391 lines
55 KiB
XML
391 lines
55 KiB
XML
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<mods:title id="A38887393B47BB7551399AD1D5450400">Sinolethrus, a new subgenus of the genus Lethrus Scopoli, 1777 from China (Coleoptera: Geotrupidae: Lethrinae) and new synonymy of the subgenus Paralethrus Nikolajev, 2003</mods:title>
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<treatment id="03C087DC790DE64DB9ECA3CCB4F762DF" ID-DOI="http://doi.org/10.5281/zenodo.7781950" ID-GBIF-Taxon="209473898" ID-Zenodo-Dep="7781950" LSID="urn:lsid:plazi:treatment:03C087DC790DE64DB9ECA3CCB4F762DF" httpUri="http://treatment.plazi.org/id/03C087DC790DE64DB9ECA3CCB4F762DF" lastPageNumber="304" pageId="3" pageNumber="304">
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<paragraph id="8BD636CA790DE64DB9ECA3CCB3E26598" blockId="3.[151,910,151,214]" box="[151,910,151,178]" pageId="3" pageNumber="304">
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<heading id="D09E81A6790DE64DB9ECA3CCB3E26598" bold="true" box="[151,910,151,178]" fontSize="11" level="1" pageId="3" pageNumber="304" reason="1">
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<taxonomicName id="4C694D49790DE64DB9ECA3CCB3E26598" ID-CoL="722P9" authority="Semenov & Medvedev, 1935" authorityName="Semenov & Medvedev" authorityYear="1935" box="[151,910,151,178]" class="Insecta" family="Geotrupidae" genus="Lethrus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="304" phylum="Arthropoda" rank="species" species="serpentifer" subGenus="Sinolethrus">
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<emphasis id="B91DEAD8790DE64DB9ECA3CCB3E26598" bold="true" box="[151,910,151,178]" pageId="3" pageNumber="304">
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<emphasis id="B91DEAD8790DE64DB9ECA3CCB09A659B" bold="true" box="[151,246,151,178]" italics="true" pageId="3" pageNumber="304">Lethrus</emphasis>
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(
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<emphasis id="B91DEAD8790DE64DB87CA3CCB1FF659B" bold="true" box="[263,403,151,178]" italics="true" pageId="3" pageNumber="304">Sinolethrus</emphasis>
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)
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<emphasis id="B91DEAD8790DE64DB8DFA3CCB24A6598" bold="true" box="[420,550,151,177]" italics="true" pageId="3" pageNumber="304">serpentifer</emphasis>
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<bibRefCitation id="EFF84B3B790DE64DBB55A3CCB3E26598" author="Semenov, A. P. & Medvedev, S." box="[558,910,151,177]" pageId="3" pageNumber="304" pagination="282 - 287" refId="ref9712" refString="Semenov, A. P. & Medvedev, S. (1935) Specierum novarum generis Lethrus Scop. (Coleoptera, Scarabaeidae) diagnoses. Entomologicheskoe Obozrenie, 25, 282 - 287. [in Latin]" type="journal article" year="1935">Semenov & Medvedev, 1935</bibRefCitation>
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</emphasis>
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</taxonomicName>
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</heading>
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</paragraph>
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<subSubSection id="C3736541790DE64DB9ECA3E0B25465FF" box="[151,568,187,214]" pageId="3" pageNumber="304" type="description">
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<paragraph id="8BD636CA790DE64DB9ECA3E0B25465FF" blockId="3.[151,910,151,214]" box="[151,568,187,214]" pageId="3" pageNumber="304">
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(
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||
<figureCitation id="13522A4F790DE64DB9E4A3E0B16765FC" box="[159,267,187,213]" captionStart="FIGURES 1–4" captionStartId="2.[151,264,1867,1891]" captionTargetBox="[166,1429,751,1823]" captionTargetId="figure-229@2.[151,1436,734,1841]" captionTargetPageId="2" captionText="FIGURES 1–4. Aedeagus, schematically. 1–2 L. (Ceratodirus) dostojewskii Semenov, 1899 (1 lateral view, 2 dorsal view); 3–4 L. (Sinolethrus) serpentifer Semenov & Medvedev, 1935 (3 lateral view, 4 dorsal view)." figureDoi="http://doi.org/10.5281/zenodo.7780395" httpUri="https://zenodo.org/record/7780395/files/figure.png" pageId="3" pageNumber="304">Figs. 3–4</figureCitation>
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,
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<figureCitation id="13522A4F790DE64DB86CA3E0B1E565FC" box="[279,393,187,213]" captionStart="FIGURES 13–17" captionStartId="7.[151,264,1945,1969]" captionTargetBox="[297,1291,1108,1909]" captionTargetId="figure-393@7.[284,1304,1096,1922]" captionTargetPageId="7" captionText="FIGURES 13–17. Head of Lethrus species with mandibular appendix, keels and hornlike apophyses of mandibles (males), schematically. 13–14, 17 L. (Sinolethrus) serpentifer Semenov & Medvedev, 1935, (13 frontal view, 14 lateral view, 17 dorsal view); 15 L. (Ceratodirus) cephalotes Pallas, 1771 (dorsal view); 16 L. (Ceratodirus) karelini Gebler, 1845 (dorsal view)." figureDoi="http://doi.org/10.5281/zenodo.7780401" httpUri="https://zenodo.org/record/7780401/files/figure.png" pageId="3" pageNumber="304">13–14, 17</figureCitation>
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,
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<figureCitation id="13522A4F790DE64DB8EEA3E0B1B165FF" box="[405,477,187,214]" captionStart="FIGURES 55–58" captionStartId="11.[151,264,1556,1580]" captionTargetBox="[171,1415,1061,1519]" captionTargetId="figure-78@11.[154,1434,1040,1532]" captionTargetPageId="11" captionText="FIGURES 55–58. Labels, apical spurs and head of males of Lethrus species, schematically. 55 apical spur of front tibia of L. (Sinolethrus) serpentifer Semenov & Medvedev, 1935; 56 apical spur of front tibia of L. (Ceratodirus) karelini Gebler, 1845; 57 label of syntype of L. serpentifer; 58 label of lectotype of L. mediocris." figureDoi="http://doi.org/10.5281/zenodo.7780421" httpUri="https://zenodo.org/record/7780421/files/figure.png" pageId="3" pageNumber="304">55, 57</figureCitation>
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,
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<figureCitation id="13522A4F790DE64DB892A3E0B25C65FF" box="[489,560,187,214]" captionStart="FIGURES 59–65" captionStartId="12.[151,264,1624,1648]" captionTargetBox="[201,1392,189,1588]" captionTargetId="figure-34@12.[195,1392,181,1600]" captionTargetPageId="12" captionText="FIGURES 59–65. Lethrus (Sinolethrus) serpentifer Semenov & Medvedev, 1935 (Huang He, China). 59 male dorsal view; 60 male ventral view; 61 female dorsal view; 62 male, lateral view, right; 63 male, lateral view, left; 64 male, frontal view; 65 head with mentum of male, schematically." figureDoi="http://doi.org/10.5281/zenodo.7780423" httpUri="https://zenodo.org/record/7780423/files/figure.png" pageId="3" pageNumber="304">59–64</figureCitation>
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)
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</paragraph>
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||
</subSubSection>
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||
<subSubSection id="C3736541790DE64DB9ECA25BB2F26724" pageId="3" pageNumber="304" type="reference_group">
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<paragraph id="8BD636CA790DE64DB9ECA25BB30A647C" blockId="3.[151,1437,255,525]" pageId="3" pageNumber="304">
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<treatmentCitationGroup id="AB7911E4790DE64DB9ECA25BB5F1641F" pageId="3" pageNumber="304">
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<treatmentCitation id="0AC810DB790DE64DB9ECA25BB312643E" author="Semenov, A. P. & Medvedev, S." box="[151,894,255,280]" page="286" pageId="3" pageNumber="304" year="1935">
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||
<taxonomicName id="4C694D49790DE64DB9ECA25BB312643E" ID-CoL="722P9" authority="Semenov & Medvedev 1935: 286" authorityName="Semenov & Medvedev" authorityPageNumber="286" authorityYear="1935" box="[151,894,255,280]" class="Insecta" family="Geotrupidae" genus="Lethrus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="304" phylum="Arthropoda" rank="species" species="serpentifer" subGenus="Heteroplistodes">
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||
<emphasis id="B91DEAD8790DE64DB9ECA25BB0886431" box="[151,228,256,280]" italics="true" pageId="3" pageNumber="304">Lethrus</emphasis>
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||
(
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||
<emphasis id="B91DEAD8790DE64DB98DA25BB1F96431" box="[246,405,256,280]" italics="true" pageId="3" pageNumber="304">Heteroplistodes</emphasis>
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||
)
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<emphasis id="B91DEAD8790DE64DB8D3A25BB27A6431" box="[424,534,256,280]" italics="true" pageId="3" pageNumber="304">serpentifer</emphasis>
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||
<bibRefCitation id="EFF84B3B790DE64DBB67A25BB312643E" author="Semenov, A. P. & Medvedev, S." box="[540,894,255,280]" pageId="3" pageNumber="304" pagination="282 - 287" refId="ref9712" refString="Semenov, A. P. & Medvedev, S. (1935) Specierum novarum generis Lethrus Scop. (Coleoptera, Scarabaeidae) diagnoses. Entomologicheskoe Obozrenie, 25, 282 - 287. [in Latin]" type="journal article" year="1935">Semenov & Medvedev 1935: 286</bibRefCitation>
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</taxonomicName>
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</treatmentCitation>
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||
(original description,
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||
<typeStatus id="54D28868790DE64DBD13A25BB4F86431" box="[1128,1172,256,280]" pageId="3" pageNumber="304">type</typeStatus>
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locality: “Mongoliae ora australis, Nan-Shanj, Tshshuniche”);
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<treatmentCitation id="0AC810DB790DE64DBB45A245B3E8641F" author="Semenov, A. P. & Medvedev, S. I." box="[574,900,286,310]" page="89" pageId="3" pageNumber="304" year="1936">
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||
<bibRefCitation id="EFF84B3B790DE64DBB45A245B3E8641F" author="Semenov, A. P. & Medvedev, S. I." box="[574,900,286,310]" pageId="3" pageNumber="304" refId="ref9752" refString="Semenov, A. P. & Medvedev, S. I. (1936) Opredelitel' zhukov-kravchikov. Synopsis des genres et especes de la tribu Lethrini (fam. Scarabaeidae). Izdatelstvo Akademii Nauk SSSR, Moskva, Leningrad, 104 pp. [in Russian]" type="book" year="1936">Semenov & Medvedev 1936: 89</bibRefCitation>
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||
</treatmentCitation>
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||
(key, distribution, iconography: Tab. II
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||
<figureCitation id="13522A4F790DE64DBC6FA245B504641F" box="[1300,1384,286,310]" captionStart="FIGURES 49–54" captionStartId="11.[151,264,920,944]" captionTargetBox="[159,1432,193,886]" captionTargetId="figure-19@11.[151,1436,181,896]" captionTargetPageId="11" captionText="FIGURES 49–54. Head of Lethrus species with mandibular appendix, keels and hornlike apophyses of mandibles (males), schematically. 49–51 L. (Mesolethrus) microbuccis Ballion, 1871 (49 frontal view, 50 lateral view, 51 dorsal view); 52–54 L. (Neolethrus) arcanus Medvedev, 1971 (52 frontal view, 53 lateral view, 54 dorsal view)." figureDoi="http://doi.org/10.5281/zenodo.7780419" httpUri="https://zenodo.org/record/7780419/files/figure.png" pageId="3" pageNumber="304">Figs. 54</figureCitation>
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, Tab.
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</treatmentCitationGroup>
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V Fig. 115, Tab. VII Fig. 176, Tab. IX Fig. 225, Tab. XI Fig. 282).
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</paragraph>
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<paragraph id="8BD636CA790DE64DB9ECA207B3DC645D" blockId="3.[151,1437,255,525]" box="[151,944,347,372]" pageId="3" pageNumber="304">
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||
<taxonomicName id="4C694D49790DE64DB9ECA207B2A8645A" ID-CoL="722P9" authority=": Nikolajev 1968: 531" authorityName="Nikolajev" authorityPageNumber="531" authorityYear="1968" box="[151,708,347,372]" class="Insecta" family="Geotrupidae" genus="Lethrus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="304" phylum="Arthropoda" rank="species" species="serpentifer" subGenus="Autolethrus">
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<emphasis id="B91DEAD8790DE64DB9ECA207B088645D" box="[151,228,348,372]" italics="true" pageId="3" pageNumber="304">Lethrus</emphasis>
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||
(
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<emphasis id="B91DEAD8790DE64DB988A207B105645D" box="[243,361,348,372]" italics="true" pageId="3" pageNumber="304">Autolethrus</emphasis>
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)
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<emphasis id="B91DEAD8790DE64DB803A207B18A645D" box="[376,486,348,372]" italics="true" pageId="3" pageNumber="304">serpentifer</emphasis>
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:
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<bibRefCitation id="EFF84B3B790DE64DB888A207B2A8645A" author="Nikolajev, G. V." box="[499,708,347,372]" pageId="3" pageNumber="304" pagination="528 - 540" refId="ref9198" refString="Nikolajev, G. V. (1968) Reviziya podroda Ceratodirus Fisch. - W. (Coleoptera, Scarabaeidae). Revision of the subgenus Ceratodirus Fisch. - W. (Coleoptera, Scarabaeidae). Entomologicheskoe Obozrenie, 47 (3), 528 - 540. [in Russian]" type="journal article" year="1968">Nikolajev 1968: 531</bibRefCitation>
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</taxonomicName>
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||
(taxonomic note, key).
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||
</paragraph>
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<paragraph id="8BD636CA790DE64DB9ECA221B50364F9" blockId="3.[151,1437,255,525]" pageId="3" pageNumber="304">
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||
<taxonomicName id="4C694D49790DE64DB9ECA221B31664BB" ID-CoL="722P9" authority=": Nikolajev & Puntsagdulam 1984: 110" authorityName="Nikolajev & Puntsagdulam" authorityPageNumber="110" authorityYear="1984" box="[151,890,378,402]" class="Insecta" family="Geotrupidae" genus="Lethrus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="304" phylum="Arthropoda" rank="species" species="serpentifer" subGenus="Ceratodirus">
|
||
<emphasis id="B91DEAD8790DE64DB9ECA221B08864BB" box="[151,228,378,402]" italics="true" pageId="3" pageNumber="304">Lethrus</emphasis>
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||
(
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||
<emphasis id="B91DEAD8790DE64DB98FA221B10264BB" box="[244,366,378,402]" italics="true" pageId="3" pageNumber="304">Ceratodirus</emphasis>
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||
)
|
||
<emphasis id="B91DEAD8790DE64DB806A221B18764BB" box="[381,491,378,402]" italics="true" pageId="3" pageNumber="304">serpentifer</emphasis>
|
||
:
|
||
<bibRefCitation id="EFF84B3B790DE64DB882A221B31664BB" author="Nikolajev, G. V. & Puntsagdulam, Z." box="[505,890,378,402]" pageId="3" pageNumber="304" pagination="90 - 294" refId="ref9339" refString="Nikolajev, G. V. & Puntsagdulam, Z. (1984) Plastinchatousye (Coleoptera, Scarabaeoidea) Mongolskoy Narodnoy Respubliki [Lamellicorns of the Mongolian People's Republic]. Insects of Mongolia, 9, 90 - 294. [in Russian]" type="journal article" year="1984">Nikolajev & Puntsagdulam 1984: 110</bibRefCitation>
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||
</taxonomicName>
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||
(key, characteristic, distribution);
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||
<bibRefCitation id="EFF84B3B790DE64DBDA3A221B5F064BB" author="Nikolajev, G. V." box="[1240,1436,378,402]" pageId="3" pageNumber="304" refId="ref9139" refString="Nikolajev, G. V. (2003) Zhuki-kravchiki (Scarabaeidae, Geotrupinae, Lethrini): biologiya, sistematika, rasprostranenie, opredelitel. [Eath-boring dung beetles (Scarabaeidae, Geotrupinae, Lethrini): biology, systematic, distribution, identification]. Kazak Univerziteti, Almaty, 254 pp. [in Russian]" type="book" year="2003">Nikolajev 2003: 93</bibRefCitation>
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||
(key, characteristic, distribution, iconography:
|
||
<figureCitation id="13522A4F790DE64DBBDEA2C2B29B6498" box="[677,759,409,433]" captionStart="FIGURES 59–65" captionStartId="12.[151,264,1624,1648]" captionTargetBox="[201,1392,189,1588]" captionTargetId="figure-34@12.[195,1392,181,1600]" captionTargetPageId="12" captionText="FIGURES 59–65. Lethrus (Sinolethrus) serpentifer Semenov & Medvedev, 1935 (Huang He, China). 59 male dorsal view; 60 male ventral view; 61 female dorsal view; 62 male, lateral view, right; 63 male, lateral view, left; 64 male, frontal view; 65 head with mentum of male, schematically." figureDoi="http://doi.org/10.5281/zenodo.7780423" httpUri="https://zenodo.org/record/7780423/files/figure.png" pageId="3" pageNumber="304">Figs. 62</figureCitation>
|
||
Nr. 1–2);
|
||
<bibRefCitation id="EFF84B3B790DE64DBA19A2C2B4196498" author="Kral, D. & Nikolajev, G. V." box="[866,1141,409,433]" pageId="3" pageNumber="304" refId="ref8832" refString="Kral, D. & Nikolajev, G. V. (2006) Geotrupidae: Lethrinae. In: Lobl, I. & Smetana, A. (Eds.), Catalogue of Palaearctic Coleoptera. Vol. 3. Apollo Books, Stenstrup, pp. 93 - 95." type="book" year="2006">Král & Nikolajev 2006: 92</bibRefCitation>
|
||
(catalogue);
|
||
<bibRefCitation id="EFF84B3B790DE64DBD86A2C2B5F16498" author="Nikolajev, G. V. & Kral, D. & Bezdek, A." box="[1277,1437,409,433]" pageId="3" pageNumber="304" refId="ref9253" refString="Nikolajev, G. V., Kral, D. & Bezdek, A. (2016) Family Geotrupidae. In: Lobl, I. & Lobl, D. (Eds.), Catalogue of Palaearctic Coleoptera. Vol. 3. Scarabaeoidea - Scirtoidea - Buprestoidea - Byrrhoidea. Revised and Updated Edition. E. J. Brill, Leiden, pp. 33 - 52. https: // doi. org / 10.1163 / 9789004309142 _ 003" type="book" year="2016">
|
||
Nikolajev
|
||
<emphasis id="B91DEAD8790DE64DBC12A2C1B5F16498" box="[1385,1437,409,433]" italics="true" pageId="3" pageNumber="304">et al.</emphasis>
|
||
</bibRefCitation>
|
||
2016: 48 (catalogue);
|
||
<bibRefCitation id="EFF84B3B790DE64DB8DDA2E3B36C64F9" author="Bagaturov, M. F. & Nikolajev, G. V." box="[422,768,439,464]" pageId="3" pageNumber="304" pagination="303 - 314" refId="ref8128" refString="Bagaturov, M. F. & Nikolajev, G. V. (2015) Obzor rasprostraneniya zhukov-kravchikov roda Lethrus, 1777 (Coleoptera: Geotrupidae). [Overview of distribution of the genus Lethrus Scopoli, 1777 (Coleoptera: Geotrupidae)]. Kavkazskiy Entomologicheskiy Byulleten, 11, 303 - 314. [in Russian, English abstract] https: // doi. org / 10.23885 / 1814 - 3326 - 2015 - 11 - 2 - 303 - 314" type="journal article" year="2015">Bagaturov & Nikolajev 2015: 309</bibRefCitation>
|
||
(note, distribution);
|
||
<bibRefCitation id="EFF84B3B790DE64DBAA9A2E3B4C664F9" author="Schoolmeesters, P." box="[978,1194,440,464]" pageId="3" pageNumber="304" pagination="2020" refId="ref9802" refString="Schoolmeesters, P. (2021) Scarabs: World Scarabaeidae Database. Version November 2020. In: Roskov, Y., Abucay, L., Orrell, T., Nicolson, D., Bailly, N., Kirk, P., Bourgoin, T.," type="book chapter" year="2021">Schoolmeesters 2021</bibRefCitation>
|
||
(online catalogue).
|
||
</paragraph>
|
||
<paragraph id="8BD636CA790DE64DB9ECA28DB30364C7" blockId="3.[151,1437,255,525]" box="[151,879,470,494]" pageId="3" pageNumber="304">
|
||
<taxonomicName id="4C694D49790DE64DB9ECA28DB2EF64C7" ID-CoL="722P9" authority=": Baraud 1992: 81" authorityName="Scopoli" authorityPageNumber="81" authorityYear="1777" box="[151,643,470,494]" class="Insecta" family="Geotrupidae" genus="Lethrus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="304" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B91DEAD8790DE64DB9ECA28DB08864C7" box="[151,228,470,494]" italics="true" pageId="3" pageNumber="304">Lethrus</emphasis>
|
||
(Subgen.?)
|
||
<emphasis id="B91DEAD8790DE64DB81BA28DB1A264C7" box="[352,462,470,494]" italics="true" pageId="3" pageNumber="304">serpentifer</emphasis>
|
||
:
|
||
<bibRefCitation id="EFF84B3B790DE64DB8A0A28DB2EF64C7" author="Baraud, J." box="[475,643,470,494]" pageId="3" pageNumber="304" refId="ref8028" refString="Baraud, J. (1992) Coleopteres Scarabaeoidea d'Europe. Faune de France. Federation francaise des Societes de Sciences naturelles, Paris & Societe linneenne de Lyon, Paris, 856 pp. https: // doi. org / 10.3406 / linly. 1992.11007" type="book" year="1992">Baraud 1992: 81</bibRefCitation>
|
||
</taxonomicName>
|
||
(key, taxonomic note).
|
||
</paragraph>
|
||
<paragraph id="8BD636CA790DE64DB9ECA2AEB2F26724" blockId="3.[151,1437,255,525]" box="[151,670,501,525]" pageId="3" pageNumber="304">
|
||
<taxonomicName id="4C694D49790DE64DB9ECA2AEB2726724" ID-CoL="722P9" authority=": Krajcik 2012: 142" authorityName="Krajcik" authorityPageNumber="142" authorityYear="2012" box="[151,542,501,525]" class="Insecta" family="Geotrupidae" genus="Lethrus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="304" phylum="Arthropoda" rank="species" species="serpentifer">
|
||
<emphasis id="B91DEAD8790DE64DB9ECA2AEB1356724" box="[151,345,501,525]" italics="true" pageId="3" pageNumber="304">Lethrus serpentifer</emphasis>
|
||
:
|
||
<bibRefCitation id="EFF84B3B790DE64DB81DA2AEB2726724" author="Krajcik, M." box="[358,542,501,525]" pageId="3" pageNumber="304" pagination="1 - 278" refId="ref8577" refString="Krajcik, M. (2012) Checklist of the World Scarabaeoidea. Animma. x (supplement), 5, 1 - 278." type="journal article" year="2012">Krajčík 2012: 142</bibRefCitation>
|
||
</taxonomicName>
|
||
(catalogue).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3736541790DE64DB9ECA160B52C66B0" pageId="3" pageNumber="304" type="materials_examined">
|
||
<paragraph id="8BD636CA790DE64DB9ECA160B1A067B4" blockId="3.[151,1437,570,2038]" pageId="3" pageNumber="304">
|
||
<emphasis id="B91DEAD8790DE64DB9ECA160B1AC677C" bold="true" box="[151,448,571,597]" pageId="3" pageNumber="304">Type material examined.</emphasis>
|
||
<materialsCitation id="3B013C97790DE64DB8B3A160B1A467B4" ID-GBIF-Occurrence="4067802302" pageId="3" pageNumber="304" typeStatus="syntype">
|
||
<emphasis id="B91DEAD8790DE64DB8B3A160B244677C" bold="true" box="[456,552,571,597]" pageId="3" pageNumber="304">
|
||
<typeStatus id="54D28868790DE64DB8B3A160B244677C" box="[456,552,571,597]" pageId="3" pageNumber="304" type="syntype">Syntype</typeStatus>
|
||
</emphasis>
|
||
, Ô (
|
||
<collectionCode id="ED78AE0F790DE64DBB20A160B2C0677C" box="[603,684,571,597]" pageId="3" pageNumber="304">ZMAS</collectionCode>
|
||
) “ДОЛ[ина] Р[еки] ЧхунихƐ/ 12 IV 09 ƐкЗ[емпЛЯР] КОЗЛОва [hw] //
|
||
<taxonomicName id="4C694D49790DE64DB9D4A104B10C6750" authorityName="B. Jakovlev" authorityYear="1890" box="[175,352,607,633]" class="Insecta" family="Geotrupidae" genus="Heteroplistodus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="305" phylum="Arthropoda" rank="genus">Heteroplistodus</taxonomicName>
|
||
/ serpentifer / sp. n. Ô [hw] / A. Semenov-Tian-Shansky det. [p] 22 [hw] // Holotypus [p] //
|
||
<collectingRegion id="49ADF828790DE64DBC05A104B16267B4" country="Russia" name="Saint Petersburg" pageId="3" pageNumber="304">St. Petersburg</collectingRegion>
|
||
/ Zool. Inst [p]”
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8BD636CA790DE64DB9BCA1FCB52C66B0" blockId="3.[151,1437,570,2038]" pageId="3" pageNumber="304">
|
||
<emphasis id="B91DEAD8790DE64DB9BCA1FCB24567E8" bold="true" box="[199,553,679,705]" pageId="3" pageNumber="304">Additional material examined</emphasis>
|
||
(
|
||
<specimenCount id="9D6FFD43790DE64DBB42A1FCB2AF67E8" box="[569,707,679,705]" pageId="3" pageNumber="304" type="generic">7 specimens</specimenCount>
|
||
).
|
||
<materialsCitation id="3B013C97790DE64DBBA2A1FCB0856620" ID-GBIF-Occurrence="4067802308" collectingDate="1996-05-26" collectingDateMax="1996-05-28" collectingDateMin="1996-05-26" collectionCode="ZSM, NMPC" collectorName="J. Halada" country="China" latitude="34.116665" location="Huang He" longLatPrecision="1201" longitude="111.11667" municipality="river Huang He" pageId="3" pageNumber="304" specimenCount="1" stateProvince="Monan">
|
||
<collectingCountry id="F37E765A790DE64DBBA2A1FCB37767E8" box="[729,795,679,705]" name="China" pageId="3" pageNumber="304">China</collectingCountry>
|
||
c.,
|
||
<collectingRegion id="49ADF828790DE64DBA38A1F3B3F867EB" box="[835,916,680,706]" pageId="3" pageNumber="304">Monan</collectingRegion>
|
||
, river
|
||
<location id="8EB66011790DE64DBAA0A1F3B42767EB" LSID="urn:lsid:plazi:treatment:03C087DC790DE64DB9ECA3CCB4F762DF:8EB66011790DE64DBAA0A1F3B42767EB" box="[987,1099,680,706]" country="China" latitude="34.116665" longLatPrecision="1201" longitude="111.11667" municipality="river Huang He" name="Huang He" pageId="3" pageNumber="304" stateProvince="Monan">Huang He</location>
|
||
(
|
||
<geoCoordinate id="EE5D500D790DE64DBD20A1FCB4DA67E8" box="[1115,1206,679,705]" degrees="34" direction="north" minutes="07" orientation="latitude" pageId="3" pageNumber="304" precision="925" value="34.116665">34°07'N</geoCoordinate>
|
||
/
|
||
<geoCoordinate id="EE5D500D790DE64DBDBDA1FCB54067E8" box="[1222,1324,679,705]" degrees="111" direction="east" minutes="07" orientation="longitude" pageId="3" pageNumber="304" precision="925" value="111.11667">111°07'E</geoCoordinate>
|
||
) [
|
||
<location id="8EB66011790DE64DBC3AA1F3B5FD67EB" LSID="urn:lsid:plazi:treatment:03C087DC790DE64DB9ECA3CCB4F762DF:8EB66011790DE64DBC3AA1F3B5FD67EB" box="[1345,1425,680,706]" country="China" latitude="34.116665" longLatPrecision="1201" longitude="111.11667" municipality="river Huang He" name="Henan" pageId="3" pageNumber="304" stateProvince="Monan">Henan</location>
|
||
],
|
||
<date id="FFD7100A790DE64DB9ECA190B12467CC" box="[151,328,715,742]" pageId="3" pageNumber="304" value="1996-05-26" valueMax="1996-05-28" valueMin="1996-05-26">
|
||
<collectingDate id="EF93E9E2790DE64DB9ECA190B12467CC" box="[151,328,715,742]" pageId="3" pageNumber="304" value="1996-05-26" valueMax="1996-05-28" valueMin="1996-05-26">26–28. V. 1996</collectingDate>
|
||
</date>
|
||
, leg.
|
||
<collectorName id="269C531C790DE64DB8F2A197B19967CC" box="[393,501,715,742]" pageId="3" pageNumber="304">J. Halada</collectorName>
|
||
, Ô (
|
||
<collectionCode id="ED78AE0F790DE64DBB51A190B20B67CC" box="[554,615,715,741]" country="Germany" httpUri="http://grbio.org/cool/4b7r-7sec" name="Bavarian State Collection of Zoology" pageId="3" pageNumber="304" type="Museum">ZSM</collectionCode>
|
||
, ex. coll. O. Hillert), Ô (
|
||
<collectionCode id="ED78AE0F790DE64DBAF3A190B3B667CC" box="[904,986,715,741]" country="Czech Republic" httpUri="http://grbio.org/cool/hj12-2xm7" name="National Museum Prague" pageId="3" pageNumber="304">NMPC</collectionCode>
|
||
)
|
||
<emphasis id="B91DEAD8790DE64DBA90A190B0856620" bold="true" pageId="3" pageNumber="304">[first records from Henan province, China]</emphasis>
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation id="3B013C97790DE64DB98DA1ABB45E6620" ID-GBIF-Occurrence="4067802305" box="[246,1074,751,778]" collectionCode="ZSM" country="China" latitude="35.416668" location="Gansu" longLatPrecision="1193" longitude="103.78333" pageId="3" pageNumber="304" specimenCount="1" specimenCount-female="1" stateProvince="Kansu">
|
||
<collectingRegion id="49ADF828790DE64DB98DA1ABB1516623" box="[246,317,752,778]" country="China" name="Gansu" pageId="3" pageNumber="304">Kansu</collectingRegion>
|
||
[
|
||
<location id="8EB66011790DE64DB832A1B4B1FB6620" LSID="urn:lsid:plazi:treatment:03C087DC790DE64DB9ECA3CCB4F762DF:8EB66011790DE64DB832A1B4B1FB6620" box="[329,407,751,777]" country="China" latitude="35.416668" longLatPrecision="1193" longitude="103.78333" name="Gansu" pageId="3" pageNumber="304" stateProvince="Kansu">Gansu</location>
|
||
],
|
||
<location id="8EB66011790DE64DB8DCA1B4B2606620" LSID="urn:lsid:plazi:treatment:03C087DC790DE64DB9ECA3CCB4F762DF:8EB66011790DE64DB8DCA1B4B2606620" box="[423,524,751,777]" country="China" latitude="35.416668" longLatPrecision="1193" longitude="103.78333" name="Tu-kiang" pageId="3" pageNumber="304" stateProvince="Kansu">Tu-kiang</location>
|
||
[
|
||
<geoCoordinate id="EE5D500D790DE64DBB63A1B4B21A6623" box="[536,630,751,778]" degrees="35" direction="north" minutes="25" orientation="latitude" pageId="3" pageNumber="304" precision="925" value="35.416668">35°25'N</geoCoordinate>
|
||
<geoCoordinate id="EE5D500D790DE64DBB07A1B4B28A6620" box="[636,742,751,777]" degrees="103" direction="east" minutes="47" orientation="longitude" pageId="3" pageNumber="304" precision="925" value="103.78333">103°47'E</geoCoordinate>
|
||
],
|
||
<specimenCount id="9D6FFD43790DE64DBB8CA1ABB3606623" box="[759,780,752,778]" pageId="3" pageNumber="304" type="female">♀</specimenCount>
|
||
(
|
||
<collectionCode id="ED78AE0F790DE64DBA62A1B4B33A6620" box="[793,854,751,777]" collectionName="Germany, Muenchen [= Munich], Zoologische Staatssammlung" country="Germany" httpUri="http://grbio.org/cool/4b7r-7sec" name="Bavarian State Collection of Zoology" pageId="3" pageNumber="304" type="Museum">ZSM</collectionCode>
|
||
, ex. coll. O. Hillert)
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation id="3B013C97790DE64DBD47A1B4B0856678" ID-GBIF-Occurrence="4067802306" collectingDate="1996-12-08" collectionCode="NMPC" collectorName="E. Kucera" country="China" latitude="35.566666" location="Dingxi" longLatPrecision="1192" longitude="104.6" pageId="3" pageNumber="304" specimenCount="1" specimenCount-female="1" stateProvince="Gansu">
|
||
<collectingCountry id="F37E765A790DE64DBD47A1B4B4126620" box="[1084,1150,751,777]" name="China" pageId="3" pageNumber="304">China</collectingCountry>
|
||
-
|
||
<collectingRegion id="49ADF828790DE64DBDE9A1B4B4B06620" box="[1170,1244,751,777]" country="China" name="Gansu" pageId="3" pageNumber="304">Gansu</collectingRegion>
|
||
,
|
||
<location id="8EB66011790DE64DBD9EA1B4B55E6620" LSID="urn:lsid:plazi:treatment:03C087DC790DE64DB9ECA3CCB4F762DF:8EB66011790DE64DBD9EA1B4B55E6620" box="[1253,1330,751,777]" country="China" latitude="35.566666" longLatPrecision="1192" longitude="104.6" name="Dingxi" pageId="3" pageNumber="304" stateProvince="Gansu">Dingxi</location>
|
||
[
|
||
<geoCoordinate id="EE5D500D790DE64DBC45A1B4B5F16623" box="[1342,1437,751,778]" degrees="35" direction="north" minutes="34" orientation="latitude" pageId="3" pageNumber="304" precision="925" value="35.566666">35°34'N</geoCoordinate>
|
||
<geoCoordinate id="EE5D500D790DE64DB9ECA048B16E6604" box="[151,258,787,813]" degrees="104" direction="east" minutes="36" orientation="longitude" pageId="3" pageNumber="304" precision="925" value="104.6">104°36'E</geoCoordinate>
|
||
], 7.8.–
|
||
<date id="FFD7100A790DE64DB836A048B1AD6604" box="[333,449,787,813]" pageId="3" pageNumber="304" value="1996-12-08">
|
||
<collectingDate id="EF93E9E2790DE64DB836A048B1AD6604" box="[333,449,787,813]" pageId="3" pageNumber="304" value="1996-12-08">12.8.1996</collectingDate>
|
||
</date>
|
||
,
|
||
<collectorName id="269C531C790DE64DB8B5A04FB22C6604" box="[462,576,787,814]" pageId="3" pageNumber="304">E. Kučera</collectorName>
|
||
leg., Ô
|
||
<specimenCount id="9D6FFD43790DE64DBBE7A04EB2C16619" box="[668,685,789,816]" pageId="3" pageNumber="304" type="female">♀</specimenCount>
|
||
(
|
||
<collectionCode id="ED78AE0F790DE64DBBC5A048B37D6604" box="[702,785,787,813]" country="Czech Republic" httpUri="http://grbio.org/cool/hj12-2xm7" name="National Museum Prague" pageId="3" pageNumber="304">NMPC</collectionCode>
|
||
, ex. coll. D. Král)
|
||
<emphasis id="B91DEAD8790DE64DBA96A048B0856678" bold="true" pageId="3" pageNumber="304">[first records from Gansu province, China]</emphasis>
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation id="3B013C97790DE64DB983A06CB27F665C" ID-GBIF-Occurrence="4067802307" collectingDate="2001-06-19" collectionCode="ZSM" collectorName="E. Kucera" country="China" isTailEnumeration="true" latitude="35.566666" location="Wenchuan" longLatPrecision="1192" longitude="104.6" municipality="Wenchuan" pageId="3" pageNumber="304" specimenCount="1" specimenCount-female="1" stateProvince="Sechuan">
|
||
<collectingCountry id="F37E765A790DE64DB983A06CB1516678" box="[248,317,823,849]" name="China" pageId="3" pageNumber="304">China</collectingCountry>
|
||
,
|
||
<collectingRegion id="49ADF828790DE64DB833A06CB1C56678" box="[328,425,823,849]" pageId="3" pageNumber="304">Sechuan</collectingRegion>
|
||
,
|
||
<location id="8EB66011790DE64DB8CFA06CB2446678" LSID="urn:lsid:plazi:treatment:03C087DC790DE64DB9ECA3CCB4F762DF:8EB66011790DE64DB8CFA06CB2446678" box="[436,552,823,849]" country="China" latitude="35.566666" longLatPrecision="1192" longitude="104.6" municipality="Wenchuan" name="Wenchuan" pageId="3" pageNumber="304" stateProvince="Sechuan">Wenchuan</location>
|
||
[
|
||
<geoCoordinate id="EE5D500D790DE64DBB4DA06CB2FA6678" box="[566,662,823,849]" degrees="31" direction="north" minutes="24" orientation="latitude" pageId="3" pageNumber="304" precision="925" value="31.4">31°24'N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE5D500D790DE64DBBD9A06CB3616678" box="[674,781,823,849]" degrees="103" direction="east" minutes="34" orientation="longitude" pageId="3" pageNumber="304" precision="925" value="103.566666">103°34'E</geoCoordinate>
|
||
], 18.6. –
|
||
<date id="FFD7100A790DE64DBA16A06CB3B26678" box="[877,990,823,849]" pageId="3" pageNumber="304" value="2001-06-19">
|
||
<collectingDate id="EF93E9E2790DE64DBA16A06CB3B26678" box="[877,990,823,849]" pageId="3" pageNumber="304" value="2001-06-19">19.6.2001</collectingDate>
|
||
</date>
|
||
, leg.
|
||
<collectorName id="269C531C790DE64DBD61A063B4E06678" box="[1050,1164,823,850]" pageId="3" pageNumber="304">E. Kučera</collectorName>
|
||
,
|
||
<specimenCount id="9D6FFD43790DE64DBDECA063B4C0667B" box="[1175,1196,824,850]" pageId="3" pageNumber="304" type="female">♀</specimenCount>
|
||
(
|
||
<collectionCode id="ED78AE0F790DE64DBDC0A06CB4956678" box="[1211,1273,823,849]" country="Germany" httpUri="http://grbio.org/cool/4b7r-7sec" name="Bavarian State Collection of Zoology" pageId="3" pageNumber="304" type="Museum">ZSM</collectionCode>
|
||
,)
|
||
<emphasis id="B91DEAD8790DE64DBC76A06CB27F665C" bold="true" pageId="3" pageNumber="304">[first record from Sichuan province, China]</emphasis>
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation id="3B013C97790DE64DBB5DA000B52C66B0" ID-GBIF-Occurrence="4067802301" collectingDate="2001-07-04" collectingDateMax="2001-07-05" collectingDateMin="2001-07-04" collectionCode="NMPC" collectorName="Jaroslav Turna" country="China" isTailEnumeration="true" latitude="31.4" location="Chao Shan" longLatPrecision="1216" longitude="103.566666" municipality="Wenchuan" pageId="3" pageNumber="304" specimenCount="1" stateProvince="Shanxi">
|
||
<collectingCountry id="F37E765A790DE64DBB5DA000B207665C" box="[550,619,859,885]" name="China" pageId="3" pageNumber="304">China</collectingCountry>
|
||
, centr.
|
||
<collectingRegion id="49ADF828790DE64DBBB8A000B37F665C" box="[707,787,859,885]" country="China" name="Shanxi" pageId="3" pageNumber="304">Shanxi</collectingRegion>
|
||
,
|
||
<location id="8EB66011790DE64DBA59A000B3CE665C" LSID="urn:lsid:plazi:treatment:03C087DC790DE64DB9ECA3CCB4F762DF:8EB66011790DE64DBA59A000B3CE665C" box="[802,930,859,885]" country="China" latitude="31.4" longLatPrecision="1216" longitude="103.566666" municipality="Wenchuan" name="Chao Shan" pageId="3" pageNumber="304" stateProvince="Shanxi">Chao Shan</location>
|
||
,
|
||
<geoCoordinate id="EE5D500D790DE64DBAC9A000B462665C" box="[946,1038,859,885]" degrees="37" direction="north" minutes="06" orientation="latitude" pageId="3" pageNumber="304" precision="925" value="37.1">37°06'N</geoCoordinate>
|
||
<geoCoordinate id="EE5D500D790DE64DBD62A000B4ED665C" box="[1049,1153,859,885]" degrees="112" direction="east" minutes="24" orientation="longitude" pageId="3" pageNumber="304" precision="925" value="112.4">112°24'E</geoCoordinate>
|
||
,
|
||
<locationDeviation id="9A642FE8790DE64DBDEAA000B50F665C" box="[1169,1379,859,886]" location="Pingyao" pageId="3" pageNumber="304">
|
||
<quantity id="4C919B2F790DE64DBDEAA000B4BC665F" box="[1169,1232,859,886]" metricMagnitude="4" metricUnit="m" metricValue="2.5" pageId="3" pageNumber="304" unit="km" value="25.0">25km</quantity>
|
||
SE Pingyao
|
||
</locationDeviation>
|
||
<date id="FFD7100A790DE64DBC15A000B17866B3" pageId="3" pageNumber="304" value="2001-07-04" valueMax="2001-07-05" valueMin="2001-07-04">
|
||
<collectingDate id="EF93E9E2790DE64DBC15A000B17866B3" pageId="3" pageNumber="304" value="2001-07-04" valueMax="2001-07-05" valueMin="2001-07-04">4. – 5.VII.2001</collectingDate>
|
||
</date>
|
||
,
|
||
<collectorName id="269C531C790DE64DB864A024B1AD66B3" box="[287,449,895,922]" pageId="3" pageNumber="304">Jaroslav Turna</collectorName>
|
||
leg., Ô (
|
||
<collectionCode id="ED78AE0F790DE64DBB59A024B21966B0" box="[546,629,895,921]" country="Czech Republic" httpUri="http://grbio.org/cool/hj12-2xm7" name="National Museum Prague" pageId="3" pageNumber="304">NMPC</collectionCode>
|
||
, ex. coll. D. Král)
|
||
<emphasis id="B91DEAD8790DE64DBA38A024B52C66B0" bold="true" box="[835,1344,895,921]" pageId="3" pageNumber="304">[first record from Shanxi province, China].</emphasis>
|
||
</materialsCitation>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3736541790DE64DB9BCA0F8B4DB62F8" pageId="3" pageNumber="304" type="description">
|
||
<paragraph id="8BD636CA790DE64DB9BCA0F8B46066C8" blockId="3.[151,1437,570,2038]" pageId="3" pageNumber="304">
|
||
<emphasis id="B91DEAD8790DE64DB9BCA0F8B1016694" bold="true" box="[199,365,931,957]" pageId="3" pageNumber="304">Redescription</emphasis>
|
||
(based on the male from Monan). Body length 16.0 mm. Dorsal surface black, without any metallic lustre or shiny parts (probably present in freshly collected specimens).
|
||
</paragraph>
|
||
<paragraph id="8BD636CA790DE64DB9BCA0B0B1D961F7" blockId="3.[151,1437,570,2038]" pageId="3" pageNumber="304">
|
||
<emphasis id="B91DEAD8790DE64DB9BCA0B0B168612C" box="[199,260,1003,1029]" italics="true" pageId="3" pageNumber="304">Head</emphasis>
|
||
(
|
||
<figureCitation id="13522A4F790DE64DB86DA0B0B1A1612C" box="[278,461,1003,1029]" captionStart="FIGURES 13–17" captionStartId="7.[151,264,1945,1969]" captionTargetBox="[297,1291,1108,1909]" captionTargetId="figure-393@7.[284,1304,1096,1922]" captionTargetPageId="7" captionText="FIGURES 13–17. Head of Lethrus species with mandibular appendix, keels and hornlike apophyses of mandibles (males), schematically. 13–14, 17 L. (Sinolethrus) serpentifer Semenov & Medvedev, 1935, (13 frontal view, 14 lateral view, 17 dorsal view); 15 L. (Ceratodirus) cephalotes Pallas, 1771 (dorsal view); 16 L. (Ceratodirus) karelini Gebler, 1845 (dorsal view)." figureDoi="http://doi.org/10.5281/zenodo.7780401" httpUri="https://zenodo.org/record/7780401/files/figure.png" pageId="3" pageNumber="304">Figs. 13–14, 17</figureCitation>
|
||
,
|
||
<figureCitation id="13522A4F790DE64DB8A7A0B0B24B612F" box="[476,551,1003,1030]" captionStart-0="FIGURES 55–58" captionStart-1="FIGURES 59–65" captionStartId-0="11.[151,264,1556,1580]" captionStartId-1="12.[151,264,1624,1648]" captionTargetBox-0="[171,1415,1061,1519]" captionTargetBox-1="[201,1392,189,1588]" captionTargetId-0="figure-78@11.[154,1434,1040,1532]" captionTargetId-1="figure-34@12.[195,1392,181,1600]" captionTargetPageId-0="11" captionTargetPageId-1="12" captionText-0="FIGURES 55–58. Labels, apical spurs and head of males of Lethrus species, schematically. 55 apical spur of front tibia of L. (Sinolethrus) serpentifer Semenov & Medvedev, 1935; 56 apical spur of front tibia of L. (Ceratodirus) karelini Gebler, 1845; 57 label of syntype of L. serpentifer; 58 label of lectotype of L. mediocris." captionText-1="FIGURES 59–65. Lethrus (Sinolethrus) serpentifer Semenov & Medvedev, 1935 (Huang He, China). 59 male dorsal view; 60 male ventral view; 61 female dorsal view; 62 male, lateral view, right; 63 male, lateral view, left; 64 male, frontal view; 65 head with mentum of male, schematically." figureDoi-0="http://doi.org/10.5281/zenodo.7780421" figureDoi-1="http://doi.org/10.5281/zenodo.7780423" httpUri-0="https://zenodo.org/record/7780421/files/figure.png" httpUri-1="https://zenodo.org/record/7780423/files/figure.png" pageId="3" pageNumber="304">58–64</figureCitation>
|
||
). Labrum bilobed, asymmetrical, right lobe slightly more developed; surface rugosely, shallowly and sparsely punctate; anterior margin with dense row of long macrosetae. Clypeus transverse, nearly semicircular. Eye canthus exceeding eyes distinctly, projecting anterolaterad, indistinctly semicircular, interrupted, anterolateral angle angular, oblique keel above eyes absent. Pleurostomal process evenly rounded, not exceeding ventrolateral mandibular outline. Punctation of clypeus, frons and occiput similarly impressed, consisting of distinct but weakly regularly distanced punctures, intermixed with fine and also regularly distributed ones; eye canthus distinctly rugose.
|
||
</paragraph>
|
||
<paragraph id="8BD636CA790DE64DB9BCA7B3B166600F" blockId="3.[151,1437,570,2038]" pageId="3" pageNumber="304">
|
||
<emphasis id="B91DEAD8790DE64DB9BCA7B3B14F6028" box="[199,291,1256,1281]" italics="true" pageId="3" pageNumber="304">Mentum</emphasis>
|
||
strongly convex in lateral aspect (
|
||
<figureCitation id="13522A4F790DE64DBBCEA7BCB3546028" box="[693,824,1255,1281]" captionStart="FIGURES 59–65" captionStartId="12.[151,264,1624,1648]" captionTargetBox="[201,1392,189,1588]" captionTargetId="figure-34@12.[195,1392,181,1600]" captionTargetPageId="12" captionText="FIGURES 59–65. Lethrus (Sinolethrus) serpentifer Semenov & Medvedev, 1935 (Huang He, China). 59 male dorsal view; 60 male ventral view; 61 female dorsal view; 62 male, lateral view, right; 63 male, lateral view, left; 64 male, frontal view; 65 head with mentum of male, schematically." figureDoi="http://doi.org/10.5281/zenodo.7780423" httpUri="https://zenodo.org/record/7780423/files/figure.png" pageId="3" pageNumber="304">Fig. 61, 64</figureCitation>
|
||
), with distinct concavity basally, shortly and densely setaceous.
|
||
</paragraph>
|
||
<paragraph id="8BD636CA790DE64DB9BCA674B4B56308" blockId="3.[151,1437,570,2038]" pageId="3" pageNumber="304">
|
||
<emphasis id="B91DEAD8790DE64DB9BCA674B1516060" box="[199,317,1327,1353]" italics="true" pageId="3" pageNumber="304">Mandibles</emphasis>
|
||
with asymmetrical mandibular appendices, left being long, angled nearly 90° downward and from half of length bending regularly forward in lateral aspect (serpent like), s-shaped in frontal aspect; right appendix only weakly developed, visibly as a weakly elevation. Exterior mandibular outline straight from base to apex in dorsal aspect (
|
||
<figureCitation id="13522A4F790DE64DB841A6C0B182609C" box="[314,494,1435,1461]" captionStart="FIGURES 13–17" captionStartId="7.[151,264,1945,1969]" captionTargetBox="[297,1291,1108,1909]" captionTargetId="figure-393@7.[284,1304,1096,1922]" captionTargetPageId="7" captionText="FIGURES 13–17. Head of Lethrus species with mandibular appendix, keels and hornlike apophyses of mandibles (males), schematically. 13–14, 17 L. (Sinolethrus) serpentifer Semenov & Medvedev, 1935, (13 frontal view, 14 lateral view, 17 dorsal view); 15 L. (Ceratodirus) cephalotes Pallas, 1771 (dorsal view); 16 L. (Ceratodirus) karelini Gebler, 1845 (dorsal view)." figureDoi="http://doi.org/10.5281/zenodo.7780401" httpUri="https://zenodo.org/record/7780401/files/figure.png" pageId="3" pageNumber="304">Figs. 13–14, 17</figureCitation>
|
||
,
|
||
<figureCitation id="13522A4F790DE64DB880A6C0B2F6609C" box="[507,666,1435,1462]" captionStart-0="FIGURES 55–58" captionStart-1="FIGURES 59–65" captionStartId-0="11.[151,264,1556,1580]" captionStartId-1="12.[151,264,1624,1648]" captionTargetBox-0="[171,1415,1061,1519]" captionTargetBox-1="[201,1392,189,1588]" captionTargetId-0="figure-78@11.[154,1434,1040,1532]" captionTargetId-1="figure-34@12.[195,1392,181,1600]" captionTargetPageId-0="11" captionTargetPageId-1="12" captionText-0="FIGURES 55–58. Labels, apical spurs and head of males of Lethrus species, schematically. 55 apical spur of front tibia of L. (Sinolethrus) serpentifer Semenov & Medvedev, 1935; 56 apical spur of front tibia of L. (Ceratodirus) karelini Gebler, 1845; 57 label of syntype of L. serpentifer; 58 label of lectotype of L. mediocris." captionText-1="FIGURES 59–65. Lethrus (Sinolethrus) serpentifer Semenov & Medvedev, 1935 (Huang He, China). 59 male dorsal view; 60 male ventral view; 61 female dorsal view; 62 male, lateral view, right; 63 male, lateral view, left; 64 male, frontal view; 65 head with mentum of male, schematically." figureDoi-0="http://doi.org/10.5281/zenodo.7780421" figureDoi-1="http://doi.org/10.5281/zenodo.7780423" httpUri-0="https://zenodo.org/record/7780421/files/figure.png" httpUri-1="https://zenodo.org/record/7780423/files/figure.png" pageId="3" pageNumber="304">58–59, 61–64</figureCitation>
|
||
), with maximum width approximately at base. Length of mandibles distinctly less than length of head, dorsal surface of right mandible absent from any elevations or protuberance, dorsal surface of left mandible with weakly arcuate protuberance at middle of length, shortly anteriorly of labrum, directed to the side of right mandible. Apex of both mandibles absent of subapical protuberances.
|
||
</paragraph>
|
||
<paragraph id="8BD636CA790DE64DB9BCA577B54063A4" blockId="3.[151,1437,570,2038]" pageId="3" pageNumber="304">
|
||
<emphasis id="B91DEAD8790DE64DB9BCA577B15A636C" box="[199,310,1580,1605]" italics="true" pageId="3" pageNumber="304">Pronotum</emphasis>
|
||
transverse, equal as base of elytra, broadest just posteriorly of middle (
|
||
<figureCitation id="13522A4F790DE64DBD05A570B4B9636F" box="[1150,1237,1579,1606]" captionStart="FIGURES 55–58" captionStartId="11.[151,264,1556,1580]" captionTargetBox="[171,1415,1061,1519]" captionTargetId="figure-78@11.[154,1434,1040,1532]" captionTargetPageId="11" captionText="FIGURES 55–58. Labels, apical spurs and head of males of Lethrus species, schematically. 55 apical spur of front tibia of L. (Sinolethrus) serpentifer Semenov & Medvedev, 1935; 56 apical spur of front tibia of L. (Ceratodirus) karelini Gebler, 1845; 57 label of syntype of L. serpentifer; 58 label of lectotype of L. mediocris." figureDoi="http://doi.org/10.5281/zenodo.7780421" httpUri="https://zenodo.org/record/7780421/files/figure.png" pageId="3" pageNumber="304">Fig. 58</figureCitation>
|
||
); margin entirely bordered, slightly crenulate. Anterior angles with evenly rounded outline. Punctation of dorsal surface simple, distinct, weakly irregular, punctures separated by one diameter, intermixed with fine simple punctuation.
|
||
</paragraph>
|
||
<paragraph id="8BD636CA790DE64DB9BCA5CCB3296398" blockId="3.[151,1437,570,2038]" box="[199,837,1687,1714]" pageId="3" pageNumber="304">
|
||
<emphasis id="B91DEAD8790DE64DB9BCA5CCB1596398" box="[199,309,1687,1713]" italics="true" pageId="3" pageNumber="304">Scutellum</emphasis>
|
||
(
|
||
<figureCitation id="13522A4F790DE64DB83EA5CCB1FA639B" box="[325,406,1687,1714]" captionStart="FIGURES 55–58" captionStartId="11.[151,264,1556,1580]" captionTargetBox="[171,1415,1061,1519]" captionTargetId="figure-78@11.[154,1434,1040,1532]" captionTargetPageId="11" captionText="FIGURES 55–58. Labels, apical spurs and head of males of Lethrus species, schematically. 55 apical spur of front tibia of L. (Sinolethrus) serpentifer Semenov & Medvedev, 1935; 56 apical spur of front tibia of L. (Ceratodirus) karelini Gebler, 1845; 57 label of syntype of L. serpentifer; 58 label of lectotype of L. mediocris." figureDoi="http://doi.org/10.5281/zenodo.7780421" httpUri="https://zenodo.org/record/7780421/files/figure.png" pageId="3" pageNumber="304">Fig. 58</figureCitation>
|
||
) widely triangular, strong shagreened.
|
||
</paragraph>
|
||
<paragraph id="8BD636CA790DE64DB9BCA5E0B37263D0" blockId="3.[151,1437,570,2038]" pageId="3" pageNumber="304">
|
||
<emphasis id="B91DEAD8790DE64DB9BCA5E0B16163FC" box="[199,269,1723,1749]" italics="true" pageId="3" pageNumber="304">Elytra</emphasis>
|
||
(
|
||
<figureCitation id="13522A4F790DE64DB867A5E0B10163FF" box="[284,365,1723,1750]" captionStart="FIGURES 55–58" captionStartId="11.[151,264,1556,1580]" captionTargetBox="[171,1415,1061,1519]" captionTargetId="figure-78@11.[154,1434,1040,1532]" captionTargetPageId="11" captionText="FIGURES 55–58. Labels, apical spurs and head of males of Lethrus species, schematically. 55 apical spur of front tibia of L. (Sinolethrus) serpentifer Semenov & Medvedev, 1935; 56 apical spur of front tibia of L. (Ceratodirus) karelini Gebler, 1845; 57 label of syntype of L. serpentifer; 58 label of lectotype of L. mediocris." figureDoi="http://doi.org/10.5281/zenodo.7780421" httpUri="https://zenodo.org/record/7780421/files/figure.png" pageId="3" pageNumber="304">Fig. 58</figureCitation>
|
||
) almost semicircular, apices not prominent, apex of each elytron forming dependent arcus, with indistinct striae, with broad, distinct shagreened intervals.
|
||
</paragraph>
|
||
<paragraph id="8BD636CA790DE64DB9BCA458B4716237" blockId="3.[151,1437,570,2038]" box="[199,1053,1795,1822]" pageId="3" pageNumber="304">
|
||
<emphasis id="B91DEAD8790DE64DB9BCA458B1286234" box="[199,324,1795,1821]" italics="true" pageId="3" pageNumber="304">Epipleuron</emphasis>
|
||
strongly narrowed apicad, epipleural keel reaching elytron apex.
|
||
</paragraph>
|
||
<paragraph id="8BD636CA790DE64DB9BCA473B5F7624F" blockId="3.[151,1437,570,2038]" pageId="3" pageNumber="304">
|
||
<emphasis id="B91DEAD8790DE64DB9BCA473B0976268" box="[199,251,1832,1857]" italics="true" pageId="3" pageNumber="304">Legs</emphasis>
|
||
. Profemur not armed, protibia with eight gradually diminishing external denticles proximad, and with row of shallow tubercles on ventromedial edge. Apical spur of protibia without subbasal process, weakly s-shaped (
|
||
<figureCitation id="13522A4F790DE64DBC46A410B5E3624F" box="[1341,1423,1867,1894]" captionStart="FIGURES 55–58" captionStartId="11.[151,264,1556,1580]" captionTargetBox="[171,1415,1061,1519]" captionTargetId="figure-78@11.[154,1434,1040,1532]" captionTargetPageId="11" captionText="FIGURES 55–58. Labels, apical spurs and head of males of Lethrus species, schematically. 55 apical spur of front tibia of L. (Sinolethrus) serpentifer Semenov & Medvedev, 1935; 56 apical spur of front tibia of L. (Ceratodirus) karelini Gebler, 1845; 57 label of syntype of L. serpentifer; 58 label of lectotype of L. mediocris." figureDoi="http://doi.org/10.5281/zenodo.7780421" httpUri="https://zenodo.org/record/7780421/files/figure.png" pageId="3" pageNumber="304">Fig. 55</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BD636CA790DE64DB9BCA434B18A62A0" blockId="3.[151,1437,570,2038]" box="[199,486,1903,1930]" pageId="3" pageNumber="304">
|
||
<emphasis id="B91DEAD8790DE64DB9BCA434B15862A0" box="[199,308,1903,1929]" italics="true" pageId="3" pageNumber="304">Aedeagus</emphasis>
|
||
as in
|
||
<figureCitation id="13522A4F790DE64DB80DA434B18D62A0" box="[374,481,1903,1929]" captionStart="FIGURES 1–4" captionStartId="2.[151,264,1867,1891]" captionTargetBox="[166,1429,751,1823]" captionTargetId="figure-229@2.[151,1436,734,1841]" captionTargetPageId="2" captionText="FIGURES 1–4. Aedeagus, schematically. 1–2 L. (Ceratodirus) dostojewskii Semenov, 1899 (1 lateral view, 2 dorsal view); 3–4 L. (Sinolethrus) serpentifer Semenov & Medvedev, 1935 (3 lateral view, 4 dorsal view)." figureDoi="http://doi.org/10.5281/zenodo.7780395" httpUri="https://zenodo.org/record/7780395/files/figure.png" pageId="3" pageNumber="304">Figs. 3–4</figureCitation>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8BD636CA790DE64DB9BCA4C8B4DB62F8" blockId="3.[151,1437,570,2038]" pageId="3" pageNumber="304">
|
||
<emphasis id="B91DEAD8790DE64DB9BCA4C8B1456284" bold="true" box="[199,297,1939,1965]" pageId="3" pageNumber="304">Females</emphasis>
|
||
differ from males as follows: body broader, ventral mandibular appendices absent; hornlike dorsal protuberans of mandibular surface absent, apical spur of protibia simple and straight (
|
||
<figureCitation id="13522A4F790DE64DBD2CA4ECB4C762F8" box="[1111,1195,1975,2001]" captionStart="FIGURES 59–65" captionStartId="12.[151,264,1624,1648]" captionTargetBox="[201,1392,189,1588]" captionTargetId="figure-34@12.[195,1392,181,1600]" captionTargetPageId="12" captionText="FIGURES 59–65. Lethrus (Sinolethrus) serpentifer Semenov & Medvedev, 1935 (Huang He, China). 59 male dorsal view; 60 male ventral view; 61 female dorsal view; 62 male, lateral view, right; 63 male, lateral view, left; 64 male, frontal view; 65 head with mentum of male, schematically." figureDoi="http://doi.org/10.5281/zenodo.7780423" httpUri="https://zenodo.org/record/7780423/files/figure.png" pageId="3" pageNumber="304">Fig. 60</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3736541790DE64DB9BCA480B4F762DF" box="[199,1179,2011,2038]" pageId="3" pageNumber="304" type="distribution">
|
||
<paragraph id="8BD636CA790DE64DB9BCA480B4F762DF" blockId="3.[151,1437,570,2038]" box="[199,1179,2011,2038]" pageId="3" pageNumber="304">
|
||
<emphasis id="B91DEAD8790DE64DB9BCA480B10D62DC" bold="true" box="[199,353,2011,2037]" pageId="3" pageNumber="304">Distribution.</emphasis>
|
||
<collectingCountry id="F37E765A790DE64DB81CA480B1C662DC" box="[359,426,2011,2037]" name="China" pageId="3" pageNumber="304">China</collectingCountry>
|
||
(
|
||
<collectingRegion id="49ADF828790DE64DB8C2A480B26962DC" box="[441,517,2011,2037]" country="China" name="Gansu" pageId="3" pageNumber="304">Gansu</collectingRegion>
|
||
,
|
||
<collectingRegion id="49ADF828790DE64DBB6BA487B23062DF" box="[528,604,2012,2038]" country="China" name="Henan" pageId="3" pageNumber="304">Henan</collectingRegion>
|
||
,
|
||
<collectingRegion id="49ADF828790DE64DBB1CA480B2A862DC" box="[615,708,2011,2037]" country="China" name="Qinghai" pageId="3" pageNumber="304">Qinghai</collectingRegion>
|
||
,
|
||
<collectingRegion id="49ADF828790DE64DBBB4A480B37162DC" box="[719,797,2011,2037]" country="China" name="Shanxi" pageId="3" pageNumber="304">Shanxi</collectingRegion>
|
||
and
|
||
<collectingRegion id="49ADF828790DE64DBA2FA480B3C262DC" box="[852,942,2011,2037]" country="China" name="Sichuan" pageId="3" pageNumber="304">Sichuan</collectingRegion>
|
||
provinces) (
|
||
<figureCitation id="13522A4F790DE64DBD41A480B4E362DF" box="[1082,1167,2011,2038]" captionStart="FIGURES 59–65" captionStartId="12.[151,264,1624,1648]" captionTargetBox="[201,1392,189,1588]" captionTargetId="figure-34@12.[195,1392,181,1600]" captionTargetPageId="12" captionText="FIGURES 59–65. Lethrus (Sinolethrus) serpentifer Semenov & Medvedev, 1935 (Huang He, China). 59 male dorsal view; 60 male ventral view; 61 female dorsal view; 62 male, lateral view, right; 63 male, lateral view, left; 64 male, frontal view; 65 head with mentum of male, schematically." figureDoi="http://doi.org/10.5281/zenodo.7780423" httpUri="https://zenodo.org/record/7780423/files/figure.png" pageId="3" pageNumber="304">Fig. 65</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |