162 lines
21 KiB
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162 lines
21 KiB
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<document id="E1A394C28CCEA83FDECF7B4255FBF514" ID-CLB-Dataset="5634" ID-DOI="10.11646/zootaxa.3786.5.1" ID-GBIF-Dataset="cdd39956-de72-43ea-afa3-cf79f805dd83" ID-ISSN="1175-5326" ID-Zenodo-Dep="4913561" ID-ZooBank="D2764982-F7D7-4922-BF3F-8314FE9FD869" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1623171832930" checkinUser="felipe" docAuthor="Schoddei, Richard & Christidis, Les" docDate="2014" docId="03C087B55B6DA840FF75FA68FCD6FE63" docLanguage="en" docName="zt03786p522.pdf" docOrigin="Zootaxa 3786 (5)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Rhagologus Stresemann & Paludan 1934" docType="treatment" docVersion="6" lastPageNumber="509" masterDocId="FFF9FFCD5B65A849FFE2FFADFF95FFB3" masterDocTitle="Relicts from Tertiary Australasia: undescribed families and subfamilies of songbirds (Passeriformes) and their zoogeographic signal" masterLastPageNumber="522" masterPageNumber="501" pageNumber="509" updateTime="1699005125857" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="1C5E9D62B3D9869FB89023DF093A2C70">Relicts from Tertiary Australasia: undescribed families and subfamilies of songbirds (Passeriformes) and their zoogeographic signal</mods:title>
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<mods:namePart id="3BB1763B343B3C2201A2E3D158AB8284">Schoddei, Richard</mods:namePart>
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<mods:namePart id="7352EF6E773D28E36077A731C4D59454">Christidis, Les</mods:namePart>
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<mods:title id="DFCA3F75007374B4B244C79185CC1CE0">Zootaxa</mods:title>
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<mods:date id="AAA883775A71BBADD44026045F008CFE">2014</mods:date>
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<mods:number id="45BD577BD4084D45EE9337EF7F0A7DEA">2014-04-14</mods:number>
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<mods:number id="A010AE07AB5C82A57A4181CA777B129F">3786</mods:number>
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<treatment id="03C087B55B6DA840FF75FA68FCD6FE63" ID-DOI="http://doi.org/10.5281/zenodo.4913567" ID-GBIF-Taxon="183401426" ID-Zenodo-Dep="4913567" LSID="urn:lsid:plazi:treatment:03C087B55B6DA840FF75FA68FCD6FE63" httpUri="http://treatment.plazi.org/id/03C087B55B6DA840FF75FA68FCD6FE63" lastPageId="9" lastPageNumber="509" pageId="8" pageNumber="509">
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<subSubSection id="C37365285B6DA841FF75FA68FED0FA6C" box="[151,325,1477,1503]" pageId="8" pageNumber="509" type="nomenclature">
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<paragraph id="8BD636A35B6DA841FF75FA68FED0FA6C" blockId="8.[151,325,1477,1503]" box="[151,325,1477,1503]" pageId="8" pageNumber="509">
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<heading id="D09E81CF5B6DA841FF75FA68FED0FA6C" bold="true" box="[151,325,1477,1503]" fontSize="11" level="1" pageId="8" pageNumber="509" reason="1">
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<emphasis id="B91DEAB15B6DA841FF75FA68FED0FA6C" bold="true" box="[151,325,1477,1503]" pageId="8" pageNumber="509">
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3.
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<taxonomicName id="4C694D205B6DA841FF56FA68FED0FA6C" ID-CoL="78BG" authorityName="Stresemann & Paludan" authorityYear="1934" box="[180,325,1477,1503]" class="Aves" family="Pachycephalidae" genus="Rhagologus" kingdom="Animalia" order="Passeriformes" pageId="8" pageNumber="509" phylum="Chordata" rank="genus">
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<emphasis id="B91DEAB15B6DA841FF56FA68FED0FA6C" bold="true" box="[180,325,1477,1503]" italics="true" pageId="8" pageNumber="509">Rhagologus</emphasis>
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</taxonomicName>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C37365285B6DA840FF75F9A1FCD6FE63" lastPageId="9" lastPageNumber="510" pageId="8" pageNumber="509" type="description">
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<paragraph id="8BD636A35B6DA841FF75F9A1FE38F83E" blockId="8.[151,1437,1548,2005]" pageId="8" pageNumber="509">
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Despite its streak-plumaged females, New Guinean
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<taxonomicName id="4C694D205B6DA841FD1EF9A1FC17F996" authorityName="Stresemann & Paludan" authorityYear="1934" box="[764,898,1548,1573]" class="Aves" family="Pachycephalidae" genus="Rhagologus" kingdom="Animalia" order="Passeriformes" pageId="8" pageNumber="509" phylum="Chordata" rank="genus">
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<emphasis id="B91DEAB15B6DA841FD1EF9A1FC17F996" box="[764,898,1548,1573]" italics="true" pageId="8" pageNumber="509">Rhagologus</emphasis>
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</taxonomicName>
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has traditionally been included in the family
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<taxonomicName id="4C694D205B6DA841FF75F982FEC6F9FB" authorityName="Swainson" authorityYear="1831" box="[151,339,1583,1608]" class="Aves" family="Pachycephalidae" kingdom="Animalia" order="Passeriformes" pageId="8" pageNumber="509" phylum="Chordata" rank="family">Pachycephalidae</taxonomicName>
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(Australasian whistlers) because of its whistler-like appearance (
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<emphasis id="B91DEAB15B6DA841FBCDF99CFBC5F9FB" box="[1071,1104,1585,1608]" italics="true" pageId="8" pageNumber="509">e.g</emphasis>
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.
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<bibRefCitation id="EFF84B525B6DA841FBBDF982FA98F9FB" author="Salvadori, T." box="[1119,1293,1583,1608]" pageId="8" pageNumber="509" pagination="896 - 976" refId="ref19108" refString="Salvadori, T. (1876) Descrizione di cinquantotto nuove specie di uccelli, ed osservazioni intorno ad altre poco note, della Nuova Guinea e di altre Isole Papuane, raccolte del Dr. Odoardo Beccari e dai cacciatore del. Sig. A. A. Bruijn. Annali del Museo civico di storia naturale di Genova, 7 (56 / 61), 896 - 976." type="journal article" year="1876">Salvadori 1876</bibRefCitation>
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;
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<bibRefCitation id="EFF84B525B6DA841FAFBF99DFA02F9FB" author="Mayr, E." box="[1305,1431,1583,1608]" pageId="8" pageNumber="509" refId="ref18185" refString="Mayr, E. (1941) List of New Guinea Birds. American Museum of Natural History, New York, 260 pp." type="book" year="1941">Mayr 1941</bibRefCitation>
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,
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<bibRefCitation id="EFF84B525B6DA841FF75F9F9FF46F9DE" author="Mayr, E." box="[151,211,1620,1645]" pageId="8" pageNumber="509" pagination="3 - 51" refId="ref18210" refString="Mayr, E. (1967) Subfamily Pachycephalinae. In: Paynter, R. A. Jr. (Ed.), Check-list of Birds of the World, A Continuation of the Work of James L. Peters. Vol. 12. Museum of Comparative Zoology, Cambridge, Massachusetts, pp. 3 - 51." type="book chapter" year="1967">1967</bibRefCitation>
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;
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<bibRefCitation id="EFF84B525B6DA841FF05F9F9FE6EF9DE" author="Rand, A. & Gilliard, E. T." box="[231,507,1620,1645]" pageId="8" pageNumber="509" refId="ref18830" refString="Rand, A. L & Gilliard, E. T. (1967) Handbook of New Guinea Birds. Weidenfeld & Nicolson, London, 611 pp." type="book" year="1967">Rand & Gilliard 1967</bibRefCitation>
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;
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<bibRefCitation id="EFF84B525B6DA841FDEDF9F9FCA5F9DE" author="Sibley, C. E. & Monroe, B. L. Jr." box="[527,816,1620,1645]" pageId="8" pageNumber="509" refId="ref19862" refString="Sibley, C. E. & Monroe, B. L. Jr. (1990) Distribution and Taxonomy of Birds of the World. Yale University Press, New Haven, 1111 pp." type="book" year="1990">Sibley & Monroe 1990</bibRefCitation>
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;
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<bibRefCitation id="EFF84B525B6DA841FCA7F9F9FB96F9DE" author="Dickinson, E. C." box="[837,1027,1620,1645]" pageId="8" pageNumber="509" refId="ref16665" refString="Dickinson, E. C. (Ed.) (2003) The Howard and Moore Complete Checklist of the Birds of the World. 3 rd Edition. Christopher Helm, London, 1039 pp." type="book" year="2003">Dickinson 2003</bibRefCitation>
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;
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<bibRefCitation id="EFF84B525B6DA841FBFAF9F9FB24F9DE" author="Boles, W. E." box="[1048,1201,1620,1645]" pageId="8" pageNumber="509" pagination="374 - 437" refId="ref15799" refString="Boles, W. E. (2007 a) Family Pachycephalidae (whistlers). In: del Hoyo, J., Elliott, A. & Christie, D. (Eds.), Handbook of the Birds of the World. Vol. 12. Lynx Edicions, Barcelona, pp. 374 - 437." type="book chapter" year="2007">Boles 2007a</bibRefCitation>
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). Early DNA-DNA hybridization work (
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<bibRefCitation id="EFF84B525B6DA841FE9CF9DAFD07F923" author="Sibley, C. G. & Ahlquist, J. E." box="[382,658,1655,1680]" pageId="8" pageNumber="509" pagination="251 - 255" refId="ref19636" refString="Sibley, C. G. & Ahlquist, J. E. (1982) The relationships of the Australasian whistlers Pachycephala as indicated by DNA hybridization. Emu, 82, 251 - 255. http: // dx. doi. org / 10.1071 / mu 9820199" type="journal article" year="1982">Sibley & Ahlquist 1982</bibRefCitation>
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,
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<bibRefCitation id="EFF84B525B6DA841FD7DF9DAFD4CF923" author="Sibley, C. G. & Ahlquist, J. E." box="[671,729,1655,1680]" pageId="8" pageNumber="509" pagination="1 - 14" refId="ref19731" refString="Sibley, C. G. & Ahlquist, J. E. (1985) The phylogeny and classification of the Australo-Papuan passerine birds. Emu, 85, 1 - 14. http: // dx. doi. org / 10.1071 / mu 9850001" type="journal article" year="1985">1985</bibRefCitation>
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, 1990) lent support to that presumption. Nevertheless, all five multi-locus DNA sequence studies that have screened
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<taxonomicName id="4C694D205B6DA841FD18F931FC15F906" authorityName="Stresemann & Paludan" authorityYear="1934" box="[762,896,1692,1717]" class="Aves" family="Pachycephalidae" genus="Rhagologus" kingdom="Animalia" order="Passeriformes" pageId="8" pageNumber="509" phylum="Chordata" rank="genus">
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<emphasis id="B91DEAB15B6DA841FD18F931FC15F906" box="[762,896,1692,1717]" italics="true" pageId="8" pageNumber="509">Rhagologus</emphasis>
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</taxonomicName>
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(
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<bibRefCitation id="EFF84B525B6DA841FC6CF931FBFFF906" author="Jonsson, K. A. & Bowie, R. C. K. & Norman, J. A. & Christidis, L. & Fjelds a, J." box="[910,1130,1692,1717]" pageId="8" pageNumber="509" pagination="71 - 74" refId="ref17586" refString="Jonsson, K. A., Bowie, R. C. K., Norman, J. A., Christidis, L. & Fjelds a, J. (2007) Polyphyletic origin of toxic pitohui birds suggests widespread occurrence of toxicity in corvoid birds. Biology Letters, 4, 71 - 74. http: // dx. doi. org / 10.1098 / rsbl. 2007.0464" type="journal article" year="2007">
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Jønsson
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<emphasis id="B91DEAB15B6DA841FC12F930FBB5F906" box="[1008,1056,1692,1717]" italics="true" pageId="8" pageNumber="509">et al</emphasis>
