431 lines
63 KiB
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431 lines
63 KiB
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<document id="E8A979855C8687881E3395422623D4F4" ID-CLB-Dataset="4210" ID-DOI="10.5852/ejt.2021.767.1495" ID-GBIF-Dataset="67bc6a73-4137-4de3-9dc4-ce87972f742d" ID-ISSN="2118-9773" ID-Zenodo-Dep="5514839" ID-ZooBank="9F019E02-95E7-4B6A-B5BC-01F8E26F3466" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1631927261920" checkinUser="felipe" docAuthor="Nahok, Benchawan, Tumpeesuwan, Sakboworn & Tumpeesuwan, Chanidaporn" docDate="2021" docId="03AC87E7C33FFFDC9CD8FACF9704B918" docLanguage="en" docName="ejt-767_142-166.pdf" docOrigin="European Journal of Taxonomy 767 (1)" docSource="http://dx.doi.org/10.5852/ejt.2021.767.1495" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Landouria diplogramma Nahok & Tumpeesuwan & Tumpeesuwan 2021, comb. nov." docType="treatment" docVersion="9" lastPageNumber="161" masterDocId="FF95FF9FC32EFFCF9D1FFF9B931BBE42" masterDocTitle="Five new species of the land snail genus Landouria Godwin-Austen, 1918 (Gastropoda, Camaenidae) from northeastern Thailand, with note on genitalia and radula morphology of Landouria diplogramma (Möllendorff, 1902)" masterLastPageNumber="166" masterPageNumber="142" pageNumber="159" updateTime="1699449512379" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="FFBD6F4630DD7599187E4817082A3149">Five new species of the land snail genus Landouria Godwin-Austen, 1918 (Gastropoda, Camaenidae) from northeastern Thailand, with note on genitalia and radula morphology of Landouria diplogramma (Möllendorff, 1902)</mods:title>
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<mods:namePart id="75DF51040174C4B4075649735A01EED0">Nahok, Benchawan</mods:namePart>
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<mods:affiliation id="6798FB214B26F3CFB861D09D71B86B5F">Department of Biology, Faculty of Science, Mahasarakham University, Kantharawichai District, Maha Sarakham, 44150 Thailand. & Educational Research Development and Demonstration Institute, Srinakharinwirot University, Ongkharak District, Nakhon Nayok, 26120 Thailand.</mods:affiliation>
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<mods:nameIdentifier id="0FF77877C0C0A36B14EA64FCD92E5903" type="email">nahok.b@gmail.com</mods:nameIdentifier>
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<mods:namePart id="BA1A71F8EE4B7A5B316311B60630049F">Tumpeesuwan, Sakboworn</mods:namePart>
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<mods:affiliation id="C6400EAC2D7B5FE725B98F9254D998E8">Department of Biology, Faculty of Science, Mahasarakham University, Kantharawichai District, Maha Sarakham, 44150 Thailand. & Palaeontological Research and Education Centre, Mahasarakham University, Kantharawichai District, Maha Sarakham, 44150 Thailand.</mods:affiliation>
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<mods:nameIdentifier id="5519F6F46B73030AF6D1510CF1B12444" type="email">stumpeesuwan@yahoo.com</mods:nameIdentifier>
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<mods:namePart id="549608BA7084EB6A753F5E97A2666C0D">Tumpeesuwan, Chanidaporn</mods:namePart>
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<mods:affiliation id="3D93ABEDC08DF1EE6A80CEF7AD75EB35">Department of Biology, Faculty of Science, Mahasarakham University, Kantharawichai District, Maha Sarakham, 44150 Thailand.</mods:affiliation>
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<mods:nameIdentifier id="FF44E5E46C241BD2500489BFA0728F3F" type="email">ctumpeesuwan@yahoo.com</mods:nameIdentifier>
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<mods:title id="3FCBE7BA7852B9D990221BB02E66E0A1">European Journal of Taxonomy</mods:title>
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<treatment id="03AC87E7C33FFFDC9CD8FACF9704B918" ID-DOI="http://doi.org/10.5281/zenodo.5528098" ID-GBIF-Taxon="190223605" ID-Zenodo-Dep="5528098" LSID="urn:lsid:plazi:treatment:03AC87E7C33FFFDC9CD8FACF9704B918" httpUri="http://treatment.plazi.org/id/03AC87E7C33FFFDC9CD8FACF9704B918" lastPageId="19" lastPageNumber="161" pageId="17" pageNumber="159">
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<heading id="D0F2819DC33FFFDE9CD8FACF9776BB2D" box="[455,1133,1364,1391]" centered="true" fontSize="11" level="2" pageId="17" pageNumber="159" reason="2">
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<taxonomicName id="4C054D72C33FFFDE9CD8FACF90F8BB2D" ID-CoL="8QG7M" authority="(Mollendorff, 1902)" authorityName="Nahok & Tumpeesuwan & Tumpeesuwan" authorityYear="2021" baseAuthorityName="Mollendorff" baseAuthorityYear="1902" box="[455,995,1364,1391]" class="Gastropoda" family="Camaenidae" genus="Landouria" kingdom="Animalia" order="Stylommatophora" pageId="17" pageNumber="159" phylum="Mollusca" rank="species" species="diplogramma" status="comb. nov.">
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<emphasis id="B971EAE3C33FFFDE9CD8FACF91EBBB2C" bold="true" box="[455,752,1364,1390]" italics="true" pageId="17" pageNumber="159">Landouria diplogramma</emphasis>
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(
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<bibRefCitation id="EF944B00C33FFFDE9E1FFACE90C0BB2D" author="Mollendorff O. F." box="[768,987,1365,1391]" pageId="17" pageNumber="159" pagination="153 - 160" refId="ref10489" refString="Mollendorff O. F. 1902. Neue Arten und Unterarten von Fruhstorfer in Siam gesammelt. Nachrichtsblatt der Deutschen Malakozoologischen Gesellschaft 34 (9 - 10): 153 - 160." type="journal article" year="1902">Möllendorff, 1902</bibRefCitation>
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<subSubSection id="C31F657AC33FFFDE9F4BFAE390C4BBD0" box="[596,991,1400,1427]" pageId="17" pageNumber="159" type="description">
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<paragraph id="8BBA36F1C33FFFDE9F4BFAE390C4BBD0" blockId="17.[455,1133,1364,1427]" box="[596,991,1400,1427]" pageId="17" pageNumber="159">
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<figureCitation id="133E2A74C33FFFDE9F4BFAE391B6BBD0" box="[596,685,1400,1426]" captionStart="Fig" captionStartId="9.[189,232,1849,1875]" captionTargetBox="[220,1368,265,1811]" captionTargetId="figure-16@9.[220,1368,265,1811]" captionTargetPageId="9" captionText="Fig. 3. External shell morphology of Landouria spp. from northeastern Thailand. A. Landouria circinata sp. nov., holotype (NHMSU-00023). B. L. tuberculata sp. nov., holotype (NHMSU-00025). C. L. trochomorphoides sp. nov., holotype (NHMSU-00027). D. L. chloritoides sp. nov., holotype (NHMSU-00029). E. L. elegans sp. nov., holotype (NHMSU-00032). F. L. diplogramma (Möllendorff, 1902) comb. nov. (NHMSU-00034). Scale bar = 10 mm." figureDoi="http://doi.org/10.5281/zenodo.5514845" httpUri="https://zenodo.org/record/5514845/files/figure.png" pageId="17" pageNumber="159">Figs 3F</figureCitation>
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,
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<figureCitation id="133E2A74C33FFFDE9FA6FAE391C2BBD0" box="[697,729,1400,1426]" captionStart="Fig" captionStartId="12.[189,232,1921,1947]" captionTargetBox="[193,1395,278,1870]" captionTargetId="figure-12@12.[193,1395,265,1883]" captionTargetPageId="12" captionText="Fig. 4. Living adults of Landouria spp. from northeastern Thailand. A. Landouria circinata sp. nov. B. L. tuberculata sp. nov. C. L. trochomorphoides sp. nov. D. L. chloritoides sp. nov. E. L. elegans sp. nov. F. L. diplogramma (Möllendorff, 1902) comb. nov. Scale bars = 10 mm." figureDoi="http://doi.org/10.5281/zenodo.5514847" httpUri="https://zenodo.org/record/5514847/files/figure.png" pageId="17" pageNumber="159">4F</figureCitation>
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,
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<figureCitation id="133E2A74C33FFFDE9FFBFAE3901FBBD1" box="[740,772,1400,1427]" captionStart="Fig" captionStartId="15.[189,232,1816,1842]" captionTargetBox="[283,1352,278,1767]" captionTargetId="figure-15@15.[232,1356,265,1778]" captionTargetPageId="15" captionText="Fig. 5. Illustrations of the genital system of Landouria spp. from northeastern Thailand. A. Landouria circinata sp. nov., paratype (NHMSU-00024). B. L. tuberculata sp. nov., paratype (NHMSU-00026). C. L. trochomorphoides sp. nov., paratype (NHMSU-00028). D. L. chloritoides sp. nov., paratype (NHMSU-00030). E. L. elegans sp. nov., paratype (NHMSU-00033). F. L. diplogramma (Möllendorff, 1902) comb. nov. (NHMSU-00034). Internal sculpture of a = flagellum; b = penis; c = vagina. Scale bars = 10 mm." figureDoi="http://doi.org/10.5281/zenodo.5514849" httpUri="https://zenodo.org/record/5514849/files/figure.png" pageId="17" pageNumber="159">5F</figureCitation>
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,
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<figureCitation id="133E2A74C33FFFDE9E0FFAE3904EBBD0" box="[784,853,1400,1426]" captionStart="Fig" captionStartId="18.[189,232,1783,1809]" captionTargetBox="[244,1343,265,1744]" captionTargetId="figure-12@18.[244,1343,265,1744]" captionTargetPageId="18" captionText="Fig. 6.Radula morphology ofLandouria spp. from northeasternThailand.A–C, G–R. Right side of central teeth. D–F. Left side of central teeth. A–C. Landouria circinata sp. nov., paratype (NHMSU-00024). D–F. L. tuberculata sp. nov., paratype (NHMSU-00026). G–I. L. trochomorphoides sp. nov., paratype (NHMSU-00028). J–L. L. chloritoides sp. nov., paratype (NHMSU-00030). M–O. L. elegans sp. nov., paratype (NHMSU-00033). P–R. L. diplogramma (Möllendorff, 1902) comb. nov. (NHMSU-00034). Numbers indicate order of lateral and marginal teeth. Abbreviation: c = central tooth. Scale bars = 25 µm." figureDoi="http://doi.org/10.5281/zenodo.5514851" httpUri="https://zenodo.org/record/5514851/files/figure.png" pageId="17" pageNumber="159">6P–R</figureCitation>
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;
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<tableCitation id="C687034AC33FFFDE9E7FFAE390C4BBD0" box="[864,991,1400,1426]" captionStart-0="Table 1" captionStart-1="Table 2" captionStart-2="Table 3" captionStart-3="Table 4" captionStartId-0="2.[185,255,857,885]" captionStartId-1="4.[186,256,600,628]" captionStartId-2="6.[185,255,695,723]" captionStartId-3="10.[185,255,544,572]" captionTargetBox-0="[195,1921,987,1444]" captionTargetBox-1="[197,1907,827,1685]" captionTargetBox-2="[212,1909,868,1619]" captionTargetBox-3="[196,1908,758,1739]" captionTargetPageId-0="2" captionTargetPageId-1="4" captionTargetPageId-2="6" captionTargetPageId-3="10" captionText-0="Table 1. Shell measurements (mm) of six new species of the genus Landouria Godwin-Austen, 1918 from northeastern Thailand." captionText-1="Table 2. Locality data and identification of specimens of Landouria Godwin-Austen, 1918 used for mtDNA sequencing, including collection catalogue numbers and GenBank accession numbers. Abbreviations: NHMSU = Natural History Museum, Mahasarakham University; HNHM = Hungarian Natural History Museum; AM = Australian Museum. References: 1 = Tumpeesuwan & Tumpeesuwan (2019); 2 = Hirano et al. (2014); 3 = Köhler et al. (2018)." captionText-2="Table 3. Pairwise p-distance values of Landouria Godwin-Austen, 1918 based on mitochondrial COI sequences." captionText-3="Table 4. Comparison of shells, radulae, and genital systems of Landouria spp. from northeastern Thailand. References: 1 = Tumpeesuwan & Tumpeesuwan (2019); 2 = Nahok (2020); 3 = this study. *In Tumpeesuwan & Tumpeesuwan (2019) indicated the number of teeth per row in Landouria strobiloides varies from 3 to 53. This lowest row has only 3 teeth. It is the uppermost row, probably lacerated and peel off. Unfortunately, we could not re-checked the complete specimens in this time being, so, this point need further investigation and proof." httpUri-0="http://table.plazi.org/id/DF7A6679C32CFFCD9DA6FCC295B0BD37" httpUri-1="http://table.plazi.org/id/DF7A6679C32AFFCB9DA5FDC39249BC9F" httpUri-2="http://table.plazi.org/id/DF7A6679C328FFC99DA6FD2C96FFBC91" httpUri-3="http://table.plazi.org/id/DF7A6679C324FFC59DA6FDBB969CBCE7" pageId="17" pageNumber="159" tableUuid-0="DF7A6679C32CFFCD9DA6FCC295B0BD37" tableUuid-1="DF7A6679C32AFFCB9DA5FDC39249BC9F" tableUuid-2="DF7A6679C328FFC99DA6FD2C96FFBC91" tableUuid-3="DF7A6679C324FFC59DA6FDBB969CBCE7">Tables 1–4</tableCitation>
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</paragraph>
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</subSubSection>
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<subSubSection id="C31F657AC33FFFDE9DA2FA22902AB819" pageId="17" pageNumber="159" type="reference_group">
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<paragraph id="8BBA36F1C33FFFDE9DA2FA229057BB96" blockId="17.[189,844,1465,1492]" box="[189,844,1465,1492]" pageId="17" pageNumber="159">
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<treatmentCitationGroup id="AB1511DFC33FFFDE9DA2FA229057BB96" box="[189,844,1465,1492]" pageId="17" pageNumber="159">
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<treatmentCitation id="0AA410E0C33FFFDE9DA2FA229053BB96" author="Mollendorff O. F." box="[189,840,1465,1492]" page="159" pageId="17" pageNumber="159" year="1902">
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<taxonomicName id="4C054D72C33FFFDE9DA2FA229053BB96" ID-CoL="7SRFC" authority="Mollendorff, 1902: 158 - 159" authorityName="Mollendorff" authorityPageNumber="158 - 159" authorityYear="1902" box="[189,840,1465,1492]" class="Gastropoda" family="Camaenidae" genus="Plectotropis" kingdom="Animalia" order="Stylommatophora" pageId="17" pageNumber="159" phylum="Mollusca" rank="species" species="diplogramme">
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<emphasis id="B971EAE3C33FFFDE9DA2FA2292F4BB91" box="[189,495,1465,1491]" italics="true" pageId="17" pageNumber="159">Plectotropis diplogramme</emphasis>
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<bibRefCitation id="EF944B00C33FFFDE9CE9FA229053BB96" author="Mollendorff O. F." box="[502,840,1465,1492]" pageId="17" pageNumber="159" pagination="153 - 160" refId="ref10489" refString="Mollendorff O. F. 1902. Neue Arten und Unterarten von Fruhstorfer in Siam gesammelt. Nachrichtsblatt der Deutschen Malakozoologischen Gesellschaft 34 (9 - 10): 153 - 160." type="journal article" year="1902">Möllendorff, 1902: 158–159</bibRefCitation>
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.
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<taxonomicName id="4C054D72C33FFFDE9DA2FA6092CCB857" box="[189,471,1531,1557]" class="Gastropoda" family="Camaenidae" genus="Ganesella" kingdom="Animalia" order="Stylommatophora" pageId="17" pageNumber="159" phylum="Mollusca" rank="species" species="diplogramme">
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<emphasis id="B971EAE3C33FFFDE9DA2FA6092CCB857" box="[189,471,1531,1557]" italics="true" pageId="17" pageNumber="159">Ganesella diplogramme</emphasis>
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</taxonomicName>
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–
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<treatmentCitation id="0AA410E0C33FFFDE9CEAFA6091B4B857" author="Panha S." box="[501,687,1531,1557]" page="36" pageId="17" pageNumber="159" year="1996">
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<bibRefCitation id="EF944B00C33FFFDE9CEAFA6091B4B857" author="Panha S." box="[501,687,1531,1557]" pageId="17" pageNumber="159" pagination="31 - 40" refId="ref10845" refString="Panha S. 1996 A checklist and classification of the terrestrial pulmonate snails of Thailand. Walkerana 8 (19): 31 - 40." type="journal article" year="1996">Panha 1996: 36</bibRefCitation>
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.
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|
||
<taxonomicName id="4C054D72C33FFFDE9DA2F9849178B87A" authorityName="Nahok & Tumpeesuwan & Tumpeesuwan" authorityYear="2021" baseAuthorityName="Mollendorff" baseAuthorityYear="1902" box="[189,611,1566,1593]" class="Gastropoda" family="Camaenidae" genus="Aegista" kingdom="Animalia" order="Stylommatophora" pageId="17" pageNumber="159" phylum="Mollusca" rank="species" species="diplogramma" subGenus="Plectotropis">
|
||
<emphasis id="B971EAE3C33FFFDE9DA2F984920EB87B" box="[189,277,1567,1593]" italics="true" pageId="17" pageNumber="159">Aegista</emphasis>
|
||
(
|
||
<emphasis id="B971EAE3C33FFFDE9C39F98592AFB87A" box="[294,436,1566,1592]" italics="true" pageId="17" pageNumber="159">Plectotropis</emphasis>
|
||
)
|
||
<emphasis id="B971EAE3C33FFFDE9CDAF9859178B87A" box="[453,611,1566,1592]" italics="true" pageId="17" pageNumber="159">diplogramma</emphasis>
|
||
</taxonomicName>
|
||
–
|
||
<treatmentCitation id="0AA410E0C33FFFDE9F9EF98490EBB87A" author="Hemmen J. & Hemmen C." box="[641,1008,1566,1593]" page="47" pageId="17" pageNumber="159" year="2001">
|
||
<bibRefCitation id="EF944B00C33FFFDE9F9EF98490EBB87A" author="Hemmen J. & Hemmen C." box="[641,1008,1566,1593]" pageId="17" pageNumber="159" pagination="35 - 70" refId="ref10248" refString="Hemmen J. & Hemmen C. 2001 Aktualisierte Liste der terrestrischen Gastropoden Thailands. Schriften zur Malakozoologie 18: 35 - 70." type="journal article" year="2001">Hemmen & Hemmen 2001: 47</bibRefCitation>
|
||
</treatmentCitation>
|
||
.
|
||
</treatmentCitationGroup>
|
||
</paragraph>
|
||
<paragraph id="8BBA36F1C33FFFDE9DA2F9DA902AB819" blockId="17.[189,1012,1531,1628]" box="[189,817,1601,1628]" pageId="17" pageNumber="159">
|
||
<treatmentCitationGroup id="AB1511DFC33FFFDE9DA2F9DA902AB819" box="[189,817,1601,1628]" pageId="17" pageNumber="159">
|
||
<taxonomicName id="4C054D72C33FFFDE9DA2F9DA9244B81E" box="[189,351,1601,1628]" class="Gastropoda" family="Camaenidae" genus="Thaitropis" kingdom="Animalia" order="Stylommatophora" pageId="17" pageNumber="159" phylum="Mollusca" rank="species" species="undetermined">
|
||
<emphasis id="B971EAE3C33FFFDE9DA2F9DA922DB819" box="[189,310,1601,1627]" italics="true" pageId="17" pageNumber="159">Thaitropis</emphasis>
|
||
sp.
|
||
</taxonomicName>
|
||
–
|
||
<treatmentCitation id="0AA410E0C33FFFDE9C63F9DA91B2B819" author="Sutcharit C. & Tongkerd P. & Panha S." box="[380,681,1601,1627]" page="249" pageId="17" pageNumber="159" year="2017">
|
||
<bibRefCitation id="EF944B00C33FFFDE9C63F9DA91B2B819" author="Sutcharit C. & Tongkerd P. & Panha S." box="[380,681,1601,1627]" pageId="17" pageNumber="159" refId="ref11218" refString="Sutcharit C., Tongkerd P. & Panha S. 2017. Land Snails the Invaluable Bio-Resources for the Kingdom of Thailand. The Thailand Research Fund, Bangkok. [In Thai.]" type="book" year="2017">
|
||
Sutcharit
|
||
<emphasis id="B971EAE3C33FFFDE9CF2F9D99133B819" box="[493,552,1601,1627]" italics="true" pageId="17" pageNumber="159">et al.</emphasis>
|
||
2017: 249
|
||
</bibRefCitation>
|
||
</treatmentCitation>
|
||
, fig. 6–32e.
|
||
</treatmentCitationGroup>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C31F657AC33FFFDE9DA2F919905EB950" pageId="17" pageNumber="159" type="diagnosis">
|
||
<paragraph id="8BBA36F1C33FFFDE9DA2F919922DB8DE" blockId="17.[189,310,1666,1692]" box="[189,310,1666,1692]" pageId="17" pageNumber="159">
|
||
<heading id="D0F2819DC33FFFDE9DA2F919922DB8DE" bold="true" box="[189,310,1666,1692]" fontSize="11" level="3" pageId="17" pageNumber="159" reason="3">
|
||
<emphasis id="B971EAE3C33FFFDE9DA2F919922DB8DE" bold="true" box="[189,310,1666,1692]" pageId="17" pageNumber="159">Diagnosis</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="8BBA36F1C33FFFDE9DA2F92A905EB950" blockId="17.[189,1399,1713,1810]" pageId="17" pageNumber="159">Shell small, low-conical, weakly keeled, with a slightly elevated spire, whitish corneous with a pale brown band above periphery. Flagellum like a short protrusion, with rounded tip. Penis short swollen. Central and the first few lateral radular teeth triangular.</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C31F657AC33FFFDE9DA2F8A297D0B98A" pageId="17" pageNumber="159" type="materials_examined">
|
||
<paragraph id="8BBA36F1C33FFFDE9DA2F8A292B1B911" blockId="17.[189,426,1849,1875]" box="[189,426,1849,1875]" pageId="17" pageNumber="159">
|
||
<heading id="D0F2819DC33FFFDE9DA2F8A292B1B911" bold="true" box="[189,426,1849,1875]" fontSize="11" level="3" pageId="17" pageNumber="159" reason="3">
|
||
<emphasis id="B971EAE3C33FFFDE9DA2F8A292B1B911" bold="true" box="[189,426,1849,1875]" pageId="17" pageNumber="159">Material examined</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="8BBA36F1C33FFFDE9DA2F8F397D0B98A" blockId="17.[189,1399,1896,1993]" pageId="17" pageNumber="159">
|
||
<materialsCitation id="3B6D3CACC33FFFDE9DA2F8F397D0B98A" ID-GBIF-Occurrence="3408781301" accessionNumber="MN449411 and MZ435752" collectingDate="2017-06-23" collectionCode="MN, MZ" collectorName="B. Nahok & U. Chanlabut" country="Thailand" elevation="451" latitude="14.615159" location="Wat Thep Phithak Punnaram" longLatPrecision="1" longitude="101.26535" municipality="Pak Chong District" pageId="17" pageNumber="159" specimenCode="NHMSU-00034" specimenCount="43" stateProvince="Nakhon Ratchasima Province">
|
||
<collectingCountry id="F3127661C33FFFDE9DA2F8F3924EB9C0" box="[189,341,1896,1922]" name="Thailand" pageId="17" pageNumber="159">THAILAND</collectingCountry>
|
||
•
|
||
<specimenCount id="9D03FD78C33FFFDE9C66F8F392F2B9C0" box="[377,489,1896,1922]" pageId="17" pageNumber="159">29 shells</specimenCount>
|
||
,
|
||
<specimenCount id="9D03FD78C33FFFDE9CE4F8F391A2B9C0" box="[507,697,1896,1922]" pageId="17" pageNumber="159">14 living specs</specimenCount>
|
||
;
|
||
<collectingRegion id="49C1F813C33FFFDE9FD4F8F3972DB9C0" box="[715,1078,1896,1922]" country="Thailand" name="Nakhon Ratchasima" pageId="17" pageNumber="159">Nakhon Ratchasima Province</collectingRegion>
|
||
,
|
||
<collectingMunicipality id="6BDEAC8BC33FFFDE9957F8F3962CB9C0" box="[1096,1335,1896,1922]" pageId="17" pageNumber="159">Pak Chong District</collectingMunicipality>
|
||
,
|
||
<location id="8EDA602AC33FFFDE9857F8F392F9B9E7" LSID="urn:lsid:plazi:treatment:03AC87E7C33FFFDC9CD8FACF9704B918:8EDA602AC33FFFDE9857F8F392F9B9E7" country="Thailand" latitude="14.615159" longLatPrecision="1" longitude="101.26535" municipality="Pak Chong District" name="Wat Thep Phithak Punnaram" pageId="17" pageNumber="159" stateProvince="Nakhon Ratchasima Province">Wat Thep Phithak Punnaram</location>
|
||
;
|
||
<geoCoordinate id="EE315036C33FFFDE9CEFF81291B6B9E7" box="[496,685,1929,1957]" degrees="14" direction="north" minutes="36" orientation="latitude" pageId="17" pageNumber="159" precision="1" seconds="54.57" value="14.615159">14°36′54.57″ N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE315036C33FFFDE9FA4F812909FB9E7" box="[699,900,1929,1957]" degrees="101" direction="east" minutes="15" orientation="longitude" pageId="17" pageNumber="159" precision="1" seconds="55.25" value="101.26535">101°15′55.25″ E</geoCoordinate>
|
||
; alt.
|
||
<quantity id="4CFD9B14C33FFFDE9EDDF8109709B9E7" box="[962,1042,1931,1958]" metricMagnitude="2" metricUnit="m" metricValue="4.51" pageId="17" pageNumber="159" unit="m" value="451.0">
|
||
<elevation id="0028D1C2C33FFFDE9EDDF8109709B9E7" box="[962,1042,1931,1958]" metricMagnitude="2" metricUnit="m" metricValue="4.51" pageId="17" pageNumber="159" unit="m" value="451.0">451 m</elevation>
|
||
</quantity>
|
||
;
|
||
<date id="FFBB1031C33FFFDE993EF81097DBB9E7" box="[1057,1216,1931,1957]" pageId="17" pageNumber="159" value="2017-06-23">
|
||
<collectingDate id="EFFFE9D9C33FFFDE993EF81097DBB9E7" box="[1057,1216,1931,1957]" pageId="17" pageNumber="159" value="2017-06-23">23 Jun. 2017</collectingDate>
|
||
</date>
|
||
;
|
||
<collectorName id="26F05327C33FFFDE99D0F8109659B9E7" box="[1231,1346,1931,1957]" pageId="17" pageNumber="159">B. Nahok</collectorName>
|
||
and
|
||
<collectorName id="26F05327C33FFFDE9DA2F8349241B98A" box="[189,346,1966,1993]" pageId="17" pageNumber="159">U. Chanlabut</collectorName>
|
||
leg.; GenBank No.
|
||
<accessionNumber id="9456AB12C33FFFDE9F5BF83591D1B98A" box="[580,714,1966,1992]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/MN449411" pageId="17" pageNumber="159">MN449411</accessionNumber>
|
||
and
|
||
<accessionNumber id="9456AB12C33FFFDE9E1CF8359090B98B" box="[771,907,1966,1993]" httpUri="https://www.ncbi.nlm.nih.gov/nuccore/MZ435752" pageId="17" pageNumber="159" type="GenBank">MZ435752</accessionNumber>
|
||
;
|
||
<specimenCode id="DBA39E8AC33FFFDE9E88F8359743B98A" box="[919,1112,1966,1992]" pageId="17" pageNumber="159">NHMSU-00034</specimenCode>
|
||
. (
|
||
<figureCitation id="133E2A74C33FFFDE9974F83597DAB98A" box="[1131,1217,1966,1992]" captionStart="Fig" captionStartId="9.[189,232,1849,1875]" captionTargetBox="[220,1368,265,1811]" captionTargetId="figure-16@9.[220,1368,265,1811]" captionTargetPageId="9" captionText="Fig. 3. External shell morphology of Landouria spp. from northeastern Thailand. A. Landouria circinata sp. nov., holotype (NHMSU-00023). B. L. tuberculata sp. nov., holotype (NHMSU-00025). C. L. trochomorphoides sp. nov., holotype (NHMSU-00027). D. L. chloritoides sp. nov., holotype (NHMSU-00029). E. L. elegans sp. nov., holotype (NHMSU-00032). F. L. diplogramma (Möllendorff, 1902) comb. nov. (NHMSU-00034). Scale bar = 10 mm." figureDoi="http://doi.org/10.5281/zenodo.5514845" httpUri="https://zenodo.org/record/5514845/files/figure.png" pageId="17" pageNumber="159">Fig. 3F</figureCitation>
|
||
)
|
||
</materialsCitation>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<caption id="DF7A6679C33CFFDD9DA2F96C920AB9A7" ID-DOI="http://doi.org/10.5281/zenodo.5514851" ID-Zenodo-Dep="5514851" httpUri="https://zenodo.org/record/5514851/files/figure.png" pageId="18" pageNumber="160" startId="18.[189,232,1783,1809]" targetBox="[244,1343,265,1744]" targetPageId="18">
|
||
<paragraph id="8BBA36F1C33CFFDD9DA2F96C920AB9A7" blockId="18.[185,1399,1783,2021]" pageId="18" pageNumber="160">
|
||
<emphasis id="B971EAE3C33CFFDD9DA2F96C9213B953" bold="true" box="[189,264,1783,1809]" pageId="18" pageNumber="160">Fig. 6.</emphasis>
|
||
Radula morphology of
|
||
<taxonomicName id="4C054D72C33CFFDD9F11F96C91DAB953" box="[526,705,1783,1809]" class="Gastropoda" family="Camaenidae" genus="Landouria" kingdom="Animalia" order="Stylommatophora" pageId="18" pageNumber="160" phylum="Mollusca" rank="species" species="undetermined">
|
||
<emphasis id="B971EAE3C33CFFDD9F11F96C9191B953" box="[526,650,1783,1809]" italics="true" pageId="18" pageNumber="160">Landouria</emphasis>
|
||
spp.
|
||
</taxonomicName>
|
||
from northeastern Thailand.
|
||
<emphasis id="B971EAE3C33CFFDD991AF96C9725B953" bold="true" box="[1029,1086,1783,1809]" pageId="18" pageNumber="160">A–C</emphasis>
|
||
,
|
||
<emphasis id="B971EAE3C33CFFDD9952F96C979CB953" bold="true" box="[1101,1159,1783,1809]" pageId="18" pageNumber="160">G–R</emphasis>
|
||
. Right side of central teeth.
|
||
<emphasis id="B971EAE3C33CFFDD9C19F8819227B976" bold="true" box="[262,316,1818,1844]" pageId="18" pageNumber="160">D–F</emphasis>
|
||
. Left side of central teeth.
|
||
<emphasis id="B971EAE3C33CFFDD9F9AF88191A7B976" bold="true" box="[645,700,1818,1844]" pageId="18" pageNumber="160">A–C</emphasis>
|
||
.
|
||
<taxonomicName id="4C054D72C33CFFDD9FD2F88190A0B976" authority="Nahok & Tumpeesuwan & Tumpeesuwan, 2021" authorityName="Nahok & Tumpeesuwan & Tumpeesuwan" authorityYear="2021" box="[717,955,1818,1844]" class="Gastropoda" family="Camaenidae" genus="Landouria" kingdom="Animalia" order="Stylommatophora" pageId="18" pageNumber="160" phylum="Mollusca" rank="species" species="circinata" status="sp. nov.">
|
||
<emphasis id="B971EAE3C33CFFDD9FD2F88190A0B976" box="[717,955,1818,1844]" italics="true" pageId="18" pageNumber="160">Landouria circinata</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A2425798C33CFFDD9EDBF8809738B977" box="[964,1059,1819,1845]" pageId="18" pageNumber="160" rank="species">sp. nov.</taxonomicNameLabel>
|
||
, paratype (NHMSU-00024).
|
||
<emphasis id="B971EAE3C33CFFDD9DA0F8A693E8B915" bold="true" box="[191,243,1853,1879]" pageId="18" pageNumber="160">D–F</emphasis>
|
||
.
|
||
<taxonomicName id="4C054D72C33CFFDD9C1DF8A592B3B915" authority="Nahok & Tumpeesuwan & Tumpeesuwan, 2021" authorityName="Nahok & Tumpeesuwan & Tumpeesuwan" authorityYear="2021" box="[258,424,1853,1879]" class="Gastropoda" family="Camaenidae" genus="Landouria" kingdom="Animalia" order="Stylommatophora" pageId="18" pageNumber="160" phylum="Mollusca" rank="species" species="tuberculata" status="sp. nov.">
|
||
<emphasis id="B971EAE3C33CFFDD9C1DF8A592B3B915" box="[258,424,1853,1879]" italics="true" pageId="18" pageNumber="160">L. tuberculata</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A2425798C33CFFDD9CAFF8A59115B91A" box="[432,526,1854,1880]" pageId="18" pageNumber="160" rank="species">sp. nov.</taxonomicNameLabel>
|
||
, paratype (NHMSU-00026).
|
||
<emphasis id="B971EAE3C33CFFDD9E76F8A69081B915" bold="true" box="[873,922,1853,1879]" pageId="18" pageNumber="160">G–I</emphasis>
|
||
.
|
||
<taxonomicName id="4C054D72C33CFFDD9EB6F8A59784B915" authority="Nahok & Tumpeesuwan & Tumpeesuwan, 2021" authorityName="Nahok & Tumpeesuwan & Tumpeesuwan" authorityYear="2021" box="[937,1183,1853,1879]" class="Gastropoda" family="Camaenidae" genus="Landouria" kingdom="Animalia" order="Stylommatophora" pageId="18" pageNumber="160" phylum="Mollusca" rank="species" species="trochomorphoides" status="sp. nov.">
|
||
<emphasis id="B971EAE3C33CFFDD9EB6F8A59784B915" box="[937,1183,1853,1879]" italics="true" pageId="18" pageNumber="160">L. trochomorphoides</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A2425798C33CFFDD99B8F8A5961EB91A" box="[1191,1285,1854,1880]" pageId="18" pageNumber="160" rank="species">sp. nov.</taxonomicNameLabel>
|
||
, paratype (NHMSU-00028).
|
||
<emphasis id="B971EAE3C33CFFDD9C83F8FA92D6B939" bold="true" box="[412,461,1889,1915]" pageId="18" pageNumber="160">J–L</emphasis>
|
||
.
|
||
<taxonomicName id="4C054D72C33CFFDD9CC3F8FA919CB938" authority="Nahok & Tumpeesuwan & Tumpeesuwan, 2021" authorityName="Nahok & Tumpeesuwan & Tumpeesuwan" authorityYear="2021" box="[476,647,1888,1914]" class="Gastropoda" family="Camaenidae" genus="Landouria" kingdom="Animalia" order="Stylommatophora" pageId="18" pageNumber="160" phylum="Mollusca" rank="species" species="chloritoides" status="sp. nov.">
|
||
<emphasis id="B971EAE3C33CFFDD9CC3F8FA919CB938" box="[476,647,1888,1914]" italics="true" pageId="18" pageNumber="160">L. chloritoides</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A2425798C33CFFDD9F90F8FA91F7B939" box="[655,748,1889,1915]" pageId="18" pageNumber="160" rank="species">sp. nov.</taxonomicNameLabel>
|
||
, paratype (NHMSU-00030).
|
||
<emphasis id="B971EAE3C33CFFDD995AF8FB979DB938" bold="true" box="[1093,1158,1888,1914]" pageId="18" pageNumber="160">M–O</emphasis>
|
||
.
|
||
<taxonomicName id="4C054D72C33CFFDD998BF8FA9615B938" authority="Nahok & Tumpeesuwan & Tumpeesuwan, 2021" authorityName="Nahok & Tumpeesuwan & Tumpeesuwan" authorityYear="2021" box="[1172,1294,1888,1914]" class="Gastropoda" family="Camaenidae" genus="Landouria" kingdom="Animalia" order="Stylommatophora" pageId="18" pageNumber="160" phylum="Mollusca" rank="species" species="elegans" status="sp. nov.">
|
||
<emphasis id="B971EAE3C33CFFDD998BF8FA9615B938" box="[1172,1294,1888,1914]" italics="true" pageId="18" pageNumber="160">L. elegans</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A2425798C33CFFDD980AF8FA9669B939" box="[1301,1394,1889,1915]" pageId="18" pageNumber="160" rank="species">sp. nov.</taxonomicNameLabel>
|
||
, paratype (NHMSU-00033).
|
||
<emphasis id="B971EAE3C33CFFDD9F11F81F9159B9DC" bold="true" box="[526,578,1924,1950]" pageId="18" pageNumber="160">P–R</emphasis>
|
||
.
|
||
<taxonomicName id="4C054D72C33CFFDD9F4CF81F9711B9DC" authority="(Mollendorff, 1902)" authorityName="Nahok & Tumpeesuwan & Tumpeesuwan" authorityYear="2021" baseAuthorityName="Mollendorff" baseAuthorityYear="1902" box="[595,1034,1924,1950]" class="Gastropoda" family="Camaenidae" genus="Aegista" kingdom="Animalia" order="Stylommatophora" pageId="18" pageNumber="160" phylum="Mollusca" rank="species" species="diplogramma" status="comb. nov.">
|
||
<emphasis id="B971EAE3C33CFFDD9F4CF81F9009B9DC" box="[595,786,1924,1950]" italics="true" pageId="18" pageNumber="160">L. diplogramma</emphasis>
|
||
(
|
||
<bibRefCitation id="EF944B00C33CFFDD9E3AF81F9719B9DC" author="Mollendorff O. F." box="[805,1026,1924,1950]" pageId="18" pageNumber="160" pagination="153 - 160" refId="ref10489" refString="Mollendorff O. F. 1902. Neue Arten und Unterarten von Fruhstorfer in Siam gesammelt. Nachrichtsblatt der Deutschen Malakozoologischen Gesellschaft 34 (9 - 10): 153 - 160." type="journal article" year="1902">Möllendorff, 1902</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A2425798C33CFFDD990BF81F9783B9DC" box="[1044,1176,1924,1950]" pageId="18" pageNumber="160" rank="species">comb. nov.</taxonomicNameLabel>
|
||
(NHMSU-00034). Numbers indicate order of lateral and marginal teeth. Abbreviation: c = central tooth. Scale bars = 25 µm.
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection id="C31F657AC33DFFDC9DA2FE9391F7BDF8" pageId="19" pageNumber="161" type="description">
|
||
<paragraph id="8BBA36F1C33DFFDC9DA2FE939255BF60" blockId="19.[189,334,264,290]" box="[189,334,264,290]" pageId="19" pageNumber="161">
|
||
<heading id="D0F2819DC33DFFDC9DA2FE939255BF60" bold="true" box="[189,334,264,290]" fontSize="11" level="3" pageId="19" pageNumber="161" reason="3">
|
||
<emphasis id="B971EAE3C33DFFDC9DA2FE939255BF60" bold="true" box="[189,334,264,290]" pageId="19" pageNumber="161">Description</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="8BBA36F1C33DFFDC9DA2FEA39248BFFC" blockId="19.[189,1400,310,446]" pageId="19" pageNumber="161">
|
||
SHELL (
|
||
<figureCitation id="133E2A74C33DFFDC9C08FEA39274BF10" box="[279,367,312,338]" captionStart="Fig" captionStartId="9.[189,232,1849,1875]" captionTargetBox="[220,1368,265,1811]" captionTargetId="figure-16@9.[220,1368,265,1811]" captionTargetPageId="9" captionText="Fig. 3. External shell morphology of Landouria spp. from northeastern Thailand. A. Landouria circinata sp. nov., holotype (NHMSU-00023). B. L. tuberculata sp. nov., holotype (NHMSU-00025). C. L. trochomorphoides sp. nov., holotype (NHMSU-00027). D. L. chloritoides sp. nov., holotype (NHMSU-00029). E. L. elegans sp. nov., holotype (NHMSU-00032). F. L. diplogramma (Möllendorff, 1902) comb. nov. (NHMSU-00034). Scale bar = 10 mm." figureDoi="http://doi.org/10.5281/zenodo.5514845" httpUri="https://zenodo.org/record/5514845/files/figure.png" pageId="19" pageNumber="161">Fig. 3F</figureCitation>
|
||
,
|
||
<tableCitation id="C687034AC33DFFDC9C61FEA392CCBF10" box="[382,471,312,338]" captionStart="Table 1" captionStartId="2.[185,255,857,885]" captionTargetBox="[195,1921,987,1444]" captionTargetPageId="2" captionText="Table 1. Shell measurements (mm) of six new species of the genus Landouria Godwin-Austen, 1918 from northeastern Thailand." httpUri="http://table.plazi.org/id/DF7A6679C32CFFCD9DA6FCC295B0BD37" pageId="19" pageNumber="161" tableUuid="DF7A6679C32CFFCD9DA6FCC295B0BD37">Table 1</tableCitation>
|
||
). Dextral, small, depressed conical. Whorls 5½, suture rather shallow, apex obtuse and depressed, whitish corneous, with a pale brown peripheral band. Protoconch almost smooth. Umbilicus moderately deep and wide. Aperture oval oblique, with thin, solid, and weakly reflected lip.
|
||
</paragraph>
|
||
<paragraph id="8BBA36F1C33DFFDC9DA2FE7590C4BCA2" blockId="19.[188,1399,493,736]" pageId="19" pageNumber="161">
|
||
GENITAL SYSTEM (n = 3) (
|
||
<figureCitation id="133E2A74C33DFFDC9CC7FE759137BC4A" box="[472,556,493,520]" captionStart="Fig" captionStartId="15.[189,232,1816,1842]" captionTargetBox="[283,1352,278,1767]" captionTargetId="figure-15@15.[232,1356,265,1778]" captionTargetPageId="15" captionText="Fig. 5. Illustrations of the genital system of Landouria spp. from northeastern Thailand. A. Landouria circinata sp. nov., paratype (NHMSU-00024). B. L. tuberculata sp. nov., paratype (NHMSU-00026). C. L. trochomorphoides sp. nov., paratype (NHMSU-00028). D. L. chloritoides sp. nov., paratype (NHMSU-00030). E. L. elegans sp. nov., paratype (NHMSU-00033). F. L. diplogramma (Möllendorff, 1902) comb. nov. (NHMSU-00034). Internal sculpture of a = flagellum; b = penis; c = vagina. Scale bars = 10 mm." figureDoi="http://doi.org/10.5281/zenodo.5514849" httpUri="https://zenodo.org/record/5514849/files/figure.png" pageId="19" pageNumber="161">Fig. 5F</figureCitation>
|
||
,
|
||
<tableCitation id="C687034AC33DFFDC9F29FE759194BC4A" box="[566,655,494,520]" captionStart="Table 4" captionStartId="10.[185,255,544,572]" captionTargetBox="[196,1908,758,1739]" captionTargetPageId="10" captionText="Table 4. Comparison of shells, radulae, and genital systems of Landouria spp. from northeastern Thailand. References: 1 = Tumpeesuwan & Tumpeesuwan (2019); 2 = Nahok (2020); 3 = this study. *In Tumpeesuwan & Tumpeesuwan (2019) indicated the number of teeth per row in Landouria strobiloides varies from 3 to 53. This lowest row has only 3 teeth. It is the uppermost row, probably lacerated and peel off. Unfortunately, we could not re-checked the complete specimens in this time being, so, this point need further investigation and proof." httpUri="http://table.plazi.org/id/DF7A6679C324FFC59DA6FDBB969CBCE7" pageId="19" pageNumber="161" tableUuid="DF7A6679C324FFC59DA6FDBB969CBCE7">Table 4</tableCitation>
|
||
). Penis divided into two short portions, the portion that connects to epiphallus is swollen and the portion that connects to atrium is constricted; internally with three corrugated longitudinal pilasters; with short, rather rounded verge. Flagellum like a short protrusion, internally with two narrow longitudinal pilasters. Vas deferens long, cylindrical, entering the epiphallus at base. Free oviduct very short. Vagina twice as long as free oviduct, internally with five thick, corrugated longitudinal pilasters. Proximal part of gametolytic sac slightly thicker than the cylindrical tube, with a medium-sized oval sax at distal end. Uterus and prostate gland long.
|
||
</paragraph>
|
||
<paragraph id="8BBA36F1C33DFFDC9DA2FC8B91F7BDF8" blockId="19.[189,1401,783,954]" pageId="19" pageNumber="161">
|
||
RADULA (n = 3). Comprises 121–125 transverse rows with 61–69 teeth per row; radular formula: (23–25)- (7–9)-1-(7–9)-(23–25). Central tooth usually symmetric, unicuspid, triangular (
|
||
<figureCitation id="133E2A74C33DFFDC9971FCA897D1BD0C" box="[1134,1226,819,846]" captionStart="Fig" captionStartId="18.[189,232,1783,1809]" captionTargetBox="[244,1343,265,1744]" captionTargetId="figure-12@18.[244,1343,265,1744]" captionTargetPageId="18" captionText="Fig. 6.Radula morphology ofLandouria spp. from northeasternThailand.A–C, G–R. Right side of central teeth. D–F. Left side of central teeth. A–C. Landouria circinata sp. nov., paratype (NHMSU-00024). D–F. L. tuberculata sp. nov., paratype (NHMSU-00026). G–I. L. trochomorphoides sp. nov., paratype (NHMSU-00028). J–L. L. chloritoides sp. nov., paratype (NHMSU-00030). M–O. L. elegans sp. nov., paratype (NHMSU-00033). P–R. L. diplogramma (Möllendorff, 1902) comb. nov. (NHMSU-00034). Numbers indicate order of lateral and marginal teeth. Abbreviation: c = central tooth. Scale bars = 25 µm." figureDoi="http://doi.org/10.5281/zenodo.5514851" httpUri="https://zenodo.org/record/5514851/files/figure.png" pageId="19" pageNumber="161">Fig. 6P</figureCitation>
|
||
). Lateral teeth longer and larger than central teeth, bicuspid, ectocone small, with sharp cusp (
|
||
<figureCitation id="133E2A74C33DFFDC9944FCCC97A2BD30" box="[1115,1209,855,882]" captionStart="Fig" captionStartId="18.[189,232,1783,1809]" captionTargetBox="[244,1343,265,1744]" captionTargetId="figure-12@18.[244,1343,265,1744]" captionTargetPageId="18" captionText="Fig. 6.Radula morphology ofLandouria spp. from northeasternThailand.A–C, G–R. Right side of central teeth. D–F. Left side of central teeth. A–C. Landouria circinata sp. nov., paratype (NHMSU-00024). D–F. L. tuberculata sp. nov., paratype (NHMSU-00026). G–I. L. trochomorphoides sp. nov., paratype (NHMSU-00028). J–L. L. chloritoides sp. nov., paratype (NHMSU-00030). M–O. L. elegans sp. nov., paratype (NHMSU-00033). P–R. L. diplogramma (Möllendorff, 1902) comb. nov. (NHMSU-00034). Numbers indicate order of lateral and marginal teeth. Abbreviation: c = central tooth. Scale bars = 25 µm." figureDoi="http://doi.org/10.5281/zenodo.5514851" httpUri="https://zenodo.org/record/5514851/files/figure.png" pageId="19" pageNumber="161">Fig. 6Q</figureCitation>
|
||
). Marginal teeth gradually changing from bicuspid to tricuspid, small endocone; mesocone large lanceolate; ectocone triangular with two to four tiny cusps (
|
||
<figureCitation id="133E2A74C33DFFDC9F9DFC0491C4BDF8" box="[642,735,927,954]" captionStart="Fig" captionStartId="18.[189,232,1783,1809]" captionTargetBox="[244,1343,265,1744]" captionTargetId="figure-12@18.[244,1343,265,1744]" captionTargetPageId="18" captionText="Fig. 6.Radula morphology ofLandouria spp. from northeasternThailand.A–C, G–R. Right side of central teeth. D–F. Left side of central teeth. A–C. Landouria circinata sp. nov., paratype (NHMSU-00024). D–F. L. tuberculata sp. nov., paratype (NHMSU-00026). G–I. L. trochomorphoides sp. nov., paratype (NHMSU-00028). J–L. L. chloritoides sp. nov., paratype (NHMSU-00030). M–O. L. elegans sp. nov., paratype (NHMSU-00033). P–R. L. diplogramma (Möllendorff, 1902) comb. nov. (NHMSU-00034). Numbers indicate order of lateral and marginal teeth. Abbreviation: c = central tooth. Scale bars = 25 µm." figureDoi="http://doi.org/10.5281/zenodo.5514851" httpUri="https://zenodo.org/record/5514851/files/figure.png" pageId="19" pageNumber="161">Fig. 6R</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C31F657AC33DFFDC9DA2FC729219B8DA" pageId="19" pageNumber="161" type="discussion">
|
||
<paragraph id="8BBA36F1C33DFFDC9DA2FC729234BA41" blockId="19.[189,303,1001,1027]" box="[189,303,1001,1027]" pageId="19" pageNumber="161">
|
||
<heading id="D0F2819DC33DFFDC9DA2FC729234BA41" bold="true" box="[189,303,1001,1027]" fontSize="11" level="3" pageId="19" pageNumber="161" reason="3">
|
||
<emphasis id="B971EAE3C33DFFDC9DA2FC729234BA41" bold="true" box="[189,303,1001,1027]" pageId="19" pageNumber="161">Remarks</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="8BBA36F1C33DFFDC9DA2FB839193BB6D" blockId="19.[187,1399,1048,1327]" pageId="19" pageNumber="161">
|
||
Specimens deposited in the Leiden Museum,
|
||
<collectingCountry id="F3127661C33DFFDC9FE7FB8390D9BA70" box="[760,962,1048,1074]" name="Netherlands" pageId="19" pageNumber="161">The Netherlands</collectingCountry>
|
||
, RMNH.MOL.309867 and RMNH. MOL.309851 (https://images.app.goo.gl/PgrpqKaWuSrgjdvN9), are identified as
|
||
<taxonomicName id="4C054D72C33DFFDC99F7FBA79122BA39" authority="Mollendorff, 1902" authorityName="Mollendorff" authorityYear="1902" class="Gastropoda" family="Camaenidae" genus="Plectotropis" kingdom="Animalia" order="Stylommatophora" pageId="19" pageNumber="161" phylum="Mollusca" rank="species" species="diplogramme">
|
||
<emphasis id="B971EAE3C33DFFDC99F7FBA79242BA38" italics="true" pageId="19" pageNumber="161">Plectotropis diplogramme</emphasis>
|
||
<bibRefCitation id="EF944B00C33DFFDC9C40FBFB9122BA39" author="Mollendorff O. F." box="[351,569,1120,1147]" pageId="19" pageNumber="161" pagination="153 - 160" refId="ref10489" refString="Mollendorff O. F. 1902. Neue Arten und Unterarten von Fruhstorfer in Siam gesammelt. Nachrichtsblatt der Deutschen Malakozoologischen Gesellschaft 34 (9 - 10): 153 - 160." type="journal article" year="1902">Möllendorff, 1902</bibRefCitation>
|
||
</taxonomicName>
|
||
. These specimens appear identical to specimens from Khao See Siad Ah in Nakhon Ratchasima Province, therefore we used the name “
|
||
<taxonomicName id="4C054D72C33DFFDC9EB4FB1F921DBA81" authority="(Mollendorff, 1902)" authorityName="Nahok & Tumpeesuwan & Tumpeesuwan" authorityYear="2021" baseAuthorityName="Mollendorff" baseAuthorityYear="1902" class="Gastropoda" family="Camaenidae" genus="Landouria" kingdom="Animalia" order="Stylommatophora" pageId="19" pageNumber="161" phylum="Mollusca" rank="species" species="diplogramma">
|
||
<emphasis id="B971EAE3C33DFFDC9EB4FB1F97D5BADC" box="[939,1230,1156,1182]" italics="true" pageId="19" pageNumber="161">Landouria diplogramma</emphasis>
|
||
(
|
||
<bibRefCitation id="EF944B00C33DFFDC99C1FB1F93E7BA81" author="Mollendorff O. F." pageId="19" pageNumber="161" pagination="153 - 160" refId="ref10489" refString="Mollendorff O. F. 1902. Neue Arten und Unterarten von Fruhstorfer in Siam gesammelt. Nachrichtsblatt der Deutschen Malakozoologischen Gesellschaft 34 (9 - 10): 153 - 160." type="journal article" year="1902">Möllendorff, 1902</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
” for this species in the present study.
|
||
<taxonomicName id="4C054D72C33DFFDC9FCAFB3390E2BA80" authorityName="Nahok & Tumpeesuwan & Tumpeesuwan" authorityYear="2021" baseAuthorityName="Mollendorff" baseAuthorityYear="1902" box="[725,1017,1192,1218]" class="Gastropoda" family="Camaenidae" genus="Landouria" kingdom="Animalia" order="Stylommatophora" pageId="19" pageNumber="161" phylum="Mollusca" rank="species" species="diplogramma">
|
||
<emphasis id="B971EAE3C33DFFDC9FCAFB3390E2BA80" box="[725,1017,1192,1218]" italics="true" pageId="19" pageNumber="161">Landouria diplogramma</emphasis>
|
||
</taxonomicName>
|
||
differs from other Thai species of
|
||
<taxonomicName id="4C054D72C33DFFDC9DC1FB579241BAA4" authorityName="Godwin-Austen" authorityYear="1918" box="[222,346,1228,1254]" class="Gastropoda" family="Camaenidae" genus="Landouria" kingdom="Animalia" order="Stylommatophora" pageId="19" pageNumber="161" phylum="Mollusca" rank="genus">
|
||
<emphasis id="B971EAE3C33DFFDC9DC1FB579241BAA4" box="[222,346,1228,1254]" italics="true" pageId="19" pageNumber="161">Landouria</emphasis>
|
||
</taxonomicName>
|
||
by its relatively small lustrous shell without peripheral keel, but with a pale brown band present above periphery (
|
||
<figureCitation id="133E2A74C33DFFDC9CF1FB6B9151BB49" box="[494,586,1264,1291]" captionStart="Fig" captionStartId="9.[189,232,1849,1875]" captionTargetBox="[220,1368,265,1811]" captionTargetId="figure-16@9.[220,1368,265,1811]" captionTargetPageId="9" captionText="Fig. 3. External shell morphology of Landouria spp. from northeastern Thailand. A. Landouria circinata sp. nov., holotype (NHMSU-00023). B. L. tuberculata sp. nov., holotype (NHMSU-00025). C. L. trochomorphoides sp. nov., holotype (NHMSU-00027). D. L. chloritoides sp. nov., holotype (NHMSU-00029). E. L. elegans sp. nov., holotype (NHMSU-00032). F. L. diplogramma (Möllendorff, 1902) comb. nov. (NHMSU-00034). Scale bar = 10 mm." figureDoi="http://doi.org/10.5281/zenodo.5514845" httpUri="https://zenodo.org/record/5514845/files/figure.png" pageId="19" pageNumber="161">Figs 3F</figureCitation>
|
||
,
|
||
<figureCitation id="133E2A74C33DFFDC9F46FB6A9167BB49" box="[601,636,1265,1291]" captionStart="Fig" captionStartId="12.[189,232,1921,1947]" captionTargetBox="[193,1395,278,1870]" captionTargetId="figure-12@12.[193,1395,265,1883]" captionTargetPageId="12" captionText="Fig. 4. Living adults of Landouria spp. from northeastern Thailand. A. Landouria circinata sp. nov. B. L. tuberculata sp. nov. C. L. trochomorphoides sp. nov. D. L. chloritoides sp. nov. E. L. elegans sp. nov. F. L. diplogramma (Möllendorff, 1902) comb. nov. Scale bars = 10 mm." figureDoi="http://doi.org/10.5281/zenodo.5514847" httpUri="https://zenodo.org/record/5514847/files/figure.png" pageId="19" pageNumber="161">4F</figureCitation>
|
||
). Its radular teeth are relatively short (
|
||
<figureCitation id="133E2A74C33DFFDC9955FB6B97D2BB49" box="[1098,1225,1264,1291]" captionStart="Fig" captionStartId="18.[189,232,1783,1809]" captionTargetBox="[244,1343,265,1744]" captionTargetId="figure-12@18.[244,1343,265,1744]" captionTargetPageId="18" captionText="Fig. 6.Radula morphology ofLandouria spp. from northeasternThailand.A–C, G–R. Right side of central teeth. D–F. Left side of central teeth. A–C. Landouria circinata sp. nov., paratype (NHMSU-00024). D–F. L. tuberculata sp. nov., paratype (NHMSU-00026). G–I. L. trochomorphoides sp. nov., paratype (NHMSU-00028). J–L. L. chloritoides sp. nov., paratype (NHMSU-00030). M–O. L. elegans sp. nov., paratype (NHMSU-00033). P–R. L. diplogramma (Möllendorff, 1902) comb. nov. (NHMSU-00034). Numbers indicate order of lateral and marginal teeth. Abbreviation: c = central tooth. Scale bars = 25 µm." figureDoi="http://doi.org/10.5281/zenodo.5514851" httpUri="https://zenodo.org/record/5514851/files/figure.png" pageId="19" pageNumber="161">Fig. 6P–R</figureCitation>
|
||
). Its penis and flagellum are relatively short (
|
||
<figureCitation id="133E2A74C33DFFDC9F3EFA8F9160BB6D" box="[545,635,1300,1327]" captionStart="Fig" captionStartId="15.[189,232,1816,1842]" captionTargetBox="[283,1352,278,1767]" captionTargetId="figure-15@15.[232,1356,265,1778]" captionTargetPageId="15" captionText="Fig. 5. Illustrations of the genital system of Landouria spp. from northeastern Thailand. A. Landouria circinata sp. nov., paratype (NHMSU-00024). B. L. tuberculata sp. nov., paratype (NHMSU-00026). C. L. trochomorphoides sp. nov., paratype (NHMSU-00028). D. L. chloritoides sp. nov., paratype (NHMSU-00030). E. L. elegans sp. nov., paratype (NHMSU-00033). F. L. diplogramma (Möllendorff, 1902) comb. nov. (NHMSU-00034). Internal sculpture of a = flagellum; b = penis; c = vagina. Scale bars = 10 mm." figureDoi="http://doi.org/10.5281/zenodo.5514849" httpUri="https://zenodo.org/record/5514849/files/figure.png" pageId="19" pageNumber="161">Fig. 5F</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BBA36F1C33DFFDC9DA2FAC59219B8DA" blockId="19.[188,1400,1374,1688]" pageId="19" pageNumber="161">
|
||
The genus
|
||
<taxonomicName id="4C054D72C33DFFDC9C5BFAC59191BB3A" authority="Schileyko, 2004" authorityName="Schileyko" authorityYear="2004" box="[324,650,1374,1400]" class="Gastropoda" family="Camaenidae" genus="Thaitropis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="19" pageNumber="161" phylum="Mollusca" rank="genus">
|
||
<emphasis id="B971EAE3C33DFFDC9C5BFAC592A6BB3A" box="[324,445,1374,1400]" italics="true" pageId="19" pageNumber="161">Thaitropis</emphasis>
|
||
<bibRefCitation id="EF944B00C33DFFDC9CD7FAC59191BB3A" author="Schileyko A. A." box="[456,650,1374,1400]" pageId="19" pageNumber="161" pagination="1627 - 1764" refId="ref11091" refString="Schileyko A. A. 2004. Treatise on recent terrestrial pulmonate molluscs. Part 12. Bradybaenidae, Monadeniidae, Xanthonychidae, Epiphragmophoridae, Helminthoglyptidae, Elonidae, Humboldtianidae, Sphincterochilidae, Cochlicellidae. Ruthenica Supplement 2: 1627 - 1764." type="journal article" year="2004">Schileyko, 2004</bibRefCitation>
|
||
</taxonomicName>
|
||
was recently reported in
|
||
<collectingCountry id="F3127661C33DFFDC9EDCFAC59730BB3A" box="[963,1067,1374,1400]" name="Thailand" pageId="19" pageNumber="161">Thailand</collectingCountry>
|
||
(
|
||
<bibRefCitation id="EF944B00C33DFFDC9922FAC597E6BB3A" author="Schileyko A. A." box="[1085,1277,1374,1400]" pageId="19" pageNumber="161" pagination="1627 - 1764" refId="ref11091" refString="Schileyko A. A. 2004. Treatise on recent terrestrial pulmonate molluscs. Part 12. Bradybaenidae, Monadeniidae, Xanthonychidae, Epiphragmophoridae, Helminthoglyptidae, Elonidae, Humboldtianidae, Sphincterochilidae, Cochlicellidae. Ruthenica Supplement 2: 1627 - 1764." type="journal article" year="2004">Schileyko 2004</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EF944B00C33DFFDC9812FAC59258BBDE" author="Sutcharit C. & Tongkerd P. & Panha S." pageId="19" pageNumber="161" refId="ref11218" refString="Sutcharit C., Tongkerd P. & Panha S. 2017. Land Snails the Invaluable Bio-Resources for the Kingdom of Thailand. The Thailand Research Fund, Bangkok. [In Thai.]" type="book" year="2017">
|
||
Sutcharit
|
||
<emphasis id="B971EAE3C33DFFDC9DA2FA1893E1BBDE" box="[189,250,1410,1436]" italics="true" pageId="19" pageNumber="161">et al.</emphasis>
|
||
2017
|
||
</bibRefCitation>
|
||
). Yet, this genus was synonymized with
|
||
<taxonomicName id="4C054D72C33DFFDC9E32FA1990B2BBDE" authorityName="Godwin-Austen" authorityYear="1918" box="[813,937,1410,1436]" class="Gastropoda" family="Camaenidae" genus="Landouria" kingdom="Animalia" order="Stylommatophora" pageId="19" pageNumber="161" phylum="Mollusca" rank="genus">
|
||
<emphasis id="B971EAE3C33DFFDC9E32FA1990B2BBDE" box="[813,937,1410,1436]" italics="true" pageId="19" pageNumber="161">Landouria</emphasis>
|
||
</taxonomicName>
|
||
by
|
||
<bibRefCitation id="EF944B00C33DFFDC9EC8FA1997E0BBDE" author="Nurinsiyah A. S. & Neiber M. T. & Hausdorf B." box="[983,1275,1410,1436]" pageId="19" pageNumber="161" pagination="1 - 73" refId="ref10615" refString="Nurinsiyah A. S., Neiber M. T. & Hausdorf B. 2019. Revision of the land snail genus Landouria Godwin- Austen, 1918 (Gastropoda, Camaenidae) from Java. European Journal of Taxonomy 526: 1 - 73. https: // doi. org / 10.5852 / ejt. 2019.526" type="journal article" year="2019">
|
||
Nurinsiyah
|
||
<emphasis id="B971EAE3C33DFFDC997AFA1897B9BBDE" box="[1125,1186,1410,1436]" italics="true" pageId="19" pageNumber="161">et al.</emphasis>
|
||
(2019)
|
||
</bibRefCitation>
|
||
. Based on the illustration provided by
|
||
<bibRefCitation id="EF944B00C33DFFDC9F0BFA3D9003BB82" author="Sutcharit C. & Tongkerd P. & Panha S." box="[532,792,1446,1472]" pageId="19" pageNumber="161" refId="ref11218" refString="Sutcharit C., Tongkerd P. & Panha S. 2017. Land Snails the Invaluable Bio-Resources for the Kingdom of Thailand. The Thailand Research Fund, Bangkok. [In Thai.]" type="book" year="2017">
|
||
Sutcharit
|
||
<emphasis id="B971EAE3C33DFFDC9F99FA3C91DEBB82" box="[646,709,1446,1472]" italics="true" pageId="19" pageNumber="161">et al.</emphasis>
|
||
(2017
|
||
</bibRefCitation>
|
||
: fig. 6–32e), we identify their Thai
|
||
<taxonomicName id="4C054D72C33DFFDC99CEFA3D966DBB83" box="[1233,1398,1446,1473]" class="Gastropoda" family="Camaenidae" genus="Thaitropis" kingdom="Animalia" order="Stylommatophora" pageId="19" pageNumber="161" phylum="Mollusca" rank="species" species="undetermined">
|
||
<emphasis id="B971EAE3C33DFFDC99CEFA3D9651BB82" box="[1233,1354,1446,1472]" italics="true" pageId="19" pageNumber="161">Thaitropis</emphasis>
|
||
sp.
|
||
</taxonomicName>
|
||
as
|
||
<taxonomicName id="4C054D72C33DFFDC9DC1FA509286BBA6" authorityName="Nahok & Tumpeesuwan & Tumpeesuwan" authorityYear="2021" baseAuthorityName="Mollendorff" baseAuthorityYear="1902" box="[222,413,1482,1508]" class="Gastropoda" family="Camaenidae" genus="Landouria" kingdom="Animalia" order="Stylommatophora" pageId="19" pageNumber="161" phylum="Mollusca" rank="species" species="diplogramma">
|
||
<emphasis id="B971EAE3C33DFFDC9DC1FA509286BBA6" box="[222,413,1482,1508]" italics="true" pageId="19" pageNumber="161">L. diplogramma</emphasis>
|
||
</taxonomicName>
|
||
, because it has the same shell morphology and colours:
|
||
<taxonomicName id="4C054D72C33DFFDC995BFA519287B84A" authority="Mollendorff, 1902" authorityName="Mollendorff" authorityYear="1902" class="Gastropoda" family="Camaenidae" genus="Plectotropis" kingdom="Animalia" order="Stylommatophora" pageId="19" pageNumber="161" phylum="Mollusca" rank="species" species="diplogramme">
|
||
<emphasis id="B971EAE3C33DFFDC995BFA519663BBA6" box="[1092,1400,1482,1508]" italics="true" pageId="19" pageNumber="161">Plectotropis diplogramme</emphasis>
|
||
<bibRefCitation id="EF944B00C33DFFDC9DA2FA759287B84A" author="Mollendorff O. F." box="[189,412,1518,1544]" pageId="19" pageNumber="161" pagination="153 - 160" refId="ref10489" refString="Mollendorff O. F. 1902. Neue Arten und Unterarten von Fruhstorfer in Siam gesammelt. Nachrichtsblatt der Deutschen Malakozoologischen Gesellschaft 34 (9 - 10): 153 - 160." type="journal article" year="1902">Möllendorff, 1902</bibRefCitation>
|
||
</taxonomicName>
|
||
, RMNH.MOL.309867, and
|
||
<taxonomicName id="4C054D72C33DFFDC9FEAFA7497AAB84A" authority="(Mollendorff, 1902)" authorityName="Nahok & Tumpeesuwan & Tumpeesuwan" authorityYear="2021" baseAuthorityName="Mollendorff" baseAuthorityYear="1902" box="[757,1201,1518,1544]" class="Gastropoda" family="Camaenidae" genus="Landouria" kingdom="Animalia" order="Stylommatophora" pageId="19" pageNumber="161" phylum="Mollusca" rank="species" species="diplogramma">
|
||
<emphasis id="B971EAE3C33DFFDC9FEAFA7490ADB84A" box="[757,950,1518,1544]" italics="true" pageId="19" pageNumber="161">L. diplogramma</emphasis>
|
||
(
|
||
<bibRefCitation id="EF944B00C33DFFDC9ED6FA7597B2B84A" author="Mollendorff O. F." box="[969,1193,1518,1544]" pageId="19" pageNumber="161" pagination="153 - 160" refId="ref10489" refString="Mollendorff O. F. 1902. Neue Arten und Unterarten von Fruhstorfer in Siam gesammelt. Nachrichtsblatt der Deutschen Malakozoologischen Gesellschaft 34 (9 - 10): 153 - 160." type="journal article" year="1902">Möllendorff, 1902</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
from Khao See Siad Ah,
|
||
<collectingRegion id="49C1F813C33DFFDC9C31F9899105B86E" box="[302,542,1554,1580]" country="Thailand" name="Nakhon Ratchasima" pageId="19" pageNumber="161">Nakhon Ratchasima</collectingRegion>
|
||
(NHMSU-00034) (
|
||
<figureCitation id="133E2A74C33DFFDC9E14F9899073B86E" box="[779,872,1554,1580]" captionStart="Fig" captionStartId="9.[189,232,1849,1875]" captionTargetBox="[220,1368,265,1811]" captionTargetId="figure-16@9.[220,1368,265,1811]" captionTargetPageId="9" captionText="Fig. 3. External shell morphology of Landouria spp. from northeastern Thailand. A. Landouria circinata sp. nov., holotype (NHMSU-00023). B. L. tuberculata sp. nov., holotype (NHMSU-00025). C. L. trochomorphoides sp. nov., holotype (NHMSU-00027). D. L. chloritoides sp. nov., holotype (NHMSU-00029). E. L. elegans sp. nov., holotype (NHMSU-00032). F. L. diplogramma (Möllendorff, 1902) comb. nov. (NHMSU-00034). Scale bar = 10 mm." figureDoi="http://doi.org/10.5281/zenodo.5514845" httpUri="https://zenodo.org/record/5514845/files/figure.png" pageId="19" pageNumber="161">Figs 3F</figureCitation>
|
||
,
|
||
<figureCitation id="133E2A74C33DFFDC9E69F9899081B86E" box="[886,922,1554,1580]" captionStart="Fig" captionStartId="12.[189,232,1921,1947]" captionTargetBox="[193,1395,278,1870]" captionTargetId="figure-12@12.[193,1395,265,1883]" captionTargetPageId="12" captionText="Fig. 4. Living adults of Landouria spp. from northeastern Thailand. A. Landouria circinata sp. nov. B. L. tuberculata sp. nov. C. L. trochomorphoides sp. nov. D. L. chloritoides sp. nov. E. L. elegans sp. nov. F. L. diplogramma (Möllendorff, 1902) comb. nov. Scale bars = 10 mm." figureDoi="http://doi.org/10.5281/zenodo.5514847" httpUri="https://zenodo.org/record/5514847/files/figure.png" pageId="19" pageNumber="161">4F</figureCitation>
|
||
). Unfortunately, the whereabouts of the
|
||
<typeStatus id="54BE8853C33DFFDC9DA2F9AC93EBB813" box="[189,240,1591,1617]" pageId="19" pageNumber="161">type</typeStatus>
|
||
specimen of
|
||
<taxonomicName id="4C054D72C33DFFDC9C96F9AD91A1B812" authorityName="Mollendorff" authorityYear="1902" box="[393,698,1590,1616]" class="Gastropoda" family="Camaenidae" genus="Plectotropis" kingdom="Animalia" order="Stylommatophora" pageId="19" pageNumber="161" phylum="Mollusca" rank="species" species="diplogramme">
|
||
<emphasis id="B971EAE3C33DFFDC9C96F9AD91A1B812" box="[393,698,1590,1616]" italics="true" pageId="19" pageNumber="161">Plectotropis diplogramme</emphasis>
|
||
</taxonomicName>
|
||
are unknown. Yet, it might be deposited in the collections of the Senckenberg Museum in Frankfurt,
|
||
<collectingCountry id="F3127661C33DFFDC9F92F9C191E1B836" box="[653,762,1626,1652]" name="Germany" pageId="19" pageNumber="161">Germany</collectingCountry>
|
||
where it should be looked up and checked in a further study.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C31F657AC33DFFDC9DA2F9539704B918" pageId="19" pageNumber="161" type="distribution">
|
||
<paragraph id="8BBA36F1C33DFFDC9DA2F953924DB8A0" blockId="19.[189,342,1736,1762]" box="[189,342,1736,1762]" pageId="19" pageNumber="161">
|
||
<heading id="D0F2819DC33DFFDC9DA2F953924DB8A0" bold="true" box="[189,342,1736,1762]" fontSize="11" level="3" pageId="19" pageNumber="161" reason="3">
|
||
<emphasis id="B971EAE3C33DFFDC9DA2F953924DB8A0" bold="true" box="[189,342,1736,1762]" pageId="19" pageNumber="161">Distribution</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="8BBA36F1C33DFFDC9DA2F96C9704B918" blockId="19.[189,1399,1783,1882]" pageId="19" pageNumber="161">
|
||
<taxonomicName id="4C054D72C33DFFDC9DA2F96C92C5B953" authorityName="Nahok & Tumpeesuwan & Tumpeesuwan" authorityYear="2021" baseAuthorityName="Mollendorff" baseAuthorityYear="1902" box="[189,478,1783,1809]" class="Gastropoda" family="Camaenidae" genus="Landouria" kingdom="Animalia" order="Stylommatophora" pageId="19" pageNumber="161" phylum="Mollusca" rank="species" species="diplogramma">
|
||
<emphasis id="B971EAE3C33DFFDC9DA2F96C92C5B953" box="[189,478,1783,1809]" italics="true" pageId="19" pageNumber="161">Landouria diplogramma</emphasis>
|
||
</taxonomicName>
|
||
was collected by local people in the mountain range known as Khao See Siad Ah, Pak Chong District,
|
||
<collectingRegion id="49C1F813C33DFFDC9CF2F8809048B977" box="[493,851,1819,1845]" country="Thailand" name="Nakhon Ratchasima" pageId="19" pageNumber="161">Nakhon Ratchasima Province</collectingRegion>
|
||
,
|
||
<collectingCountry id="F3127661C33DFFDC9E7DF88090D5B977" box="[866,974,1819,1845]" name="Thailand" pageId="19" pageNumber="161">Thailand</collectingCountry>
|
||
. Other reported localities comprise
|
||
<collectingRegion id="49C1F813C33DFFDC9DA2F8A49231B91B" box="[189,298,1855,1881]" country="Thailand" name="Krung Thep Mahanakhon" pageId="19" pageNumber="161">Bangkok</collectingRegion>
|
||
, Hinlap, Muoklek, Kanburi (
|
||
<collectingRegion id="49C1F813C33DFFDC9F63F8A4903FB918" box="[636,804,1855,1882]" country="Thailand" name="Kanchanaburi" pageId="19" pageNumber="161">Kanchanaburi</collectingRegion>
|
||
) (
|
||
<bibRefCitation id="EF944B00C33DFFDC9E24F8A49709B918" author="Mollendorff O. F." box="[827,1042,1855,1882]" pageId="19" pageNumber="161" pagination="153 - 160" refId="ref10489" refString="Mollendorff O. F. 1902. Neue Arten und Unterarten von Fruhstorfer in Siam gesammelt. Nachrichtsblatt der Deutschen Malakozoologischen Gesellschaft 34 (9 - 10): 153 - 160." type="journal article" year="1902">Möllendorff 1902</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |