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<document id="1B8D633596C871796136C2FD622A5609" ID-CLB-Dataset="21234" ID-DOI="10.11646/zootaxa.4560.3.12" ID-GBIF-Dataset="e1c7a0c0-f975-4b96-9bb4-8bef87d84860" ID-ISSN="1175-5326" ID-Zenodo-Dep="3993466" ID-ZooBank="FC4CC45F-8261-4FFA-82EC-A27AC0301944" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1597949354476" checkinUser="felipe" docAuthor="Figueroa, Luis &amp; Neita-Moreno, Jhon César" docDate="2019" docId="1510E54A6B500A19FF1AFAABFE8496AD" docLanguage="en" docName="zootaxa.4560.3.12.pdf" docOrigin="Zootaxa 4560 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Compsodactylus vallejoi Figueroa &amp; Neita-Moreno 2019, new species" docType="treatment" docVersion="8" lastPageNumber="591" masterDocId="E9299D326B500A1DFF8DFF97FF98967C" masterDocTitle="A new Compsodactylus Fuhrmann (Coleoptera: Scarabaeidae: Melolonthinae) from Peru" masterLastPageNumber="591" masterPageNumber="587" pageNumber="587" updateTime="1698844952502" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
<mods:mods id="AE5FDDBE749E625F663C6F6621067E9D" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="29A219901318B248703ADF00BAC48C37">
<mods:title id="69815FFDA5EC87FBB6B6E7C3B72707A5">A new Compsodactylus Fuhrmann (Coleoptera: Scarabaeidae: Melolonthinae) from Peru</mods:title>
</mods:titleInfo>
<mods:name id="753E9E8BB7617BD9C510C487B6089767" type="personal">
<mods:role id="1F7E6EF6213E2F704C2AB361E2EFB1B5">
<mods:roleTerm id="DA4FBC5D2159F050BFFCD1205750B0D9">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="162AAC163B2EC57802FD7E8B987D0C70">Figueroa, Luis</mods:namePart>
<mods:affiliation id="6278FEF4A9071EE9F499CE4A43619626">Departamento de Entomología, Museo de Historia Natural, Universidad Nacional Mayor de San Marcos, Av. Arenales 1256 Jesús María, Lima 14, Perú. &amp; Bosque Llaqta, Av. Confraternidad Internacional Este Nº 364, Huaraz, Perú</mods:affiliation>
</mods:name>
<mods:name id="0D5F35F3189EFF337302B49045A8FCF3" type="personal">
<mods:role id="2FF463A15045DF054428F3F3B8E98603">
<mods:roleTerm id="EB09B9AC22FFBB2C64A60D1191E5DEB8">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="1826906ECB10B56A6EDAC8DCDB0A2781">Neita-Moreno, Jhon César</mods:namePart>
<mods:affiliation id="D97CDC6D5BEF712527944FBC1EB9DB72">Instituto de Investigaciones de Recursos Biológicos Alexander von Humboldt. Claustro de San Agustín, Villa de Leyva, Boyacá, Colombia</mods:affiliation>
</mods:name>
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<mods:title id="A511160F91A7364C9DFC5D6D9188EC45">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="5F91AA4D626AC99E04C75608BB1F7C87">
<mods:date id="6034EB27358323E518E136D32D43E837">2019</mods:date>
<mods:detail id="7C81F24F335C80E98E26DF5DB1FBF122" type="pubDate">
<mods:number id="1421A7B968989F479B4CAB4A4A150FB1">2019-02-26</mods:number>
</mods:detail>
<mods:detail id="6140E8627ECF85075C9EAFF330E06417" type="volume">
<mods:number id="A2892ADCC1E2C5EDDD3E3FF6D52A3580">4560</mods:number>
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<mods:number id="5D8437C1900D9B5E3C324E7A9CB50556">3</mods:number>
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<mods:start id="113F2432F45B9C52C7E9E1668076D8CE">587</mods:start>
<mods:end id="495E701D0EB3617999E89210069CDB01">591</mods:end>
</mods:extent>
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<mods:classification id="27F16737C4DFA89B58A9CF0A0ACD0B6A">journal article</mods:classification>
<mods:identifier id="61C03F7AC1EF52176022EDC7D0E4F446" type="CLB-Dataset">21234</mods:identifier>
<mods:identifier id="DCEE58DEB30C7CD3F3F173B4BF6996A1" type="DOI">10.11646/zootaxa.4560.3.12</mods:identifier>
<mods:identifier id="71A5E61F1D7FC28C952ECE7EB626CE74" type="GBIF-Dataset">e1c7a0c0-f975-4b96-9bb4-8bef87d84860</mods:identifier>
<mods:identifier id="B405B425739634D3049DAC49AC88FEE5" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="025F2F4D038680CB92DC85478A7A7BEA" type="Zenodo-Dep">3993466</mods:identifier>
<mods:identifier id="A9FCAC0FE4AA8FE52B89F2480BC8DAB6" type="ZooBank">FC4CC45F-8261-4FFA-82EC-A27AC0301944</mods:identifier>
</mods:mods>
<treatment id="1510E54A6B500A19FF1AFAABFE8496AD" ID-DOI="http://doi.org/10.5281/zenodo.4323786" ID-GBIF-Taxon="167284737" ID-Zenodo-Dep="4323786" LSID="urn:lsid:plazi:treatment:1510E54A6B500A19FF1AFAABFE8496AD" httpUri="http://treatment.plazi.org/id/1510E54A6B500A19FF1AFAABFE8496AD" lastPageId="4" lastPageNumber="591" pageId="0" pageNumber="587">
<subSubSection id="D5A307D76B500A1DFF1AFAABFAB1932A" box="[151,1321,1340,1366]" pageId="0" pageNumber="587" type="nomenclature">
<paragraph id="9D06545C6B500A1DFF1AFAABFAB1932A" blockId="0.[151,1436,1340,2027]" box="[151,1321,1340,1366]" pageId="0" pageNumber="587">
<heading id="C64EE3306B500A1DFF1AFAABFAB1932A" bold="true" box="[151,1321,1340,1366]" fontSize="11" level="1" pageId="0" pageNumber="587" reason="1">
<emphasis id="AFCD884E6B500A1DFF1AFAABFAB1932A" bold="true" box="[151,1321,1340,1366]" pageId="0" pageNumber="587">
<taxonomicName id="5AB92FDF6B500A1DFF1AFAABFC92932A" authority="Figueroa &amp; Neita-Moreno" authorityName="Figueroa &amp; Neita-Moreno" authorityYear="2019" box="[151,778,1340,1366]" class="Insecta" family="Melolonthidae" genus="Compsodactylus" kingdom="Animalia" order="Coleoptera" pageId="0" pageNumber="587" phylum="Arthropoda" rank="species" species="vallejoi" status="sp. nov.">
<emphasis id="AFCD884E6B500A1DFF1AFAABFE23932A" bold="true" box="[151,443,1340,1366]" italics="true" pageId="0" pageNumber="587">Compsodactylus vallejoi</emphasis>
Figueroa &amp; Neita-Moreno
</taxonomicName>
,
<taxonomicNameLabel id="B4FE35356B500A1DFC9AFAABFC3F932A" box="[791,935,1340,1366]" pageId="0" pageNumber="587" rank="species">new species</taxonomicNameLabel>
(
<figureCitation id="058248D96B500A1DFC3AFAABFBB1932A" box="[951,1065,1340,1366]" captionStart="FIGURES 1–2" captionStartId="1.[151,258,2008,2030]" captionTargetBox="[229,1378,905,1946]" captionTargetId="figure@1.[151,1436,853,1984]" captionTargetPageId="1" captionText="FIGURES 1–2. Compsodactylus vallejoi holotype. 1, Dorsal habitus; 2, lateral habitus." figureDoi="http://doi.org/10.5281/zenodo.3993468" httpUri="https://zenodo.org/record/3993468/files/figure.png" pageId="0" pageNumber="587">Figs. 1–2</figureCitation>
,
<figureCitation id="058248D96B500A1DFBBBFAABFB52932A" box="[1078,1226,1340,1366]" captionStart="FIGURES 3–10" captionStartId="2.[151,258,1937,1959]" captionTargetBox="[158,1432,229,1880]" captionTargetId="figure@2.[151,1435,229,1891]" captionTargetPageId="2" captionText="FIGURES 3–10. Compsodactylus martinezi and C. vallejoi leg characters. 3, Compsodactylus martinezi prothoracic legs (dorsal); 4, C. vallejoi prothoracic legs (dorsal); 5, C. vallejoi prothoracic legs (dorsal); 6, C. martinezi metafemur (dorsal); 7, C. vallejoi metafemur (dorsal); 8, C. vallejoi metafemur (dorsal); 9, C. martinezi apex of metatibia; 10, C. vallejoi apex of metatibia." figureDoi="http://doi.org/10.5281/zenodo.3993470" httpUri="https://zenodo.org/record/3993470/files/figure.png" pageId="0" pageNumber="587">4–5, 7–8, 10</figureCitation>
,
<figureCitation id="058248D96B500A1DFB5AFAABFAB9932A" box="[1239,1313,1340,1366]" captionStart-0="FIGURES 11–16" captionStart-1="FIGURES 17–19" captionStartId-0="3.[151,258,1171,1193]" captionStartId-1="3.[151,258,1987,2009]" captionTargetBox-0="[267,1329,240,1109]" captionTargetBox-1="[261,1328,1295,1965]" captionTargetId-0="figure@3.[248,1346,191,1157]" captionTargetId-1="figure@3.[261,1328,1295,1965]" captionTargetPageId-0="3" captionTargetPageId-1="3" captionText-0="FIGURES 11–16. Compsodactylus martinezi and C. vallejoi genitalic characters. 11, Compsodactylus martinezi phallobase and parameres (lateral); 12, C. martinezi parameres (frontal); 13, C. martinezi female genital plates; 14, C. vallejoi phallobase and parameres (lateral); 15, C. vallejoi parameres (frontal); 16, C. vallejoi female genital plates." captionText-1="FIGURES 17–19. Compsodactylus vallejoi. 17, distribution; 18–19, habitat." figureDoi-0="http://doi.org/10.5281/zenodo.3993472" figureDoi-1="http://doi.org/10.5281/zenodo.3993474" httpUri-0="https://zenodo.org/record/3993472/files/figure.png" httpUri-1="https://zenodo.org/record/3993474/files/figure.png" pageId="0" pageNumber="587">14–19</figureCitation>
)
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="D5A307D76B500A1DFF1AFAC8FDE59012" pageId="0" pageNumber="587" type="materials_examined">
<paragraph id="9D06545C6B500A1DFF1AFAC8FDE59012" blockId="0.[151,1436,1340,2027]" pageId="0" pageNumber="587">
<emphasis id="AFCD884E6B500A1DFF1AFAC8FEA5930A" box="[151,317,1375,1398]" inLineHeading="true" pageId="0" pageNumber="587" reason="1">
<emphasis id="AFCD884E6B500A1DFF1AFAC8FEAD930A" bold="true" box="[151,309,1375,1398]" pageId="0" pageNumber="587">Type material</emphasis>
.
</emphasis>
<materialsCitation id="2DD15E016B500A1DFECEFAC8FA0493E7" ID-GBIF-Occurrence="2846548301" collectionCode="MUSM" collectorName="P. Ancajima" country="Peru" elevation="1785" latitude="-7.7770023" location="Museo de Historia Natural de la Universidad Nacional Mayor" municipality="Historia Natural de la Universidad Nacional Mayor de San Marcos" pageId="0" pageNumber="587" specimenCount="8" stateProvince="La Libertad" typeStatus="holotype">
<typeStatus id="4202EAFE6B500A1DFECEFAC8FE3E930A" box="[323,422,1375,1398]" pageId="0" pageNumber="587" type="holotype">Holotype</typeStatus>
and allotype labeled: “
<collectingCountry id="E5AE14CC6B500A1DFD11FAC8FD7E930A" box="[668,742,1375,1398]" name="Peru" pageId="0" pageNumber="587">PERÚ</collectingCountry>
. LL. [
<collectingRegion id="5F7D9ABE6B500A1DFCA9FAC8FC3C930A" box="[804,932,1375,1398]" country="Peru" name="La Libertad" pageId="0" pageNumber="587">La Libertad</collectingRegion>
Departament] Pataz, // Unidad Minera Santa // María. Rio Hualanga //
<geoCoordinate id="F88D329B6B500A1DFE2DFA13FDB393E7" box="[416,555,1410,1435]" degrees="7" direction="south" minutes="46" orientation="latitude" pageId="0" pageNumber="587" precision="1" seconds="37.21" value="-7.7770023">
7
<superScript id="6ACCF9146B500A1DFE20FA15FE2C93F2" attach="none" box="[429,436,1410,1422]" fontSize="5" pageId="0" pageNumber="587">o</superScript>
46’37.21”S
</geoCoordinate>
/
<geoCoordinate id="F88D329B6B500A1DFDB8FA13FD6793E7" box="[565,767,1410,1435]" pageId="0" pageNumber="587">
77
<superScript id="6ACCF9146B500A1DFDDCFA15FDC093F2" attach="none" box="[593,600,1410,1422]" fontSize="5" pageId="0" pageNumber="587">o</superScript>
36’46.3 // 5”W
</geoCoordinate>
<quantity id="5A41F9B96B500A1DFC84FA13FCCB93E7" box="[777,851,1412,1435]" metricMagnitude="3" metricUnit="m" metricValue="1.785" pageId="0" pageNumber="587" unit="m" value="1785.0">
<elevation id="1694B36F6B500A1DFC84FA13FCCB93E7" box="[777,851,1412,1435]" metricMagnitude="3" metricUnit="m" metricValue="1.785" pageId="0" pageNumber="587" unit="m" value="1785.0">1785m</elevation>
</quantity>
07.iii.17[2017] //
<collectorName id="304C318A6B500A1DFB92FA13FB3D93E7" box="[1055,1189,1412,1435]" pageId="0" pageNumber="587">P. Ancajima</collectorName>
” both with red labels.
</materialsCitation>
<materialsCitation id="2DD15E016B500A1DFF1AFA30FAFB9054" ID-GBIF-Occurrence="2846548302" collectionCode="MUSM" collectorName="P. Ancajima" country="Peru" elevation="1785" latitude="-7.7770023" location="Museo de Historia Natural de la Universidad Nacional Mayor" municipality="Historia Natural de la Universidad Nacional Mayor de San Marcos" pageId="0" pageNumber="587" specimenCount="8" stateProvince="La Libertad" typeStatus="holotype">
Three male paratypes with same data as the holotype, and one female paratype labeled: “
<collectingCountry id="E5AE14CC6B500A1DFB0AFA30FB4993C2" box="[1159,1233,1447,1470]" name="Peru" pageId="0" pageNumber="587">PERÚ</collectingCountry>
. LL. [
<collectingRegion id="5F7D9ABE6B500A1DFA9AFA30FA0393C2" box="[1303,1435,1447,1470]" country="Peru" name="La Libertad" pageId="0" pageNumber="587">La Libertad</collectingRegion>
Departament] Pataz, // Unidad Minera Santa // María. Rio Hualanga //
<geoCoordinate id="F88D329B6B500A1DFC69FA5BFBE8939F" box="[996,1136,1482,1507]" degrees="7" direction="south" minutes="45" orientation="latitude" pageId="0" pageNumber="587" precision="1" seconds="45.23" value="-7.7625637">
7
<superScript id="6ACCF9146B500A1DFC7FFA5DFC6193AA" attach="right" box="[1010,1017,1482,1494]" fontSize="5" pageId="0" pageNumber="587">o</superScript>
45’45.23”S
</geoCoordinate>
/
<geoCoordinate id="F88D329B6B500A1DFBF4FA5BFADA939F" box="[1145,1346,1482,1507]" pageId="0" pageNumber="587">
77
<superScript id="6ACCF9146B500A1DFB18FA5DFB0493AA" attach="none" box="[1173,1180,1482,1494]" fontSize="5" pageId="0" pageNumber="587">o</superScript>
38’30.2// 4’’W
</geoCoordinate>
<quantity id="5A41F9B96B500A1DFADFFA5BFA04939F" box="[1362,1436,1484,1507]" metricMagnitude="3" metricUnit="m" metricValue="1.413" pageId="0" pageNumber="587" unit="m" value="1413.0">
<elevation id="1694B36F6B500A1DFADFFA5BFA04939F" box="[1362,1436,1484,1507]" metricMagnitude="3" metricUnit="m" metricValue="1.413" pageId="0" pageNumber="587" unit="m" value="1413.0">1413m</elevation>
</quantity>
04.iii.17[2017] //
<collectorName id="304C318A6B500A1DFEDFFA78FE40907A" box="[338,472,1519,1542]" pageId="0" pageNumber="587">P. Ancajima</collectorName>
”; paratypes with yellow labels. The holotype, allotype, and two paratypes (both males) are deposited at the
<location id="986602876B500A1DFECEF986FC499054" LSID="urn:lsid:plazi:treatment:1510E54A6B500A19FF1AFAABFE8496AD:986602876B500A1DFECEF986FC499054" box="[323,977,1553,1576]" country="Peru" latitude="-7.7770023" municipality="Historia Natural de la Universidad Nacional Mayor de San Marcos" name="Museo de Historia Natural de la Universidad Nacional Mayor" pageId="0" pageNumber="587" stateProvince="La Libertad">
Museo de
<collectingMunicipality id="7D62CE266B500A1DFE3CF986FC499054" box="[433,977,1553,1576]" pageId="0" pageNumber="587">Historia Natural de la Universidad Nacional Mayor</collectingMunicipality>
</location>
de San Marcos (
<collectionCode id="FBA8CC996B500A1DFB0CF986FB409054" box="[1153,1240,1553,1576]" pageId="0" pageNumber="587">MUSM</collectionCode>
),
<collectingRegion id="5F7D9ABE6B500A1DFB67F986FABD9054" box="[1258,1317,1553,1576]" country="Peru" name="Lima" pageId="0" pageNumber="587">Lima</collectingRegion>
, Peru
</materialsCitation>
.
<materialsCitation id="2DD15E016B500A1DFAE1F986FDE09012" ID-GBIF-Occurrence="2846548303" collectionCode="SEMC" collectorName="Two &amp; Snow Entomological Collection &amp; University" country="United States of America" location="Kansas" pageId="0" pageNumber="587" specimenCount="1" stateProvince="Kansas" typeStatus="holotype">
<collectorName id="304C318A6B500A1DFAE1F986FA039054" box="[1388,1435,1553,1576]" pageId="0" pageNumber="587">Two</collectorName>
paratypes (male and female) are deposited in the
<collectorName id="304C318A6B500A1DFD43F9A3FBAA9037" box="[718,1074,1588,1611]" pageId="0" pageNumber="587">Snow Entomological Collection</collectorName>
,
<collectorName id="304C318A6B500A1DFBCFF9A3FB2B9037" box="[1090,1203,1588,1611]" pageId="0" pageNumber="587">University</collectorName>
of
<collectingRegion id="5F7D9ABE6B500A1DFB6CF9A3FAB69037" box="[1249,1326,1588,1611]" country="United States of America" name="Kansas" pageId="0" pageNumber="587">Kansas</collectingRegion>
(
<collectionCode id="FBA8CC996B500A1DFACFF9A3FA089037" box="[1346,1424,1588,1611]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34264" name="University of Kansas - Biodiversity Institute" pageId="0" pageNumber="587" type="Museum">SEMC</collectionCode>
), Lawrence,
<collectingRegion id="5F7D9ABE6B500A1DFE80F9C0FEC59012" box="[269,349,1623,1646]" country="United States of America" name="Kansas" pageId="0" pageNumber="587">Kansas</collectingRegion>
,
<collectingCountry id="E5AE14CC6B500A1DFEEAF9C0FDE09012" box="[359,632,1623,1646]" name="United States of America" pageId="0" pageNumber="587">United States of America</collectingCountry>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="D5A307D76B500A1CFF4AF9EDFB649745" lastPageId="1" lastPageNumber="588" pageId="0" pageNumber="587" type="description">
<paragraph id="9D06545C6B500A1DFF4AF9EDFA0491B5" blockId="0.[151,1436,1340,2027]" pageId="0" pageNumber="587">
<emphasis id="AFCD884E6B500A1DFF4AF9EDFECA90ED" box="[199,338,1658,1681]" inLineHeading="true" pageId="0" pageNumber="587" reason="1">
<emphasis id="AFCD884E6B500A1DFF4AF9EDFED390ED" bold="true" box="[199,331,1658,1681]" pageId="0" pageNumber="587">Description</emphasis>
.
</emphasis>
<emphasis id="AFCD884E6B500A1DFED5F9EDFE5F90ED" bold="true" box="[344,455,1658,1681]" pageId="0" pageNumber="587">
<typeStatus id="4202EAFE6B500A1DFED5F9EDFE5A90ED" box="[344,450,1658,1681]" pageId="0" pageNumber="587" type="holotype">Holotype</typeStatus>
.
</emphasis>
Male (
<figureCitation id="058248D96B500A1DFD9EF9EDFDE290ED" box="[531,634,1658,1681]" captionStart="FIGURES 1–2" captionStartId="1.[151,258,2008,2030]" captionTargetBox="[229,1378,905,1946]" captionTargetId="figure@1.[151,1436,853,1984]" captionTargetPageId="1" captionText="FIGURES 1–2. Compsodactylus vallejoi holotype. 1, Dorsal habitus; 2, lateral habitus." figureDoi="http://doi.org/10.5281/zenodo.3993468" httpUri="https://zenodo.org/record/3993468/files/figure.png" pageId="0" pageNumber="587">Figs. 1–2</figureCitation>
). Length
<quantity id="5A41F9B96B500A1DFD6DF9EDFCAC90ED" box="[736,820,1658,1681]" metricMagnitude="-3" metricUnit="m" metricValue="6.9" pageId="0" pageNumber="587" unit="mm" value="6.9">6.9 mm</quantity>
, width
<quantity id="5A41F9B96B500A1DFC0FF9EDFC4C90ED" box="[898,980,1658,1681]" metricMagnitude="-3" metricUnit="m" metricValue="3.4" pageId="0" pageNumber="587" unit="mm" value="3.4">3.4 mm</quantity>
at middle of elytra. Head black; pronotum black with lateral margins and posterior third brownish testaceous (
<figureCitation id="058248D96B500A1DFC0FF90BFC7390CF" box="[898,1003,1692,1715]" captionStart="FIGURES 1–2" captionStartId="1.[151,258,2008,2030]" captionTargetBox="[229,1378,905,1946]" captionTargetId="figure@1.[151,1436,853,1984]" captionTargetPageId="1" captionText="FIGURES 1–2. Compsodactylus vallejoi holotype. 1, Dorsal habitus; 2, lateral habitus." figureDoi="http://doi.org/10.5281/zenodo.3993468" httpUri="https://zenodo.org/record/3993468/files/figure.png" pageId="0" pageNumber="587">Figs. 1–2</figureCitation>
); elytra testaceous with black margins; scutellum testaceous; femur brownish testaceous; protibia black, laterodistally transitioning to brown; mesotibia brownish black, dorsally transitioning to testaceous; metatibia brownish testaceus with dark maculae.
<emphasis id="AFCD884E6B500A1DFB65F975FAB69085" bold="true" box="[1256,1326,1762,1785]" pageId="0" pageNumber="587">Head:</emphasis>
frons and clypeus with denticle-like setae, epistomal suture indistinct, interocular distance about 5.7 times as wide as width of eye in dorsal view.
<emphasis id="AFCD884E6B500A1DFEB1F8B0FE209142" bold="true" box="[316,440,1831,1854]" pageId="0" pageNumber="587">Pronotum:</emphasis>
with long setae (
<figureCitation id="058248D96B500A1DFDFCF8B0FD299142" box="[625,689,1831,1854]" captionStart="FIGURES 1–2" captionStartId="1.[151,258,2008,2030]" captionTargetBox="[229,1378,905,1946]" captionTargetId="figure@1.[151,1436,853,1984]" captionTargetPageId="1" captionText="FIGURES 1–2. Compsodactylus vallejoi holotype. 1, Dorsal habitus; 2, lateral habitus." figureDoi="http://doi.org/10.5281/zenodo.3993468" httpUri="https://zenodo.org/record/3993468/files/figure.png" pageId="0" pageNumber="587">Fig. 1</figureCitation>
).
<emphasis id="AFCD884E6B500A1DFD45F8B0FC829142" bold="true" box="[712,794,1831,1854]" pageId="0" pageNumber="587">Elytra:</emphasis>
anterior margin of scutellum 0.4 times as wide as length of anterior margin of elytra, elytral striae distinctly punctate.
<emphasis id="AFCD884E6B500A1DFC8AF8DDFCA2911D" bold="true" box="[775,826,1866,1889]" pageId="0" pageNumber="587">Leg:</emphasis>
protibia bidentate; protarsomere 1 widened distally (
<figureCitation id="058248D96B500A1DFAFDF8DDFF3E91FF" captionStart="FIGURES 3–10" captionStartId="2.[151,258,1937,1959]" captionTargetBox="[158,1432,229,1880]" captionTargetId="figure@2.[151,1435,229,1891]" captionTargetPageId="2" captionText="FIGURES 3–10. Compsodactylus martinezi and C. vallejoi leg characters. 3, Compsodactylus martinezi prothoracic legs (dorsal); 4, C. vallejoi prothoracic legs (dorsal); 5, C. vallejoi prothoracic legs (dorsal); 6, C. martinezi metafemur (dorsal); 7, C. vallejoi metafemur (dorsal); 8, C. vallejoi metafemur (dorsal); 9, C. martinezi apex of metatibia; 10, C. vallejoi apex of metatibia." figureDoi="http://doi.org/10.5281/zenodo.3993470" httpUri="https://zenodo.org/record/3993470/files/figure.png" pageId="0" pageNumber="587">Fig. 7</figureCitation>
), 2.3 times as long as protarsomere II; protarsal claws with anterior tooth narrower and longer than internal tooth; metafemur enlarged with some long, erecte setae (
<figureCitation id="058248D96B500A1DFD6CF818FCB191DA" box="[737,809,1935,1958]" captionStart="FIGURES 3–10" captionStartId="2.[151,258,1937,1959]" captionTargetBox="[158,1432,229,1880]" captionTargetId="figure@2.[151,1435,229,1891]" captionTargetPageId="2" captionText="FIGURES 3–10. Compsodactylus martinezi and C. vallejoi leg characters. 3, Compsodactylus martinezi prothoracic legs (dorsal); 4, C. vallejoi prothoracic legs (dorsal); 5, C. vallejoi prothoracic legs (dorsal); 6, C. martinezi metafemur (dorsal); 7, C. vallejoi metafemur (dorsal); 8, C. vallejoi metafemur (dorsal); 9, C. martinezi apex of metatibia; 10, C. vallejoi apex of metatibia." figureDoi="http://doi.org/10.5281/zenodo.3993470" httpUri="https://zenodo.org/record/3993470/files/figure.png" pageId="0" pageNumber="587">Fig. 7</figureCitation>
); metatibial apex enlarged, distally forming an obtuse projection. Pygidium glabrous or sparsely setose, setae short and slender.
<emphasis id="AFCD884E6B500A1DFC2FF825FBBD91B5" bold="true" box="[930,1061,1970,1993]" pageId="0" pageNumber="587">Parameres:</emphasis>
simple and enlarged (
<figureCitation id="058248D96B500A1DFA83F825FA0891B5" box="[1294,1424,1970,1993]" captionStart="FIGURES 11–16" captionStartId="3.[151,258,1171,1193]" captionTargetBox="[267,1329,240,1109]" captionTargetId="figure@3.[248,1346,191,1157]" captionTargetPageId="3" captionText="FIGURES 11–16. Compsodactylus martinezi and C. vallejoi genitalic characters. 11, Compsodactylus martinezi phallobase and parameres (lateral); 12, C. martinezi parameres (frontal); 13, C. martinezi female genital plates; 14, C. vallejoi phallobase and parameres (lateral); 15, C. vallejoi parameres (frontal); 16, C. vallejoi female genital plates." figureDoi="http://doi.org/10.5281/zenodo.3993472" httpUri="https://zenodo.org/record/3993472/files/figure.png" pageId="0" pageNumber="587">Figs. 14–16</figureCitation>
).
</paragraph>
<paragraph id="9D06545C6B500A1CFF1AF843FF2C9688" blockId="0.[151,1436,1340,2027]" lastBlockId="1.[151,1436,152,799]" lastPageId="1" lastPageNumber="588" pageId="0" pageNumber="587">
<emphasis id="AFCD884E6B500A1DFF1AF843FF669197" bold="true" box="[151,254,2004,2027]" inLineHeading="true" pageId="0" pageNumber="587" reason="1">
<typeStatus id="4202EAFE6B500A1DFF1AF843FF619197" box="[151,249,2004,2027]" pageId="0" pageNumber="587" type="allotype">Allotype</typeStatus>
.
</emphasis>
Female. Length
<quantity id="5A41F9B96B500A1DFE3AF843FD959197" box="[439,525,2004,2027]" metricMagnitude="-3" metricUnit="m" metricValue="6.8" pageId="0" pageNumber="587" unit="mm" value="6.8">6.8 mm</quantity>
, width
<quantity id="5A41F9B96B500A1DFDD3F843FD299197" box="[606,689,2004,2027]" metricMagnitude="-3" metricUnit="m" metricValue="3.5" pageId="0" pageNumber="587" unit="mm" value="3.5">3.5 mm</quantity>
at middle of elytra. Coloration similar to
<typeStatus id="4202EAFE6B500A1DFBF7F843FB429197" box="[1146,1242,2004,2027]" pageId="0" pageNumber="587" type="holotype">holotype</typeStatus>
, except pronotum black and with only the posterior margin brownish testaceous; protarsomere 1 thin (
<figureCitation id="058248D96B510A1CFBD9FF0FFB0296D3" box="[1108,1178,152,175]" captionStart="FIGURES 3–10" captionStartId="2.[151,258,1937,1959]" captionTargetBox="[158,1432,229,1880]" captionTargetId="figure@2.[151,1435,229,1891]" captionTargetPageId="2" captionText="FIGURES 3–10. Compsodactylus martinezi and C. vallejoi leg characters. 3, Compsodactylus martinezi prothoracic legs (dorsal); 4, C. vallejoi prothoracic legs (dorsal); 5, C. vallejoi prothoracic legs (dorsal); 6, C. martinezi metafemur (dorsal); 7, C. vallejoi metafemur (dorsal); 8, C. vallejoi metafemur (dorsal); 9, C. martinezi apex of metatibia; 10, C. vallejoi apex of metatibia." figureDoi="http://doi.org/10.5281/zenodo.3993470" httpUri="https://zenodo.org/record/3993470/files/figure.png" pageId="1" pageNumber="588">Fig. 5</figureCitation>
), 1.3 times as long as protarsomere 2; metafemur only slightly enlarged without long, erect setae, and without enlarged metatibial apex (
<figureCitation id="058248D96B510A1CFAFDFF2DFF3E9688" captionStart="FIGURES 3–10" captionStartId="2.[151,258,1937,1959]" captionTargetBox="[158,1432,229,1880]" captionTargetId="figure@2.[151,1435,229,1891]" captionTargetPageId="2" captionText="FIGURES 3–10. Compsodactylus martinezi and C. vallejoi leg characters. 3, Compsodactylus martinezi prothoracic legs (dorsal); 4, C. vallejoi prothoracic legs (dorsal); 5, C. vallejoi prothoracic legs (dorsal); 6, C. martinezi metafemur (dorsal); 7, C. vallejoi metafemur (dorsal); 8, C. vallejoi metafemur (dorsal); 9, C. martinezi apex of metatibia; 10, C. vallejoi apex of metatibia." figureDoi="http://doi.org/10.5281/zenodo.3993470" httpUri="https://zenodo.org/record/3993470/files/figure.png" pageId="1" pageNumber="588">Fig. 8</figureCitation>
).
</paragraph>
<paragraph id="9D06545C6B510A1CFF4AFE97FB649745" blockId="1.[151,1436,152,799]" pageId="1" pageNumber="588">
<emphasis id="AFCD884E6B510A1CFF4AFE97FEA1976B" bold="true" box="[199,313,256,279]" inLineHeading="true" pageId="1" pageNumber="588" reason="1">Variation.</emphasis>
Males: length
<quantity id="5A41F9B96B510A1CFE5AFE97FDC2976B" box="[471,602,256,279]" metricMagnitude="-3" metricUnit="m" metricValue="6.1" metricValueMax="6.4" metricValueMin="5.8" pageId="1" pageNumber="588" unit="mm" value="6.1" valueMax="6.4" valueMin="5.8">5.8–6.4 mm</quantity>
, width at middle of elytra 3.0–
<quantity id="5A41F9B96B510A1CFC2CFE97FC6D976B" box="[929,1013,256,279]" metricMagnitude="-3" metricUnit="m" metricValue="3.3" pageId="1" pageNumber="588" unit="mm" value="3.3">3.3 mm</quantity>
. Females: the
<typeStatus id="4202EAFE6B510A1CFB00FE97FB71976B" box="[1165,1257,256,279]" pageId="1" pageNumber="588" type="paratype">paratype</typeStatus>
is slightly larger than the
<typeStatus id="4202EAFE6B510A1CFF7EFEB5FED19745" box="[243,329,290,313]" pageId="1" pageNumber="588" type="allotype">allotype</typeStatus>
(length
<quantity id="5A41F9B96B510A1CFE2CFEB5FE6D9745" box="[417,501,290,313]" metricMagnitude="-3" metricUnit="m" metricValue="7.3" pageId="1" pageNumber="588" unit="mm" value="7.3">7.3 mm</quantity>
, width
<quantity id="5A41F9B96B510A1CFDCEFEB5FD0C9745" box="[579,660,290,313]" metricMagnitude="-3" metricUnit="m" metricValue="3.7" pageId="1" pageNumber="588" unit="mm" value="3.7">3.7 mm</quantity>
at middle of elytra) and the pronotum is uniformly black.
</paragraph>
</subSubSection>
<subSubSection id="D5A307D76B510A1CFF4AFED2FEA194CB" pageId="1" pageNumber="588" type="diagnosis">
<paragraph id="9D06545C6B510A1CFF4AFED2FEA194CB" blockId="1.[151,1436,152,799]" pageId="1" pageNumber="588">
<emphasis id="AFCD884E6B510A1CFF4AFED2FEA49720" bold="true" box="[199,316,325,348]" inLineHeading="true" pageId="1" pageNumber="588" reason="1">Diagnosis.</emphasis>
In the
<bibRefCitation id="F92829AD6B510A1CFE09FED2FDA69720" author="Fuhrmann, J." box="[388,574,325,348]" pageId="1" pageNumber="588" pagination="43 - 57" refId="ref1551" refString="Fuhrmann, J. (2012) Compsodactylus, a new South American genus with one new species and two new combinations (Coleoptera: Scarabaeidae: Melolonthinae). Zootaxa, 3577, 43 - 57." type="journal article" year="2012">Fuhrmann (2012)</bibRefCitation>
key to
<taxonomicName id="5AB92FDF6B510A1CFD01FED2FCA49720" authorityName="Fuhrmann" authorityYear="2012" box="[652,828,325,348]" class="Insecta" family="Melolonthidae" genus="Compsodactylus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="588" phylum="Arthropoda" rank="genus">
<emphasis id="AFCD884E6B510A1CFD01FED2FCA49720" box="[652,828,325,348]" italics="true" pageId="1" pageNumber="588">Compsodactylus</emphasis>
</taxonomicName>
species,
<taxonomicName id="5AB92FDF6B510A1CFC11FED2FB939720" authorityName="Figueroa &amp; Neita-Moreno" authorityYear="2019" box="[924,1035,325,348]" class="Insecta" family="Melolonthidae" genus="Compsodactylus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="588" phylum="Arthropoda" rank="species" species="vallejoi">
<emphasis id="AFCD884E6B510A1CFC11FED2FC369720" box="[924,942,325,348]" italics="true" pageId="1" pageNumber="588">C</emphasis>
.
<emphasis id="AFCD884E6B510A1CFC36FED2FB939720" box="[955,1035,325,348]" italics="true" pageId="1" pageNumber="588">vallejoi</emphasis>
</taxonomicName>
will key out to couplet 2 based on the character states: body not dorsoventrally flattened, posterior area of clypeus with denticle-like setae, elytral striae strongly punctate with
<taxonomicName id="5AB92FDF6B510A1CFE07FE1DFD9597DD" baseAuthorityName="Frey" baseAuthorityYear="1972" box="[394,525,394,417]" class="Insecta" family="Melolonthidae" genus="Compsodactylus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="588" phylum="Arthropoda" rank="species" species="martinezi">
<emphasis id="AFCD884E6B510A1CFE07FE1DFD9597DD" box="[394,525,394,417]" italics="true" pageId="1" pageNumber="588">C. martinezi</emphasis>
</taxonomicName>
and
<taxonomicName id="5AB92FDF6B510A1CFDCDFE1DFD5F97DD" authorityName="Fuhrmann" authorityYear="2012" box="[576,711,394,417]" class="Insecta" family="Melolonthidae" genus="Compsodactylus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="588" phylum="Arthropoda" rank="species" species="scabrosus">
<emphasis id="AFCD884E6B510A1CFDCDFE1DFD5F97DD" box="[576,711,394,417]" italics="true" pageId="1" pageNumber="588">C. scabrosus</emphasis>
</taxonomicName>
.
<taxonomicName id="5AB92FDF6B510A1CFD59FE1DFC4397DD" authorityName="Figueroa &amp; Neita-Moreno" authorityYear="2019" box="[724,987,394,417]" class="Insecta" family="Melolonthidae" genus="Compsodactylus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="588" phylum="Arthropoda" rank="species" species="vallejoi">
<emphasis id="AFCD884E6B510A1CFD59FE1DFC4397DD" box="[724,987,394,417]" italics="true" pageId="1" pageNumber="588">Compsodactylus vallejoi</emphasis>
</taxonomicName>
differs from the other species in the genus by having protarsomere 1 of males enlarged (
<figureCitation id="058248D96B510A1CFD0DFE3AFD5B97B8" box="[640,707,429,452]" captionStart="FIGURES 3–10" captionStartId="2.[151,258,1937,1959]" captionTargetBox="[158,1432,229,1880]" captionTargetId="figure@2.[151,1435,229,1891]" captionTargetPageId="2" captionText="FIGURES 3–10. Compsodactylus martinezi and C. vallejoi leg characters. 3, Compsodactylus martinezi prothoracic legs (dorsal); 4, C. vallejoi prothoracic legs (dorsal); 5, C. vallejoi prothoracic legs (dorsal); 6, C. martinezi metafemur (dorsal); 7, C. vallejoi metafemur (dorsal); 8, C. vallejoi metafemur (dorsal); 9, C. martinezi apex of metatibia; 10, C. vallejoi apex of metatibia." figureDoi="http://doi.org/10.5281/zenodo.3993470" httpUri="https://zenodo.org/record/3993470/files/figure.png" pageId="1" pageNumber="588">Fig. 4</figureCitation>
).
<taxonomicName id="5AB92FDF6B510A1CFD5AFE3AFC4797B8" authorityName="Figueroa &amp; Neita-Moreno" authorityYear="2019" box="[727,991,429,452]" class="Insecta" family="Melolonthidae" genus="Compsodactylus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="588" phylum="Arthropoda" rank="species" species="vallejoi">
<emphasis id="AFCD884E6B510A1CFD5AFE3AFC4797B8" box="[727,991,429,452]" italics="true" pageId="1" pageNumber="588">Compsodactylus vallejoi</emphasis>
</taxonomicName>
is similar to
<taxonomicName id="5AB92FDF6B510A1CFBE3FE3AFB6A97B8" baseAuthorityName="Frey" baseAuthorityYear="1972" box="[1134,1266,429,452]" class="Insecta" family="Melolonthidae" genus="Compsodactylus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="588" phylum="Arthropoda" rank="species" species="martinezi">
<emphasis id="AFCD884E6B510A1CFBE3FE3AFB1897B8" box="[1134,1152,429,452]" italics="true" pageId="1" pageNumber="588">C</emphasis>
.
<emphasis id="AFCD884E6B510A1CFB03FE3AFB6A97B8" box="[1166,1266,429,452]" italics="true" pageId="1" pageNumber="588">martinezi</emphasis>
</taxonomicName>
as both species have the metafemur of both sexes slightly enlarged (
<figureCitation id="058248D96B510A1CFD62FE47FCC2979B" box="[751,858,464,487]" captionStart="FIGURES 3–10" captionStartId="2.[151,258,1937,1959]" captionTargetBox="[158,1432,229,1880]" captionTargetId="figure@2.[151,1435,229,1891]" captionTargetPageId="2" captionText="FIGURES 3–10. Compsodactylus martinezi and C. vallejoi leg characters. 3, Compsodactylus martinezi prothoracic legs (dorsal); 4, C. vallejoi prothoracic legs (dorsal); 5, C. vallejoi prothoracic legs (dorsal); 6, C. martinezi metafemur (dorsal); 7, C. vallejoi metafemur (dorsal); 8, C. vallejoi metafemur (dorsal); 9, C. martinezi apex of metatibia; 10, C. vallejoi apex of metatibia." figureDoi="http://doi.org/10.5281/zenodo.3993470" httpUri="https://zenodo.org/record/3993470/files/figure.png" pageId="1" pageNumber="588">Figs. 6–7</figureCitation>
) and both species with a gonostyle (
<figureCitation id="058248D96B510A1CFA8FFE47FA17979B" box="[1282,1423,464,487]" captionStart="FIGURES 11–16" captionStartId="3.[151,258,1171,1193]" captionTargetBox="[267,1329,240,1109]" captionTargetId="figure@3.[248,1346,191,1157]" captionTargetPageId="3" captionText="FIGURES 11–16. Compsodactylus martinezi and C. vallejoi genitalic characters. 11, Compsodactylus martinezi phallobase and parameres (lateral); 12, C. martinezi parameres (frontal); 13, C. martinezi female genital plates; 14, C. vallejoi phallobase and parameres (lateral); 15, C. vallejoi parameres (frontal); 16, C. vallejoi female genital plates." figureDoi="http://doi.org/10.5281/zenodo.3993472" httpUri="https://zenodo.org/record/3993472/files/figure.png" pageId="1" pageNumber="588">Figs. 13, 16</figureCitation>
).
<taxonomicName id="5AB92FDF6B510A1CFF1AFE65FE079475" authorityName="Figueroa &amp; Neita-Moreno" authorityYear="2019" box="[151,415,498,521]" class="Insecta" family="Melolonthidae" genus="Compsodactylus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="588" phylum="Arthropoda" rank="species" species="vallejoi">
<emphasis id="AFCD884E6B510A1CFF1AFE65FE079475" box="[151,415,498,521]" italics="true" pageId="1" pageNumber="588">Compsodactylus vallejoi</emphasis>
</taxonomicName>
is distinguished by the black head without metallic reflections (
<taxonomicName id="5AB92FDF6B510A1CFBD7FE65FB789475" baseAuthorityName="Frey" baseAuthorityYear="1972" box="[1114,1248,498,521]" class="Insecta" family="Melolonthidae" genus="Compsodactylus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="588" phylum="Arthropoda" rank="species" species="martinezi">
<emphasis id="AFCD884E6B510A1CFBD7FE65FBF49475" box="[1114,1132,498,521]" italics="true" pageId="1" pageNumber="588">C</emphasis>
.
<emphasis id="AFCD884E6B510A1CFBF6FE65FB789475" box="[1147,1248,498,521]" italics="true" pageId="1" pageNumber="588">martinezi</emphasis>
</taxonomicName>
has a black head with metallic green reflections), the pronotum is black with margins brownish testaceous (
<taxonomicName id="5AB92FDF6B510A1CFB06FD82FA8B9450" baseAuthorityName="Frey" baseAuthorityYear="1972" box="[1163,1299,533,556]" class="Insecta" family="Melolonthidae" genus="Compsodactylus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="588" phylum="Arthropoda" rank="species" species="martinezi">
<emphasis id="AFCD884E6B510A1CFB06FD82FA8B9450" box="[1163,1299,533,556]" italics="true" pageId="1" pageNumber="588">C. martinezi</emphasis>
</taxonomicName>
has a black pronotum with metallic green reflections), and elytra without metallic green reflections (elytra have metallic green reflections in
<taxonomicName id="5AB92FDF6B510A1CFEA2FDCDFE2E940D" baseAuthorityName="Frey" baseAuthorityYear="1972" box="[303,438,602,625]" class="Insecta" family="Melolonthidae" genus="Compsodactylus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="588" phylum="Arthropoda" rank="species" species="martinezi">
<emphasis id="AFCD884E6B510A1CFEA2FDCDFE2E940D" box="[303,438,602,625]" italics="true" pageId="1" pageNumber="588">C. martinezi</emphasis>
</taxonomicName>
). Finally, the metafemur of the male of
<taxonomicName id="5AB92FDF6B510A1CFCF8FDCDFC7F940D" authorityName="Figueroa &amp; Neita-Moreno" authorityYear="2019" box="[885,999,602,625]" class="Insecta" family="Melolonthidae" genus="Compsodactylus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="588" phylum="Arthropoda" rank="species" species="vallejoi">
<emphasis id="AFCD884E6B510A1CFCF8FDCDFC1F940D" box="[885,903,602,625]" italics="true" pageId="1" pageNumber="588">C</emphasis>
.
<emphasis id="AFCD884E6B510A1CFC1AFDCDFC7F940D" box="[919,999,602,625]" italics="true" pageId="1" pageNumber="588">vallejoi</emphasis>
</taxonomicName>
lacks an apical tooth (
<figureCitation id="058248D96B510A1CFB68FDCDFAB0940D" box="[1253,1320,602,625]" captionStart="FIGURES 3–10" captionStartId="2.[151,258,1937,1959]" captionTargetBox="[158,1432,229,1880]" captionTargetId="figure@2.[151,1435,229,1891]" captionTargetPageId="2" captionText="FIGURES 3–10. Compsodactylus martinezi and C. vallejoi leg characters. 3, Compsodactylus martinezi prothoracic legs (dorsal); 4, C. vallejoi prothoracic legs (dorsal); 5, C. vallejoi prothoracic legs (dorsal); 6, C. martinezi metafemur (dorsal); 7, C. vallejoi metafemur (dorsal); 8, C. vallejoi metafemur (dorsal); 9, C. martinezi apex of metatibia; 10, C. vallejoi apex of metatibia." figureDoi="http://doi.org/10.5281/zenodo.3993470" httpUri="https://zenodo.org/record/3993470/files/figure.png" pageId="1" pageNumber="588">Fig. 7</figureCitation>
), which is present in
<taxonomicName id="5AB92FDF6B510A1CFE85FDEAFE1694E8" baseAuthorityName="Frey" baseAuthorityYear="1972" box="[264,398,637,660]" class="Insecta" family="Melolonthidae" genus="Compsodactylus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="588" phylum="Arthropoda" rank="species" species="martinezi">
<emphasis id="AFCD884E6B510A1CFE85FDEAFE1694E8" box="[264,398,637,660]" italics="true" pageId="1" pageNumber="588">C. martinezi</emphasis>
</taxonomicName>
(
<figureCitation id="058248D96B510A1CFE10FDEAFE7894E8" box="[413,480,637,660]" captionStart="FIGURES 3–10" captionStartId="2.[151,258,1937,1959]" captionTargetBox="[158,1432,229,1880]" captionTargetId="figure@2.[151,1435,229,1891]" captionTargetPageId="2" captionText="FIGURES 3–10. Compsodactylus martinezi and C. vallejoi leg characters. 3, Compsodactylus martinezi prothoracic legs (dorsal); 4, C. vallejoi prothoracic legs (dorsal); 5, C. vallejoi prothoracic legs (dorsal); 6, C. martinezi metafemur (dorsal); 7, C. vallejoi metafemur (dorsal); 8, C. vallejoi metafemur (dorsal); 9, C. martinezi apex of metatibia; 10, C. vallejoi apex of metatibia." figureDoi="http://doi.org/10.5281/zenodo.3993470" httpUri="https://zenodo.org/record/3993470/files/figure.png" pageId="1" pageNumber="588">Fig. 6</figureCitation>
). The male and female genitalia (
<figureCitation id="058248D96B510A1CFCC1FDEAFC5594E8" box="[844,973,637,660]" captionStart="FIGURES 11–16" captionStartId="3.[151,258,1171,1193]" captionTargetBox="[267,1329,240,1109]" captionTargetId="figure@3.[248,1346,191,1157]" captionTargetPageId="3" captionText="FIGURES 11–16. Compsodactylus martinezi and C. vallejoi genitalic characters. 11, Compsodactylus martinezi phallobase and parameres (lateral); 12, C. martinezi parameres (frontal); 13, C. martinezi female genital plates; 14, C. vallejoi phallobase and parameres (lateral); 15, C. vallejoi parameres (frontal); 16, C. vallejoi female genital plates." figureDoi="http://doi.org/10.5281/zenodo.3993472" httpUri="https://zenodo.org/record/3993472/files/figure.png" pageId="1" pageNumber="588">Figs. 11–16</figureCitation>
) are diagnostic (see Fuhmrann 2012: Figs. 70–71, 73–75).
</paragraph>
</subSubSection>
<subSubSection id="D5A307D76B510A1CFF4AFD55FC739563" pageId="1" pageNumber="588" type="distribution">
<paragraph id="9D06545C6B510A1CFF4AFD55FC739563" blockId="1.[151,1436,152,799]" pageId="1" pageNumber="588">
<emphasis id="AFCD884E6B510A1CFF4AFD55FEC194A5" bold="true" box="[199,345,706,729]" inLineHeading="true" pageId="1" pageNumber="588" reason="1">Distribution.</emphasis>
Specimens were collected in two locations, at approximately
<quantity id="5A41F9B96B510A1CFB8EFD55FBA294A5" box="[1027,1082,706,729]" metricMagnitude="3" metricUnit="m" metricValue="3.0" pageId="1" pageNumber="588" unit="km" value="3.0">3 km</quantity>
apart along the Hualanga River, which is a tributary of the Marañón River (
<figureCitation id="058248D96B510A1CFDECFD72FD419480" box="[609,729,741,764]" captionStart="FIGURES 17–19" captionStartId="3.[151,258,1987,2009]" captionTargetBox="[261,1328,1295,1965]" captionTargetId="figure@3.[261,1328,1295,1965]" captionTargetPageId="3" captionText="FIGURES 17–19. Compsodactylus vallejoi. 17, distribution; 18–19, habitat." figureDoi="http://doi.org/10.5281/zenodo.3993474" httpUri="https://zenodo.org/record/3993474/files/figure.png" pageId="1" pageNumber="588">Fig. 17–19</figureCitation>
). It is an area that corresponds to a dry inter-Andean forest, which depends strictly on seasonal rains, as well as moisture from the Hualanga River.
</paragraph>
</subSubSection>
<caption id="C9C604D46B510A1CFF1AF84FFC479191" ID-DOI="http://doi.org/10.5281/zenodo.3993468" ID-Zenodo-Dep="3993468" box="[151,991,2008,2030]" httpUri="https://zenodo.org/record/3993468/files/figure.png" pageId="1" pageNumber="588" startId="1.[151,258,2008,2030]" targetBox="[229,1378,905,1946]" targetPageId="1">
<paragraph id="9D06545C6B510A1CFF1AF84FFC479191" blockId="1.[151,991,2008,2030]" box="[151,991,2008,2030]" pageId="1" pageNumber="588">
<emphasis id="AFCD884E6B510A1CFF1AF84FFEAA9192" bold="true" box="[151,306,2008,2030]" pageId="1" pageNumber="588">FIGURES 1–2.</emphasis>
<taxonomicName id="5AB92FDF6B510A1CFEBAF84FFDBB9191" authorityName="Figueroa &amp; Neita-Moreno" authorityYear="2019" box="[311,547,2008,2029]" class="Insecta" family="Melolonthidae" genus="Compsodactylus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="588" phylum="Arthropoda" rank="species" species="vallejoi">
<emphasis id="AFCD884E6B510A1CFEBAF84FFDBB9191" box="[311,547,2008,2029]" italics="true" pageId="1" pageNumber="588">Compsodactylus vallejoi</emphasis>
</taxonomicName>
holotype.
<emphasis id="AFCD884E6B510A1CFD04F84FFD0D9192" bold="true" box="[649,661,2008,2030]" pageId="1" pageNumber="588">1</emphasis>
, Dorsal habitus;
<emphasis id="AFCD884E6B510A1CFCB4F84FFCDD9192" bold="true" box="[825,837,2008,2030]" pageId="1" pageNumber="588">2</emphasis>
, lateral habitus.
</paragraph>
</caption>
<caption id="C9C604D46B520A1FFF1AF806FABE919D" ID-DOI="http://doi.org/10.5281/zenodo.3993470" ID-Zenodo-Dep="3993470" httpUri="https://zenodo.org/record/3993470/files/figure.png" pageId="2" pageNumber="589" startId="2.[151,258,1937,1959]" targetBox="[158,1432,229,1880]" targetPageId="2">
<paragraph id="9D06545C6B520A1FFF1AF806FABE919D" blockId="2.[151,1436,1937,2017]" pageId="2" pageNumber="589">
<emphasis id="AFCD884E6B520A1FFF1AF806FEA691DB" bold="true" box="[151,318,1937,1959]" pageId="2" pageNumber="589">FIGURES 3–10.</emphasis>
<taxonomicName id="5AB92FDF6B520A1FFEC9F806FDDD91DA" baseAuthorityName="Frey" baseAuthorityYear="1972" box="[324,581,1937,1958]" class="Insecta" family="Melolonthidae" genus="Compsodactylus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="589" phylum="Arthropoda" rank="species" species="martinezi">
<emphasis id="AFCD884E6B520A1FFEC9F806FDDD91DA" box="[324,581,1937,1958]" italics="true" pageId="2" pageNumber="589">Compsodactylus martinezi</emphasis>
</taxonomicName>
and
<taxonomicName id="5AB92FDF6B520A1FFDF9F806FD4191DA" authorityName="Figueroa &amp; Neita-Moreno" authorityYear="2019" box="[628,729,1937,1958]" class="Insecta" family="Melolonthidae" genus="Compsodactylus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="589" phylum="Arthropoda" rank="species" species="vallejoi">
<emphasis id="AFCD884E6B520A1FFDF9F806FD4191DA" box="[628,729,1937,1958]" italics="true" pageId="2" pageNumber="589">C. vallejoi</emphasis>
</taxonomicName>
leg characters.
<emphasis id="AFCD884E6B520A1FFCFFF806FCE691DB" bold="true" box="[882,894,1937,1959]" pageId="2" pageNumber="589">3</emphasis>
,
<taxonomicName id="5AB92FDF6B520A1FFC07F806FB1291DA" baseAuthorityName="Frey" baseAuthorityYear="1972" box="[906,1162,1937,1958]" class="Insecta" family="Melolonthidae" genus="Compsodactylus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="589" phylum="Arthropoda" rank="species" species="martinezi">
<emphasis id="AFCD884E6B520A1FFC07F806FB1291DA" box="[906,1162,1937,1958]" italics="true" pageId="2" pageNumber="589">Compsodactylus martinezi</emphasis>
</taxonomicName>
prothoracic legs (dorsal);
<emphasis id="AFCD884E6B520A1FFA07F806FA0E91DB" bold="true" box="[1418,1430,1937,1959]" pageId="2" pageNumber="589">4</emphasis>
,
<taxonomicName id="5AB92FDF6B520A1FFF1AF838FE9891B8" authorityName="Figueroa &amp; Neita-Moreno" authorityYear="2019" box="[151,256,1967,1988]" class="Insecta" family="Melolonthidae" genus="Compsodactylus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="589" phylum="Arthropoda" rank="species" species="vallejoi">
<emphasis id="AFCD884E6B520A1FFF1AF838FE9891B8" box="[151,256,1967,1988]" italics="true" pageId="2" pageNumber="589">C. vallejoi</emphasis>
</taxonomicName>
prothoracic legs (dorsal);
<emphasis id="AFCD884E6B520A1FFD82F839FD8391B8" bold="true" box="[527,539,1966,1988]" pageId="2" pageNumber="589">5</emphasis>
,
<taxonomicName id="5AB92FDF6B520A1FFDA1F838FD0D91B8" authorityName="Figueroa &amp; Neita-Moreno" authorityYear="2019" box="[556,661,1967,1988]" class="Insecta" family="Melolonthidae" genus="Compsodactylus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="589" phylum="Arthropoda" rank="species" species="vallejoi">
<emphasis id="AFCD884E6B520A1FFDA1F838FD0D91B8" box="[556,661,1967,1988]" italics="true" pageId="2" pageNumber="589">C. vallejoi</emphasis>
</taxonomicName>
prothoracic legs (dorsal);
<emphasis id="AFCD884E6B520A1FFC28F839FC2991B8" bold="true" box="[933,945,1966,1988]" pageId="2" pageNumber="589">6</emphasis>
,
<taxonomicName id="5AB92FDF6B520A1FFC4FF838FBA591B8" baseAuthorityName="Frey" baseAuthorityYear="1972" box="[962,1085,1967,1988]" class="Insecta" family="Melolonthidae" genus="Compsodactylus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="589" phylum="Arthropoda" rank="species" species="martinezi">
<emphasis id="AFCD884E6B520A1FFC4FF838FBA591B8" box="[962,1085,1967,1988]" italics="true" pageId="2" pageNumber="589">C. martinezi</emphasis>
</taxonomicName>
metafemur (dorsal);
<emphasis id="AFCD884E6B520A1FFA9BF839FABA91B8" bold="true" box="[1302,1314,1966,1988]" pageId="2" pageNumber="589">7</emphasis>
,
<taxonomicName id="5AB92FDF6B520A1FFABEF838FA0491B8" authorityName="Figueroa &amp; Neita-Moreno" authorityYear="2019" box="[1331,1436,1967,1988]" class="Insecta" family="Melolonthidae" genus="Compsodactylus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="589" phylum="Arthropoda" rank="species" species="vallejoi">
<emphasis id="AFCD884E6B520A1FFABEF838FA0491B8" box="[1331,1436,1967,1988]" italics="true" pageId="2" pageNumber="589">C. vallejoi</emphasis>
</taxonomicName>
metafemur (dorsal);
<emphasis id="AFCD884E6B520A1FFED1F85CFEF0919D" bold="true" box="[348,360,1995,2017]" pageId="2" pageNumber="589">8</emphasis>
,
<taxonomicName id="5AB92FDF6B520A1FFEF9F85BFE40919D" authorityName="Figueroa &amp; Neita-Moreno" authorityYear="2019" box="[372,472,1996,2017]" class="Insecta" family="Melolonthidae" genus="Compsodactylus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="589" phylum="Arthropoda" rank="species" species="vallejoi">
<emphasis id="AFCD884E6B520A1FFEF9F85BFE40919D" box="[372,472,1996,2017]" italics="true" pageId="2" pageNumber="589">C. vallejoi</emphasis>
</taxonomicName>
metafemur (dorsal);
<emphasis id="AFCD884E6B520A1FFD2EF85CFD37919D" bold="true" box="[675,687,1995,2017]" pageId="2" pageNumber="589">9</emphasis>
,
<taxonomicName id="5AB92FDF6B520A1FFD36F85BFCAA919D" baseAuthorityName="Frey" baseAuthorityYear="1972" box="[699,818,1996,2017]" class="Insecta" family="Melolonthidae" genus="Compsodactylus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="589" phylum="Arthropoda" rank="species" species="martinezi">
<emphasis id="AFCD884E6B520A1FFD36F85BFCAA919D" box="[699,818,1996,2017]" italics="true" pageId="2" pageNumber="589">C. martinezi</emphasis>
</taxonomicName>
apex of metatibia;
<emphasis id="AFCD884E6B520A1FFC67F85CFB9A919D" bold="true" box="[1002,1026,1995,2017]" pageId="2" pageNumber="589">10</emphasis>
,
<taxonomicName id="5AB92FDF6B520A1FFB83F85BFBEA919D" authorityName="Figueroa &amp; Neita-Moreno" authorityYear="2019" box="[1038,1138,1996,2017]" class="Insecta" family="Melolonthidae" genus="Compsodactylus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="589" phylum="Arthropoda" rank="species" species="vallejoi">
<emphasis id="AFCD884E6B520A1FFB83F85BFBEA919D" box="[1038,1138,1996,2017]" italics="true" pageId="2" pageNumber="589">C. vallejoi</emphasis>
</taxonomicName>
apex of metatibia.
</paragraph>
</caption>
<caption id="C9C604D46B530A1EFF1AFB04FB9D929F" ID-DOI="http://doi.org/10.5281/zenodo.3993472" ID-Zenodo-Dep="3993472" httpUri="https://zenodo.org/record/3993472/files/figure.png" pageId="3" pageNumber="590" startId="3.[151,258,1171,1193]" targetBox="[267,1329,240,1109]" targetPageId="3">
<paragraph id="9D06545C6B530A1EFF1AFB04FB9D929F" blockId="3.[151,1436,1171,1252]" pageId="3" pageNumber="590">
<emphasis id="AFCD884E6B530A1EFF1AFB04FED492D5" bold="true" box="[151,332,1171,1193]" pageId="3" pageNumber="590">FIGURES 11–16.</emphasis>
<taxonomicName id="5AB92FDF6B530A1EFEDAFB03FDC392D5" baseAuthorityName="Frey" baseAuthorityYear="1972" box="[343,603,1172,1193]" class="Insecta" family="Melolonthidae" genus="Compsodactylus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="590" phylum="Arthropoda" rank="species" species="martinezi">
<emphasis id="AFCD884E6B530A1EFEDAFB03FDC392D5" box="[343,603,1172,1193]" italics="true" pageId="3" pageNumber="590">Compsodactylus martinezi</emphasis>
</taxonomicName>
and
<taxonomicName id="5AB92FDF6B530A1EFD1FFB03FD6292D5" authorityName="Figueroa &amp; Neita-Moreno" authorityYear="2019" box="[658,762,1172,1193]" class="Insecta" family="Melolonthidae" genus="Compsodactylus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="590" phylum="Arthropoda" rank="species" species="vallejoi">
<emphasis id="AFCD884E6B530A1EFD1FFB03FD6292D5" box="[658,762,1172,1193]" italics="true" pageId="3" pageNumber="590">C. vallejoi</emphasis>
</taxonomicName>
genitalic characters.
<emphasis id="AFCD884E6B530A1EFC5EFB04FC6892D5" bold="true" box="[979,1008,1171,1193]" pageId="3" pageNumber="590">11,</emphasis>
<taxonomicName id="5AB92FDF6B530A1EFC77FB03FB6692D5" baseAuthorityName="Frey" baseAuthorityYear="1972" box="[1018,1278,1172,1193]" class="Insecta" family="Melolonthidae" genus="Compsodactylus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="590" phylum="Arthropoda" rank="species" species="martinezi">
<emphasis id="AFCD884E6B530A1EFC77FB03FB6692D5" box="[1018,1278,1172,1193]" italics="true" pageId="3" pageNumber="590">Compsodactylus martinezi</emphasis>
</taxonomicName>
phallobase and parameres (lateral);
<emphasis id="AFCD884E6B530A1EFEECFB27FEE792BA" bold="true" box="[353,383,1200,1222]" pageId="3" pageNumber="590">12,</emphasis>
<taxonomicName id="5AB92FDF6B530A1EFE07FB26FD9D92BA" baseAuthorityName="Frey" baseAuthorityYear="1972" box="[394,517,1201,1222]" class="Insecta" family="Melolonthidae" genus="Compsodactylus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="590" phylum="Arthropoda" rank="species" species="martinezi">
<emphasis id="AFCD884E6B530A1EFE07FB26FD9D92BA" box="[394,517,1201,1222]" italics="true" pageId="3" pageNumber="590">C. martinezi</emphasis>
</taxonomicName>
parameres (frontal);
<emphasis id="AFCD884E6B530A1EFD50FB27FD6392BA" bold="true" box="[733,763,1200,1222]" pageId="3" pageNumber="590">13,</emphasis>
<taxonomicName id="5AB92FDF6B530A1EFC8BFB26FC1A92BA" baseAuthorityName="Frey" baseAuthorityYear="1972" box="[774,898,1201,1222]" class="Insecta" family="Melolonthidae" genus="Compsodactylus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="590" phylum="Arthropoda" rank="species" species="martinezi">
<emphasis id="AFCD884E6B530A1EFC8BFB26FC1A92BA" box="[774,898,1201,1222]" italics="true" pageId="3" pageNumber="590">C. martinezi</emphasis>
</taxonomicName>
female genital plates;
<emphasis id="AFCD884E6B530A1EFBE1FB27FB1C92BA" bold="true" box="[1132,1156,1200,1222]" pageId="3" pageNumber="590">14</emphasis>
,
<taxonomicName id="5AB92FDF6B530A1EFB18FB26FB6692BA" authorityName="Figueroa &amp; Neita-Moreno" authorityYear="2019" box="[1173,1278,1201,1222]" class="Insecta" family="Melolonthidae" genus="Compsodactylus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="590" phylum="Arthropoda" rank="species" species="vallejoi">
<emphasis id="AFCD884E6B530A1EFB18FB26FB6692BA" box="[1173,1278,1201,1222]" italics="true" pageId="3" pageNumber="590">C. vallejoi</emphasis>
</taxonomicName>
phallobase and parameres (lateral);
<emphasis id="AFCD884E6B530A1EFED5FB59FEE89298" bold="true" box="[344,368,1230,1252]" pageId="3" pageNumber="590">15</emphasis>
,
<taxonomicName id="5AB92FDF6B530A1EFEF1FB59FE47929F" authorityName="Figueroa &amp; Neita-Moreno" authorityYear="2019" box="[380,479,1230,1251]" class="Insecta" family="Melolonthidae" genus="Compsodactylus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="590" phylum="Arthropoda" rank="species" species="vallejoi">
<emphasis id="AFCD884E6B530A1EFEF1FB59FE47929F" box="[380,479,1230,1251]" italics="true" pageId="3" pageNumber="590">C. vallejoi</emphasis>
</taxonomicName>
parameres (frontal);
<emphasis id="AFCD884E6B530A1EFD26FB59FD5B9298" bold="true" box="[683,707,1230,1252]" pageId="3" pageNumber="590">16</emphasis>
,
<taxonomicName id="5AB92FDF6B530A1EFD43FB59FCAB929F" authorityName="Figueroa &amp; Neita-Moreno" authorityYear="2019" box="[718,819,1230,1251]" class="Insecta" family="Melolonthidae" genus="Compsodactylus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="590" phylum="Arthropoda" rank="species" species="vallejoi">
<emphasis id="AFCD884E6B530A1EFD43FB59FCAB929F" box="[718,819,1230,1251]" italics="true" pageId="3" pageNumber="590">C. vallejoi</emphasis>
</taxonomicName>
female genital plates.
</paragraph>
</caption>
<caption id="C9C604D46B530A1EFF1AF854FCEE91A4" ID-DOI="http://doi.org/10.5281/zenodo.3993474" ID-Zenodo-Dep="3993474" box="[151,886,1987,2009]" httpUri="https://zenodo.org/record/3993474/files/figure.png" pageId="3" pageNumber="590" startId="3.[151,258,1987,2009]" targetBox="[261,1328,1295,1965]" targetPageId="3">
<paragraph id="9D06545C6B530A1EFF1AF854FCEE91A4" blockId="3.[151,886,1987,2009]" box="[151,886,1987,2009]" pageId="3" pageNumber="590">
<emphasis id="AFCD884E6B530A1EFF1AF854FED191A5" bold="true" box="[151,329,1987,2009]" pageId="3" pageNumber="590">FIGURES 17–19.</emphasis>
<taxonomicName id="5AB92FDF6B530A1EFEC2F854FDA391A4" authorityName="Figueroa &amp; Neita-Moreno" authorityYear="2019" box="[335,571,1987,2008]" class="Insecta" family="Melolonthidae" genus="Compsodactylus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="590" phylum="Arthropoda" rank="species" species="vallejoi">
<emphasis id="AFCD884E6B530A1EFEC2F854FDA391A4" box="[335,571,1987,2008]" italics="true" pageId="3" pageNumber="590">Compsodactylus vallejoi</emphasis>
</taxonomicName>
.
<emphasis id="AFCD884E6B530A1EFDCAF854FDC791A5" bold="true" box="[583,607,1987,2009]" pageId="3" pageNumber="590">17</emphasis>
, distribution;
<emphasis id="AFCD884E6B530A1EFD6AF854FCBA91A5" bold="true" box="[743,802,1987,2009]" pageId="3" pageNumber="590">18–19</emphasis>
, habitat.
</paragraph>
</caption>
<subSubSection id="D5A307D76B540A19FF4AFF0FFE8496AD" pageId="4" pageNumber="591" type="etymology">
<paragraph id="9D06545C6B540A19FF4AFF0FFE8496AD" blockId="4.[151,1435,152,209]" pageId="4" pageNumber="591">
<emphasis id="AFCD884E6B540A19FF4AFF0FFEDF96D3" bold="true" box="[199,327,152,175]" inLineHeading="true" pageId="4" pageNumber="591" reason="1">Etymology.</emphasis>
The specific name
<taxonomicName id="5AB92FDF6B540A19FD93FF0FFDF696D3" authorityName="Figueroa &amp; Neita-Moreno" authorityYear="2019" box="[542,622,152,175]" class="Insecta" family="Melolonthidae" genus="Compsodactylus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="591" phylum="Arthropoda" rank="species" species="vallejoi">
<emphasis id="AFCD884E6B540A19FD93FF0FFDF696D3" box="[542,622,152,175]" italics="true" pageId="4" pageNumber="591">vallejoi</emphasis>
</taxonomicName>
is in honor of the great poet and writer
<collectingRegion id="5F7D9ABE6B540A19FBAAFF0FFBFC96D3" box="[1063,1124,152,175]" country="Colombia" name="Cesar" pageId="4" pageNumber="591">Cesar</collectingRegion>
Vallejo, born in
<collectingRegion id="5F7D9ABE6B540A19FA90FF0FFA0396D3" box="[1309,1435,152,175]" country="Peru" name="La Libertad" pageId="4" pageNumber="591">La Libertad</collectingRegion>
Department.
</paragraph>
</subSubSection>
</treatment>
</document>
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