treatments-xml/data/BB/3D/87/BB3D87DA0112FF9FFF6ECFCF9C62C213.xml
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<document id="A92808929A94DE0D420977C2E6416927" ID-CLB-Dataset="53630" ID-DOI="10.11646/phytotaxa.542.2.9" ID-GBIF-Dataset="d0b52ac6-2df6-4e54-9302-656f49298295" ID-ISSN="1179-3163" ID-Zenodo-Dep="6418029" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1649142044537" checkinUser="plazi" docAuthor="Zografidis, Aris, Esser, Hans-Joachim, Dimopoulos, Panayotis &amp; Raus, Thomas" docDate="2022" docId="BB3D87DA0112FF9FFF6ECFCF9C62C213" docLanguage="en" docName="phytotaxa.542.2.9.pdf" docOrigin="Phytotaxa 542 (2)" docSource="http://dx.doi.org/10.11646/phytotaxa.542.2.9" docStyle="DocumentStyle:96748F8F1B6C902996E134952A3A36B9.7:Phytotaxa.2014-.journal_article" docStyleId="96748F8F1B6C902996E134952A3A36B9" docStyleName="Phytotaxa.2014-.journal_article" docStyleVersion="7" docTitle="Verbascum sipiadense Zograf." docType="treatment" docVersion="7" lastPageNumber="218" masterDocId="4704FFA20110FF9AFFD3CA019D42C337" masterDocTitle="Typification of the names Verbascum limnense and Celsia tomentosa (Scrophulariaceae) and a new nothospecies, V. × sipiadense, with the hybrid formula V. limnense × V. sinuatum" masterLastPageNumber="220" masterPageNumber="214" pageNumber="216" updateTime="1699319337158" updateUser="ExternalLinkService" zenodo-license-figures="UNSPECIFIED" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="081CC99A025A6CFCFDBDC7AC658AF048">Typification of the names Verbascum limnense and Celsia tomentosa (Scrophulariaceae) and a new nothospecies, V. × sipiadense, with the hybrid formula V. limnense × V. sinuatum</mods:title>
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<mods:namePart id="D14613989C08E7D8F2CA00B240618167">Zografidis, Aris</mods:namePart>
<mods:nameIdentifier id="A5431462E6A2B3158F9187F711AB44F3" type="ORCID">0000-0002-1559-4900</mods:nameIdentifier>
<mods:affiliation id="474559AAFD5EFFDDBF123B7193C45D88">Laboratory of Botany, Faculty of Biology, University of Patras, 26504, Greece &amp; azografidis @ upatras. gr; https: // orcid. org / 0000 - 0002 - 1559 - 4900</mods:affiliation>
<mods:nameIdentifier id="F5D61E08B26749D873A04E71212A71B7" type="email">azografidis@upatras.gr</mods:nameIdentifier>
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<mods:namePart id="2E743AE6A0F8941BA51A32AFF3510858">Esser, Hans-Joachim</mods:namePart>
<mods:nameIdentifier id="8CCE21297C7D57A031A680E08DD2EDD8" type="ORCID">0000-0001-6518-8612</mods:nameIdentifier>
<mods:affiliation id="97EC333CF9D33915052AD8D03AEE98F4">SNSB-Botanische Staatssammlung München, 80638 München, Germany &amp; esser @ snsb. de; https: // orcid. org / 0000 - 0001 - 6518 - 8612</mods:affiliation>
<mods:nameIdentifier id="C9C2A4F7B37880518279C499A003DCE4" type="email">esser@snsb.de</mods:nameIdentifier>
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<mods:namePart id="7F60EE5C3F3323A75833007466A0F9C6">Dimopoulos, Panayotis</mods:namePart>
<mods:nameIdentifier id="1714ABAE367614467BE0881EC7B3835B" type="ORCID">0000-0003-0699-9954</mods:nameIdentifier>
<mods:affiliation id="EA54012AABFA195C8020E57C1EC518BB">Laboratory of Botany, Faculty of Biology, University of Patras, 26504, Greece &amp; pdimopoulos @ upatras. gr; https: // orcid. org / 0000 - 0003 - 0699 - 9954</mods:affiliation>
<mods:nameIdentifier id="A32841262DBC3F520BFCD2C276418975" type="email">pdimopoulos@upatras.gr</mods:nameIdentifier>
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<mods:namePart id="3B751456879FCB68850549DAA73FF2AC">Raus, Thomas</mods:namePart>
<mods:nameIdentifier id="471A218FB01A5F4A1E3E699CAEC7B4C2" type="ORCID">0000-0001-5778-4705</mods:nameIdentifier>
<mods:affiliation id="EEC3EED9F328DAC1EB79E4DB29F73D3A">Botanischer Garten und Botanisches Museum Berlin, Freie Universität Berlin, Königin-Luise-Strasse 6 - 8, 14195 Berlin, Germany &amp; t. raus @ bo. berlin; https: // orcid. org / 0000 - 0001 - 5778 - 4705</mods:affiliation>
