230 lines
32 KiB
XML
230 lines
32 KiB
XML
<document id="A4039CF07E63AB1CDB438833C67ED410" ID-CLB-Dataset="27702" ID-DOI="10.11646/zootaxa.4543.2.7" ID-GBIF-Dataset="c5d46c52-27c5-4b8d-a9d3-b4b3ff22de8f" ID-ISSN="1175-5326" ID-Zenodo-Dep="2617833" ID-ZooBank="0CDAB733-EA1B-4793-873D-609CC50F83B4" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1553928371931" checkinUser="plazi" docAuthor="Simone, Luiz Ricardo L." docDate="2019" docId="E37222277575FF8EE3C41FA5FE9DF9BD" docLanguage="en" docName="zootaxa.4543.2.7.pdf" docOrigin="Zootaxa 4543 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Habeastrum omphalium Simone 2019, new species" docType="treatment" docUuid="524FD91C-2855-40F8-8419-747E459DB3D1" docUuidSource="ZooBank" docVersion="9" lastPageNumber="288" masterDocId="1F4B5A5F7574FF8FE3531D41FF92FFE2" masterDocTitle="The new genus Habeastrum, with two new species (Gastropoda, Diplommatinidae) in Mato Grosso do Sul caves, Brazil" masterLastPageNumber="290" masterPageNumber="287" pageNumber="288" updateTime="1698715407212" updateUser="ExternalLinkService">
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<mods:title id="0946E2E179ECFB0DEC98113F30F46CB6">The new genus Habeastrum, with two new species (Gastropoda, Diplommatinidae) in Mato Grosso do Sul caves, Brazil</mods:title>
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<mods:name id="EC44787692123BBCA4E9D572394A9D49" type="personal">
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<mods:roleTerm id="328367E0BC28D66C740AB809302234B9">Author</mods:roleTerm>
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<mods:namePart id="AD206635721218DEDB86308F8808260F">Simone, Luiz Ricardo L.</mods:namePart>
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<mods:title id="42D973E8F3A94FA8413D9B8654270038">Zootaxa</mods:title>
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<mods:part id="663EEE0108C0F6ED7E051FBBDEF86A4E">
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<mods:date id="53583E31434EDCB971C193B99050AE61">2019</mods:date>
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<mods:number id="6CB91B06C2187F7ABDB2349033128C48">2019-01-07</mods:number>
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<mods:number id="5F1178F55785977E03435646C9A3E638">4543</mods:number>
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<mods:identifier id="4A20A796E916B90DFAAF33A34499EC4B" type="DOI">10.11646/zootaxa.4543.2.7</mods:identifier>
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<treatment id="E37222277575FF8EE3C41FA5FE9DF9BD" ID-DOI="http://doi.org/10.5281/zenodo.5941069" ID-GBIF-Taxon="154529779" ID-Zenodo-Dep="5941069" LSID="urn:lsid:zoobank.org:act:524FD91C-2855-40F8-8419-747E459DB3D1" httpUri="http://treatment.plazi.org/id/E37222277575FF8EE3C41FA5FE9DF9BD" lastPageNumber="288" pageId="1" pageNumber="288">
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<subSubSection id="23C1C0BA7575FF8EE3C41FA5FC02FCFD" pageId="1" pageNumber="288" type="nomenclature">
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<paragraph id="6B6493317575FF8EE3C41FA5FC8DFD1D" blockId="1.[151,1437,740,1631]" box="[151,799,740,767]" pageId="1" pageNumber="288">
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<heading id="302C245D7575FF8EE3C41FA5FC8DFD1D" bold="true" box="[151,799,740,767]" fontSize="11" level="1" pageId="1" pageNumber="288" reason="1">
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<emphasis id="59AF4F237575FF8EE3C41FA5FC8DFD1D" bold="true" box="[151,799,740,767]" pageId="1" pageNumber="288">
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<taxonomicName id="ACDBE8B27575FF8EE3C41FA5FE2DFD1C" ID-CoL="3J3L2" authority="Simone, 2019" authorityName="Simone" authorityYear="2019" box="[151,447,740,766]" class="Gastropoda" family="Diplommatinidae" genus="Habeastrum" higherTaxonomySource="GBIF" kingdom="Animalia" order="Architaenioglossa" pageId="1" pageNumber="288" phylum="Mollusca" rank="species" species="omphalium" status="sp. nov.">
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<emphasis id="59AF4F237575FF8EE3C41FA5FE2DFD1C" bold="true" box="[151,447,740,766]" italics="true" pageId="1" pageNumber="288">Habeastrum omphalium</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="429CF2587575FF8EE2961FA4FDC7FD1D" box="[453,597,741,767]" pageId="1" pageNumber="288" rank="species">new species</taxonomicNameLabel>
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(
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<figureCitation id="F3E08FB47575FF8EE1361FA4FD4AFD1D" box="[613,728,741,767]" captionStart="FIGURE 1" captionStartId="2.[151,244,1877,1899]" captionTargetBox="[150,1436,330,1838]" captionTargetId="figure@2.[150,1436,330,1838]" captionTargetPageId="2" captionText="FIGURE 1. Habeastrum spp. shells. A–J. Habeastrum parafusum, n. sp. A. Holotype MZSP 134301, L 1.4 mm. B–J. Paratypes MZSP 135584, L 1.7 mm. C. L 1.6 mm. D. L 1.6 mm. E. L 1.4 mm. F. SEM, L 1.1 mm. G. L. 1.2 mm. H. same specimen, showing protoconch and first teleoconch whorls, SEM, scale = 0.1 mm. I. same specimen of F, scale = 0.1 mm. J. Young specimen, L 0.6 mm. K–R. H. omphalium n. sp. K–N. Holotype MZSP 135583, W 1.1 mm, L 1.6 mm. O–R. Paratypes MZSP 134324. O. L 1.0 mm. P. SEM, L 1.0 mm. Q–R. Scale = 0.1 mm." httpUri="https://zenodo.org/record/2617835/files/figure.png" pageId="1" pageNumber="288">Figures 1</figureCitation>
