No. 1799-2 [CCHH], holotype), habitus in dorso-lateral (
<emphasis id="B9450311FFD14B0E0F824D1BFD50F893" bold="true" box="[661,674,1838,1861]" pageId="10" pageNumber="11">1</emphasis>
) and ventro-lateral (
<emphasis id="B9450311FFD14B0E0E6E4D1BFC74F893" bold="true" box="[889,902,1838,1861]" pageId="10" pageNumber="11">2</emphasis>
) views, and a reconstruction of the habitus in dorsal view (
<emphasis id="B9450311FFD14B0E0C3D4D7EFEC5F8B4" bold="true" box="[298,311,1867,1890]" pageId="10" pageNumber="11">3</emphasis>
).
<emphasis id="B9450311FFD14B0E0C5B4D7EFE93F8B4" bold="true" box="[332,353,1867,1890]" pageId="10" pageNumber="11">4,</emphasis>
<taxonomicName id="4C31A480FFD14B0E0C704D7EFE0DF8B4" authority="Alekseev &amp; Tomaszewska, 2018" authorityName="Alekseev &amp; Tomaszewska" authorityYear="2018" box="[359,511,1867,1890]" class="Insecta" family="Endomychidae" genus="Laima" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="andreei" status="gen. et sp. nov.">
<emphasis id="B9450311FFD14B0E0C704D7EFE0DF8B4" box="[359,511,1867,1890]" italics="true" pageId="10" pageNumber="11">Laima andreei</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A276BE6AFFD14B0E0F114D7EFD5EF8B4" box="[518,684,1867,1890]" pageId="10" pageNumber="11" rank="species">gen. et sp. nov.</taxonomicNameLabel>
, a reconstruction of the habitus in dorsal view.
</paragraph>
</caption>
<paragraph id="8B8EDF03FFD04B0F0DED481CFD59FC14" blockId="11.[192,793,169,1730]" pageId="11" pageNumber="12">Pronotum widest at middle, strongly transverse (2.0 times wider than long); pronotal anterior and posterior margins almost straight, anterior angles obtuse, and basal angles right-angled. Lateral margins of pronotum arcuate, broad, raised, and lateral edges smooth. Median region convex, base with well-marked transverse sulcus, lateral sulci large, triangularly longitudinal, reaching anteriorly nearly one-half length of pronotum. Pronotal disc punctate: coarsely and moderately densely at sides (interspaces 1.02.0 times puncture diameter), more finely and sparsely near middle (interspaces 2.04.0 times puncture diameter).</paragraph>
<paragraph id="8B8EDF03FFD04B0F0DED49FCFE0EFAB4" blockId="11.[192,793,169,1730]" pageId="11" pageNumber="12">Elytra comparatively long (3.0 times longer than pronotum), oval, decidedly wider than prothorax, attaining their greatest width within about one-third of their basal length; with explanate margins. Elytral disc coarsely and densely punctate; elytral striae absent except subsutural ones. Punctures irregular, interspaces 0.20.5 of diameter, denser laterally and sparser medially. Elytral epipleura pubescent. Subsutural stria distinct, broadly curved near scutellum and reaching middle of elytral base. Elytral apices conjointly rounded. Scutellum visible, transverse (about two times wider than long), widely rounded posteriorly.</paragraph>
<paragraph id="8B8EDF03FFD04B0F0DED4F5CFEC5F994" blockId="11.[192,793,169,1730]" pageId="11" pageNumber="12">Legs with procoxal cavities open externally. Metacoxae transverse, widely separated. Femur widest in middle, more than twice as wide as tibia. All tibiae simple. Tarsal formula 4-4-4. Tarsomeres simple, cylindrical; tarsomere 4 longest. Tarsal claws simple, almost 0.3 times as long as tarsomere 4.</paragraph>
<paragraph id="8B8EDF03FFD04B0F0DED4C7CFD50F914" blockId="11.[192,793,169,1730]" pageId="11" pageNumber="12">Abdomen with five ventrites. Ventrite 1 as long as 2 and 3 combined; ventrites 2, 3, and 4 subequal in length; ventrite 5, 1.5 times longer than 4. Apex of terminal ventrite widely rounded.</paragraph>
</subSubSection>
</treatment>
</document>