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<mods:title id="908AB6789083DE0C058430E881BE9EAD">Phytotaxa</mods:title>
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<mods:date id="4D0B32963E4743019E8DDC16D0781467">2022</mods:date>
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<mods:number id="7BA787414A6DDE5B1AD8F8990786C26C">542</mods:number>
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<treatment id="BB3D87DA0112FF9FFF6ECFCF9C62C213" ID-DOI="http://doi.org/10.5281/zenodo.6418033" ID-GBIF-Taxon="194466555" ID-Zenodo-Dep="6418033" LSID="urn:lsid:plazi:treatment:BB3D87DA0112FF9FFF6ECFCF9C62C213" httpUri="http://treatment.plazi.org/id/BB3D87DA0112FF9FFF6ECFCF9C62C213" lastPageId="5" lastPageNumber="218" pageId="2" pageNumber="216">
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<paragraph id="332B36CC0112FF98FF6ECFCF9F70C6DF" blockId="2.[136,1452,1378,2089]" box="[189,562,1486,1513]" pageId="2" pageNumber="216">
<taxonomicName id="F4944D4F0112FF98FF6ECFCF9F70C6DF" authority="Zograf." authorityName="Zograf." box="[189,562,1486,1513]" class="Magnoliopsida" family="Scrophulariaceae" genus="Verbascum" isHybrid="true" kingdom="Plantae" order="Lamiales" pageId="2" pageNumber="216" phylum="Tracheophyta" rank="species" species="sipiadense">
<emphasis id="01E0EADE0112FF98FF6ECFCF9C7FC6DE" bold="true" box="[189,317,1486,1513]" italics="true" pageId="2" pageNumber="216">Verbascum</emphasis>
×
<emphasis id="01E0EADE0112FF98FE8DCFCF9C94C6DE" bold="true" box="[350,470,1486,1513]" italics="true" pageId="2" pageNumber="216">sipiadense</emphasis>
Zograf.
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</paragraph>
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<paragraph id="332B36CC0112FF98FDE9CFCF9963C6DF" blockId="2.[136,1452,1378,2089]" box="[570,1057,1486,1513]" pageId="2" pageNumber="216">
(
<taxonomicName id="F4944D4F0112FF98FD90CFCE9F87C6DF" ID-CoL="7FQBH" authorityName="Fraas" authorityYear="1845" box="[579,709,1486,1512]" class="Magnoliopsida" family="Scrophulariaceae" genus="Verbascum" kingdom="Plantae" order="Lamiales" pageId="2" pageNumber="216" phylum="Tracheophyta" rank="species" species="limnense">
<emphasis id="01E0EADE0112FF98FD90CFCE9F87C6DF" box="[579,709,1486,1512]" italics="true" pageId="2" pageNumber="216">V. limnense</emphasis>
</taxonomicName>
×
<taxonomicName id="F4944D4F0112FF98FD36CFCE9E2BC6DF" ID-CoL="5B24N" authorityName="Linnaeus" authorityYear="1753" box="[741,873,1487,1512]" class="Magnoliopsida" family="Scrophulariaceae" genus="Verbascum" kingdom="Plantae" order="Lamiales" pageId="2" pageNumber="216" phylum="Tracheophyta" rank="species" species="sinuatum">
<emphasis id="01E0EADE0112FF98FD36CFCE9E2BC6DF" box="[741,873,1487,1512]" italics="true" pageId="2" pageNumber="216">V. sinuatum</emphasis>
</taxonomicName>
),
<emphasis id="01E0EADE0112FF98FC51CFCF9963C6DF" bold="true" box="[898,1057,1486,1512]" pageId="2" pageNumber="216">nothosp. nov.</emphasis>
</paragraph>
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<paragraph id="332B36CC0112FF98FBF2CFCE9E85C507" blockId="2.[136,1452,1378,2089]" pageId="2" pageNumber="216">
<emphasis id="01E0EADE0112FF98FBF2CFCE9978C6DF" box="[1057,1082,1487,1512]" italics="true" pageId="2" pageNumber="216"></emphasis>
<materialsCitation id="83FC3C910112FF98FBE9CFCE9F84C507" ID-GBIF-Occurrence="3747628303" collectingDate="2018-05-05" collectionCode="UPA" collectorName="Zografidis" country="Greece" county="Sipiada" elevation="30" latitude="38.766666" location="Nomos Evvias" longLatPrecision="1173" longitude="23.266666" municipality="Island Evvia" pageId="2" pageNumber="216" specimenCount="1" typeStatus="holotype">
Type:
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.
