treatments-xml/data/03/3A/95/033A9525464BFFF077DBA4AFBBB9A6A5.xml
2024-06-21 12:22:17 +02:00

455 lines
55 KiB
XML
Raw Blame History

This file contains ambiguous Unicode characters

This file contains Unicode characters that might be confused with other characters. If you think that this is intentional, you can safely ignore this warning. Use the Escape button to reveal them.

<document id="DDC28AAF657A7872D1A5B9D23F96539D" ID-DOI="10.5281/zenodo.186587" ID-GBIF-Dataset="f979f9ce-284c-46d4-a6b0-25dfc8f77007" ID-ISSN="1175-5326" ID-Zenodo-Dep="186587" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1461053329372" checkinUser="plazi" docAuthor="Gawor, Anna, Hendrix, Ralf, Vences, Miguel, Böhme, Wolfgang &amp; Ziegler, Thomas" docDate="2009" docId="033A9525464BFFF077DBA4AFBBB9A6A5" docLanguage="en" docName="zt02051p025.pdf" docOrigin="Zootaxa 2051" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Hylarana maosonensis Bourret 1937" docType="treatment" docVersion="7" lastPageNumber="11" masterDocId="FF03ED5D464CFFFA774CA101BB6AA157" masterDocTitle="Larval morphology in four species of Hylarana from Vietnam and Thailand with comments on the taxonomy of H. nigrovittata sensu latu (Anura: Ranidae)" masterLastPageNumber="25" masterPageNumber="1" pageNumber="8" updateTime="1698580311479" updateUser="plazi">
<mods:mods id="7FC52488B6BB7BBED067323695E67D44" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="D72A2998D49F01F3524015AA8048D15A">
<mods:title id="6E1CE7422246910537CD35FD84D9A5C1">Larval morphology in four species of Hylarana from Vietnam and Thailand with comments on the taxonomy of H. nigrovittata sensu latu (Anura: Ranidae)</mods:title>
</mods:titleInfo>
<mods:name id="40DB5E94E6D5E9124CDD43D064C62053" type="personal">
<mods:role id="CEC704ED89DBC9F650E54DDC1D39719C">
<mods:roleTerm id="2277DE68AE7B18E1478CCA888767B1CD">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="56BF32A73E613C64CBE794867E52EB70">Gawor, Anna</mods:namePart>
</mods:name>
<mods:name id="31EBCEE6DAAA21AA2817D982EA20FFBA" type="personal">
<mods:role id="AB6F8277F65141C6A26C18227223F889">
<mods:roleTerm id="93E1450712C72A51C83FF0D9DFE20C5C">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="2202404834B880C07E22BB456ECC10D8">Hendrix, Ralf</mods:namePart>
</mods:name>
<mods:name id="DB21C8F5AC383434442288375D5B58B4" type="personal">
<mods:role id="28232D37B8CF95A06C2B767F15D70214">
<mods:roleTerm id="EBF220EA67225A90F7DB07212E028AAD">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="9B2F7A4D432A22E8723D6B9C4E4E1736">Vences, Miguel</mods:namePart>
</mods:name>
<mods:name id="A85DE4041F41F66B61549F5BCA793C12" type="personal">
<mods:role id="92DD239F3D7252DADA647B7D56A53D24">
<mods:roleTerm id="8A7995ED9DCED0025B4EDA81CD46AB40">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="64598CCF11BD13074A8D4F9E6B83FDAE">Böhme, Wolfgang</mods:namePart>
</mods:name>
<mods:name id="91984C6FF982C6BA6698EBB542DF46A7" type="personal">
<mods:role id="E2D47FED3E7592E1FCB3083AE04D69CC">
<mods:roleTerm id="27751FF131A49897F67F167952024C00">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="2286EFC473D5CE55ACCA0015C6984C1A">Ziegler, Thomas</mods:namePart>
</mods:name>
<mods:typeOfResource id="E250E7C9F12FB232F572BBACF972F459">text</mods:typeOfResource>
<mods:relatedItem id="43261E8547F42D67115C56ED815F3CA1" type="host">
<mods:titleInfo id="48FF0383A70F303BF6D5DDA7DFF4E5D4">
<mods:title id="06D609D94E6F81CEF3D73385E1166EDC">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="FDF23200606A7B77CB0E2C8673B0E435">
<mods:date id="3A823D56A09DEB9C3D94EE65A5915308">2009</mods:date>
<mods:detail id="F632B77C120828476AC2EB8BD5DFF930" type="volume">
<mods:number id="34B26C58235C6DDF8E8C962C82B863B2">2051</mods:number>
</mods:detail>
<mods:extent id="A145537E3DC102B46E4740AE4FCC8D7D" unit="page">
<mods:start id="EFD3743D17723C5732C97E90051BB88C">1</mods:start>
<mods:end id="2719C7A1210E6FF6864CBF4A4E2A2BAB">25</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification id="84F1A80AC17C0B1A8B81CB02F34A3787">journal article</mods:classification>
<mods:identifier id="20D1F04CD9A38E8EAAB13EF41FC3E5BF" type="DOI">10.5281/zenodo.186587</mods:identifier>
<mods:identifier id="0BE98668367CE95140EE25F409117D2D" type="GBIF-Dataset">f979f9ce-284c-46d4-a6b0-25dfc8f77007</mods:identifier>
<mods:identifier id="7195B0462D22BDD4F4994F6FF473968E" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="26063B1799F26FE8D1D615C0297DD544" type="Zenodo-Dep">186587</mods:identifier>
</mods:mods>
<treatment id="033A9525464BFFF077DBA4AFBBB9A6A5" ID-DOI="http://doi.org/10.5281/zenodo.6217272" ID-GBIF-Taxon="119612532" ID-Zenodo-Dep="6217272" LSID="urn:lsid:plazi:treatment:033A9525464BFFF077DBA4AFBBB9A6A5" httpUri="http://treatment.plazi.org/id/033A9525464BFFF077DBA4AFBBB9A6A5" lastPageId="10" lastPageNumber="11" pageId="7" pageNumber="8">
<subSubSection id="C38977B8464BFFFD77DBA4AFB92EA722" pageId="7" pageNumber="8" type="nomenclature">
<paragraph id="8B2C2433464BFFFD77DBA4AFB935A49F" blockId="7.[151,607,1454,1480]" box="[151,607,1454,1480]" pageId="7" pageNumber="8">
<heading id="D064935F464BFFFD77DBA4AFB935A49F" bold="true" box="[151,607,1454,1480]" fontSize="11" level="1" pageId="7" pageNumber="8" reason="1">
<taxonomicName id="4C935FB0464BFFFD77DBA4AFB935A49F" ID-CoL="6MMSB" authority="Bourret, 1937" authorityName="Bourret" authorityYear="1937" box="[151,607,1454,1480]" class="Amphibia" family="Ranidae" genus="Hylarana" kingdom="Animalia" order="Anura" pageId="7" pageNumber="8" phylum="Chordata" rank="species" species="maosonensis">
<emphasis id="B9E7F821464BFFFD77DBA4AFB935A49F" bold="true" box="[151,607,1454,1480]" pageId="7" pageNumber="8">
<emphasis id="B9E7F821464BFFFD77DBA4AFBAC7A49F" bold="true" box="[151,429,1454,1480]" italics="true" pageId="7" pageNumber="8">Hylarana maosonensis</emphasis>
Bourret, 1937
</emphasis>
</taxonomicName>
</heading>
</paragraph>
<paragraph id="8B2C2433464BFFFD77DBA4F2B92EA722" blockId="7.[151,1437,1523,2035]" pageId="7" pageNumber="8">
<emphasis id="B9E7F821464BFFFD77DBA4F2BA02A75A" bold="true" box="[151,360,1523,1549]" pageId="7" pageNumber="8">Series examined.</emphasis>
Tadpoles of
<taxonomicName id="4C935FB0464BFFFD76B2A4F5B861A75A" box="[510,779,1524,1549]" class="Amphibia" family="Ranidae" genus="Hylarana" kingdom="Animalia" order="Anura" pageId="7" pageNumber="8" phylum="Chordata" rank="species" species="maosonensis">
<emphasis id="B9E7F821464BFFFD76B2A4F5B861A75A" box="[510,779,1524,1549]" italics="true" pageId="7" pageNumber="8">Hylarana maosonensis</emphasis>
</taxonomicName>
(ZFMK 87287-87305: n = 19) were collected in PNKB, U Bo region,
<collectingCountry id="F38464A3464BFFFD7678A717BAFDA767" box="[308,407,1558,1584]" name="Vietnam" pageId="7" pageNumber="8">Vietnam</collectingCountry>
by R. Hendrix from
<date id="FF2D02F3464BFFFD75C6A717B817A767" box="[650,893,1558,1584]" pageId="7" pageNumber="8" value="2006-06" valueMax="2006-08-31" valueMin="2006-06-00">June to August 2006</date>
. The larvae were collected throughout day and night time in primary forest streams, in ponds and in a catchment basin next to a forest road at altitudes between
<quantity id="4C6B89D6464BFFFD764CA75ABAE8A722" box="[256,386,1627,1653]" metricMagnitude="2" metricUnit="m" metricValue="6.15" metricValueMax="7.0" metricValueMin="5.3" pageId="7" pageNumber="8" unit="m" value="615.0" valueMax="700.0" valueMin="530.0">530700 m</quantity>
above sea level.
</paragraph>
</subSubSection>
<subSubSection id="C38977B8464BFFF07789A77FBBB9A6A5" lastPageId="10" lastPageNumber="11" pageId="7" pageNumber="8" type="description">
<paragraph id="8B2C2433464BFFFD7789A77FBF9BA7EC" blockId="7.[151,1437,1523,2035]" pageId="7" pageNumber="8">
Adult specimens (ZFMK 86374, 86399-86400) were collected at night near primary forest streams and ponds within U Bo region of PNKB,
<collectingCountry id="F38464A3464BFFFD7507A7A0B9DBA7EC" box="[587,689,1697,1723]" name="Vietnam" pageId="7" pageNumber="8">Vietnam</collectingCountry>
, at altitudes between
<quantity id="4C6B89D6464BFFFD74E0A7A0BF45A7EC" box="[940,1071,1697,1723]" metricMagnitude="2" metricUnit="m" metricValue="6.15" metricValueMax="7.0" metricValueMin="5.3" pageId="7" pageNumber="8" unit="m" value="615.0" valueMax="700.0" valueMin="530.0">530700 m</quantity>
above sea level.
</paragraph>
<paragraph id="8B2C2433464BFFF27789A7C2B8DEA04D" blockId="7.[151,1437,1523,2035]" lastBlockId="8.[151,1437,152,282]" lastPageId="8" lastPageNumber="9" pageId="7" pageNumber="8">
<emphasis id="B9E7F821464BFFFD7789A7C2BAEFA78A" bold="true" box="[197,389,1731,1757]" pageId="7" pageNumber="8">Identification.</emphasis>
Adult specimens (
<figureCitation id="13A838B6464BFFFD753AA7C2B9BFA78A" box="[630,725,1731,1757]" captionStart="FIGURE 1 a d" captionStartId="4.[151,255,1913,1937]" captionTargetBox="[153,1430,359,1912]" captionTargetId="figure@4.[151,1436,348,1923]" captionTargetPageId="4" captionText="FIGURE 1 a d adult Hylarana from Phong Nha Ke Bang National Park, Quang Binh Province, Vietnam in life; a) H. attigua (ZFMK 86404, field number RH 06074), b) H. maosonensis (ZFMK 86374), c) H. guentheri (ZFMK 86372, field number RH 06007), d) H. nigrovittata. Photos: Ralf Hendrix. e) adult H. nigrovittata in life from near Chiang Mai, northern Thailand. Photo: Thomas Ziegler" httpUri="https://zenodo.org/record/186588/files/figure.png" pageId="7" pageNumber="8">Fig. 1b</figureCitation>
; for measurements see
<tableCitation id="C6111188464BFFFD734FA7C2BF09A78A" box="[1027,1123,1731,1757]" captionStart="TABLE 4" captionStartId="8.[151,232,328,349]" captionText="TABLE 4. Measurements (min. - max., mean and standard deviation, in mm) of the adult Hylarana maosonensis series from Phong Nha Ke Bang National Park, Quang Binh Province, Vietnam (ZFMK 86374, 86399 - 86400); for abbreviations see material and methods." pageId="7" pageNumber="8">Table 4</tableCitation>
) largely agreed with the descriptions by
<bibRefCitation id="EF0259C2464BFFFD7612A7E7B976A657" author="Bourret" box="[350,540,1766,1792]" pageId="7" pageNumber="24" refString="Bourret, R. (1942) Les batraciens de l'Indochine. Institut Oceanographique de l'Indochine, Hanoi, 547 pp." type="book" year="1942">Bourret (1942)</bibRefCitation>
,
<bibRefCitation id="EF0259C2464BFFFD7566A7E7B87EA657" box="[554,788,1766,1792]" pageId="7" pageNumber="25" refString="Inger, R. F., Orlov, N. L. &amp; Darevsky, I. (1999) Frogs of Vietnam: A report on new collections. Fieldiana Zoology, N. S., 92, 1 - 46." type="journal article">
Inger
<emphasis id="B9E7F821464BFFFD7539A7E6B9DFA657" box="[629,693,1767,1792]" italics="true" pageId="7" pageNumber="8">et al.</emphasis>
(1999)
</bibRefCitation>
,
<bibRefCitation id="EF0259C2464BFFFD746EA7E7B8B2A657" author="Ziegler" box="[802,984,1766,1792]" pageId="7" pageNumber="25" refString="Ziegler, T. (2002) Die Amphibien und Reptilien eines Tieflandfeuchtwald-Schutzgebietes in Vietnam. Natur &amp; Tier Verlag, Munster, 342 pp." type="book" year="2002">Ziegler (2002)</bibRefCitation>
and
<bibRefCitation id="EF0259C2464BFFFD735AA7E7BE73A657" box="[1046,1305,1766,1792]" pageId="7" pageNumber="25" refString="Ziegler, T., Ohler, A., Vu, N. T., Le, K. Q., Nguyen, X. T., Dinh, H. T. &amp; Bui, N. T. (2006) Review of the amphibian and reptile diversity of Phong Nha - Ke Bang National Park and adjacent areas, central Truong Son, Vietnam. In: Vences, M., Kohler, J., Ziegler, T. &amp; Bohme, W. (Eds.), Herpetologia Bonnensis II. Proceedings of the 13 th Ordinary General Meeting of the Societas Europaea Herpetologica, Bonn, pp. 247 - 262." type="proceedings paper">
Ziegler
<emphasis id="B9E7F821464BFFFD7335A7E6BFD3A657" box="[1145,1209,1767,1792]" italics="true" pageId="7" pageNumber="8">et al.</emphasis>
(2006)
</bibRefCitation>
. Sequence congruence between adults (ZFMK 86373, 86399-400) and tadpoles (ZFMK 87293-295, 87300, 87305;
<figureCitation id="13A838B6464BFFFD7221A608BBDCA612" captionStart="FIGURE 2 a d" captionStartId="5.[151,255,1718,1742]" captionTargetBox="[153,1430,186,1694]" captionTargetId="figure@5.[151,1436,183,1758]" captionTargetPageId="5" captionText="FIGURE 2 a d larvae of Hylarana from Phong Nha Ke Bang National Park, Quang Binh Province, Vietnam in life; a) H. attigua (ZFMK 87286, field number RH 06230), b) H. maosonensis (ZFMK 87300), c) H. guentheri (ZFMK 86988), d) H. nigrovittata. Photos: Ralf Hendrix. e) larva of H. nigrovittata from near Chiang Mai, northern Thailand in life. Photo: Thomas Ziegler" httpUri="https://zenodo.org/record/186589/files/figure.png" pageId="7" pageNumber="8">Fig. 2b</figureCitation>
) was 100 %. The following larval description is based on a single tadpole in Gosner stage 36 (
<figureCitation id="13A838B6464BFFFD726EA62ABE06A612" box="[1314,1388,1835,1861]" captionStart="FIGURE 4" captionStartId="9.[151,259,1534,1558]" captionTargetBox="[157,1426,1119,1496]" captionTargetId="figure@9.[151,1436,1094,1511]" captionTargetPageId="9" captionText="FIGURE 4. Drawings of the preserved tadpole of Hylarana maosonensis (stage 36) from Phong Nha Ke Bang National Park, Quang Binh Province, Vietnam; (A) dorsal view, (B) lateral view, (C) oral disc. Drawings by Anna Gawor" httpUri="https://zenodo.org/record/186591/files/figure.png" pageId="7" pageNumber="8">Fig. 4</figureCitation>
; for measurements see
<tableCitation id="C6111188464BFFFD763FA64FBAA7A63F" box="[371,461,1870,1896]" captionStart="TABLE 5" captionStartId="8.[151,239,1057,1081]" captionText="TABLE 5. Measurements (min. max., mean and standard deviation, in mm) of the Hylarana maosonensis larvae series from Phong Nha Ke Bang National Park, Quang Binh Province, Vietnam (ZFMK 87287 - 87305); for abbreviations see material and methods." pageId="7" pageNumber="8">Table 5</tableCitation>
). The tadpole specimen used for description and drawing originates from the same water body as the DNA voucher specimens (characters of the five DNA voucher specimens of the series ZFMK 87300, 87305, and 87293-87295: stages 28-40, BL
<quantity id="4C6B89D6464BFFFD7415A692B89FA6FA" box="[857,1013,1939,1965]" metricMagnitude="-2" metricUnit="m" metricValue="1.1099999999999999" metricValueMax="1.51" metricValueMin="0.71" pageId="7" pageNumber="8" unit="mm" value="11.1" valueMax="15.1" valueMin="7.1">7.1-15.1 mm</quantity>
, NP
<quantity id="4C6B89D6464BFFFD7363A692BF06A6FA" box="[1071,1132,1939,1965]" metricMagnitude="0" metricUnit="m" metricValue="1.3969999999999998" pageId="7" pageNumber="8" unit="in" value="55.0">55 in</quantity>
stage 28, 78-
<quantity id="4C6B89D6464BFFFD7242A692BE21A6FA" box="[1294,1355,1939,1965]" metricMagnitude="0" metricUnit="m" metricValue="2.3622" pageId="7" pageNumber="8" unit="in" value="93.0">93 in</quantity>
stages 35-40, NK 26-29 per
<quantity id="4C6B89D6464BFFFD76DFA6B7BA98A687" box="[403,498,1974,2000]" metricMagnitude="-4" metricUnit="m" metricValue="5.0" pageId="7" pageNumber="8" unit="mm" value="0.5">0.5 mm</quantity>
). In this water body, also
<taxonomicName id="4C935FB0464BFFFD7457A6B6B8C5A687" box="[795,943,1975,2000]" class="Amphibia" family="Ranidae" genus="Hylarana" kingdom="Animalia" order="Anura" pageId="7" pageNumber="8" phylum="Chordata" rank="species" species="guentheri">
<emphasis id="B9E7F821464BFFFD7457A6B6B8C5A687" box="[795,943,1975,2000]" italics="true" pageId="7" pageNumber="8">H. guentheri</emphasis>
</taxonomicName>
(adults only) and
<taxonomicName id="4C935FB0464BFFFD73CFA6B6BF90A687" box="[1155,1274,1975,2000]" class="Amphibia" family="Ranidae" genus="Hylarana" kingdom="Animalia" order="Anura" pageId="7" pageNumber="8" phylum="Chordata" rank="species" species="attigua">
<emphasis id="B9E7F821464BFFFD73CFA6B6BF90A687" box="[1155,1274,1975,2000]" italics="true" pageId="7" pageNumber="8">H. attigua</emphasis>
</taxonomicName>
(larvae) were found. The DNA voucher specimens fully agree morphologically with the described and drawn specimen, including the characters that diagnose
<taxonomicName id="4C935FB04644FFF27526A199B840A1E6" box="[618,810,152,177]" class="Amphibia" family="Ranidae" genus="Hylarana" kingdom="Animalia" order="Anura" pageId="8" pageNumber="9" phylum="Chordata" rank="species" species="maosonensis">
<emphasis id="B9E7F8214644FFF27526A199B840A1E6" box="[618,810,152,177]" italics="true" pageId="8" pageNumber="9">H. maosonensis</emphasis>
</taxonomicName>
from
<taxonomicName id="4C935FB04644FFF27438A199B885A1E6" box="[884,1007,152,177]" class="Amphibia" family="Ranidae" genus="Hylarana" kingdom="Animalia" order="Anura" pageId="8" pageNumber="9" phylum="Chordata" rank="species" species="attigua">
<emphasis id="B9E7F8214644FFF27438A199B885A1E6" box="[884,1007,152,177]" italics="true" pageId="8" pageNumber="9">H. attigua</emphasis>
</taxonomicName>
and
<taxonomicName id="4C935FB04644FFF27360A199BFAEA1E6" box="[1068,1220,152,177]" class="Amphibia" family="Ranidae" genus="Hylarana" kingdom="Animalia" order="Anura" pageId="8" pageNumber="9" phylum="Chordata" rank="species" species="guentheri">
<emphasis id="B9E7F8214644FFF27360A199BFAEA1E6" box="[1068,1220,152,177]" italics="true" pageId="8" pageNumber="9">H. guentheri</emphasis>
</taxonomicName>
(broad upper jaw sheath, finger-shaped marginal papillae, two submarginal rows of papillae, papillae strongly pigmented in their distal halfs, dark olive brown colour with broad dark spots and bright blotch on throat). The identification of the described specimen is therefore well supported.
</paragraph>
<caption id="DFEC74BB4644FFF277DBA049BB85A0CF" pageId="8" pageNumber="9">
<paragraph id="8B2C24334644FFF277DBA049BB85A0CF" blockId="8.[151,1437,328,995]" pageId="8" pageNumber="9">
<emphasis id="B9E7F8214644FFF277DBA049BA6BA00A" bold="true" box="[151,257,328,349]" pageId="8" pageNumber="9">TABLE 4.</emphasis>
Measurements (min.-max., mean and standard deviation, in mm) of the adult
<taxonomicName id="4C935FB04644FFF2734EA048BF88A009" box="[1026,1250,329,350]" class="Amphibia" family="Ranidae" genus="Hylarana" kingdom="Animalia" order="Anura" pageId="8" pageNumber="9" phylum="Chordata" rank="species" species="maosonensis">
<emphasis id="B9E7F8214644FFF2734EA048BF88A009" box="[1026,1250,329,350]" italics="true" pageId="8" pageNumber="9">Hylarana maosonensis</emphasis>
</taxonomicName>
series from Phong Nha Ke Bang National Park, Quang Binh Province, Vietnam (ZFMK 86374, 86399-86400); for abbreviations see material and methods.
</paragraph>
</caption>
<paragraph id="8B2C24334644FFF277EDA0B3B99FA09E" blockId="8.[151,1437,328,995]" box="[161,757,434,457]" pageId="8" pageNumber="9">Measurements Males (n = 3)</paragraph>
<paragraph id="8B2C24334644FFF277EDA0DDB839A34B" blockId="8.[151,1437,328,995]" pageId="8" pageNumber="9">SVL 35.5 37.5 (36.50 ± 1.00) SL 6.0 7.0 (6.17 ± 0.76)</paragraph>
<paragraph id="8B2C24334644FFF277EDA32FB839A312" blockId="8.[151,1437,328,995]" box="[161,851,558,581]" pageId="8" pageNumber="9">NL 2.5 3.0 (2.77 ± 0.25)</paragraph>
<paragraph id="8B2C24334644FFF277EDA359B839A338" blockId="8.[151,1437,328,995]" box="[161,851,600,623]" pageId="8" pageNumber="9">EN 3.0 3.2 (3.07 ± 0.12)</paragraph>
<paragraph id="8B2C24334644FFF277EDA380B839A3BC" blockId="8.[151,1437,328,995]" pageId="8" pageNumber="9">HW 12.5 13.8 (13.27 ± 0.68) HL 13.0 14.5 (13.57 ± 0.81) EE 7.0 8.5 (7.83 ± 0.76)</paragraph>
<paragraph id="8B2C24334644FFF277EDA3FCB839A243" blockId="8.[151,1437,328,995]" box="[161,851,765,788]" pageId="8" pageNumber="9">ED 5.5 6.0 (5.83 ± 0.29)</paragraph>
<paragraph id="8B2C24334644FFF277EDA227B839A26A" blockId="8.[151,1437,328,995]" box="[161,851,806,829]" pageId="8" pageNumber="9">TD 3.5 4.0 (3.67 ± 0.29)</paragraph>
<paragraph id="8B2C24334644FFF277EDA251B839A2B4" blockId="8.[151,1437,328,995]" pageId="8" pageNumber="9">HaL 10.0 13.0 (11.17 ± 1.61) LL 70.0 75.5 (72.33 ± 2.84) FL 20.0 23.0 (21.67 ± 1.53) IMT 1.5 2.2 (1.90 ± 0.36)</paragraph>
<caption id="DFEC74BB4644FFF277DBA520BAEEA52A" pageId="8" pageNumber="9">
<paragraph id="8B2C24334644FFF277DBA520BAEEA52A" blockId="8.[151,1436,1057,1937]" pageId="8" pageNumber="9">
<emphasis id="B9E7F8214644FFF277DBA520BA63A56E" bold="true" box="[151,265,1057,1081]" pageId="8" pageNumber="9">TABLE 5.</emphasis>
Measurements (min.max., mean and standard deviation, in mm) of the
<taxonomicName id="4C935FB04644FFF27355A520BE65A56F" box="[1049,1295,1057,1080]" class="Amphibia" family="Ranidae" genus="Hylarana" kingdom="Animalia" order="Anura" pageId="8" pageNumber="9" phylum="Chordata" rank="species" species="maosonensis">
<emphasis id="B9E7F8214644FFF27355A520BE65A56F" box="[1049,1295,1057,1080]" italics="true" pageId="8" pageNumber="9">Hylarana maosonensis</emphasis>
</taxonomicName>
larvae series from Phong Nha Ke Bang National Park, Quang Binh Province, Vietnam (ZFMK 87287-87305); for abbreviations see material and methods.
</paragraph>
</caption>
<paragraph id="8B2C24334644FFF277D3A595BE20A59C" blockId="8.[151,1436,1057,1937]" pageId="8" pageNumber="9">
Stage BH BL BW ED
<collectingCountry id="F38464A34644FFF2759BA595B868A5FE" box="[727,770,1172,1193]" name="India" pageId="8" pageNumber="9">IND</collectingCountry>
IP LF MTH NK NP 28 2.8 7.1 4.5 0.8 1.9 2.7 - - 29 55
</paragraph>
<paragraph id="8B2C24334644FFF277A7A5D6BE12A459" blockId="8.[151,1436,1057,1937]" pageId="8" pageNumber="9">3.74.3 8.89.2 3.05.7 1.01.1 1.01.1 3.13.3 0.91.0 4.34.5 2628 4860 30 (4. 0 0 (9. 0 0 (4. 7 3 (1. 0 3 (1. 0 3 (3. 1 7 (0. 9 5 (4. 40 (27 ±1) (54 ±8)</paragraph>
<paragraph id="8B2C24334644FFF277A7A416BFE9A47B" blockId="8.[151,1436,1057,1937]" box="[235,1155,1303,1324]" pageId="8" pageNumber="9">±0.30) ±0.20) ±1.50) ±0.06) ±0.06) ±0.08) ±0.07) ±0.14)</paragraph>
<paragraph id="8B2C24334644FFF277A7A435BE20A43C" blockId="8.[151,1436,1057,1937]" pageId="8" pageNumber="9">(n=3) (n=3) (n=3) (n=3) (n=3) (n=3) (n=2) (n=2) (n=3) (n=2) 32 4.7 10.0 6.3 1.8 2.1 3.7 1.3 5.7 32 87</paragraph>
<paragraph id="8B2C24334644FFF277A7A479BE13A4F8" blockId="8.[151,1436,1057,1937]" pageId="8" pageNumber="9">4.95.1 11.112.6 6.87.0 1.51.6 2.22.6 3.84.3 2930 7780 35 (5. 0 0 (11. 8 5 (6. 9 0 (1. 5 5 (2. 4 0 (4. 0 5 1.3 5.8 (30 ±1) (79 ±2)</paragraph>
<paragraph id="8B2C24334644FFF277A7A4B6B8FAA49B" blockId="8.[151,1436,1057,1937]" box="[235,912,1463,1484]" pageId="8" pageNumber="9">±0.14) ±1.06) ±0.14) ±0.07) ±0.28) ±0.35)</paragraph>
<paragraph id="8B2C24334644FFF277A7A4D4BE20A707" blockId="8.[151,1436,1057,1937]" pageId="8" pageNumber="9">(n=2) (n=2) (n=2) (n=2) (n=2) (n=2) (n=2) (n=2) 36 4.7 12.7 7.2 1.6 2.7 4.4 1.2 6.1 34 85 37 6.6 12.8 7.8 1.8 2.8 4.5 1.9 7.2 29 - 38 5.9 12.1 7.6 1.6 2.6 4.6 1.6 6.7 27 90</paragraph>
<paragraph id="8B2C24334644FFF277A7A75CBE12A7C3" blockId="8.[151,1436,1057,1937]" pageId="8" pageNumber="9">6.07.5 13.715.1 8.410.8 1.82.1 2.83.1 5.36.2 1.02.2 7.28.0 2730 7898 40 (6. 8 2 (1 4. 5 2 (9. 5 8 (1. 9 0 (2. 9 6 (5. 6 2 (1. 6 4 (7. 58 (28 ±1) (89 ±9)</paragraph>
<paragraph id="8B2C24334644FFF277A7A79DBFEEA7E6" blockId="8.[151,1436,1057,1937]" box="[235,1156,1692,1713]" pageId="8" pageNumber="9">±0.63) ±0.54) ±0.90) ±0.14) ±0.11) ±0.36) ±0.46) ±0.30)</paragraph>
<paragraph id="8B2C24334644FFF277A7A7B8BE50A644" blockId="8.[151,1436,1057,1937]" pageId="8" pageNumber="9">(n=5) (n=5) (n=5) (n=5) (n=5) (n=5) (n=5) (n=5) (n=5) (n=5) 41 6.5 12.3 8.1 1.9 2.3 5.0 0.8 6.0 29 - 42 6.1 12.3 8.2 2.1 2.3 5.0 1.4 5.2 28 -</paragraph>
<paragraph id="8B2C24334644FFF277A7A61EBEEEA601" blockId="8.[151,1436,1057,1937]" pageId="8" pageNumber="9">5.15.2 11.111.5 6.40 1.92.0 1.31.4 3.73.8 0.50.7 4.14.2 - 4562 43 (5. 1 5 (11. 3 0 (6. 3 8 (1. 9 3 (1. 3 5 (3. 7 5 (0. 6 0 (4. 13 - (54 ±12)</paragraph>
<paragraph id="8B2C24334644FFF277A7A65EBFE9A623" blockId="8.[151,1436,1057,1937]" box="[235,1155,1887,1908]" pageId="8" pageNumber="9">±0.07) ±0.28) ±0.04) ±0.11) ±0.07) ±0.07) ±0.14) ±0.04)</paragraph>
<paragraph id="8B2C24334644FFF377A7A67DBE02A38B" blockId="8.[151,1436,1057,1937]" lastBlockId="9.[159,1431,711,802]" lastPageId="9" lastPageNumber="10" pageId="8" pageNumber="9">(n=2) (n=2) (n=2) (n=2) (n=2) (n=2) (n=2) (n=2) - (n=2) 2.63.2 3.64.1 1.01.5 9.410.4 33.035.0 47.749.8 3.84.2 3.84.1 2.63.0</paragraph>
<paragraph id="8B2C24334645FFF377DBA199BEFDA000" pageId="9" pageNumber="10">
<table id="F993D6934645000577DBA199BEFDA023" box="[151,1431,152,372]" gridcols="9" gridrows="5" pageId="9" pageNumber="10">
<tr id="35A326714645000577DBA199BEFDA1F8" box="[151,1431,152,175]" gridrow="0" pageId="9" pageNumber="10" rowspan-1="1" rowspan-2="1" rowspan-3="1" rowspan-4="1" rowspan-5="1" rowspan-6="1" rowspan-7="1" rowspan-8="1">
<th id="76724F0D4645000577DBA199BA31A1F8" box="[151,347,152,175]" gridcol="0" gridrow="0" pageId="9" pageNumber="10">continued.</th>
</tr>
<tr id="35A326714645000577DBA1CBBEFDA188" box="[151,1431,202,223]" gridrow="1" pageId="9" pageNumber="10">
<th id="76724F0D4645000577DBA1CBBA31A188" box="[151,347,202,223]" gridcol="0" gridrow="1" pageId="9" pageNumber="10">Stage NPD</th>
<td id="76724F0D464500057627A1CBBA88A188" box="[363,482,202,223]" gridcol="1" gridrow="1" pageId="9" pageNumber="10">ODW</td>
<td id="76724F0D4645000576BEA1CBB903A188" box="[498,617,202,223]" gridcol="2" gridrow="1" pageId="9" pageNumber="10">RND</td>
<td id="76724F0D464500057535A1CBB99AA188" box="[633,752,202,223]" gridcol="3" gridrow="1" pageId="9" pageNumber="10">SS</td>
<td id="76724F0D46450005744CA1CBB80AA188" box="[768,864,202,223]" gridcol="4" gridrow="1" pageId="9" pageNumber="10">TAL</td>
<td id="76724F0D4645000574C5A1CBB883A188" box="[905,1001,202,223]" gridcol="5" gridrow="1" pageId="9" pageNumber="10">TL</td>
<td id="76724F0D46450005735EA1CBBFE3A188" box="[1042,1161,202,223]" gridcol="6" gridrow="1" pageId="9" pageNumber="10">TMH</td>
<td id="76724F0D4645000573D5A1CBBE7AA188" box="[1177,1296,202,223]" gridcol="7" gridrow="1" pageId="9" pageNumber="10">TMW</td>
<td id="76724F0D46450005726CA1CBBEFDA188" box="[1312,1431,202,223]" gridcol="8" gridrow="1" pageId="9" pageNumber="10">UF</td>
</tr>
<tr id="35A326714645000577DBA1F3BEFDA050" box="[151,1431,242,263]" gridrow="2" pageId="9" pageNumber="10">
<th id="76724F0D4645000577DBA1F3BA31A050" box="[151,347,242,263]" gridcol="0" gridrow="2" pageId="9" pageNumber="10">28 1.3</th>
<td id="76724F0D464500057627A1F3BA88A050" box="[363,482,242,263]" gridcol="1" gridrow="2" pageId="9" pageNumber="10">2.0</td>
<td id="76724F0D4645000576BEA1F3B903A050" box="[498,617,242,263]" gridcol="2" gridrow="2" pageId="9" pageNumber="10">0.8</td>
<td id="76724F0D464500057535A1F3B99AA050" box="[633,752,242,263]" gridcol="3" gridrow="2" pageId="9" pageNumber="10">5.0</td>
<td id="76724F0D46450005744CA1F3B80AA050" box="[768,864,242,263]" gridcol="4" gridrow="2" pageId="9" pageNumber="10">-</td>
<td id="76724F0D4645000574C5A1F3B883A050" box="[905,1001,242,263]" gridcol="5" gridrow="2" pageId="9" pageNumber="10">-</td>
<td id="76724F0D46450005735EA1F3BFE3A050" box="[1042,1161,242,263]" gridcol="6" gridrow="2" pageId="9" pageNumber="10">1.9</td>
<td id="76724F0D4645000573D5A1F3BE7AA050" box="[1177,1296,242,263]" gridcol="7" gridrow="2" pageId="9" pageNumber="10">1.8</td>
<td id="76724F0D46450005726CA1F3BEFDA050" box="[1312,1431,242,263]" gridcol="8" gridrow="2" pageId="9" pageNumber="10">-</td>
</tr>
<tr id="35A326714645000577DBA01BBEFDA078" box="[151,1431,282,303]" gridrow="3" pageId="9" pageNumber="10">
<th id="76724F0D4645000577DBA01BBA31A078" box="[151,347,282,303]" gridcol="0" gridrow="3" pageId="9" pageNumber="10">1.41.6</th>
<td id="76724F0D464500057627A01BBA88A078" box="[363,482,282,303]" gridcol="1" gridrow="3" pageId="9" pageNumber="10">2.02.2</td>
<td id="76724F0D4645000576BEA01BB903A078" box="[498,617,282,303]" gridcol="2" gridrow="3" pageId="9" pageNumber="10">0.71.0</td>
<td id="76724F0D464500057535A01BB99AA078" box="[633,752,282,303]" gridcol="3" gridrow="3" pageId="9" pageNumber="10">5.26.0</td>
<td id="76724F0D46450005744CA01BB80AA078" box="[768,864,282,303]" gridcol="4" gridrow="3" pageId="9" pageNumber="10">13.414.6</td>
<td id="76724F0D4645000574C5A01BB883A078" box="[905,1001,282,303]" gridcol="5" gridrow="3" pageId="9" pageNumber="10">22.223.6</td>
<td id="76724F0D46450005735EA01BBFE3A078" box="[1042,1161,282,303]" gridcol="6" gridrow="3" pageId="9" pageNumber="10">2.32.5</td>
<td id="76724F0D4645000573D5A01BBE7AA078" box="[1177,1296,282,303]" gridcol="7" gridrow="3" pageId="9" pageNumber="10">2.12.2</td>
<td id="76724F0D46450005726CA01BBEFDA078" box="[1312,1431,282,303]" gridcol="8" gridrow="3" pageId="9" pageNumber="10">1.51.7</td>
</tr>
<tr id="35A326714645000577DBA043BEFDA023" box="[151,1431,322,372]" gridrow="4" pageId="9" pageNumber="10">
<th id="76724F0D4645000577DBA043BA31A023" box="[151,347,322,372]" gridcol="0" gridrow="4" pageId="9" pageNumber="10">30 (1.47 ±0.12)</th>
<td id="76724F0D464500057627A043BA88A023" box="[363,482,322,372]" gridcol="1" gridrow="4" pageId="9" pageNumber="10">(2.13 ±0.12)</td>
<td id="76724F0D4645000576BEA043B903A023" box="[498,617,322,372]" gridcol="2" gridrow="4" pageId="9" pageNumber="10">(0.87 ±0.15)</td>
<td id="76724F0D464500057535A043B99AA023" box="[633,752,322,372]" gridcol="3" gridrow="4" pageId="9" pageNumber="10">(5.67 ±0.42)</td>
<td id="76724F0D46450005744CA043B80AA023" box="[768,864,322,372]" gridcol="4" gridrow="4" pageId="9" pageNumber="10">(14.00 ±0.85)</td>
<td id="76724F0D4645000574C5A043B883A023" box="[905,1001,322,372]" gridcol="5" gridrow="4" pageId="9" pageNumber="10">(22.90 ±0.99)</td>
<td id="76724F0D46450005735EA043BFE3A023" box="[1042,1161,322,372]" gridcol="6" gridrow="4" pageId="9" pageNumber="10">(2.40 ±0.10)</td>
<td id="76724F0D4645000573D5A043BE7AA023" box="[1177,1296,322,372]" gridcol="7" gridrow="4" pageId="9" pageNumber="10">(2.13 ±0.06)</td>
<td id="76724F0D46450005726CA043BEFDA023" box="[1312,1431,322,372]" gridcol="8" gridrow="4" pageId="9" pageNumber="10">(1.60 ±0.14)</td>
</tr>
</table>
</paragraph>
<paragraph id="8B2C24334645FFF377A8A086BF8BA0BB" pageId="9" pageNumber="10">
<table id="F993D6934645000577DBA086BEFDA0BB" box="[151,1431,391,492]" gridcols="9" gridrows="3" pageId="9" pageNumber="10">
<tr id="35A326714645000577DBA086BEFDA0CB" box="[151,1431,391,412]" gridrow="0" pageId="9" pageNumber="10">
<th id="76724F0D4645000577DBA086BA46A0CB" box="[151,300,391,412]" gridcol="0" gridrow="0" pageId="9" pageNumber="10">(n=3)</th>
<th id="76724F0D464500057627A086BAD9A0CB" box="[363,435,391,412]" gridcol="1" gridrow="0" pageId="9" pageNumber="10">(n=3)</th>
<th id="76724F0D4645000576BEA086B94DA0CB" box="[498,551,391,412]" gridcol="2" gridrow="0" pageId="9" pageNumber="10">(n=3)</th>
<th id="76724F0D464500057535A086B9ABA0CB" box="[633,705,391,412]" gridcol="3" gridrow="0" pageId="9" pageNumber="10">(n=3)</th>
<th id="76724F0D46450005744CA086B85FA0CB" box="[768,821,391,412]" gridcol="4" gridrow="0" pageId="9" pageNumber="10">(n=2)</th>
<th id="76724F0D4645000574C5A086B8D4A0CB" box="[905,958,391,412]" gridcol="5" gridrow="0" pageId="9" pageNumber="10">(n=2)</th>
<th id="76724F0D46450005735EA086BF2DA0CB" box="[1042,1095,391,412]" gridcol="6" gridrow="0" pageId="9" pageNumber="10">(n=3)</th>
<th id="76724F0D4645000573D5A086BF8BA0CB" box="[1177,1249,391,412]" gridcol="7" gridrow="0" pageId="9" pageNumber="10">(n=3)</th>
<th id="76724F0D46450005726CA086BEFDA0CB" box="[1312,1431,391,412]" gridcol="8" gridrow="0" pageId="9" pageNumber="10">(n=2)</th>
</tr>
<tr id="35A326714645000577DBA0AEBEFDA093" box="[151,1431,431,452]" gridrow="1" pageId="9" pageNumber="10">
<th id="76724F0D4645000577DBA0AEBA46A093" box="[151,300,431,452]" gridcol="0" gridrow="1" pageId="9" pageNumber="10">32 1.8</th>
<td id="76724F0D464500057627A0AEBAD9A093" box="[363,435,431,452]" gridcol="1" gridrow="1" pageId="9" pageNumber="10">2.5</td>
<td id="76724F0D4645000576BEA0AEB94DA093" box="[498,551,431,452]" gridcol="2" gridrow="1" pageId="9" pageNumber="10">0.9</td>
<td id="76724F0D464500057535A0AEB9ABA093" box="[633,705,431,452]" gridcol="3" gridrow="1" pageId="9" pageNumber="10">7.2</td>
<td id="76724F0D46450005744CA0AEB85FA093" box="[768,821,431,452]" gridcol="4" gridrow="1" pageId="9" pageNumber="10">-</td>
<td id="76724F0D4645000574C5A0AEB8D4A093" box="[905,958,431,452]" gridcol="5" gridrow="1" pageId="9" pageNumber="10">-</td>
<td id="76724F0D46450005735EA0AEBF2DA093" box="[1042,1095,431,452]" gridcol="6" gridrow="1" pageId="9" pageNumber="10">3.0</td>
<td id="76724F0D4645000573D5A0AEBF8BA093" box="[1177,1249,431,452]" gridcol="7" gridrow="1" pageId="9" pageNumber="10">2.1</td>
<td id="76724F0D46450005726CA0AEBEFDA093" box="[1312,1431,431,452]" gridcol="8" gridrow="1" pageId="9" pageNumber="10">2.1</td>
</tr>
<tr id="35A326714645000577DBA0D6BEFDA0BB" box="[151,1431,471,492]" gridrow="2" pageId="9" pageNumber="10" rowspan-2="1" rowspan-4="1" rowspan-5="1" rowspan-6="1" rowspan-8="1">
<th id="76724F0D4645000577DBA0D6BA46A0BB" box="[151,300,471,492]" gridcol="0" gridrow="2" pageId="9" pageNumber="10">2.02.3</th>
<td id="76724F0D464500057627A0D6BAD9A0BB" box="[363,435,471,492]" gridcol="1" gridrow="2" pageId="9" pageNumber="10">2.83.3</td>
<td id="76724F0D464500057535A0D6B9ABA0BB" box="[633,705,471,492]" gridcol="3" gridrow="2" pageId="9" pageNumber="10">7.18.0</td>
<td id="76724F0D4645000573D5A0D6BF8BA0BB" box="[1177,1249,471,492]" gridcol="7" gridrow="2" pageId="9" pageNumber="10">3.03.1</td>
</tr>
</table>
</paragraph>
<paragraph id="8B2C24334645FFF377D3A0FEBE54A3E3" pageId="9" pageNumber="10">
<table id="F993D6934645000577DBA0FEBEFDA3E3" box="[151,1431,511,692]" gridcols="5" gridrows="5" pageId="9" pageNumber="10">
<tr id="35A326714645000577DBA0FEBEFDA343" box="[151,1431,511,532]" gridrow="0" pageId="9" pageNumber="10">
<th id="76724F0D4645000577DBA0FEB94DA343" box="[151,551,511,532]" gridcol="0" gridrow="0" pageId="9" pageNumber="10">35 (2.15 ±0.21) (3.05 ±0.35) 1.0</th>
<th id="76724F0D464500057535A0FEB840A343" box="[633,810,511,532]" gridcol="1" gridrow="0" pageId="9" pageNumber="10">(7.55 ±0.64) 20.5</th>
<th id="76724F0D4645000574C5A0FEB8D9A343" box="[905,947,511,532]" gridcol="2" gridrow="0" pageId="9" pageNumber="10">31.6</th>
<th id="76724F0D46450005735EA0FEBF2DA343" box="[1042,1095,511,532]" gridcol="3" gridrow="0" pageId="9" pageNumber="10">3.1</th>
<th id="76724F0D4645000573D5A0FEBEFDA343" box="[1177,1431,511,532]" gridcol="4" gridrow="0" pageId="9" pageNumber="10">(3.05 ±0.07) 2.1</th>
</tr>
<tr id="35A326714645000577DBA326BEFDA36B" box="[151,1431,551,572]" gridrow="1" pageId="9" pageNumber="10" rowspan-2="1">
<th id="76724F0D4645000577DBA326B94DA36B" box="[151,551,551,572]" gridcol="0" gridrow="1" pageId="9" pageNumber="10">(n=2) (n=2) (n=2)</th>
<td id="76724F0D464500057535A326B840A36B" box="[633,810,551,572]" gridcol="1" gridrow="1" pageId="9" pageNumber="10">(n=2)</td>
<td id="76724F0D46450005735EA326BF2DA36B" box="[1042,1095,551,572]" gridcol="3" gridrow="1" pageId="9" pageNumber="10">(n=2)</td>
<td id="76724F0D4645000573D5A326BEFDA36B" box="[1177,1431,551,572]" gridcol="4" gridrow="1" pageId="9" pageNumber="10">(n=2)</td>
</tr>
<tr id="35A326714645000577DBA34EBEFDA333" box="[151,1431,591,612]" gridrow="2" pageId="9" pageNumber="10">
<th id="76724F0D4645000577DBA34EB94DA333" box="[151,551,591,612]" gridcol="0" gridrow="2" pageId="9" pageNumber="10">36 2.2 3.3 1.1</th>
<td id="76724F0D464500057535A34EB840A333" box="[633,810,591,612]" gridcol="1" gridrow="2" pageId="9" pageNumber="10">8.5 22.6</td>
<td id="76724F0D4645000574C5A34EB8D9A333" box="[905,947,591,612]" gridcol="2" gridrow="2" pageId="9" pageNumber="10">35.3</td>
<td id="76724F0D46450005735EA34EBF2DA333" box="[1042,1095,591,612]" gridcol="3" gridrow="2" pageId="9" pageNumber="10">3.3</td>
<td id="76724F0D4645000573D5A34EBEFDA333" box="[1177,1431,591,612]" gridcol="4" gridrow="2" pageId="9" pageNumber="10">3.2 2.2</td>
</tr>
<tr id="35A326714645000577DBA376BEFDA3DB" box="[151,1431,631,652]" gridrow="3" pageId="9" pageNumber="10">
<th id="76724F0D4645000577DBA376B94DA3DB" box="[151,551,631,652]" gridcol="0" gridrow="3" pageId="9" pageNumber="10">37 2.2 3.1 1.0</th>
<td id="76724F0D464500057535A376B840A3DB" box="[633,810,631,652]" gridcol="1" gridrow="3" pageId="9" pageNumber="10">8.5 25.2</td>
<td id="76724F0D4645000574C5A376B8D9A3DB" box="[905,947,631,652]" gridcol="2" gridrow="3" pageId="9" pageNumber="10">38.0</td>
<td id="76724F0D46450005735EA376BF2DA3DB" box="[1042,1095,631,652]" gridcol="3" gridrow="3" pageId="9" pageNumber="10">4.0</td>
<td id="76724F0D4645000573D5A376BEFDA3DB" box="[1177,1431,631,652]" gridcol="4" gridrow="3" pageId="9" pageNumber="10">4.5 2.5</td>
</tr>
<tr id="35A326714645000577DBA39EBEFDA3E3" box="[151,1431,671,692]" gridrow="4" pageId="9" pageNumber="10">
<th id="76724F0D4645000577DBA39EB94DA3E3" box="[151,551,671,692]" gridcol="0" gridrow="4" pageId="9" pageNumber="10">38 2.0 3.0 1.1</th>
<td id="76724F0D464500057535A39EB840A3E3" box="[633,810,671,692]" gridcol="1" gridrow="4" pageId="9" pageNumber="10">7.8 -</td>
<td id="76724F0D4645000574C5A39EB8D9A3E3" box="[905,947,671,692]" gridcol="2" gridrow="4" pageId="9" pageNumber="10">-</td>
<td id="76724F0D46450005735EA39EBF2DA3E3" box="[1042,1095,671,692]" gridcol="3" gridrow="4" pageId="9" pageNumber="10">3.6</td>
<td id="76724F0D4645000573D5A39EBEFDA3E3" box="[1177,1431,671,692]" gridcol="4" gridrow="4" pageId="9" pageNumber="10">3.5 2.4</td>
</tr>
</table>
</paragraph>
<paragraph id="8B2C24334645FFF377D3A3EEBEFDA253" blockId="9.[159,1431,711,802]" box="[159,1431,751,772]" pageId="9" pageNumber="10">40 (2.80 ±0.23) (3.86 ±0.21) (1.18 ±0.25) (9.80 ±0.41) (34.00 (48.75 (4.02 ±0.20) (4.02 ±0.13) (2.84 ±0.18)</paragraph>
<paragraph id="8B2C24334645FFF3744CA20CB8ADA275" blockId="9.[159,1431,711,802]" box="[768,967,781,802]" pageId="9" pageNumber="10">±1.41) ±1.48)</paragraph>
<paragraph id="8B2C24334645FFF377A8A234BE02A295" pageId="9" pageNumber="10">
<table id="F993D6934645000577DBA234BEFDA295" box="[151,1431,821,962]" gridcols="9" gridrows="4" pageId="9" pageNumber="10">
<tr id="35A326714645000577DBA234BEFDA21D" box="[151,1431,821,842]" gridrow="0" pageId="9" pageNumber="10">
<th id="76724F0D4645000577DBA234BA46A21D" box="[151,300,821,842]" gridcol="0" gridrow="0" pageId="9" pageNumber="10">(n=5)</th>
<th id="76724F0D464500057627A234BAD9A21D" box="[363,435,821,842]" gridcol="1" gridrow="0" pageId="9" pageNumber="10">(n=5)</th>
<th id="76724F0D4645000576BEA234B950A21D" box="[498,570,821,842]" gridcol="2" gridrow="0" pageId="9" pageNumber="10">(n=5)</th>
<th id="76724F0D464500057535A234B9C4A21D" box="[633,686,821,842]" gridcol="3" gridrow="0" pageId="9" pageNumber="10">(n=5)</th>
<th id="76724F0D46450005744CA234B85FA21D" box="[768,821,821,842]" gridcol="4" gridrow="0" pageId="9" pageNumber="10">(n=2)</th>
<th id="76724F0D4645000574C5A234B8D4A21D" box="[905,958,821,842]" gridcol="5" gridrow="0" pageId="9" pageNumber="10">(n=2)</th>
<th id="76724F0D46450005735EA234BF30A21D" box="[1042,1114,821,842]" gridcol="6" gridrow="0" pageId="9" pageNumber="10">(n=5)</th>
<th id="76724F0D4645000573D5A234BF8BA21D" box="[1177,1249,821,842]" gridcol="7" gridrow="0" pageId="9" pageNumber="10">(n=5)</th>
<th id="76724F0D46450005726CA234BEFDA21D" box="[1312,1431,821,842]" gridcol="8" gridrow="0" pageId="9" pageNumber="10">(n=5)</th>
</tr>
<tr id="35A326714645000577DBA25CBEFDA225" box="[151,1431,861,882]" gridrow="1" pageId="9" pageNumber="10">
<th id="76724F0D4645000577DBA25CBA46A225" box="[151,300,861,882]" gridcol="0" gridrow="1" pageId="9" pageNumber="10">41 2.3</th>
<td id="76724F0D464500057627A25CBAD9A225" box="[363,435,861,882]" gridcol="1" gridrow="1" pageId="9" pageNumber="10">2.9</td>
<td id="76724F0D4645000576BEA25CB950A225" box="[498,570,861,882]" gridcol="2" gridrow="1" pageId="9" pageNumber="10">0.6</td>
<td id="76724F0D464500057535A25CB9C4A225" box="[633,686,861,882]" gridcol="3" gridrow="1" pageId="9" pageNumber="10">8.3</td>
<td id="76724F0D46450005744CA25CB85FA225" box="[768,821,861,882]" gridcol="4" gridrow="1" pageId="9" pageNumber="10">26.0</td>
<td id="76724F0D4645000574C5A25CB8D4A225" box="[905,958,861,882]" gridcol="5" gridrow="1" pageId="9" pageNumber="10">38.3</td>
<td id="76724F0D46450005735EA25CBF30A225" box="[1042,1114,861,882]" gridcol="6" gridrow="1" pageId="9" pageNumber="10">3.1</td>
<td id="76724F0D4645000573D5A25CBF8BA225" box="[1177,1249,861,882]" gridcol="7" gridrow="1" pageId="9" pageNumber="10">3.8</td>
<td id="76724F0D46450005726CA25CBEFDA225" box="[1312,1431,861,882]" gridcol="8" gridrow="1" pageId="9" pageNumber="10">2.6</td>
</tr>
<tr id="35A326714645000577DBA284BEFDA2CD" box="[151,1431,901,922]" gridrow="2" pageId="9" pageNumber="10">
<th id="76724F0D4645000577DBA284BA46A2CD" box="[151,300,901,922]" gridcol="0" gridrow="2" pageId="9" pageNumber="10">42 2.3</th>
<td id="76724F0D464500057627A284BAD9A2CD" box="[363,435,901,922]" gridcol="1" gridrow="2" pageId="9" pageNumber="10">5.6</td>
<td id="76724F0D4645000576BEA284B950A2CD" box="[498,570,901,922]" gridcol="2" gridrow="2" pageId="9" pageNumber="10">0.5</td>
<td id="76724F0D464500057535A284B9C4A2CD" box="[633,686,901,922]" gridcol="3" gridrow="2" pageId="9" pageNumber="10">-</td>
<td id="76724F0D46450005744CA284B85FA2CD" box="[768,821,901,922]" gridcol="4" gridrow="2" pageId="9" pageNumber="10">27.0</td>
<td id="76724F0D4645000574C5A284B8D4A2CD" box="[905,958,901,922]" gridcol="5" gridrow="2" pageId="9" pageNumber="10">39.3</td>
<td id="76724F0D46450005735EA284BF30A2CD" box="[1042,1114,901,922]" gridcol="6" gridrow="2" pageId="9" pageNumber="10">3.7</td>
<td id="76724F0D4645000573D5A284BF8BA2CD" box="[1177,1249,901,922]" gridcol="7" gridrow="2" pageId="9" pageNumber="10">4.2</td>
<td id="76724F0D46450005726CA284BEFDA2CD" box="[1312,1431,901,922]" gridcol="8" gridrow="2" pageId="9" pageNumber="10">1.9</td>
</tr>
<tr id="35A326714645000577DBA2ACBEFDA295" box="[151,1431,941,962]" gridrow="3" pageId="9" pageNumber="10" rowspan-4="1" rowspan-5="1">
<th id="76724F0D4645000577DBA2ACBA46A295" box="[151,300,941,962]" gridcol="0" gridrow="3" pageId="9" pageNumber="10">2.22.3</th>
<td id="76724F0D464500057627A2ACBAD9A295" box="[363,435,941,962]" gridcol="1" gridrow="3" pageId="9" pageNumber="10">1.62.1</td>
<td id="76724F0D4645000576BEA2ACB950A295" box="[498,570,941,962]" gridcol="2" gridrow="3" pageId="9" pageNumber="10">0.50.6</td>
<td id="76724F0D464500057535A2ACB9C4A295" box="[633,686,941,962]" gridcol="3" gridrow="3" pageId="9" pageNumber="10">-</td>
<td id="76724F0D46450005735EA2ACBF30A295" box="[1042,1114,941,962]" gridcol="6" gridrow="3" pageId="9" pageNumber="10">2.42.8</td>
<td id="76724F0D4645000573D5A2ACBF8BA295" box="[1177,1249,941,962]" gridcol="7" gridrow="3" pageId="9" pageNumber="10">2.82.9</td>
<td id="76724F0D46450005726CA2ACBEFDA295" box="[1312,1431,941,962]" gridcol="8" gridrow="3" pageId="9" pageNumber="10">1.41.5</td>
</tr>
</table>
</paragraph>
<paragraph id="8B2C24334645FFF377D3A2D4BF06A615" blockId="9.[159,1431,981,1042]" lastBlockId="9.[151,1436,1639,2013]" pageId="9" pageNumber="10">
43 (2.23 ±0.11) (1.86 ±0.34) (0.58 ±0.04) - 21.0 32.5 (2.60 ±0.28) (2.85 ±0.07) (1.48 ±0.04) (n=2) (n=2) (n=2) - (n=2) (n=2) (n=2)
<emphasis id="B9E7F8214645FFF37789A766BAE2A7D6" bold="true" box="[197,392,1639,1665]" pageId="9" pageNumber="10">Colour pattern.</emphasis>
Colouration of tadpole in preservative: body semitransparent, dorsolaterally light brown, greyish to yellowish, ventrally whitish. Body marbled with dark brown to dark green irregular spots. Dorsal and dorsolateral pigmentation of body is more dense than on the venter. On the ventral side, the throat and chest region is somewhat stronger pigmented especially around the oral disc. Tail musculature is yellowish, upper and lower tail fin almost transparent. The tail is covered with green to grey pigments and single dark brown irregular spots. Dorsolateral parts of hind limbs with green to grey pigments.
</paragraph>
<caption id="DFEC74BB4645FFF377DBA4FFBEF6A76E" httpUri="https://zenodo.org/record/186591/files/figure.png" pageId="9" pageNumber="10" targetBox="[157,1426,1119,1496]" targetPageId="9">
<paragraph id="8B2C24334645FFF377DBA4FFBEF6A76E" blockId="9.[151,1437,1534,1593]" pageId="9" pageNumber="10">
<emphasis id="B9E7F8214645FFF377DBA4FFBA4AA741" bold="true" box="[151,288,1534,1558]" pageId="9" pageNumber="10">FIGURE 4.</emphasis>
Drawings of the preserved tadpole of
<taxonomicName id="4C935FB04645FFF37595A4FEB8B7A741" box="[729,989,1535,1558]" class="Amphibia" family="Ranidae" genus="Hylarana" kingdom="Animalia" order="Anura" pageId="9" pageNumber="10" phylum="Chordata" rank="species" species="maosonensis">
<emphasis id="B9E7F8214645FFF37595A4FEB8B7A741" box="[729,989,1535,1558]" italics="true" pageId="9" pageNumber="10">Hylarana maosonensis</emphasis>
</taxonomicName>
(stage 36) from Phong Nha Ke Bang National Park, Quang Binh Province, Vietnam; (A) dorsal view, (B) lateral view, (C) oral disc. Drawings by Anna Gawor
</paragraph>
</caption>
<paragraph id="8B2C24334645FFF37789A64EB8EBA68A" blockId="9.[151,1436,1639,2013]" pageId="9" pageNumber="10">Colouration in life: body olive-coloured to brownish with gold-coloured pigments. Furthermore, body and tail covered with remarkable dark flecks. Iris of eye marbled with four black flecks at the outermost margin. Ventral side of body is transparent to white, intestinal spiral slightly visible, with a bright and slightly pigmented fleck at the throat. Internal gills reddish glimmering.</paragraph>
<paragraph id="8B2C24334646FFF07789A196B8A5A0CD" blockId="10.[151,1437,151,2034]" pageId="10" pageNumber="11">
<emphasis id="B9E7F8214646FFF07789A196B979A1E6" bold="true" box="[197,531,151,177]" pageId="10" pageNumber="11">Description in dorsal view:</emphasis>
Body of tadpole reaches a length of
<quantity id="4C6B89D64646FFF074F2A199BF43A1E5" box="[958,1065,152,178]" metricMagnitude="-2" metricUnit="m" metricValue="1.27" pageId="10" pageNumber="11" unit="mm" value="12.7">12.7 mm</quantity>
, oval elongated, with a rounded snout. Maximum body width is between eye and spiracle (maximum body width 0.57 of body length). Nares anterodorsally positioned and directed, nearer to the tip of the snout than to the pupil (rostro-narial distance 0.50 of naro-pupilar distance). Internarial distance 0.61 of interpupilar distance. Eyes dorsolaterally positioned and directed, at the end of the first body third. Eyes are of remarkable size (maximum diameter of eye 0.13 of body length). Maximum interpupilar distance is 0.61 of body width. Muscles of tail are widest at base (width of tail musculature at base 0.44 of maximum body width).
</paragraph>
<paragraph id="8B2C24334646FFF07789A0A7B9B5A3A1" blockId="10.[151,1437,151,2034]" pageId="10" pageNumber="11">
<emphasis id="B9E7F8214646FFF07789A0A7B949A097" bold="true" box="[197,547,422,448]" pageId="10" pageNumber="11">Description in lateral view:</emphasis>
Body laterally depressed (maximum body height 0.65 of maximum body width). Cone-shaped spiracle sinistral, laterally positioned at the beginning of the last third of body, posterodorsally directed. Vent tube separated from body. Distance of the spiracle opening from centre of eye is
<quantity id="4C6B89D64646FFF077DBA31BBB9CA363" box="[151,246,538,564]" metricMagnitude="-3" metricUnit="m" metricValue="5.4" pageId="10" pageNumber="11" unit="mm" value="5.4">5.4 mm</quantity>
. Vent tube ventromedially located between hind limbs. Outer wall of vent tube adhered to lower fin. Cloacal aperture is situated dextral in dorsal view. Tail length is 1.78 of body length, with a pointed tail tip. Tail musculature distinct (height of the tail musculature at base 0.70 of body height), running parallel from the base to the end of the first third of tail fin, then gradually tapering to the tip of the tail. Upper tail fin situated behind the base of tail, slightly convex, with maximum height at the last third of tail fin. Maximum height of lower fin is 0.55 of maximum height of upper fin.
</paragraph>
<paragraph id="8B2C24334646FFF07789A203BE6EA553" blockId="10.[151,1437,151,2034]" pageId="10" pageNumber="11">
<emphasis id="B9E7F8214646FFF07789A203BA22A24B" bold="true" box="[197,328,770,796]" pageId="10" pageNumber="11">Oral disc.</emphasis>
Oral disc anteroventrally positioned (oral disc width 0.46 of maximum body width) and laterally emarginated. Oral disc framed by finger-shaped, marginal papillae except for a large medial gap on the upper labium. Marginal papillae of the lower labium elongated. Single submarginal papillae situated laterally at both ends of the beak between the upper and lower labium. Two additional rows of submarginal papillae situated on the lower labium. Pigmentation is somewhat stronger at the end of papillae. Keratodont row formula 2(2)/3(1), with about 34 keratodonts per
<quantity id="4C6B89D64646FFF0745DA2C5B805A289" box="[785,879,964,990]" metricMagnitude="-4" metricUnit="m" metricValue="5.0" pageId="10" pageNumber="11" unit="mm" value="0.5">0.5 mm</quantity>
. Jaw sheaths black, robust, and serrated. Upper jaw sheath remarkably developed and curved with long appendices. Lower jaw sheath V-shaped.
</paragraph>
<paragraph id="8B2C24334646FFF07789A511B8EDA5C8" blockId="10.[151,1437,151,2034]" pageId="10" pageNumber="11">
<emphasis id="B9E7F8214646FFF07789A511BAC9A57D" bold="true" box="[197,419,1040,1066]" pageId="10" pageNumber="11">Glandular Zones.</emphasis>
Areas with a high density of secretory glands are oval-shaped, whitish and with dark brown pigments. Glands are situated on each side of the body, dorsally between the anterior corner of the eyes, dorsolaterally at midbody and laterally at the end of body. Additional secretory glands are situated ventrally on the throat and ventrolaterally before the hind limbs.
</paragraph>
<paragraph id="8B2C24334646FFF0778AA5AABF50A5BB" blockId="10.[151,1437,151,2034]" pageId="10" pageNumber="11">
<emphasis id="B9E7F8214646FFF0778AA5AABA9BA592" bold="true" box="[198,497,1195,1221]" pageId="10" pageNumber="11">Measurements (in mm).</emphasis>
BH 4.7; BL 12.7; BW 7.2; ED 1.6;
<collectingCountry id="F38464A34646FFF074ECA5ADB8BEA591" box="[928,980,1196,1222]" name="India" pageId="10" pageNumber="11">IND</collectingCountry>
2.7; IP 4.4; LF 1.2; MTH 6.1; NK 34; NP 85; NPD 2.2; ODW 3.3; RND 1.0; TAL 22.6; TMH 3.3; TMW 3.2; UF 2.2.
</paragraph>
<paragraph id="8B2C24334646FFF0778AA5F9BFE9A483" blockId="10.[151,1437,151,2034]" pageId="10" pageNumber="11">
<emphasis id="B9E7F8214646FFF0778AA5F9B976A445" bold="true" box="[198,540,1272,1298]" pageId="10" pageNumber="11">Variation within the series.</emphasis>
Series with 19 tadpoles in stages 2843. Proportions vary as follows: BW 0.330.73 of BL, ED 0.110.18 of BL, IP 0.531.08 of BW, RND 0.220.69 of NPD,
<collectingCountry id="F38464A34646FFF073EDA421BFBFA46D" box="[1185,1237,1312,1338]" name="India" pageId="10" pageNumber="11">IND</collectingCountry>
0.340.70 of IP, TMW 0.330.73 of BW, BH 0.611.33 of BW, TAL 1.522.36 of BL, TMH 0.470.70 of 0.70, TMH 0.480.71 of MTH, UF 0.330.43 of MTH, LF 0.310.76 of UF, ODW 0.250.73 of BW. Keratodont row formula is homogenous within specimens. In earlier stages, oral papillae are less developed, the skin of body is more transparent and the internal organs are well visible in lateral and ventral view.
</paragraph>
<paragraph id="8B2C24334646FFF07789A4E1BBB9A6A5" blockId="10.[151,1437,151,2034]" pageId="10" pageNumber="11">
<emphasis id="B9E7F8214646FFF07789A4E1BAE6A4AD" bold="true" box="[197,396,1504,1530]" pageId="10" pageNumber="11">Natural history.</emphasis>
Larvae of
<taxonomicName id="4C935FB04646FFF07542A4E0B9A2A4AD" box="[526,712,1505,1530]" class="Amphibia" family="Ranidae" genus="Hylarana" kingdom="Animalia" order="Anura" pageId="10" pageNumber="11" phylum="Chordata" rank="species" species="maosonensis">
<emphasis id="B9E7F8214646FFF07542A4E0B9A2A4AD" box="[526,712,1505,1530]" italics="true" pageId="10" pageNumber="11">H. maosonensis</emphasis>
</taxonomicName>
were collected in a primary forest stream pool below a small cascade during the dry season between
<date id="FF2D02F34646FFF07530A709B84AA775" box="[636,800,1544,1570]" pageId="10" pageNumber="11" value="2006-06-20">20 June 2006</date>
and
<date id="FF2D02F34646FFF07410A709BF6AA775" box="[860,1024,1544,1570]" pageId="10" pageNumber="11" value="2006-07-30">30 July 2006</date>
. The tadpoles were hidden on the ground between leaves and rock crevices. Further tadpoles were found in a water catchment basin next to the highway at km 51 and
<quantity id="4C6B89D64646FFF076E7A754BA82A738" box="[427,488,1621,1647]" metricMagnitude="0" metricUnit="m" metricValue="1.3462" pageId="10" pageNumber="11" unit="in" value="53.0">53 in</quantity>
the U Bo region, Phong Nha Ke Bang National Park,
<collectingCountry id="F38464A34646FFF073C2A754BF9CA738" box="[1166,1270,1621,1647]" name="Vietnam" pageId="10" pageNumber="11">Vietnam</collectingCountry>
, at
<date id="FF2D02F34646FFF0726DA754BEEBA738" box="[1313,1409,1621,1647]" pageId="10" pageNumber="11" value="1920-06">20 June</date>
,
<date id="FF2D02F34646FFF072C1A754BAD6A7C1" pageId="10" pageNumber="11" value="2006-07-07" valueMax="2006-08-22" valueMin="2006-07-07">7 July and 22 August 2006</date>
. These larvae were hidden between algae on the ground of the water body during day time (if algae were present, otherwise they were exposed on the sand or gravel of the ground), but were found swimming in the open water during night time. Once disturbed, the larvae tried to escape deep into the mud on the ground. During night time (08:
<specimenCount id="9D95EFBA4646FFF07565A7F1B90CA65D" box="[553,614,1776,1802]" pageId="10" pageNumber="11" type="generic">30 p.</specimenCount>
m.) of
<date id="FF2D02F34646FFF075FAA7F1B828A65D" box="[694,834,1776,1802]" pageId="10" pageNumber="11" value="2006-07-07">7 July 2006</date>
, water temperature was 26°C. During day time, the water catchment basin was at least in part exposed to the sun. At day time of
<date id="FF2D02F34646FFF07366A617BF8DA667" box="[1066,1255,1814,1840]" pageId="10" pageNumber="11" value="2006-08-22">22 August 2006</date>
(12:00) the
<collectingCountry id="F38464A34646FFF07234A617BEF6A667" box="[1400,1436,1814,1840]" name="Philippines" pageId="10" pageNumber="11">pH</collectingCountry>
value was 5.5. Adult
<taxonomicName id="4C935FB04646FFF076DDA63CB921A601" box="[401,587,1853,1878]" class="Amphibia" family="Ranidae" genus="Hylarana" kingdom="Animalia" order="Anura" pageId="10" pageNumber="11" phylum="Chordata" rank="species" species="maosonensis">
<emphasis id="B9E7F8214646FFF076DDA63CB921A601" box="[401,587,1853,1878]" italics="true" pageId="10" pageNumber="11">H. maosonensis</emphasis>
</taxonomicName>
(including calling males) were found during night time on forest stream rocks and cascades as well as around water catchment basins next to the highway at km 51 and
<quantity id="4C6B89D64646FFF073BDA665BE46A629" box="[1265,1324,1892,1918]" metricMagnitude="0" metricUnit="m" metricValue="1.3462" pageId="10" pageNumber="11" unit="in" value="53.0">53 in</quantity>
the U Bo region between
<date id="FF2D02F34646FFF07618A68BB912A6F3" box="[340,632,1930,1956]" pageId="10" pageNumber="11" value="2006-06-20" valueMax="2006-07-08" valueMin="2006-06-20">20 June and 8 July 2006</date>
. Air temperature at night of
<date id="FF2D02F34646FFF07487A68BBF3CA6F3" box="[971,1110,1930,1956]" pageId="10" pageNumber="11" value="2006-07-08">8 July 2006</date>
next to a forest stream was 29°C. Once disturbed, the adult frogs tried to jump into the water where they remained motionless for some time.
</paragraph>
</subSubSection>
</treatment>
</document>