treatments-xml/data/F7/38/87/F738879E600F9C254084F9D57E7FEDA1.xml

<document id="7710B7DFD5F6FECCC8196EB57292EBF6" ID-DOI="10.5281/zenodo.177568" ID-GBIF-Dataset="e50c438f-5703-46d8-b182-6378a55e288d" ID-ISSN="1175-5326" ID-Zenodo-Dep="177568" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1459868792221" checkinUser="plazi" docAuthor="Nik, Grzegorz Pa Ś" docDate="2007" docId="F738879E600F9C254084F9D57E7FEDA1" docLanguage="en" docName="zt01527p038.pdf" docOrigin="Zootaxa 1527" docStyle="DocumentStyle:6581673A57F01A3145754A1E615EDFF0.4:Zootaxa.2007-2008.journal_article" docStyleId="6581673A57F01A3145754A1E615EDFF0" docStyleName="Zootaxa.2007-2008.journal_article" docStyleVersion="4" docTitle="Thendelecrotona Nik, 2007, gen. n." docType="treatment" docVersion="8" lastPageNumber="36" masterDocId="0B01FFE6600F9C204013FFB07C74EE73" masterDocTitle="A new genus and species of the aleocharine tribe Thamiaraeini (Coleoptera, Staphylinidae) from South Africa" masterLastPageNumber="38" masterPageNumber="31" pageNumber="31" updateTime="1698226395473" updateUser="plazi">
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<mods:title id="972C2AA7B400317BB46386496FD80D45">A new genus and species of the aleocharine tribe Thamiaraeini (Coleoptera, Staphylinidae) from South Africa</mods:title>
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<mods:namePart id="1036193B86D984D249AFBC033A712320">Nik, Grzegorz Pa Ś</mods:namePart>
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<mods:date id="D45D1FB9BF93C47E5E9700D22D730B3F">2007</mods:date>
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<treatment id="F738879E600F9C254084F9D57E7FEDA1" ID-DOI="http://doi.org/10.5281/zenodo.6240759" ID-GBIF-Taxon="119369052" ID-Zenodo-Dep="6240759" LSID="urn:lsid:plazi:treatment:F738879E600F9C254084F9D57E7FEDA1" httpUri="http://treatment.plazi.org/id/F738879E600F9C254084F9D57E7FEDA1" lastPageId="5" lastPageNumber="36" pageId="0" pageNumber="31">
<subSubSection id="378B6503600F9C204084F9D57D63E8D1" pageId="0" pageNumber="31" type="nomenclature">
<paragraph id="7F2E3688600F9C204084F9D57DCAE80C" blockId="0.[151,446,1637,1698]" box="[151,446,1637,1663]" pageId="0" pageNumber="31">
<heading id="246681E4600F9C204084F9D57DCAE80C" bold="true" box="[151,446,1637,1663]" fontSize="11" level="1" pageId="0" pageNumber="31" reason="1">
<emphasis id="4DE5EA9A600F9C204084F9D57DCAE80C" bold="true" box="[151,446,1637,1663]" pageId="0" pageNumber="31">
<taxonomicName id="B8914D0B600F9C204084F9D57D10E80C" ID-CoL="8X7WW" box="[151,356,1637,1663]" class="Insecta" family="Staphylinidae" genus="Thendelecrotona" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="0" pageNumber="31" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="4DE5EA9A600F9C204084F9D57D10E80C" bold="true" box="[151,356,1637,1663]" italics="true" pageId="0" pageNumber="31">Thendelecrotona</emphasis>
</taxonomicName>
<taxonomicNameLabel id="56D657E1600F9C204178F9D57DCAE80C" box="[363,446,1637,1663]" pageId="0" pageNumber="31" rank="genus">gen. n.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="7F2E3688600F9C204084F9387D63E8D1" blockId="0.[151,446,1637,1698]" box="[151,279,1672,1698]" pageId="0" pageNumber="31">
(
<figureCitation id="E7AA2A0D600F9C20408CF9387D7AE8D1" box="[159,270,1672,1698]" captionStart-0="FIGURE 1" captionStart-1="FIGURES 2 – 6" captionStartId-0="1.[151,255,1832,1856]" captionStartId-1="2.[151,269,1956,1980]" captionTargetBox-0="[453,1118,195,1794]" captionTargetBox-1="[185,1408,204,1925]" captionTargetId-0="figure@1.[431,1155,169,1806]" captionTargetId-1="figure@2.[169,1418,194,1933]" captionTargetPageId-0="1" captionTargetPageId-1="2" captionText-0="FIGURE 1. Habitus of Thendelecrotona natalica gen. &amp; sp. n. Scale bar 0.5 mm." captionText-1="FIGURES 2 – 6. Mouthparts of Thendelecrotona natalica gen. &amp; sp. n. 2 — right maxilla, 3 — labium, 4 — right mandible, 5 — labrum, 6 — mentum. Scale bars 0.1 mm." httpUri-0="https://zenodo.org/record/177569/files/figure.png" httpUri-1="https://zenodo.org/record/177570/files/figure.png" pageId="0" pageNumber="31">Figs. 1–6</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="378B6503600F9C204084F9677EC2E881" box="[151,694,1751,1778]" pageId="0" pageNumber="31" type="materials_examined">
<paragraph id="7F2E3688600F9C204084F9677EC2E881" blockId="0.[151,1436,1751,2018]" box="[151,694,1751,1778]" pageId="0" pageNumber="31">
<emphasis id="4DE5EA9A600F9C204084F9677D4FE882" bold="true" box="[151,315,1751,1777]" pageId="0" pageNumber="31">
<typeStatus id="A02A882A600F9C204084F9677CA0E882" box="[151,212,1751,1777]" pageId="0" pageNumber="31">Type</typeStatus>
species:
</emphasis>
<taxonomicName id="B8914D0B600F9C204150F9687E1BE882" box="[323,623,1752,1777]" class="Insecta" family="Staphylinidae" genus="Thendelecrotona" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="0" pageNumber="31" phylum="Arthropoda" rank="species" species="natalica" status="sp. nov.">
<emphasis id="4DE5EA9A600F9C204150F9687E1BE882" box="[323,623,1752,1777]" italics="true" pageId="0" pageNumber="31">Thendelecrotona natalica</emphasis>
</taxonomicName>
<taxonomicNameLabel id="56D657E1600F9C204264F9687EC2E881" box="[631,694,1752,1778]" pageId="0" pageNumber="31" rank="species">sp. n.</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection id="378B6503600F9C2040D5F94F7F76E9C9" pageId="0" pageNumber="31" type="diagnosis">
<paragraph id="7F2E3688600F9C2040D5F94F7F76E9C9" blockId="0.[151,1436,1751,2018]" pageId="0" pageNumber="31">
<emphasis id="4DE5EA9A600F9C2040D5F94F7D32E96A" bold="true" box="[198,326,1791,1817]" pageId="0" pageNumber="31">Diagnosis.</emphasis>
<taxonomicName id="B8914D0B600F9C20415CF8B07E60E96A" box="[335,532,1792,1817]" class="Insecta" family="Staphylinidae" genus="Thendelecrotona" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="0" pageNumber="31" phylum="Arthropoda" rank="genus">
<emphasis id="4DE5EA9A600F9C20415CF8B07E60E96A" box="[335,532,1792,1817]" italics="true" pageId="0" pageNumber="31">Thendelecrotona</emphasis>
</taxonomicName>
can be recognized by the combination of the following characters: the head with infraorbital ridge, the maxillary palpus with 4 articles and a moderately defined apical pseudosegment, the labial palpi 2-jointed, the ligula entire (not divided into lobes), the mentum with produced anterior angles, the pronotal hypomera not visible in lateral view, the mesocoxae narrowly separated, the elytral postero-lateral angles strongly sinuate, the tarsal formula 4-5-5.
</paragraph>
</subSubSection>
<subSubSection id="378B6503600F9C2540D5F87779F0EDF1" lastPageId="5" lastPageNumber="36" pageId="0" pageNumber="31" type="description">
<paragraph id="7F2E3688600F9C2040D5F87778C0E991" blockId="0.[151,1436,1751,2018]" box="[198,1204,1991,2018]" pageId="0" pageNumber="31">
<emphasis id="4DE5EA9A600F9C2040D5F8777D2AE992" bold="true" box="[198,350,1991,2017]" pageId="0" pageNumber="31">Description.</emphasis>
Body length 2.2–2.4 mm, broad, convex and moderately glossy (
<figureCitation id="E7AA2A0D600F9C20444EF87878D6E991" box="[1117,1186,1992,2018]" captionStart="FIGURE 1" captionStartId="1.[151,255,1832,1856]" captionTargetBox="[453,1118,195,1794]" captionTargetId="figure@1.[431,1155,169,1806]" captionTargetPageId="1" captionText="FIGURE 1. Habitus of Thendelecrotona natalica gen. &amp; sp. n. Scale bar 0.5 mm." httpUri="https://zenodo.org/record/177569/files/figure.png" pageId="0" pageNumber="31">Fig. 1</figureCitation>
).
</paragraph>
<caption id="2BEE6600600E9C214084F8987870E933" box="[151,1028,1832,1856]" httpUri="https://zenodo.org/record/177569/files/figure.png" pageId="1" pageNumber="32" targetBox="[453,1118,195,1794]" targetPageId="1">
<paragraph id="7F2E3688600E9C214084F8987870E933" blockId="1.[151,1028,1832,1856]" box="[151,1028,1832,1856]" pageId="1" pageNumber="32">
<emphasis id="4DE5EA9A600E9C214084F8987D6DE933" bold="true" box="[151,281,1832,1856]" pageId="1" pageNumber="32">FIGURE 1.</emphasis>
Habitus of 
<taxonomicName id="B8914D0B600E9C214185F8997EDCE933" box="[406,680,1833,1856]" class="Insecta" family="Staphylinidae" genus="Thendelecrotona" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="32" phylum="Arthropoda" rank="species" species="natalica">
<emphasis id="4DE5EA9A600E9C214185F8997EDCE933" box="[406,680,1833,1856]" italics="true" pageId="1" pageNumber="32">Thendelecrotona natalica</emphasis>
</taxonomicName>
<emphasis id="4DE5EA9A600E9C2142BDF8987EA9E933" bold="true" box="[686,733,1832,1856]" pageId="1" pageNumber="32">gen.</emphasis>
&amp; 
<emphasis id="4DE5EA9A600E9C2142ECF8987F4FE933" bold="true" box="[767,827,1832,1856]" pageId="1" pageNumber="32">sp. n.</emphasis>
Scale bar 0.5 mm.