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K–R)
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</emphasis>
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</heading>
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</paragraph>
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<paragraph id="6B6493317575FF8EE3C41E49FC02FCFD" blockId="1.[151,1437,740,1631]" box="[151,912,776,799]" pageId="1" pageNumber="288">
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<uri id="1F4A9F337575FF8EE3C41E49FC02FCFD" box="[151,912,776,799]" pageId="1" pageNumber="288">
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urn:lsid:zoobank.org:act:
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<uuid id="1F7DA9E47575FF8EE2CD1E49FC02FCFD" box="[414,912,776,799]" pageId="1" pageNumber="288">524FD91C-2855-40F8-8419-747E459DB3D1</uuid>
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</uri>
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</paragraph>
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||
</subSubSection>
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<subSubSection id="23C1C0BA7575FF8EE3C41E6BFB76FCA3" box="[151,1252,810,833]" pageId="1" pageNumber="288" type="description">
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<paragraph id="6B6493317575FF8EE3C41E6BFB76FCA3" blockId="1.[151,1437,740,1631]" box="[151,1252,810,833]" pageId="1" pageNumber="288">
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<emphasis id="59AF4F237575FF8EE3C41E6BFF4CFCA3" bold="true" box="[151,222,810,833]" pageId="1" pageNumber="288">Types.</emphasis>
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<materialsCitation id="DBB3996C7575FF8EE3B51E6BFDECFCA3" ID-GBIF-Occurrence="2147525213" box="[230,638,810,833]" collectionCode="MZSP" pageId="1" pageNumber="288" specimenCode="MZSP 135583" specimenCount="1" typeStatus="holotype">
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<typeStatus id="B4602D937575FF8EE3B51E6BFEDBFCA3" box="[230,329,810,833]" pageId="1" pageNumber="288" type="holotype">Holotype</typeStatus>
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<specimenCode id="3B7D3B4A7575FF8EE2031E6BFE7EFCA3" box="[336,492,810,833]" collectionCode="MZSP" country="Brazil" httpUri="http://grbio.org/cool/9yp6-zxp9" name="Sao Paulo, Museu de Zoologia da Universidade de Sao Paulo" pageId="1" pageNumber="288" type="Museum">MZSP 135583</specimenCode>
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(
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<figureCitation id="F3E08FB47575FF8EE2A91E6BFDA8FCA3" box="[506,570,810,833]" captionStart="FIGURE 1" captionStartId="2.[151,244,1877,1899]" captionTargetBox="[150,1436,330,1838]" captionTargetId="figure@2.[150,1436,330,1838]" captionTargetPageId="2" captionText="FIGURE 1. Habeastrum spp. shells. A–J. Habeastrum parafusum, n. sp. A. Holotype MZSP 134301, L 1.4 mm. B–J. Paratypes MZSP 135584, L 1.7 mm. C. L 1.6 mm. D. L 1.6 mm. E. L 1.4 mm. F. SEM, L 1.1 mm. G. L. 1.2 mm. H. same specimen, showing protoconch and first teleoconch whorls, SEM, scale = 0.1 mm. I. same specimen of F, scale = 0.1 mm. J. Young specimen, L 0.6 mm. K–R. H. omphalium n. sp. K–N. Holotype MZSP 135583, W 1.1 mm, L 1.6 mm. O–R. Paratypes MZSP 134324. O. L 1.0 mm. P. SEM, L 1.0 mm. Q–R. Scale = 0.1 mm." httpUri="https://zenodo.org/record/2617835/files/figure.png" pageId="1" pageNumber="288">Figs 1</figureCitation>
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K–N)
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</materialsCitation>
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.
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<materialsCitation id="DBB3996C7575FF8EE1DB1E6BFB72FCA3" ID-GBIF-Occurrence="2147525052" box="[648,1248,810,833]" collectionCode="MZSP" pageId="1" pageNumber="288" specimenCode="MZSP 134324, 8" specimenCount="1" typeStatus="paratype">
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<typeStatus id="B4602D937575FF8EE1DB1E6BFD66FCA3" box="[648,756,810,833]" pageId="1" pageNumber="288" type="paratype">Paratypes</typeStatus>
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:
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<specimenCode id="3B7D3B4A7575FF8EE1AE1E6BFC0EFCA3" box="[765,924,810,833]" collectionCode="MZSP" country="Brazil" httpUri="http://grbio.org/cool/9yp6-zxp9" name="Sao Paulo, Museu de Zoologia da Universidade de Sao Paulo" pageId="1" pageNumber="288" type="Museum">MZSP 134324</specimenCode>
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,
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<specimenCode id="3B7D3B4A7575FF8EE0F41E6BFC26FCA3" box="[935,948,810,833]" collectionCode="MZSP" country="Brazil" httpUri="http://grbio.org/cool/9yp6-zxp9" name="Sao Paulo, Museu de Zoologia da Universidade de Sao Paulo" pageId="1" pageNumber="288" type="Museum">8</specimenCode>
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shells all from type locality
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</materialsCitation>
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.
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</paragraph>
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</subSubSection>
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<subSubSection id="23C1C0BA7575FF8EE3941E0CFE2EFC65" pageId="1" pageNumber="288" type="materials_examined">
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<paragraph id="6B6493317575FF8EE3941E0CFE2EFC65" blockId="1.[151,1437,740,1631]" pageId="1" pageNumber="288">
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||
<materialsCitation id="DBB3996C7575FF8EE3941E0CFE25FC65" ID-GBIF-Occurrence="2147525051" collectingDate="1992-07-16" collectorName="Trajano & Gnaspini" country="Brazil" latitude="-21.11028" location="Gruta Pitangueiras" longLatPrecision="21" longitude="-56.581112" municipality="Bonito" pageId="1" pageNumber="288" specimenCount="1" stateProvince="Mato Grosso do Sul" typeStatus="holotype">
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<emphasis id="59AF4F237575FF8EE3941E0CFEF2FC86" bold="true" box="[199,352,845,868]" pageId="1" pageNumber="288">
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<typeStatus id="B4602D937575FF8EE3941E0CFF6DFC86" box="[199,255,845,868]" pageId="1" pageNumber="288">Type</typeStatus>
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locality.
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</emphasis>
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||
<collectingCountry id="13CCD3A17575FF8EE23F1E0CFE42FC86" box="[364,464,845,868]" name="Brazil" pageId="1" pageNumber="288">BRAZIL</collectingCountry>
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.
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<collectingRegion id="A91F5DD37575FF8EE28E1E0CFD53FC86" box="[477,705,845,868]" country="Brazil" name="Mato Grosso do Sul" pageId="1" pageNumber="288">Mato Grosso do Sul</collectingRegion>
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,
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<collectingMunicipality id="8B00094B7575FF8EE1831E0CFC89FC86" box="[720,795,845,868]" pageId="1" pageNumber="288">Bonito</collectingMunicipality>
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,
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<location id="6E04C5EA7575FF8EE0791E0CFC6AFC86" LSID="urn:lsid:plazi:treatment:E37222277575FF8EE3C41FA5FE9DF9BD:6E04C5EA7575FF8EE0791E0CFC6AFC86" box="[810,1016,845,868]" country="Brazil" latitude="-21.11028" longLatPrecision="21" longitude="-56.581112" municipality="Bonito" name="Gruta Pitangueiras" pageId="1" pageNumber="288" stateProvince="Mato Grosso do Sul">Gruta Pitangueiras</location>
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,
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<geoCoordinate id="0EEFF5F67575FF8EE7551E0CFB11FC86" box="[1030,1155,845,868]" direction="south" orientation="latitude" pageId="1" pageNumber="288" precision="15" value="-21.11028">21°06’37”S</geoCoordinate>
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<geoCoordinate id="0EEFF5F67575FF8EE7DD1E0CFA85FC86" box="[1166,1303,845,868]" direction="west" orientation="longitude" pageId="1" pageNumber="288" precision="15" value="-56.581112">56°34’52”W</geoCoordinate>
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(
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<collectorName id="C62EF6E77575FF8EE67A1E0CFAEFFC86" box="[1321,1405,845,868]" pageId="1" pageNumber="288">Trajano</collectorName>
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&
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<collectorName id="C62EF6E77575FF8EE3C41E31FF65FC65" box="[151,247,880,903]" pageId="1" pageNumber="288">Gnaspini</collectorName>
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col.,
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<date id="1F65B5F17575FF8EE2611E31FE3DFC65" box="[306,431,880,903]" pageId="1" pageNumber="288" value="1992-07-16">
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<collectingDate id="0F214C197575FF8EE2611E31FE3DFC65" box="[306,431,880,903]" pageId="1" pageNumber="288" value="1992-07-16">16.vii.1992</collectingDate>
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</date>
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)
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</materialsCitation>
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.
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</paragraph>
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</subSubSection>
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<subSubSection id="23C1C0BA7575FF8EE3941ED3FBBDFC4B" box="[199,1071,914,937]" pageId="1" pageNumber="288" type="diagnosis">
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<paragraph id="6B6493317575FF8EE3941ED3FBBDFC4B" blockId="1.[151,1437,740,1631]" box="[199,1071,914,937]" pageId="1" pageNumber="288">
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<emphasis id="59AF4F237575FF8EE3941ED3FEAEFC4B" bold="true" box="[199,316,914,937]" pageId="1" pageNumber="288">Diagnosis.</emphasis>
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Shell with about 5 whorls growing uniformly widely. Umbilicus wide.