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,
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:
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, just outside the settlement of
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(near Limni) on roadsides,
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,
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,
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,
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<collectingDate id="576EE9E40112FF98FF1CCC179C1BC507" box="[207,345,1558,1585]" pageId="2" pageNumber="216" value="2018-05-05">5 May 2018</collectingDate>
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,
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360
</emphasis>
(
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!
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;
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<typeStatus id="EC2F886E0112FF98FD02CC179E60C507" box="[721,802,1558,1584]" pageId="2" pageNumber="216" type="isotype">isotype</typeStatus>
<collectionCode id="5585AE090112FF98FCF4CC179E2AC507" box="[807,872,1558,1584]" pageId="2" pageNumber="216">ATH</collectionCode>
!).
<figureCitation id="ABAF2A490112FF98FCADCC179E83C507" box="[894,961,1558,1584]" captionStart="FIGURE 3" captionStartId="4.[136,229,1953,1975]" captionTargetBox="[203,1385,190,1922]" captionTargetId="figure-18@4.[203,1385,190,1928]" captionTargetPageId="4" captionText="FIGURE 3. Holotype of Verbascum × sipiadense Zograf. kept in UPA (UPA-29510)." figureDoi="http://doi.org/10.5281/zenodo.6418080" httpUri="https://zenodo.org/record/6418080/files/figure.png" pageId="2" pageNumber="216">Fig. 3</figureCitation>
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.
</paragraph>
</subSubSection>
<subSubSection id="7B8E65470112FF98FF6ECC3B9EEEC43E" pageId="2" pageNumber="216" type="diagnosis">
<paragraph id="332B36CC0112FF98FF6ECC3B9EEEC43E" blockId="2.[136,1452,1378,2089]" pageId="2" pageNumber="216">
<emphasis id="01E0EADE0112FF98FF6ECC3B9C72C563" bold="true" box="[189,304,1594,1620]" pageId="2" pageNumber="216">Diagnosis</emphasis>
: Intermediate between the parental species
<taxonomicName id="F4944D4F0112FF98FCCECC3A9EDFC563" authorityName="Fraas" authorityYear="1845" box="[797,925,1594,1620]" class="Magnoliopsida" family="Scrophulariaceae" genus="Verbascum" kingdom="Plantae" order="Lamiales" pageId="2" pageNumber="216" phylum="Tracheophyta" rank="species" species="limnense">
<emphasis id="01E0EADE0112FF98FCCECC3A9EDFC563" box="[797,925,1594,1620]" italics="true" pageId="2" pageNumber="216">V. limnense</emphasis>
</taxonomicName>
and
<taxonomicName id="F4944D4F0112FF98FC07CC3A9915C563" authorityName="Linnaeus" authorityYear="1753" box="[980,1111,1595,1620]" class="Magnoliopsida" family="Scrophulariaceae" genus="Verbascum" kingdom="Plantae" order="Lamiales" pageId="2" pageNumber="216" phylum="Tracheophyta" rank="species" species="sinuatum">
<emphasis id="01E0EADE0112FF98FC07CC3A9915C563" box="[980,1111,1595,1620]" italics="true" pageId="2" pageNumber="216">V. sinuatum</emphasis>
</taxonomicName>
but sterile. Differing from the former by having the lamina of basal leaves coarsely crenate to lobed (vs. crenate to subentire), leafy flowering stems (vs. subaphyllous), up to 4-flowered glomerules (vs. solitary flowers), bracteoles present (vs. bracteoles absent) and bicoloured, white-violet stamen hairs (vs. white). Differing from the latter by the presence of long eglandular hairs at the base of the petioles of rosette leaves, 2- to 7-flowered glomerules (vs. 1- to 4-flowered), glabrous outer surface of corolla (vs. stellate-tomentose) and bicoloured stamen hairs (vs. violet).
</paragraph>
</subSubSection>
<subSubSection id="7B8E65470112FF99FF6ECD139F35C237" lastPageId="3" lastPageNumber="217" pageId="2" pageNumber="216" type="description">
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<emphasis id="01E0EADE0112FF98FF6ECD139C05C41B" bold="true" box="[189,327,1810,1836]" pageId="2" pageNumber="216">Description</emphasis>
: Sterile biennial herb producing a basal leaf rosette in the first year of growth and an erect, terete, leafy flowering stem
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of height in the second year. Indumentum covering the aerial part of the plant eglandular, grayish-yellowish, persistent. Rosette-leaves petiolate with petioles
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long, base of petioles with longer hairs; lamina obovate to obovate-elliptic, coarsely crenate to lobed, attenuate to petiole, obtuse at apex, 730 ×
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. Lower cauline leaves similar but ±sessile. Middle and upper cauline leaves sessile, progressively smaller, ovate-oblong to ovate-triangular, crenulate, subacute to acute, the upper ones ca. 2 ×
<quantity id="F46C9B290112FF98FC47CDC79E8FC4D6" box="[916,973,1990,2017]" metricMagnitude="-2" metricUnit="m" metricValue="1.0" pageId="2" pageNumber="216" unit="cm" value="1.0">1 cm</quantity>
. Inflorescence freely branched from below, lax. Flowers solitary or in groups (glomerules) of 24; bracts c. 2 ×
<quantity id="F46C9B290112FF98FCADCDEB9E83CB32" box="[894,961,2026,2053]" metricMagnitude="-3" metricUnit="m" metricValue="1.0" pageId="2" pageNumber="216" unit="mm" value="1.0">1 mm</quantity>
, ovate-lanceolate, acute; bracteoles present, lanceolate; pedicels
<quantity id="F46C9B290112FF98FEBEC20F9C8FCB1E" box="[365,461,2062,2089]" metricMagnitude="-3" metricUnit="m" metricValue="2.0" metricValueMax="3.0" metricValueMin="1.0" pageId="2" pageNumber="216" unit="mm" value="2.0" valueMax="3.0" valueMin="1.0">13 mm</quantity>
. Calyx
<quantity id="F46C9B290112FF98FDF0C20F9FC1CB1E" box="[547,643,2062,2089]" metricMagnitude="-3" metricUnit="m" metricValue="2.5" metricValueMax="3.0" metricValueMin="2.0" pageId="2" pageNumber="216" unit="mm" value="2.5" valueMax="3.0" valueMin="2.0">23 mm</quantity>
, segments lanceolate-oblong, acute, divided almost to the base. Corolla yellow,
<quantity id="F46C9B290113FF99FF33CA9F9C0AC38E" box="[224,328,158,185]" metricMagnitude="-2" metricUnit="m" metricValue="2.25" metricValueMax="2.5" metricValueMin="2.0" pageId="3" pageNumber="217" unit="cm" value="2.25" valueMax="2.5" valueMin="2.0">22.5 cm</quantity>
in diameter, with pellucid glands, outer surface glabrous, inner surface glabrous or with simple hairs on the proximal part of upper lobes (on the central vein of the lobe); stamens 4(5) with medifixed anthers; filament hairs white and violet. Capsules not formed.
</paragraph>
</subSubSection>
<caption id="67EB66440113FF99FF5BC20A9E15CB15" ID-DOI="http://doi.org/10.5281/zenodo.6418045" ID-Zenodo-Dep="6418045" box="[136,855,2059,2082]" httpUri="https://zenodo.org/record/6418045/files/figure.png" pageId="3" pageNumber="217" startId="3.[136,229,2059,2081]" targetBox="[219,1368,281,2035]" targetPageId="3">
<paragraph id="332B36CC0113FF99FF5BC20A9E15CB15" blockId="3.[136,855,2059,2082]" box="[136,855,2059,2082]" pageId="3" pageNumber="217">
<emphasis id="01E0EADE0113FF99FF5BC20A9DB5CB16" bold="true" box="[136,247,2059,2081]" pageId="3" pageNumber="217">FIGURE 2</emphasis>
. Lectotype of
<taxonomicName id="F4944D4F0113FF99FE50C20D9F24CB16" authority="Zucc." authorityName="Zuccarini" authorityYear="1837" box="[387,614,2060,2081]" class="Magnoliopsida" family="Scrophulariaceae" genus="Celsia" higherTaxonomySource="GBIF" kingdom="Plantae" order="Lamiales" pageId="3" pageNumber="217" phylum="Tracheophyta" rank="species" species="tomentosa">
<emphasis id="01E0EADE0113FF99FE50C20D9F68CB16" box="[387,554,2060,2081]" italics="true" pageId="3" pageNumber="217">Celsia tomentosa</emphasis>
Zucc.
</taxonomicName>
kept in M (M-0244375).
</paragraph>
</caption>
<caption id="67EB66440114FF9EFF5BCDA09EFCC480" ID-DOI="http://doi.org/10.5281/zenodo.6418080" ID-Zenodo-Dep="6418080" box="[136,958,1953,1975]" httpUri="https://zenodo.org/record/6418080/files/figure.png" pageId="4" pageNumber="218" startId="4.[136,229,1953,1975]" targetBox="[203,1385,190,1922]" targetPageId="4">
<paragraph id="332B36CC0114FF9EFF5BCDA09EFCC480" blockId="4.[136,958,1953,1975]" box="[136,958,1953,1975]" pageId="4" pageNumber="218">
<emphasis id="01E0EADE0114FF9EFF5BCDA09DB5C481" bold="true" box="[136,247,1953,1975]" pageId="4" pageNumber="218">FIGURE 3</emphasis>
. Holotype of
<emphasis id="01E0EADE0114FF9EFEAECDA09F25C481" box="[381,615,1953,1975]" italics="true" pageId="4" pageNumber="218">
<taxonomicName id="F4944D4F0114FF9EFEAECDA09CA4C481" box="[381,486,1953,1974]" class="Magnoliopsida" family="Scrophulariaceae" genus="Verbascum" kingdom="Plantae" order="Lamiales" pageId="4" pageNumber="218" phylum="Tracheophyta" rank="genus">Verbascum</taxonomicName>
× sipiadense
</emphasis>
Zograf. kept in UPA (UPA-29510).
</paragraph>
</caption>
<subSubSection id="7B8E65470114FF9FFF6ECDE49C62C213" lastPageId="5" lastPageNumber="219" pageId="4" pageNumber="218" type="discussion">
<paragraph id="332B36CC0114FF9FFF6ECDE49C62C213" blockId="4.[136,1451,2021,2084]" lastBlockId="5.[136,1452,158,292]" lastPageId="5" lastPageNumber="219" pageId="4" pageNumber="218">
<emphasis id="01E0EADE0114FF9EFF6ECDE49C42C4C8" bold="true" box="[189,256,2021,2047]" pageId="4" pageNumber="218">Notes</emphasis>
: Natural hybrids between
<taxonomicName id="F4944D4F0114FF9EFDFFCDE49F9FCB37" box="[556,733,2021,2048]" class="Magnoliopsida" family="Scrophulariaceae" genus="Verbascum" kingdom="Plantae" order="Lamiales" pageId="4" pageNumber="218" phylum="Tracheophyta" rank="species" species="undetermined">
<emphasis id="01E0EADE0114FF9EFDFFCDE49FE5C4C8" box="[556,679,2021,2047]" italics="true" pageId="4" pageNumber="218">Verbascum</emphasis>
spp.
</taxonomicName>
are readily produced when species coexist, whereas all hitherto described hybrid taxa are invariably sterile (e.g.,
<bibRefCitation id="57054B3D0114FF9EFD6BC2089E23CB14" author="Murbeck, S. S." box="[696,865,2057,2083]" pageId="4" pageNumber="218" refId="ref3647" refString="Murbeck, S. S. (1933) Monographie der Gattung Verbascum. H. Ohlsson, Lund, 630 pp." type="book" year="1933">Murbeck 1933</bibRefCitation>
,
<bibRefCitation id="57054B3D0114FF9EFCBDC20899AFCB13" author="Huber-Morath, A. &amp; Rechinger, K. H." box="[878,1261,2057,2084]" pageId="4" pageNumber="218" pagination="42 - 55" refId="ref3458" refString="Huber-Morath, A. &amp; Rechinger, K. H. (1960) Zur Kenntnis der Gattungen Verbascum und Celsia in Griechenland. Mitteilungen des Thuringischen Botanischen Vereins 2: 42 - 55." type="journal article" year="1960">Huber-Morath &amp; Rechinger 1960</bibRefCitation>
). It is also noted that, unlike
<emphasis id="01E0EADE0115FF9FFED9CA9E9CF1C38F" box="[266,435,158,184]" italics="true" pageId="5" pageNumber="219">V. × sipiadense</emphasis>
, the parental species
<taxonomicName id="F4944D4F0115FF9FFD4CCA9E9E5FC38F" authorityName="Fraas" authorityYear="1845" box="[671,797,158,184]" class="Magnoliopsida" family="Scrophulariaceae" genus="Verbascum" kingdom="Plantae" order="Lamiales" pageId="5" pageNumber="219" phylum="Tracheophyta" rank="species" species="limnense">
<emphasis id="01E0EADE0115FF9FFD4CCA9E9E5FC38F" box="[671,797,158,184]" italics="true" pageId="5" pageNumber="219">V. limnense</emphasis>
</taxonomicName>
always exhibits white stamen hairs, not violet ones as cited in former literature (
<bibRefCitation id="57054B3D0115FF9FFEA2CAC39F5DC3EB" author="Ferguson, I. K." box="[369,543,194,220]" pageId="5" pageNumber="219" pagination="205 - 216" refId="ref3020" refString="Ferguson, I. K. (1972) Verbascum L. In: Tutin, T. G., Heywood, V. H., Burges, N. A., Moore, D. M., Valentine, D. H., Walters, S. M. &amp; Webb, D. A. (Eds.) Flora Europaea 3. Cambridge University Press, Cambridge, pp. 205 - 216." type="book chapter" year="1972">Ferguson 1972</bibRefCitation>
,
<bibRefCitation id="57054B3D0115FF9FFDFFCAC39FF4C3EB" author="Trigas, P." box="[556,694,194,220]" pageId="5" pageNumber="219" refId="ref3910" refString="Trigas, P. (2003) Contribution to the study of the endemism of the flora of the island of Evvia. PhD Thesis, University of Patras, Greece, 341 pp. [in Greek with an English summary]." type="book" year="2003">Trigas 2003</bibRefCitation>
,
<bibRefCitation id="57054B3D0115FF9FFD10CAC39E0EC3EA" author="Strid, A." box="[707,844,194,221]" pageId="5" pageNumber="219" pagination="1 - 700" refId="ref3850" refString="Strid, A. (2016 a) Atlas of the Aegean flora, Part 1: Texts and Plates. Englera 33 (1): 1 - 700." type="journal article" year="2016">Strid 2016a</bibRefCitation>
). It is thus hypothesized that the various descriptions of
<taxonomicName id="F4944D4F0115FF9FFF79CAE69C6FC237" authorityName="Fraas" authorityYear="1845" box="[170,301,230,256]" class="Magnoliopsida" family="Scrophulariaceae" genus="Verbascum" kingdom="Plantae" order="Lamiales" pageId="5" pageNumber="219" phylum="Tracheophyta" rank="species" species="limnense">
<emphasis id="01E0EADE0115FF9FFF79CAE69C6FC237" box="[170,301,230,256]" italics="true" pageId="5" pageNumber="219">V. limnense</emphasis>
</taxonomicName>
in previous floras might have been partially based on heterologous material consisting of the new nothospecies.
</paragraph>
</subSubSection>
</treatment>
</document>