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. 2007
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</bibRefCitation>
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,
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<bibRefCitation id="EFF84B525B6DA841FB97F931FB25F906" author="Jonsson, K. A. & Bowie, R. C. K. & Nylander, J. A. A. & Christidis, L. & Norman, J. A. & Fjeldsa, J." box="[1141,1200,1692,1717]" pageId="8" pageNumber="509" pagination="1767 - 1781" refId="ref17767" refString="Jonsson, K. A., Bowie, R. C. K., Nylander, J. A. A., Christidis, L., Norman, J. A. & Fjeldsa, J. (2010) Biogeographical history of cuckoo-shrikes (Aves: Passeriformes): transoceanic colonization of Africa from Australo-Papua. Journal of Biogeography, 37, 1767 - 1781. http: // dx. doi. org / 10.1111 / j. 1365 - 2699.2010.02328. x" type="journal article" year="2010">2010</bibRefCitation>
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,
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<bibRefCitation id="EFF84B525B6DA841FB59F931FB60F906" author="Jonsson, K. A. & Fabre, P. - H. & Ricklefs, R. E. & Fjelds a, J." box="[1211,1269,1692,1717]" pageId="8" pageNumber="509" pagination="2328 - 2333" refId="ref17856" refString="Jonsson, K. A., Fabre, P. - H., Ricklefs, R. E. & Fjelds a, J. (2011) Major global radiation of corvoid birds originated in the proto- Papuan archipelago. Proceedings of the National Academy of Sciences, USA, 108, 2328 - 2333. http: // dx. doi. org / 10.1073 / pnas. 1018956108" type="journal article" year="2011">2011</bibRefCitation>
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;
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<bibRefCitation id="EFF84B525B6DA841FAE3F931FF75F96B" author="Norman, J. A. & Ericson, P. G. & Jonsson, K. A. & Fjelds a, J. & Christidis, L." pageId="8" pageNumber="509" pagination="488 - 497" refId="ref18462" refString="Norman, J. A., Ericson, P. G. P, Jonsson, K. A., Fjelds a, J. & Christidis, L. (2009 b) A multi-gene phylogeny reveals novel relationships for aberrant genera of Australo-Papuan core Corvoidea and polyphyly of the Pachycephalidae and Psophodidae (Aves: Passeriformes). Molecular Phylogenetics and Evolution, 52, 488 - 497. http: // dx. doi. org / 10.1016 / j. ympev. 2009.03.019" type="journal article" year="2009">
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Norman
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<emphasis id="B91DEAB15B6DA841FA86F930FA00F906" box="[1380,1429,1692,1717]" italics="true" pageId="8" pageNumber="509">et al</emphasis>
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. 2009b
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</bibRefCitation>
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;
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<bibRefCitation id="EFF84B525B6DA841FF0EF912FE7FF96B" author="Aggerbeck, M. & Fjelds a, J. & Christidis, L. & Fabre, P. - H. & Jonsson, K. A." box="[236,490,1727,1752]" pageId="8" pageNumber="509" pagination="272 - 285" refId="ref15184" refString="Aggerbeck, M., Fjelds a, J., Christidis, L., Fabre, P. - H. & Jonsson, K. A. (2014) Resolving deep lineage divergences in core corvoid passerine birds supports a proto-Papuan island origin. Molecular Phylogenetics and Evolution, 70, 272 - 285. http: // dx. doi. org / 10.1016 / j. ympev. 2013.09.027" type="journal article" year="2014">
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Aggerbeck
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<emphasis id="B91DEAB15B6DA841FE8DF96CFE34F96B" box="[367,417,1727,1752]" italics="true" pageId="8" pageNumber="509">et al</emphasis>
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. 2014
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</bibRefCitation>
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) place it firmly elsewhere, among a complex of corvoid songbirds that includes the Australasian butcherbirds and woodswallows (
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<taxonomicName id="4C694D205B6DA841FD2DF949FCD9F94E" authorityName="Vigors" authorityYear="1825" box="[719,844,1764,1789]" class="Aves" family="Artamidae" kingdom="Animalia" order="Passeriformes" pageId="8" pageNumber="509" phylum="Chordata" rank="family">Artamidae</taxonomicName>
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), African bush-shrikes, wattle-eyes and vangids (
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<taxonomicName id="4C694D205B6DA841FF7CF8AAFEDCF893" authorityName="Swainson" authorityYear="1824" box="[158,329,1799,1824]" class="Aves" family="Malaconotidae" kingdom="Animalia" order="Passeriformes" pageId="8" pageNumber="509" phylum="Chordata" rank="family">Malaconotidae</taxonomicName>
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,
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<taxonomicName id="4C694D205B6DA841FEBBF8AAFE61F893" box="[345,500,1799,1824]" class="Aves" family="Platysteiridae" kingdom="Animalia" order="Passeriformes" pageId="8" pageNumber="509" phylum="Chordata" rank="family">Platysteiridae</taxonomicName>
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,
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<taxonomicName id="4C694D205B6DA841FDE6F8AAFDE4F893" authorityName="Swainson" authorityYear="1831" box="[516,625,1799,1824]" class="Aves" family="Vangidae" kingdom="Animalia" order="Passeriformes" pageId="8" pageNumber="509" phylum="Chordata" rank="family">Vangidae</taxonomicName>
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), southeast Asian ioras (
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<taxonomicName id="4C694D205B6DA841FC70F8AAFBB3F893" authorityName="G.R.Gray" authorityYear="1869" box="[914,1062,1799,1824]" class="Aves" family="Aegithinidae" kingdom="Animalia" order="Passeriformes" pageId="8" pageNumber="509" phylum="Chordata" rank="family">Aegithinidae</taxonomicName>
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) and Old World cuckoo-shrikes (
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<taxonomicName id="4C694D205B6DA841FF7CF881FECFF8F6" authorityName="Vigors" authorityYear="1825" box="[158,346,1836,1861]" class="Aves" family="Campephagidae" kingdom="Animalia" order="Passeriformes" pageId="8" pageNumber="509" phylum="Chordata" rank="family">Campephagidae</taxonomicName>
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). Nearest relatives within that complex are still unclear. The ioras (
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<bibRefCitation id="EFF84B525B6DA841FB84F881FAD2F8F6" author="Jonsson, K. A. & Fabre, P. - H. & Ricklefs, R. E. & Fjelds a, J." box="[1126,1351,1836,1861]" pageId="8" pageNumber="509" pagination="2328 - 2333" refId="ref17856" refString="Jonsson, K. A., Fabre, P. - H., Ricklefs, R. E. & Fjelds a, J. (2011) Major global radiation of corvoid birds originated in the proto- Papuan archipelago. Proceedings of the National Academy of Sciences, USA, 108, 2328 - 2333. http: // dx. doi. org / 10.1073 / pnas. 1018956108" type="journal article" year="2011">
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Jønsson
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<emphasis id="B91DEAB15B6DA841FB28F880FB6BF8F6" box="[1226,1278,1836,1861]" italics="true" pageId="8" pageNumber="509">et al</emphasis>
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. 2011
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</bibRefCitation>
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) or the cracticid-artamid group (
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<bibRefCitation id="EFF84B525B6DA841FE53F8E2FD20F8DB" author="Aggerbeck, M. & Fjelds a, J. & Christidis, L. & Fabre, P. - H. & Jonsson, K. A." box="[433,693,1871,1896]" pageId="8" pageNumber="509" pagination="272 - 285" refId="ref15184" refString="Aggerbeck, M., Fjelds a, J., Christidis, L., Fabre, P. - H. & Jonsson, K. A. (2014) Resolving deep lineage divergences in core corvoid passerine birds supports a proto-Papuan island origin. Molecular Phylogenetics and Evolution, 70, 272 - 285. http: // dx. doi. org / 10.1016 / j. ympev. 2013.09.027" type="journal article" year="2014">
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Aggerbeck
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<emphasis id="B91DEAB15B6DA841FDD5F8FCFDFFF8DB" box="[567,618,1871,1896]" italics="true" pageId="8" pageNumber="509">et al</emphasis>
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. 2014
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</bibRefCitation>
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) have been indicated, but support values are weak, and depth of divergence considerable.
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</paragraph>
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|
<paragraph id="8BD636A35B6DA840FF25F835FCD6FE63" blockId="8.[151,1437,1548,2005]" lastBlockId="9.[151,1437,151,464]" lastPageId="9" lastPageNumber="510" pageId="8" pageNumber="509">
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|
As diagnosed, frugivorous
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|
<taxonomicName id="4C694D205B6DA841FDF1F835FD0CF802" authorityName="Stresemann & Paludan" authorityYear="1934" box="[531,665,1944,1969]" class="Aves" family="Pachycephalidae" genus="Rhagologus" kingdom="Animalia" order="Passeriformes" pageId="8" pageNumber="509" phylum="Chordata" rank="genus">
|
|
<emphasis id="B91DEAB15B6DA841FDF1F835FD0CF802" box="[531,665,1944,1969]" italics="true" pageId="8" pageNumber="509">Rhagologus</emphasis>
|
|
</taxonomicName>
|
|
has little in common with the insectivorous ioras, which are predominantly yellow-, green- and black-plumaged with silken-plumed flanks, display aerobatically and build perched cup-shaped nests bound smoothly with cobweb. Nor do its rather broad palate, open nares and unstructured temporal region of the skull resemble the narrowed palate, heavily ossified nasal cavity and compound zygomatic processes found in the Australasian butcherbirds and woodswallows (
|
|
<bibRefCitation id="EFF84B525B6CA840FB20FF72FE99FEAE" author="Schodde, R. & Mason, I. J." pageId="9" pageNumber="510" refId="ref19607" refString="Schodde, R. & Mason, I. J. (1999) The Directory of Australian Birds Passerines. CSIRO Publishing, Melbourne, 551 pp." type="book" year="1999">Schodde & Mason 1999: 533</bibRefCitation>
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|
) and, in part, vangas. The insectivorous pachycephalids have a broad palate and internally perforate nasal cavity similar in form to those of
|
|
<taxonomicName id="4C694D205B6CA840FDB3FE8AFD42FEF3" authorityName="Stresemann & Paludan" authorityYear="1934" box="[593,727,295,320]" class="Aves" family="Pachycephalidae" genus="Rhagologus" kingdom="Animalia" order="Passeriformes" pageId="9" pageNumber="510" phylum="Chordata" rank="genus">
|
|
<emphasis id="B91DEAB15B6CA840FDB3FE8AFD42FEF3" box="[593,727,295,320]" italics="true" pageId="9" pageNumber="510">Rhagologus</emphasis>
|
|
</taxonomicName>
|
|
, but the temporal region of the skull differs markedly: its fossa is much narrower, deeper and more clearly defined in pachycephalids, and both postorbital and simple zygomatic processes are well-developed and directed ventrally at an angle of
|
|
<emphasis id="B91DEAB15B6CA840FC77FEDCFC34FE3B" box="[917,929,369,392]" italics="true" pageId="9" pageNumber="510">c</emphasis>
|
|
. 45º. Combined morphological, behavioural and DNA sequence evidence reveal
|
|
<taxonomicName id="4C694D205B6CA840FDCFFE39FD26FE1E" authorityName="Stresemann & Paludan" authorityYear="1934" box="[557,691,404,429]" class="Aves" family="Pachycephalidae" genus="Rhagologus" kingdom="Animalia" order="Passeriformes" pageId="9" pageNumber="510" phylum="Chordata" rank="genus">
|
|
<emphasis id="B91DEAB15B6CA840FDCFFE39FD26FE1E" box="[557,691,404,429]" italics="true" pageId="9" pageNumber="510">Rhagologus</emphasis>
|
|
</taxonomicName>
|
|
as a deeply divergent corvoid lineage that cannot be placed in any other family. Accordingly, it is described at family rank here.
|
|
</paragraph>
|
|
</subSubSection>
|
|
</treatment>
|
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</document> |