127 lines
15 KiB
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127 lines
15 KiB
XML
<document id="01E09177ADDDFFC92E204BCA9F99EE0D" ID-DOI="10.5281/zenodo.176960" ID-GBIF-Dataset="a67acf3d-f43a-4d16-b437-aacf9f0d09ec" ID-ISSN="1175-5326" ID-Zenodo-Dep="176960" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1459844304451" checkinUser="plazi" docAuthor="Ziegler, Thomas, Hendrix, Ralf, Thanh, Vu Ngoc, Vogt, Martina, Forster, Bernhard & Kien, Dang Ngoc" docDate="2007" docId="03AC87D0B113FF99FF469690FC9E8985" docLanguage="en" docName="zt01493p040.pdf" docOrigin="Zootaxa 1493" docStyle="DocumentStyle:6581673A57F01A3145754A1E615EDFF0.4:Zootaxa.2007-2008.journal_article" docStyleId="6581673A57F01A3145754A1E615EDFF0" docStyleName="Zootaxa.2007-2008.journal_article" docStyleVersion="4" docTitle="Calamaria thanhi Ziegler & Le 2005" docType="treatment" docVersion="9" lastPageNumber="8" masterDocId="FF95FFA8B115FF9EFFD19607FFD28827" masterDocTitle="The diversity of a snake community in a karst forest ecosystem in the central Truong Son, Vietnam, with an identification key" masterLastPageNumber="40" masterPageNumber="1" pageNumber="7" updateTime="1698224724941" updateUser="plazi">
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<mods:titleInfo id="CC15B229A6FB153C9E09545F16CD73B4">
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<mods:title id="DA7DE20BC0C564F223892D5276ACA57F">The diversity of a snake community in a karst forest ecosystem in the central Truong Son, Vietnam, with an identification key</mods:title>
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</mods:titleInfo>
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<mods:name id="863E7076D0947F6F54507E9419B0A000" type="personal">
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<mods:roleTerm id="54D630418FC0A09AC00E37813043B57C">Author</mods:roleTerm>
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<mods:namePart id="B3C9F5745021005749F619B928F35413">Ziegler, Thomas</mods:namePart>
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<mods:roleTerm id="FFDCAAE0F436923DCEB61E9B73749039">Author</mods:roleTerm>
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<mods:namePart id="67B7590132399BBCFFBEC59B613F7ACB">Hendrix, Ralf</mods:namePart>
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<mods:roleTerm id="CB1429D739A0723140E9F09E26C1112F">Author</mods:roleTerm>
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<mods:namePart id="DEEC06ED1453A6EE3B171A347EB95237">Thanh, Vu Ngoc</mods:namePart>
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<mods:namePart id="A5C000A946EF32A7B587C56B9FC7F2FF">Vogt, Martina</mods:namePart>
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<mods:roleTerm id="A9B1B3A7A884CDCABE1F88C301AFC6CD">Author</mods:roleTerm>
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<mods:namePart id="475674EF1F7341F39EA06F0434501B6A">Forster, Bernhard</mods:namePart>
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<mods:roleTerm id="CB61783200E7307F46EE7035FEAA08B7">Author</mods:roleTerm>
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<mods:namePart id="3DBB56D776D5748D33B56ED67D0EDDDE">Kien, Dang Ngoc</mods:namePart>
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<mods:title id="F6FBE5E3C5418A42BC9B83F677D0B863">Zootaxa</mods:title>
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<mods:part id="81AC7989AC3DF1CF66E41D87972851D8">
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<mods:date id="58D720418E7D051685D7595600840785">2007</mods:date>
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<mods:detail id="C5A7B22BEEA441CF38893F76068167D1" type="volume">
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<mods:number id="6F3DA7185ACA3F576D5C0CF0B8C89524">1493</mods:number>
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<mods:classification id="DFE04F0B350E01E6ACF740F2B0D533F0">journal article</mods:classification>
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<mods:identifier id="C619271271367A8BF4134B3611161687" type="DOI">10.5281/zenodo.176960</mods:identifier>
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<mods:identifier id="FAC50414EF64EA465ADF6F0839061C8C" type="ISSN">1175-5326</mods:identifier>
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<mods:identifier id="038D1CF4F1E300434117EA06E0CD7CBF" type="Zenodo-Dep">176960</mods:identifier>
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<treatment id="03AC87D0B113FF99FF469690FC9E8985" ID-DOI="http://doi.org/10.5281/zenodo.6247451" ID-GBIF-Taxon="119361345" ID-Zenodo-Dep="6247451" LSID="urn:lsid:plazi:treatment:03AC87D0B113FF99FF469690FC9E8985" httpUri="http://treatment.plazi.org/id/03AC87D0B113FF99FF469690FC9E8985" lastPageId="7" lastPageNumber="8" pageId="6" pageNumber="7">
