258 lines
32 KiB
XML
258 lines
32 KiB
XML
<document id="33C2C34A206BF51B797823109BB6EB82" ID-CLB-Dataset="27631" ID-DOI="10.11646/zootaxa.4545.3.11" ID-GBIF-Dataset="e1d69bec-cc4a-49aa-815e-e1e74f34f6b6" ID-ISSN="1175-5326" ID-Zenodo-Dep="2618912" ID-ZooBank="63E4840E-2172-496B-A186-4C2D271DEA04" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1553930850276" checkinUser="plazi" docAuthor="Jałoszyński, Paweł" docDate="2019" docId="ED452153FFF8FFCE4395FE4AFCABFF3F" docLanguage="en" docName="zootaxa.4545.3.11.pdf" docOrigin="Zootaxa 4545 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Siamites sarawakensis Jałoszyński 2019, sp. n." docType="treatment" docVersion="8" lastPageNumber="448" masterDocId="117C592BFFF9FFCC4302FF85FFA1FFCB" masterDocTitle="The first species of Siamites Franz (Coleoptera, Staphylinidae, Scydmaeninae) in East Malaysia" masterLastPageNumber="450" masterPageNumber="447" pageNumber="448" updateTime="1698715945016" updateUser="ExternalLinkService">
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<mods:titleInfo id="B5557E6B6F2B92071FD04C99DEAC44A1">
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<mods:title id="38FEE33882BF42DBDFEDC21E4AEF7082">The first species of Siamites Franz (Coleoptera, Staphylinidae, Scydmaeninae) in East Malaysia</mods:title>
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</mods:titleInfo>
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<mods:name id="B7544671C8F093BC8E6E2F1817D7C29B" type="personal">
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<mods:roleTerm id="E204E519E0DA6A436635A216587AD230">Author</mods:roleTerm>
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<mods:namePart id="7577FD5A55181B407E267DCEE5CF5275">Jałoszyński, Paweł</mods:namePart>
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<mods:typeOfResource id="500CC0FA5BCC19E02527260F570EE377">text</mods:typeOfResource>
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<mods:title id="03D43FCE287409CA24ED191CF72A027E">Zootaxa</mods:title>
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<mods:part id="9165C07D556688109E92C7587BFEC467">
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<mods:date id="BE6995D2FE3F25D592D5926F811D648D">2019</mods:date>
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<mods:detail id="3317BD2DAEFB361D1AA411437DBC99B6" type="pubDate">
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<mods:number id="36620F8DAB054C552D74E449C6EE4680">2019-01-18</mods:number>
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<mods:number id="85B4AB68BC147F36763C934E492CBA2B">4545</mods:number>
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<mods:detail id="3EA4EF06C15E8EFF13E0CB5A6D6116BC" type="issue">
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<mods:identifier id="D55016F78C6F2CE3275DD00D435A5780" type="DOI">10.11646/zootaxa.4545.3.11</mods:identifier>
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<treatment id="ED452153FFF8FFCE4395FE4AFCABFF3F" ID-DOI="http://doi.org/10.5281/zenodo.5932592" ID-GBIF-Taxon="154531104" ID-Zenodo-Dep="5932592" LSID="urn:lsid:plazi:treatment:ED452153FFF8FFCE4395FE4AFCABFF3F" httpUri="http://treatment.plazi.org/id/ED452153FFF8FFCE4395FE4AFCABFF3F" lastPageId="2" lastPageNumber="448" pageId="1" pageNumber="448">
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<subSubSection id="2DF6C3CEFFF8FFCD4395FE4AFDD8FE21" box="[151,633,463,490]" pageId="1" pageNumber="448" type="nomenclature">
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<paragraph id="65539045FFF8FFCD4395FE4AFDD8FE21" blockId="1.[151,1437,429,2012]" box="[151,633,463,490]" pageId="1" pageNumber="448">
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<heading id="3E1B2729FFF8FFCD4395FE4AFDD8FE21" bold="true" box="[151,633,463,490]" fontSize="11" level="1" pageId="1" pageNumber="448" reason="1">
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<emphasis id="57984C57FFF8FFCD4395FE4AFDD8FE21" bold="true" box="[151,633,463,490]" pageId="1" pageNumber="448">
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<taxonomicName id="A2ECEBC6FFF8FFCD4395FE4AFE07FE22" ID-CoL="8Y9KZ" authority="Jałoszyński, 2019" authorityName="Jałoszyński" authorityYear="2019" box="[151,422,463,489]" class="Insecta" family="Staphylinidae" genus="Siamites" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="448" phylum="Arthropoda" rank="species" species="sarawakensis" status="sp. nov.">
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<emphasis id="57984C57FFF8FFCD4395FE4AFE07FE22" bold="true" box="[151,422,463,489]" italics="true" pageId="1" pageNumber="448">Siamites sarawakensis</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="4CABF12CFFF8FFCD42AFFE55FE4EFE21" box="[429,495,464,490]" pageId="1" pageNumber="448" rank="species">sp. n.</taxonomicNameLabel>
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(
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<figureCitation id="FDD78CC0FFF8FFCD42FDFE55FDD1FE21" box="[511,624,464,490]" captionStart="FIGURES 1–7" captionStartId="2.[151,258,1962,1984]" captionTargetBox="[151,1436,315,1938]" captionTargetId="figure@2.[151,1436,298,1938]" captionTargetPageId="2" captionText="FIGURES 1–7. Siamites sarawakensis sp. n., dorsal habitus of holotype male (1); head in anterior view (2); aedeagus in ventral (3, 5) and lateral (4, 6) views; and distribution of nominal Siamites species illustrated with their aedeagi in ventral view (7)." httpUri="https://zenodo.org/record/2618914/files/figure.png" pageId="1" pageNumber="448">Figs. 1–7</figureCitation>
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)
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="2DF6C3CEFFF8FFCD4395FE77FC91FD84" pageId="1" pageNumber="448" type="materials_examined">
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<paragraph id="65539045FFF8FFCD4395FE77FC91FD84" blockId="1.[151,1437,429,2012]" pageId="1" pageNumber="448">
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<emphasis id="57984C57FFF8FFCD4395FE77FD73FDC2" bold="true" box="[151,722,498,521]" pageId="1" pageNumber="448">
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Type material.
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<materialsCitation id="D5849A18FFF8FFCD4246FE77FD73FDC2" ID-GBIF-Occurrence="2147526481" box="[324,722,498,521]" collectingDate="1994-05-19" collectionCode="MHNG" collectorName="E. Kapit" country="Malaysia" determinerName="P. Jaloszynski" elevation="150" location="Mujong" municipality="Suan Oyan" pageId="1" pageNumber="448" specimenCount="1" specimenCount-male="1" stateProvince="Sarawak" typeStatus="holotype">
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<typeStatus id="BA572EE7FFF8FFCD4246FE77FE11FDC2" box="[324,432,498,521]" pageId="1" pageNumber="448" type="holotype">Holotype</typeStatus>
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:
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<collectingCountry id="1DFBD0D5FFF8FFCD42BEFE77FDEAFDC2" box="[444,587,498,521]" name="Malaysia" pageId="1" pageNumber="448">MALAYSIA</collectingCountry>
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(
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<collectingRegion id="A7285EA7FFF8FFCD4158FE77FD64FDC2" box="[602,709,498,521]" country="Malaysia" name="Sarawak" pageId="1" pageNumber="448">Sarawak</collectingRegion>
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):
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</materialsCitation>
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</emphasis>
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<specimenCount id="73EA5BCCFFF8FFCD41DBFE77FD4EFDC2" box="[729,751,498,521]" pageId="1" pageNumber="448" type="male">♂</specimenCount>
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, two labels: "E.
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<collectingCountry id="1DFBD0D5FFF8FFCD4091FE77FB86FDC2" box="[915,1063,498,521]" name="Malaysia" pageId="1" pageNumber="448">MALAYSIA</collectingCountry>
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:
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<collectingRegion id="A7285EA7FFF8FFCD4730FE77FB2FFDC2" box="[1074,1166,498,521]" country="Malaysia" name="Sarawak" pageId="1" pageNumber="448">Sarawak</collectingRegion>
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/ confl.
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<collectingMunicipality id="85370A3FFFF8FFCD47EAFE77FAFEFDC2" box="[1256,1375,498,521]" pageId="1" pageNumber="448">Suan Oyan</collectingMunicipality>
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and /
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<location id="6033C69EFFF8FFCD4395FD90FF4AFDE7" LSID="urn:lsid:plazi:treatment:ED452153FFF8FFCE4395FE4AFCABFF3F:6033C69EFFF8FFCD4395FD90FF4AFDE7" box="[151,235,533,556]" country="Malaysia" municipality="Suan Oyan" name="Mujong" pageId="1" pageNumber="448" stateProvince="Sarawak">Mujong</location>
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riv.,
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<collectorName id="C819F593FFF8FFCD421BFD90FECFFDE7" box="[281,366,533,556]" pageId="1" pageNumber="448">E.Kapit</collectorName>
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/
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<quantity id="A2143DA0FFF8FFCD4281FD90FE65FDE7" box="[387,452,533,556]" metricMagnitude="2" metricUnit="m" metricValue="1.5" pageId="1" pageNumber="448" unit="m" value="150.0">
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<elevation id="EEC17776FFF8FFCD4281FD90FE65FDE7" box="[387,452,533,556]" metricMagnitude="2" metricUnit="m" metricValue="1.5" pageId="1" pageNumber="448" unit="m" value="150.0">150m</elevation>
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</quantity>
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,
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<date id="1152B685FFF8FFCD42CAFD90FD96FDE7" box="[456,567,533,556]" pageId="1" pageNumber="448" value="1994-05-19">
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<collectingDate id="01164F6DFFF8FFCD42CAFD90FD96FDE7" box="[456,567,533,556]" pageId="1" pageNumber="448" value="1994-05-19">19.V.1994</collectingDate>
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</date>
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, # 6a / Löbl & Burckhardt" [white, printed]; "
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<taxonomicName id="A2ECEBC6FFF8FFCD4727FD90FB36FDE7" authorityName="Franz" authorityYear="1989" box="[1061,1175,533,556]" class="Insecta" family="Staphylinidae" genus="Siamites" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="448" phylum="Arthropoda" rank="genus">
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<emphasis id="57984C57FFF8FFCD4727FD90FB36FDE7" box="[1061,1175,533,556]" italics="true" pageId="1" pageNumber="448">SIAMITES</emphasis>
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</taxonomicName>
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/
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<taxonomicName id="A2ECEBC6FFF8FFCD47AEFD90FA9DFDE7" authorityName="Jałoszyński" authorityYear="2019" box="[1196,1340,533,556]" class="Insecta" family="Staphylinidae" genus="Siamites" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="448" phylum="Arthropoda" rank="species" species="sarawakensis">
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<emphasis id="57984C57FFF8FFCD47AEFD90FA9DFDE7" box="[1196,1340,533,556]" italics="true" pageId="1" pageNumber="448">sarawakensis</emphasis>
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</taxonomicName>
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m. / det.
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<determinerName id="908F7C81FFF8FFCD4395FDBDFE8FFD84" box="[151,302,568,591]" pageId="1" pageNumber="448">P. Jałoszyński</determinerName>
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, 2018 /
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<typeStatus id="BA572EE7FFF8FFCD4280FDBDFD82FD84" box="[386,547,568,591]" pageId="1" pageNumber="448" type="holotype">HOLOTYPUS</typeStatus>
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" [red, printed] (MHNG).
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</paragraph>
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</subSubSection>
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<subSubSection id="2DF6C3CEFFF8FFCD43C5FDDFFE95FD5F" pageId="1" pageNumber="448" type="diagnosis">
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<paragraph id="65539045FFF8FFCD43C5FDDFFE95FD5F" blockId="1.[151,1437,429,2012]" pageId="1" pageNumber="448">
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<emphasis id="57984C57FFF8FFCD43C5FDDFFE9DFDBA" bold="true" box="[199,316,602,625]" pageId="1" pageNumber="448">Diagnosis.</emphasis>
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Male macrophthalmous; aedeagus pear-shaped; subapical endophallic plate with concave lateral and distal margins.
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</paragraph>
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</subSubSection>
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<subSubSection id="2DF6C3CEFFF8FFCD43C5FD25FE32FA1F" pageId="1" pageNumber="448" type="description">
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<paragraph id="65539045FFF8FFCD43C5FD25FE5CFD12" blockId="1.[151,1437,429,2012]" pageId="1" pageNumber="448">
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<emphasis id="57984C57FFF8FFCD43C5FD25FEF3FD7C" bold="true" box="[199,338,672,695]" pageId="1" pageNumber="448">Description.</emphasis>
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Body of male (
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<figureCitation id="FDD78CC0FFF8FFCD42FDFD25FD9FFD7C" box="[511,574,672,695]" captionStart="FIGURES 1–7" captionStartId="2.[151,258,1962,1984]" captionTargetBox="[151,1436,315,1938]" captionTargetId="figure@2.[151,1436,298,1938]" captionTargetPageId="2" captionText="FIGURES 1–7. Siamites sarawakensis sp. n., dorsal habitus of holotype male (1); head in anterior view (2); aedeagus in ventral (3, 5) and lateral (4, 6) views; and distribution of nominal Siamites species illustrated with their aedeagi in ventral view (7)." httpUri="https://zenodo.org/record/2618914/files/figure.png" pageId="1" pageNumber="448">Fig. 1</figureCitation>
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) strongly convex and moderately slender, uniformly light brown, covered with yellowish vestiture. BL
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<quantity id="A2143DA0FFF8FFCD4295FD47FE58FD12" box="[407,505,706,729]" metricMagnitude="-3" metricUnit="m" metricValue="1.16" pageId="1" pageNumber="448" unit="mm" value="1.16">1.16 mm</quantity>
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.
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</paragraph>
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<paragraph id="65539045FFF8FFCD43C5FD60FD62FBFF" blockId="1.[151,1437,429,2012]" pageId="1" pageNumber="448">
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Head (
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<figureCitation id="FDD78CC0FFF8FFCD420CFD60FEEEFD37" box="[270,335,741,764]" captionStart="FIGURES 1–7" captionStartId="2.[151,258,1962,1984]" captionTargetBox="[151,1436,315,1938]" captionTargetId="figure@2.[151,1436,298,1938]" captionTargetPageId="2" captionText="FIGURES 1–7. Siamites sarawakensis sp. n., dorsal habitus of holotype male (1); head in anterior view (2); aedeagus in ventral (3, 5) and lateral (4, 6) views; and distribution of nominal Siamites species illustrated with their aedeagi in ventral view (7)." httpUri="https://zenodo.org/record/2618914/files/figure.png" pageId="1" pageNumber="448">Fig. 2</figureCitation>
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) broadest at large, strongly convex and coarsely faceted eyes, HL
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<quantity id="A2143DA0FFF8FFCD471BFD60FBDAFD37" box="[1049,1147,741,764]" metricMagnitude="-4" metricUnit="m" metricValue="2.1" pageId="1" pageNumber="448" unit="mm" value="0.21">0.21 mm</quantity>
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, HW
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<quantity id="A2143DA0FFF8FFCD47B8FD60FABCFD37" box="[1210,1309,741,764]" metricMagnitude="-4" metricUnit="m" metricValue="2.3" pageId="1" pageNumber="448" unit="mm" value="0.23">0.23 mm</quantity>
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; tempora in lateral and dorsal view as long as eyes; vertex strongly impressed at middle, strongly transverse; frons confluent with vertex, with shallow median longitudinal impression; supraantennal tubercles weakly marked. Punctures on vertex obscured by bristles, those on frons very fine, inconspicuous. Vestiture of head dorsum (
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<figureCitation id="FDD78CC0FFF8FFCD4765FCC8FB69FCAF" box="[1127,1224,845,868]" captionStart="FIGURES 1–7" captionStartId="2.[151,258,1962,1984]" captionTargetBox="[151,1436,315,1938]" captionTargetId="figure@2.[151,1436,298,1938]" captionTargetPageId="2" captionText="FIGURES 1–7. Siamites sarawakensis sp. n., dorsal habitus of holotype male (1); head in anterior view (2); aedeagus in ventral (3, 5) and lateral (4, 6) views; and distribution of nominal Siamites species illustrated with their aedeagi in ventral view (7)." httpUri="https://zenodo.org/record/2618914/files/figure.png" pageId="1" pageNumber="448">Figs 1–2</figureCitation>
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) composed of four large tufts of dense bristles forming a transverse arc on vertex, lateral pairs are so close to each other that appear as one transverse group of bristles, median asetose area between bristles relatively broad; additionally vertex at each side and tempora with long and sparse erect bristles flanking the tufts, and frons with thin, sparse and suberect setae. Antennae (
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<figureCitation id="FDD78CC0FFF8FFCD439DFC5DFF44FC24" box="[159,229,984,1007]" captionStart="FIGURES 1–7" captionStartId="2.[151,258,1962,1984]" captionTargetBox="[151,1436,315,1938]" captionTargetId="figure@2.[151,1436,298,1938]" captionTargetPageId="2" captionText="FIGURES 1–7. Siamites sarawakensis sp. n., dorsal habitus of holotype male (1); head in anterior view (2); aedeagus in ventral (3, 5) and lateral (4, 6) views; and distribution of nominal Siamites species illustrated with their aedeagi in ventral view (7)." httpUri="https://zenodo.org/record/2618914/files/figure.png" pageId="1" pageNumber="448">Fig. 1</figureCitation>
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) short, with strongly broadened tetramerous club, AnL
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||
<quantity id="A2143DA0FFF8FFCD4079FC5DFC44FC24" box="[891,997,984,1007]" metricMagnitude="-4" metricUnit="m" metricValue="4.5" pageId="1" pageNumber="448" unit="mm" value="0.45">0.45 mm</quantity>
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; scape and pedicel strongly elongate; antennomeres III–V and VII each about as long as broad, VI indistinctly elongate, VIII–X each strongly transverse, XI slightly narrower than X, slightly longer than broad.
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</paragraph>
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<paragraph id="65539045FFF8FFCD43C5FBC5FB40FB74" blockId="1.[151,1437,429,2012]" pageId="1" pageNumber="448">
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Pronotum semioval, broadest in posterior third but indistinctly narrowing posterad; PL
|
||
<quantity id="A2143DA0FFF8FFCD478CFBC5FB53FB9C" box="[1166,1266,1088,1111]" metricMagnitude="-4" metricUnit="m" metricValue="3.0" pageId="1" pageNumber="448" unit="mm" value="0.3">0.30 mm</quantity>
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, PW
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<quantity id="A2143DA0FFF8FFCD4630FBC5FA39FB9C" box="[1330,1432,1088,1111]" metricMagnitude="-4" metricUnit="m" metricValue="3.0" pageId="1" pageNumber="448" unit="mm" value="0.3">0.30 mm</quantity>
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||
. Anterior and posterior margins weakly arcuate; sides nearly evenly rounded; pronotum with distinct carinate but relatively blunt lateral edges in posterior 3/4 and arcuate transverse ante-basal groove distinctly deepened at each end. Punctures on pronotum fine and sparse, unremarkable; setae sparse, suberect to erect, moderately long.
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</paragraph>
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<paragraph id="65539045FFF8FFCD43C5FB4FFCB1FA82" blockId="1.[151,1437,429,2012]" pageId="1" pageNumber="448">
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Elytra slightly flattened, oval, broadest distinctly in front of middle; EL
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||
<quantity id="A2143DA0FFF8FFCD40D4FB4FFB98FB2A" box="[982,1081,1226,1249]" metricMagnitude="-4" metricUnit="m" metricValue="6.5" pageId="1" pageNumber="448" unit="mm" value="0.65">0.65 mm</quantity>
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, EW
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<quantity id="A2143DA0FFF8FFCD4774FB4FFB78FB2A" box="[1142,1241,1226,1249]" metricMagnitude="-4" metricUnit="m" metricValue="4.5" pageId="1" pageNumber="448" unit="mm" value="0.45">0.45 mm</quantity>
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, EI 1.44; humeral calli weakly marked, each with indistinct humeral carina not exceeding 1/4 of elytral length; basal impressions shallow and short; apices of elytra separately rounded. Punctures on elytra slightly more distinct than those on pronotum but superficial and inconspicuous; setae sparse, short, suberect.
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</paragraph>
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<paragraph id="65539045FFF8FFCD43C5FAD0FBFEFA7A" blockId="1.[151,1437,429,2012]" pageId="1" pageNumber="448">
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Aedeagus (
|
||
<figureCitation id="FDD78CC0FFF8FFCD4240FAD0FE04FAA7" box="[322,421,1365,1388]" captionStart="FIGURES 1–7" captionStartId="2.[151,258,1962,1984]" captionTargetBox="[151,1436,315,1938]" captionTargetId="figure@2.[151,1436,298,1938]" captionTargetPageId="2" captionText="FIGURES 1–7. Siamites sarawakensis sp. n., dorsal habitus of holotype male (1); head in anterior view (2); aedeagus in ventral (3, 5) and lateral (4, 6) views; and distribution of nominal Siamites species illustrated with their aedeagi in ventral view (7)." httpUri="https://zenodo.org/record/2618914/files/figure.png" pageId="1" pageNumber="448">Figs 3–6</figureCitation>
|
||
) stout and small; AeL
|
||
<quantity id="A2143DA0FFF8FFCD41ACFAD0FCB5FAA7" box="[686,788,1365,1388]" metricMagnitude="-5" metricUnit="m" metricValue="9.0" pageId="1" pageNumber="448" unit="mm" value="0.09">0.09 mm</quantity>
|
||
; median lobe pear-shaped, broadest near distal third, with sinuate sides and rounded apical margin; internal sclerotization simple, with subapical endophallic plate-like structure, its sides and distal margin concave; parameres short and broad, each with five apical setae.
|
||
</paragraph>
|
||
<paragraph id="65539045FFF8FFCD43C5FA38FE32FA1F" blockId="1.[151,1437,429,2012]" box="[199,403,1469,1492]" pageId="1" pageNumber="448">Female. Unknown.</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="2DF6C3CEFFF8FFCD43C5FA65FD62FA3C" box="[199,707,1504,1527]" pageId="1" pageNumber="448" type="distribution">
|
||
<paragraph id="65539045FFF8FFCD43C5FA65FD62FA3C" blockId="1.[151,1437,429,2012]" box="[199,707,1504,1527]" pageId="1" pageNumber="448">
|
||
<emphasis id="57984C57FFF8FFCD43C5FA65FEF8FA3C" bold="true" box="[199,345,1504,1527]" pageId="1" pageNumber="448">Distribution.</emphasis>
|
||
Central-northern Borneo (
|
||
<figureCitation id="FDD78CC0FFF8FFCD4177FA65FD17FA3C" box="[629,694,1504,1527]" captionStart="FIGURES 1–7" captionStartId="2.[151,258,1962,1984]" captionTargetBox="[151,1436,315,1938]" captionTargetId="figure@2.[151,1436,298,1938]" captionTargetPageId="2" captionText="FIGURES 1–7. Siamites sarawakensis sp. n., dorsal habitus of holotype male (1); head in anterior view (2); aedeagus in ventral (3, 5) and lateral (4, 6) views; and distribution of nominal Siamites species illustrated with their aedeagi in ventral view (7)." httpUri="https://zenodo.org/record/2618914/files/figure.png" pageId="1" pageNumber="448">Fig. 7</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="2DF6C3CEFFF8FFCD43C5F987FCC7F9D2" box="[199,870,1538,1561]" pageId="1" pageNumber="448" type="etymology">
|
||
<paragraph id="65539045FFF8FFCD43C5F987FCC7F9D2" blockId="1.[151,1437,429,2012]" box="[199,870,1538,1561]" pageId="1" pageNumber="448">
|
||
<emphasis id="57984C57FFF8FFCD43C5F987FEE6F9D2" bold="true" box="[199,327,1538,1561]" pageId="1" pageNumber="448">Etymology.</emphasis>
|
||
Locotypical, after the Malaysian state of
|
||
<collectingCountry id="1DFBD0D5FFF8FFCD4000F987FCC0F9D2" box="[770,865,1538,1561]" name="Malaysia" pageId="1" pageNumber="448">Sarawak</collectingCountry>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="2DF6C3CEFFF8FFCE43C5F9A0FCABFF3F" lastPageId="2" lastPageNumber="449" pageId="1" pageNumber="448" type="discussion">
|
||
<paragraph id="65539045FFF8FFCD43C5F9A0FABDF89A" blockId="1.[151,1437,429,2012]" pageId="1" pageNumber="448">
|
||
<emphasis id="57984C57FFF8FFCD43C5F9A0FE94F9F7" bold="true" box="[199,309,1573,1596]" pageId="1" pageNumber="448">Remarks.</emphasis>
|
||
Adults of
|
||
<taxonomicName id="A2ECEBC6FFF8FFCD42AEF9A0FDA7F9F7" authorityName="Franz" authorityYear="1989" box="[428,518,1573,1596]" class="Insecta" family="Staphylinidae" genus="Siamites" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="448" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="57984C57FFF8FFCD42AEF9A0FDA7F9F7" box="[428,518,1573,1596]" italics="true" pageId="1" pageNumber="448">Siamites</emphasis>
|
||
</taxonomicName>
|
||
species are morphologically relatively uniform and only males of
|
||
<taxonomicName id="A2ECEBC6FFF8FFCD47DCF9A0FAD2F9F7" authorityName="Jaloszynski" authorityYear="2017" box="[1246,1395,1573,1596]" class="Insecta" family="Staphylinidae" genus="Siamites" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="448" phylum="Arthropoda" rank="species" species="deharvengi">
|
||
<emphasis id="57984C57FFF8FFCD47DCF9A0FAD2F9F7" box="[1246,1395,1573,1596]" italics="true" pageId="1" pageNumber="448">S. deharvengi</emphasis>
|
||
</taxonomicName>
|
||
are conspicuous by having vestigial eyes, whereas males of all remaining species are macrophthalmous. Consequently, the aedeagus must be examined to identify species, and male genital organs within this small genus show greater diversity of shapes than externally visible structures of adults (
|
||
<figureCitation id="FDD78CC0FFF8FFCD41AFF908FD4CF96F" box="[685,749,1677,1700]" captionStart="FIGURES 1–7" captionStartId="2.[151,258,1962,1984]" captionTargetBox="[151,1436,315,1938]" captionTargetId="figure@2.[151,1436,298,1938]" captionTargetPageId="2" captionText="FIGURES 1–7. Siamites sarawakensis sp. n., dorsal habitus of holotype male (1); head in anterior view (2); aedeagus in ventral (3, 5) and lateral (4, 6) views; and distribution of nominal Siamites species illustrated with their aedeagi in ventral view (7)." httpUri="https://zenodo.org/record/2618914/files/figure.png" pageId="1" pageNumber="448">Fig. 7</figureCitation>
|
||
). The aedeagi of
|
||
<taxonomicName id="A2ECEBC6FFF8FFCD40A6F908FC5BF96F" authorityName="Franz" authorityYear="1989" box="[932,1018,1677,1700]" class="Insecta" family="Staphylinidae" genus="Siamites" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="448" phylum="Arthropoda" rank="species" species="loebli">
|
||
<emphasis id="57984C57FFF8FFCD40A6F908FC5BF96F" box="[932,1018,1677,1700]" italics="true" pageId="1" pageNumber="448">S. loebli</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="A2ECEBC6FFF8FFCD4705F908FB38F96F" authorityName="Jaloszynski" authorityYear="2017" box="[1031,1177,1677,1700]" class="Insecta" family="Staphylinidae" genus="Siamites" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="448" phylum="Arthropoda" rank="species" species="deharvengi">
|
||
<emphasis id="57984C57FFF8FFCD4705F908FB38F96F" box="[1031,1177,1677,1700]" italics="true" pageId="1" pageNumber="448">S. deharvengi</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="A2ECEBC6FFF8FFCD47CEF908FAD5F96F" authorityName="Jałoszyński" authorityYear="2019" box="[1228,1396,1677,1700]" class="Insecta" family="Staphylinidae" genus="Siamites" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="448" phylum="Arthropoda" rank="species" species="sarawakensis">
|
||
<emphasis id="57984C57FFF8FFCD47CEF908FAD5F96F" box="[1228,1396,1677,1700]" italics="true" pageId="1" pageNumber="448">S. sarawakensis</emphasis>
|
||
</taxonomicName>
|
||
are relatively simple, but they clearly differ in ventral view in the shape of median lobe and endophallic structures. The aedeagus of
|
||
<taxonomicName id="A2ECEBC6FFF8FFCD421FF957FE18F922" authorityName="Jaloszynski" authorityYear="2013" box="[285,441,1746,1769]" class="Insecta" family="Staphylinidae" genus="Siamites" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="448" phylum="Arthropoda" rank="species" species="complicatus">
|
||
<emphasis id="57984C57FFF8FFCD421FF957FE18F922" box="[285,441,1746,1769]" italics="true" pageId="1" pageNumber="448">S. complicatus</emphasis>
|
||
</taxonomicName>
|
||
is strikingly complex, with a pair of distolateral corkscrew-like projections and a very large, darkly sclerotized endophallus; none of the remaining species has similar structures. The aedeagus of
|
||
<taxonomicName id="A2ECEBC6FFF8FFCD4623F970FADCF8C7" authorityName="Franz" authorityYear="1989" box="[1313,1405,1781,1804]" class="Insecta" family="Staphylinidae" genus="Siamites" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="448" phylum="Arthropoda" rank="species" species="loebli">
|
||
<emphasis id="57984C57FFF8FFCD4623F970FADCF8C7" box="[1313,1405,1781,1804]" italics="true" pageId="1" pageNumber="448">S. loebli</emphasis>
|
||
</taxonomicName>
|
||
is conspicuously stout and has a distinctly bisinuate distal margin; the aedeagus of
|
||
<taxonomicName id="A2ECEBC6FFF8FFCD4717F89DFB0AF8E4" authorityName="Jaloszynski" authorityYear="2017" box="[1045,1195,1816,1839]" class="Insecta" family="Staphylinidae" genus="Siamites" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="448" phylum="Arthropoda" rank="species" species="deharvengi">
|
||
<emphasis id="57984C57FFF8FFCD4717F89DFB0AF8E4" box="[1045,1195,1816,1839]" italics="true" pageId="1" pageNumber="448">S. deharvengi</emphasis>
|
||
</taxonomicName>
|
||
is more slender, with distal margin truncated; and the aedeagus of
|
||
<taxonomicName id="A2ECEBC6FFF8FFCD4176F8BFFCBCF89A" authorityName="Jałoszyński" authorityYear="2019" box="[628,797,1850,1873]" class="Insecta" family="Staphylinidae" genus="Siamites" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="448" phylum="Arthropoda" rank="species" species="sarawakensis">
|
||
<emphasis id="57984C57FFF8FFCD4176F8BFFCBCF89A" box="[628,797,1850,1873]" italics="true" pageId="1" pageNumber="448">S. sarawakensis</emphasis>
|
||
</taxonomicName>
|
||
is drop-shaped, with the distal margin rounded.
|
||
</paragraph>
|
||
<paragraph id="65539045FFF8FFCD43C5F8D8FAC6F85C" blockId="1.[151,1437,429,2012]" pageId="1" pageNumber="448">
|
||
<taxonomicName id="A2ECEBC6FFF8FFCD43C5F8D8FE19F8BF" authorityName="Jałoszyński" authorityYear="2019" box="[199,440,1885,1908]" class="Insecta" family="Staphylinidae" genus="Siamites" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="448" phylum="Arthropoda" rank="species" species="sarawakensis">
|
||
<emphasis id="57984C57FFF8FFCD43C5F8D8FE19F8BF" box="[199,440,1885,1908]" italics="true" pageId="1" pageNumber="448">Siamites sarawakensis</emphasis>
|
||
</taxonomicName>
|
||
was found over
|
||
<quantity id="A2143DA0FFF8FFCD416FF8D8FD6AF8BF" box="[621,715,1885,1908]" metricMagnitude="6" metricUnit="m" metricValue="1.2" pageId="1" pageNumber="448" unit="km" value="1200.0">1200 km</quantity>
|
||
southeast of the collecting site of
|
||
<taxonomicName id="A2ECEBC6FFF8FFCD473CF8D8FB70F8BF" authorityName="Jaloszynski" authorityYear="2017" box="[1086,1233,1885,1908]" class="Insecta" family="Staphylinidae" genus="Siamites" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="448" phylum="Arthropoda" rank="species" species="deharvengi">
|
||
<emphasis id="57984C57FFF8FFCD473CF8D8FB70F8BF" box="[1086,1233,1885,1908]" italics="true" pageId="1" pageNumber="448">S. deharvengi</emphasis>
|
||
</taxonomicName>
|
||
and over
|
||
<quantity id="A2143DA0FFF8FFCD463FF8D8FA3DF8BF" box="[1341,1436,1885,1908]" metricMagnitude="6" metricUnit="m" metricValue="1.5" pageId="1" pageNumber="448" unit="km" value="1500.0">1500 km</quantity>
|
||
SE of the collecting site of
|
||
<taxonomicName id="A2ECEBC6FFF8FFCD42BBF805FDF4F85C" authorityName="Jaloszynski" authorityYear="2013" box="[441,597,1920,1943]" class="Insecta" family="Staphylinidae" genus="Siamites" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="448" phylum="Arthropoda" rank="species" species="complicatus">
|
||
<emphasis id="57984C57FFF8FFCD42BBF805FDF4F85C" box="[441,597,1920,1943]" italics="true" pageId="1" pageNumber="448">S. complicatus</emphasis>
|
||
</taxonomicName>
|
||
; this discovery significantly broadens the known distribution of
|
||
<taxonomicName id="A2ECEBC6FFF8FFCD4604F805FAC1F85C" authorityName="Franz" authorityYear="1989" box="[1286,1376,1920,1943]" class="Insecta" family="Staphylinidae" genus="Siamites" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="448" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="57984C57FFF8FFCD4604F805FAC1F85C" box="[1286,1376,1920,1943]" italics="true" pageId="1" pageNumber="448">Siamites</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="65539045FFF8FFCE43C5F827FCABFF3F" blockId="1.[151,1437,429,2012]" lastBlockId="2.[151,1436,152,244]" lastPageId="2" lastPageNumber="449" pageId="1" pageNumber="448">
|
||
A variously distinctly delimited trimerous club was included in the diagnosis of
|
||
<taxonomicName id="A2ECEBC6FFF8FFCD4726F827FBDFF872" authorityName="Franz" authorityYear="1989" box="[1060,1150,1954,1977]" class="Insecta" family="Staphylinidae" genus="Siamites" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="448" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="57984C57FFF8FFCD4726F827FBDFF872" box="[1060,1150,1954,1977]" italics="true" pageId="1" pageNumber="448">Siamites</emphasis>
|
||
</taxonomicName>
|
||
by
|
||
<bibRefCitation id="017DEDB4FFF8FFCD47A5F827FAD5F872" author="Jaloszynski, P." box="[1191,1396,1954,1977]" pageId="1" pageNumber="448" pagination="385 - 390" refId="ref1682" refString="Jaloszynski, P. (2013) A new species of Siamites Franz (Coleoptera, Staphylinidae, Scydmaeninae) from Thailand. Zootaxa, 3734 (3), 385 - 390. https: // doi. org / 10.11646 / zootaxa. 3734.3.8" type="journal article" year="2013">Jałoszyński (2013)</bibRefCitation>
|
||
. As the antennomere VIII in all species is broader than VII and narrower than IX, the club can be rather subjectively interpreted as tri- or tetramerous, depending on the width difference between antennomeres VII and VIII. In
|
||
<taxonomicName id="A2ECEBC6FFFBFFCE468AFF1DFE86FF1A" authorityName="Jałoszyński" authorityYear="2019" class="Insecta" family="Staphylinidae" genus="Siamites" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="449" phylum="Arthropoda" rank="species" species="sarawakensis">
|
||
<emphasis id="57984C57FFFBFFCE468AFF1DFE86FF1A" italics="true" pageId="2" pageNumber="449">S. sarawakensis</emphasis>
|
||
</taxonomicName>
|
||
the club is relatively strongly broadened and clearly tetramerous; the diagnosis of
|
||
<taxonomicName id="A2ECEBC6FFFBFFCE4792FF3FFB4BFF1A" authorityName="Franz" authorityYear="1989" box="[1168,1258,186,209]" class="Insecta" family="Staphylinidae" genus="Siamites" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="449" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="57984C57FFFBFFCE4792FF3FFB4BFF1A" box="[1168,1258,186,209]" italics="true" pageId="2" pageNumber="449">Siamites</emphasis>
|
||
</taxonomicName>
|
||
is here emended to include species with tri- and tetramerous antennal clubs.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |