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<document id="2B903D86C50D3B7C8CB3AA2B299234EA" ID-CLB-Dataset="56303" ID-DOI="10.11646/phytotaxa.547.2.3" ID-GBIF-Dataset="69414458-b7e4-4a54-bcb7-1e43b0e2d184" ID-ISSN="1179-3163" ID-Zenodo-Dep="6571439" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1653287346468" checkinUser="plazi" docAuthor="Smith, Gideon F. &amp; Figueiredo, Estrela" docDate="2022" docId="FF38BA54FF8F5C6AFF2213BFFD963E17" docLanguage="en" docName="phytotaxa.547.2.3.pdf" docOrigin="Phytotaxa 547 (2)" docStyle="DocumentStyle:96748F8F1B6C902996E134952A3A36B9.7:Phytotaxa.2014-.journal_article" docStyleId="96748F8F1B6C902996E134952A3A36B9" docStyleName="Phytotaxa.2014-.journal_article" docStyleVersion="7" docTitle="Kalanchoe beharensis Drake" docType="treatment" docVersion="7" lastPageNumber="163" masterDocId="0301C22CFF8D5C6FFFAA1635FFBC3D57" masterDocTitle="Nomenclatural and taxonomic notes on the names published in Kalanchoe (Crassulaceae subfam. Kalanchooideae) by Drake del Castillo in 1903" masterLastPageNumber="166" masterPageNumber="158" pageNumber="160" updateTime="1699337851642" updateUser="ExternalLinkService" zenodo-license-figures="UNSPECIFIED" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="B8FC2E184D5C36E84980BCB19B91BD04">Nomenclatural and taxonomic notes on the names published in Kalanchoe (Crassulaceae subfam. Kalanchooideae) by Drake del Castillo in 1903</mods:title>
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<mods:namePart id="2DC902F7F0EC1191FD01B406AB0537E5">Smith, Gideon F.</mods:namePart>
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<mods:affiliation id="AA6209C9AB9AF10BAA0B37B400C4BE83">Ria Olivier Herbarium, Department of Botany, Nelson Mandela University, P. O. Box 77000, Gqeberha, 6031 South Africa. &amp; smithgideon 1 @ gmail. com; https: // orcid. org / 0000 - 0002 - 5417 - 9208</mods:affiliation>
<mods:nameIdentifier id="5F9A7162297C20A507854DB2D5F33BA8" type="email">smithgideon1@gmail.com</mods:nameIdentifier>
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<mods:namePart id="89B8A640D8982EE3E586A216166C5ACB">Figueiredo, Estrela</mods:namePart>
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<mods:affiliation id="10AD40CB985C9356927CE9EA3C19E1D1">Ria Olivier Herbarium, Department of Botany, Nelson Mandela University, P. O. Box 77000, Gqeberha, 6031 South Africa. &amp; epnfigueiredo @ gmail. com; https: // orcid. org / 0000 - 0002 - 8511 - 8213</mods:affiliation>
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<mods:title id="51714015513B4413815F84BCB4941695">Phytotaxa</mods:title>
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<mods:date id="14B08E9C93BFFFEAB5A579D1B066BF23">2022</mods:date>
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<treatment id="FF38BA54FF8F5C6AFF2213BFFD963E17" ID-DOI="http://doi.org/10.5281/zenodo.6576563" ID-GBIF-Taxon="195582137" ID-Zenodo-Dep="6576563" LSID="urn:lsid:plazi:treatment:FF38BA54FF8F5C6AFF2213BFFD963E17" httpUri="http://treatment.plazi.org/id/FF38BA54FF8F5C6AFF2213BFFD963E17" lastPageId="5" lastPageNumber="163" pageId="2" pageNumber="160">
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<paragraph id="772E0B42FF8F5C6DFF2213BFFCF338F3" blockId="2.[136,847,1418,1445]" box="[136,847,1418,1445]" pageId="2" pageNumber="160">
<heading id="2C66BC2EFF8F5C6DFF2213BFFCF338F3" box="[136,847,1418,1445]" fontSize="11" level="2" pageId="2" pageNumber="160" reason="4">
2.
<taxonomicName id="B09170C1FF8F5C6DFF0E13BFFE5338F3" ID-CoL="3QYGG" authority="Drake" authorityName="Drake" box="[164,495,1418,1445]" class="Magnoliopsida" family="Crassulaceae" genus="Kalanchoe" kingdom="Plantae" order="Saxifragales" pageId="2" pageNumber="160" phylum="Tracheophyta" rank="species" species="beharensis">
<emphasis id="45E5D750FF8F5C6DFF0E13BFFE1838F2" bold="true" box="[164,420,1418,1445]" italics="true" pageId="2" pageNumber="160">Kalanchoe beharensis</emphasis>
Drake
</taxonomicName>
del
<bibRefCitation id="130076B3FF8F5C6DFD8A13BFFD4F38F3" author="Castillo, E." box="[544,755,1418,1444]" pageId="2" pageNumber="160" pagination="35 - 46" refId="ref5654" refString="Drake del Castillo, E. (1903) Note sur les plantes recueillies par M. Guillaume Grandidier dans le sud de Madagascar, en 1898 et 1901. (A suivre). Bulletin du Museum d'Histoire Naturelle 9: 35 - 46. [https: // www. biodiversitylibrary. org / page / 42893852 # page / 45 / mode / 1 up]" type="journal article" year="1903">Castillo (1903: 41)</bibRefCitation>
(
<figureCitation id="EFAA17C7FF8F5C6DFCA813BFFCFA38F3" box="[770,838,1418,1444]" captionStart="FIGURE 2" captionStartId="3.[136,229,1969,1991]" captionTargetBox="[164,1423,245,1944]" captionTargetId="figure-56@3.[164,1423,245,1944]" captionTargetPageId="3" captionText="FIGURE 2. Kalanchoe beharensis is one of the few truly arborescent species of Kalanchoe. Photograph: Gideon F. Smith." figureDoi="http://doi.org/10.5281/zenodo.6571443" httpUri="https://zenodo.org/record/6571443/files/figure.png" pageId="2" pageNumber="160">Fig. 2</figureCitation>
)
</heading>
</paragraph>
</subSubSection>
<subSubSection id="3F8B58C9FF8F5C6DFF2213E3FAB83BB0" pageId="2" pageNumber="160" type="materials_examined">
<paragraph id="772E0B42FF8F5C6DFF2213E3FB783B00" blockId="2.[136,1452,1493,1768]" pageId="2" pageNumber="160">
<emphasis id="45E5D750FF8F5C6DFF2213E3FF0638BC" bold="true" box="[136,186,1494,1515]" pageId="2" pageNumber="160">Type</emphasis>
:—
<materialsCitation id="C7F9011FFF8F5C6DFF7C13E3FAE538BB" ID-GBIF-Occurrence="3780763304" box="[214,1369,1493,1516]" collectionCode="G" country="Madagascar" location="Southeastern Madagascar. Anosy Region. Behara" pageId="2" pageNumber="160" specimenCount="1" typeStatus="lectotype">
<collectingCountry id="0F864BD2FF8F5C6DFF7C13E3FE3B38BC" box="[214,391,1494,1515]" name="Madagascar" pageId="2" pageNumber="160">MADAGASCAR</collectingCountry>
.
<location id="724E5D99FF8F5C6DFE3A13E3FCD738BB" LSID="urn:lsid:plazi:treatment:FF38BA54FF8F5C6AFF2213BFFD963E17:724E5D99FF8F5C6DFE3A13E3FCD738BB" box="[400,875,1494,1516]" country="Madagascar" name="Southeastern Madagascar. Anosy Region. Behara" pageId="2" pageNumber="160">Southeastern Madagascar. Anosy Region. Behara</location>
, 8 juillet [July] 1901,
<emphasis id="45E5D750FF8F5C6DFBEA13E3FB5738BB" box="[1088,1259,1494,1516]" italics="true" pageId="2" pageNumber="160">
<collectionCode id="11809387FF8F5C6DFBEA13E3FBE838BC" box="[1088,1108,1494,1515]" country="Switzerland" lsid="urn:lsid:biocol.org:col:15706" name="Conservatoire et Jardin botaniques de la Ville de Genève" pageId="2" pageNumber="160" type="Herbarium">G</collectionCode>
. Grandidier s.n
</emphasis>
. (
<typeStatus id="A82AB5E0FF8F5C6DFB5713E3FAE538BB" box="[1277,1369,1494,1516]" pageId="2" pageNumber="160" type="lectotype">lectotype</typeStatus>
</materialsCitation>
,
<materialsCitation id="C7F9011FFF8F5C6DFAC913E3FDB83B0F" ID-GBIF-Occurrence="3780763303" collectionCode="P" country="Madagascar" pageId="2" pageNumber="160" specimenCode="P00374194, P00374195" specimenCount="1" typeStatus="holotype">
Herb.
<collectionCode id="11809387FF8F5C6DFA3513E3FA1738BC" box="[1439,1451,1494,1515]" country="France" lsid="urn:lsid:biocol.org:col:15763" name="Museum National d' Histoire Naturelle, Paris (MNHN) - Vascular Plants" pageId="2" pageNumber="160" type="Herbarium">P</collectionCode>
barcode
<specimenCode id="2737A339FF8F5C6DFEA413CFFE3D3B58" box="[270,385,1530,1551]" collectionCode="P" country="France" lsid="urn:lsid:biocol.org:col:15763" name="Museum National d' Histoire Naturelle, Paris (MNHN) - Vascular Plants" pageId="2" pageNumber="160" type="Herbarium">P00374194</specimenCode>
! and Herb.
<collectionCode id="11809387FF8F5C6DFE4113CFFE4B3B58" box="[491,503,1530,1551]" country="France" lsid="urn:lsid:biocol.org:col:15763" name="Museum National d' Histoire Naturelle, Paris (MNHN) - Vascular Plants" pageId="2" pageNumber="160" type="Herbarium">P</collectionCode>
barcode
<specimenCode id="2737A339FF8F5C6DFDE713CFFD7C3B47" box="[589,704,1530,1552]" collectionCode="P" country="France" lsid="urn:lsid:biocol.org:col:15763" name="Museum National d' Histoire Naturelle, Paris (MNHN) - Vascular Plants" pageId="2" pageNumber="160" type="Herbarium">P00374195</specimenCode>
! [Images available at: http://coldb.mnhn.fr/catalognumber/mnhn/p/p00374194 (inflorescence) and http://coldb.mnhn.fr/catalognumber/mnhn/p/p00374195 (leaves in envelopes and inflorescence)]; consisting of two sheets interpreted as
<typeStatus id="A82AB5E0FF8F5C6DFE1A1077FDB83B0F" box="[432,516,1602,1624]" pageId="2" pageNumber="160" type="holotype">holotype</typeStatus>
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by
<bibRefCitation id="130076B3FF8F5C6DFD821077FC183B0F" author="Boiteau, P. &amp; Allorge-Boiteau, L." box="[552,932,1602,1624]" pageId="2" pageNumber="160" refId="ref5454" refString="Boiteau, P. &amp; Allorge-Boiteau, L. (1995) Kalanchoe (Crassulacees) de Madagascar. Systematique, ecophysiologie et phytochimie. ICSN, CNRS, 91198 Gif-sur-Yvette, Editions Karthala, Paris, 252 pp." type="book" year="1995">Boiteau &amp; Allorge-Boiteau (1995: 154)</bibRefCitation>
,
<emphasis id="45E5D750FF8F5C6DFC041077FB023B00" bold="true" box="[942,1214,1602,1623]" pageId="2" pageNumber="160">
here corrected to
<typeStatus id="A82AB5E0FF8F5C6DFBC81077FB023B00" box="[1122,1214,1602,1623]" pageId="2" pageNumber="160" type="lectotype">lectotype</typeStatus>
</emphasis>
.
</paragraph>
<paragraph id="772E0B42FF8F5C6DFF221053FAB83BB0" blockId="2.[136,1452,1493,1768]" pageId="2" pageNumber="160">
<emphasis id="45E5D750FF8F5C6DFF221053FEDE3B2C" bold="true" box="[136,354,1638,1660]" pageId="2" pageNumber="160">Homotypic synonym</emphasis>
:—
<taxonomicName id="B09170C1FF8F5C6DFE2B1053FC803B2C" authority="Hamet (1906: 110" authorityName="Hamet" authorityYear="1906" box="[385,828,1638,1660]" class="Magnoliopsida" family="Crassulaceae" genus="Kalanchoe" kingdom="Plantae" order="Saxifragales" pageId="2" pageNumber="160" phylum="Tracheophyta" rank="species" species="vantieghemii">
<emphasis id="45E5D750FF8F5C6DFE2B1053FDCE3B2C" box="[385,626,1638,1659]" italics="true" pageId="2" pageNumber="160">Kalanchoe vantieghemii</emphasis>
<bibRefCitation id="130076B3FF8F5C6DFDD41053FC803B2C" author="Hamet, R." box="[638,828,1638,1660]" pageId="2" pageNumber="160" pagination="109 - 111" refId="ref5772" refString="Hamet, R. (1906 [Janvier - Mars]) Note sur une nouvelle espece de Kalanchoe [Kalanchoe vantieghemii, as ' Van Tieghemi ']. Journal de Botanique [Morot] XX [20 e [annee] (No 1 - 3)]: 109 - 111. [https: // www. biodiversitylibrary. org / item / 22106 # page / 113 / mode / 1 up]" type="book chapter" year="1906">Hamet (1906: 110</bibRefCitation>
</taxonomicName>
, as
<taxonomicName id="B09170C1FF8F5C6DFCDF1052FC423B2C" authority="Tieghemi" authorityName="Tieghemi" box="[885,1022,1638,1660]" class="Pennatae" family="Surirellaceae" genus="Van" higherTaxonomySource="GBIF" kingdom="Chromista" order="Surirellales" pageId="2" pageNumber="160" phylum="Diatomeae" rank="genus">
<emphasis id="45E5D750FF8F5C6DFCDF1052FC423B2C" box="[885,1022,1638,1660]" italics="true" pageId="2" pageNumber="160">Van Tieghemi</emphasis>
</taxonomicName>
).
<emphasis id="45E5D750FF8F5C6DFB8B1053FBEF3B2C" bold="true" box="[1057,1107,1638,1659]" pageId="2" pageNumber="160">
<location id="724E5D99FF8F5C6DFB8B1053FBEF3B2C" LSID="urn:lsid:plazi:treatment:FF38BA54FF8F5C6AFF2213BFFD963E17:724E5D99FF8F5C6DFB8B1053FBEF3B2C" box="[1057,1107,1638,1659]" country="Madagascar" county="Grandidier" municipality="Southeastern Madagascar. Anosy Region. Behara" name="Type" pageId="2" pageNumber="160">Type</location>
</emphasis>
:—
<collectingCountry id="0F864BD2FF8F5C6DFBC51053FA9C3B2C" box="[1135,1312,1638,1659]" name="Madagascar" pageId="2" pageNumber="160">MADAGASCAR</collectingCountry>
.
<collectingMunicipality id="974A9138FF8F5C6DFA851053FDA63BF7" pageId="2" pageNumber="160">Southeastern Madagascar. Anosy Region. Behara</collectingMunicipality>
, 8 juillet [July] 1901,
<emphasis id="45E5D750FF8F5C6DFD5810BFFC233BF7" box="[754,927,1674,1696]" italics="true" pageId="2" pageNumber="160">
<collectionCode id="11809387FF8F5C6DFD5810BFFCBA3BC8" box="[754,774,1674,1695]" country="Switzerland" lsid="urn:lsid:biocol.org:col:15706" name="Conservatoire et Jardin botaniques de la Ville de Genève" pageId="2" pageNumber="160" type="Herbarium">G</collectionCode>
.
<collectingCounty id="9E4F73CEFF8F5C6DFCBB10BFFCC13BC8" box="[785,893,1674,1695]" pageId="2" pageNumber="160">Grandidier</collectingCounty>
s.n
</emphasis>
. (
<typeStatus id="A82AB5E0FF8F5C6DFC1810BFFBB23BF7" box="[946,1038,1674,1696]" pageId="2" pageNumber="160" type="lectotype">lectotype</typeStatus>
, Herb.
<collectionCode id="11809387FF8F5C6DFBFC10BFFBDE3BC8" box="[1110,1122,1674,1695]" country="France" lsid="urn:lsid:biocol.org:col:15763" name="Museum National d' Histoire Naturelle, Paris (MNHN) - Vascular Plants" pageId="2" pageNumber="160" type="Herbarium">P</collectionCode>
barcode
<specimenCode id="2737A339FF8F5C6DFB1610BFFA933BC8" box="[1212,1327,1674,1695]" collectionCode="P" country="France" lsid="urn:lsid:biocol.org:col:15763" name="Museum National d' Histoire Naturelle, Paris (MNHN) - Vascular Plants" pageId="2" pageNumber="160" type="Herbarium">P00374194</specimenCode>
! and Herb.
<collectionCode id="11809387FF8F5C6DFA3510BFFA173BC8" box="[1439,1451,1674,1695]" country="France" lsid="urn:lsid:biocol.org:col:15763" name="Museum National d' Histoire Naturelle, Paris (MNHN) - Vascular Plants" pageId="2" pageNumber="160" type="Herbarium">P</collectionCode>
barcode
<specimenCode id="2737A339FF8F5C6DFEA5109BFE3E3B93" box="[271,386,1710,1732]" collectionCode="P" country="France" lsid="urn:lsid:biocol.org:col:15763" name="Museum National d' Histoire Naturelle, Paris (MNHN) - Vascular Plants" pageId="2" pageNumber="160" type="Herbarium">P00374195</specimenCode>
! [Images available at: http://coldb.mnhn.fr/catalognumber/mnhn/p/p00374194 (inflorescence) and http://coldb. mnhn.fr/catalognumber/mnhn/p/p00374195 (leaves in envelopes and inflorescence)];
<emphasis id="45E5D750FF8F5C6DFC5C10E7FB423BBF" bold="true" box="[1014,1278,1746,1768]" pageId="2" pageNumber="160">
<typeStatus id="A82AB5E0FF8F5C6DFC5C10E7FBEF3BB0" box="[1014,1107,1746,1767]" pageId="2" pageNumber="160" type="lectotype">lectotype</typeStatus>
, here designated
</emphasis>
.
</paragraph>
</subSubSection>
<subSubSection id="3F8B58C9FF8F5C6BFF221123FAEC38F2" lastPageId="4" lastPageNumber="162" pageId="2" pageNumber="160" type="discussion">
<paragraph id="772E0B42FF8F5C6DFF221123FCDA3A67" blockId="2.[136,1452,1814,2093]" box="[136,870,1814,1841]" pageId="2" pageNumber="160">
<emphasis id="45E5D750FF8F5C6DFF221123FEF93A67" bold="true" box="[136,325,1814,1840]" pageId="2" pageNumber="160">Taxonomic note</emphasis>
:—
<taxonomicName id="B09170C1FF8F5C6DFEC31123FDDE3A67" box="[361,610,1814,1840]" class="Magnoliopsida" family="Crassulaceae" genus="Kalanchoe" kingdom="Plantae" order="Saxifragales" pageId="2" pageNumber="160" phylum="Tracheophyta" rank="species" species="beharensis">
<emphasis id="45E5D750FF8F5C6DFEC31123FDDE3A67" box="[361,610,1814,1840]" italics="true" pageId="2" pageNumber="160">Kalanchoe beharensis</emphasis>
</taxonomicName>
is an accepted species.
</paragraph>
<paragraph id="772E0B42FF8F5C6CFF17110FFB6B3D8B" blockId="2.[136,1452,1814,2093]" lastBlockId="3.[136,1451,158,221]" lastPageId="3" lastPageNumber="161" pageId="2" pageNumber="160">
<emphasis id="45E5D750FF8F5C6DFF17110FFE113A03" bold="true" box="[189,429,1850,1876]" pageId="2" pageNumber="160">Nomenclature notes</emphasis>
:—The only material Drake del
<bibRefCitation id="130076B3FF8F5C6DFCB0110FFC4C3A03" author="Castillo, E." box="[794,1008,1850,1876]" pageId="2" pageNumber="160" pagination="35 - 46" refId="ref5654" refString="Drake del Castillo, E. (1903) Note sur les plantes recueillies par M. Guillaume Grandidier dans le sud de Madagascar, en 1898 et 1901. (A suivre). Bulletin du Museum d'Histoire Naturelle 9: 35 - 46. [https: // www. biodiversitylibrary. org / page / 42893852 # page / 45 / mode / 1 up]" type="journal article" year="1903">Castillo (1903: 41)</bibRefCitation>
cited in the protologue of the name
<taxonomicName id="B09170C1FF8F5C6DFA38110EFEBC3A2F" class="Magnoliopsida" family="Crassulaceae" genus="Kalanchoe" kingdom="Plantae" order="Saxifragales" pageId="2" pageNumber="160" phylum="Tracheophyta" rank="species" species="beharensis">
<emphasis id="45E5D750FF8F5C6DFA38110EFEBC3A2F" italics="true" pageId="2" pageNumber="160">K. beharensis</emphasis>
</taxonomicName>
, “Behara (
<date id="032F2D82FF8F5C6DFEDD116BFDB23A2F" box="[375,526,1886,1912]" pageId="2" pageNumber="160" value="1901-07-08">8 juillet 1901</date>
)”, was a collection without a number made by Guillaume Grandidier on
<date id="032F2D82FF8F5C6DFA86116BFA173A2F" box="[1324,1451,1886,1912]" pageId="2" pageNumber="160" value="1901-07-08">
8 July
<quantity id="B069A6A7FF8F5C6DFAD9116BFA173A2F" box="[1395,1451,1886,1912]" metricMagnitude="1" metricUnit="m" metricValue="4.828539999999999" pageId="2" pageNumber="160" unit="in" value="1901.0">1901</quantity>
</date>
in southeastern
<collectingCountry id="0F864BD2FF8F5C6DFE9C11B7FE033ACB" box="[310,447,1922,1948]" name="Madagascar" pageId="2" pageNumber="160">Madagascar</collectingCountry>
. Further, when publishing the name
<taxonomicName id="B09170C1FF8F5C6DFCFB11B6FB863ACB" authority=", Drake" authorityName="Drake" box="[849,1082,1922,1949]" class="Magnoliopsida" family="Crassulaceae" genus="Kalanchoe" kingdom="Plantae" order="Saxifragales" pageId="2" pageNumber="160" phylum="Tracheophyta" rank="species" species="beharensis">
<emphasis id="45E5D750FF8F5C6DFCFB11B6FC543ACB" box="[849,1000,1922,1948]" italics="true" pageId="2" pageNumber="160">K. beharensis</emphasis>
, Drake
</taxonomicName>
del
<bibRefCitation id="130076B3FF8F5C6DFBC211B7FA863ACB" author="Castillo, E." box="[1128,1338,1922,1948]" pageId="2" pageNumber="160" pagination="35 - 46" refId="ref5654" refString="Drake del Castillo, E. (1903) Note sur les plantes recueillies par M. Guillaume Grandidier dans le sud de Madagascar, en 1898 et 1901. (A suivre). Bulletin du Museum d'Histoire Naturelle 9: 35 - 46. [https: // www. biodiversitylibrary. org / page / 42893852 # page / 45 / mode / 1 up]" type="journal article" year="1903">Castillo (1903: 41)</bibRefCitation>
described the leaves as “carnosis magnis (1020 centim.) late ovatis lanceolatis glabris” [English: “fleshy, large (
<quantity id="B069A6A7FF8F5C6DFA821193FA233A96" box="[1320,1439,1958,1985]" metricMagnitude="-1" metricUnit="m" metricValue="1.5" metricValueMax="2.0" metricValueMin="1.0" pageId="2" pageNumber="160" unit="cm" value="15.0" valueMax="20.0" valueMin="10.0">1020 cm</quantity>
), broadly ovate, lanceolate, glabrous”]. The leaves being described as both “broadly ovate” and as “lanceolate” seem to present a morphological conflict (further discussed below). Critically, although horticultural selections of the species do have glabrous leaves, for example cultivated material known under the designation
<taxonomicName id="B09170C1FF8F5C6DFBDA1E26FA18357B" box="[1136,1444,2066,2093]" class="Magnoliopsida" family="Crassulaceae" genus="Kalanchoe" kingdom="Plantae" order="Saxifragales" pageId="2" pageNumber="160" phylum="Tracheophyta" rank="variety" species="beharensis" variety="subnuda">
<emphasis id="45E5D750FF8F5C6DFBDA1E26FAB6357B" box="[1136,1290,2066,2092]" italics="true" pageId="2" pageNumber="160">K. beharensis</emphasis>
var.
<emphasis id="45E5D750FF8F5C6DFAEF1E27FA18357B" box="[1349,1444,2066,2092]" italics="true" pageId="2" pageNumber="160">subnuda</emphasis>
</taxonomicName>
or under the monikers
<emphasis id="45E5D750FF8E5C6CFE2F16AAFE243DEF" box="[389,408,159,184]" italics="true" pageId="3" pageNumber="161">K</emphasis>
. Subnuda and
<emphasis id="45E5D750FF8E5C6CFDE516AAFDDE3DEF" box="[591,610,159,184]" italics="true" pageId="3" pageNumber="161">K</emphasis>
. Babys Bottom, the leaves of
<taxonomicName id="B09170C1FF8E5C6CFC6716AAFBDA3DEF" box="[973,1126,158,184]" class="Magnoliopsida" family="Crassulaceae" genus="Kalanchoe" kingdom="Plantae" order="Saxifragales" pageId="3" pageNumber="161" phylum="Tracheophyta" rank="species" species="beharensis">
<emphasis id="45E5D750FF8E5C6CFC6716AAFBDA3DEF" box="[973,1126,158,184]" italics="true" pageId="3" pageNumber="161">K. beharensis</emphasis>
</taxonomicName>
are most often variously and very densely tomentose, not glabrous, giving them a distinctly felt-like texture (
<bibRefCitation id="130076B3FF8E5C6CFBAE16F7FB763D8B" author="Smith, G. F. &amp; Figueiredo, E. &amp; Walker, C. C." box="[1028,1226,194,221]" pageId="3" pageNumber="161" pagination="50 - 59" refId="ref6831" refString="Smith, G. F., Figueiredo, E. &amp; Walker, C. C. (2021) Kalanchoe beharensis. Crassulaceae subfam. Kalanchooideae. Madagascar. Flowering Plants of Africa 67: 50 - 59, plate 2365." type="journal article" year="2021">
Smith
<emphasis id="45E5D750FF8E5C6CFBE516F6FB3D3D8B" box="[1103,1153,194,220]" italics="true" pageId="3" pageNumber="161">et al</emphasis>
. 2021
</bibRefCitation>
).
</paragraph>
<caption id="23EE5BCAFF8E5C6CFF221184FA9A3A90" ID-DOI="http://doi.org/10.5281/zenodo.6571443" ID-Zenodo-Dep="6571443" box="[136,1318,1969,1991]" httpUri="https://zenodo.org/record/6571443/files/figure.png" pageId="3" pageNumber="161" startId="3.[136,229,1969,1991]" targetBox="[164,1423,245,1944]" targetPageId="3">
<paragraph id="772E0B42FF8E5C6CFF221184FA9A3A90" blockId="3.[136,1318,1969,1991]" box="[136,1318,1969,1991]" pageId="3" pageNumber="161">
<emphasis id="45E5D750FF8E5C6CFF221184FF423A91" bold="true" box="[136,254,1969,1991]" pageId="3" pageNumber="161">FIGURE 2.</emphasis>
<taxonomicName id="B09170C1FF8E5C6CFEA91184FE643A91" box="[259,472,1969,1990]" class="Magnoliopsida" family="Crassulaceae" genus="Kalanchoe" kingdom="Plantae" order="Saxifragales" pageId="3" pageNumber="161" phylum="Tracheophyta" rank="species" species="beharensis">
<emphasis id="45E5D750FF8E5C6CFEA91184FE643A91" box="[259,472,1969,1990]" italics="true" pageId="3" pageNumber="161">Kalanchoe beharensis</emphasis>
</taxonomicName>
is one of the few truly arborescent species of
<taxonomicName id="B09170C1FF8E5C6CFC381184FC463A91" authorityName="Adanson" authorityYear="1763" box="[914,1018,1969,1990]" class="Magnoliopsida" family="Crassulaceae" genus="Kalanchoe" kingdom="Plantae" order="Saxifragales" pageId="3" pageNumber="161" phylum="Tracheophyta" rank="genus">
<emphasis id="45E5D750FF8E5C6CFC381184FC463A91" box="[914,1018,1969,1990]" italics="true" pageId="3" pageNumber="161">Kalanchoe</emphasis>
</taxonomicName>
. Photograph: Gideon F. Smith.
</paragraph>
</caption>
<paragraph id="772E0B42FF895C6BFF1716ABFC8E3CE3" blockId="4.[136,1452,158,2057]" pageId="4" pageNumber="162">
Then, three years after
<taxonomicName id="B09170C1FF895C6BFE1016AAFDED3DEF" box="[442,593,158,184]" class="Magnoliopsida" family="Crassulaceae" genus="Kalanchoe" kingdom="Plantae" order="Saxifragales" pageId="4" pageNumber="162" phylum="Tracheophyta" rank="species" species="beharensis">
<emphasis id="45E5D750FF895C6BFE1016AAFDED3DEF" box="[442,593,158,184]" italics="true" pageId="4" pageNumber="162">K. beharensis</emphasis>
</taxonomicName>
was described by Drake del
<bibRefCitation id="130076B3FF895C6BFC2116ABFBE23DEF" author="Castillo, E." box="[907,1118,158,184]" pageId="4" pageNumber="162" pagination="35 - 46" refId="ref5654" refString="Drake del Castillo, E. (1903) Note sur les plantes recueillies par M. Guillaume Grandidier dans le sud de Madagascar, en 1898 et 1901. (A suivre). Bulletin du Museum d'Histoire Naturelle 9: 35 - 46. [https: // www. biodiversitylibrary. org / page / 42893852 # page / 45 / mode / 1 up]" type="journal article" year="1903">Castillo (1903: 41)</bibRefCitation>
,
<bibRefCitation id="130076B3FF895C6BFBC216AAFA853DEF" author="Hamet, R." box="[1128,1337,158,185]" pageId="4" pageNumber="162" pagination="109 - 111" refId="ref5772" refString="Hamet, R. (1906 [Janvier - Mars]) Note sur une nouvelle espece de Kalanchoe [Kalanchoe vantieghemii, as ' Van Tieghemi ']. Journal de Botanique [Morot] XX [20 e [annee] (No 1 - 3)]: 109 - 111. [https: // www. biodiversitylibrary. org / item / 22106 # page / 113 / mode / 1 up]" type="book chapter" year="1906">Hamet (1906: 110)</bibRefCitation>
published the name
<taxonomicName id="B09170C1FF895C6BFF5F16F6FE1B3D8B" authorityName="Hamet" authorityYear="1906" box="[245,423,194,220]" class="Magnoliopsida" family="Crassulaceae" genus="Kalanchoe" kingdom="Plantae" order="Saxifragales" pageId="4" pageNumber="162" phylum="Tracheophyta" rank="species" species="vantieghemii">
<emphasis id="45E5D750FF895C6BFF5F16F6FE1B3D8B" box="[245,423,194,220]" italics="true" pageId="4" pageNumber="162">K. vantieghemii</emphasis>
</taxonomicName>
, as
<taxonomicName id="B09170C1FF895C6BFE7716F6FDCA3D8B" authority="Tieghemi" authorityName="Tieghemi" box="[477,630,194,220]" class="Pennatae" family="Surirellaceae" genus="Van" higherTaxonomySource="GBIF" kingdom="Chromista" order="Surirellales" pageId="4" pageNumber="162" phylum="Diatomeae" rank="genus">
<emphasis id="45E5D750FF895C6BFE7716F6FDCA3D8B" box="[477,630,194,220]" italics="true" pageId="4" pageNumber="162">Van Tieghemi</emphasis>
</taxonomicName>
, based on “
<collectingCountry id="0F864BD2FF895C6BFCAB16F7FC333D8B" box="[769,911,194,220]" name="Madagascar" pageId="4" pageNumber="162">Madagascar</collectingCountry>
: Behara, 6 à [to]
<date id="032F2D82FF895C6BFBFA16F7FB5A3D8B" box="[1104,1254,194,220]" pageId="4" pageNumber="162" value="1901-07-08">8 juillet 1901</date>
(Grandidier fils). Dans lherbier Drake del Castillo.” This was, presumably, at least partly, the same material that Drake del Castillo cited in the protologue of the name
<taxonomicName id="B09170C1FF895C6BFDBF173EFD113C73" box="[533,685,266,292]" class="Magnoliopsida" family="Crassulaceae" genus="Kalanchoe" kingdom="Plantae" order="Saxifragales" pageId="4" pageNumber="162" phylum="Tracheophyta" rank="species" species="beharensis">
<emphasis id="45E5D750FF895C6BFDBF173EFD113C73" box="[533,685,266,292]" italics="true" pageId="4" pageNumber="162">K. beharensis</emphasis>
</taxonomicName>
, but with the date in
<date id="032F2D82FF895C6BFC33173FFBBB3C73" box="[921,1031,266,292]" pageId="4" pageNumber="162" value="1901-07">July 1901</date>
when Grandidier made the collection extended to “6 à [to] 8”, not only “8”. He might therefore have had material collected on 6 and 7 as well as on
<date id="032F2D82FF895C6BFACD171BFF7C3C3B" pageId="4" pageNumber="162" value="1901-07-08">8 July 1901</date>
at his disposal. The large herbarium of Drake del Castillo was bequeathed to the
<emphasis id="45E5D750FF895C6BFBC11766FF503CC7" italics="true" pageId="4" pageNumber="162">Muséum national dHistoire naturelle</emphasis>
after his death in 1904. Hamet, who was 16 years old when he described
<taxonomicName id="B09170C1FF895C6BFBB51742FB6C3CC7" authorityName="Hamet" authorityYear="1906" box="[1055,1232,374,400]" class="Magnoliopsida" family="Crassulaceae" genus="Kalanchoe" kingdom="Plantae" order="Saxifragales" pageId="4" pageNumber="162" phylum="Tracheophyta" rank="species" species="vantieghemii">
<emphasis id="45E5D750FF895C6BFBB51742FB6C3CC7" box="[1055,1232,374,400]" italics="true" pageId="4" pageNumber="162">K. vantieghemii</emphasis>
</taxonomicName>
, was to become the global expert on
<taxonomicName id="B09170C1FF895C6BFEEF17AFFE023CE3" authorityName="Adanson" authorityYear="1763" box="[325,446,410,436]" class="Magnoliopsida" family="Crassulaceae" genus="Kalanchoe" kingdom="Plantae" order="Saxifragales" pageId="4" pageNumber="162" phylum="Tracheophyta" rank="genus">
<emphasis id="45E5D750FF895C6BFEEF17AFFE023CE3" box="[325,446,410,436]" italics="true" pageId="4" pageNumber="162">Kalanchoe</emphasis>
</taxonomicName>
in the 20
<superScript id="80E4A60AFF895C6BFD8D17AFFD883CFF" attach="left" box="[551,564,410,424]" fontSize="6" pageId="4" pageNumber="162">th</superScript>
century (
<bibRefCitation id="130076B3FF895C6BFD3417AFFC9A3CE3" author="Smith, G. F." box="[670,806,410,436]" pageId="4" pageNumber="162" pagination="120 - 140" refId="ref6732" refString="Smith, G. F. (2020) The kalanchoes (Crassulaceae subfam. Kalanchooideae) of Raymond-Hamet (1890 - 1972). Bradleya 38: 120 - 140. https: // doi. org / 10.25223 / brad. n 38.2020. a 15" type="journal article" year="2020">Smith 2020</bibRefCitation>
).
</paragraph>
<paragraph id="772E0B42FF895C6BFF17178AFAE639FF" blockId="4.[136,1452,158,2057]" pageId="4" pageNumber="162">
In contrast to Drake del Castillos statement that the leaves of
<taxonomicName id="B09170C1FF895C6BFC22178AFB9E3C8F" box="[904,1058,446,472]" class="Magnoliopsida" family="Crassulaceae" genus="Kalanchoe" kingdom="Plantae" order="Saxifragales" pageId="4" pageNumber="162" phylum="Tracheophyta" rank="species" species="beharensis">
<emphasis id="45E5D750FF895C6BFC22178AFB9E3C8F" box="[904,1058,446,472]" italics="true" pageId="4" pageNumber="162">K. beharensis</emphasis>
</taxonomicName>
were glabrous (Drake del
<bibRefCitation id="130076B3FF895C6BFAFE178BFF483CAB" author="Castillo, E." pageId="4" pageNumber="162" pagination="35 - 46" refId="ref5654" refString="Drake del Castillo, E. (1903) Note sur les plantes recueillies par M. Guillaume Grandidier dans le sud de Madagascar, en 1898 et 1901. (A suivre). Bulletin du Museum d'Histoire Naturelle 9: 35 - 46. [https: // www. biodiversitylibrary. org / page / 42893852 # page / 45 / mode / 1 up]" type="journal article" year="1903">Castillo 1903: 41</bibRefCitation>
),
<bibRefCitation id="130076B3FF895C6BFEA417D6FE4C3CAB" author="Hamet, R." box="[270,496,482,509]" pageId="4" pageNumber="162" pagination="109 - 111" refId="ref5772" refString="Hamet, R. (1906 [Janvier - Mars]) Note sur une nouvelle espece de Kalanchoe [Kalanchoe vantieghemii, as ' Van Tieghemi ']. Journal de Botanique [Morot] XX [20 e [annee] (No 1 - 3)]: 109 - 111. [https: // www. biodiversitylibrary. org / item / 22106 # page / 113 / mode / 1 up]" type="book chapter" year="1906">Hamet (1906: 109)</bibRefCitation>
noted that: “Cette espèce [
<taxonomicName id="B09170C1FF895C6BFC9717D6FC493CAB" authorityName="Hamet" authorityYear="1906" box="[829,1013,482,508]" class="Magnoliopsida" family="Crassulaceae" genus="Kalanchoe" kingdom="Plantae" order="Saxifragales" pageId="4" pageNumber="162" phylum="Tracheophyta" rank="species" species="vantieghemii">
<emphasis id="45E5D750FF895C6BFC9717D6FC493CAB" box="[829,1013,482,508]" italics="true" pageId="4" pageNumber="162">K. vantieghemii</emphasis>
</taxonomicName>
] appartient au groupe malgache fort remarquable constitué par le
<taxonomicName id="B09170C1FF895C6BFE611433FD023F77" box="[459,702,518,544]" class="Magnoliopsida" family="Crassulaceae" genus="Kalanchoe" kingdom="Plantae" order="Saxifragales" pageId="4" pageNumber="162" phylum="Tracheophyta" rank="species" species="eriophylla">
<emphasis id="45E5D750FF895C6BFE611433FD023F77" box="[459,702,518,544]" italics="true" pageId="4" pageNumber="162">Kalanchoe eriophylla</emphasis>
</taxonomicName>
et le
<taxonomicName id="B09170C1FF895C6BFD501432FC313F77" box="[762,909,519,544]" class="Magnoliopsida" family="Crassulaceae" genus="Kalanchoe" kingdom="Plantae" order="Saxifragales" pageId="4" pageNumber="162" phylum="Tracheophyta" rank="species" species="tomentosa">
<emphasis id="45E5D750FF895C6BFD501432FC313F77" box="[762,909,519,544]" italics="true" pageId="4" pageNumber="162">K. tomentosa</emphasis>
</taxonomicName>
groupe caractérisé par la présence, sur toutes les parties des espèces qui le composent, de poils très serrés formant une sorte de velours.” [English: “This species [
<taxonomicName id="B09170C1FF895C6BFA38141EFEA53F3F" authorityName="Hamet" authorityYear="1906" class="Magnoliopsida" family="Crassulaceae" genus="Kalanchoe" kingdom="Plantae" order="Saxifragales" pageId="4" pageNumber="162" phylum="Tracheophyta" rank="species" species="vantieghemii">
<emphasis id="45E5D750FF895C6BFA38141EFEA53F3F" italics="true" pageId="4" pageNumber="162">K. vantieghemii</emphasis>
</taxonomicName>
] belongs to the very remarkable Malagasy group, comprising the
<taxonomicName id="B09170C1FF895C6BFC58147BFB583F3F" box="[1010,1252,590,616]" class="Magnoliopsida" family="Crassulaceae" genus="Kalanchoe" kingdom="Plantae" order="Saxifragales" pageId="4" pageNumber="162" phylum="Tracheophyta" rank="species" species="eriophylla">
<emphasis id="45E5D750FF895C6BFC58147BFB583F3F" box="[1010,1252,590,616]" italics="true" pageId="4" pageNumber="162">Kalanchoe eriophylla</emphasis>
</taxonomicName>
and
<taxonomicName id="B09170C1FF895C6BFAB3147AFA173F3F" box="[1305,1451,591,616]" class="Magnoliopsida" family="Crassulaceae" genus="Kalanchoe" kingdom="Plantae" order="Saxifragales" pageId="4" pageNumber="162" phylum="Tracheophyta" rank="species" species="tomentosa">
<emphasis id="45E5D750FF895C6BFAB3147AFA173F3F" box="[1305,1451,591,616]" italics="true" pageId="4" pageNumber="162">K. tomentosa</emphasis>
</taxonomicName>
group, which is characterised by the presence, on all the plant parts of the species, of very dense hairs forming a sort of velvet.”] Regarding the leaves he though stated in the Latin description of
<taxonomicName id="B09170C1FF895C6BFC5D14A2FB153FE7" authorityName="Hamet" authorityYear="1906" box="[1015,1193,662,688]" class="Magnoliopsida" family="Crassulaceae" genus="Kalanchoe" kingdom="Plantae" order="Saxifragales" pageId="4" pageNumber="162" phylum="Tracheophyta" rank="species" species="vantieghemii">
<emphasis id="45E5D750FF895C6BFC5D14A2FB153FE7" box="[1015,1193,662,688]" italics="true" pageId="4" pageNumber="162">K. vantieghemii</emphasis>
</taxonomicName>
that they are “ignota”, i.e., “unknown”. Two years later,
<bibRefCitation id="130076B3FF895C6BFDA9148EFD643F83" author="Hamet, R." box="[515,728,698,725]" pageId="4" pageNumber="162" pagination="17 - 48" refId="ref5980" refString="Hamet, R. (1908 a) Monographie du genre Kalanchoe. Suite et fin. Bulletin de l'Herbier Boissier, ser. 2, 8: 17 - 48. [https: // www. biodiversitylibrary. org / item / 104945 # page / 31 / mode / 1 up]" type="journal article" year="1908">Hamet (1908a: 29)</bibRefCitation>
explained that he did not describe the leaves of
<taxonomicName id="B09170C1FF895C6BFB53148EFA103F83" authorityName="Hamet" authorityYear="1906" box="[1273,1452,698,724]" class="Magnoliopsida" family="Crassulaceae" genus="Kalanchoe" kingdom="Plantae" order="Saxifragales" pageId="4" pageNumber="162" phylum="Tracheophyta" rank="species" species="vantieghemii">
<emphasis id="45E5D750FF895C6BFB53148EFA103F83" box="[1273,1452,698,724]" italics="true" pageId="4" pageNumber="162">K. vantieghemii</emphasis>
</taxonomicName>
in 1906, because even though Drake del
<bibRefCitation id="130076B3FF895C6BFDE414EBFD4A3FAF" author="Castillo, E." box="[590,758,734,760]" pageId="4" pageNumber="162" pagination="35 - 46" refId="ref5654" refString="Drake del Castillo, E. (1903) Note sur les plantes recueillies par M. Guillaume Grandidier dans le sud de Madagascar, en 1898 et 1901. (A suivre). Bulletin du Museum d'Histoire Naturelle 9: 35 - 46. [https: // www. biodiversitylibrary. org / page / 42893852 # page / 45 / mode / 1 up]" type="journal article" year="1903">Castillo (1903)</bibRefCitation>
described
<taxonomicName id="B09170C1FF895C6BFCC514EAFBB43FAF" box="[879,1032,734,760]" class="Magnoliopsida" family="Crassulaceae" genus="Kalanchoe" kingdom="Plantae" order="Saxifragales" pageId="4" pageNumber="162" phylum="Tracheophyta" rank="species" species="beharensis">
<emphasis id="45E5D750FF895C6BFCC514EAFBB43FAF" box="[879,1032,734,760]" italics="true" pageId="4" pageNumber="162">K. beharensis</emphasis>
</taxonomicName>
as having “broadly ovate, lanceolate, glabrous” leaves, the leaf fragments that were part of the Grandidiers unnumbered collection from Behara appeared to have a different shape and there were doubts if they belonged to the same material as that from which the inflorescence and flowers were obtained and described. This, then, was the reason put forward by Hamet as to why he opted to not consider the leaf material in his description. However, a further two years later
<bibRefCitation id="130076B3FF895C6BFB9C155AFAA53EDF" author="Hamet, R." box="[1078,1305,878,905]" pageId="4" pageNumber="162" pagination="191 - 194" refId="ref6223" refString="Hamet, R. (1910 c) Kalanchoe aliciae sp. nova et K. beharensis Drake del Castillo. Bulletin de la Societe Botanique de France 57: 191 - 194. [https: // www. biodiversitylibrary. org / item / 8679 # page / 241 / mode / 1 up]" type="journal article" year="1910">Hamet (1910c: 194)</bibRefCitation>
revisited the material and concluded that the leaf remains attached to
<emphasis id="45E5D750FF895C6BFCA715A7FC0F3EFB" box="[781,947,914,940]" italics="true" pageId="4" pageNumber="162">Grandidier s.n</emphasis>
. “Behara (
<date id="032F2D82FF895C6BFB9A15A7FB773EFB" box="[1072,1227,914,940]" pageId="4" pageNumber="162" value="1901-07-08">8 juillet 1901</date>
)” belonged to three taxa (an admixture, therefore), with these being: (1)
<taxonomicName id="B09170C1FF895C6BFD761582FB3D3E87" authority="Scott Elliot (1891: 14)" authorityName="Scott Elliot" authorityPageNumber="14" authorityYear="1891" box="[732,1153,950,976]" class="Magnoliopsida" family="Crassulaceae" genus="Kalanchoe" kingdom="Plantae" order="Saxifragales" pageId="4" pageNumber="162" phylum="Tracheophyta" rank="species" species="verticillata">
<emphasis id="45E5D750FF895C6BFD761582FCC53E87" box="[732,889,950,976]" italics="true" pageId="4" pageNumber="162">K. verticillata</emphasis>
<bibRefCitation id="130076B3FF895C6BFC2B1583FB3D3E87" author="Scott Elliot, G. F." box="[897,1153,950,976]" pageId="4" pageNumber="162" refId="ref6639" refString="Scott Elliot, G. F. (1891 [22 August]) New and little known Madagascar plants, collected and enumerated by G. F. Scott Elliot, M. A., B. Sc. F. L. S. The Journal of the Linnean Society, Botany XXIX (197): 1 - 67, with Plates I - XII. [https: // www. biodiversitylibrary. org / item / 8386 # page / 9 / mode / 1 up]" type="url" year="1891">Scott Elliot (1891: 14)</bibRefCitation>
</taxonomicName>
[i.e.,
<taxonomicName id="B09170C1FF895C6BFB691582FE6B3EA3" authority="(Harvey 1862: 380) Hamet (1912: 44)" authorityName="Hamet" authorityPageNumber="44" authorityYear="1912" baseAuthorityName="Harvey" baseAuthorityPageNumber="380" baseAuthorityYear="1862" class="Magnoliopsida" family="Crassulaceae" genus="Kalanchoe" kingdom="Plantae" order="Saxifragales" pageId="4" pageNumber="162" phylum="Tracheophyta" rank="species" species="tubiflora">
<emphasis id="45E5D750FF895C6BFB691582FAFB3E87" box="[1219,1351,950,976]" italics="true" pageId="4" pageNumber="162">K. tubiflora</emphasis>
(
<bibRefCitation id="130076B3FF895C6BFAFD1583FF403EA3" author="Harvey, W. H." pageId="4" pageNumber="162" refId="ref6337" refString="Harvey, W. H. (1862) Order LIII. Crassulaceae, D. C. VIII. Kalanchoe, Adans. In: W. H. Harvey &amp; O. W. Sonder (Eds.) Flora capensis [being a systematic description of the plants of the Cape Colony, Caffraria, &amp; Port Natal]. Vol. 2. L. Reeve &amp; Co., Ltd, Kent, pp. 378 - 380. [https: // www. biodiversitylibrary. org / item / 15233 # page / 386 / mode / 1 up]" type="url" year="1862">Harvey 1862: 380</bibRefCitation>
)
<bibRefCitation id="130076B3FF895C6BFEA615EEFE6B3EA3" author="Hamet, R." box="[268,471,986,1013]" pageId="4" pageNumber="162" pagination="41 - 44" refId="ref6280" refString="Hamet, R. (1912 [12 April]) Observations sur le Kalanchoe tubiflora nom. nov. Beihefte zum Botanischen Centralblatt 29 (Heft 1): 41 - 44. [https: // www. biodiversitylibrary. org / item / 27107 # page / 523 / mode / 1 up]" type="journal article" year="1912">Hamet (1912: 44)</bibRefCitation>
</taxonomicName>
, see
<bibRefCitation id="130076B3FF895C6BFDBA15EFFCCF3EA2" author="Figueiredo, E. &amp; Smith, G. F." box="[528,883,986,1013]" pageId="4" pageNumber="162" pagination="717" refId="ref5723" refString="Figueiredo, E. &amp; Smith, G. F. (2017) (56) Request for a binding decision on the descriptive statement associated with Kalanchoe delagoensis (Crassulaceae). Taxon 66: 717. https: // doi. org / 10.12705 / 663.37" type="journal article" year="2017">Figueiredo &amp; Smith 2017: 717</bibRefCitation>
)]; (2)
<taxonomicName id="B09170C1FF895C6BFC1215EEFABE3EA3" authority="Baillon (1888" authorityName="Baillon" authorityYear="1888" box="[952,1282,986,1012]" class="Magnoliopsida" family="Crassulaceae" genus="Kalanchoe" kingdom="Plantae" order="Saxifragales" pageId="4" pageNumber="162" phylum="Tracheophyta" rank="species" species="grandidieri">
<emphasis id="45E5D750FF895C6BFC1215EEFBE63EA3" box="[952,1114,986,1012]" italics="true" pageId="4" pageNumber="162">K. grandidieri</emphasis>
<bibRefCitation id="130076B3FF895C6BFBC815EFFABE3EA3" author="Baillon, M. H." box="[1122,1282,986,1012]" pageId="4" pageNumber="162" pagination="51 - 88" refId="ref5276" refString="Baillon, M. H. (1888) Kalanchoe grandidieri, Plate 57 [of Plates 51 - 88]. In: A. Grandidier, Histoire physique, naturelle et politique Madagascar publiee par Alfred Grandidier. Vol. XXX. Histoire naturelle des plantes par M. E. Drake del Castillo. Tome II - Texte - 1, partie. 50 Fascicule. Imprime par Autorisation de M. Le Garde des Sceaux. A l'Imprimerie Nationale, Paris, 130 plates. [https: // www. biodiversitylibrary. org / item / 41106 # page / 71 / mode / 1 up]" type="book chapter" year="1888">Baillon (1888</bibRefCitation>
</taxonomicName>
: t. 57); and (3)
<taxonomicName id="B09170C1FF895C6BFF2215CAFE9E394F" box="[136,290,1022,1048]" class="Magnoliopsida" family="Crassulaceae" genus="Kalanchoe" kingdom="Plantae" order="Saxifragales" pageId="4" pageNumber="162" phylum="Tracheophyta" rank="species" species="beharensis">
<emphasis id="45E5D750FF895C6BFF2215CAFE9E394F" box="[136,290,1022,1048]" italics="true" pageId="4" pageNumber="162">K. beharensis</emphasis>
</taxonomicName>
, and he was then able to describe the leaves of the latter. It is noteworthy that the natural geographical distribution ranges of both
<taxonomicName id="B09170C1FF895C6BFE1D1216FD86396B" authorityName="Hamet" authorityYear="1912" baseAuthorityName="Harvey" baseAuthorityYear="1862" box="[439,570,1058,1084]" class="Magnoliopsida" family="Crassulaceae" genus="Kalanchoe" kingdom="Plantae" order="Saxifragales" pageId="4" pageNumber="162" phylum="Tracheophyta" rank="species" species="tubiflora">
<emphasis id="45E5D750FF895C6BFE1D1216FD86396B" box="[439,570,1058,1084]" italics="true" pageId="4" pageNumber="162">K. tubiflora</emphasis>
</taxonomicName>
and
<taxonomicName id="B09170C1FF895C6BFDDA1216FCAD396B" authorityName="Baillon" authorityYear="1888" box="[624,785,1058,1084]" class="Magnoliopsida" family="Crassulaceae" genus="Kalanchoe" kingdom="Plantae" order="Saxifragales" pageId="4" pageNumber="162" phylum="Tracheophyta" rank="species" species="grandidieri">
<emphasis id="45E5D750FF895C6BFDDA1216FCAD396B" box="[624,785,1058,1084]" italics="true" pageId="4" pageNumber="162">K. grandidieri</emphasis>
</taxonomicName>
overlap with that of
<taxonomicName id="B09170C1FF895C6BFC501216FB2F396B" box="[1018,1171,1058,1084]" class="Magnoliopsida" family="Crassulaceae" genus="Kalanchoe" kingdom="Plantae" order="Saxifragales" pageId="4" pageNumber="162" phylum="Tracheophyta" rank="species" species="beharensis">
<emphasis id="45E5D750FF895C6BFC501216FB2F396B" box="[1018,1171,1058,1084]" italics="true" pageId="4" pageNumber="162">K. beharensis</emphasis>
</taxonomicName>
, which could have given rise to the admixture. To maintain the interpretation of the name
<taxonomicName id="B09170C1FF895C6BFC2E1272FB853937" authorityName="Hamet" authorityYear="1906" box="[900,1081,1094,1120]" class="Magnoliopsida" family="Crassulaceae" genus="Kalanchoe" kingdom="Plantae" order="Saxifragales" pageId="4" pageNumber="162" phylum="Tracheophyta" rank="species" species="vantieghemii">
<emphasis id="45E5D750FF895C6BFC2E1272FB853937" box="[900,1081,1094,1120]" italics="true" pageId="4" pageNumber="162">K. vantieghemii</emphasis>
</taxonomicName>
as a homotypic synonym of
<taxonomicName id="B09170C1FF895C6BFA381272FEBC39D3" class="Magnoliopsida" family="Crassulaceae" genus="Kalanchoe" kingdom="Plantae" order="Saxifragales" pageId="4" pageNumber="162" phylum="Tracheophyta" rank="species" species="beharensis">
<emphasis id="45E5D750FF895C6BFA381272FEBC39D3" italics="true" pageId="4" pageNumber="162">K. beharensis</emphasis>
</taxonomicName>
, as has been done consistently in the literature, we here lectotypify it on the same two-sheet specimen on which the name
<taxonomicName id="B09170C1FF895C6BFEEA12BAFE6539FF" box="[320,473,1166,1192]" class="Magnoliopsida" family="Crassulaceae" genus="Kalanchoe" kingdom="Plantae" order="Saxifragales" pageId="4" pageNumber="162" phylum="Tracheophyta" rank="species" species="beharensis">
<emphasis id="45E5D750FF895C6BFEEA12BAFE6539FF" box="[320,473,1166,1192]" italics="true" pageId="4" pageNumber="162">K. beharensis</emphasis>
</taxonomicName>
is lectotypified (see above), which agrees with the views of
<bibRefCitation id="130076B3FF895C6BFBD512BAFAEA39FF" author="Hamet, R." box="[1151,1366,1166,1193]" pageId="4" pageNumber="162" pagination="17 - 48" refId="ref5980" refString="Hamet, R. (1908 a) Monographie du genre Kalanchoe. Suite et fin. Bulletin de l'Herbier Boissier, ser. 2, 8: 17 - 48. [https: // www. biodiversitylibrary. org / item / 104945 # page / 31 / mode / 1 up]" type="journal article" year="1908">Hamet (1908a: 30)</bibRefCitation>
.
