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<document id="24F1ED418D560252F3CA6EFBD02BFD9E" ID-DOI="10.5281/zenodo.179290" ID-GBIF-Dataset="3218835d-a188-406e-96ec-346bd5911888" ID-ISSN="1175-5326" ID-Zenodo-Dep="179290" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1459961374711" checkinUser="plazi" docAuthor="Anker, Arthur, Hurt, Carla &amp; Knowlton, Nancy" docDate="2007" docId="038E6046FFEBFF9D37E3FA87FC50ACC0" docLanguage="en" docName="zt01626p023.pdf" docOrigin="Zootaxa 1626" docStyle="DocumentStyle:6581673A57F01A3145754A1E615EDFF0.4:Zootaxa.2007-2008.journal_article" docStyleId="6581673A57F01A3145754A1E615EDFF0" docStyleName="Zootaxa.2007-2008.journal_article" docStyleVersion="4" docTitle="Alpheus aequus Kim &amp; Abele 1988" docType="treatment" docVersion="10" lastPageNumber="19" masterDocId="FFB7183EFFFBFF8F3774FF98FFC6AB24" masterDocTitle="Three transisthmian snapping shrimps (Crustacea: Decapoda: Alpheidae: Alpheus) associated with innkeeper worms (Echiura: Thalassematidae) in Panama" masterLastPageNumber="23" masterPageNumber="1" pageNumber="17" updateTime="1698230876662" updateUser="plazi">
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<mods:title id="E262F8B4EB88FF4341A0BEA480BB7DB2">Three transisthmian snapping shrimps (Crustacea: Decapoda: Alpheidae: Alpheus) associated with innkeeper worms (Echiura: Thalassematidae) in Panama</mods:title>
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<mods:namePart id="6D88B3C4BAF8048F5DB04A435696FAC6">Anker, Arthur</mods:namePart>
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<mods:namePart id="F98EEA1328CC10B422C228F6E203A7AB">Hurt, Carla</mods:namePart>
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<mods:namePart id="2F6A55728F1834BD1DE8E38EA08D3083">Knowlton, Nancy</mods:namePart>
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<mods:title id="F664D9D491979367149DBF8BC09AE925">Zootaxa</mods:title>
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<mods:date id="04DDD5C97BB51855358037FA6E44E232">2007</mods:date>
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<mods:number id="14F1DE1E9B160343993D930CCB7C2F67">1626</mods:number>
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<treatment id="038E6046FFEBFF9D37E3FA87FC50ACC0" ID-DOI="http://doi.org/10.5281/zenodo.6247389" ID-GBIF-Taxon="119376643" ID-Zenodo-Dep="6247389" LSID="urn:lsid:plazi:treatment:038E6046FFEBFF9D37E3FA87FC50ACC0" httpUri="http://treatment.plazi.org/id/038E6046FFEBFF9D37E3FA87FC50ACC0" lastPageId="18" lastPageNumber="19" pageId="16" pageNumber="17">
<subSubSection id="C33D82DBFFEBFF9F37E3FA87FEECAE78" pageId="16" pageNumber="17" type="nomenclature">
<paragraph id="8B98D150FFEBFF9F37E3FA87FD88AE1D" blockId="16.[151,590,1311,1372]" box="[151,590,1311,1337]" pageId="16" pageNumber="17">
<heading id="D0D0663CFFEBFF9F37E3FA87FD88AE1D" bold="true" box="[151,590,1311,1337]" fontSize="11" level="1" pageId="16" pageNumber="17" reason="1">
<taxonomicName id="4C27AAD3FFEBFF9F37E3FA87FD88AE1D" ID-CoL="85LDH" authority="Kim &amp; Abele, 1988" authorityName="Kim &amp; Abele" authorityYear="1988" box="[151,590,1311,1337]" class="Malacostraca" family="Alpheidae" genus="Alpheus" kingdom="Animalia" order="Decapoda" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="aequus">
<emphasis id="B9530D42FFEBFF9F37E3FA87FD88AE1D" bold="true" box="[151,590,1311,1337]" pageId="16" pageNumber="17">
<emphasis id="B9530D42FFEBFF9F37E3FA87FE91AE1D" bold="true" box="[151,343,1311,1337]" italics="true" pageId="16" pageNumber="17">Alpheus aequus</emphasis>
<bibRefCitation id="EFB6ACA1FFEBFF9F362BFA87FD88AE1D" author="Kim" box="[351,590,1311,1337]" pageId="16" pageNumber="23" refString="Kim, W. &amp; Abele, L. G. (1988) The snapping shrimp genus Alpheus from the eastern Pacific (Decapoda: Caridea: Alpheidae). Smithsonian Contributions to Zoology, 454, 1 - 119." type="journal article" year="1988">Kim &amp; Abele, 1988</bibRefCitation>
</emphasis>
</taxonomicName>
</heading>
</paragraph>
<paragraph id="8B98D150FFEBFF9F37E3FADAFEECAE78" blockId="16.[151,590,1311,1372]" box="[151,298,1346,1372]" pageId="16" pageNumber="17">
<figureCitation id="131CCDD5FFEBFF9F37E3FADAFF31AE78" box="[151,247,1346,1372]" captionStart="FIGURE 10" captionStartId="17.[151,255,1460,1484]" captionTargetBox="[172,1431,204,1436]" captionTargetId="figure@17.[151,1436,194,1437]" captionTargetPageId="17" captionText="FIGURE 10. Alpheus aequus Kim &amp; Abeke, 1988, female from Coibita, Coiba Marine National Park, Panama (MNHN 16389): a, frontal margin, dorsal view; b, major cheliped, lateral view; c, same, chela, mesial view; d, minor chela, lateral view; e, second pereiopod, lateral view; f, third pereiopod, lateral view. Scale bars: 1 mm." httpUri="https://zenodo.org/record/179300/files/figure.png" pageId="16" pageNumber="17">Figs. 10</figureCitation>
,
<figureCitation id="131CCDD5FFEBFF9F3670FADAFED8AE78" box="[260,286,1346,1372]" captionStart="FIGURE 11" captionStartId="18.[151,255,1495,1519]" captionTargetBox="[151,1434,434,1471]" captionTargetId="figure@18.[151,1436,434,1471]" captionTargetPageId="18" captionText="FIGURE 11. Color patterns in life. Alpheus christofferseni n. sp., male from Bocas del Toro, Panama (USNM 1100667) (a – c): a, b, dorsal view (note partly faded chromatopores in a); c, same, lateral view. Alpheus naos n. sp., male holotype from Punta Culebra, Isla Naos, Panama (USNM 1100668) (d, e): d, dorsal view; e, lateral view. Alpheus aequus Kim &amp; Abele, 1988, female from Coibita, Coiba Marine National Park, Panama (MNHN-Na 16389) (f): dorsal view." httpUri="https://zenodo.org/record/179301/files/figure.png" pageId="16" pageNumber="17">11</figureCitation>
f
</paragraph>
</subSubSection>
<subSubSection id="C33D82DBFFEBFF9F37E3FA0AFD3AAEED" pageId="16" pageNumber="17" type="reference_group">
<paragraph id="8B98D150FFEBFF9F37E3FA0AFD3AAEED" blockId="16.[151,1436,1426,1481]" pageId="16" pageNumber="17">
<treatmentCitationGroup id="AB37F67EFFEBFF9F37E3FA0AFD3AAEED" pageId="16" pageNumber="17">
<taxonomicName id="4C27AAD3FFEBFF9F37E3FA0AFD8BAE8D" ID-CoL="85LDH" authority="Kim &amp; Abele, 1988: 55" authorityName="Kim &amp; Abele" authorityPageNumber="55" authorityYear="1988" box="[151,589,1426,1449]" class="Malacostraca" family="Alpheidae" genus="Alpheus" kingdom="Animalia" order="Decapoda" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="aequus">
<emphasis id="B9530D42FFEBFF9F37E3FA0AFE86AE8D" box="[151,320,1426,1449]" italics="true" pageId="16" pageNumber="17">Alpheus aequus</emphasis>
<treatmentCitation id="0A86F741FFEBFF9F363CFA0AFD8BAE8D" author="Kim" box="[328,589,1426,1449]" page="55" pageId="16" pageNumber="17" year="1988">
<bibRefCitation id="EFB6ACA1FFEBFF9F363CFA0AFDE2AE8D" author="Kim" box="[328,548,1426,1449]" pageId="16" pageNumber="23" refString="Kim, W. &amp; Abele, L. G. (1988) The snapping shrimp genus Alpheus from the eastern Pacific (Decapoda: Caridea: Alpheidae). Smithsonian Contributions to Zoology, 454, 1 - 119." type="journal article" year="1988">Kim &amp; Abele, 1988</bibRefCitation>
: 55
</treatmentCitation>
</taxonomicName>
, fig. 23;
<treatmentCitation id="0A86F741FFEBFF9F35C4FA0AFBCBAE8D" author="Wicksten" box="[688,1037,1426,1449]" page="4" pageId="16" pageNumber="17" year="1992">
<bibRefCitation id="EFB6ACA1FFEBFF9F35C4FA0AFC37AE8D" author="Wicksten" box="[688,1009,1426,1449]" pageId="16" pageNumber="23" refString="Wicksten, M. K. &amp; Hendrickx, M. E. (1992) Checklist of penaeoid and caridean shrimps (Decapoda: Penaeoidea, Caridea) from the eastern tropical Pacific. Proceedings of San Diego Society of Natural History, 9, 1 - 11." type="journal article" year="1992">Wicksten &amp; Hendrickx, 1992</bibRefCitation>
: 4
</treatmentCitation>
;
<treatmentCitation id="0A86F741FFEBFF9F336DFA0AFB27AE8D" author="Ramos" box="[1049,1249,1426,1449]" page="145" pageId="16" pageNumber="17" year="1995">
<bibRefCitation id="EFB6ACA1FFEBFF9F336DFA0AFB6DAE8D" author="Ramos" box="[1049,1195,1426,1449]" pageId="16" pageNumber="23" refString="Ramos, G. (1995) Nuevos registros de camarones alfeidos (Crustacea, Decapoda, Alpheidae) para el Pacifico de Colombia. In: Cantera, J. R. &amp; Restrepo, J. D. (eds.), Delta del Rio San Juan, Bahias de Malaga y Buenaventura, Pacifico Colombiano. Tomo II, pp. 127 - 153. Colciencias, Univeridad EAFIT y Universidad del Valle, Colombia." type="book chapter" year="1995">Ramos, 1995</bibRefCitation>
: 145
</treatmentCitation>
, fig. 9;
<treatmentCitation id="0A86F741FFEBFF9F3243FA0AFE4EAEED" author="Vargas" page="898" pageId="16" pageNumber="17" year="1999">
<bibRefCitation id="EFB6ACA1FFEBFF9F3243FA0AFE95AEED" author="Vargas" pageId="16" pageNumber="23" refString="Vargas, R. &amp; Cortes, J. (1999) Biodiversidad marina de Costa Rica: Crustacea: Decapoda (Penaeoidea, Sergestoidea, Stenopodidea, Caridea, Thalassinidea, Palinura) del Caribe. Revista de Biologia Tropical, 47, 887 - 911." type="journal article" year="1999">Vargas &amp; Cortés, 1999</bibRefCitation>
: 898
</treatmentCitation>
;
<treatmentCitation id="0A86F741FFEBFF9F36E7FA2AFD3EAEED" author="Wicksten" box="[403,760,1458,1481]" page="63" pageId="16" pageNumber="17" year="2003">
<bibRefCitation id="EFB6ACA1FFEBFF9F36E7FA2AFD16AEED" author="Wicksten" box="[403,720,1458,1481]" pageId="16" pageNumber="23" refString="Wicksten, M. K. &amp; Hendrickx, M. E. (2003) An updated checklist of benthic marine and brackish water shrimps (Decapoda: Penaeoidea, Stenopodidea, Caridea) from the eastern tropical Pacific. In: Hendrickx, M. (ed.), Contribuciones al estudio de los crustaceos del Pacifico Este, Instituto de Ciencias del Mar y Limnologia, Universidad Nacional Autonoma de Mexico, Mexico, pp. 49 - 76." type="book chapter" year="2003">Wicksten &amp; Hendrickx, 2003</bibRefCitation>
: 63
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C33D82DBFFEBFF9F37E3FA62FD32ADF8" pageId="16" pageNumber="17" type="materials_examined">
<paragraph id="8B98D150FFEBFF9F37E3FA62FD32ADF8" blockId="16.[151,1437,1530,2036]" pageId="16" pageNumber="17">
<emphasis id="B9530D42FFEBFF9F37E3FA62FE40AD30" bold="true" box="[151,390,1530,1556]" pageId="16" pageNumber="17">Material examined</emphasis>
.
<specimenCount id="9D211AD9FFEBFF9F36E2FA62FE38AD30" box="[406,510,1530,1556]" pageId="16" pageNumber="17" type="female">1 female</specimenCount>
(CL 4.7, TL 14.3), MNHN-Na 16388,
<collectingCountry id="F33091C0FFEBFF9F34A3FA62FBF1AD30" box="[983,1079,1530,1556]" name="Panama" pageId="16" pageNumber="17">Panama</collectingCountry>
, Coiba Marine National Park, Coibita, mud-rock intertidal, under rock, in burrow of
<taxonomicName id="4C27AAD3FFEBFF9F344FF9BBFBDFAD18" box="[827,1049,1571,1596]" class="Polychaeta" family="Echiuridae" genus="Ochetostoma" higherTaxonomySource="CoL" kingdom="Animalia" order="Echiuroidea" pageId="16" pageNumber="17" phylum="Echiura" rank="species" species="edax">
<emphasis id="B9530D42FFEBFF9F344FF9BBFBDFAD18" box="[827,1049,1571,1596]" italics="true" pageId="16" pageNumber="17">Ochetostoma edax</emphasis>
</taxonomicName>
(host collected and preserved), extreme low tide,
<date id="FF99F790FFEBFF9F3618F9D2FDCEAD40" box="[364,520,1610,1636]" pageId="16" pageNumber="17" value="2007-03-20">20 Mar 2007</date>
, coll. A. Anker, I. Marin, J. Jara, E. Gómez and E. Tóth [fcn 07-109]; 1 ovig. female (CL 4.9, TL 14.3), MNHN-Na 16389,
<collectingCountry id="F33091C0FFEBFF9F35C2F9EAFCD0ADA8" box="[694,790,1650,1676]" name="Panama" pageId="16" pageNumber="17">Panama</collectingCountry>
, Coiba Marine National Park, Coibita, mud-rock intertidal near
<collectionCode id="ED364995FFEBFF9F367BF902FE89AD90" box="[271,335,1690,1716]" pageId="16" pageNumber="17">STRI</collectionCode>
station, under rock, in burrow of
<taxonomicName id="4C27AAD3FFEBFF9F3590F903FC79AD90" box="[740,959,1691,1716]" class="Polychaeta" family="Echiuridae" genus="Ochetostoma" higherTaxonomySource="CoL" kingdom="Animalia" order="Echiuroidea" pageId="16" pageNumber="17" phylum="Echiura" rank="species" species="edax">
<emphasis id="B9530D42FFEBFF9F3590F903FC79AD90" box="[740,959,1691,1716]" italics="true" pageId="16" pageNumber="17">Ochetostoma edax</emphasis>
</taxonomicName>
(host collected and preserved), extreme low tide,
<date id="FF99F790FFEBFF9F3671F95AFE58ADF8" box="[261,414,1730,1756]" pageId="16" pageNumber="17" value="2007-03-22">22 Mar 2007</date>
, coll. A. Anker [fcn 07-177].
</paragraph>
</subSubSection>
<subSubSection id="C33D82DBFFEBFF9F37B2F972FB58ACD0" pageId="16" pageNumber="17" type="description">
<paragraph id="8B98D150FFEBFF9F37B2F972FC72AC20" blockId="16.[151,1437,1530,2036]" box="[198,948,1770,1796]" pageId="16" pageNumber="17">
<emphasis id="B9530D42FFEBFF9F37B2F972FE91AC20" bold="true" box="[198,343,1770,1796]" pageId="16" pageNumber="17">Description</emphasis>
. For detailed description see
<bibRefCitation id="EFB6ACA1FFEBFF9F35C3F972FC76AC20" author="Kim" box="[695,944,1770,1796]" pageId="16" pageNumber="23" refString="Kim, W. &amp; Abele, L. G. (1988) The snapping shrimp genus Alpheus from the eastern Pacific (Decapoda: Caridea: Alpheidae). Smithsonian Contributions to Zoology, 454, 1 - 119." type="journal article" year="1988">Kim &amp; Abele (1988)</bibRefCitation>
.
</paragraph>
<paragraph id="8B98D150FFEBFF9F37B2F88AFC88AC70" blockId="16.[151,1437,1530,2036]" pageId="16" pageNumber="17">
<emphasis id="B9530D42FFEBFF9F37B2F88AFECAAC08" bold="true" box="[198,268,1810,1836]" pageId="16" pageNumber="17">Color</emphasis>
. Mostly ivory-whitish with pinkish tinge in some areas due to presence of scattered reddish chromatophores (
<figureCitation id="131CCDD5FFEBFF9F365BF8A2FE44AC70" box="[303,386,1850,1876]" captionStart="FIGURE 11" captionStartId="18.[151,255,1495,1519]" captionTargetBox="[151,1434,434,1471]" captionTargetId="figure@18.[151,1436,434,1471]" captionTargetPageId="18" captionText="FIGURE 11. Color patterns in life. Alpheus christofferseni n. sp., male from Bocas del Toro, Panama (USNM 1100667) (a – c): a, b, dorsal view (note partly faded chromatopores in a); c, same, lateral view. Alpheus naos n. sp., male holotype from Punta Culebra, Isla Naos, Panama (USNM 1100668) (d, e): d, dorsal view; e, lateral view. Alpheus aequus Kim &amp; Abele, 1988, female from Coibita, Coiba Marine National Park, Panama (MNHN-Na 16389) (f): dorsal view." httpUri="https://zenodo.org/record/179301/files/figure.png" pageId="16" pageNumber="17">Fig. 11</figureCitation>
f); one female had yellow-orange eggs.
</paragraph>
<paragraph id="8B98D150FFEBFF9F37B2F8FAFB58ACD0" blockId="16.[151,1437,1530,2036]" pageId="16" pageNumber="17">
<emphasis id="B9530D42FFEBFF9F37B2F8FAFF3FAC58" bold="true" box="[198,249,1890,1916]" pageId="16" pageNumber="17">Size</emphasis>
. The CL of the two females is
<quantity id="4CDF7CB5FFEBFF9F3514F8FAFD7CAC58" box="[608,698,1890,1916]" metricMagnitude="-3" metricUnit="m" metricValue="4.7" pageId="16" pageNumber="17" unit="mm" value="4.7">4.7 mm</quantity>
and
<quantity id="4CDF7CB5FFEBFF9F3585F8FAFC8BAC58" box="[753,845,1890,1916]" metricMagnitude="-3" metricUnit="m" metricValue="4.9" pageId="16" pageNumber="17" unit="mm" value="4.9">4.9 mm</quantity>
, the TL for both is
<quantity id="4CDF7CB5FFEBFF9F3358F8FAFB5EAC58" box="[1068,1176,1890,1916]" metricMagnitude="-2" metricUnit="m" metricValue="1.43" pageId="16" pageNumber="17" unit="mm" value="14.3">14.3 mm</quantity>
. The single male originally reported by
<bibRefCitation id="EFB6ACA1FFEBFF9F3606F812FDADAC80" author="Kim" box="[370,619,1930,1956]" pageId="16" pageNumber="23" refString="Kim, W. &amp; Abele, L. G. (1988) The snapping shrimp genus Alpheus from the eastern Pacific (Decapoda: Caridea: Alpheidae). Smithsonian Contributions to Zoology, 454, 1 - 119." type="journal article" year="1988">Kim &amp; Abele (1988)</bibRefCitation>
had a CL of
<quantity id="4CDF7CB5FFEBFF9F347FF812FCAFAC80" box="[779,873,1930,1956]" metricMagnitude="-3" metricUnit="m" metricValue="4.3" pageId="16" pageNumber="17" unit="mm" value="4.3">4.3 mm</quantity>
, whereas the two females both had a CL of
<quantity id="4CDF7CB5FFEBFF9F320CF812FF0FACE8" metricMagnitude="-3" metricUnit="m" metricValue="6.8" pageId="16" pageNumber="17" unit="mm" value="6.8">6.8 mm</quantity>
. Ramos’ (1995) three specimens were smaller, with
<quantity id="4CDF7CB5FFEBFF9F345AF82AFC4EACE8" box="[814,904,1970,1996]" metricMagnitude="-3" metricUnit="m" metricValue="2.9" pageId="16" pageNumber="17" unit="mm" value="2.9">2.9 mm</quantity>
CL in the single male and
<quantity id="4CDF7CB5FFEBFF9F33B1F82AFA94ACE8" box="[1221,1362,1970,1996]" metricMagnitude="-3" metricUnit="m" metricValue="4.35" metricValueMax="4.4" metricValueMin="4.3" pageId="16" pageNumber="17" unit="mm" value="4.35" valueMax="4.4" valueMin="4.3">4.3–4.4 mm</quantity>
in the two females. Based on these data, the CL of adult
<taxonomicName id="4C27AAD3FFEBFF9F3590F843FC9EACD0" box="[740,856,2011,2036]" class="Malacostraca" family="Alpheidae" genus="Alpheus" kingdom="Animalia" order="Decapoda" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="aequus">
<emphasis id="B9530D42FFEBFF9F3590F843FC9EACD0" box="[740,856,2011,2036]" italics="true" pageId="16" pageNumber="17">A. aequus</emphasis>
</taxonomicName>
ranges from
<quantity id="4CDF7CB5FFEBFF9F3487F842FB5CACD0" box="[1011,1178,2010,2036]" metricMagnitude="-3" metricUnit="m" metricValue="4.85" metricValueMax="6.8" metricValueMin="2.9" pageId="16" pageNumber="17" unit="mm" value="4.85" valueMax="6.8" valueMin="2.9">2.9 to 6.8 mm</quantity>
.
</paragraph>
</subSubSection>
<caption id="DF5881D8FFEAFF9E37E3FA2CFB93AD35" httpUri="https://zenodo.org/record/179300/files/figure.png" pageId="17" pageNumber="18" targetBox="[172,1431,204,1436]" targetPageId="17">
<paragraph id="8B98D150FFEAFF9E37E3FA2CFB93AD35" blockId="17.[151,1436,1460,1553]" pageId="17" pageNumber="18">
<emphasis id="B9530D42FFEAFF9E37E3FA2CFEE1AEE8" bold="true" box="[151,295,1460,1484]" pageId="17" pageNumber="18">FIGURE 10.</emphasis>
<taxonomicName id="4C27AAD3FFEAFF9E3659FA2DFD7CAEE8" authority="Kim &amp; Abeke, 1988" authorityName="Kim &amp; Abeke" authorityYear="1988" box="[301,698,1461,1484]" class="Malacostraca" family="Alpheidae" genus="Alpheus" kingdom="Animalia" order="Decapoda" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="aequus">
<emphasis id="B9530D42FFEAFF9E3659FA2DFE13AEE8" box="[301,469,1461,1484]" italics="true" pageId="17" pageNumber="18">Alpheus aequus</emphasis>
Kim &amp; Abeke, 1988
</taxonomicName>
, female from Coibita, Coiba Marine National Park, Panama (MNHN 16389): a, frontal margin, dorsal view; b, major cheliped, lateral view; c, same, chela, mesial view; d, minor chela, lateral view; e, second pereiopod, lateral view; f, third pereiopod, lateral view. Scale bars: 1 mm.
</paragraph>
</caption>
<subSubSection id="C33D82DBFFEAFF9E37B2F9DDFC7FAD7B" box="[198,953,1605,1631]" pageId="17" pageNumber="18" type="materials_examined">
<paragraph id="8B98D150FFEAFF9E37B2F9DDFC7FAD7B" blockId="17.[151,1437,1605,2031]" box="[198,953,1605,1631]" pageId="17" pageNumber="18">
<emphasis id="B9530D42FFEAFF9E37B2F9DDFEA2AD7B" bold="true" box="[198,356,1605,1631]" pageId="17" pageNumber="18">
<typeStatus id="549C6FF2FFEAFF9E37B2F9DDFEC2AD7B" box="[198,260,1605,1631]" pageId="17" pageNumber="18">Type</typeStatus>
locality
</emphasis>
. Playa Blancas,
<collectingCountry id="F33091C0FFEAFF9E355AF9DDFD68AD7B" box="[558,686,1605,1631]" name="Costa Rica" pageId="17" pageNumber="18">Costa Rica</collectingCountry>
(
<bibRefCitation id="EFB6ACA1FFEAFF9E35C9F9DDFC6AAD7B" author="Kim" box="[701,940,1605,1631]" pageId="17" pageNumber="23" refString="Kim, W. &amp; Abele, L. G. (1988) The snapping shrimp genus Alpheus from the eastern Pacific (Decapoda: Caridea: Alpheidae). Smithsonian Contributions to Zoology, 454, 1 - 119." type="journal article" year="1988">Kim &amp; Abele, 1988</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C33D82DBFFEAFF9E37B2F9F5FA5AACCA" pageId="17" pageNumber="18" type="distribution">
<paragraph id="8B98D150FFEAFF9E37B2F9F5FABCAD8B" blockId="17.[151,1437,1605,2031]" pageId="17" pageNumber="18">
<emphasis id="B9530D42FFEAFF9E37B2F9F5FE99ADA3" bold="true" box="[198,351,1645,1671]" pageId="17" pageNumber="18">Distribution</emphasis>
. Eastern Pacific:
<collectingCountry id="F33091C0FFEAFF9E354CF9F5FD06ADA3" box="[568,704,1645,1671]" name="Costa Rica" pageId="17" pageNumber="18">Costa Rica</collectingCountry>
: Playa Blancas; Galapagos: Santa Fe (
<bibRefCitation id="EFB6ACA1FFEAFF9E33E1F9F5FA48ADA3" author="Kim" box="[1173,1422,1645,1671]" pageId="17" pageNumber="23" refString="Kim, W. &amp; Abele, L. G. (1988) The snapping shrimp genus Alpheus from the eastern Pacific (Decapoda: Caridea: Alpheidae). Smithsonian Contributions to Zoology, 454, 1 - 119." type="journal article" year="1988">Kim &amp; Abele, 1988</bibRefCitation>
);
<collectingCountry id="F33091C0FFEAFF9E37E3F90DFF3FAD8B" box="[151,249,1685,1711]" name="Panama" pageId="17" pageNumber="18">Panama</collectingCountry>
: Coibita (Coiba Marine National Park) (present study);
<collectingCountry id="F33091C0FFEAFF9E34F1F90DFC3BAD8B" box="[901,1021,1685,1711]" name="Colombia" pageId="17" pageNumber="18">Colombia</collectingCountry>
: Gorgona Island (
<bibRefCitation id="EFB6ACA1FFEAFF9E33B9F90DFAABAD8B" author="Ramos" box="[1229,1389,1685,1711]" pageId="17" pageNumber="23" refString="Ramos, G. (1995) Nuevos registros de camarones alfeidos (Crustacea, Decapoda, Alpheidae) para el Pacifico de Colombia. In: Cantera, J. R. &amp; Restrepo, J. D. (eds.), Delta del Rio San Juan, Bahias de Malaga y Buenaventura, Pacifico Colombiano. Tomo II, pp. 127 - 153. Colciencias, Univeridad EAFIT y Universidad del Valle, Colombia." type="book chapter" year="1995">Ramos, 1995</bibRefCitation>
).
</paragraph>
<paragraph id="8B98D150FFEAFF9E37B2F925FA5AACCA" blockId="17.[151,1437,1605,2031]" pageId="17" pageNumber="18">
<emphasis id="B9530D42FFEAFF9E37B2F925FEEEADF3" bold="true" box="[198,296,1725,1751]" pageId="17" pageNumber="18">Ecology</emphasis>
. The habitat of the
<typeStatus id="549C6FF2FFEAFF9E3560F925FDBCADF3" box="[532,634,1725,1751]" pageId="17" pageNumber="18" type="holotype">holotype</typeStatus>
from Playa Blancas,
<collectingCountry id="F33091C0FFEAFF9E340EF925FC39ADF3" box="[890,1023,1725,1751]" name="Costa Rica" pageId="17" pageNumber="18">Costa Rica</collectingCountry>
, was described as “shore to 5.0 m; shale beach” and “shale beach between beach and rocky reef” (
<bibRefCitation id="EFB6ACA1FFEAFF9E340FF97DFBADADDB" author="Kim" box="[891,1131,1765,1791]" pageId="17" pageNumber="23" refString="Kim, W. &amp; Abele, L. G. (1988) The snapping shrimp genus Alpheus from the eastern Pacific (Decapoda: Caridea: Alpheidae). Smithsonian Contributions to Zoology, 454, 1 - 119." type="journal article" year="1988">Kim &amp; Abele, 1988</bibRefCitation>
). Other alpheids collected in this area were
<taxonomicName id="4C27AAD3FFEAFF9E361EF895FCD7AC03" authority="Lockington 1878" authorityName="Lockington" authorityYear="1878" box="[362,785,1805,1831]" class="Malacostraca" family="Alpheidae" genus="Alpheus" kingdom="Animalia" order="Decapoda" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="fasciatus">
<emphasis id="B9530D42FFEAFF9E361EF895FDFDAC02" box="[362,571,1805,1830]" italics="true" pageId="17" pageNumber="18">Alpheus fasciatus</emphasis>
Lockington 1878
</taxonomicName>
(reported as
<taxonomicName id="4C27AAD3FFEAFF9E34C7F895FAC4AC03" authority="Miers, 1888" authorityName="Miers" authorityYear="1888" box="[947,1282,1805,1831]" class="Malacostraca" family="Alpheidae" genus="Alpheus" kingdom="Animalia" order="Decapoda" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="paracrinitus">
<emphasis id="B9530D42FFEAFF9E34C7F895FBA1AC02" box="[947,1127,1805,1830]" italics="true" pageId="17" pageNumber="18">A. paracrinitus</emphasis>
Miers, 1888
</taxonomicName>
) and
<taxonomicName id="4C27AAD3FFEAFF9E323DF895FE09AC6B" authority="Sivertsen, 1933" authorityName="Sivertsen" authorityYear="1933" class="Malacostraca" family="Alpheidae" genus="Alpheus" kingdom="Animalia" order="Decapoda" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="galapagensis">
<emphasis id="B9530D42FFEAFF9E323DF895FECAAC6A" italics="true" pageId="17" pageNumber="18">A. galapagensis</emphasis>
Sivertsen, 1933
</taxonomicName>
(reported as
<taxonomicName id="4C27AAD3FFEAFF9E3505F8ADFC2EAC6B" authority="Kim &amp; Abele, 1988" authorityName="Kim &amp; Abele" authorityYear="1988" box="[625,1000,1845,1871]" class="Malacostraca" family="Alpheidae" genus="Alpheus" kingdom="Animalia" order="Decapoda" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="canalis">
<emphasis id="B9530D42FFEAFF9E3505F8ADFD2FAC6A" box="[625,745,1845,1870]" italics="true" pageId="17" pageNumber="18">A. canalis</emphasis>
<bibRefCitation id="EFB6ACA1FFEAFF9E3587F8ADFC2EAC6B" author="Kim" box="[755,1000,1845,1871]" pageId="17" pageNumber="23" refString="Kim, W. &amp; Abele, L. G. (1988) The snapping shrimp genus Alpheus from the eastern Pacific (Decapoda: Caridea: Alpheidae). Smithsonian Contributions to Zoology, 454, 1 - 119." type="journal article" year="1988">Kim &amp; Abele, 1988</bibRefCitation>
</taxonomicName>
). The two specimens from Santa Fe (Barrington)
<collectingCountry id="F33091C0FFEAFF9E3644F8C5FEBDAC53" box="[304,379,1885,1911]" name="Iceland" pageId="17" pageNumber="18">Island</collectingCountry>
, Galapagos, were collected while “diving in bay, [at] about
<quantity id="4CDF7CB5FFEAFF9E3346F8C5FBA2AC53" box="[1074,1124,1885,1911]" metricMagnitude="0" metricUnit="m" metricValue="5.0" pageId="17" pageNumber="18" unit="m" value="5.0">5 m</quantity>
”. Other alpheids occurring in this locality were
<taxonomicName id="4C27AAD3FFEAFF9E36E2F81DFCA6ACBB" authority="Lockington, 1878" authorityName="Lockington" authorityYear="1878" box="[406,864,1925,1951]" class="Malacostraca" family="Alpheidae" genus="Alpheus" kingdom="Animalia" order="Decapoda" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="bellimanus">
<emphasis id="B9530D42FFEAFF9E36E2F81DFD46ACBA" box="[406,640,1925,1950]" italics="true" pageId="17" pageNumber="18">Alpheus bellimanus</emphasis>
Lockington, 1878
</taxonomicName>
and
<taxonomicName id="4C27AAD3FFEAFF9E34D5F81DFAF4ACBB" authority="Kim &amp; Abele, 1988" authorityName="Kim &amp; Abele" authorityYear="1988" box="[929,1330,1925,1951]" class="Malacostraca" family="Alpheidae" genus="Alpheus" kingdom="Animalia" order="Decapoda" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="rostratus">
<emphasis id="B9530D42FFEAFF9E34D5F81DFBEBACBA" box="[929,1069,1925,1950]" italics="true" pageId="17" pageNumber="18">A. rostratus</emphasis>
<bibRefCitation id="EFB6ACA1FFEAFF9E334DF81DFAF4ACBB" author="Kim" box="[1081,1330,1925,1951]" pageId="17" pageNumber="23" refString="Kim, W. &amp; Abele, L. G. (1988) The snapping shrimp genus Alpheus from the eastern Pacific (Decapoda: Caridea: Alpheidae). Smithsonian Contributions to Zoology, 454, 1 - 119." type="journal article" year="1988">Kim &amp; Abele, 1988</bibRefCitation>
</taxonomicName>
. Ramos’ (1995) specimens came from a tidal beach with coral sand (“playas de arena coralina”). No particular associations were noted by
<bibRefCitation id="EFB6ACA1FFEAFF9E36F3F84DFDB8ACCB" author="Kim" box="[391,638,2005,2031]" pageId="17" pageNumber="23" refString="Kim, W. &amp; Abele, L. G. (1988) The snapping shrimp genus Alpheus from the eastern Pacific (Decapoda: Caridea: Alpheidae). Smithsonian Contributions to Zoology, 454, 1 - 119." type="journal article" year="1988">Kim &amp; Abele (1988)</bibRefCitation>
or
<bibRefCitation id="EFB6ACA1FFEAFF9E35D2F84DFC94ACCB" author="Ramos" box="[678,850,2005,2031]" pageId="17" pageNumber="23" refString="Ramos, G. (1995) Nuevos registros de camarones alfeidos (Crustacea, Decapoda, Alpheidae) para el Pacifico de Colombia. In: Cantera, J. R. &amp; Restrepo, J. D. (eds.), Delta del Rio San Juan, Bahias de Malaga y Buenaventura, Pacifico Colombiano. Tomo II, pp. 127 - 153. Colciencias, Univeridad EAFIT y Universidad del Valle, Colombia." type="book chapter" year="1995">Ramos (1995)</bibRefCitation>
. In contrast, the two Panamanian specimens of
<emphasis id="B9530D42FFEAFF9E32F7F84DFA5AACCA" box="[1411,1436,2005,2030]" italics="true" pageId="17" pageNumber="18">A.</emphasis>
</paragraph>
</subSubSection>
<subSubSection id="C33D82DBFFE9FF9D37E3FF00FC50ACC0" pageId="18" pageNumber="19" type="description">
<paragraph id="8B98D150FFE9FF9D37E3FF00FC19AA5E" blockId="18.[151,1436,152,378]" pageId="18" pageNumber="19">
<taxonomicName id="4C27AAD3FFE9FF9D37E3FF00FF2CAB95" box="[151,234,152,177]" class="Malacostraca" family="Alpheidae" genus="Alpheus" kingdom="Animalia" order="Decapoda" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="aequus">
<emphasis id="B9530D42FFE9FF9D37E3FF00FF2CAB95" box="[151,234,152,177]" italics="true" pageId="18" pageNumber="19">aequus</emphasis>
</taxonomicName>
were found associated with medium-sized (body length
<quantity id="4CDF7CB5FFE9FF9D34F4FF00FC10AB96" box="[896,982,152,178]" metricMagnitude="-2" metricUnit="m" metricValue="2.8" pageId="18" pageNumber="19" unit="mm" value="28.0">28 mm</quantity>
, diameter
<quantity id="4CDF7CB5FFE9FF9D3324FF00FB16AB96" box="[1104,1232,152,178]" metricMagnitude="-2" metricUnit="m" metricValue="1.15" metricValueMax="1.2" metricValueMin="1.1" pageId="18" pageNumber="19" unit="mm" value="11.5" valueMax="12.0" valueMin="11.0">11–12 mm</quantity>
, proboscis length
<quantity id="4CDF7CB5FFE9FF9D37E3FF58FF2BABFE" box="[151,237,192,218]" metricMagnitude="-2" metricUnit="m" metricValue="1.5" pageId="18" pageNumber="19" unit="mm" value="15.0">15 mm</quantity>
, proboscis diameter
<quantity id="4CDF7CB5FFE9FF9D3694FF58FD82ABFE" box="[480,580,192,218]" metricMagnitude="-3" metricUnit="m" metricValue="4.0" metricValueMax="5.0" metricValueMin="3.0" pageId="18" pageNumber="19" unit="mm" value="4.0" valueMax="5.0" valueMin="3.0">3–5 mm</quantity>
) thalassematid echiurans,
<taxonomicName id="4C27AAD3FFE9FF9D340EFF58FAC7ABFE" authority="Fisher, 1926" authorityName="Fisher" authorityYear="1926" box="[890,1281,192,218]" class="Polychaeta" family="Echiuridae" genus="Ochetostoma" higherTaxonomySource="CoL" kingdom="Animalia" order="Echiuroidea" pageId="18" pageNumber="19" phylum="Echiura" rank="species" species="edax">
<emphasis id="B9530D42FFE9FF9D340EFF58FB92ABFD" box="[890,1108,192,217]" italics="true" pageId="18" pageNumber="19">Ochetostoma edax</emphasis>
(Fisher, 1926)
</taxonomicName>
(
<figureCitation id="131CCDD5FFE9FF9D3266FF58FAA1ABFE" box="[1298,1383,192,218]" captionStart="FIGURE 12" captionStartId="19.[151,255,1786,1810]" captionTargetBox="[151,1436,194,1760]" captionTargetId="figure@19.[151,1436,194,1762]" captionTargetPageId="19" captionText="FIGURE 12. Habitats and hosts: a, shallow subtidal sandflat with patches of flotting algae and seagrass at Carenero Island, Bocas del Toro, Panama, type locality of Alpheus christofferseni n. sp.; b, c, Alpheus christofferseni n. sp. with host, Ochetostoma cf. edax (Fisher, 1946) (Echiura, Thalassematidae), a few hours after collection; d, partly exposed rock-mud intertidal at Coibita, Coiba Marine National Park, Panama, one of collection sites of Alpheus aequus Kim &amp; Abele, 1988; e, Ochetostoma edax (Echiura, Thalassematidae), host of the Coibita specimen (MNHN-Na 16389)." httpUri="https://zenodo.org/record/179302/files/figure.png" pageId="18" pageNumber="19">Fig. 12</figureCitation>
e) in a mixed mud-rock intertidal of Coibita (
<figureCitation id="131CCDD5FFE9FF9D351BFF70FD03AA26" box="[623,709,232,258]" captionStart="FIGURE 12" captionStartId="19.[151,255,1786,1810]" captionTargetBox="[151,1436,194,1760]" captionTargetId="figure@19.[151,1436,194,1762]" captionTargetPageId="19" captionText="FIGURE 12. Habitats and hosts: a, shallow subtidal sandflat with patches of flotting algae and seagrass at Carenero Island, Bocas del Toro, Panama, type locality of Alpheus christofferseni n. sp.; b, c, Alpheus christofferseni n. sp. with host, Ochetostoma cf. edax (Fisher, 1946) (Echiura, Thalassematidae), a few hours after collection; d, partly exposed rock-mud intertidal at Coibita, Coiba Marine National Park, Panama, one of collection sites of Alpheus aequus Kim &amp; Abele, 1988; e, Ochetostoma edax (Echiura, Thalassematidae), host of the Coibita specimen (MNHN-Na 16389)." httpUri="https://zenodo.org/record/179302/files/figure.png" pageId="18" pageNumber="19">Fig. 12</figureCitation>
d). The burrows of
<taxonomicName id="4C27AAD3FFE9FF9D34DCFF70FBC2AA25" box="[936,1028,232,257]" class="Polychaeta" family="Echiuridae" genus="Ochetostoma" higherTaxonomySource="CoL" kingdom="Animalia" order="Echiuroidea" pageId="18" pageNumber="19" phylum="Echiura" rank="species" species="edax">
<emphasis id="B9530D42FFE9FF9D34DCFF70FBC2AA25" box="[936,1028,232,257]" italics="true" pageId="18" pageNumber="19">O. edax</emphasis>
</taxonomicName>
are smoothly lined horizontal tunnels dug in muddy sand under partly mud-embedded rocks. The shrimps were observed sitting just next to their hosts. Notably, in both cases, only one shrimp per host was found, despite some efforts to find its potentially present mate by prospecting the immediate vicinity of the burrow.
</paragraph>
<caption id="DF5881D8FFE9FF9D37E3FA4FFAE1AD73" httpUri="https://zenodo.org/record/179301/files/figure.png" pageId="18" pageNumber="19" targetBox="[151,1434,434,1471]" targetPageId="18">
<paragraph id="8B98D150FFE9FF9D37E3FA4FFAE1AD73" blockId="18.[151,1436,1495,1623]" pageId="18" pageNumber="19">
<emphasis id="B9530D42FFE9FF9D37E3FA4FFED8AECB" bold="true" box="[151,286,1495,1519]" pageId="18" pageNumber="19">FIGURE 11</emphasis>
. Color patterns in life.
<taxonomicName id="4C27AAD3FFE9FF9D3562FA40FCCAAECB" box="[534,780,1496,1519]" class="Malacostraca" family="Alpheidae" genus="Alpheus" kingdom="Animalia" order="Decapoda" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="christofferseni" status="sp. nov.">
<emphasis id="B9530D42FFE9FF9D3562FA40FCCAAECB" box="[534,780,1496,1519]" italics="true" pageId="18" pageNumber="19">Alpheus christofferseni</emphasis>
</taxonomicName>
<emphasis id="B9530D42FFE9FF9D3467FA4FFC89AECB" bold="true" box="[787,847,1495,1519]" pageId="18" pageNumber="19">
<taxonomicNameLabel id="A260B039FFE9FF9D3467FA4FFC89AECB" box="[787,847,1495,1519]" pageId="18" pageNumber="19" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
, male from Bocas del Toro, Panama (USNM 1100667) (a–c): a, b, dorsal view (note partly faded chromatopores in a); c, same, lateral view.
<taxonomicName id="4C27AAD3FFE9FF9D3350FA62FB74AD35" box="[1060,1202,1530,1553]" class="Malacostraca" family="Alpheidae" genus="Alpheus" kingdom="Animalia" order="Decapoda" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="naos" status="sp. nov.">
<emphasis id="B9530D42FFE9FF9D3350FA62FB74AD35" box="[1060,1202,1530,1553]" italics="true" pageId="18" pageNumber="19">Alpheus naos</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A260B039FFE9FF9D33CEFA62FB30AD36" box="[1210,1270,1530,1554]" pageId="18" pageNumber="19" rank="species">
<emphasis id="B9530D42FFE9FF9D33CEFA62FB30AD36" bold="true" box="[1210,1270,1530,1554]" pageId="18" pageNumber="19">n. sp.</emphasis>
</taxonomicNameLabel>
, male holotype from Punta Culebra, Isla
<taxonomicName id="4C27AAD3FFE9FF9D36D2F985FE19AD10" box="[422,479,1565,1588]" class="Malacostraca" family="Alpheidae" genus="Alpheus" kingdom="Animalia" order="Decapoda" pageId="18" pageNumber="22" phylum="Arthropoda" rank="species" species="naos" status="sp. nov.">Naos</taxonomicName>
, Panama (USNM 1100668) (d, e): d, dorsal view; e, lateral view.
<taxonomicName id="4C27AAD3FFE9FF9D33D5F985FEDAAD73" authority="Kim &amp; Abele, 1988" authorityName="Kim &amp; Abele" authorityYear="1988" class="Malacostraca" family="Alpheidae" genus="Alpheus" kingdom="Animalia" order="Decapoda" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="aequus">
<emphasis id="B9530D42FFE9FF9D33D5F985FA8CAD10" box="[1185,1354,1565,1588]" italics="true" pageId="18" pageNumber="19">Alpheus aequus</emphasis>
Kim &amp; Abele, 1988
</taxonomicName>
, female from Coibita, Coiba Marine National Park, Panama (MNHN-Na 16389) (f): dorsal view.
</paragraph>
</caption>
<paragraph id="8B98D150FFE9FF9D37B2F912FE76AC38" blockId="18.[151,1436,1674,2020]" pageId="18" pageNumber="19">
<emphasis id="B9530D42FFE9FF9D37B2F912FDE6AD80" bold="true" box="[198,544,1674,1700]" pageId="18" pageNumber="19">Taxonomic remarks on host</emphasis>
. The finding of two specimens of
<taxonomicName id="4C27AAD3FFE9FF9D34C4F913FB4FAD80" box="[944,1161,1675,1700]" class="Polychaeta" family="Echiuridae" genus="Ochetostoma" higherTaxonomySource="CoL" kingdom="Animalia" order="Echiuroidea" pageId="18" pageNumber="19" phylum="Echiura" rank="species" species="edax">
<emphasis id="B9530D42FFE9FF9D34C4F913FB4FAD80" box="[944,1161,1675,1700]" italics="true" pageId="18" pageNumber="19">Ochetostoma edax</emphasis>
</taxonomicName>
(identification by G.-V. Murina) at Coibita represents a considerable extension of the distribution range of this species from Baja California to
<collectingCountry id="F33091C0FFE9FF9D367AF942FEA9ADD0" box="[270,367,1754,1780]" name="Panama" pageId="18" pageNumber="19">Panama</collectingCountry>
. The identity of
<taxonomicName id="4C27AAD3FFE9FF9D3541F943FD7FADD0" box="[565,697,1754,1780]" class="Polychaeta" family="Echiuridae" genus="Ochetostoma" higherTaxonomySource="CoL" kingdom="Animalia" order="Echiuroidea" pageId="18" pageNumber="19" phylum="Echiura" rank="species" species="edax">
<emphasis id="B9530D42FFE9FF9D3541F943FD8CADD0" box="[565,586,1755,1780]" italics="true" pageId="18" pageNumber="19">O</emphasis>
. cf.
<emphasis id="B9530D42FFE9FF9D35F6F943FD7FADD0" box="[642,697,1755,1780]" italics="true" pageId="18" pageNumber="19">edax</emphasis>
</taxonomicName>
from the Atlantic (Caribbean) coast of
<collectingCountry id="F33091C0FFE9FF9D33E1F942FB34ADD0" box="[1173,1266,1754,1780]" name="Panama" pageId="18" pageNumber="19">Panama</collectingCountry>
remains to be determined (see above).
</paragraph>
<paragraph id="8B98D150FFE9FF9D37B2F8B2FD6AAC98" blockId="18.[151,1436,1674,2020]" pageId="18" pageNumber="19">
<emphasis id="B9530D42FFE9FF9D37B2F8B2FEFFAC60" bold="true" box="[198,313,1834,1860]" pageId="18" pageNumber="19">Variation</emphasis>
. The two female specimens from Coibita agree well with the description and illustrations of
<taxonomicName id="4C27AAD3FFE9FF9D32F7F8B3FF2CAC48" class="Malacostraca" family="Alpheidae" genus="Alpheus" kingdom="Animalia" order="Decapoda" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="aequus">
<emphasis id="B9530D42FFE9FF9D32F7F8B3FF2CAC48" italics="true" pageId="18" pageNumber="19">A. aequus</emphasis>
</taxonomicName>
provided by
<bibRefCitation id="EFB6ACA1FFE9FF9D36FBF8CAFD57AC48" author="Kim" box="[399,657,1874,1900]" pageId="18" pageNumber="23" refString="Kim, W. &amp; Abele, L. G. (1988) The snapping shrimp genus Alpheus from the eastern Pacific (Decapoda: Caridea: Alpheidae). Smithsonian Contributions to Zoology, 454, 1 - 119." type="journal article" year="1988">Kim &amp; Abele (1988)</bibRefCitation>
, except for the length of the dactylus of the major chela, which reaches only slightly beyond the tip of the pollex in the Coibita specimens, and distinctly beyond the pollex in the
<typeStatus id="549C6FF2FFE9FF9D37B6F83AFEEEAC98" box="[194,296,1954,1980]" pageId="18" pageNumber="19" type="holotype">holotype</typeStatus>
(cf.
<bibRefCitation id="EFB6ACA1FFE9FF9D362BF83AFD85AC98" author="Kim" box="[351,579,1954,1980]" pageId="18" pageNumber="23" refString="Kim, W. &amp; Abele, L. G. (1988) The snapping shrimp genus Alpheus from the eastern Pacific (Decapoda: Caridea: Alpheidae). Smithsonian Contributions to Zoology, 454, 1 - 119." type="journal article" year="1988">Kim &amp; Abele 1988</bibRefCitation>
, fig. 23).
</paragraph>
<paragraph id="8B98D150FFE9FF9D37B2F852FC50ACC0" blockId="18.[151,1436,1674,2020]" box="[198,918,1994,2020]" pageId="18" pageNumber="19">
<emphasis id="B9530D42FFE9FF9D37B2F852FE60ACC0" bold="true" box="[198,422,1994,2020]" pageId="18" pageNumber="19">GenBank number</emphasis>
.
<accessionNumber id="94744CB3FFE9FF9D36C8F852FDFDACC0" box="[444,571,1994,2020]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/EU084881" pageId="18" pageNumber="19">EU084881</accessionNumber>
(fcn 07-109, MNHN 16388).
</paragraph>
</subSubSection>
</treatment>
</document>
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