</paragraph>
</caption>
<paragraph id="7F2E3688600E9C2140D5F8C37F77E9AE" blockId="1.[151,1436,1907,2013]" pageId="1" pageNumber="32">Head. Circular in outline, temples arcuately narrowed to hind angles; eyes weakly protruding from lateral contours of head; head with infraorbital ridge complete to maxillary insertion. Antennae short, clearly increasing in width apically, extending to base of pronotum.</paragraph>
<caption id="2BEE6600600D9C224084F8147E05E9AD" httpUri="https://zenodo.org/record/177570/files/figure.png" pageId="2" pageNumber="33" targetBox="[185,1408,204,1925]" targetPageId="2">
<paragraph id="7F2E3688600D9C224084F8147E05E9AD" blockId="2.[151,1436,1956,2014]" pageId="2" pageNumber="33">
<emphasis id="4DE5EA9A600D9C224084F8147D37E9CF" bold="true" box="[151,323,1956,1980]" pageId="2" pageNumber="33">FIGURES 2–6.</emphasis>
Mouthparts of 
<taxonomicName id="B8914D0B600D9C2241FBF8157E8EE9CF" box="[488,762,1957,1980]" class="Insecta" family="Staphylinidae" genus="Thendelecrotona" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="33" phylum="Arthropoda" rank="species" species="natalica">
<emphasis id="4DE5EA9A600D9C2241FBF8157E8EE9CF" box="[488,762,1957,1980]" italics="true" pageId="2" pageNumber="33">Thendelecrotona natalica</emphasis>
</taxonomicName>
<emphasis id="4DE5EA9A600D9C224313F8147F5BE9CF" bold="true" box="[768,815,1956,1980]" pageId="2" pageNumber="33">gen.</emphasis>
&amp; 
<emphasis id="4DE5EA9A600D9C224342F8147FF8E9CF" bold="true" box="[849,908,1956,1980]" pageId="2" pageNumber="33">sp. n.</emphasis>
2—right maxilla, 3—labium, 4—right mandible, 5—labrum, 6—mentum. Scale bars 0.1 mm.
</paragraph>
</caption>
<caption id="2BEE6600600C9C234084FA6E7D9FE86B" httpUri="https://zenodo.org/record/177571/files/figure.png" pageId="3" pageNumber="34" targetBox="[159,1413,190,1473]" targetPageId="3">
<paragraph id="7F2E3688600C9C234084FA6E7D9FE86B" blockId="3.[151,1436,1502,1560]" pageId="3" pageNumber="34">
<emphasis id="4DE5EA9A600C9C234084FA6E7D30EB85" bold="true" box="[151,324,1502,1526]" pageId="3" pageNumber="34">FIGURES 7–8.</emphasis>
Genitalia of 
<taxonomicName id="B8914D0B600C9C2341C5FA6E7E9DEB86" box="[470,745,1502,1525]" class="Insecta" family="Staphylinidae" genus="Thendelecrotona" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="34" phylum="Arthropoda" rank="species" species="natalica">
<emphasis id="4DE5EA9A600C9C2341C5FA6E7E9DEB86" box="[470,745,1502,1525]" italics="true" pageId="3" pageNumber="34">Thendelecrotona natalica</emphasis>
</taxonomicName>
<emphasis id="4DE5EA9A600C9C2342E1FA6E7F54EB85" bold="true" box="[754,800,1502,1526]" pageId="3" pageNumber="34">gen.</emphasis>
&amp; 
<emphasis id="4DE5EA9A600C9C234355FA6E7FF7EB85" bold="true" box="[838,899,1502,1526]" pageId="3" pageNumber="34">sp. n.</emphasis>
7—median lobe of aedeagus in lateral view, 8— spermatheca. Scale bar 0.1 mm.
</paragraph>
</caption>
<paragraph id="7F2E3688600C9C2440D5F9C37FCEEFD1" blockId="3.[151,1437,1651,2037]" lastBlockId="4.[151,1436,152,418]" lastPageId="4" lastPageNumber="35" pageId="3" pageNumber="34">
Mouthparts. Maxilla (
<figureCitation id="E7AA2A0D600C9C2341D8F9C37E66E8FE" box="[459,530,1651,1677]" captionStart="FIGURES 2 – 6" captionStartId="2.[151,269,1956,1980]" captionTargetBox="[185,1408,204,1925]" captionTargetId="figure@2.[169,1418,194,1933]" captionTargetPageId="2" captionText="FIGURES 2 – 6. Mouthparts of Thendelecrotona natalica gen. &amp; sp. n. 2 — right maxilla, 3 — labium, 4 — right mandible, 5 — labrum, 6 — mentum. Scale bars 0.1 mm." httpUri="https://zenodo.org/record/177570/files/figure.png" pageId="3" pageNumber="34">Fig. 2</figureCitation>
) with lacinia longer than galea; galea narrow, rounded apically, subequal in width to lacinia; apex of galea with membranous lobe in apical 1/3; apical and outer margin of lobe covered with several long setae, inner and middle areas of lobe with two longitudinal rows of dense setae (
<figureCitation id="E7AA2A0D600C9C234551F97379F9E8AE" box="[1346,1421,1731,1757]" captionStart="FIGURES 2 – 6" captionStartId="2.[151,269,1956,1980]" captionTargetBox="[185,1408,204,1925]" captionTargetId="figure@2.[169,1418,194,1933]" captionTargetPageId="2" captionText="FIGURES 2 – 6. Mouthparts of Thendelecrotona natalica gen. &amp; sp. n. 2 — right maxilla, 3 — labium, 4 — right mandible, 5 — labrum, 6 — mentum. Scale bars 0.1 mm." httpUri="https://zenodo.org/record/177570/files/figure.png" pageId="3" pageNumber="34">Fig. 2</figureCitation>
); inner margin of lacinia with spines and long setae; maxillary palpus with 4 articles and a moderately defined apical pseudosegment, article 1 very short, article 2 elongate, slightly widened apically, article 3 distinctly widened apically, longer than article 2, article 4 relatively long, about ½ length of article 3, article 4 with pseudosegment. Labrum (
<figureCitation id="E7AA2A0D600C9C23418BF8D37D95E90E" box="[408,481,1891,1917]" captionStart="FIGURES 2 – 6" captionStartId="2.[151,269,1956,1980]" captionTargetBox="[185,1408,204,1925]" captionTargetId="figure@2.[169,1418,194,1933]" captionTargetPageId="2" captionText="FIGURES 2 – 6. Mouthparts of Thendelecrotona natalica gen. &amp; sp. n. 2 — right maxilla, 3 — labium, 4 — right mandible, 5 — labrum, 6 — mentum. Scale bars 0.1 mm." httpUri="https://zenodo.org/record/177570/files/figure.png" pageId="3" pageNumber="34">Fig. 5</figureCitation>
) transverse, widely emarginate apically and with characteristic arrangement of setae. Labium (
<figureCitation id="E7AA2A0D600C9C23415CF83B7DE3E9D6" box="[335,407,1931,1957]" captionStart="FIGURES 2 – 6" captionStartId="2.[151,269,1956,1980]" captionTargetBox="[185,1408,204,1925]" captionTargetId="figure@2.[169,1418,194,1933]" captionTargetPageId="2" captionText="FIGURES 2 – 6. Mouthparts of Thendelecrotona natalica gen. &amp; sp. n. 2 — right maxilla, 3 — labium, 4 — right mandible, 5 — labrum, 6 — mentum. Scale bars 0.1 mm." httpUri="https://zenodo.org/record/177570/files/figure.png" pageId="3" pageNumber="34">Fig. 3</figureCitation>
) with ligula relatively short and broad, about ¾ length of labial palpomere 1; apex of ligula with several pseudopores; prementum with two moderately long medial setae situated one behind the other, insertions of setae almost adjacent; median pseudopore field narrow and more or less linear, lateral pore field with a single setose pore and two asetose pores; labial palpi 2-jointed, article 1 elongate, narrowed towards apex, with 6 long setae, article 2 slightly shorter than article 1, distinctly widened apically; apex with several very short spinules and numerous pseudopores. Mentum strongly transverse (
<figureCitation id="E7AA2A0D600B9C24449FFF5878A1EF71" box="[1164,1237,232,258]" captionStart="FIGURES 2 – 6" captionStartId="2.[151,269,1956,1980]" captionTargetBox="[185,1408,204,1925]" captionTargetId="figure@2.[169,1418,194,1933]" captionTargetPageId="2" captionText="FIGURES 2 – 6. Mouthparts of Thendelecrotona natalica gen. &amp; sp. n. 2 — right maxilla, 3 — labium, 4 — right mandible, 5 — labrum, 6 — mentum. Scale bars 0.1 mm." httpUri="https://zenodo.org/record/177570/files/figure.png" pageId="4" pageNumber="35">Fig. 6</figureCitation>
), widest at base, apical margin deeply concave, anterior angles strongly produced, 8 long setae on each side present. Mandibles curved towards apex, right mandible (
<figureCitation id="E7AA2A0D600B9C244245FE887EE9EF21" box="[598,669,312,338]" captionStart="FIGURES 2 – 6" captionStartId="2.[151,269,1956,1980]" captionTargetBox="[185,1408,204,1925]" captionTargetId="figure@2.[169,1418,194,1933]" captionTargetPageId="2" captionText="FIGURES 2 – 6. Mouthparts of Thendelecrotona natalica gen. &amp; sp. n. 2 — right maxilla, 3 — labium, 4 — right mandible, 5 — labrum, 6 — mentum. Scale bars 0.1 mm." httpUri="https://zenodo.org/record/177570/files/figure.png" pageId="4" pageNumber="35">Fig. 4</figureCitation>
) with large median tooth at middle of inner margin, strongly serrate in dorsal molar region, “velvety patch” area large, wider than half of mandibular base, composed of combination of anterior row of teeth and basal area of very fine spinules.
</paragraph>
<caption id="2BEE6600600B9C244084FAF67F00EB2D" box="[151,884,1350,1374]" httpUri="https://zenodo.org/record/177572/files/figure.png" pageId="4" pageNumber="35" targetBox="[155,1431,474,1326]" targetPageId="4">
<paragraph id="7F2E3688600B9C244084FAF67F00EB2D" blockId="4.[151,884,1350,1374]" box="[151,884,1350,1374]" pageId="4" pageNumber="35">
<emphasis id="4DE5EA9A600B9C244084FAF67D6DEB2D" bold="true" box="[151,281,1350,1374]" pageId="4" pageNumber="35">FIGURE 9.</emphasis>
Type locality of 
<taxonomicName id="B8914D0B600B9C2441DCFAF77E94EB2D" box="[463,736,1351,1374]" class="Insecta" family="Staphylinidae" genus="Thendelecrotona" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="35" phylum="Arthropoda" rank="species" species="natalica">
<emphasis id="4DE5EA9A600B9C2441DCFAF77E94EB2D" box="[463,736,1351,1374]" italics="true" pageId="4" pageNumber="35">Thendelecrotona natalica</emphasis>
</taxonomicName>
<emphasis id="4DE5EA9A600B9C2442F4FAF67F62EB2D" bold="true" box="[743,790,1350,1374]" pageId="4" pageNumber="35">gen.</emphasis>
&amp; sp. n.
</paragraph>
</caption>
<paragraph id="7F2E3688600B9C2440D5FA0A7D7FE8B7" blockId="4.[151,1436,1466,1972]" pageId="4" pageNumber="35">
Thorax. Pronotum transverse, strongly convex, moderately glossy, widest in the middle, lateral sides arcuate, more strongly narrowed to apex than to base. Hypomera not visible in lateral view. Mesocoxae narrowly separated. Mesoventral process, often referred to as “mesosternal” (see 
<bibRefCitation id="1B004B79600B9C2443F0F9BA781AE857" author="Baehr" box="[995,1134,1546,1572]" pageId="4" pageNumber="37" refString="Baehr, M. (1975) Skelett und Muskulatur des Thorax von Priacma serrata Leconte (Coleoptera, Cupedidae). Zeitschrift fur Morphologie der Tiere, 81, 55 - 101." type="journal article" year="1975">Baehr 1975</bibRefCitation>
, 
<bibRefCitation id="1B004B79600B9C244468F9BA7979E857" author="Beutel" box="[1147,1293,1546,1572]" pageId="4" pageNumber="37" refString="Beutel, R. (1986) Skelet und Muskulatur des Kopfes und Thorax von Hygrobia tarda (Herbst). Ein Beitrag zur Klarung der phylogenetischen Beziehungen der Hydradephaga (Insecta: Coleoptera). Stuttgarter Beitrage zur Naturkunde (A), 388, 1 - 54." type="journal article" year="1986">Beutel 1986</bibRefCitation>
, 
<bibRefCitation id="1B004B79600B9C244509F9BA7CA0E83F" author="Belkaceme" pageId="4" pageNumber="37" refString="Belkaceme, T. (1991) Skelet und Muskulatur des Kopfes und Thorax von Noterus laevis Sturm. Ein Beitrag zur Morphologie und Phylogenie der Noteridae (Coleoptera: Adephaga). Stuttgarter Beitrage zur Naturkunde (A), 462, 1 - 94." type="journal article" year="1991">Belkaceme 1991</bibRefCitation>
, 
<bibRefCitation id="1B004B79600B9C2440F3F9827DEDE83F" author="Lawrence" box="[224,409,1586,1612]" pageId="4" pageNumber="37" refString="Lawrence, J. F. (1999) The Australian Ommatidae (Coleoptera: Archostemata): new species, larva and discussion of relationships. Invertebrate Taxonomy, 13, 369 - 390." type="journal article" year="1999">Lawrence 1999</bibRefCitation>
, 
<bibRefCitation id="1B004B79600B9C2441B6F9827EECE83F" author="Beutel" box="[421,664,1586,1612]" pageId="4" pageNumber="37" refString="Beutel, R. G. &amp; Haas, F. (2000) Phylogenetic relationships of the suborders of Coleoptera (Insecta). Cladistics, 16, 1 - 39." type="journal article" year="2000">Beutel &amp; Haas 2000</bibRefCitation>
, 
<bibRefCitation id="1B004B79600B9C2442B0F9827FBAE83F" author="Beutel" box="[675,974,1586,1612]" pageId="4" pageNumber="37" refString="Beutel, R. G. &amp; Lawrence, J. F. (2005) 4. Coleoptera, Morphology. In: Beutel, G. R. &amp; Leschen, R. A. B. Handbook of Zoology. A Natural History of the Phyla of the Animal Kingdom. Coleoptera, Beetles. Volume 1: Morphology and Systematics (Archostemata, Adephaga, Myxophaga, Polyphaga partim). Walter de Gruyter, Berlin, pp. 23 - 27." type="book chapter" year="2005">Beutel &amp; Lawrence 2005</bibRefCitation>
, 
<bibRefCitation id="1B004B79600B9C2443C9F9827888E83F" author="Friedrich" box="[986,1276,1586,1612]" pageId="4" pageNumber="37" refString="Friedrich, F. &amp; Beutel R. G. (2006) The pterothoracic skeletomuscular system of Scirtoidea (Coleoptera: Polyphaga) and its implications for the high-level phylogeny of beetles. Journal of Zoological Systematics and Evolutionary Research, 44 (4), 290 - 315." type="journal article" year="2006">Friedrich &amp; Beutel 2006</bibRefCitation>
) triangular in outline, pointed at apex, extending to about 2/3 of mesocoxae; metaventral process very short, triangular in outline, broadly rounded at apex, extending to 1/3 of mesocoxae. Mesocoxal cavities margined posteriorly by fine bead.
</paragraph>
<paragraph id="7F2E3688600B9C2440D5F962796FE89F" blockId="4.[151,1436,1466,1972]" box="[198,1307,1746,1772]" pageId="4" pageNumber="35">Elytra. Transverse, lateral sides straight, widest at base, postero-lateral angles strongly sinuate.</paragraph>
<paragraph id="7F2E3688600B9C2440D5F94A7FBBE94F" blockId="4.[151,1436,1466,1972]" pageId="4" pageNumber="35">Abdomen. Acuminate, widest at base, tergites III–IV weakly and shallowly transversely impressed at base, impressions smooth, impunctate, sternites not impressed at base.</paragraph>
<paragraph id="7F2E3688600B9C2440D5F8FA787DE9C7" blockId="4.[151,1436,1466,1972]" pageId="4" pageNumber="35">Legs elongate, metatarsus long, approximately 3/4 the length of metatibia; basal article of metatarsus long, subequal in length to articles 2 and 3 combined, articles 2–4 relatively short, subequal in length, article 5 elongate, subequal in length to articles 3–4 combined; tarsal formula 4-5-5.</paragraph>
<paragraph id="7F2E3688600A9C2540D5FF277DB9EFD1" blockId="5.[151,1437,151,978]" pageId="5" pageNumber="36">
<emphasis id="4DE5EA9A600A9C2540D5FF277D92EEC2" bold="true" box="[198,486,151,177]" pageId="5" pageNumber="36">Comparative remarks.</emphasis>
Based on the tarsal formula (4-5-5) and the number of segments of the labial palpi, 
<taxonomicName id="B8914D0B600A9C2540F0FF707DDCEEAA" box="[227,424,192,217]" class="Insecta" family="Staphylinidae" genus="Thendelecrotona" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="36" phylum="Arthropoda" rank="genus">
<emphasis id="4DE5EA9A600A9C2540F0FF707DDCEEAA" box="[227,424,192,217]" italics="true" pageId="5" pageNumber="36">Thendelecrotona</emphasis>
</taxonomicName>
is assigned to the tribe Thamiaraeini 
<bibRefCitation id="1B004B79600A9C254364FF70786DEEA9" author="Fenyes" box="[887,1049,192,218]" pageId="5" pageNumber="37" refString="Fenyes, A. (1921) New genera and species of Aleocharinae with polytomic synopsis of the tribes. Bulletin of the Museum of Comparative Zoology, 65, 17 - 36." type="journal article" year="1921">Fenyes, 1921</bibRefCitation>
. 
<taxonomicName id="B8914D0B600A9C25443BFF707899EEAA" box="[1064,1261,192,217]" class="Insecta" family="Staphylinidae" genus="Thendelecrotona" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="36" phylum="Arthropoda" rank="genus">
<emphasis id="4DE5EA9A600A9C25443BFF707899EEAA" box="[1064,1261,192,217]" italics="true" pageId="5" pageNumber="36">Thendelecrotona</emphasis>
</taxonomicName>
can be distinguished from the other genera of that tribe by the combination of the following characters: the 4-jointed maxillary palpi with distinct apical pseudosegment (present also in 
<taxonomicName id="B8914D0B600A9C254384FEA07857EF5A" box="[919,1059,272,297]" class="Insecta" family="Staphylinidae" genus="Oxypodinus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="36" phylum="Arthropoda" rank="genus">
<emphasis id="4DE5EA9A600A9C254384FEA07857EF5A" box="[919,1059,272,297]" italics="true" pageId="5" pageNumber="36">Oxypodinus</emphasis>
</taxonomicName>
)(
<figureCitation id="E7AA2A0D600A9C254426FEA078F4EF59" box="[1077,1152,272,298]" captionStart="FIGURES 2 – 6" captionStartId="2.[151,269,1956,1980]" captionTargetBox="[185,1408,204,1925]" captionTargetId="figure@2.[169,1418,194,1933]" captionTargetPageId="2" captionText="FIGURES 2 – 6. Mouthparts of Thendelecrotona natalica gen. &amp; sp. n. 2 — right maxilla, 3 — labium, 4 — right mandible, 5 — labrum, 6 — mentum. Scale bars 0.1 mm." httpUri="https://zenodo.org/record/177570/files/figure.png" pageId="5" pageNumber="36">Fig. 2</figureCitation>
), the apical lobe of the galea with two parallel to each other, longitudinal rows of long setae (
<figureCitation id="E7AA2A0D600A9C2543DEFE887861EF21" box="[973,1045,312,338]" captionStart="FIGURES 2 – 6" captionStartId="2.[151,269,1956,1980]" captionTargetBox="[185,1408,204,1925]" captionTargetId="figure@2.[169,1418,194,1933]" captionTargetPageId="2" captionText="FIGURES 2 – 6. Mouthparts of Thendelecrotona natalica gen. &amp; sp. n. 2 — right maxilla, 3 — labium, 4 — right mandible, 5 — labrum, 6 — mentum. Scale bars 0.1 mm." httpUri="https://zenodo.org/record/177570/files/figure.png" pageId="5" pageNumber="36">Fig. 2</figureCitation>
), and by the mandibles with well defined “velvety patch” area, which is composed of combination of anterior row of teeth and basal area of very fine spinules (
<figureCitation id="E7AA2A0D600A9C254164FE387DB4EFD1" box="[375,448,392,418]" captionStart="FIGURES 2 – 6" captionStartId="2.[151,269,1956,1980]" captionTargetBox="[185,1408,204,1925]" captionTargetId="figure@2.[169,1418,194,1933]" captionTargetPageId="2" captionText="FIGURES 2 – 6. Mouthparts of Thendelecrotona natalica gen. &amp; sp. n. 2 — right maxilla, 3 — labium, 4 — right mandible, 5 — labrum, 6 — mentum. Scale bars 0.1 mm." httpUri="https://zenodo.org/record/177570/files/figure.png" pageId="5" pageNumber="36">Fig. 4</figureCitation>
).
</paragraph>
<paragraph id="7F2E3688600A9C2540D4FE007F14EC31" blockId="5.[151,1437,151,978]" pageId="5" pageNumber="36">
The new genus most closely resembles species of the genus 
<taxonomicName id="B8914D0B600A9C25438CFE007975EFB9" authority="Cameron, 1920" authorityName="Cameron" authorityYear="1920" box="[927,1281,432,458]" class="Insecta" family="Staphylinidae" genus="Mimacrotona" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="36" phylum="Arthropoda" rank="genus">
<emphasis id="4DE5EA9A600A9C25438CFE00784AEFBA" box="[927,1086,432,457]" italics="true" pageId="5" pageNumber="36">Mimacrotona</emphasis>
<bibRefCitation id="1B004B79600A9C254454FE007975EFB9" author="Cameron" box="[1095,1281,432,458]" pageId="5" pageNumber="37" refString="Cameron, M. (1920) New species of Staphylinidae from Singapore. Part III. Transactions of the Entomological Society of London, 1920, 212 - 284." type="journal article" year="1920">
<collectingCountry id="07867618600A9C254454FE0078C2EFB9" box="[1095,1206,432,458]" name="Cameroon" pageId="5" pageNumber="36">Cameron</collectingCountry>
, 1920
</bibRefCitation>
</taxonomicName>
, but, besides the characters mentioned above, is distinguished by the tergites III and IV transversely impressed at base (only tergite III is impressed in 
<taxonomicName id="B8914D0B600A9C254216FDB07ED7EC6A" box="[517,675,512,537]" class="Insecta" family="Staphylinidae" genus="Mimacrotona" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="36" phylum="Arthropoda" rank="genus">
<emphasis id="4DE5EA9A600A9C254216FDB07ED7EC6A" box="[517,675,512,537]" italics="true" pageId="5" pageNumber="36">Mimacrotona</emphasis>
</taxonomicName>
), the entire ligula (divided to the middle in 
<taxonomicName id="B8914D0B600A9C25448CFDB07949EC6A" box="[1183,1341,512,537]" class="Insecta" family="Staphylinidae" genus="Mimacrotona" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="36" phylum="Arthropoda" rank="genus">
<emphasis id="4DE5EA9A600A9C25448CFDB07949EC6A" box="[1183,1341,512,537]" italics="true" pageId="5" pageNumber="36">Mimacrotona</emphasis>
</taxonomicName>
) and by the medial setae of prementum situated one behind the other.
</paragraph>
<paragraph id="7F2E3688600A9C2540D4FDE079F0EDF1" blockId="5.[151,1437,151,978]" pageId="5" pageNumber="36">
Among other genera with transverse pronotum and entire ligula, the new genus most closely resembles 
<taxonomicName id="B8914D0B600A9C254084FDC87C9BECE2" box="[151,239,632,657]" class="Insecta" family="Staphylinidae" genus="Peliusa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="36" phylum="Arthropoda" rank="genus">
<emphasis id="4DE5EA9A600A9C254084FDC87C9BECE2" box="[151,239,632,657]" italics="true" pageId="5" pageNumber="36">Peliusa</emphasis>
</taxonomicName>
, 
<taxonomicName id="B8914D0B600A9C2540EEFDC87DFBECE2" box="[253,399,632,657]" class="Insecta" family="Staphylinidae" genus="Tyloplatyola" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="36" phylum="Arthropoda" rank="genus">
<emphasis id="4DE5EA9A600A9C2540EEFDC87DFBECE2" box="[253,399,632,657]" italics="true" pageId="5" pageNumber="36">Tyloplatyola</emphasis>
</taxonomicName>
and 
<taxonomicName id="B8914D0B600A9C2541DAFDC87E21ECE2" box="[457,597,632,657]" class="Insecta" family="Staphylinidae" genus="Oxypodinus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="36" phylum="Arthropoda" rank="genus">
<emphasis id="4DE5EA9A600A9C2541DAFDC87E21ECE2" box="[457,597,632,657]" italics="true" pageId="5" pageNumber="36">Oxypodinus</emphasis>
</taxonomicName>
, but is distinguished from them by the medial setae of prementum situated one behind the other. Additionally, 
<taxonomicName id="B8914D0B600A9C25427AFD107E8FECCA" box="[617,763,672,697]" class="Insecta" family="Staphylinidae" genus="Tyloplatyola" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="36" phylum="Arthropoda" rank="genus">
<emphasis id="4DE5EA9A600A9C25427AFD107E8FECCA" box="[617,763,672,697]" italics="true" pageId="5" pageNumber="36">Tyloplatyola</emphasis>
</taxonomicName>
differs from 
<taxonomicName id="B8914D0B600A9C254387FD10782DECCA" box="[916,1113,672,697]" class="Insecta" family="Staphylinidae" genus="Thendelecrotona" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="36" phylum="Arthropoda" rank="genus">
<emphasis id="4DE5EA9A600A9C254387FD10782DECCA" box="[916,1113,672,697]" italics="true" pageId="5" pageNumber="36">Thendelecrotona</emphasis>
</taxonomicName>
by the mentum with lateroanterior angles not produced, the article 2 of labial palpi not widened apically and the very elongate article 1 of labial palpi. 
<taxonomicName id="B8914D0B600A9C25415BFD407DA0ED7A" box="[328,468,752,777]" class="Insecta" family="Staphylinidae" genus="Oxypodinus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="36" phylum="Arthropoda" rank="genus">
<emphasis id="4DE5EA9A600A9C25415BFD407DA0ED7A" box="[328,468,752,777]" italics="true" pageId="5" pageNumber="36">Oxypodinus</emphasis>
</taxonomicName>
is distinguished by the very long and not widened apically article 2 of labial palpi, the quadrate mentum and by the parallel-sided ligula. The members of the genus 
<taxonomicName id="B8914D0B600A9C254445FCA878DAED42" box="[1110,1198,792,817]" class="Insecta" family="Staphylinidae" genus="Peliusa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="36" phylum="Arthropoda" rank="genus">
<emphasis id="4DE5EA9A600A9C254445FCA878DAED42" box="[1110,1198,792,817]" italics="true" pageId="5" pageNumber="36">Peliusa</emphasis>
</taxonomicName>
differ from the new genus in the build of the labium (short first article of the labial palpi, weakly dilated apically article 2, wide median pseudopore field), the less transverse pronotum and by the elytral postero-lateral angles not sinuate.
</paragraph>
</subSubSection>
<subSubSection id="378B6503600A9C2540D4FC3F7E7FEDA1" pageId="5" pageNumber="36" type="etymology">
<paragraph id="7F2E3688600A9C2540D4FC3F7E7FEDA1" blockId="5.[151,1437,151,978]" pageId="5" pageNumber="36">
<emphasis id="4DE5EA9A600A9C2540D4FC3F7D20EDDA" bold="true" box="[199,340,911,937]" pageId="5" pageNumber="36">Etymology.</emphasis>
The name (gender: feminine) is composed of the word 
<emphasis id="4DE5EA9A600A9C2543F8FC207822EDDA" box="[1003,1110,912,937]" italics="true" pageId="5" pageNumber="36">Thendele</emphasis>
(refers to the 
<typeStatus id="A02A882A600A9C2544ECFC207946EDD9" box="[1279,1330,912,938]" pageId="5" pageNumber="36">type</typeStatus>
locality) and the generic name 
<taxonomicName id="B8914D0B600A9C25418BFC087E70EDA2" box="[408,516,952,977]" class="Insecta" family="Staphylinidae" genus="Acrotona" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="36" phylum="Arthropoda" rank="genus">
<emphasis id="4DE5EA9A600A9C25418BFC087E70EDA2" box="[408,516,952,977]" italics="true" pageId="5" pageNumber="36">Acrotona</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
</document>