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</paragraph>
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</subSubSection>
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<subSubSection id="23C1C0BA7575FF8EE3941EF4FDEFFB7E" pageId="1" pageNumber="288" type="description">
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||
<paragraph id="6B6493317575FF8EE3941EF4FDEFFB7E" blockId="1.[151,1437,740,1631]" pageId="1" pageNumber="288">
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||
<emphasis id="59AF4F237575FF8EE3941EF4FEC0FC2E" bold="true" box="[199,338,949,972]" pageId="1" pageNumber="288">Description.</emphasis>
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Shell relatively trochiform, conical. Total of about 5 whorls, uniformly widening, spire angle about 50° to 60° (
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<figureCitation id="F3E08FB47575FF8EE3BF1E99FEBCFC0D" box="[236,302,984,1007]" captionStart="FIGURE 1" captionStartId="2.[151,244,1877,1899]" captionTargetBox="[150,1436,330,1838]" captionTargetId="figure@2.[150,1436,330,1838]" captionTargetPageId="2" captionText="FIGURE 1. Habeastrum spp. shells. A–J. Habeastrum parafusum, n. sp. A. Holotype MZSP 134301, L 1.4 mm. B–J. Paratypes MZSP 135584, L 1.7 mm. C. L 1.6 mm. D. L 1.6 mm. E. L 1.4 mm. F. SEM, L 1.1 mm. G. L. 1.2 mm. H. same specimen, showing protoconch and first teleoconch whorls, SEM, scale = 0.1 mm. I. same specimen of F, scale = 0.1 mm. J. Young specimen, L 0.6 mm. K–R. H. omphalium n. sp. K–N. Holotype MZSP 135583, W 1.1 mm, L 1.6 mm. O–R. Paratypes MZSP 134324. O. L 1.0 mm. P. SEM, L 1.0 mm. Q–R. Scale = 0.1 mm." httpUri="https://zenodo.org/record/2617835/files/figure.png" pageId="1" pageNumber="288">Figs 1</figureCitation>
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L–P). Protoconch
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<quantity id="AC233ED47575FF8EE2A81E99FDCEFC0D" box="[507,604,984,1007]" metricMagnitude="-4" metricUnit="m" metricValue="4.2" pageId="1" pageNumber="288" unit="mm" value="0.42">0.42 mm</quantity>
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wide, smooth, except for minute radial wrinkles, translucent (
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<figureCitation id="F3E08FB47575FF8EE65D1E99FA1CFC0D" box="[1294,1422,984,1007]" captionStart="FIGURE 1" captionStartId="2.[151,244,1877,1899]" captionTargetBox="[150,1436,330,1838]" captionTargetId="figure@2.[150,1436,330,1838]" captionTargetPageId="2" captionText="FIGURE 1. Habeastrum spp. shells. A–J. Habeastrum parafusum, n. sp. A. Holotype MZSP 134301, L 1.4 mm. B–J. Paratypes MZSP 135584, L 1.7 mm. C. L 1.6 mm. D. L 1.6 mm. E. L 1.4 mm. F. SEM, L 1.1 mm. G. L. 1.2 mm. H. same specimen, showing protoconch and first teleoconch whorls, SEM, scale = 0.1 mm. I. same specimen of F, scale = 0.1 mm. J. Young specimen, L 0.6 mm. K–R. H. omphalium n. sp. K–N. Holotype MZSP 135583, W 1.1 mm, L 1.6 mm. O–R. Paratypes MZSP 134324. O. L 1.0 mm. P. SEM, L 1.0 mm. Q–R. Scale = 0.1 mm." httpUri="https://zenodo.org/record/2617835/files/figure.png" pageId="1" pageNumber="288">Figs 1O, Q</figureCitation>
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); occupying about 35% of shell width, about 20% of shell length. Shell varying from wide (
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<figureCitation id="F3E08FB47575FF8EE7051EBBFB06FBF3" box="[1110,1172,1018,1041]" captionStart="FIGURE 1" captionStartId="2.[151,244,1877,1899]" captionTargetBox="[150,1436,330,1838]" captionTargetId="figure@2.[150,1436,330,1838]" captionTargetPageId="2" captionText="FIGURE 1. Habeastrum spp. shells. A–J. Habeastrum parafusum, n. sp. A. Holotype MZSP 134301, L 1.4 mm. B–J. Paratypes MZSP 135584, L 1.7 mm. C. L 1.6 mm. D. L 1.6 mm. E. L 1.4 mm. F. SEM, L 1.1 mm. G. L. 1.2 mm. H. same specimen, showing protoconch and first teleoconch whorls, SEM, scale = 0.1 mm. I. same specimen of F, scale = 0.1 mm. J. Young specimen, L 0.6 mm. K–R. H. omphalium n. sp. K–N. Holotype MZSP 135583, W 1.1 mm, L 1.6 mm. O–R. Paratypes MZSP 134324. O. L 1.0 mm. P. SEM, L 1.0 mm. Q–R. Scale = 0.1 mm." httpUri="https://zenodo.org/record/2617835/files/figure.png" pageId="1" pageNumber="288">Figs 1</figureCitation>
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L–N) to very wide (
|
||