</paragraph>
<paragraph id="772E0B42FF895C6BFF171287FAEC38F2" blockId="4.[136,1452,158,2057]" pageId="4" pageNumber="162">
Apart from the morphological conflict in the description of
<taxonomicName id="B09170C1FF895C6BFCCA1286FC46399B" box="[864,1018,1202,1228]" class="Magnoliopsida" family="Crassulaceae" genus="Kalanchoe" kingdom="Plantae" order="Saxifragales" pageId="4" pageNumber="162" phylum="Tracheophyta" rank="species" species="beharensis">
<emphasis id="45E5D750FF895C6BFCCA1286FC46399B" box="[864,1018,1202,1228]" italics="true" pageId="4" pageNumber="162">K. beharensis</emphasis>
</taxonomicName>
, a further complication in interpreting the nomenclature of
<taxonomicName id="B09170C1FF895C6BFEC712E2FDB939A7" box="[365,517,1238,1264]" class="Magnoliopsida" family="Crassulaceae" genus="Kalanchoe" kingdom="Plantae" order="Saxifragales" pageId="4" pageNumber="162" phylum="Tracheophyta" rank="species" species="beharensis">
<emphasis id="45E5D750FF895C6BFEC712E2FDB939A7" box="[365,517,1238,1264]" italics="true" pageId="4" pageNumber="162">K. beharensis</emphasis>
</taxonomicName>
, is that, as noted above, Drake del
<bibRefCitation id="130076B3FF895C6BFC2212E3FBE039A7" author="Castillo, E." box="[904,1116,1238,1264]" pageId="4" pageNumber="162" pagination="35 - 46" refId="ref5654" refString="Drake del Castillo, E. (1903) Note sur les plantes recueillies par M. Guillaume Grandidier dans le sud de Madagascar, en 1898 et 1901. (A suivre). Bulletin du Museum d'Histoire Naturelle 9: 35 - 46. [https: // www. biodiversitylibrary. org / page / 42893852 # page / 45 / mode / 1 up]" type="journal article" year="1903">Castillo (1903: 41)</bibRefCitation>
, when publishing the name
<taxonomicName id="B09170C1FF895C6BFA3812E2FEBC3843" class="Magnoliopsida" family="Crassulaceae" genus="Kalanchoe" kingdom="Plantae" order="Saxifragales" pageId="4" pageNumber="162" phylum="Tracheophyta" rank="species" species="beharensis">
<emphasis id="45E5D750FF895C6BFA3812E2FEBC3843" italics="true" pageId="4" pageNumber="162">K. beharensis</emphasis>
</taxonomicName>
, only stated: “Behara (
<date id="032F2D82FF895C6BFDAC12CFFD1C3843" box="[518,672,1274,1300]" pageId="4" pageNumber="162" value="1901-07-08">8 juillet 1901</date>
)” where he cited material in the protologue of the name. Two further voucher specimens cited under other species names treated by Drake del
<bibRefCitation id="130076B3FF895C6BFC79132BFB6B386F" author="Castillo, E." box="[979,1239,1310,1336]" pageId="4" pageNumber="162" pagination="35 - 46" refId="ref5654" refString="Drake del Castillo, E. (1903) Note sur les plantes recueillies par M. Guillaume Grandidier dans le sud de Madagascar, en 1898 et 1901. (A suivre). Bulletin du Museum d'Histoire Naturelle 9: 35 - 46. [https: // www. biodiversitylibrary. org / page / 42893852 # page / 45 / mode / 1 up]" type="journal article" year="1903">Castillo (1903: 4243)</bibRefCitation>
, but not originally described by him, were identified in an identical way, i.e., as “Behara (
<date id="032F2D82FF895C6BFC031377FBF9380B" box="[937,1093,1346,1372]" pageId="4" pageNumber="162" value="1901-07-08">8 juillet 1901</date>
)”. Apart from “Behara (
<date id="032F2D82FF895C6BFAFE1377FF7F38D7" pageId="4" pageNumber="162" value="1901-07-08">8 juillet 1901</date>
)” referring to material today interpreted as being of
<taxonomicName id="B09170C1FF895C6BFCBD1352FC0D38D7" box="[791,945,1382,1408]" class="Magnoliopsida" family="Crassulaceae" genus="Kalanchoe" kingdom="Plantae" order="Saxifragales" pageId="4" pageNumber="162" phylum="Tracheophyta" rank="species" species="beharensis">
<emphasis id="45E5D750FF895C6BFCBD1352FC0D38D7" box="[791,945,1382,1408]" italics="true" pageId="4" pageNumber="162">K. beharensis</emphasis>
</taxonomicName>
(1.1 and 1.2, below), this specimen citation, “Behara (
<date id="032F2D82FF895C6BFF5E13BFFE3138F3" box="[244,397,1418,1444]" pageId="4" pageNumber="162" value="1901-07-08">8 juillet 1901</date>
)”, was done in exactly the same way under two other species names (2 and 3, below):
</paragraph>
</subSubSection>
<subSubSection id="3F8B58C9FF895C6BFF17139BFCB03BBE" pageId="4" pageNumber="162" type="materials_examined">
<paragraph id="772E0B42FF895C6BFF17139BFEE538BB" blockId="4.[136,1452,158,2057]" pageId="4" pageNumber="162">
1.
<taxonomicName id="B09170C1FF895C6BFF73139BFE6C389F" box="[217,464,1454,1480]" class="Magnoliopsida" family="Crassulaceae" genus="Kalanchoe" kingdom="Plantae" order="Saxifragales" pageId="4" pageNumber="162" phylum="Tracheophyta" rank="species" species="beharensis">
<emphasis id="45E5D750FF895C6BFF73139BFE6C389F" box="[217,464,1454,1480]" italics="true" pageId="4" pageNumber="162">Kalanchoe beharensis</emphasis>
</taxonomicName>
, a single specimen mounted on two sheets, i.e., as more than one preparation (
<bibRefCitation id="130076B3FF895C6BFA9D139BFF5438BB" author="Turland, N. J. &amp; Wiersema, J. H. &amp; Barrie, F. R. &amp; Greuter, W. &amp; Hawksworth, D. L. &amp; Herendeen, P. S. &amp; Knapp, S. &amp; Kusber, W. - H. &amp; Li, D. - Z. &amp; Marhold, K. &amp; May, T. W. &amp; McNeill, J. &amp; Monro, A. M. &amp; Prado, J. &amp; Price, M. J. &amp; Smith, G. F." pageId="4" pageNumber="162" refId="ref6952" refString="Turland, N. J., Wiersema, J. H., Barrie, F. R., Greuter, W., Hawksworth, D. L., Herendeen, P. S., Knapp, S., Kusber, W. - H., Li, D. - Z., Marhold, K., May, T. W., McNeill, J., Monro, A. M., Prado, J., Price, M. J. &amp; Smith, G. F. (2018) International Code of Nomenclature for algae, fungi, and plants (Shenzhen Code) adopted by the Nineteenth International Botanical Congress Shenzhen, China, July 2017. Koeltz Botanical Books, Glashutten. [Regnum Vegetabile 159], 254 pp. https: // doi. org / 10.12705 / Code. 2018" type="book" year="2018">
Turland
<emphasis id="45E5D750FF895C6BFA3D139AFF2238BB" italics="true" pageId="4" pageNumber="162">et al</emphasis>
. 2018
</bibRefCitation>
: Art. 8.3):
</paragraph>
<paragraph id="772E0B42FF895C6BFF1713C3FAD23B46" blockId="4.[136,1452,158,2057]" box="[189,1390,1526,1553]" pageId="4" pageNumber="162">
1.1 Herb.
<collectionCode id="11809387FF895C6BFE8713C2FE803B46" box="[301,316,1527,1553]" country="France" lsid="urn:lsid:biocol.org:col:15763" name="Museum National d' Histoire Naturelle, Paris (MNHN) - Vascular Plants" pageId="4" pageNumber="162" type="Herbarium">P</collectionCode>
[barcode
<collectionCode id="11809387FF895C6BFE0613C3FE003B47" box="[428,444,1526,1552]" country="France" lsid="urn:lsid:biocol.org:col:15763" name="Museum National d' Histoire Naturelle, Paris (MNHN) - Vascular Plants" pageId="4" pageNumber="162" type="Herbarium">P</collectionCode>
00374194! http://coldb.mnhn.fr/catalognumber/mnhn/p/p00374194 (inflorescence)];
</paragraph>
<paragraph id="772E0B42FF895C6BFF17102FFEDB3B0E" blockId="4.[136,1452,158,2057]" pageId="4" pageNumber="162">
1.2 Herb.
<collectionCode id="11809387FF895C6BFE98102EFEFD3B62" box="[306,321,1563,1589]" country="France" lsid="urn:lsid:biocol.org:col:15763" name="Museum National d' Histoire Naturelle, Paris (MNHN) - Vascular Plants" pageId="4" pageNumber="162" type="Herbarium">P</collectionCode>
[barcode
<collectionCode id="11809387FF895C6BFE1F102FFE793B62" box="[437,453,1562,1589]" country="France" lsid="urn:lsid:biocol.org:col:15763" name="Museum National d' Histoire Naturelle, Paris (MNHN) - Vascular Plants" pageId="4" pageNumber="162" type="Herbarium">P</collectionCode>
00374195! http://coldb.mnhn.fr/catalognumber/mnhn/p/p00374195 (leaves in envelopes and inflorescence)];
</paragraph>
<paragraph id="772E0B42FF895C6BFF171057FA8F3BF7" blockId="4.[136,1452,158,2057]" pageId="4" pageNumber="162">
2. “(?)”
<taxonomicName id="B09170C1FF895C6BFEB21056FC893B2B" authority="Hils. &amp; Bojer ex Tulasne (1857: 149)" authorityName="Hils. &amp; Bojer ex Tulasne" authorityPageNumber="149" authorityYear="1857" box="[280,821,1634,1661]" class="Magnoliopsida" family="Crassulaceae" genus="Kalanchoe" kingdom="Plantae" order="Saxifragales" pageId="4" pageNumber="162" phylum="Tracheophyta" rank="species" species="miniata">
<emphasis id="45E5D750FF895C6BFEB21056FE323B2B" box="[280,398,1635,1660]" italics="true" pageId="4" pageNumber="162">K. miniata</emphasis>
Hils. &amp; Bojer ex
<bibRefCitation id="130076B3FF895C6BFDF91057FC893B2B" author="Tulasne, L. - R." box="[595,821,1634,1661]" pageId="4" pageNumber="162" pagination="44 - 163" refId="ref6875" refString="Tulasne, L. - R. [' Edmond']. (1857) Fragmentum alterum (1). Florae madagascariensis. II. Calanchoe [sic, Kalanchoe] pp. 149 - 150. Annales des Sciences Naturelles; Botanique ser. 4, 8: 44 - 163. [https: // www. biodiversitylibrary. org / item / 129817 # page / 153 / mode / 1 up]" type="journal article" year="1857">Tulasne (1857: 149)</bibRefCitation>
</taxonomicName>
(see Drake del
<bibRefCitation id="130076B3FF895C6BFC4E1057FB1B3B2B" author="Castillo, E." box="[996,1191,1634,1660]" pageId="4" pageNumber="162" pagination="35 - 46" refId="ref5654" refString="Drake del Castillo, E. (1903) Note sur les plantes recueillies par M. Guillaume Grandidier dans le sud de Madagascar, en 1898 et 1901. (A suivre). Bulletin du Museum d'Histoire Naturelle 9: 35 - 46. [https: // www. biodiversitylibrary. org / page / 42893852 # page / 45 / mode / 1 up]" type="journal article" year="1903">Castillo 1903: 42</bibRefCitation>
; Herb.
<collectionCode id="11809387FF895C6BFB521056FABB3B2A" box="[1272,1287,1635,1661]" country="France" lsid="urn:lsid:biocol.org:col:15763" name="Museum National d' Histoire Naturelle, Paris (MNHN) - Vascular Plants" pageId="4" pageNumber="162" type="Herbarium">P</collectionCode>
[specimen not accessible online; possibly a voucher to record the unlikely natural presence of
<taxonomicName id="B09170C1FF895C6BFBA810B2FBC43BF7" authorityName="Hils. &amp; Bojer ex Tulasne" authorityYear="1857" box="[1026,1144,1671,1696]" class="Magnoliopsida" family="Crassulaceae" genus="Kalanchoe" kingdom="Plantae" order="Saxifragales" pageId="4" pageNumber="162" phylum="Tracheophyta" rank="species" species="miniata">
<emphasis id="45E5D750FF895C6BFBA810B2FBC43BF7" box="[1026,1144,1671,1696]" italics="true" pageId="4" pageNumber="162">K. miniata</emphasis>
</taxonomicName>
at Behara]); and
</paragraph>
<paragraph id="772E0B42FF895C6BFF17109FFCB03BBE" blockId="4.[136,1452,158,2057]" pageId="4" pageNumber="162">
3.
<taxonomicName id="B09170C1FF895C6BFF76109FFD7F3B92" authority="Baillon (1887: 672)" authorityName="Baillon" authorityPageNumber="672" authorityYear="1887" box="[220,707,1706,1733]" class="Magnoliopsida" family="Euphorbiaceae" genus="Euphorbia" kingdom="Plantae" order="Malpighiales" pageId="4" pageNumber="162" phylum="Tracheophyta" rank="species" species="stenoclada">
<emphasis id="45E5D750FF895C6BFF76109FFE6B3B93" box="[220,471,1706,1732]" italics="true" pageId="4" pageNumber="162">Euphorbia stenoclada</emphasis>
<bibRefCitation id="130076B3FF895C6BFE4B109FFD7F3B92" author="Baillon, M. H." box="[481,707,1706,1733]" pageId="4" pageNumber="162" pagination="671 - 672" refId="ref5212" refString="Baillon, M. H. (1887) Liste des plantes de Madagascar (suite de la page 624). Bulletin Mensuel de la Societe Linneenne de Paris 1 (No. 84): 671 - 672. [https: // www. biodiversitylibrary. org / page / 11162078 # page / 24 / mode / 1 up]" type="book chapter" year="1887">Baillon (1887: 672)</bibRefCitation>
</taxonomicName>
(see Drake del
<bibRefCitation id="130076B3FF895C6BFCD4109FFBFB3B93" author="Castillo, E." box="[894,1095,1706,1732]" pageId="4" pageNumber="162" pagination="35 - 46" refId="ref5654" refString="Drake del Castillo, E. (1903) Note sur les plantes recueillies par M. Guillaume Grandidier dans le sud de Madagascar, en 1898 et 1901. (A suivre). Bulletin du Museum d'Histoire Naturelle 9: 35 - 46. [https: // www. biodiversitylibrary. org / page / 42893852 # page / 45 / mode / 1 up]" type="journal article" year="1903">Castillo 1903: 43</bibRefCitation>
; Herb.
<collectionCode id="11809387FF895C6BFB34109EFB113B92" box="[1182,1197,1707,1733]" country="France" lsid="urn:lsid:biocol.org:col:15763" name="Museum National d' Histoire Naturelle, Paris (MNHN) - Vascular Plants" pageId="4" pageNumber="162" type="Herbarium">P</collectionCode>
[barcode
<collectionCode id="11809387FF895C6BFA88109FFA8E3B93" box="[1314,1330,1706,1732]" country="France" lsid="urn:lsid:biocol.org:col:15763" name="Museum National d' Histoire Naturelle, Paris (MNHN) - Vascular Plants" pageId="4" pageNumber="162" type="Herbarium">P</collectionCode>
00226016! http://coldb.mnhn.fr/catalognumber/mnhn/p/p00226016].
</paragraph>
</subSubSection>
<subSubSection id="3F8B58C9FF895C6AFF1710C7FD963E17" lastPageId="5" lastPageNumber="163" pageId="4" pageNumber="162" type="discussion">
<paragraph id="772E0B42FF895C6BFF1710C7FB913A2E" blockId="4.[136,1452,158,2057]" pageId="4" pageNumber="162">
The effect of at least three
<taxonomicName id="B09170C1FF895C6BFE4310C7FDDE3A5B" authorityName="Adanson" authorityYear="1763" box="[489,610,1778,1804]" class="Magnoliopsida" family="Crassulaceae" genus="Kalanchoe" kingdom="Plantae" order="Saxifragales" pageId="4" pageNumber="162" phylum="Tracheophyta" rank="genus">
<emphasis id="45E5D750FF895C6BFE4310C7FDDE3A5B" box="[489,610,1778,1804]" italics="true" pageId="4" pageNumber="162">Kalanchoe</emphasis>
</taxonomicName>
sheets, numbers 1.1, 1.2, and 2, above, and, for that matter, a fourth one of a
<taxonomicName id="B09170C1FF895C6BFF361123FEA83A67" box="[156,276,1814,1840]" class="Magnoliopsida" family="Euphorbiaceae" genus="Euphorbia" kingdom="Plantae" order="Malpighiales" pageId="4" pageNumber="162" phylum="Tracheophyta" rank="genus">
<emphasis id="45E5D750FF895C6BFF361123FEA83A67" box="[156,276,1814,1840]" italics="true" pageId="4" pageNumber="162">Euphorbia</emphasis>
</taxonomicName>
, having been cited and identified in exactly the same way, i.e., as [
<emphasis id="45E5D750FF895C6BFBA21122FB6C3A67" box="[1032,1232,1814,1841]" italics="true" pageId="4" pageNumber="162">G. Grandidier s.n</emphasis>
.] “Behara (
<date id="032F2D82FF895C6BFAFE1123FF7F3A03" pageId="4" pageNumber="162" value="1901-07-08">8 juillet 1901</date>
)” by Drake del
<bibRefCitation id="130076B3FF895C6BFED9110FFDFA3A03" author="Castillo, E." box="[371,582,1850,1876]" pageId="4" pageNumber="162" pagination="35 - 46" refId="ref5654" refString="Drake del Castillo, E. (1903) Note sur les plantes recueillies par M. Guillaume Grandidier dans le sud de Madagascar, en 1898 et 1901. (A suivre). Bulletin du Museum d'Histoire Naturelle 9: 35 - 46. [https: // www. biodiversitylibrary. org / page / 42893852 # page / 45 / mode / 1 up]" type="journal article" year="1903">Castillo (1903: 41)</bibRefCitation>
was that considerable confusion was caused and contributed to
<bibRefCitation id="130076B3FF895C6BFAA5110EFA173A02" author="Hamet, R." box="[1295,1451,1850,1877]" pageId="4" pageNumber="162" pagination="109 - 111" refId="ref5772" refString="Hamet, R. (1906 [Janvier - Mars]) Note sur une nouvelle espece de Kalanchoe [Kalanchoe vantieghemii, as ' Van Tieghemi ']. Journal de Botanique [Morot] XX [20 e [annee] (No 1 - 3)]: 109 - 111. [https: // www. biodiversitylibrary. org / item / 22106 # page / 113 / mode / 1 up]" type="book chapter" year="1906">Hamet (1906)</bibRefCitation>
redescribing, as
<taxonomicName id="B09170C1FF895C6BFE95116AFE4C3A2F" authorityName="Hamet" authorityYear="1906" box="[319,496,1886,1912]" class="Magnoliopsida" family="Crassulaceae" genus="Kalanchoe" kingdom="Plantae" order="Saxifragales" pageId="4" pageNumber="162" phylum="Tracheophyta" rank="species" species="vantieghemii">
<emphasis id="45E5D750FF895C6BFE95116AFE4C3A2F" box="[319,496,1886,1912]" italics="true" pageId="4" pageNumber="162">K. vantieghemii</emphasis>
</taxonomicName>
, material of what today is known as
<taxonomicName id="B09170C1FF895C6BFC27116AFB9A3A2F" box="[909,1062,1886,1912]" class="Magnoliopsida" family="Crassulaceae" genus="Kalanchoe" kingdom="Plantae" order="Saxifragales" pageId="4" pageNumber="162" phylum="Tracheophyta" rank="species" species="beharensis">
<emphasis id="45E5D750FF895C6BFC27116AFB9A3A2F" box="[909,1062,1886,1912]" italics="true" pageId="4" pageNumber="162">K. beharensis</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="772E0B42FF895C6BFF1711B7FB74355F" blockId="4.[136,1452,158,2057]" pageId="4" pageNumber="162">
While we accept that the type of the name
<taxonomicName id="B09170C1FF895C6BFD2411B6FC993ACB" box="[654,805,1922,1948]" class="Magnoliopsida" family="Crassulaceae" genus="Kalanchoe" kingdom="Plantae" order="Saxifragales" pageId="4" pageNumber="162" phylum="Tracheophyta" rank="species" species="beharensis">
<emphasis id="45E5D750FF895C6BFD2411B6FC993ACB" box="[654,805,1922,1948]" italics="true" pageId="4" pageNumber="162">K. beharensis</emphasis>
</taxonomicName>
consists of
<specimenCount id="6197C0CBFF895C6BFC0F11B6FBF63ACB" box="[933,1098,1922,1949]" pageId="4" pageNumber="162" type="generic">two specimens</specimenCount>
(see below), we do not interpret this as the
<typeStatus id="A82AB5E0FF895C6BFF561193FEDD3A97" box="[252,353,1958,1984]" pageId="4" pageNumber="162" type="holotype">holotype</typeStatus>
, given that
<specimenCount id="6197C0CBFF895C6BFE751193FD343A97" box="[479,648,1958,1984]" pageId="4" pageNumber="162" type="generic">four specimens</specimenCount>
representing three species were cited with identical information. Rather, we interpret “Behara (
<date id="032F2D82FF895C6BFE2B11FFFDA43AB3" box="[385,536,1994,2020]" pageId="4" pageNumber="162" value="1901-07-08">
<collectingDate id="136BD46AFF895C6BFE2B11FFFDA43AB3" box="[385,536,1994,2020]" pageId="4" pageNumber="162" value="1901-07-08">8 juillet 1901</collectingDate>
</date>
)” represented by Herb. P [barcode
<specimenCode id="2737A339FF895C6BFC0A11FFFB9E3AB2" box="[928,1058,1994,2021]" collectionCode="P" country="France" lsid="urn:lsid:biocol.org:col:15763" name="Museum National d' Histoire Naturelle, Paris (MNHN) - Vascular Plants" pageId="4" pageNumber="162" type="Herbarium">P00374194</specimenCode>
] and Herb. P [barcode
<specimenCode id="2737A339FF895C6BFA8811FFFA193AB2" box="[1314,1445,1994,2021]" collectionCode="P" country="France" lsid="urn:lsid:biocol.org:col:15763" name="Museum National d' Histoire Naturelle, Paris (MNHN) - Vascular Plants" pageId="4" pageNumber="162" type="Herbarium">P00374195</specimenCode>
] as original material that qualify for designation as
<typeStatus id="A82AB5E0FF895C6BFD1711DBFC9B355F" box="[701,807,2030,2056]" pageId="4" pageNumber="162" type="lectotype">lectotype</typeStatus>
. We here affect this lectotypification.
</paragraph>
<paragraph id="772E0B42FF885C6AFF1716ABFDAE3FE6" blockId="5.[136,1452,158,832]" pageId="5" pageNumber="163">
The
<typeStatus id="A82AB5E0FF885C6AFF5B16AAFE9D3DEE" box="[241,289,159,185]" pageId="5" pageNumber="163">type</typeStatus>
of the name
<taxonomicName id="B09170C1FF885C6AFE1F16AAFDF33DEF" box="[437,591,158,184]" class="Magnoliopsida" family="Crassulaceae" genus="Kalanchoe" kingdom="Plantae" order="Saxifragales" pageId="5" pageNumber="163" phylum="Tracheophyta" rank="species" species="beharensis">
<emphasis id="45E5D750FF885C6AFE1F16AAFDF33DEF" box="[437,591,158,184]" italics="true" pageId="5" pageNumber="163">K. beharensis</emphasis>
</taxonomicName>
consists of two preparations, in this case two herbarium sheets, i.e., Herb. P [barcode P00374194] and Herb. P [barcode P00374195], which together constitute
<specimenCount id="6197C0CBFF885C6AFB8216F6FB7C3D8B" box="[1064,1216,194,221]" pageId="5" pageNumber="163" type="generic">one specimen</specimenCount>
(
<bibRefCitation id="130076B3FF885C6AFB6416F7FA143D8B" author="Turland, N. J. &amp; Wiersema, J. H. &amp; Barrie, F. R. &amp; Greuter, W. &amp; Hawksworth, D. L. &amp; Herendeen, P. S. &amp; Knapp, S. &amp; Kusber, W. - H. &amp; Li, D. - Z. &amp; Marhold, K. &amp; May, T. W. &amp; McNeill, J. &amp; Monro, A. M. &amp; Prado, J. &amp; Price, M. J. &amp; Smith, G. F." box="[1230,1448,194,221]" pageId="5" pageNumber="163" refId="ref6952" refString="Turland, N. J., Wiersema, J. H., Barrie, F. R., Greuter, W., Hawksworth, D. L., Herendeen, P. S., Knapp, S., Kusber, W. - H., Li, D. - Z., Marhold, K., May, T. W., McNeill, J., Monro, A. M., Prado, J., Price, M. J. &amp; Smith, G. F. (2018) International Code of Nomenclature for algae, fungi, and plants (Shenzhen Code) adopted by the Nineteenth International Botanical Congress Shenzhen, China, July 2017. Koeltz Botanical Books, Glashutten. [Regnum Vegetabile 159], 254 pp. https: // doi. org / 10.12705 / Code. 2018" type="book" year="2018">
Turland
<emphasis id="45E5D750FF885C6AFA8416F6FAE23D8B" box="[1326,1374,194,220]" italics="true" pageId="5" pageNumber="163">et al</emphasis>
. 2018
</bibRefCitation>
: Art. 8.3). One of the sheets [P barcode P00374194] has an inflorescence mounted on it and the second sheet [P barcode P00374195] has an inflorescence and separate leaf fragments kept in two envelopes attached to it. The former specimen has an original label stating, inter alia, “feuilles à la page suivante”, i.e., “leaves on the next page”. Some collections of Grandidier have
<emphasis id="45E5D750FF885C6AFEEE1767FE293C3B" box="[324,405,338,364]" italics="true" pageId="5" pageNumber="163">printed</emphasis>
labels [not to be confused with
<emphasis id="45E5D750FF885C6AFD511767FCE93C3B" box="[763,853,338,364]" italics="true" pageId="5" pageNumber="163">original</emphasis>
labels] that are horizontal-rectangular with a narrowlined, black box inset a few millimetres from the edge, and with the words “Mission Grandidier. 1901” printed at the top and “
<collectingCountry id="0F864BD2FF885C6AFF4317AFFECA3CE3" box="[233,374,410,436]" name="Madagascar" pageId="5" pageNumber="163">Madagascar</collectingCountry>
” at the bottom (see for example bottom left corner of P00226016, the specimen of
<taxonomicName id="B09170C1FF885C6AFAAE17AEFA273CE3" authorityName="Baillon" authorityYear="1887" box="[1284,1435,410,436]" class="Magnoliopsida" family="Euphorbiaceae" genus="Euphorbia" kingdom="Plantae" order="Malpighiales" pageId="5" pageNumber="163" phylum="Tracheophyta" rank="species" species="stenoclada">
<emphasis id="45E5D750FF885C6AFAAE17AEFA273CE3" box="[1284,1435,410,436]" italics="true" pageId="5" pageNumber="163">E. stenoclada</emphasis>
</taxonomicName>
). Nevertheless, the majority of the specimens we examined only have handwritten labels attached to them. Although the specimen P00374194 lacks a printed label, the original label written in pencil is in the same handwriting as that which appears on other specimens collected by Grandidier and on the printed labels (see for example P00226016). The two sheets P00374194 and P00374195 therefore represent a single gathering and the two sheets have “a single, original label in common” (
<bibRefCitation id="130076B3FF885C6AFECB147BFD823F3F" author="Turland, N. J. &amp; Wiersema, J. H. &amp; Barrie, F. R. &amp; Greuter, W. &amp; Hawksworth, D. L. &amp; Herendeen, P. S. &amp; Knapp, S. &amp; Kusber, W. - H. &amp; Li, D. - Z. &amp; Marhold, K. &amp; May, T. W. &amp; McNeill, J. &amp; Monro, A. M. &amp; Prado, J. &amp; Price, M. J. &amp; Smith, G. F." box="[353,574,590,617]" pageId="5" pageNumber="163" refId="ref6952" refString="Turland, N. J., Wiersema, J. H., Barrie, F. R., Greuter, W., Hawksworth, D. L., Herendeen, P. S., Knapp, S., Kusber, W. - H., Li, D. - Z., Marhold, K., May, T. W., McNeill, J., Monro, A. M., Prado, J., Price, M. J. &amp; Smith, G. F. (2018) International Code of Nomenclature for algae, fungi, and plants (Shenzhen Code) adopted by the Nineteenth International Botanical Congress Shenzhen, China, July 2017. Koeltz Botanical Books, Glashutten. [Regnum Vegetabile 159], 254 pp. https: // doi. org / 10.12705 / Code. 2018" type="book" year="2018">
Turland
<emphasis id="45E5D750FF885C6AFE68147AFE483F3F" box="[450,500,590,616]" italics="true" pageId="5" pageNumber="163">et al</emphasis>
. 2018
</bibRefCitation>
: Art. 8.3). It is further assumed that the foreign leaf material [that gave rise to the admixture] noticed and mentioned by
<bibRefCitation id="130076B3FF885C6AFDF11446FC813FDB" author="Hamet, R." box="[603,829,626,653]" pageId="5" pageNumber="163" pagination="49 - 54" refId="ref6153" refString="Hamet, R. (1910 b [Janvier]) Sur quelques Kalanchoe peu connus (Suite). Bulletin de la Societe Botanique de France [Tome Cinquante- Septieme], Quatrieme serie - Tome X, Quatrieme series 57: 49 - 54. [https: // www. biodiversitylibrary. org / item / 8679 # page / 87 / mode / 1 up]" type="book chapter" year="1910">Hamet (1910b: 194)</bibRefCitation>
has since been removed from the envelopes attached to the sheet with barcode P00374195.
</paragraph>
<paragraph id="772E0B42FF885C6AFF17148EFD963E17" blockId="5.[136,1452,158,832]" pageId="5" pageNumber="163">
We therefore concur with
<bibRefCitation id="130076B3FF885C6AFE49148FFC223F82" author="Boiteau, P. &amp; Allorge-Boiteau, L." box="[483,926,698,725]" pageId="5" pageNumber="163" refId="ref5454" refString="Boiteau, P. &amp; Allorge-Boiteau, L. (1995) Kalanchoe (Crassulacees) de Madagascar. Systematique, ecophysiologie et phytochimie. ICSN, CNRS, 91198 Gif-sur-Yvette, Editions Karthala, Paris, 252 pp." type="book" year="1995">Boiteau &amp; Allorge-Boiteau (1995: 154)</bibRefCitation>
where they stated that the type of the name
<taxonomicName id="B09170C1FF885C6AFA38148EFEBC3FAF" class="Magnoliopsida" family="Crassulaceae" genus="Kalanchoe" kingdom="Plantae" order="Saxifragales" pageId="5" pageNumber="163" phylum="Tracheophyta" rank="species" species="beharensis">
<emphasis id="45E5D750FF885C6AFA38148EFEBC3FAF" italics="true" pageId="5" pageNumber="163">K. beharensis</emphasis>
</taxonomicName>
is “Grandidier Guillaume s.
<superScript id="80E4A60AFF885C6AFDE414EAFDD83FBB" attach="left" box="[590,612,734,761]" fontSize="6" pageId="5" pageNumber="163">no</superScript>
, Behara,
<date id="032F2D82FF885C6AFD7E14EBFC8A3FAF" box="[724,822,734,760]" pageId="5" pageNumber="163" value="1901-07-08">8.7.1901</date>
(fl.)
<typeStatus id="A82AB5E0FF885C6AFCD914EBFC693FAF" box="[883,981,734,760]" pageId="5" pageNumber="163" type="holotype">holotype</typeStatus>
P, 2 parts”, but we correct “
<typeStatus id="A82AB5E0FF885C6AFAB114EBFA3E3FAF" box="[1307,1410,734,760]" pageId="5" pageNumber="163" type="holotype">holotype</typeStatus>
” to “
<typeStatus id="A82AB5E0FF885C6AFF381537FEBD3E4B" box="[146,257,770,796]" pageId="5" pageNumber="163" type="lectotype">lectotype</typeStatus>
”. This lectotypification maintains the current wide application of the name
<taxonomicName id="B09170C1FF885C6AFBED1536FB633E4B" box="[1095,1247,770,796]" class="Magnoliopsida" family="Crassulaceae" genus="Kalanchoe" kingdom="Plantae" order="Saxifragales" pageId="5" pageNumber="163" phylum="Tracheophyta" rank="species" species="beharensis">
<emphasis id="45E5D750FF885C6AFBED1536FB633E4B" box="[1095,1247,770,796]" italics="true" pageId="5" pageNumber="163">K. beharensis</emphasis>
</taxonomicName>
to an arborescent, felt-leaved species from
<collectingCountry id="0F864BD2FF885C6AFE361513FD993E17" box="[412,549,806,832]" name="Madagascar" pageId="5" pageNumber="163">Madagascar</collectingCountry>
.
</paragraph>
</subSubSection>
</treatment>
</document>