treatments-xml/data/21/9A/93/219A9352B86056889DBEB0947CC84DEA.xml

<document ID-DOI="http://dx.doi.org/10.3897/zookeys.1048.67564" ID-GBIF-Dataset="57d9ade8-67e7-4d65-9581-b85a80e88893" ID-PMC="PMC8292283" ID-Pensoft-Pub="1313-2970-1048-145" ID-Pensoft-UUID="93FE96A4127A5D8FAA3A39BCB3E4B729" ID-PubMed="34316267" ID-ZooBank="578795FD41BF4807B0016F7B5AC4D4B9" ModsDocID="1313-2970-1048-145" checkinTime="1626283267302" checkinUser="pensoft" docAuthor="Lantsov, Vladimir I. &amp; Pilipenko, Valentin E." docDate="2021" docId="219A9352B86056889DBEB0947CC84DEA" docLanguage="en" docName="ZooKeys 1048: 145-175" docOrigin="ZooKeys 1048" docPubDate="2021-07-13" docSource="http://dx.doi.org/10.3897/zookeys.1048.67564" docTitle="Tipula (Lunatipula) caucasica Riedel 1920" docType="treatment" docVersion="4" id="93FE96A4127A5D8FAA3A39BCB3E4B729" lastPageNumber="145" masterDocId="93FE96A4127A5D8FAA3A39BCB3E4B729" masterDocTitle="A new Palaearctic species of the subgenus Lunatipula (Diptera, Tipulidae) from the West Caucasus with a survey of the caucasica species group" masterLastPageNumber="175" masterPageNumber="145" pageNumber="145" updateTime="1668150580932" updateUser="ExternalLinkService">
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<mods:titleInfo>
<mods:title>A new Palaearctic species of the subgenus Lunatipula (Diptera, Tipulidae) from the West Caucasus with a survey of the caucasica species group</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Lantsov, Vladimir I.</mods:namePart>
<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0002-8275-496X</mods:nameIdentifier>
<mods:affiliation>Tembotov Institute of Ecology of Mountain Territories of Russian Academy of Sciences, I. Armand str., 37 a, Nalchik 360051, Russia</mods:affiliation>
<mods:nameIdentifier type="email">lantsov@megalog.ru</mods:nameIdentifier>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Pilipenko, Valentin E.</mods:namePart>
<mods:affiliation>Lomonosov Moscow State University, GSP- 1, Leninskie Gory, Moscow 119991, Russia</mods:affiliation>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
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<mods:title>ZooKeys</mods:title>
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<mods:date>2021</mods:date>
<mods:detail type="pubDate">
<mods:number>2021-07-13</mods:number>
</mods:detail>
<mods:detail type="volume">
<mods:number>1048</mods:number>
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<mods:start>145</mods:start>
<mods:end>175</mods:end>
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<mods:url>http://dx.doi.org/10.3897/zookeys.1048.67564</mods:url>
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<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.1048.67564</mods:identifier>
<mods:identifier type="Pensoft-Pub">1313-2970-1048-145</mods:identifier>
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<treatment ID-GBIF-Taxon="183543016" LSID="urn:lsid:plazi:treatment:219A9352B86056889DBEB0947CC84DEA" httpUri="http://treatment.plazi.org/id/219A9352B86056889DBEB0947CC84DEA" lastPageNumber="145" pageId="0" pageNumber="145">
<subSubSection pageId="0" pageNumber="145" type="nomenclature">
<paragraph pageId="0" pageNumber="145">
<taxonomicName LSID="219A9352-B860-5688-9DBE-B0947CC84DEA" authority="Riedel, 1920" authorityName="Riedel" authorityYear="1920" class="Insecta" family="Tipulidae" genus="Tipula" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tipula (Lunatipula) caucasica" order="Diptera" pageId="0" pageNumber="145" phylum="Arthropoda" rank="species" species="caucasica" subGenus="Lunatipula">Tipula (Lunatipula) caucasica Riedel, 1920</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="145" type="description">
<paragraph pageId="0" pageNumber="145">
<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. T. (Lunatipula) caucasica male habitus A general lateral view B head and thorax, dorsal view. Scale bars: 10 mm (A), 1 mm (B)." figureDoi="10.3897/zookeys.1048.67564.figure1" httpUri="https://binary.pensoft.net/fig/566410" pageId="0" pageNumber="145">Figs 1</figureCitation>
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. T. (Lunatipula) caucasica A terminal segments of male abdomen, lateral view (dry) B-L male terminalia (after KOH 10 % treatment) B hypopygium, caudal view C hypopygium, lateral view D sternite 8, ventral view E tergite 9, caudal view F tergite 9, dorsal view G inner and outer gonostylus, lateral view H inner and outer gonostylus, caudal view I gonocoxal fragment, dorsal view J semen pump and aedeagus, lateral view K adminiculum and paired apical appendages of sternite 9, caudal view L adminiculum and paired apical appendages of sternite 9, lateral view. Abbreviations: adm-adminiculum; as 8 - paired appendages of sternite 8; as 9 - apical appendages of sternite 9; ig-inner gonostylus; gcx-gonocoxite; og-outer gonostylus; s 8 - sternite 8; s 9 - sternite 9; t 9 - tergite 9. Scale bar: 1 mm." figureDoi="10.3897/zookeys.1048.67564.figure2" httpUri="https://binary.pensoft.net/fig/566411" pageId="0" pageNumber="145">, 2</figureCitation>
<figureCitation captionStart="Figure 12" captionStartId="F12" captionText="Figure 12. Female terminalia of Tipula caucasica species group A-C T. (Lunatipula) caucasica D-F T. (Lunatipula) quadridentata G-I T. (Lunatipula) talyshensis A, D, G ovipositor, lateral view B, E, H ovipositor ventral view C, F, I sternite 9 and furca, lateral view. Abbreviations: c-cerci; f-furca; hv-hypogynial valve; s-sternite; t-tergite. Scale bar: 1 mm." figureDoi="10.3897/zookeys.1048.67564.figure12" httpUri="https://binary.pensoft.net/fig/566421" pageId="0" pageNumber="145">, 12A-C</figureCitation>
<figureCitation captionStart="Figure 13" captionStartId="F13" captionText="Figure 13. Heads of Tipula caucasica species group, dorsal view A T. (Lunatipula) caucasica B T. (Lunatipula) eleniya sp. nov. C T. (Lunatipula) paupera D T. (Lunatipula) quadridentata E T. (Lunatipula) Tipula talyshensis. Scale bar: 1 mm." figureDoi="10.3897/zookeys.1048.67564.figure13" httpUri="https://binary.pensoft.net/fig/566422" pageId="0" pageNumber="145">, 13A</figureCitation>
<figureCitation captionStart="Figure 14" captionStartId="F14" captionText="Figure 14. Comparisons of some components of male terminalia of the Tipula caucasica species group A, F T. (Lunatipula) caucasica B, G T. (Lunatipula) eleniya sp. nov. C, I T. (Lunatipula) paupera D, J T. (Lunatipula) quadridentata E, K T. (Lunatipula) talyshensis A-E tergite 9, dorsal view F-K inner and outer gonostylus, lateral view. Arrows point to central notch at apex of tergite 9 (C, D), to the wedge-shaped projection posteriorly on inner gonostylus (F), to the rod-like outgrowth behind anterior beak of inner gonostylus (G), to the anterior (dorsal) margin of gonocoxite (I, J, K). Scale bar: 1 mm." figureDoi="10.3897/zookeys.1048.67564.figure14" httpUri="https://binary.pensoft.net/fig/566423" pageId="0" pageNumber="145">, 14A,F</figureCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="145" type="reference_group">
<paragraph pageId="0" pageNumber="145">
<taxonomicName authorityName="Riedel" authorityYear="1920" class="Insecta" family="Tipulidae" genus="Tipula" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tipula (Lunatipula) caucasica" order="Diptera" pageId="0" pageNumber="145" phylum="Arthropoda" rank="species" species="caucasica" subGenus="Lunatipula">Tipula (Lunatipula) caucasica</taxonomicName>
<bibRefCitation author="Riedel, MP" journalOrPublisher="Zoologische Jahrbucher, Abteilung fur Systematik, Geographie und Biologie der Tiere" pageId="0" pageNumber="145" pagination="13 - 22" refId="B9" refString="Riedel, MP, 1920. Nematocera Polyneura (Dipt.) aus dem Kaukasus. Zoologische Jahrbucher, Abteilung fur Systematik, Geographie und Biologie der Tiere 43: 13 - 22" title="Nematocera Polyneura (Dipt.) aus dem Kaukasus." volume="43" year="1920">Riedel 1920</bibRefCitation>
: 17 (type locality: Georgia, 
<normalizedToken originalValue="Mts’chet">Mts'chet</normalizedToken>
, prope Tiflis); 
<bibRefCitation author="Savchenko, EN" journalOrPublisher="Zoologicheskiy Zhurnal" pageId="0" pageNumber="145" refId="B12" refString="Savchenko, EN, 1964. Crane-flies (Diptera, Tipulidae), Subfam. Tipulinae, Genus TipulaL., 2. Fauna USSR, N.S. 89, Nasekomye Dvukrylye [Diptera] 2(4): 1-503. [in Russian]" title="Crane-flies (Diptera, Tipulidae), Subfam. Tipulinae, Genus TipulaL., 2. Fauna USSR, N. S. 89, Nasekomye Dvukrylye [Diptera] 2 (4): 1 - 503. [in Russian]" year="1964">Savchenko 1964</bibRefCitation>
: 390; Oosterbroek, Theowald 1992:104; 
<bibRefCitation DOI="https://doi.org/10.47640/978-5-00105-586-0_2020_292" author="Lantsov, VI" journalOrPublisher="Fauna of the Stavropol region" pageId="0" pageNumber="145" refId="B7" refString="Lantsov, VI, 2020. Biological diversity of crane flies (Diptera, Tipuloidea) in the natural landscapes of Dagestan (North-East Caucasus, Russia) // XI All-Russian Dipterological Symposium (with international participation). Voronezh. August 24-29, 2020. Proceedings of conference / Responsible editors: O.G.Ovchinnikova, I.V.Shamshev. St.Petersburg: Russian Entomological Society, LEMA, 292-296. https://doi.org/10.47640/978-5-00105-586-0_2020_292" title="Biological diversity of crane flies (Diptera, Tipuloidea) in the natural landscapes of Dagestan (North-East Caucasus, Russia) // XI All-Russian Dipterological Symposium (with international participation). Voronezh. August 24 - 29, 2020. Proceedings of conference / Responsible editors: O. G. Ovchinnikova, I. V. Shamshev. St. Petersburg: Russian Entomological Society, LEMA, 292 - 296." url="https://doi.org/10.47640/978-5-00105-586-0_2020_292" year="2020">Lantsov 2020</bibRefCitation>
: 294; 
<bibRefCitation author="Oosterbroek, P" journalOrPublisher="Fauna of the Stavropol region" pageId="0" pageNumber="145" publicationUrl="http://ccw.naturalis.nl/detail.php" refId="B8" refString="Oosterbroek, P, 2021. Catalogue of the Craneflies of the World. http://ccw.naturalis.nl/detail.php" title="Catalogue of the Craneflies of the World." url="http://ccw.naturalis.nl/detail.php" year="2021">Oosterbroek 2021</bibRefCitation>
.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="145" type="materials_examined">
<paragraph pageId="0" pageNumber="145">Material examined.</paragraph>
<paragraph pageId="0" pageNumber="145">
<materialsCitation collectingDate="1958-05-23" collectingDateMax="1959-07-17" collectingDateMin="1958-05-23" collectorName="Kurnakov, Savenko" country="Georgia" location="Bani-Khevi" specimenCount="10" specimenCount-female="4" specimenCount-male="6">
<collectingCountry name="Georgia">Georgia</collectingCountry>
• 
<specimenCount type="male">1 male</specimenCount>
, 
<specimenCount type="female">3 females</specimenCount>
; 
<normalizedToken originalValue="“ущ">&quot;ushch</normalizedToken>
. 
<normalizedToken originalValue="Бани-Хеви">
<location LSID="urn:lsid:plazi:treatment:219A9352B86056889DBEB0947CC84DEA:01AD7850430DC2BAF6BC3CC4422828FD" country="Georgia" name="Bani-Khevi">Bani-Khevi</location>
</normalizedToken>
<normalizedToken originalValue="бл">bl</normalizedToken>
. 
<normalizedToken originalValue="Боржоми">Borzhomi</normalizedToken>
, 
<normalizedToken originalValue="Аджаро-Имеретинск">Adzharo-Imeretinsk</normalizedToken>
. 
<normalizedToken originalValue="хр">khr</normalizedToken>
. [Gorge Bani-Khevi near Borjomi, Adjaro-Imeretinsk. Ridge]; 
<collectingDate value="1958-06-26">26 Jun. 1958</collectingDate>
; 
<collectorName>Kurnakov</collectorName>
leg.; ZISP • 
<specimenCount type="male">1 male</specimenCount>
; 
<normalizedToken originalValue="“Цхра-Цхаро">&quot;Tskhra-Tskharo</normalizedToken>
<normalizedToken originalValue="близ">bliz</normalizedToken>
<normalizedToken originalValue="Бакуриани”">Bakuriani&quot;</normalizedToken>
[Tskhra-Tskharo near Bakuriani]; 
<collectingDate value="1958-06-29">29 Jun. 1958</collectingDate>
; 
<collectorName>Kurnakov</collectorName>
leg.; ZISP • 
<specimenCount type="male">1 male</specimenCount>
; 
<normalizedToken originalValue="“Леберде">&quot;Leberde</normalizedToken>
, 
<normalizedToken originalValue="Менгрелия">Mengreliya</normalizedToken>
, 
<normalizedToken originalValue="Грузия”">Gruziya&quot;</normalizedToken>
[Leberde, Mengrelia, 
<collectingCountry name="Georgia">Georgia</collectingCountry>
]; 
<collectingDate value="1959-07-17">17 Jul. 1959</collectingDate>
; 
<collectorName>Savenko</collectorName>
leg.; ZISP • 
<specimenCount type="male">1 male</specimenCount>
; 
<normalizedToken originalValue="“Аджаро-Имеретинск">&quot;Adzharo-Imeretinsk</normalizedToken>
. 
<normalizedToken originalValue="хр">khr</normalizedToken>
. 
<normalizedToken originalValue="Горы">Gory</normalizedToken>
<normalizedToken originalValue="у">u</normalizedToken>
<normalizedToken originalValue="Саирме">Sairme</normalizedToken>
, 
<normalizedToken originalValue="Грузия”">Gruziya&quot;</normalizedToken>
[Adjaro-Imeretinsk. Ridge, mountains near Sairme, 
<collectingCountry name="Georgia">Georgia</collectingCountry>
]; 
<collectingDate value="1958-05-23">23 May 1958</collectingDate>
; Kurnakov; ZISP • 
<specimenCount type="male">2 males</specimenCount>
, 
<specimenCount type="female">1 female</specimenCount>
; 
<normalizedToken originalValue="”ущ">&quot;ushch</normalizedToken>
. 
<normalizedToken originalValue="Цваниат-Хеви">Tsvaniat-Khevi</normalizedToken>
<normalizedToken originalValue="Аджаро-Имеретинск">Adzharo-Imeretinsk</normalizedToken>
. 
<normalizedToken originalValue="хр">khr</normalizedToken>
</materialsCitation>
. 
<materialsCitation collectingDate="1958-05-20" collectingDateMax="1958-06-30" collectingDateMin="1958-05-20" collectorName="Kurnakov" country="Georgia" location="Oak-hornbeam forest" municipality="Gudautsk. District" specimenCount="4" specimenCount-male="4" stateProvince="Abkhazia">
<normalizedToken originalValue="Грузия”">Gruziya&quot;</normalizedToken>
[Tsvaniat-Khevi Gorge, Adjaro-Imeretinsk. Ridge, 
<collectingCountry name="Georgia">Georgia</collectingCountry>
]; 
<collectingDate value="1958-06-23">23 Jun. 1958</collectingDate>
; 
<collectorName>Kurnakov</collectorName>
leg; / 
<normalizedToken originalValue="“Высокотравье">&quot;Vysokotrave</normalizedToken>
<normalizedToken originalValue="у">u</normalizedToken>
<normalizedToken originalValue="верхн">verkhn</normalizedToken>
. 
<normalizedToken originalValue="границы">granitsy</normalizedToken>
<normalizedToken originalValue="леса">lesa</normalizedToken>
<normalizedToken originalValue="у">u</normalizedToken>
<normalizedToken originalValue="ручья”">ruchya&quot;</normalizedToken>
[High grass at the edge of the border of the forest by the stream]; ZISP • 
<specimenCount type="male">2 males</specimenCount>
; 
<normalizedToken originalValue="“с">&quot;s</normalizedToken>
. 
<normalizedToken originalValue="Отхара">Otkhara</normalizedToken>
, 
<normalizedToken originalValue="Гудаутск">Gudautsk</normalizedToken>
. 
<normalizedToken originalValue="р-н">r-n</normalizedToken>
, 
<normalizedToken originalValue="Абхазия">Abkhaziya</normalizedToken>
, 
<normalizedToken originalValue="бер">ber</normalizedToken>
. 
<normalizedToken originalValue="речки">rechki</normalizedToken>
, 
<normalizedToken originalValue="кусты.”">kusty.&quot;</normalizedToken>
[village Otkhara, 
<collectingMunicipality>Gudautsk. District</collectingMunicipality>
, 
<collectingRegion country="Georgia" name="Abkhazia">Abkhazia</collectingRegion>
, river bank, bushes]; 
<collectingDate value="1958-06-30">30 Jun. 1958</collectingDate>
; Kurnakov; ZISP • 
<specimenCount type="male">2 males</specimenCount>
; 
<normalizedToken originalValue="“Абхазия">&quot;Abkhaziya</normalizedToken>
, 
<normalizedToken originalValue="с">s</normalizedToken>
. 
<normalizedToken originalValue="Отхара">Otkhara</normalizedToken>
<normalizedToken originalValue="предгорья">predgorya</normalizedToken>
<normalizedToken originalValue="Бзыбского">Bzybskogo</normalizedToken>
<normalizedToken originalValue="хр.”">khr.&quot;</normalizedToken>
[
<collectingRegion country="Georgia" name="Abkhazia">Abkhazia</collectingRegion>
, village Otkhara foothills of the Bzybsk Ridge]; 
<collectingDate value="1958-05-20">20 May 1958</collectingDate>
; Kurnakov; / 
<normalizedToken originalValue="“Дубово-грабовый">&quot;Dubovo-grabovyi</normalizedToken>
<normalizedToken originalValue="лес">les</normalizedToken>
, 
<normalizedToken originalValue="кусты">kusty</normalizedToken>
Asalia&quot; [
<location LSID="urn:lsid:plazi:treatment:219A9352B86056889DBEB0947CC84DEA:61555C28CE56949C9D2B78972D9F174E" country="Georgia" municipality="Gudautsk. District" name="Oak-hornbeam forest" stateProvince="Abkhazia">Oak-hornbeam forest</location>
, bushes Asalia&quot;]; ZISP
</materialsCitation>
; 
<materialsCitation collectingDate="1964-06-15" collectorName="Pastukhov" country="Azerbaijan" elevation="200" location="Akhkemal Ridge. Azerb." municipality="Belokany" specimenCount="1" specimenCount-male="1">
<collectingCountry name="Azerbaijan">Azerbaijan</collectingCountry>
• 
<specimenCount type="male">1 male</specimenCount>
; 
<normalizedToken originalValue="“Белоканы">&quot;Belokany</normalizedToken>
, 
<normalizedToken originalValue="хр">khr</normalizedToken>
. 
<normalizedToken originalValue="Ахкемал">Akhkemal</normalizedToken>
. 
<normalizedToken originalValue="Азерб">Azerb</normalizedToken>
. 
<quantity metricMagnitude="2" metricUnit="m" metricValue="2.0" unit="m" value="200.0">
<elevation metricMagnitude="2" metricUnit="m" metricValue="2.0" unit="m" value="200.0">
200 
<normalizedToken originalValue="м”">m&quot;</normalizedToken>
</elevation>
</quantity>
[
<collectingMunicipality>Belokany</collectingMunicipality>
, 
<location LSID="urn:lsid:plazi:treatment:219A9352B86056889DBEB0947CC84DEA:F13E52C84FF7BA3AFF9E97401FDB0E61" country="Azerbaijan" municipality="Belokany" name="Akhkemal Ridge. Azerb.">Akhkemal Ridge. Azerb.</location>
[aidjan] alt. 
<quantity metricMagnitude="2" metricUnit="m" metricValue="2.0" unit="m" value="200.0">
<elevation metricMagnitude="2" metricUnit="m" metricValue="2.0" unit="m" value="200.0">200 m</elevation>
</quantity>
]; 
<collectingDate value="1964-06-15">15 Jun. 1964</collectingDate>
; 
<collectorName>Pastukhov</collectorName>
leg.; ZISP
</materialsCitation>
; 
<materialsCitation collectingDate="1932-05-06" collectingDateMax="2018-05-10" collectingDateMin="1932-05-06" collectorName="B. Rohdendorf, V. Lantsov, V. Pilipenko" country="Russia" county="Krasnodarskiy" determinerName="Ried" elevation="54" location="Pontica" municipality="Sotshi prov." specimenCount="37" specimenCount-female="16" specimenCount-male="21">
<collectingCountry name="Russia">Russia</collectingCountry>
- 
<emphasis bold="true" pageId="0" pageNumber="145">
<collectingCounty>Krasnodarskiy</collectingCounty>
kray
</emphasis>
• 
<specimenCount type="male">1 male</specimenCount>
; 
<collectingMunicipality>Sotshi prov.</collectingMunicipality>
<location LSID="urn:lsid:plazi:treatment:219A9352B86056889DBEB0947CC84DEA:FD55ACAB2034F90A598D387613A1A4C0" country="Russia" county="Krasnodarskiy" municipality="Sotshi prov." name="Pontica">Pontica</location>
; 
<collectingDate value="1932-05-06">6 May 1932</collectingDate>
; 
<collectorName>B.Rohdendorf</collectorName>
leg.; / &quot; 
<taxonomicName genus="T." lsidName="T." pageId="0" pageNumber="145" rank="genus">T.</taxonomicName>
<taxonomicName authorityName="Riedel" authorityYear="1920" class="Insecta" family="Tipulidae" genus="Tipula" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tipula caucasica" order="Diptera" pageId="0" pageNumber="145" phylum="Arthropoda" rank="species" species="caucasica">Tipula caucasica</taxonomicName>
<determinerName>Ried</determinerName>
det. Lacksch.&quot; [terminalia in 
<collectingCountry name="Canada">Canada</collectingCountry>
balsam pinned with specimen]; ZISP • 
<specimenCount type="female">1 female</specimenCount>
, Khosta, 
<location LSID="urn:lsid:plazi:treatment:219A9352B86056889DBEB0947CC84DEA:6D6A33030C8E654AED854694996317AC" country="Russia" county="Krasnodarskiy" municipality="Sotshi prov." name="Caucasian Reserve">Caucasian Reserve</location>
, Tiso-samshitovaya rosha [Yew-and-Boxwood Tree Grove]; 43°31'656&quot;N, 39°52'467&quot;E; alt. 
<quantity metricMagnitude="1" metricUnit="m" metricValue="5.4" unit="m" value="54.0">
<elevation metricMagnitude="1" metricUnit="m" metricValue="5.4" unit="m" value="54.0">54 m</elevation>
</quantity>
; collected at light, 
<collectingDate value="2018-05-08">8 May 2018</collectingDate>
; 
<collectorName>V. Lantsov</collectorName>
leg.; IEMT • 
<specimenCount type="male">2 males</specimenCount>
, 
<specimenCount type="female">2 females</specimenCount>
; same collection data as for preceding, 43°31'688&quot;N, 39°52'561&quot;E; alt. 
<quantity metricMagnitude="1" metricUnit="m" metricValue="3.5" unit="m" value="35.0">
<elevation metricMagnitude="1" metricUnit="m" metricValue="3.5" unit="m" value="35.0">35 m</elevation>
</quantity>
; 
<collectingDate value="2018-05-08">8 May 2018</collectingDate>
; 
<collectorName>V. Lantsov</collectorName>
leg. / Lime-beech (
<taxonomicName class="Dicotyledoneae" family="Tiliaceae" genus="Tilia" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="Tilia begoniifolia" order="Malvales" pageId="0" pageNumber="145" phylum="Angiospermae" rank="species" species="begoniifolia">
<emphasis italics="true" pageId="0" pageNumber="145">Tilia begoniifolia</emphasis>
</taxonomicName>
and 
<taxonomicName authorityName="Lipsky" authorityYear="1898" class="Dicotyledoneae" family="Fagaceae" genus="Fagus" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="Fagus orientalis" order="Fagales" pageId="0" pageNumber="145" phylum="Angiospermae" rank="species" species="orientalis">
<emphasis italics="true" pageId="0" pageNumber="145">Fagus orientalis</emphasis>
</taxonomicName>
) forest with yew (
<taxonomicName class="Pinopsida" family="Taxaceae" genus="Taxus" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="Taxus baccata" order="Taxales" pageId="0" pageNumber="145" phylum="Coniferophyta" rank="species" species="baccata">
<emphasis italics="true" pageId="0" pageNumber="145">Taxus baccata</emphasis>
</taxonomicName>
) and hornbeam (
<taxonomicName class="Dicotyledoneae" family="Betulaceae" genus="Carpinus" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="Carpinus betulus" order="Fagales" pageId="0" pageNumber="145" phylum="Angiospermae" rank="species" species="betulus">
<emphasis italics="true" pageId="0" pageNumber="145">Carpinus betulus</emphasis>
</taxonomicName>
) in tree layer and 
<normalizedToken originalValue="butcher’s-broom">butcher's-broom</normalizedToken>
(
<taxonomicName authorityName="Yeo" authorityYear="1966" class="Monocotyledoneae" family="Liliaceae" genus="Ruscus" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="Ruscus colchicus" order="Liliiflorae" pageId="0" pageNumber="145" phylum="Angiospermae" rank="species" species="colchicus">
<emphasis italics="true" pageId="0" pageNumber="145">Ruscus colchicus</emphasis>
</taxonomicName>
- dominant and 
<taxonomicName class="Monocotyledoneae" family="Liliaceae" genus="Ruscus" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="Ruscus aculeatus" order="Liliiflorae" pageId="0" pageNumber="145" phylum="Angiospermae" rank="species" species="aculeatus">
<emphasis italics="true" pageId="0" pageNumber="145">Ruscus aculeatus</emphasis>
</taxonomicName>
) in ground layer [one of the females was collected in the same community in a damp place near a yew log]; IEMT • 
<specimenCount type="male">2 males</specimenCount>
, 
<specimenCount type="female">2 females</specimenCount>
, same collection data as for preceding, 43°31'780&quot;N, 39°52'518&quot;E; alt. 
<quantity metricMagnitude="1" metricUnit="m" metricValue="9.8" unit="m" value="98.0">
<elevation metricMagnitude="1" metricUnit="m" metricValue="9.8" unit="m" value="98.0">98 m</elevation>
</quantity>
; 
<collectingDate value="2018-05-09">9 May 2018</collectingDate>
; 
<collectorName>V. Lantsov</collectorName>
leg.; / Hornbeam - ash (
<taxonomicName authorityName="Mill" authorityYear="1768" class="Dicotyledoneae" family="Betulaceae" genus="Carpinus" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="Carpinus orientalis" order="Fagales" pageId="0" pageNumber="145" phylum="Angiospermae" rank="species" species="orientalis">
<emphasis italics="true" pageId="0" pageNumber="145">Carpinus orientalis</emphasis>
</taxonomicName>
, 
<taxonomicName class="Dicotyledoneae" family="Oleaceae" genus="Fraxinus" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="Fraxinus excelsior" order="Oleales" pageId="0" pageNumber="145" phylum="Angiospermae" rank="species" species="excelsior">
<emphasis italics="true" pageId="0" pageNumber="145">Fraxinus excelsior</emphasis>
</taxonomicName>
) forest with addition of yew (
<taxonomicName class="Pinopsida" family="Taxaceae" genus="Taxus" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="Taxus baccata" order="Taxales" pageId="0" pageNumber="145" phylum="Coniferophyta" rank="species" species="baccata">
<emphasis italics="true" pageId="0" pageNumber="145">Taxus baccata</emphasis>
</taxonomicName>
) and lime (
<taxonomicName class="Dicotyledoneae" family="Tiliaceae" genus="Tilia" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="Tilia begoniifolia" order="Malvales" pageId="0" pageNumber="145" phylum="Angiospermae" rank="species" species="begoniifolia">
<emphasis italics="true" pageId="0" pageNumber="145">Tilia begoniifolia</emphasis>
</taxonomicName>
) with blackberry (
<taxonomicName class="Magnoliopsida" family="Rosaceae" genus="Rubus" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Rubus sanctus" order="Rosales" pageId="0" pageNumber="145" phylum="Tracheophyta" rank="species" species="sanctus">
<emphasis italics="true" pageId="0" pageNumber="145">Rubus sanctus</emphasis>
</taxonomicName>
) and fig (
<taxonomicName class="Gastropoda" family="Ficidae" genus="Ficus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ficus carica" order="Littorinimorpha" pageId="0" pageNumber="145" phylum="Mollusca" rank="species" species="carica">
<emphasis italics="true" pageId="0" pageNumber="145">Ficus carica</emphasis>
</taxonomicName>
) in shrub layer and with 
<normalizedToken originalValue="butcher’s-broom">butcher's-broom</normalizedToken>
(
<taxonomicName authorityName="Yeo" authorityYear="1966" class="Monocotyledoneae" family="Liliaceae" genus="Ruscus" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="Ruscus colchicus" order="Liliiflorae" pageId="0" pageNumber="145" phylum="Angiospermae" rank="species" species="colchicus">
<emphasis italics="true" pageId="0" pageNumber="145">Ruscus colchicus</emphasis>
</taxonomicName>
) in ground layer; ZISP • 
<specimenCount type="male">4 males</specimenCount>
, 
<specimenCount type="female">4 females</specimenCount>
, same collection data as for preceding, 43°31'775&quot;N, 39°52'423&quot;E; alt. 
<quantity metricMagnitude="2" metricUnit="m" metricValue="1.01" unit="m" value="101.0">
<elevation metricMagnitude="2" metricUnit="m" metricValue="1.01" unit="m" value="101.0">101 m</elevation>
</quantity>
; 
<collectingDate value="2018-05-09">9 May 2018</collectingDate>
; 
<collectorName>V. Lantsov</collectorName>
leg.; / Hornbeam (
<taxonomicName authorityName="Mill" authorityYear="1768" class="Dicotyledoneae" family="Betulaceae" genus="Carpinus" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="Carpinus orientalis" order="Fagales" pageId="0" pageNumber="145" phylum="Angiospermae" rank="species" species="orientalis">
<emphasis italics="true" pageId="0" pageNumber="145">Carpinus orientalis</emphasis>
</taxonomicName>
) forest with 
<normalizedToken originalValue="butcher’s-broom">butcher's-broom</normalizedToken>
(
<taxonomicName authorityName="Yeo" authorityYear="1966" class="Monocotyledoneae" family="Liliaceae" genus="Ruscus" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="Ruscus colchicus" order="Liliiflorae" pageId="0" pageNumber="145" phylum="Angiospermae" rank="species" species="colchicus">
<emphasis italics="true" pageId="0" pageNumber="145">Ruscus colchicus</emphasis>
</taxonomicName>
) and fern (
<taxonomicName class="Polypodiopsida" family="Aspleniaceae" genus="Asplenium" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Asplenium scolopendrium" order="Polypodiales" pageId="0" pageNumber="145" phylum="Polypodiopsida" rank="species" species="scolopendrium">
<emphasis italics="true" pageId="0" pageNumber="145">Asplenium scolopendrium</emphasis>
</taxonomicName>
) in ground layer; ZISP • 
<specimenCount type="male">1 male</specimenCount>
and 
<specimenCount type="female">1 female</specimenCount>
(in copula), 
<specimenCount type="female">1 female</specimenCount>
; same collection data as for preceding, 43°31'913&quot;N, 39°52'463&quot;E; alt. 
<quantity metricMagnitude="2" metricUnit="m" metricValue="1.53" unit="m" value="153.0">
<elevation metricMagnitude="2" metricUnit="m" metricValue="1.53" unit="m" value="153.0">153 m</elevation>
</quantity>
; 
<collectingDate value="2018-05-09">9 May 2018</collectingDate>
, 
<collectorName>V. Lantsov</collectorName>
leg. / Ash-lime (
<taxonomicName class="Dicotyledoneae" family="Oleaceae" genus="Fraxinus" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="Fraxinus excelsior" order="Oleales" pageId="0" pageNumber="145" phylum="Angiospermae" rank="species" species="excelsior">
<emphasis italics="true" pageId="0" pageNumber="145">Fraxinus excelsior</emphasis>
</taxonomicName>
+ 
<taxonomicName class="Dicotyledoneae" family="Tiliaceae" genus="Tilia" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="Tilia begoniifolia" order="Malvales" pageId="0" pageNumber="145" phylum="Angiospermae" rank="species" species="begoniifolia">
<emphasis italics="true" pageId="0" pageNumber="145">Tilia begoniifolia</emphasis>
</taxonomicName>
) forest with hornbeam (
<taxonomicName class="Dicotyledoneae" family="Betulaceae" genus="Carpinus" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="Carpinus betulus" order="Fagales" pageId="0" pageNumber="145" phylum="Angiospermae" rank="species" species="betulus">
<emphasis italics="true" pageId="0" pageNumber="145">Carpinus betulus</emphasis>
</taxonomicName>
) and with black berry (
<taxonomicName class="Magnoliopsida" family="Rosaceae" genus="Rubus" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Rubus anatolicus" order="Rosales" pageId="0" pageNumber="145" phylum="Tracheophyta" rank="species" species="anatolicus">
<emphasis italics="true" pageId="0" pageNumber="145">Rubus anatolicus</emphasis>
</taxonomicName>
) and miscellaneous herbs in ground layer; IEMT • 
<specimenCount type="male">2 males</specimenCount>
, 
<specimenCount type="female">1 female</specimenCount>
, same collection data as for preceding, 43°31'825&quot;N, 39°52'653&quot;E; alt. 
<quantity metricMagnitude="1" metricUnit="m" metricValue="9.9" unit="m" value="99.0">
<elevation metricMagnitude="1" metricUnit="m" metricValue="9.9" unit="m" value="99.0">99 m</elevation>
</quantity>
; 
<collectingDate value="2018-05-10">10 May 2018</collectingDate>
, 
<collectorName>V. Lantsov</collectorName>
leg.; IEMT • 
<specimenCount type="male">1 male</specimenCount>
, same collection data as for preceding, 43°31'656&quot;N, 39°52'467&quot;E; alt. 
<quantity metricMagnitude="1" metricUnit="m" metricValue="5.4" unit="m" value="54.0">
<elevation metricMagnitude="1" metricUnit="m" metricValue="5.4" unit="m" value="54.0">54 m</elevation>
</quantity>
; 
<collectorName>V. Lantsov</collectorName>
leg.; ZISP • 
<specimenCount type="male">2 males</specimenCount>
, 
<specimenCount type="female">1 female</specimenCount>
, same collection data as for preceding, 43°32'257&quot;N, 39°52'682&quot;E; alt. 
<quantity metricMagnitude="1" metricUnit="m" metricValue="9.4" unit="m" value="94.0">
<elevation metricMagnitude="1" metricUnit="m" metricValue="9.4" unit="m" value="94.0">94 m</elevation>
</quantity>
; 
<collectingDate value="2018-05-10">10 May 2018</collectingDate>
, 
<collectorName>V. Lantsov</collectorName>
leg.; / beech (
<taxonomicName authorityName="Lipsky" authorityYear="1898" class="Dicotyledoneae" family="Fagaceae" genus="Fagus" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="Fagus orientalis" order="Fagales" pageId="0" pageNumber="145" phylum="Angiospermae" rank="species" species="orientalis">
<emphasis italics="true" pageId="0" pageNumber="145">Fagus orientalis</emphasis>
</taxonomicName>
) forest with addition of yew (
<taxonomicName class="Pinopsida" family="Taxaceae" genus="Taxus" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="Taxus baccata" order="Taxales" pageId="0" pageNumber="145" phylum="Coniferophyta" rank="species" species="baccata">
<emphasis italics="true" pageId="0" pageNumber="145">Taxus baccata</emphasis>
</taxonomicName>
) and hornbeam (
<taxonomicName class="Dicotyledoneae" family="Betulaceae" genus="Carpinus" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="Carpinus betulus" order="Fagales" pageId="0" pageNumber="145" phylum="Angiospermae" rank="species" species="betulus">
<emphasis italics="true" pageId="0" pageNumber="145">Carpinus betulus</emphasis>
</taxonomicName>
) with bunch grass (
<taxonomicName class="Monocotyledoneae" family="Poaceae" genus="Calamagrostis" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="Calamagrostis arundinacea" order="Graminales" pageId="0" pageNumber="145" phylum="Angiospermae" rank="species" species="arundinacea">
<emphasis italics="true" pageId="0" pageNumber="145">Calamagrostis arundinacea</emphasis>
</taxonomicName>
) in ground layer; IEMT • 
<specimenCount type="male">6 males</specimenCount>
, 
<specimenCount type="female">3 females</specimenCount>
; Krasnodar Territory, environs 
<location LSID="urn:lsid:plazi:treatment:219A9352B86056889DBEB0947CC84DEA:B80E7705F8EFD3C39DECBD708528B115" country="Russia" county="Krasnodarskiy" municipality="Sotshi prov." name="Shakhe River">Shakhe River</location>
higher than Salokh-Aul; 
<collectingDate value="2011-06-05" valueMax="2011-06-08" valueMin="2011-06-05">5-8 Jun. 2011</collectingDate>
; 
<collectorName>V. Pilipenko</collectorName>
leg.; VPMC
</materialsCitation>
; 
<materialsCitation collectingDate="1909-06-29" collectingDateMax="2016-07-05" collectingDateMin="1909-06-29" collectorName="Mlokosevich, V. Lantsov" country="Russia" county="Tlyaratinsky district" determinerName="Ried" elevation="1792" location="river Dzhurmut" municipality="Salda" specimenCount="11" specimenCount-female="3" specimenCount-male="8" stateProvince="Dagestan">
<collectingCountry name="Russia">Russia</collectingCountry>
- 
<emphasis bold="true" pageId="0" pageNumber="145">
<collectingRegion country="Russia" name="Dagestan">Dagestan</collectingRegion>
</emphasis>
• 
<specimenCount type="female">1 female</specimenCount>
, 
<normalizedToken originalValue="“Хочал-Даг">&quot;Khochal-Dag</normalizedToken>
, 
<normalizedToken originalValue="альп">alp</normalizedToken>
. 
<normalizedToken originalValue="обл">obl</normalizedToken>
. 
<normalizedToken originalValue="Дагст">Dagst</normalizedToken>
. [
<normalizedToken originalValue="Дагестан]”">
<collectingRegion country="Russia" name="Dagestan">Dagestan</collectingRegion>
]&quot;
</normalizedToken>
[Hochal-Dag, alp. Region Dagst. [
<collectingRegion country="Russia" name="Dagestan">Dagestan</collectingRegion>
]]; 
<collectingDate value="1909-06-29">29 Jun. 1909</collectingDate>
<collectorName>Mlokosevich</collectorName>
leg.; ZISP • 
<specimenCount type="female">1 female</specimenCount>
; 
<collectingCounty>Tlyaratinsky district</collectingCounty>
, near village 
<collectingMunicipality>Salda</collectingMunicipality>
, stream on the right bank of the 
<location LSID="urn:lsid:plazi:treatment:219A9352B86056889DBEB0947CC84DEA:10A03F8FDE20845170E21CD52183F14F" country="Russia" county="Tlyaratinsky district" municipality="Salda" name="river Dzhurmut" stateProvince="Dagestan">river Dzhurmut</location>
; 41°58'388&quot;N, 46°30'446&quot;E; alt. 
<quantity metricMagnitude="3" metricUnit="m" metricValue="1.7919999999999998" unit="m" value="1792.0">
<elevation metricMagnitude="3" metricUnit="m" metricValue="1.7919999999999998" unit="m" value="1792.0">1792 m</elevation>
</quantity>
; 
<collectingDate value="2016-07-02">2 Jul. 2016</collectingDate>
, 
<collectorName>V. Lantsov</collectorName>
leg.; / &quot;Oak (
<taxonomicName class="Dicotyledoneae" family="Fagaceae" genus="Quercus" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="Quercus iberica" order="Fagales" pageId="0" pageNumber="145" phylum="Angiospermae" rank="species" species="iberica">
<emphasis italics="true" pageId="0" pageNumber="145">Quercus iberica</emphasis>
</taxonomicName>
) sparse forest with 
<taxonomicName class="Dicotyledoneae" family="Salicaceae" genus="Populus" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="Populus tremula" order="Salicales" pageId="0" pageNumber="145" phylum="Angiospermae" rank="species" species="tremula">
<emphasis italics="true" pageId="0" pageNumber="145">Populus tremula</emphasis>
</taxonomicName>
and 
<taxonomicName class="Dicotyledoneae" family="Aceraceae" genus="Acer" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="Acer platanoides" order="Sapindales" pageId="0" pageNumber="145" phylum="Angiospermae" rank="species" species="platanoides">
<emphasis italics="true" pageId="0" pageNumber="145">Acer platanoides</emphasis>
</taxonomicName>
in tree layer, 
<emphasis italics="true" pageId="0" pageNumber="145">Rosa canina</emphasis>
, 
<taxonomicName class="Dicotyledoneae" family="Rosaceae" genus="Spiraea" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="Spiraea crenata" order="Rosales" pageId="0" pageNumber="145" phylum="Angiospermae" rank="species" species="crenata">
<emphasis italics="true" pageId="0" pageNumber="145">Spiraea crenata</emphasis>
</taxonomicName>
and 
<taxonomicName class="Dicotyledoneae" family="Rosaceae" genus="Cotoneaster" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="Cotoneaster integerrimus" order="Rosales" pageId="0" pageNumber="145" phylum="Angiospermae" rank="species" species="integerrimus">
<emphasis italics="true" pageId="0" pageNumber="145">Cotoneaster integerrimus</emphasis>
</taxonomicName>
in shrub, and clover (
<taxonomicName class="Magnoliopsida" family="Fabaceae" genus="Trifolium" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Trifolium canescens" order="Fabales" pageId="0" pageNumber="145" phylum="Tracheophyta" rank="species" species="canescens">
<emphasis italics="true" pageId="0" pageNumber="145">Trifolium canescens</emphasis>
</taxonomicName>
) and miscellaneous herbs in ground layer; ZISP • 
<specimenCount type="male">2 males</specimenCount>
, 
<specimenCount type="female">1 female</specimenCount>
; near village Betelda, stream on the left bank of the 
<location LSID="urn:lsid:plazi:treatment:219A9352B86056889DBEB0947CC84DEA:FED2A7EFB21D4CAE46D06E4881F53C13" country="Russia" county="Tlyaratinsky district" municipality="Salda" name="river Dzhurmut" stateProvince="Dagestan">river Dzhurmut</location>
; 41°56'853&quot;N, 46°32'424&quot;E; alt. 
<quantity metricMagnitude="3" metricUnit="m" metricValue="1.85" unit="m" value="1850.0">
<elevation metricMagnitude="3" metricUnit="m" metricValue="1.85" unit="m" value="1850.0">1850 m</elevation>
</quantity>
; 
<collectingDate value="2016-07-03">3 Jul. 2016</collectingDate>
, 
<collectorName>V. Lantsov</collectorName>
leg.; / Willow (
<taxonomicName class="Ascidiacea" family="Polycitoridae" genus="Salix" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Salix caprea" order="Aplousobranchia" pageId="0" pageNumber="145" phylum="Chordata" rank="species" species="caprea">
<emphasis italics="true" pageId="0" pageNumber="145">Salix caprea</emphasis>
</taxonomicName>
) shrub with herbaceous layer (
<taxonomicName class="Dicotyledoneae" family="Onagraceae" genus="Epilobium" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="Epilobium montanum" order="Myrtiflorae" pageId="0" pageNumber="145" phylum="Angiospermae" rank="species" species="montanum">
<emphasis italics="true" pageId="0" pageNumber="145">Epilobium montanum</emphasis>
</taxonomicName>
) (dominant), 
<taxonomicName authorityName="Albov" authorityYear="1894" class="Dicotyledoneae" family="Brassicaceae" genus="Cardamine" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="Cardamine seidlitziana" order="Papaverales" pageId="0" pageNumber="145" phylum="Angiospermae" rank="species" species="seidlitziana">
<emphasis italics="true" pageId="0" pageNumber="145">Cardamine seidlitziana</emphasis>
</taxonomicName>
, 
<taxonomicName class="Dicotyledoneae" family="Lamiaceae" genus="Mentha" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="Mentha caucasica" order="Tubiflorae" pageId="0" pageNumber="145" phylum="Angiospermae" rank="species" species="caucasica">
<emphasis italics="true" pageId="0" pageNumber="145">Mentha caucasica</emphasis>
</taxonomicName>
, 
<taxonomicName class="Dicotyledoneae" family="Scrophulariaceae" genus="Veronica" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="Veronica anagallis-aquatica" order="Tubiflorae" pageId="0" pageNumber="145" phylum="Angiospermae" rank="species" species="anagallis-aquatica">
<emphasis italics="true" pageId="0" pageNumber="145">Veronica anagallis-aquatica</emphasis>
</taxonomicName>
and 
<taxonomicName authorityName="Moench" authorityYear="1794" baseAuthorityName="L." class="Dicotyledoneae" family="Caryophyllaceae" genus="Myosoton" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="Myosoton aquaticum" order="Centrospermae" pageId="0" pageNumber="145" phylum="Angiospermae" rank="species" species="aquaticum">
<emphasis italics="true" pageId="0" pageNumber="145">Myosoton aquaticum</emphasis>
</taxonomicName>
community near spring above the flood plain of the left bank of the river; ZISP • 
<specimenCount type="male">2 males</specimenCount>
; near village Genekolob, on the high left bank of the river Dzhurmut; 41°57'392&quot;N, 46°31'276&quot;E; alt. 
<quantity metricMagnitude="3" metricUnit="m" metricValue="2.011" unit="m" value="2011.0">
<elevation metricMagnitude="3" metricUnit="m" metricValue="2.011" unit="m" value="2011.0">2011 m</elevation>
</quantity>
; 
<collectingDate value="2016-07-04">4 Jul. 2016</collectingDate>
, 
<collectorName>V. Lantsov</collectorName>
leg.; / Oak (
<taxonomicName class="Dicotyledoneae" family="Fagaceae" genus="Quercus" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="Quercus iberica" order="Fagales" pageId="0" pageNumber="145" phylum="Angiospermae" rank="species" species="iberica">
<emphasis italics="true" pageId="0" pageNumber="145">Quercus iberica</emphasis>
</taxonomicName>
) forest with mountain-ash (
<taxonomicName class="Dicotyledoneae" family="Rosaceae" genus="Sorbus" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="Sorbus aucuparia" order="Rosales" pageId="0" pageNumber="145" phylum="Angiospermae" rank="species" species="aucuparia">
<emphasis italics="true" pageId="0" pageNumber="145">Sorbus aucuparia</emphasis>
</taxonomicName>
), wayfaring tree (
<taxonomicName class="Dicotyledoneae" family="Caprifoliaceae" genus="Viburnum" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="Viburnum lantana" order="Dipsacales" pageId="0" pageNumber="145" phylum="Angiospermae" rank="species" species="lantana">
<emphasis italics="true" pageId="0" pageNumber="145">Viburnum lantana</emphasis>
</taxonomicName>
) in shrub layer [the lower border of a forest and a forb-cereal meadow]; ZISP • 
<specimenCount type="male">2 males</specimenCount>
; same collection data as for preceding, 41°57'449&quot;N, 46°31'228&quot;E; alt. 
<quantity metricMagnitude="3" metricUnit="m" metricValue="1.9769999999999999" unit="m" value="1977.0">
<elevation metricMagnitude="3" metricUnit="m" metricValue="1.9769999999999999" unit="m" value="1977.0">1977 m</elevation>
</quantity>
; 
<collectingDate value="2016-07-04">4 Jul. 2016</collectingDate>
, 
<collectorName>V. Lantsov</collectorName>
leg.; ZISP • 
<specimenCount type="male">2 males</specimenCount>
, between the villages of Gerel and Genekolob, the right bank of the river Dzhurmut, 41°55'619&quot;N, 46°34'468&quot;E; alt. 
<quantity metricMagnitude="3" metricUnit="m" metricValue="1.925" unit="m" value="1925.0">
<elevation metricMagnitude="3" metricUnit="m" metricValue="1.925" unit="m" value="1925.0">1925 m</elevation>
</quantity>
; 
<collectingDate value="2016-07-05">5 Jul. 2016</collectingDate>
, 
<collectorName>V. Lantsov</collectorName>
leg.; / Above the floodplain terrace with a rare herbaceous birch (
<taxonomicName class="Dicotyledoneae" family="Betulaceae" genus="Betula" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="Betula raddeana" order="Fagales" pageId="0" pageNumber="145" phylum="Angiospermae" rank="species" species="raddeana">
<emphasis italics="true" pageId="0" pageNumber="145">Betula raddeana</emphasis>
</taxonomicName>
) forest with willow (
<taxonomicName class="Ascidiacea" family="Polycitoridae" genus="Salix" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Salix caprea" order="Aplousobranchia" pageId="0" pageNumber="145" phylum="Chordata" rank="species" species="caprea">
<emphasis italics="true" pageId="0" pageNumber="145">Salix caprea</emphasis>
</taxonomicName>
) (coppice forest); ZISP
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="145" type="diagnosis">
<paragraph pageId="0" pageNumber="145">Diagnosis.</paragraph>
<paragraph pageId="0" pageNumber="145">Dark stripes medially on lighter background extending entire length of scutum. Inner gonostylus posteriorly with wedge-shaped projection directed anteriorly. Tergite 9 at apex with three rounded notches, middle one usually deepest and widest.</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="145" type="redescription">
<paragraph pageId="0" pageNumber="145">Redescription.</paragraph>
<paragraph pageId="0" pageNumber="145">
Adult male (Fig. 
<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. T. (Lunatipula) caucasica male habitus A general lateral view B head and thorax, dorsal view. Scale bars: 10 mm (A), 1 mm (B)." figureDoi="10.3897/zookeys.1048.67564.figure1" httpUri="https://binary.pensoft.net/fig/566410" pageId="0" pageNumber="145">1A, B</figureCitation>
). General color grey. Body length 14-17 mm, wing 14-17.5 mm.
</paragraph>
<caption doi="10.3897/zookeys.1048.67564.figure1" httpUri="https://binary.pensoft.net/fig/566410" pageId="0" pageNumber="145" start="Figure 1" startId="F1">
<paragraph pageId="0" pageNumber="145">
<emphasis bold="true" pageId="0" pageNumber="145">Figure 1.</emphasis>
<taxonomicName lsidName="T. (Lunatipula) caucasica" pageId="0" pageNumber="145" rank="species" species="caucasica" subGenus="Lunatipula">T. (Lunatipula) caucasica</taxonomicName>
male habitus 
<emphasis bold="true" pageId="0" pageNumber="145">A</emphasis>
general lateral view 
<emphasis bold="true" pageId="0" pageNumber="145">B</emphasis>
head and thorax, dorsal view. Scale bars: 10 mm (
<emphasis bold="true" pageId="0" pageNumber="145">A</emphasis>
), 1 mm (
<emphasis bold="true" pageId="0" pageNumber="145">B</emphasis>
).
</paragraph>
</caption>
<paragraph pageId="0" pageNumber="145">
<emphasis bold="true" italics="true" pageId="0" pageNumber="145">Head.</emphasis>
Light grey, sometimes bluish (Figs 
<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. T. (Lunatipula) caucasica male habitus A general lateral view B head and thorax, dorsal view. Scale bars: 10 mm (A), 1 mm (B)." figureDoi="10.3897/zookeys.1048.67564.figure1" httpUri="https://binary.pensoft.net/fig/566410" pageId="0" pageNumber="145">1B</figureCitation>
, 
<figureCitation captionStart="Figure 13" captionStartId="F13" captionText="Figure 13. Heads of Tipula caucasica species group, dorsal view A T. (Lunatipula) caucasica B T. (Lunatipula) eleniya sp. nov. C T. (Lunatipula) paupera D T. (Lunatipula) quadridentata E T. (Lunatipula) Tipula talyshensis. Scale bar: 1 mm." figureDoi="10.3897/zookeys.1048.67564.figure13" httpUri="https://binary.pensoft.net/fig/566422" pageId="0" pageNumber="145">13A</figureCitation>
). Rostrum whitish dorsally, flanks rusty or brownish with whitish bristles; nasus distinct, with long, whitish yellow setae; palpus grey. Frons with rows of brown setae along inner edge of eye. Vertex with faint dark median line, posteriorly with dark grey striation, with group of small rusty setae arranged fan-like on either side of glabrous medial area. Genae rusty; wide dark median stripe ventrally on rostrum, continuing along gula midline. Pharyngeal foramen framed with long, rusty hair-like setae. Postgenae light grey.
</paragraph>
<paragraph pageId="0" pageNumber="145">
<emphasis bold="true" italics="true" pageId="0" pageNumber="145">Antenna</emphasis>
(Fig. 
<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. T. (Lunatipula) caucasica male habitus A general lateral view B head and thorax, dorsal view. Scale bars: 10 mm (A), 1 mm (B)." figureDoi="10.3897/zookeys.1048.67564.figure1" httpUri="https://binary.pensoft.net/fig/566410" pageId="0" pageNumber="145">1B</figureCitation>
) 13-segmented, if bent backwards just reaching wing base. Scape light brown or grey, depending on angle of rotation to light source, with slightly pronounced silvery pruinescence and brown horizontal striations. Pedicel yellow, sometimes brownish at base. Flagellomeres brownish, thickened at base, verticils subequal in length to corresponding flagellomeres.
</paragraph>
<paragraph pageId="0" pageNumber="145">
<emphasis bold="true" italics="true" pageId="0" pageNumber="145">Thorax</emphasis>
(Fig. 
<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. T. (Lunatipula) caucasica male habitus A general lateral view B head and thorax, dorsal view. Scale bars: 10 mm (A), 1 mm (B)." figureDoi="10.3897/zookeys.1048.67564.figure1" httpUri="https://binary.pensoft.net/fig/566410" pageId="0" pageNumber="145">1B</figureCitation>
). Grey or blue-grey. Pronotum with dark grey median stripe. Katepisternum, anepisternum and meron grey, glabrous. Scutum with four longitudinal wide brownish stripes on lighter background; medial stripes extending length of scutum. Scutal lobes, scutellum and mediotergite grey with light bristles in dark brown sockets.
</paragraph>
<paragraph pageId="0" pageNumber="145">
<emphasis bold="true" italics="true" pageId="0" pageNumber="145">Wings</emphasis>
(Fig. 
<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. T. (Lunatipula) caucasica male habitus A general lateral view B head and thorax, dorsal view. Scale bars: 10 mm (A), 1 mm (B)." figureDoi="10.3897/zookeys.1048.67564.figure1" httpUri="https://binary.pensoft.net/fig/566410" pageId="0" pageNumber="145">1A</figureCitation>
) greyish with a bluish tinge in transmitted light, stigma brown; narrow whitish area (lunule) extending to base of discal cell.
</paragraph>
<paragraph pageId="0" pageNumber="145">
<taxonomicName genus="Halteres" lsidName="Halteres" pageId="0" pageNumber="145" rank="genus">
<emphasis bold="true" italics="true" pageId="0" pageNumber="145">Halteres</emphasis>
</taxonomicName>
(Fig. 
<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. T. (Lunatipula) caucasica male habitus A general lateral view B head and thorax, dorsal view. Scale bars: 10 mm (A), 1 mm (B)." figureDoi="10.3897/zookeys.1048.67564.figure1" httpUri="https://binary.pensoft.net/fig/566410" pageId="0" pageNumber="145">1A</figureCitation>
) with light brown or dark-yellow stem, lighter yellow at base, covered with short small pale setae; knob dark brown to black.
</paragraph>
<paragraph pageId="0" pageNumber="145">
<emphasis bold="true" italics="true" pageId="0" pageNumber="145">Legs</emphasis>
with coxae light grey with long light hairs. Coxae may appear darker when specimen is rotated. Trochanters yellow. Femora yellow at base, rest of femora, tibiae, and tarsi dark brown.
</paragraph>
<paragraph pageId="0" pageNumber="145">
<emphasis bold="true" italics="true" pageId="0" pageNumber="145">Abdomen</emphasis>
(Figs 
<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. T. (Lunatipula) caucasica male habitus A general lateral view B head and thorax, dorsal view. Scale bars: 10 mm (A), 1 mm (B)." figureDoi="10.3897/zookeys.1048.67564.figure1" httpUri="https://binary.pensoft.net/fig/566410" pageId="0" pageNumber="145">1A</figureCitation>
, 
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. T. (Lunatipula) caucasica A terminal segments of male abdomen, lateral view (dry) B-L male terminalia (after KOH 10 % treatment) B hypopygium, caudal view C hypopygium, lateral view D sternite 8, ventral view E tergite 9, caudal view F tergite 9, dorsal view G inner and outer gonostylus, lateral view H inner and outer gonostylus, caudal view I gonocoxal fragment, dorsal view J semen pump and aedeagus, lateral view K adminiculum and paired apical appendages of sternite 9, caudal view L adminiculum and paired apical appendages of sternite 9, lateral view. Abbreviations: adm-adminiculum; as 8 - paired appendages of sternite 8; as 9 - apical appendages of sternite 9; ig-inner gonostylus; gcx-gonocoxite; og-outer gonostylus; s 8 - sternite 8; s 9 - sternite 9; t 9 - tergite 9. Scale bar: 1 mm." figureDoi="10.3897/zookeys.1048.67564.figure2" httpUri="https://binary.pensoft.net/fig/566411" pageId="0" pageNumber="145">2A</figureCitation>
). Grey or dark grey; tergites with dark lateral stripe narrowly interrupted with whitish at posterior margins, lateral margins with wider whitish edge.
</paragraph>
<caption doi="10.3897/zookeys.1048.67564.figure2" httpUri="https://binary.pensoft.net/fig/566411" pageId="0" pageNumber="145" start="Figure 2" startId="F2">
<paragraph pageId="0" pageNumber="145">
<emphasis bold="true" pageId="0" pageNumber="145">Figure 2.</emphasis>
<taxonomicName lsidName="T. (Lunatipula) caucasica" pageId="0" pageNumber="145" rank="species" species="caucasica" subGenus="Lunatipula">T. (Lunatipula) caucasica</taxonomicName>
<emphasis bold="true" pageId="0" pageNumber="145">A</emphasis>
terminal segments of male abdomen, lateral view (dry) 
<emphasis bold="true" pageId="0" pageNumber="145">B-L</emphasis>
male terminalia (after KOH 10% treatment) 
<emphasis bold="true" pageId="0" pageNumber="145">B</emphasis>
hypopygium, caudal view 
<emphasis bold="true" pageId="0" pageNumber="145">C</emphasis>
hypopygium, lateral view 
<emphasis bold="true" pageId="0" pageNumber="145">D</emphasis>
sternite 8, ventral view 
<emphasis bold="true" pageId="0" pageNumber="145">E</emphasis>
tergite 9, caudal view 
<emphasis bold="true" pageId="0" pageNumber="145">F</emphasis>
tergite 9, dorsal view 
<emphasis bold="true" pageId="0" pageNumber="145">G</emphasis>
inner and outer gonostylus, lateral view 
<emphasis bold="true" pageId="0" pageNumber="145">H</emphasis>
inner and outer gonostylus, caudal view 
<emphasis bold="true" pageId="0" pageNumber="145">I</emphasis>
gonocoxal fragment, dorsal view 
<emphasis bold="true" pageId="0" pageNumber="145">J</emphasis>
semen pump and aedeagus, lateral view 
<emphasis bold="true" pageId="0" pageNumber="145">K</emphasis>
adminiculum and paired apical appendages of sternite 9, caudal view 
<emphasis bold="true" pageId="0" pageNumber="145">L</emphasis>
adminiculum and paired apical appendages of sternite 9, lateral view. Abbreviations: adm-adminiculum; as 8-paired appendages of sternite 8; as 9-apical appendages of sternite 9; ig-inner gonostylus; gcx-gonocoxite; og-outer gonostylus; s8-sternite 8; s9-sternite 9; t9-tergite 9. Scale bar: 1 mm.
</paragraph>
</caption>
<paragraph pageId="0" pageNumber="145">
<emphasis bold="true" italics="true" pageId="0" pageNumber="145">Terminalia</emphasis>
(Figs 
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. T. (Lunatipula) caucasica A terminal segments of male abdomen, lateral view (dry) B-L male terminalia (after KOH 10 % treatment) B hypopygium, caudal view C hypopygium, lateral view D sternite 8, ventral view E tergite 9, caudal view F tergite 9, dorsal view G inner and outer gonostylus, lateral view H inner and outer gonostylus, caudal view I gonocoxal fragment, dorsal view J semen pump and aedeagus, lateral view K adminiculum and paired apical appendages of sternite 9, caudal view L adminiculum and paired apical appendages of sternite 9, lateral view. Abbreviations: adm-adminiculum; as 8 - paired appendages of sternite 8; as 9 - apical appendages of sternite 9; ig-inner gonostylus; gcx-gonocoxite; og-outer gonostylus; s 8 - sternite 8; s 9 - sternite 9; t 9 - tergite 9. Scale bar: 1 mm." figureDoi="10.3897/zookeys.1048.67564.figure2" httpUri="https://binary.pensoft.net/fig/566411" pageId="0" pageNumber="145">2</figureCitation>
, 
<figureCitation captionStart="Figure 14" captionStartId="F14" captionText="Figure 14. Comparisons of some components of male terminalia of the Tipula caucasica species group A, F T. (Lunatipula) caucasica B, G T. (Lunatipula) eleniya sp. nov. C, I T. (Lunatipula) paupera D, J T. (Lunatipula) quadridentata E, K T. (Lunatipula) talyshensis A-E tergite 9, dorsal view F-K inner and outer gonostylus, lateral view. Arrows point to central notch at apex of tergite 9 (C, D), to the wedge-shaped projection posteriorly on inner gonostylus (F), to the rod-like outgrowth behind anterior beak of inner gonostylus (G), to the anterior (dorsal) margin of gonocoxite (I, J, K). Scale bar: 1 mm." figureDoi="10.3897/zookeys.1048.67564.figure14" httpUri="https://binary.pensoft.net/fig/566423" pageId="0" pageNumber="145">14A, F</figureCitation>
). Hypopygium (Fig. 
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. T. (Lunatipula) caucasica A terminal segments of male abdomen, lateral view (dry) B-L male terminalia (after KOH 10 % treatment) B hypopygium, caudal view C hypopygium, lateral view D sternite 8, ventral view E tergite 9, caudal view F tergite 9, dorsal view G inner and outer gonostylus, lateral view H inner and outer gonostylus, caudal view I gonocoxal fragment, dorsal view J semen pump and aedeagus, lateral view K adminiculum and paired apical appendages of sternite 9, caudal view L adminiculum and paired apical appendages of sternite 9, lateral view. Abbreviations: adm-adminiculum; as 8 - paired appendages of sternite 8; as 9 - apical appendages of sternite 9; ig-inner gonostylus; gcx-gonocoxite; og-outer gonostylus; s 8 - sternite 8; s 9 - sternite 9; t 9 - tergite 9. Scale bar: 1 mm." figureDoi="10.3897/zookeys.1048.67564.figure2" httpUri="https://binary.pensoft.net/fig/566411" pageId="0" pageNumber="145">2B, C</figureCitation>
) not much thicker than rest of abdomen, brown with moderately dense grey pruinescence, with golden yellow hairs. Gonocoxite (Fig. 
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. T. (Lunatipula) caucasica A terminal segments of male abdomen, lateral view (dry) B-L male terminalia (after KOH 10 % treatment) B hypopygium, caudal view C hypopygium, lateral view D sternite 8, ventral view E tergite 9, caudal view F tergite 9, dorsal view G inner and outer gonostylus, lateral view H inner and outer gonostylus, caudal view I gonocoxal fragment, dorsal view J semen pump and aedeagus, lateral view K adminiculum and paired apical appendages of sternite 9, caudal view L adminiculum and paired apical appendages of sternite 9, lateral view. Abbreviations: adm-adminiculum; as 8 - paired appendages of sternite 8; as 9 - apical appendages of sternite 9; ig-inner gonostylus; gcx-gonocoxite; og-outer gonostylus; s 8 - sternite 8; s 9 - sternite 9; t 9 - tergite 9. Scale bar: 1 mm." figureDoi="10.3897/zookeys.1048.67564.figure2" httpUri="https://binary.pensoft.net/fig/566411" pageId="0" pageNumber="145">2C, G</figureCitation>
) with broad tooth ventrally. Tergite 9 (Figs 
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. T. (Lunatipula) caucasica A terminal segments of male abdomen, lateral view (dry) B-L male terminalia (after KOH 10 % treatment) B hypopygium, caudal view C hypopygium, lateral view D sternite 8, ventral view E tergite 9, caudal view F tergite 9, dorsal view G inner and outer gonostylus, lateral view H inner and outer gonostylus, caudal view I gonocoxal fragment, dorsal view J semen pump and aedeagus, lateral view K adminiculum and paired apical appendages of sternite 9, caudal view L adminiculum and paired apical appendages of sternite 9, lateral view. Abbreviations: adm-adminiculum; as 8 - paired appendages of sternite 8; as 9 - apical appendages of sternite 9; ig-inner gonostylus; gcx-gonocoxite; og-outer gonostylus; s 8 - sternite 8; s 9 - sternite 9; t 9 - tergite 9. Scale bar: 1 mm." figureDoi="10.3897/zookeys.1048.67564.figure2" httpUri="https://binary.pensoft.net/fig/566411" pageId="0" pageNumber="145">2C, E, F</figureCitation>
; 
<figureCitation captionStart="Figure 14" captionStartId="F14" captionText="Figure 14. Comparisons of some components of male terminalia of the Tipula caucasica species group A, F T. (Lunatipula) caucasica B, G T. (Lunatipula) eleniya sp. nov. C, I T. (Lunatipula) paupera D, J T. (Lunatipula) quadridentata E, K T. (Lunatipula) talyshensis A-E tergite 9, dorsal view F-K inner and outer gonostylus, lateral view. Arrows point to central notch at apex of tergite 9 (C, D), to the wedge-shaped projection posteriorly on inner gonostylus (F), to the rod-like outgrowth behind anterior beak of inner gonostylus (G), to the anterior (dorsal) margin of gonocoxite (I, J, K). Scale bar: 1 mm." figureDoi="10.3897/zookeys.1048.67564.figure14" httpUri="https://binary.pensoft.net/fig/566423" pageId="0" pageNumber="145">14A</figureCitation>
) transverse, lateral corner broadly rounded, at apex with three rounded notches, middle one usually deepest and widest, bounded laterally by two large dentate projections with blunt apices, clearly visible when viewed from above (Figs 
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. T. (Lunatipula) caucasica A terminal segments of male abdomen, lateral view (dry) B-L male terminalia (after KOH 10 % treatment) B hypopygium, caudal view C hypopygium, lateral view D sternite 8, ventral view E tergite 9, caudal view F tergite 9, dorsal view G inner and outer gonostylus, lateral view H inner and outer gonostylus, caudal view I gonocoxal fragment, dorsal view J semen pump and aedeagus, lateral view K adminiculum and paired apical appendages of sternite 9, caudal view L adminiculum and paired apical appendages of sternite 9, lateral view. Abbreviations: adm-adminiculum; as 8 - paired appendages of sternite 8; as 9 - apical appendages of sternite 9; ig-inner gonostylus; gcx-gonocoxite; og-outer gonostylus; s 8 - sternite 8; s 9 - sternite 9; t 9 - tergite 9. Scale bar: 1 mm." figureDoi="10.3897/zookeys.1048.67564.figure2" httpUri="https://binary.pensoft.net/fig/566411" pageId="0" pageNumber="145">2F</figureCitation>
, 
<figureCitation captionStart="Figure 14" captionStartId="F14" captionText="Figure 14. Comparisons of some components of male terminalia of the Tipula caucasica species group A, F T. (Lunatipula) caucasica B, G T. (Lunatipula) eleniya sp. nov. C, I T. (Lunatipula) paupera D, J T. (Lunatipula) quadridentata E, K T. (Lunatipula) talyshensis A-E tergite 9, dorsal view F-K inner and outer gonostylus, lateral view. Arrows point to central notch at apex of tergite 9 (C, D), to the wedge-shaped projection posteriorly on inner gonostylus (F), to the rod-like outgrowth behind anterior beak of inner gonostylus (G), to the anterior (dorsal) margin of gonocoxite (I, J, K). Scale bar: 1 mm." figureDoi="10.3897/zookeys.1048.67564.figure14" httpUri="https://binary.pensoft.net/fig/566423" pageId="0" pageNumber="145">14A</figureCitation>
) (
<bibRefCitation author="Riedel, MP" journalOrPublisher="Zoologische Jahrbucher, Abteilung fur Systematik, Geographie und Biologie der Tiere" pageId="0" pageNumber="145" pagination="13 - 22" refId="B9" refString="Riedel, MP, 1920. Nematocera Polyneura (Dipt.) aus dem Kaukasus. Zoologische Jahrbucher, Abteilung fur Systematik, Geographie und Biologie der Tiere 43: 13 - 22" title="Nematocera Polyneura (Dipt.) aus dem Kaukasus." volume="43" year="1920">Riedel 1920</bibRefCitation>
; 
<bibRefCitation author="Savchenko, EN" journalOrPublisher="Zoologicheskiy Zhurnal" pageId="0" pageNumber="145" refId="B12" refString="Savchenko, EN, 1964. Crane-flies (Diptera, Tipulidae), Subfam. Tipulinae, Genus TipulaL., 2. Fauna USSR, N.S. 89, Nasekomye Dvukrylye [Diptera] 2(4): 1-503. [in Russian]" title="Crane-flies (Diptera, Tipulidae), Subfam. Tipulinae, Genus TipulaL., 2. Fauna USSR, N. S. 89, Nasekomye Dvukrylye [Diptera] 2 (4): 1 - 503. [in Russian]" year="1964">Savchenko 1964</bibRefCitation>
). When viewed from behind, three pairs of very short projections are visible (Fig. 
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. T. (Lunatipula) caucasica A terminal segments of male abdomen, lateral view (dry) B-L male terminalia (after KOH 10 % treatment) B hypopygium, caudal view C hypopygium, lateral view D sternite 8, ventral view E tergite 9, caudal view F tergite 9, dorsal view G inner and outer gonostylus, lateral view H inner and outer gonostylus, caudal view I gonocoxal fragment, dorsal view J semen pump and aedeagus, lateral view K adminiculum and paired apical appendages of sternite 9, caudal view L adminiculum and paired apical appendages of sternite 9, lateral view. Abbreviations: adm-adminiculum; as 8 - paired appendages of sternite 8; as 9 - apical appendages of sternite 9; ig-inner gonostylus; gcx-gonocoxite; og-outer gonostylus; s 8 - sternite 8; s 9 - sternite 9; t 9 - tergite 9. Scale bar: 1 mm." figureDoi="10.3897/zookeys.1048.67564.figure2" httpUri="https://binary.pensoft.net/fig/566411" pageId="0" pageNumber="145">2E</figureCitation>
). Outer gonostylus (Figs 
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. T. (Lunatipula) caucasica A terminal segments of male abdomen, lateral view (dry) B-L male terminalia (after KOH 10 % treatment) B hypopygium, caudal view C hypopygium, lateral view D sternite 8, ventral view E tergite 9, caudal view F tergite 9, dorsal view G inner and outer gonostylus, lateral view H inner and outer gonostylus, caudal view I gonocoxal fragment, dorsal view J semen pump and aedeagus, lateral view K adminiculum and paired apical appendages of sternite 9, caudal view L adminiculum and paired apical appendages of sternite 9, lateral view. Abbreviations: adm-adminiculum; as 8 - paired appendages of sternite 8; as 9 - apical appendages of sternite 9; ig-inner gonostylus; gcx-gonocoxite; og-outer gonostylus; s 8 - sternite 8; s 9 - sternite 9; t 9 - tergite 9. Scale bar: 1 mm." figureDoi="10.3897/zookeys.1048.67564.figure2" httpUri="https://binary.pensoft.net/fig/566411" pageId="0" pageNumber="145">2G</figureCitation>
, 
<figureCitation captionStart="Figure 14" captionStartId="F14" captionText="Figure 14. Comparisons of some components of male terminalia of the Tipula caucasica species group A, F T. (Lunatipula) caucasica B, G T. (Lunatipula) eleniya sp. nov. C, I T. (Lunatipula) paupera D, J T. (Lunatipula) quadridentata E, K T. (Lunatipula) talyshensis A-E tergite 9, dorsal view F-K inner and outer gonostylus, lateral view. Arrows point to central notch at apex of tergite 9 (C, D), to the wedge-shaped projection posteriorly on inner gonostylus (F), to the rod-like outgrowth behind anterior beak of inner gonostylus (G), to the anterior (dorsal) margin of gonocoxite (I, J, K). Scale bar: 1 mm." figureDoi="10.3897/zookeys.1048.67564.figure14" httpUri="https://binary.pensoft.net/fig/566423" pageId="0" pageNumber="145">14F</figureCitation>
) very small, wineglass shaped, elongated distally. Inner gonostylus posteriorly, with wedge-shaped projection (Figs 
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. T. (Lunatipula) caucasica A terminal segments of male abdomen, lateral view (dry) B-L male terminalia (after KOH 10 % treatment) B hypopygium, caudal view C hypopygium, lateral view D sternite 8, ventral view E tergite 9, caudal view F tergite 9, dorsal view G inner and outer gonostylus, lateral view H inner and outer gonostylus, caudal view I gonocoxal fragment, dorsal view J semen pump and aedeagus, lateral view K adminiculum and paired apical appendages of sternite 9, caudal view L adminiculum and paired apical appendages of sternite 9, lateral view. Abbreviations: adm-adminiculum; as 8 - paired appendages of sternite 8; as 9 - apical appendages of sternite 9; ig-inner gonostylus; gcx-gonocoxite; og-outer gonostylus; s 8 - sternite 8; s 9 - sternite 9; t 9 - tergite 9. Scale bar: 1 mm." figureDoi="10.3897/zookeys.1048.67564.figure2" httpUri="https://binary.pensoft.net/fig/566411" pageId="0" pageNumber="145">2G</figureCitation>
, 
<figureCitation captionStart="Figure 14" captionStartId="F14" captionText="Figure 14. Comparisons of some components of male terminalia of the Tipula caucasica species group A, F T. (Lunatipula) caucasica B, G T. (Lunatipula) eleniya sp. nov. C, I T. (Lunatipula) paupera D, J T. (Lunatipula) quadridentata E, K T. (Lunatipula) talyshensis A-E tergite 9, dorsal view F-K inner and outer gonostylus, lateral view. Arrows point to central notch at apex of tergite 9 (C, D), to the wedge-shaped projection posteriorly on inner gonostylus (F), to the rod-like outgrowth behind anterior beak of inner gonostylus (G), to the anterior (dorsal) margin of gonocoxite (I, J, K). Scale bar: 1 mm." figureDoi="10.3897/zookeys.1048.67564.figure14" httpUri="https://binary.pensoft.net/fig/566423" pageId="0" pageNumber="145">14F</figureCitation>
). Paired appendage of sternite 8 (Fig. 
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. T. (Lunatipula) caucasica A terminal segments of male abdomen, lateral view (dry) B-L male terminalia (after KOH 10 % treatment) B hypopygium, caudal view C hypopygium, lateral view D sternite 8, ventral view E tergite 9, caudal view F tergite 9, dorsal view G inner and outer gonostylus, lateral view H inner and outer gonostylus, caudal view I gonocoxal fragment, dorsal view J semen pump and aedeagus, lateral view K adminiculum and paired apical appendages of sternite 9, caudal view L adminiculum and paired apical appendages of sternite 9, lateral view. Abbreviations: adm-adminiculum; as 8 - paired appendages of sternite 8; as 9 - apical appendages of sternite 9; ig-inner gonostylus; gcx-gonocoxite; og-outer gonostylus; s 8 - sternite 8; s 9 - sternite 9; t 9 - tergite 9. Scale bar: 1 mm." figureDoi="10.3897/zookeys.1048.67564.figure2" httpUri="https://binary.pensoft.net/fig/566411" pageId="0" pageNumber="145">2D</figureCitation>
) with base ~ 2 
<normalizedToken originalValue="×">x</normalizedToken>
as long as wide, with thin rigid yellow setae along edge, not obscuring gap in middle; long, glabrous red-brown spine arising from base, bent, and intersecting medially. Paired appendage of sternite 9 (Fig. 
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. T. (Lunatipula) caucasica A terminal segments of male abdomen, lateral view (dry) B-L male terminalia (after KOH 10 % treatment) B hypopygium, caudal view C hypopygium, lateral view D sternite 8, ventral view E tergite 9, caudal view F tergite 9, dorsal view G inner and outer gonostylus, lateral view H inner and outer gonostylus, caudal view I gonocoxal fragment, dorsal view J semen pump and aedeagus, lateral view K adminiculum and paired apical appendages of sternite 9, caudal view L adminiculum and paired apical appendages of sternite 9, lateral view. Abbreviations: adm-adminiculum; as 8 - paired appendages of sternite 8; as 9 - apical appendages of sternite 9; ig-inner gonostylus; gcx-gonocoxite; og-outer gonostylus; s 8 - sternite 8; s 9 - sternite 9; t 9 - tergite 9. Scale bar: 1 mm." figureDoi="10.3897/zookeys.1048.67564.figure2" httpUri="https://binary.pensoft.net/fig/566411" pageId="0" pageNumber="145">2K, L</figureCitation>
) with thick pubescent apex. Gonocoxal fragment (Fig. 
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. T. (Lunatipula) caucasica A terminal segments of male abdomen, lateral view (dry) B-L male terminalia (after KOH 10 % treatment) B hypopygium, caudal view C hypopygium, lateral view D sternite 8, ventral view E tergite 9, caudal view F tergite 9, dorsal view G inner and outer gonostylus, lateral view H inner and outer gonostylus, caudal view I gonocoxal fragment, dorsal view J semen pump and aedeagus, lateral view K adminiculum and paired apical appendages of sternite 9, caudal view L adminiculum and paired apical appendages of sternite 9, lateral view. Abbreviations: adm-adminiculum; as 8 - paired appendages of sternite 8; as 9 - apical appendages of sternite 9; ig-inner gonostylus; gcx-gonocoxite; og-outer gonostylus; s 8 - sternite 8; s 9 - sternite 9; t 9 - tergite 9. Scale bar: 1 mm." figureDoi="10.3897/zookeys.1048.67564.figure2" httpUri="https://binary.pensoft.net/fig/566411" pageId="0" pageNumber="145">2I</figureCitation>
) flattened at the base. Semen pump and aedeagus as in Figure 
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. T. (Lunatipula) caucasica A terminal segments of male abdomen, lateral view (dry) B-L male terminalia (after KOH 10 % treatment) B hypopygium, caudal view C hypopygium, lateral view D sternite 8, ventral view E tergite 9, caudal view F tergite 9, dorsal view G inner and outer gonostylus, lateral view H inner and outer gonostylus, caudal view I gonocoxal fragment, dorsal view J semen pump and aedeagus, lateral view K adminiculum and paired apical appendages of sternite 9, caudal view L adminiculum and paired apical appendages of sternite 9, lateral view. Abbreviations: adm-adminiculum; as 8 - paired appendages of sternite 8; as 9 - apical appendages of sternite 9; ig-inner gonostylus; gcx-gonocoxite; og-outer gonostylus; s 8 - sternite 8; s 9 - sternite 9; t 9 - tergite 9. Scale bar: 1 mm." figureDoi="10.3897/zookeys.1048.67564.figure2" httpUri="https://binary.pensoft.net/fig/566411" pageId="0" pageNumber="145">2J</figureCitation>
.
</paragraph>
<paragraph pageId="0" pageNumber="145">
<emphasis bold="true" pageId="0" pageNumber="145">Female.</emphasis>
Body length 17-21.5 mm, wing 15.5-18 mm. Antennae with scape and pedicel brownish yellow. Legs stocky, black to brown, yellow at base, femora grey. Wings with noticeable bluish tint; whitish area (lunule) of wings barely reaching base of discal cell. Female terminalia (Fig. 
<figureCitation captionStart="Figure 12" captionStartId="F12" captionText="Figure 12. Female terminalia of Tipula caucasica species group A-C T. (Lunatipula) caucasica D-F T. (Lunatipula) quadridentata G-I T. (Lunatipula) talyshensis A, D, G ovipositor, lateral view B, E, H ovipositor ventral view C, F, I sternite 9 and furca, lateral view. Abbreviations: c-cerci; f-furca; hv-hypogynial valve; s-sternite; t-tergite. Scale bar: 1 mm." figureDoi="10.3897/zookeys.1048.67564.figure12" httpUri="https://binary.pensoft.net/fig/566421" pageId="0" pageNumber="145">12A-C</figureCitation>
) comparatively short. Tip of cercus upcurved, length of cercus ~ 0.8 mm. Hypogynial valve only slightly longer than width at base; length of hypogynial valve 0.2 mm. Sternite 9 and furca as in Figure 
<figureCitation captionStart="Figure 12" captionStartId="F12" captionText="Figure 12. Female terminalia of Tipula caucasica species group A-C T. (Lunatipula) caucasica D-F T. (Lunatipula) quadridentata G-I T. (Lunatipula) talyshensis A, D, G ovipositor, lateral view B, E, H ovipositor ventral view C, F, I sternite 9 and furca, lateral view. Abbreviations: c-cerci; f-furca; hv-hypogynial valve; s-sternite; t-tergite. Scale bar: 1 mm." figureDoi="10.3897/zookeys.1048.67564.figure12" httpUri="https://binary.pensoft.net/fig/566421" pageId="0" pageNumber="145">12A, C</figureCitation>
.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="145" type="comparisons">
<paragraph pageId="0" pageNumber="145">Comparison with closely related species.</paragraph>
<paragraph pageId="0" pageNumber="145">
This species differs from other species of the 
<taxonomicName authorityName="Riedel" authorityYear="1920" class="Insecta" family="Tipulidae" genus="Tipula" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tipula caucasica" order="Diptera" pageId="0" pageNumber="145" phylum="Arthropoda" rank="species" species="caucasica">
<emphasis italics="true" pageId="0" pageNumber="145">Tipula caucasica</emphasis>
</taxonomicName>
group primarily by the presence of a large, anteriorly directed sharp wedge-shaped projection located posteriorly on the inner gonostylus.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="145" type="elevation">
<paragraph pageId="0" pageNumber="145">Elevation.</paragraph>
<paragraph pageId="0" pageNumber="145">Adults were collected at altitudes ranging from sea level (Black Sea coast at 35 m) to 2011 m (in Dagestan).</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="145" type="flight period">
<paragraph pageId="0" pageNumber="145">Flight period.</paragraph>
<paragraph pageId="0" pageNumber="145">From end of April to the beginning of July.</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="145" type="habitat">
<paragraph pageId="0" pageNumber="145">Habitat.</paragraph>
<paragraph pageId="0" pageNumber="145">
Specimens were captured in diverse forest habitats (see above) predominantly in mesophytic moderately moist deciduous or mixed communities with 
<taxonomicName authorityName="Lipsky" authorityYear="1898" class="Dicotyledoneae" family="Fagaceae" genus="Fagus" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="Fagus orientalis" order="Fagales" pageId="0" pageNumber="145" phylum="Angiospermae" rank="species" species="orientalis">
<emphasis italics="true" pageId="0" pageNumber="145">Fagus orientalis</emphasis>
</taxonomicName>
, 
<taxonomicName class="Pinopsida" family="Taxaceae" genus="Taxus" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="Taxus baccata" order="Taxales" pageId="0" pageNumber="145" phylum="Coniferophyta" rank="species" species="baccata">
<emphasis italics="true" pageId="0" pageNumber="145">Taxus baccata</emphasis>
</taxonomicName>
, 
<taxonomicName authorityName="Mill" authorityYear="1768" class="Dicotyledoneae" family="Betulaceae" genus="Carpinus" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="Carpinus orientalis" order="Fagales" pageId="0" pageNumber="145" phylum="Angiospermae" rank="species" species="orientalis">
<emphasis italics="true" pageId="0" pageNumber="145">Carpinus orientalis</emphasis>
</taxonomicName>
, 
<taxonomicName class="Dicotyledoneae" family="Fagaceae" genus="Quercus" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="Quercus iberica" order="Fagales" pageId="0" pageNumber="145" phylum="Angiospermae" rank="species" species="iberica">
<emphasis italics="true" pageId="0" pageNumber="145">Quercus iberica</emphasis>
</taxonomicName>
, 
<taxonomicName class="Dicotyledoneae" family="Salicaceae" genus="Populus" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="Populus tremula" order="Salicales" pageId="0" pageNumber="145" phylum="Angiospermae" rank="species" species="tremula">
<emphasis italics="true" pageId="0" pageNumber="145">Populus tremula</emphasis>
</taxonomicName>
, 
<taxonomicName class="Dicotyledoneae" family="Aceraceae" genus="Acer" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="Acer platanoides" order="Sapindales" pageId="0" pageNumber="145" phylum="Angiospermae" rank="species" species="platanoides">
<emphasis italics="true" pageId="0" pageNumber="145">Acer platanoides</emphasis>
</taxonomicName>
, etc., sometimes in communities near springs with 
<taxonomicName class="Ascidiacea" family="Polycitoridae" genus="Salix" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Salix caprea" order="Aplousobranchia" pageId="0" pageNumber="145" phylum="Chordata" rank="species" species="caprea">
<emphasis italics="true" pageId="0" pageNumber="145">Salix caprea</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="145" type="distribution">
<paragraph pageId="0" pageNumber="145">Distribution.</paragraph>
<paragraph pageId="0" pageNumber="145">
Endemic to the Caucasus. It was recently noted for Russia (Dagestan) for the first time (
<bibRefCitation DOI="https://doi.org/10.47640/978-5-00105-586-0_2020_292" author="Lantsov, VI" journalOrPublisher="Fauna of the Stavropol region" pageId="0" pageNumber="145" refId="B7" refString="Lantsov, VI, 2020. Biological diversity of crane flies (Diptera, Tipuloidea) in the natural landscapes of Dagestan (North-East Caucasus, Russia) // XI All-Russian Dipterological Symposium (with international participation). Voronezh. August 24-29, 2020. Proceedings of conference / Responsible editors: O.G.Ovchinnikova, I.V.Shamshev. St.Petersburg: Russian Entomological Society, LEMA, 292-296. https://doi.org/10.47640/978-5-00105-586-0_2020_292" title="Biological diversity of crane flies (Diptera, Tipuloidea) in the natural landscapes of Dagestan (North-East Caucasus, Russia) // XI All-Russian Dipterological Symposium (with international participation). Voronezh. August 24 - 29, 2020. Proceedings of conference / Responsible editors: O. G. Ovchinnikova, I. V. Shamshev. St. Petersburg: Russian Entomological Society, LEMA, 292 - 296." url="https://doi.org/10.47640/978-5-00105-586-0_2020_292" year="2020">Lantsov 2020</bibRefCitation>
). Currently known from the West Caucasus (first record), East Caucasus and Transcaucasus.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="145" type="remarks">
<paragraph pageId="0" pageNumber="145">Remarks.</paragraph>
<paragraph pageId="0" pageNumber="145">
According to 
<bibRefCitation author="Riedel, MP" journalOrPublisher="Zoologische Jahrbucher, Abteilung fur Systematik, Geographie und Biologie der Tiere" pageId="0" pageNumber="145" pagination="13 - 22" refId="B9" refString="Riedel, MP, 1920. Nematocera Polyneura (Dipt.) aus dem Kaukasus. Zoologische Jahrbucher, Abteilung fur Systematik, Geographie und Biologie der Tiere 43: 13 - 22" title="Nematocera Polyneura (Dipt.) aus dem Kaukasus." volume="43" year="1920">Riedel (1920)</bibRefCitation>
, a series of type specimens, four males, two females. 
<normalizedToken originalValue="“Mts’chet">&quot;Mts'chet</normalizedToken>
prope Tiflis [Georgia, Tbilisi], 6. 5. 1913 (Mus. Caucas. Ph. Zaitzev)&quot; were deposited in the &quot;Museum Caucasicum in Tiflis&quot;. In the 
<normalizedToken originalValue="author’s">author's</normalizedToken>
publication, neither the holotype nor the paratypes are indicated. Presumably, they are not indicated on the material itself. Because this material turned out to be inaccessible to the authors of this article - it is not yet known whether this material has survived at all - there was no opportunity to designate lectotype and paralectotypes.
</paragraph>
</subSubSection>
</treatment>
</document>