<figureCitation id="F3E08FB47575FF8EE63F1EBBFF45FBD6" captionStart="FIGURE 1" captionStartId="2.[151,244,1877,1899]" captionTargetBox="[150,1436,330,1838]" captionTargetId="figure@2.[150,1436,330,1838]" captionTargetPageId="2" captionText="FIGURE 1. Habeastrum spp. shells. A–J. Habeastrum parafusum, n. sp. A. Holotype MZSP 134301, L 1.4 mm. B–J. Paratypes MZSP 135584, L 1.7 mm. C. L 1.6 mm. D. L 1.6 mm. E. L 1.4 mm. F. SEM, L 1.1 mm. G. L. 1.2 mm. H. same specimen, showing protoconch and first teleoconch whorls, SEM, scale = 0.1 mm. I. same specimen of F, scale = 0.1 mm. J. Young specimen, L 0.6 mm. K–R. H. omphalium n. sp. K–N. Holotype MZSP 135583, W 1.1 mm, L 1.6 mm. O–R. Paratypes MZSP 134324. O. L 1.0 mm. P. SEM, L 1.0 mm. Q–R. Scale = 0.1 mm." httpUri="https://zenodo.org/record/2617835/files/figure.png" pageId="1" pageNumber="288">Figs 1O, P</figureCitation>
|
||
). Teleoconch sculpture as for genus (
|
||
<figureCitation id="F3E08FB47575FF8EE130195CFD25FBD6" box="[611,695,1053,1076]" captionStart="FIGURE 1" captionStartId="2.[151,244,1877,1899]" captionTargetBox="[150,1436,330,1838]" captionTargetId="figure@2.[150,1436,330,1838]" captionTargetPageId="2" captionText="FIGURE 1. Habeastrum spp. shells. A–J. Habeastrum parafusum, n. sp. A. Holotype MZSP 134301, L 1.4 mm. B–J. Paratypes MZSP 135584, L 1.7 mm. C. L 1.6 mm. D. L 1.6 mm. E. L 1.4 mm. F. SEM, L 1.1 mm. G. L. 1.2 mm. H. same specimen, showing protoconch and first teleoconch whorls, SEM, scale = 0.1 mm. I. same specimen of F, scale = 0.1 mm. J. Young specimen, L 0.6 mm. K–R. H. omphalium n. sp. K–N. Holotype MZSP 135583, W 1.1 mm, L 1.6 mm. O–R. Paratypes MZSP 134324. O. L 1.0 mm. P. SEM, L 1.0 mm. Q–R. Scale = 0.1 mm." httpUri="https://zenodo.org/record/2617835/files/figure.png" pageId="1" pageNumber="288">Fig. 1Q</figureCitation>
|
||
), with about 75 axial cords on penultimate whorl. Suture deep, whorl profile rounded. Aperture rounded, not deviating from whorl profile, slightly orthocline (
|
||
<figureCitation id="F3E08FB47575FF8EE7021901FB3EFBB5" box="[1105,1196,1088,1111]" captionStart="FIGURE 1" captionStartId="2.[151,244,1877,1899]" captionTargetBox="[150,1436,330,1838]" captionTargetId="figure@2.[150,1436,330,1838]" captionTargetPageId="2" captionText="FIGURE 1. Habeastrum spp. shells. A–J. Habeastrum parafusum, n. sp. A. Holotype MZSP 134301, L 1.4 mm. B–J. Paratypes MZSP 135584, L 1.7 mm. C. L 1.6 mm. D. L 1.6 mm. E. L 1.4 mm. F. SEM, L 1.1 mm. G. L. 1.2 mm. H. same specimen, showing protoconch and first teleoconch whorls, SEM, scale = 0.1 mm. I. same specimen of F, scale = 0.1 mm. J. Young specimen, L 0.6 mm. K–R. H. omphalium n. sp. K–N. Holotype MZSP 135583, W 1.1 mm, L 1.6 mm. O–R. Paratypes MZSP 134324. O. L 1.0 mm. P. SEM, L 1.0 mm. Q–R. Scale = 0.1 mm." httpUri="https://zenodo.org/record/2617835/files/figure.png" pageId="1" pageNumber="288">Fig. 1M</figureCitation>
|
||
); minimally deformed by penultimate whorl; occupying about 42% of shell width, about 25% of shell length. Umbilicus deep, wide, occupying about 20% of inferior shell area (
|
||
<figureCitation id="F3E08FB47575FF8EE2A419C4FDE2FB7E" box="[503,624,1157,1180]" captionStart="FIGURE 1" captionStartId="2.[151,244,1877,1899]" captionTargetBox="[150,1436,330,1838]" captionTargetId="figure@2.[150,1436,330,1838]" captionTargetPageId="2" captionText="FIGURE 1. Habeastrum spp. shells. A–J. Habeastrum parafusum, n. sp. A. Holotype MZSP 134301, L 1.4 mm. B–J. Paratypes MZSP 135584, L 1.7 mm. C. L 1.6 mm. D. L 1.6 mm. E. L 1.4 mm. F. SEM, L 1.1 mm. G. L. 1.2 mm. H. same specimen, showing protoconch and first teleoconch whorls, SEM, scale = 0.1 mm. I. same specimen of F, scale = 0.1 mm. J. Young specimen, L 0.6 mm. K–R. H. omphalium n. sp. K–N. Holotype MZSP 135583, W 1.1 mm, L 1.6 mm. O–R. Paratypes MZSP 134324. O. L 1.0 mm. P. SEM, L 1.0 mm. Q–R. Scale = 0.1 mm." httpUri="https://zenodo.org/record/2617835/files/figure.png" pageId="1" pageNumber="288">Figs 1K, R</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="23C1C0BA7575FF8EE39419E9FD69FB03" pageId="1" pageNumber="288" type="distribution">
|
||
<paragraph id="6B6493317575FF8EE39419E9FC0FFB5D" blockId="1.[151,1437,740,1631]" box="[199,925,1192,1215]" pageId="1" pageNumber="288">
|
||
<emphasis id="59AF4F237575FF8EE39419E9FECBFB5D" bold="true" box="[199,345,1192,1215]" pageId="1" pageNumber="288">Distribution.</emphasis>
|
||
Known only from the
|
||
<typeStatus id="B4602D937575FF8EE11F19E9FDE8FB5D" box="[588,634,1192,1215]" pageId="1" pageNumber="288">type</typeStatus>
|
||
locality, possibly endemic.
|
||
</paragraph>
|
||
<paragraph id="6B6493317575FF8EE394198BFD69FB03" blockId="1.[151,1437,740,1631]" box="[199,763,1226,1249]" pageId="1" pageNumber="288">
|
||
<emphasis id="59AF4F237575FF8EE394198BFEB7FB03" bold="true" box="[199,293,1226,1249]" pageId="1" pageNumber="288">Habitat.</emphasis>
|
||
In soil in caves (no living specimen found).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="23C1C0BA7575FF8EE39419ACFCAAFAC5" pageId="1" pageNumber="288" type="description">
|
||
<paragraph id="6B6493317575FF8EE39419ACFCAAFAC5" blockId="1.[151,1437,740,1631]" pageId="1" pageNumber="288">
|
||
<emphasis id="59AF4F237575FF8EE39419ACFEFCFAE6" bold="true" box="[199,366,1261,1284]" pageId="1" pageNumber="288">Measurements</emphasis>
|
||
(length and width in mm).
|
||
<typeStatus id="B4602D937575FF8EE1DC19ACFD64FAE6" box="[655,758,1261,1284]" pageId="1" pageNumber="288" type="holotype">Holotype</typeStatus>
|
||
:
|
||
<quantity id="AC233ED47575FF8EE05319ACFCC3FAE6" box="[768,849,1261,1284]" metricMagnitude="-3" metricUnit="m" metricValue="1.6" pageId="1" pageNumber="288" unit="mm" value="1.6">1.6 mm</quantity>
|
||
by
|
||
<quantity id="AC233ED47575FF8EE02B19ACFC5BFAE6" box="[888,969,1261,1284]" metricMagnitude="-3" metricUnit="m" metricValue="1.1" pageId="1" pageNumber="288" unit="mm" value="1.1">1.1 mm</quantity>
|
||
(
|
||
<figureCitation id="F3E08FB47575FF8EE08419ACFB87FAE6" box="[983,1045,1261,1284]" captionStart="FIGURE 1" captionStartId="2.[151,244,1877,1899]" captionTargetBox="[150,1436,330,1838]" captionTargetId="figure@2.[150,1436,330,1838]" captionTargetPageId="2" captionText="FIGURE 1. Habeastrum spp. shells. A–J. Habeastrum parafusum, n. sp. A. Holotype MZSP 134301, L 1.4 mm. B–J. Paratypes MZSP 135584, L 1.7 mm. C. L 1.6 mm. D. L 1.6 mm. E. L 1.4 mm. F. SEM, L 1.1 mm. G. L. 1.2 mm. H. same specimen, showing protoconch and first teleoconch whorls, SEM, scale = 0.1 mm. I. same specimen of F, scale = 0.1 mm. J. Young specimen, L 0.6 mm. K–R. H. omphalium n. sp. K–N. Holotype MZSP 135583, W 1.1 mm, L 1.6 mm. O–R. Paratypes MZSP 134324. O. L 1.0 mm. P. SEM, L 1.0 mm. Q–R. Scale = 0.1 mm." httpUri="https://zenodo.org/record/2617835/files/figure.png" pageId="1" pageNumber="288">Figs 1</figureCitation>
|
||
K–N);
|
||
<typeStatus id="B4602D937575FF8EE73719ACFB5EFAE6" box="[1124,1228,1261,1284]" pageId="1" pageNumber="288" type="paratype">Paratypes</typeStatus>
|
||
MZSP 134324, #2, 1.0 mm by
|
||
<quantity id="AC233ED47575FF8EE2431851FEF3FAC5" box="[272,353,1296,1319]" metricMagnitude="-4" metricUnit="m" metricValue="9.0" pageId="1" pageNumber="288" unit="mm" value="0.9">0.9 mm</quantity>
|
||
(
|
||
<figureCitation id="F3E08FB47575FF8EE2231851FE56FAC5" box="[368,452,1296,1319]" captionStart="FIGURE 1" captionStartId="2.[151,244,1877,1899]" captionTargetBox="[150,1436,330,1838]" captionTargetId="figure@2.[150,1436,330,1838]" captionTargetPageId="2" captionText="FIGURE 1. Habeastrum spp. shells. A–J. Habeastrum parafusum, n. sp. A. Holotype MZSP 134301, L 1.4 mm. B–J. Paratypes MZSP 135584, L 1.7 mm. C. L 1.6 mm. D. L 1.6 mm. E. L 1.4 mm. F. SEM, L 1.1 mm. G. L. 1.2 mm. H. same specimen, showing protoconch and first teleoconch whorls, SEM, scale = 0.1 mm. I. same specimen of F, scale = 0.1 mm. J. Young specimen, L 0.6 mm. K–R. H. omphalium n. sp. K–N. Holotype MZSP 135583, W 1.1 mm, L 1.6 mm. O–R. Paratypes MZSP 134324. O. L 1.0 mm. P. SEM, L 1.0 mm. Q–R. Scale = 0.1 mm." httpUri="https://zenodo.org/record/2617835/files/figure.png" pageId="1" pageNumber="288">Fig. 1O</figureCitation>
|
||
); #3, 1.0 mm by
|
||
<quantity id="AC233ED47575FF8EE1281851FD5FFAC5" box="[635,717,1296,1319]" metricMagnitude="-4" metricUnit="m" metricValue="9.0" pageId="1" pageNumber="288" unit="mm" value="0.9">0.9 mm</quantity>
|
||
(
|
||
<figureCitation id="F3E08FB47575FF8EE1881851FCB9FAC5" box="[731,811,1296,1319]" captionStart="FIGURE 1" captionStartId="2.[151,244,1877,1899]" captionTargetBox="[150,1436,330,1838]" captionTargetId="figure@2.[150,1436,330,1838]" captionTargetPageId="2" captionText="FIGURE 1. Habeastrum spp. shells. A–J. Habeastrum parafusum, n. sp. A. Holotype MZSP 134301, L 1.4 mm. B–J. Paratypes MZSP 135584, L 1.7 mm. C. L 1.6 mm. D. L 1.6 mm. E. L 1.4 mm. F. SEM, L 1.1 mm. G. L. 1.2 mm. H. same specimen, showing protoconch and first teleoconch whorls, SEM, scale = 0.1 mm. I. same specimen of F, scale = 0.1 mm. J. Young specimen, L 0.6 mm. K–R. H. omphalium n. sp. K–N. Holotype MZSP 135583, W 1.1 mm, L 1.6 mm. O–R. Paratypes MZSP 134324. O. L 1.0 mm. P. SEM, L 1.0 mm. Q–R. Scale = 0.1 mm." httpUri="https://zenodo.org/record/2617835/files/figure.png" pageId="1" pageNumber="288">Fig. 1P</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="23C1C0BA7575FF8EE3941873FE9DF9BD" pageId="1" pageNumber="288" type="etymology">
|
||
<paragraph id="6B6493317575FF8EE3941873FE97FA8E" blockId="1.[151,1437,740,1631]" pageId="1" pageNumber="288">
|
||
<emphasis id="59AF4F237575FF8EE3941873FED5FAAB" bold="true" box="[199,327,1330,1353]" pageId="1" pageNumber="288">Etymology.</emphasis>
|
||
The specific epithet refers to wide umbilicus, result of the wider growth of the shell, from the Greek
|
||
<emphasis id="59AF4F237575FF8EE3C41815FF6DFA8E" box="[151,255,1364,1388]" italics="true" pageId="1" pageNumber="288">omphalos</emphasis>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="6B6493317575FF8EE3941839FE9DF9BD" blockId="1.[151,1437,740,1631]" pageId="1" pageNumber="288">
|
||
<emphasis id="59AF4F237575FF8EE3941839FE5FFA6D" bold="true" box="[199,461,1400,1423]" pageId="1" pageNumber="288">Taxonomic discussion.</emphasis>
|
||
<taxonomicName id="ACDBE8B27575FF8EE2881836FD4DFA6D" authorityName="Simone" authorityYear="2019" box="[475,735,1399,1423]" class="Gastropoda" family="Diplommatinidae" genus="Habeastrum" higherTaxonomySource="GBIF" kingdom="Animalia" order="Architaenioglossa" pageId="1" pageNumber="288" phylum="Mollusca" rank="species" species="parafusum">
|
||
<emphasis id="59AF4F237575FF8EE2881836FD4DFA6D" box="[475,735,1399,1423]" italics="true" pageId="1" pageNumber="288">Habeastrum parafusum</emphasis>
|
||
</taxonomicName>
|
||
can be distinguished from
|
||
<taxonomicName id="ACDBE8B27575FF8EE77B1839FB5BFA6D" authorityName="Simone" authorityYear="2019" box="[1064,1225,1399,1423]" class="Gastropoda" family="Diplommatinidae" genus="Habeastrum" higherTaxonomySource="GBIF" kingdom="Animalia" order="Architaenioglossa" pageId="1" pageNumber="288" phylum="Mollusca" rank="species" species="omphalium">
|
||
<emphasis id="59AF4F237575FF8EE77B1839FB5BFA6D" box="[1064,1225,1399,1423]" italics="true" pageId="1" pageNumber="288">H. omphalium</emphasis>
|
||
</taxonomicName>
|
||
by the characters described above. The main difference is the degree of inflation during the shell growth, much larger in
|
||
<taxonomicName id="ACDBE8B27575FF8EE65218DBFA0EFA53" authorityName="Simone" authorityYear="2019" box="[1281,1436,1433,1457]" class="Gastropoda" family="Diplommatinidae" genus="Habeastrum" higherTaxonomySource="GBIF" kingdom="Animalia" order="Architaenioglossa" pageId="1" pageNumber="288" phylum="Mollusca" rank="species" species="omphalium">
|
||
<emphasis id="59AF4F237575FF8EE65218DBFA0EFA53" box="[1281,1436,1433,1457]" italics="true" pageId="1" pageNumber="288">H. omphalium</emphasis>
|
||
</taxonomicName>
|
||
than that of
|
||
<taxonomicName id="ACDBE8B27575FF8EE27318FCFE27FA36" authorityName="Simone" authorityYear="2019" box="[288,437,1468,1492]" class="Gastropoda" family="Diplommatinidae" genus="Habeastrum" higherTaxonomySource="GBIF" kingdom="Animalia" order="Architaenioglossa" pageId="1" pageNumber="288" phylum="Mollusca" rank="species" species="parafusum">
|
||
<emphasis id="59AF4F237575FF8EE27318FCFE27FA36" box="[288,437,1468,1492]" italics="true" pageId="1" pageNumber="288">H. parafusum</emphasis>
|
||
</taxonomicName>
|
||
. This difference produces, in specimens of a same size, shells with different shape (rather cylindrical in
|
||
<taxonomicName id="ACDBE8B27575FF8EE27818A1FE2CFA15" authorityName="Simone" authorityYear="2019" box="[299,446,1503,1527]" class="Gastropoda" family="Diplommatinidae" genus="Habeastrum" higherTaxonomySource="GBIF" kingdom="Animalia" order="Architaenioglossa" pageId="1" pageNumber="288" phylum="Mollusca" rank="species" species="parafusum">
|
||
<emphasis id="59AF4F237575FF8EE27818A1FE2CFA15" box="[299,446,1503,1527]" italics="true" pageId="1" pageNumber="288">H. parafusum</emphasis>
|
||
</taxonomicName>
|
||
and conical in
|
||
<taxonomicName id="ACDBE8B27575FF8EE13218A1FD68FA15" authorityName="Simone" authorityYear="2019" box="[609,762,1503,1527]" class="Gastropoda" family="Diplommatinidae" genus="Habeastrum" higherTaxonomySource="GBIF" kingdom="Animalia" order="Architaenioglossa" pageId="1" pageNumber="288" phylum="Mollusca" rank="species" species="omphalium">
|
||
<emphasis id="59AF4F237575FF8EE13218A1FD68FA15" box="[609,762,1503,1527]" italics="true" pageId="1" pageNumber="288">H. omphalium</emphasis>
|
||
</taxonomicName>
|
||
), different number of whorls (about
|
||
<quantity id="AC233ED47575FF8EE7D118A1FB39FA15" box="[1154,1195,1504,1527]" metricMagnitude="-1" metricUnit="m" metricValue="1.778" pageId="1" pageNumber="288" unit="in" value="7.0">7 in</quantity>
|
||
<taxonomicName id="ACDBE8B27575FF8EE7E118A1FAD7FA15" authorityName="Simone" authorityYear="2019" box="[1202,1349,1503,1527]" class="Gastropoda" family="Diplommatinidae" genus="Habeastrum" higherTaxonomySource="GBIF" kingdom="Animalia" order="Architaenioglossa" pageId="1" pageNumber="288" phylum="Mollusca" rank="species" species="parafusum">
|
||
<emphasis id="59AF4F237575FF8EE7E118A1FAD7FA15" box="[1202,1349,1503,1527]" italics="true" pageId="1" pageNumber="288">H. parafusum</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<quantity id="AC233ED47575FF8EE60118A1FAE9FA15" box="[1362,1403,1504,1527]" metricMagnitude="-1" metricUnit="m" metricValue="1.27" pageId="1" pageNumber="288" unit="in" value="5.0">5 in</quantity>
|
||
<taxonomicName id="ACDBE8B27575FF8EE6D118A1FE9DF9FB" authorityName="Simone" authorityYear="2019" class="Gastropoda" family="Diplommatinidae" genus="Habeastrum" higherTaxonomySource="GBIF" kingdom="Animalia" order="Architaenioglossa" pageId="1" pageNumber="288" phylum="Mollusca" rank="species" species="omphalium">
|
||
<emphasis id="59AF4F237575FF8EE6D118A1FE9DF9FB" italics="true" pageId="1" pageNumber="288">H. omphalium</emphasis>
|
||
</taxonomicName>
|
||
), and distinct umbilicus width (wider in
|
||
<taxonomicName id="ACDBE8B27575FF8EE1811B43FCFFF9FB" authorityName="Simone" authorityYear="2019" box="[722,877,1537,1561]" class="Gastropoda" family="Diplommatinidae" genus="Habeastrum" higherTaxonomySource="GBIF" kingdom="Animalia" order="Architaenioglossa" pageId="1" pageNumber="288" phylum="Mollusca" rank="species" species="omphalium">
|
||
<emphasis id="59AF4F237575FF8EE1811B43FCFFF9FB" box="[722,877,1537,1561]" italics="true" pageId="1" pageNumber="288">H. omphalium</emphasis>
|
||
</taxonomicName>
|
||
). Despite having been found in the same cave, the relatively large amount of specimens and the lack of intermediates between both shell shapes ensure the species’ distinction.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |