treatments-xml/data/E7/20/6B/E7206B3ED33D716193E8201E2751DF25.xml
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<document ID-DOI="http://dx.doi.org/10.3897/zookeys.78.800" ID-GBIF-Dataset="824a277f-3e29-43cb-b3b4-797aa5895c03" ID-PMC="PMC3088061" ID-Pensoft-Pub="1313-2970-78-15" ID-PubMed="21594154" ModsDocAuthor="" ModsDocDate="2011" ModsDocID="1313-2970-78-15" ModsDocOrigin="ZooKeys 78" ModsDocTitle="On the identity of the fossil aquatic beetles from the Tertiary localities in the southern part of the Upper Rhine Graben (Coleoptera, Hydrophilidae, Dytiscidae)" checkinTime="1451250817608" checkinUser="pensoft" docAuthor="Martin, Fikacek, Hajek, Jiri &amp; Schmied, Heiko" docDate="2011" docId="E7206B3ED33D716193E8201E2751DF25" docLanguage="en" docName="ZooKeys 78: 15-25" docOrigin="ZooKeys 78" docSource="http://dx.doi.org/10.3897/zookeys.78.800" docTitle="Escheria punctulata Foerster 1891" docType="treatment" docVersion="4" lastPageNumber="20" masterDocId="0E71FFE5A5468A341831690CFF9AFFD7" masterDocTitle="On the identity of the fossil aquatic beetles from the Tertiary localities in the southern part of the Upper Rhine Graben (Coleoptera, Hydrophilidae, Dytiscidae)" masterLastPageNumber="25" masterPageNumber="15" pageNumber="20" updateTime="1668166156320" updateUser="ExternalLinkService">
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<mods:title>On the identity of the fossil aquatic beetles from the Tertiary localities in the southern part of the Upper Rhine Graben (Coleoptera, Hydrophilidae, Dytiscidae)</mods:title>
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<mods:name type="personal">
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Martin, Fikacek</mods:namePart>
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Hajek, Jiri</mods:namePart>
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<mods:name type="personal">
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Schmied, Heiko</mods:namePart>
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<mods:title>ZooKeys</mods:title>
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<mods:date>2011</mods:date>
<mods:detail type="volume">
<mods:number>78</mods:number>
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<mods:extent unit="page">
<mods:start>15</mods:start>
<mods:end>25</mods:end>
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<mods:url>http://dx.doi.org/10.3897/zookeys.78.800</mods:url>
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<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.78.800</mods:identifier>
<mods:identifier type="Pensoft-Pub">1313-2970-78-15</mods:identifier>
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<treatment ID-GBIF-Taxon="159361186" LSID="urn:lsid:plazi:treatment:E7206B3ED33D716193E8201E2751DF25" httpUri="http://treatment.plazi.org/id/E7206B3ED33D716193E8201E2751DF25" lastPageId="7" lastPageNumber="20" pageId="5" pageNumber="20">
<subSubSection pageId="5" pageNumber="20" type="nomenclature">
<paragraph pageId="5" pageNumber="20">
<taxonomicName authority="Foerster, 1891" authorityName="Foerster" authorityYear="1891" class="Insecta" family="Hydrophilidae" genus="Escheria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Escheria punctulata" order="Coleoptera" pageId="5" pageNumber="20" phylum="Arthropoda" rank="species" species="punctulata">
Escheria punctulata
<normalizedToken originalValue="Förster">Foerster</normalizedToken>
, 1891
</taxonomicName>
Fig. 9
</paragraph>
</subSubSection>
<subSubSection pageId="5" pageNumber="20" type="reference_group">
<paragraph pageId="5" pageNumber="20">
<taxonomicName class="Insecta" family="Hydrophilidae" genus="Escheria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Escheria punctulata" order="Coleoptera" pageId="5" pageNumber="20" phylum="Arthropoda" rank="species" species="punctulata">Escheria punctulata</taxonomicName>
<bibRefCitation author="Foerster, B" journalOrPublisher="Abhandlungen zur geologischen Spezialkarte von Elsass-Lothringen" pageId="7" pageNumber="22" pagination="333 - 594" title="Die Insekten des &quot; Plattigen Steinmergels &quot; von Brunnstatt" volume="5" year="1891">
<normalizedToken originalValue="Förster">Foerster</normalizedToken>
1891
</bibRefCitation>
: 361; plate XI, Figs 8a,b (original description from Brunstatt);
<bibRefCitation author="Handlirsch, A" journalOrPublisher="Wilhelm Engelmann, Leipzig" pageId="8" pageNumber="23" title="Die fossilen Insekten und die Phylogenie der rezenten Formen" volumeTitle="Ein Handbuch fuer Palaeontologen und Zoologen" year="1908">Handlirsch 1908</bibRefCitation>
: 767 (catalogue).
</paragraph>
<paragraph pageId="5" pageNumber="20">
<taxonomicName class="Insecta" family="Hydrophilidae" genus="Hydrobius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hydrobius punctulatus" order="Coleoptera" pageId="5" pageNumber="20" phylum="Arthropoda" rank="species" species="punctulatus">Hydrobius punctulatus</taxonomicName>
:
<bibRefCitation author="Theobald, N" journalOrPublisher="G. Thomas, Nancy" pageId="8" pageNumber="23" title="Les insectes fossiles des terrains oligocenes de France" year="1937">
<normalizedToken originalValue="Théobald">Theobald</normalizedToken>
1937
</bibRefCitation>
: 169 (transferred to
<taxonomicName class="Insecta" family="Hydrophilidae" genus="Hydrobius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hydrobius" order="Coleoptera" pageId="5" pageNumber="20" phylum="Arthropoda" rank="genus">Hydrobius</taxonomicName>
);
<bibRefCitation author="Hansen, M" journalOrPublisher="Apollo Books, Stenstrup" pageId="8" pageNumber="23" title="World Catalogue of Insects. Volume 2. Hydrophiloidea (s. str.) (Coleoptera)" year="1999">Hansen 1999</bibRefCitation>
: 319 (catalogue).
</paragraph>
</subSubSection>
<subSubSection lastPageId="7" lastPageNumber="22" pageId="5" pageNumber="20" type="taxonomic note">
<paragraph pageId="5" pageNumber="20">Taxonomic note.</paragraph>
<paragraph lastPageId="7" lastPageNumber="22" pageId="5" pageNumber="20">
Based on the drawing by
<bibRefCitation author="Foerster, B" journalOrPublisher="Abhandlungen zur geologischen Spezialkarte von Elsass-Lothringen" pageId="7" pageNumber="22" pagination="333 - 594" title="Die Insekten des &quot; Plattigen Steinmergels &quot; von Brunnstatt" volume="5" year="1891">
<normalizedToken originalValue="Förster">Foerster</normalizedToken>
(1891)
</bibRefCitation>
, the ventral morphology of this species agrees with that of
<taxonomicName family="Hydrophilidae" lsidName="" pageId="5" pageNumber="20" rank="family">Hydrophilidae</taxonomicName>
:
<taxonomicName lsidName="" pageId="5" pageNumber="20" rank="subFamily" subFamily="Hydrophilinae">Hydrophilinae</taxonomicName>
in many characters: (i) mesocoxae transverse and very narrowly separated, (ii) mesepimeron well separated, triangular, (iii) metanepisternum rather narrow; (iv) abdomen with five ventrites.
<pageBreakToken pageId="6" pageNumber="21" start="start">However</pageBreakToken>
, none of these characters or their combination is unique for the
<taxonomicName class="Magnoliopsida" family="Malvaceae" genus="Hydrophiloidea" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="Hydrophiloidea" order="Malvales" pageId="6" pageNumber="21" phylum="Tracheophyta" rank="genus">Hydrophiloidea</taxonomicName>
and may be found in other beetle families as well (see
<bibRefCitation author="Lawrence, JF" journalOrPublisher="CSIRO Publishing, Melbourne" pageId="8" pageNumber="23" title="Beetles of the world. A key and information system for families and subfamilies. CD-ROM, Version 1.0 for MS-Windows" year="1999">Lawrence et al. 1999</bibRefCitation>
). Moreover, two characters illustrated on the drawing and/or mentioned in the original description exclude the placement of
<taxonomicName class="Insecta" family="Hydrophilidae" genus="Escheria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Escheria punctulata" order="Coleoptera" pageId="6" pageNumber="21" phylum="Arthropoda" rank="species" species="punctulata">Escheria punctulata</taxonomicName>
in the
<taxonomicName class="Magnoliopsida" family="Malvaceae" genus="Hydrophiloidea" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="Hydrophiloidea" order="Malvales" pageId="6" pageNumber="21" phylum="Tracheophyta" rank="genus">Hydrophiloidea</taxonomicName>
: (i) elytra bear only 6 deeply impressed striae [9-11 striae are present in all
<taxonomicName class="Magnoliopsida" family="Malvaceae" genus="Hydrophiloidea" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="Hydrophiloidea" order="Malvales" pageId="6" pageNumber="21" phylum="Tracheophyta" rank="genus">Hydrophiloidea</taxonomicName>
with striate elytra, only rarely is the number of series higher but in such cases they are never impressed to striae]; (ii) mesoventrite fused with mesepisternal (i.e. not separated from them by sutures) [in
<taxonomicName class="Magnoliopsida" family="Malvaceae" genus="Hydrophiloidea" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="Hydrophiloidea" order="Malvales" pageId="6" pageNumber="21" phylum="Tracheophyta" rank="genus">Hydrophiloidea</taxonomicName>
, the mesoventrite is fused to mesepisterna only in derived groups of the
<taxonomicName lsidName="" pageId="6" pageNumber="21" rank="subFamily" subFamily="Sphaeridiinae">Sphaeridiinae</taxonomicName>
which are characterized by a highly elevated median portion of the mesoventrite; the elevated median elevation is missing from the fossil]. For these reasons,
<taxonomicName class="Insecta" family="Hydrophilidae" genus="Escheria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Escheria punctulata" order="Coleoptera" pageId="6" pageNumber="21" phylum="Arthropoda" rank="species" species="punctulata">Escheria punctulata</taxonomicName>
is removed from the fossil record of
<pageBreakToken pageId="7" pageNumber="22" start="start">the</pageBreakToken>
<taxonomicName class="Magnoliopsida" family="Malvaceae" genus="Hydrophiloidea" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="Hydrophiloidea" order="Malvales" pageId="7" pageNumber="22" phylum="Tracheophyta" rank="genus">Hydrophiloidea</taxonomicName>
and is placed into
<taxonomicName class="Insecta" family="Corydiidae" genus="Polyphaga" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Polyphaga" order="Blattodea" pageId="7" pageNumber="22" phylum="Arthropoda" rank="genus">Polyphaga</taxonomicName>
incertae sedis; its family placement remains unclear.
</paragraph>
</subSubSection>
</treatment>
</document>