202 lines
20 KiB
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202 lines
20 KiB
XML
<document id="4AE5A39E6010BBEE5D78D143529F0F07" ID-DOI="10.11646/zootaxa.3999.2.3" ID-GBIF-Dataset="8f47b303-98a7-43ce-b5bb-d1b4bf02555b" ID-ISSN="1175-5326" ID-Zenodo-Dep="237045" ID-ZooBank="E4B0B754-1C94-4B61-B612-848804B059A8" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1461280765185" checkinUser="plazi" docAuthor="Mcmahan, Caleb D., Matamoros, Wilfredo A., Piller, Kyle R. & Chakrabarty, Prosanta" docDate="2015" docId="36162773FFADFFADAB8EFC9EFAB102C8" docLanguage="en" docName="zt03999p234.pdf" docOrigin="Zootaxa 3999 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Oscura Mcmahan, Matamoros, Piller & Chakrabarty, 2015, gen. nov." docType="treatment" docUuid="85579A02-0524-4578-AC87-9D0C2CE1F3EC" docUuidSource="ZooBank" docVersion="9" lastPageNumber="226" masterDocId="CA2F5F0BFFA3FFA2AB19FF8EFF850318" masterDocTitle="Taxonomy and systematics of the herichthyins (Cichlidae: Tribe Heroini), with the description of eight new Middle American Genera" masterLastPageNumber="234" masterPageNumber="211" pageNumber="225" updateTime="1698613983299" updateUser="plazi">
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<mods:titleInfo id="3C0D807EE4BD449EC4BE22A39A9CDC3E">
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<mods:title id="F42D329B112453940A2F7DDD8A63CE3A">Taxonomy and systematics of the herichthyins (Cichlidae: Tribe Heroini), with the description of eight new Middle American Genera</mods:title>
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<mods:name id="93F719BEE53850EB9419372B1FEFD2D8" type="personal">
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<mods:roleTerm id="F14BF241A97D20E12D839F1C57AF734E">Author</mods:roleTerm>
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<mods:namePart id="96370705444554391C7B78867302D3F8">Mcmahan, Caleb D.</mods:namePart>
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<mods:roleTerm id="33CCB938C3E16726FBD9BF7F5174BE8C">Author</mods:roleTerm>
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<mods:namePart id="E548C5395B92640094FA491448CB2132">Matamoros, Wilfredo A.</mods:namePart>
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<mods:name id="F1C041D3FBFA2FC16702D7264C30D609" type="personal">
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<mods:roleTerm id="C693C6F413AC01307C5E7293C478B729">Author</mods:roleTerm>
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<mods:namePart id="81CF93E627C89D444AA503F57B591BC8">Piller, Kyle R.</mods:namePart>
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</mods:name>
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<mods:name id="16153CB5C679E1611542189644FF534D" type="personal">
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<mods:roleTerm id="17836D45EAA544E065857A215B329441">Author</mods:roleTerm>
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<mods:namePart id="E5409DA530D08EDD5C15EED5C94141AE">Chakrabarty, Prosanta</mods:namePart>
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<mods:typeOfResource id="9ECC4AFDDD5FA0F04E1334C202B0D01C">text</mods:typeOfResource>
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<mods:title id="70BF0B635D78C2533ED77F206970A043">Zootaxa</mods:title>
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<mods:date id="77EECC980DFA6638A100150E65E53456">2015</mods:date>
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<mods:detail id="C35F75F0E32BF9D23A10C77CD8EA6940" type="volume">
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<mods:number id="18C7D7C0002C43103BAAC181E9805C9E">3999</mods:number>
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</mods:detail>
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<mods:detail id="8DE4522C79FBEC98F334144FFEBA01F6" type="issue">
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<mods:number id="E3E94FDCA5F3508E9E56E685D7CED63C">2</mods:number>
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<mods:classification id="81D2F5CC3646BB84398DA204FADF4491">journal article</mods:classification>
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<mods:identifier id="E6F90CAB1F14B9D188A72D10C5C3A245" type="DOI">10.11646/zootaxa.3999.2.3</mods:identifier>
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<treatment id="36162773FFADFFADAB8EFC9EFAB102C8" ID-DOI="http://doi.org/10.5281/zenodo.5672264" ID-GBIF-Taxon="119640620" ID-Zenodo-Dep="5672264" LSID="urn:lsid:zoobank.org:act:85579A02-0524-4578-AC87-9D0C2CE1F3EC" httpUri="http://treatment.plazi.org/id/36162773FFADFFADAB8EFC9EFAB102C8" lastPageId="15" lastPageNumber="226" pageId="14" pageNumber="225">
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<subSubSection id="F6A5C5EEFFADFFACAB8EFC9EFD73008A" pageId="14" pageNumber="225" type="nomenclature">
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<paragraph id="BE009665FFADFFACAB8EFC9EFCF40032" blockId="14.[151,881,784,844]" box="[151,881,784,810]" pageId="14" pageNumber="225">
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<heading id="E5482109FFADFFACAB8EFC9EFCF40032" bold="true" box="[151,881,784,810]" fontSize="11" level="1" pageId="14" pageNumber="225" reason="1">
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<emphasis id="8CCB4A77FFADFFACAB8EFC9EFCF40032" bold="true" box="[151,881,784,810]" pageId="14" pageNumber="225">
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Genus
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<taxonomicName id="79BFEDE6FFADFFACABF7FC9EFEC30032" ID-CoL="63GNB" ID-ENA="2302648" box="[238,326,784,810]" class="Actinopterygii" family="Cichlidae" genus="Oscura" kingdom="Animalia" order="Perciformes" pageId="14" pageNumber="225" phylum="Chordata" rank="genus" status="gen. nov.">
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<emphasis id="8CCB4A77FFADFFACABF7FC9EFEC30032" bold="true" box="[238,326,784,810]" italics="true" pageId="14" pageNumber="225">Oscura</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="97F8F70CFFADFFACAA54FC9EFE390032" box="[333,444,784,810]" pageId="14" pageNumber="225" rank="genus">gen. nov.</taxonomicNameLabel>
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,
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<bibRefCitation id="DA2EEB94FFADFFACAAD3FC9EFCF40032" author="McMahan" box="[458,881,784,810]" pageId="14" pageNumber="225" refString="Matamoros, W. A., McMahan, C. D., Chakrabarty, P., Albert, J. S. & Schaefer, J. F. (2015) Derivation of the freshwater fish fauna of Central America revisited: Myers's hypothesis in the twenty-first century. Cladistics, 31 (2), 177 - 188. http: // dx. doi. org / 10.1111 / cla. 12081" type="journal article" year="2015">McMahan and Chakrabarty 2015</bibRefCitation>
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</emphasis>
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</heading>
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</paragraph>
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<paragraph id="BE009665FFADFFACAB8EFCBCFF780054" blockId="14.[151,881,784,844]" box="[151,253,818,844]" pageId="14" pageNumber="225">
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(
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<figureCitation id="26848AE0FFADFFACAB86FCBCFF710054" box="[159,244,818,844]" captionStart="FIGURE 13" captionStartId="14.[151,250,1863,1885]" captionTargetBox="[169,1411,1189,1814]" captionTargetId="figure@14.[151,1436,1164,1841]" captionTargetPageId="14" captionText="FIGURE 13. Oscura heterospila, FMNH 108978, 98.79 mm SL." httpUri="https://zenodo.org/record/237058/files/figure.png" pageId="14" pageNumber="225">Fig. 13</figureCitation>
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)
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</paragraph>
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<paragraph id="BE009665FFADFFACAB8EFCF7FD73008A" blockId="14.[151,1437,889,1130]" box="[151,758,889,914]" pageId="14" pageNumber="225">
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<emphasis id="8CCB4A77FFADFFACAB8EFCF7FEE6008A" bold="true" box="[151,355,889,914]" pageId="14" pageNumber="225">Inclusive species.</emphasis>
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<taxonomicName id="79BFEDE6FFADFFACAA73FCF4FD8C008A" box="[362,521,889,914]" class="Actinopterygii" family="Cichlidae" genus="Oscura" kingdom="Animalia" order="Perciformes" pageId="14" pageNumber="225" phylum="Chordata" rank="species" species="heterospila">
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<emphasis id="8CCB4A77FFADFFACAA73FCF4FEFB0089" box="[362,382,890,913]" italics="true" pageId="14" pageNumber="225">
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<collectingCountry id="C6A8D6F5FFADFFACAA73FCF4FEFB0089" box="[362,382,890,913]" name="Iceland" pageId="14" pageNumber="225">O</collectingCountry>
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</emphasis>
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.
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<emphasis id="8CCB4A77FFADFFACAA95FCF7FD8C008A" box="[396,521,889,914]" italics="true" pageId="14" pageNumber="225">heterospila</emphasis>
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</taxonomicName>
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(
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<typeStatus id="610428C7FFADFFACA90EFCF4FDCC008A" box="[535,585,890,914]" pageId="14" pageNumber="225">type</typeStatus>
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by monotypy).
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</paragraph>
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</subSubSection>
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<subSubSection id="F6A5C5EEFFADFFACABDEFC13FEE60771" pageId="14" pageNumber="225" type="diagnosis">
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<paragraph id="BE009665FFADFFACABDEFC13FEE60771" blockId="14.[151,1437,889,1130]" pageId="14" pageNumber="225">
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<emphasis id="8CCB4A77FFADFFACABDEFC13FEC400AE" bold="true" box="[199,321,925,950]" pageId="14" pageNumber="225">Diagnosis.</emphasis>
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<taxonomicName id="79BFEDE6FFADFFACAA55FC10FE1A00AD" box="[332,415,926,949]" class="Actinopterygii" family="Cichlidae" genus="Oscura" kingdom="Animalia" order="Perciformes" pageId="14" pageNumber="232" phylum="Chordata" rank="genus" status="gen. nov.">
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<emphasis id="8CCB4A77FFADFFACAA55FC10FE1A00AD" box="[332,415,926,949]" italics="true" pageId="14" pageNumber="225">Oscura</emphasis>
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</taxonomicName>
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is diagnosed by a large, round or oblong, dark blotch filling the entirety (or nearly the entirety) of the caudal peduncle and the presence of small black spots on scales covering the sides of the body. Species of this genus possess an overall dark body coloration. This genus is most similar to
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<taxonomicName id="79BFEDE6FFADFFACAF8CFC68FB4900E5" box="[1173,1228,998,1021]" family="Cichlidae" genus="Vieja" higherTaxonomySource="CoL" kingdom="Animalia" order="Perciformes" pageId="14" pageNumber="225" phylum="Chordata" rank="genus">
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<emphasis id="8CCB4A77FFADFFACAF8CFC68FB4900E5" box="[1173,1228,998,1021]" italics="true" pageId="14" pageNumber="225">Vieja</emphasis>
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</taxonomicName>
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; however,
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<taxonomicName id="79BFEDE6FFADFFACAE50FC68FA1E00E5" box="[1353,1435,998,1021]" class="Actinopterygii" family="Cichlidae" genus="Oscura" kingdom="Animalia" order="Perciformes" pageId="14" pageNumber="232" phylum="Chordata" rank="genus" status="gen. nov.">
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<emphasis id="8CCB4A77FFADFFACAE50FC68FA1E00E5" box="[1353,1435,998,1021]" italics="true" pageId="14" pageNumber="225">Oscura</emphasis>
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</taxonomicName>
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possesses dark bars down the body (typically five) and a blotch that fills the entirety of the caudal peduncle, versus the absence of broad bars down the body and a caudal blotch not filling the entirety of the caudal peduncle in members of
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<taxonomicName id="79BFEDE6FFADFFACAA3AFBDDFEDF0772" box="[291,346,1107,1130]" family="Cichlidae" genus="Vieja" higherTaxonomySource="CoL" kingdom="Animalia" order="Perciformes" pageId="14" pageNumber="225" phylum="Chordata" rank="genus">
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<emphasis id="8CCB4A77FFADFFACAA3AFBDDFEDF0772" box="[291,346,1107,1130]" italics="true" pageId="14" pageNumber="225">Vieja</emphasis>
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</taxonomicName>
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.
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</paragraph>
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</subSubSection>
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<caption id="EAC0C6EDFFADFFACAB8EF8C9FCA20445" box="[151,807,1863,1885]" httpUri="https://zenodo.org/record/237058/files/figure.png" pageId="14" pageNumber="225" targetBox="[169,1411,1189,1814]" targetPageId="14">
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<paragraph id="BE009665FFADFFACAB8EF8C9FCA20445" blockId="14.[151,807,1863,1885]" box="[151,807,1863,1885]" pageId="14" pageNumber="225">
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<emphasis id="8CCB4A77FFADFFACAB8EF8C9FE9C0445" bold="true" box="[151,281,1863,1885]" pageId="14" pageNumber="225">FIGURE 13</emphasis>
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.
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<taxonomicName id="79BFEDE6FFADFFACAA3CF8C6FE620445" box="[293,487,1863,1885]" class="Actinopterygii" family="Cichlidae" genus="Oscura" kingdom="Animalia" order="Perciformes" pageId="14" pageNumber="225" phylum="Chordata" rank="species" species="heterospila">
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<emphasis id="8CCB4A77FFADFFACAA3CF8C6FE620445" box="[293,487,1863,1885]" italics="true" pageId="14" pageNumber="225">Oscura heterospila</emphasis>
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</taxonomicName>
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, FMNH 108978, 98.79mm SL.
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</paragraph>
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</caption>
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<subSubSection id="F6A5C5EEFFADFFACABDEF80AFC9C04FE" pageId="14" pageNumber="225" type="distribution">
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<paragraph id="BE009665FFADFFACABDEF80AFC9C04FE" blockId="14.[151,1436,1924,2022]" pageId="14" pageNumber="225">
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<emphasis id="8CCB4A77FFADFFACABDEF80AFEE50485" bold="true" box="[199,352,1924,1949]" pageId="14" pageNumber="225">Distribution.</emphasis>
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Atlantic slope of
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<collectingCountry id="C6A8D6F5FFADFFACA930F80AFDFB0485" box="[553,638,1924,1949]" name="Mexico" pageId="14" pageNumber="225">Mexico</collectingCountry>
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and
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<collectingCountry id="C6A8D6F5FFADFFACA9ACF80AFCAB0485" box="[693,814,1924,1949]" name="Guatemala" pageId="14" pageNumber="225">Guatemala</collectingCountry>
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in the Río Usumacinta drainage.
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<emphasis id="8CCB4A77FFADFFACABDEF826FECB04D9" bold="true" box="[199,334,1960,1985]" pageId="14" pageNumber="225">Etymology.</emphasis>
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Gender feminine. “
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<taxonomicName id="79BFEDE6FFADFFACA926F827FD1604D9" box="[575,659,1961,1985]" class="Actinopterygii" family="Cichlidae" genus="Oscura" kingdom="Animalia" order="Perciformes" pageId="14" pageNumber="232" phylum="Chordata" rank="genus" status="gen. nov.">Oscura</taxonomicName>
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” is the Spanish word for dark, in reference to the overall dark coloration that is characteristic of members of this genus.
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</paragraph>
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</subSubSection>
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<subSubSection id="F6A5C5EEFFACFFADABDEFF19FEF4027D" pageId="15" pageNumber="226" type="discussion">
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<paragraph id="BE009665FFACFFADABDEFF19FEF4027D" blockId="15.[151,1436,151,464]" pageId="15" pageNumber="226">
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<emphasis id="8CCB4A77FFACFFADABDEFF19FECA03A8" bold="true" box="[199,335,151,176]" pageId="15" pageNumber="226">Comments.</emphasis>
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The body shape and large, dark caudal blotch of this species initially allied this species in
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<emphasis id="8CCB4A77FFACFFADAE45FF17FA1F03A8" box="[1372,1434,153,176]" italics="true" pageId="15" pageNumber="226">
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<taxonomicName id="79BFEDE6FFACFFADAE45FF17FA1603A8" box="[1372,1427,153,176]" family="Cichlidae" genus="Vieja" higherTaxonomySource="CoL" kingdom="Animalia" order="Perciformes" pageId="15" pageNumber="226" phylum="Chordata" rank="genus">Vieja</taxonomicName>
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,
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</emphasis>
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but all phylogenetic studies to date have recovered them to be distantly related. The increased taxon sampling of the present study, as well as the analysis of
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<bibRefCitation id="DA2EEB94FFACFFADA964FF51FCCB03E0" box="[637,846,223,248]" pageId="15" pageNumber="226" refString="Rican, O., Pialek, L., Zardoya, R., Doadrio, I. & Zrzav, Y. (2013) Biogeography of the Mesoamerican Cichlidae (Teleostei: Heroini): colonization through the GAARlandia land bridge and early diversification. Journal of Biogeography, 40 (3), 579 - 593. http: // dx. doi. org / 10.1111 / jbi. 12023" type="journal article">
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Říčan
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<emphasis id="8CCB4A77FFACFFADA9DDFF6FFD7303E0" box="[708,758,223,248]" italics="true" pageId="15" pageNumber="226">et al</emphasis>
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. (2013)
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</bibRefCitation>
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recovered this species in its own clade and the sister group to
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<taxonomicName id="79BFEDE6FFACFFADAA1FFE8AFEF80205" box="[262,381,260,285]" class="Actinopterygii" family="Cichlidae" genus="Rheoheros" kingdom="Animalia" order="Perciformes" pageId="15" pageNumber="232" phylum="Chordata" rank="genus" status="gen. nov.">
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<emphasis id="8CCB4A77FFACFFADAA1FFE8AFEF80205" box="[262,381,260,285]" italics="true" pageId="15" pageNumber="226">Rheoheros</emphasis>
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</taxonomicName>
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. Given the recovered phylogenetic position and the morphological distinctiveness of this species compared to its closest relatives (
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<taxonomicName id="79BFEDE6FFACFFADA96BFEA7FC9F0258" box="[626,794,295,320]" class="Actinopterygii" family="Cichlidae" genus="Rheoheros" kingdom="Animalia" order="Perciformes" pageId="15" pageNumber="226" phylum="Chordata" rank="species" species="lentiginosus">
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<emphasis id="8CCB4A77FFACFFADA96BFEA7FD060258" box="[626,643,297,320]" italics="true" pageId="15" pageNumber="226">R</emphasis>
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.
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<emphasis id="8CCB4A77FFACFFADA98AFEA9FC9F0258" box="[659,794,295,320]" italics="true" pageId="15" pageNumber="226">lentiginosus</emphasis>
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</taxonomicName>
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and
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<taxonomicName id="79BFEDE6FFACFFADA84CFEA7FC610258" box="[853,996,295,320]" class="Actinopterygii" family="Cichlidae" genus="Rheoheros" kingdom="Animalia" order="Perciformes" pageId="15" pageNumber="226" phylum="Chordata" rank="species" species="coeruleus">
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<emphasis id="8CCB4A77FFACFFADA84CFEA7FCE30258" box="[853,870,297,320]" italics="true" pageId="15" pageNumber="226">R</emphasis>
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.
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<emphasis id="8CCB4A77FFACFFADA86EFEA9FC610258" box="[887,996,295,320]" italics="true" pageId="15" pageNumber="226">coeruleus</emphasis>
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</taxonomicName>
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), we recognize this species in its own (monotypic) genus.
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</paragraph>
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</subSubSection>
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<subSubSection id="F6A5C5EEFFACFFADABDEFEE1FAB102C8" pageId="15" pageNumber="226" type="materials_examined">
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<paragraph id="BE009665FFACFFADABDEFEE1FAB102C8" blockId="15.[151,1436,151,464]" pageId="15" pageNumber="226">
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<emphasis id="8CCB4A77FFACFFADABDEFEE1FE340290" bold="true" box="[199,433,367,392]" pageId="15" pageNumber="226">Material examined.</emphasis>
|
|
<collectionCode id="D8AE0EA0FFACFFADAAA1FEFEFD920290" box="[440,535,368,392]" httpUri="http://grbio.org/cool/ew6z-dwdm" name="Louisiana State University, Musuem of Zoology" pageId="15" pageNumber="226">LSUMZ</collectionCode>
|
|
16229 [n=5,
|
|
<collectingCountry id="C6A8D6F5FFACFFADA9A8FEE1FCAA0290" box="[689,815,367,392]" name="Guatemala" pageId="15" pageNumber="226">Guatemala</collectingCountry>
|
|
: Río La Pasion],
|
|
<collectionCode id="D8AE0EA0FFACFFADA8EBFEFEFBD40290" box="[1010,1105,368,392]" httpUri="http://grbio.org/cool/ew6z-dwdm" name="Louisiana State University, Musuem of Zoology" pageId="15" pageNumber="226">LSUMZ</collectionCode>
|
|
16471 [n=1,
|
|
<collectingCountry id="C6A8D6F5FFACFFADAFF3FEE1FAED0290" box="[1258,1384,367,392]" name="Guatemala" pageId="15" pageNumber="226">Guatemala</collectingCountry>
|
|
: Río La Pasion],
|
|
<collectionCode id="D8AE0EA0FFACFFADAA39FE1BFEF202B4" box="[288,375,405,428]" httpUri="http://grbio.org/cool/ra64-3eva" name="University of Michigan, Museum of Zoology" pageId="15" pageNumber="226">UMMZ</collectionCode>
|
|
144313 [n=5,
|
|
<collectingCountry id="C6A8D6F5FFACFFADA906FE1AFD1802B5" box="[543,669,404,429]" name="Guatemala" pageId="15" pageNumber="226">Guatemala</collectingCountry>
|
|
: Río
|
|
<collectingCountry id="C6A8D6F5FFACFFADA9C0FE1AFCD702B5" box="[729,850,404,429]" name="Saint Pierre and Miquelon" pageId="15" pageNumber="226">San Pedro</collectingCountry>
|
|
],
|
|
<collectionCode id="D8AE0EA0FFACFFADA87CFE1BFC3302B4" LSID="urn:lsid:biocol.org:col:34795" box="[869,950,405,428]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34795" name="Field Museum of Natural History" pageId="15" pageNumber="226">FMNH</collectionCode>
|
|
108978 [n=1,
|
|
<collectingCountry id="C6A8D6F5FFACFFADAF46FE1AFB5802B5" box="[1119,1245,404,429]" name="Guatemala" pageId="15" pageNumber="226">Guatemala</collectingCountry>
|
|
: Río
|
|
<collectingCountry id="C6A8D6F5FFACFFADAE01FE1AFA1702B5" box="[1304,1426,404,429]" name="Saint Pierre and Miquelon" pageId="15" pageNumber="226">San Pedro</collectingCountry>
|
|
],
|
|
<collectionCode id="D8AE0EA0FFACFFADAB8EFE36FF6D02D7" LSID="urn:lsid:biocol.org:col:34795" box="[151,232,440,463]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34795" name="Field Museum of Natural History" pageId="15" pageNumber="226">FMNH</collectionCode>
|
|
108993 [n=18,
|
|
<collectingCountry id="C6A8D6F5FFACFFADAA82FE39FD9C02C8" box="[411,537,439,464]" name="Guatemala" pageId="15" pageNumber="226">Guatemala</collectingCountry>
|
|
: Río Usumacinta],
|
|
<collectionCode id="D8AE0EA0FFACFFADA9E9FE36FCC402D7" LSID="urn:lsid:biocol.org:col:34795" box="[752,833,440,463]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34795" name="Field Museum of Natural History" pageId="15" pageNumber="226">FMNH</collectionCode>
|
|
108990 [n=3,
|
|
<collectingCountry id="C6A8D6F5FFACFFADA8FFFE39FBE102C8" box="[998,1124,439,464]" name="Guatemala" pageId="15" pageNumber="226">Guatemala</collectingCountry>
|
|
: Río Usumacinta].
|
|
</paragraph>
|
|
</subSubSection>
|
|
</treatment>
|
|
</document> |