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<subSubSection id="C31F654DB113FF98FF469690FBD789ED" pageId="6" pageNumber="7" type="nomenclature">
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<paragraph id="8BBA36C6B113FF98FF469690FBAC8896" blockId="6.[151,1150,151,177]" box="[151,1150,151,177]" pageId="6" pageNumber="7">
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<heading id="D0F281AAB113FF98FF469690FBAC8896" bold="true" box="[151,1150,151,177]" fontSize="11" level="1" pageId="6" pageNumber="7" reason="1">
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<emphasis id="B971EAD4B113FF98FF469690FBAC8896" bold="true" box="[151,1150,151,177]" pageId="6" pageNumber="7">
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<taxonomicName id="4C054D45B113FF98FF469690FDB78896" authority="Ziegler & Le, 2005" authorityName="Ziegler & Le" authorityYear="2005" box="[151,613,151,177]" class="Reptilia" family="Colubridae" genus="Calamaria" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="7" phylum="Chordata" rank="species" species="thanhi">
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<emphasis id="B971EAD4B113FF98FF469690FEBD8896" bold="true" box="[151,367,151,177]" italics="true" pageId="6" pageNumber="7">Calamaria thanhi</emphasis>
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<bibRefCitation id="EF944B37B113FF98FEA79690FDB78896" author="Ziegler" box="[374,613,151,177]" pageId="6" pageNumber="40" refString="Ziegler, T. & Le, K. Q. (2005) A new species of reed snake, Calamaria (Squamata: Colubridae), from the central Truong Son (Annamite mountain range), Vietnam. Zootaxa, 1042, 27 - 38." type="journal article" year="2005">Ziegler & Le, 2005</bibRefCitation>
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</taxonomicName>
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: Thanh's reed snake / Ran mai gam Thanh
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</emphasis>
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</heading>
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</paragraph>
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<paragraph id="8BBA36C6B113FF98FF4696EFFBD789ED" blockId="6.[151,1436,232,458]" pageId="6" pageNumber="7">
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<bibRefCitation id="EF944B37B113FF98FF4696EFFE578925" author="Ziegler" box="[151,389,232,258]" pageId="6" pageNumber="40" refString="Ziegler, T. & Le, K. Q. (2005) A new species of reed snake, Calamaria (Squamata: Colubridae), from the central Truong Son (Annamite mountain range), Vietnam. Zootaxa, 1042, 27 - 38." type="journal article" year="2005">Ziegler & Le (2005)</bibRefCitation>
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described
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<taxonomicName id="4C054D45B113FF98FDD596EFFDA08926" box="[516,626,232,257]" class="Reptilia" family="Colubridae" genus="Calamaria" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="7" phylum="Chordata" rank="species" species="thanhi">
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<emphasis id="B971EAD4B113FF98FDD596EFFDA08926" box="[516,626,232,257]" italics="true" pageId="6" pageNumber="7">C. thanhi</emphasis>
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</taxonomicName>
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based on a single female specimen from adjacent to Phong Nha - Ke Bang National Park, Dan Hoa commune, Minh Hoa District, Quang Binh Province. Meanwhile a second specimen came to our attention, that was found during
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<date id="FFBB1006B113FF98FCF4973FFC748975" box="[805,934,312,338]" pageId="6" pageNumber="7" value="2005-04">April 2005</date>
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close to the National Park's border during excavations in a secondary shrub area. Because the species was formerly only known from the heavily destroyed
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<typeStatus id="54BE8864B113FF98FEC1978FFEAB8985" box="[272,377,392,418]" pageId="6" pageNumber="7" type="holotype">holotype</typeStatus>
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, we herein provide first pictures of a better preserved and at the same time of the first male as well as also largest specimen of the barely known species (
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<figureCitation id="133E2A43B113FF98FCB697B7FC2A89ED" box="[871,1016,432,458]" captionStart-0="FIGURE 6" captionStart-1="FIGURE 7" captionStart-2="FIGURE 8" captionStart-3="FIGURE 9" captionStartId-0="6.[151,255,1395,1419]" captionStartId-1="7.[151,255,1255,1279]" captionStartId-2="7.[151,255,1968,1992]" captionStartId-3="8.[151,255,844,868]" captionTargetBox-0="[151,1434,514,1371]" captionTargetBox-1="[151,1434,458,1247]" captionTargetBox-2="[151,1434,1351,1960]" captionTargetBox-3="[151,1435,178,836]" captionTargetId-0="figure@6.[151,1436,514,1371]" captionTargetId-1="figure@7.[151,1436,442,1299]" captionTargetId-2="figure@7.[151,1436,1348,1962]" captionTargetId-3="figure@8.[151,1436,176,838]" captionTargetPageId-0="6" captionTargetPageId-1="7" captionTargetPageId-2="7" captionTargetPageId-3="8" captionText-0="FIGURE 6. Thanh's reed snake (Calamaria thanhi), preserved male from Phong Nha - Ke Bang in dorsal view (SRC PNKB III ’ 06 - 1). Photo: Thomas Ziegler" captionText-1="FIGURE 7. Thanh's reed snake (Calamaria thanhi), preserved male from Phong Nha - Ke Bang in ventral view (SRC" captionText-2="FIGURE 8. Portrait of the preserved male Calamaria thanhi from Phong Nha - Ke Bang (SRC PNKB III ’ 06 - 1). Photo: Thomas Ziegler" captionText-3="FIGURE 9. Ventral view of the head of the preserved male Calamaria thanhi from Phong Nha - Ke Bang (SRC PNKB III ’ 06 - 1). Photo: Thomas Ziegler" httpUri-0="https://zenodo.org/record/176966/files/figure.png" httpUri-1="https://zenodo.org/record/176967/files/figure.png" httpUri-2="https://zenodo.org/record/176968/files/figure.png" httpUri-3="https://zenodo.org/record/176969/files/figure.png" pageId="6" pageNumber="7">Figures 6–9</figureCitation>
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).
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</paragraph>
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</subSubSection>
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<caption id="DF7A664EB113FF98FF469374FD998D8A" httpUri="https://zenodo.org/record/176966/files/figure.png" pageId="6" pageNumber="7" targetBox="[151,1434,514,1371]" targetPageId="6">
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<paragraph id="8BBA36C6B113FF98FF469374FD998D8A" blockId="6.[151,1436,1395,1453]" pageId="6" pageNumber="7">
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<emphasis id="B971EAD4B113FF98FF469374FEC98DAC" bold="true" box="[151,283,1395,1419]" pageId="6" pageNumber="7">FIGURE 6.</emphasis>
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Thanh's reed snake (
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<taxonomicName id="4C054D45B113FF98FDD29374FD118DAD" box="[515,707,1395,1418]" class="Reptilia" family="Colubridae" genus="Calamaria" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="7" phylum="Chordata" rank="species" species="thanhi">
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<emphasis id="B971EAD4B113FF98FDD29374FD118DAD" box="[515,707,1395,1418]" italics="true" pageId="6" pageNumber="7">Calamaria thanhi</emphasis>
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</taxonomicName>
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), preserved male from Phong Nha - Ke Bang in dorsal view (SRC PNKB III ’06-1). Photo: Thomas Ziegler
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</paragraph>
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</caption>
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<subSubSection id="C31F654DB113FF99FF1793DCFC9E8985" lastPageId="7" lastPageNumber="8" pageId="6" pageNumber="7" type="description">
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<paragraph id="8BBA36C6B113FF99FF1793DCFC86890D" blockId="6.[151,1437,1499,2008]" lastBlockId="7.[151,1436,152,418]" lastPageId="7" lastPageNumber="8" pageId="6" pageNumber="7">
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The new specimen (SVL 415, TaL 46, TL
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<quantity id="4CFD9B23B113FF98FD1593DCFCF88DD2" box="[708,810,1499,1525]" metricMagnitude="-1" metricUnit="m" metricValue="4.61" pageId="6" pageNumber="7" unit="mm" value="461.0">461 mm</quantity>
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), that is deposited in the Science Research Centre of the Phong Nha - Ke Bang National Park (SRC PNKB III '06-1), largely agrees with the
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<typeStatus id="54BE8864B113FF98FB7A9004FAC38E3A" box="[1195,1297,1539,1565]" pageId="6" pageNumber="7" type="holotype">holotype</typeStatus>
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description in pattern and scalation (see
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<tableCitation id="C687037DB113FF98FE20902CFD9E8E62" box="[497,588,1579,1605]" captionStart="TABLE 2" captionStartId="8.[151,239,953,977]" captionTargetBox="[159,1276,1079,1990]" captionTargetPageId="8" captionText="TABLE 2. Important measurements, scalation, coloration and dentition characteristics of the female holotype (data from Ziegler & Le 2005) and the newly collected male specimen of Calamaria thanhi from Quang Binh Province; for abbreviations see material and methods." httpUri="http://table.plazi.org/id/DF7A664EB11DFF96FF4695BEFE218C31" pageId="6" pageNumber="7" tableUuid="DF7A664EB11DFF96FF4695BEFE218C31">Table 2</tableCitation>
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), except for: 1) the rostral portion visible from above being somewhat shorter (1.5 mm) than the prefrontal suture (1.6 mm); 2) the left paraparietal being surrounded by seven instead of six shields and scales, as it is the case in the
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<typeStatus id="54BE8864B113FF98FCF5907CFC5C8EB2" box="[804,910,1659,1685]" pageId="6" pageNumber="7" type="holotype">holotype</typeStatus>
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; 3) the third supralabial being about half the size (instead ¾) of second; 4) the new specimen having 184 instead of 198 ventral scales and 28 instead of 21 divided subcaudals; 5) the number of dorsal scale rows is reduced to eight in the male specimen instead of six rows on tail opposite 8th subcaudal posterior to cloaca; 6) the ratio of tail to total length is 0.099 instead of
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<quantity id="4CFD9B23B113FF98FF469119FF258F1F" box="[151,247,1822,1848]" metricMagnitude="0" metricUnit="m" metricValue="1.7271999999999998" pageId="6" pageNumber="7" unit="in" value="68.0">0.068 in</quantity>
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the female
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<typeStatus id="54BE8864B113FF98FEAF9119FE3A8F1F" box="[382,488,1822,1848]" pageId="6" pageNumber="7" type="holotype">holotype</typeStatus>
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; 7) the male specimen bears five instead of four distinct beige to yellowish dorsal bands in the anterior two body thirds, followed by a sixth, but more vestigial light transversal band, that is only discernible from dorsal view; 8) the light dorsal bands subsequent to the first band follow each with distances of
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<quantity id="4CFD9B23B113FF98FEDB9191FEB08F97" box="[266,354,1942,1968]" metricMagnitude="-2" metricUnit="m" metricValue="4.0" metricValueMax="5.0" metricValueMin="3.0" pageId="6" pageNumber="7" unit="cm" value="4.0" valueMax="5.0" valueMin="3.0">3–5 cm</quantity>
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in the male specimen (distances are extending posteriorly); 9) both yellowish markings on the dorsal tail base and on its end are better developed in the male specimen than it is the case in the female
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<typeStatus id="54BE8864B112FF99FF46969FFED38895" box="[151,257,152,178]" pageId="7" pageNumber="8" type="holotype">holotype</typeStatus>
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, although both transversal bands do not completely encircle the dorsum; 10) the yellowish tail base bears only few tiny dark flecks in the male; 11) the underside of the head is not brownish-grey as it is the case in the female
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<typeStatus id="54BE8864B112FF99FEED96EFFE708925" box="[316,418,232,258]" pageId="7" pageNumber="8" type="holotype">holotype</typeStatus>
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but resembles the overall yellow venter coloration. We interpret the afore mentioned differences due to individual variation / sexual dimorphism.
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</paragraph>
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<paragraph id="8BBA36C6B112FF99FF17973FFC9E8985" blockId="7.[151,1436,152,418]" pageId="7" pageNumber="8">In addition, we removed the inverted left hemipenis of the male specimen and dissected the thin organ in the inverted condition. Pedicel and truncus of the forked organ were smooth and without detectable spines, and both lobes were covered with a calyces ornamentation.</paragraph>
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</subSubSection>
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</treatment>
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</document> |