treatments-xml/data/66/48/87/664887AB93723651FF35FE50354EFBD3.xml

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<mods:title id="C6928F5C816C8086D08C443B511A968E">Parathelphusa pardus, a new species of lowland freshwater crab from swamps in central Sumatra, Indonesia (Crustacea: Brachyura: Gecarcinucidae)</mods:title>
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<mods:namePart id="23FAE6B2C1BE2A762E3D05E627A1B05D">Ng, Peter K. L.</mods:namePart>
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<mods:namePart id="10452BC4E9220C2F4ECA79D0A29D62A8">Riady, Rikhi</mods:namePart>
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<treatment id="664887AB93723651FF35FE50354EFBD3" ID-DOI="http://doi.org/10.5281/zenodo.6056815" ID-GBIF-Taxon="137308690" ID-Zenodo-Dep="6056815" LSID="urn:lsid:plazi:treatment:664887AB93723651FF35FE50354EFBD3" httpUri="http://treatment.plazi.org/id/664887AB93723651FF35FE50354EFBD3" lastPageId="10" lastPageNumber="505" pageId="1" pageNumber="496">
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<paragraph id="EE5E36BD9372365AFF35FE503431FDCC" blockId="1.[151,520,501,560]" box="[151,520,501,527]" pageId="1" pageNumber="496">
<heading id="B51681D19372365AFF35FE503431FDCC" bold="true" box="[151,520,501,527]" fontSize="11" level="1" pageId="1" pageNumber="496" reason="1">
<emphasis id="DC95EAAF9372365AFF35FE503431FDCC" bold="true" box="[151,520,501,527]" pageId="1" pageNumber="496">
<taxonomicName id="29E14D3E9372365AFF35FE50379AFDCC" ID-CoL="4DFBC" authority="Ng &amp; Riady, 2016" authorityName="Ng &amp; Riady" authorityYear="2016" box="[151,419,501,527]" class="Malacostraca" family="Gecarcinucidae" genus="Parathelphusa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="496" phylum="Arthropoda" rank="species" species="pardus" status="sp. nov.">
<emphasis id="DC95EAAF9372365AFF35FE50379AFDCC" bold="true" box="[151,419,501,527]" italics="true" pageId="1" pageNumber="496">Parathelphusa pardus</emphasis>
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<taxonomicNameLabel id="C7A657D49372365AFE08FE503431FDCC" box="[426,520,501,527]" pageId="1" pageNumber="496" rank="species">sp. nov.</taxonomicNameLabel>
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<paragraph id="EE5E36BD9372365AFF35FDB2372BFDF3" blockId="1.[151,520,501,560]" box="[151,274,535,560]" pageId="1" pageNumber="496">
(
<figureCitation id="76DA2A389372365AFF3DFDB23730FDF3" box="[159,265,535,560]" captionStart-0="FIGURE 1" captionStart-1="FIGURE 2" captionStart-2="FIGURE 3" captionStart-3="FIGURE 4" captionStart-4="FIGURE 5" captionStart-5="FIGURE 6" captionStartId-0="2.[151,250,1966,1988]" captionStartId-1="3.[151,250,1942,1964]" captionStartId-2="4.[151,250,1626,1648]" captionStartId-3="5.[151,250,1933,1955]" captionStartId-4="6.[151,250,1907,1929]" captionStartId-5="7.[151,250,1796,1818]" captionTargetBox-0="[235,1330,193,1888]" captionTargetBox-1="[236,1352,193,1921]" captionTargetBox-2="[151,1436,193,1605]" captionTargetBox-3="[163,1423,503,1907]" captionTargetBox-4="[224,1324,206,1867]" captionTargetBox-5="[151,1436,213,1748]" captionTargetId-0="figure@2.[235,1330,193,1888]" captionTargetId-1="figure@3.[236,1352,193,1921]" captionTargetId-2="figure@4.[151,1436,193,1605]" captionTargetId-3="figure@5.[163,1423,503,1907]" captionTargetId-4="figure@6.[206,1359,193,1881]" captionTargetId-5="figure@7.[151,1436,213,1748]" captionTargetPageId-0="2" captionTargetPageId-1="3" captionTargetPageId-2="4" captionTargetPageId-3="5" captionTargetPageId-4="6" captionTargetPageId-5="7" captionText-0="FIGURE 1. Parathelphusa pardus sp. nov., holotype male (37.6 × 28.2 mm) (MZB Cru 4388), Kampar River, Sumatra. A, overall dorsal view; B, anterior thoraCiC sternum and male abdominal somites 5, 6 and telson; C, outer view of Chelae." captionText-1="FIGURE 2. Parathelphusa pardus sp. nov., paratype male (34.9 × 26.6 mm) (ZRC 2015.0452), Kampar River, Sumatra. A, overall dorsal view; B, dorsal view of CarapaCe; C, frontal view of CarapaCe." captionText-2="FIGURE 3. Parathelphusa pardus sp. nov., paratype male (34.9 × 26.6 mm) (ZRC 2015.0452), Kampar River, Sumatra. A, frontal view showing frontal median triangle of front; B, left third maxilliped; C, thoraCiC sternites 1 – 4 and telson; D, anterior thoraCiC sternum and male abdominal somites 5, 6 and telson; E, posterior thoraCiC sternums and male abdominal somites 1 – 5; F, right minor Chela; G, left major Chela." captionText-3="FIGURE 4. Parathelphusa pardus sp. nov. A – C, paratype male (34.9 × 26.6 mm) (ZRC 2015.0452), Kampar River, Sumatra; D, paratype female (34.7 × 26.9 mm) (ZRC 2015.0452), Kampar River, Sumatra. A, right fourth ambulatory leg; B, right third ambulatory leg; C, D, right seCond ambulatory legs." captionText-4="FIGURE 5. Parathelphusa pardus sp. nov. A – D, holotype male (37.6 × 28.2 mm) (MZB Cru 4388), Kampar River, Sumatra; E – H, paratype male (34.9 × 26.6 mm) (ZRC 2015.0452), Kampar River, Sumatra. A, E, ventral view of left G 1; B, F, ventral view of distal part of left G 1; C, G, dorsal view of distal part of left G 1; D, H, left G 2 s. SCales: A, D = 1.0 mm; B, C = 0.5 mm." captionText-5="FIGURE 6. Parathelphusa pardus sp. nov. A, paratype female (34.7 × 26.9 mm) (ZRC 2015.0452), Kampar River, Sumatra; B, paratype female (46.1 × 34.7 mm) (ZRC 2015.0451), Kampar River, Sumatra. A, overall dorsal view; B, thoraCiC sternum showing vulvae." httpUri-0="https://zenodo.org/record/1052295/files/figure.png" httpUri-1="https://zenodo.org/record/1052297/files/figure.png" httpUri-2="https://zenodo.org/record/1052299/files/figure.png" httpUri-3="https://zenodo.org/record/1052301/files/figure.png" httpUri-4="https://zenodo.org/record/1052303/files/figure.png" httpUri-5="https://zenodo.org/record/1052305/files/figure.png" pageId="1" pageNumber="496">Figs. 1–6</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="A6FB65369372365AFF35FDFA35B3FCCB" pageId="1" pageNumber="496" type="materials_examined">
<paragraph id="EE5E36BD9372365AFF35FDFA35B3FCCB" blockId="1.[151,1437,607,2037]" pageId="1" pageNumber="496">
<materialsCitation id="5E893CE09372365AFF35FDFA35B3FCCB" ID-GBIF-Occurrence="1701200763" collectionCode="MZB" country="Indonesia" county="Pekanbaru" location="Material" municipality="Kampar River" pageId="1" pageNumber="496" specimenCode="ZRC 2015.0451, ZRC 2015.0452" specimenCount="76" specimenCount-female="23" specimenCount-juvenile="38" specimenCount-male="15" stateProvince="Riau Province" typeStatus="paratype">
<emphasis id="DC95EAAF9372365AFF35FDFA3747FDBB" bold="true" box="[151,382,607,632]" pageId="1" pageNumber="496">
<location id="EB3E60669372365AFF35FDFA36C6FDBB" LSID="urn:lsid:plazi:treatment:664887AB93723651FF35FE50354EFBD3:EB3E60669372365AFF35FDFA36C6FDBB" box="[151,255,607,632]" country="Indonesia" county="Pekanbaru" municipality="Kampar River" name="Material" pageId="1" pageNumber="496" stateProvince="Riau Province">Material</location>
examined
</emphasis>
. 
<location id="EB3E60669372365AFE33FDFA3475FDBB" LSID="urn:lsid:plazi:treatment:664887AB93723651FF35FE50354EFBD3:EB3E60669372365AFE33FDFA3475FDBB" box="[401,588,607,632]" country="Indonesia" county="Pekanbaru" municipality="Kampar River" name="Holotype-male" pageId="1" pageNumber="496" stateProvince="Riau Province">Holotype—male</location>
(37.6 × 
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) (
<collectionCode id="88F0AE789372365AFC9DFDC53544FDBB" box="[831,893,608,632]" country="Indonesia" httpUri="http://grbio.org/cool/b9gz-izvg" name="Museum Zoologicum Bogoriense" pageId="1" pageNumber="496">MZB</collectionCode>
<location id="EB3E60669372365AFC2BFDC5358AFDBB" LSID="urn:lsid:plazi:treatment:664887AB93723651FF35FE50354EFBD3:EB3E60669372365AFC2BFDC5358AFDBB" box="[905,947,608,632]" country="Indonesia" county="Pekanbaru" municipality="Kampar River" name="Cru" pageId="1" pageNumber="496" stateProvince="Riau Province">Cru</location>
4388), tributary of the 
<collectingMunicipality id="0E3AACC79372365AFB72FDC5334FFDBB" box="[1232,1398,607,632]" pageId="1" pageNumber="496">Kampar River</collectingMunicipality>
, in swamp, 
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, 
<collectingRegion id="2C25F85F9372365AFEDAFD213422FD5E" box="[376,539,644,669]" country="Indonesia" name="Riau" pageId="1" pageNumber="496">Riau Province</collectingRegion>
, in central-eastern 
<collectingRegion id="2C25F85F9372365AFD4DFD213576FD5F" box="[751,847,644,668]" country="Indonesia" name="Sumatera Utara" pageId="1" pageNumber="496">Sumatra</collectingRegion>
, 
<collectingCountry id="96F6762D9372365AFCF8FD2135F3FD5E" box="[858,970,644,669]" name="Indonesia" pageId="1" pageNumber="496">Indonesia</collectingCountry>
, coll. 
<bibRefCitation id="8A704B4C9372365AFBAFFD203280FD5F" author="Riady" box="[1037,1209,644,669]" pageId="1" pageNumber="496" refString="Riady, R., Mahatma, R. &amp; Windarti (2014) Inventarisasi kepiting air tawar di KeCamatan Kampar Utara Kabupaten Kampar Provinsi Riau. JOM (Jurnal Online Mahasiswa) FMIPA (Fakultas Matematika dan Ilmu Pengetahuan Alam), 1 (2), 471 - 479. [Universitas Riau]" type="journal article" year="2014">R. Riady, 2014</bibRefCitation>
. 
<typeStatus id="315A881F9372365AFB61FD20330DFD5E" box="[1219,1332,645,669]" pageId="1" pageNumber="496" type="paratype">Paratypes</typeStatus>
: 
<specimenCount id="F8E7FD349372365AFA9DFD2033A1FD5E" box="[1343,1432,644,669]" pageId="1" pageNumber="496" type="male">7 males</specimenCount>
, 
<specimenCount id="F8E7FD349372365AFF35FD0D373CFD03" box="[151,261,679,704]" pageId="1" pageNumber="496" type="female">8 females</specimenCount>
(
<collectionCode id="88F0AE789372365AFEB4FD0D376DFD03" box="[278,340,680,704]" country="Indonesia" httpUri="http://grbio.org/cool/b9gz-izvg" name="Museum Zoologicum Bogoriense" pageId="1" pageNumber="496">MZB</collectionCode>
Cru 4389), 
<specimenCount id="F8E7FD349372365AFE40FD0D3402FD03" box="[482,571,679,704]" pageId="1" pageNumber="496" type="male">7 males</specimenCount>
(largest 41.0 × 
<quantity id="29199B589372365AFD56FD0D3563FD7C" box="[756,858,680,704]" metricMagnitude="-2" metricUnit="m" metricValue="3.01" pageId="1" pageNumber="496" unit="mm" value="30.1">30.1 mm</quantity>
[poor condition]), 
<specimenCount id="F8E7FD349372365AFB94FD0D328BFD03" box="[1078,1202,679,704]" pageId="1" pageNumber="496" type="female">14 females</specimenCount>
(largest 46.1 × 
<quantity id="29199B589372365AFAC9FD0D36FEFD27" metricMagnitude="-2" metricUnit="m" metricValue="3.47" pageId="1" pageNumber="496" unit="mm" value="34.7">34.7 mm</quantity>
), 1 brooding female (41.5 × 
<quantity id="29199B589372365AFDB3FD693440FD27" box="[529,633,716,740]" metricMagnitude="-2" metricUnit="m" metricValue="3.13" pageId="1" pageNumber="496" unit="mm" value="31.3">31.3 mm</quantity>
, with ca. 
<specimenCount id="F8E7FD349372365AFD4BFD69354CFD26" box="[745,885,716,741]" pageId="1" pageNumber="496" type="juvenile">38 juveniles</specimenCount>
) (
<specimenCode id="BE479EC69372365AFC2FFD693270FD26" box="[909,1097,716,741]" collectionCode="ZRC" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="1" pageNumber="496">ZRC 2015.0451</specimenCode>
), 
<specimenCount id="F8E7FD349372365AFBFCFD693292FD26" box="[1118,1195,716,741]" pageId="1" pageNumber="496" type="male">1 male</specimenCount>
(34.9 × 
<quantity id="29199B589372365AFAB2FD693343FD27" box="[1296,1402,716,741]" metricMagnitude="-2" metricUnit="m" metricValue="2.66" pageId="1" pageNumber="496" unit="mm" value="26.6">26.6 mm</quantity>
), 
<specimenCount id="F8E7FD349372365AFA2CFD6936DAFCCB" pageId="1" pageNumber="496" type="female">1 female</specimenCount>
(34.7 × 
<quantity id="29199B589372365AFEE0FD4A3791FCCB" box="[322,424,751,776]" metricMagnitude="-2" metricUnit="m" metricValue="2.69" pageId="1" pageNumber="496" unit="mm" value="26.9">26.9 mm</quantity>
) (
<specimenCode id="BE479EC69372365AFE1CFD553440FCCB" box="[446,633,751,776]" collectionCode="ZRC" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="1" pageNumber="496">ZRC 2015.0452</specimenCode>
), same data as holotype.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="A6FB65369372365AFF65FCB1343EFB1E" pageId="1" pageNumber="496" type="diagnosis">
<paragraph id="EE5E36BD9372365AFF65FCB1343EFB1E" blockId="1.[151,1437,607,2037]" pageId="1" pageNumber="496">
<emphasis id="DC95EAAF9372365AFF65FCB13703FCEE" bold="true" box="[199,314,788,813]" pageId="1" pageNumber="496">Diagnosis</emphasis>
. Carapace with dorsal carapace surface distinctly convex (
<figureCitation id="76DA2A389372365AFBA2FCB13253FCEF" box="[1024,1130,788,813]" captionStart="FIGURE 1" captionStartId="2.[151,250,1966,1988]" captionTargetBox="[235,1330,193,1888]" captionTargetId="figure@2.[235,1330,193,1888]" captionTargetPageId="2" captionText="FIGURE 1. Parathelphusa pardus sp. nov., holotype male (37.6 × 28.2 mm) (MZB Cru 4388), Kampar River, Sumatra. A, overall dorsal view; B, anterior thoraCiC sternum and male abdominal somites 5, 6 and telson; C, outer view of Chelae." httpUri="https://zenodo.org/record/1052295/files/figure.png" pageId="1" pageNumber="496">Figs. 1A</figureCitation>
, 
<figureCitation id="76DA2A389372365AFBDFFCB132F0FCEF" box="[1149,1225,788,812]" captionStart="FIGURE 2" captionStartId="3.[151,250,1942,1964]" captionTargetBox="[236,1352,193,1921]" captionTargetId="figure@3.[236,1352,193,1921]" captionTargetPageId="3" captionText="FIGURE 2. Parathelphusa pardus sp. nov., paratype male (34.9 × 26.6 mm) (ZRC 2015.0452), Kampar River, Sumatra. A, overall dorsal view; B, dorsal view of CarapaCe; C, frontal view of CarapaCe." httpUri="https://zenodo.org/record/1052297/files/figure.png" pageId="1" pageNumber="496">2A, B</figureCitation>
, 
<figureCitation id="76DA2A389372365AFB79FCB13339FCEE" box="[1243,1280,788,813]" captionStart="FIGURE 6" captionStartId="7.[151,250,1796,1818]" captionTargetBox="[151,1436,213,1748]" captionTargetId="figure@7.[151,1436,213,1748]" captionTargetPageId="7" captionText="FIGURE 6. Parathelphusa pardus sp. nov. A, paratype female (34.7 × 26.9 mm) (ZRC 2015.0452), Kampar River, Sumatra; B, paratype female (46.1 × 34.7 mm) (ZRC 2015.0451), Kampar River, Sumatra. A, overall dorsal view; B, thoraCiC sternum showing vulvae." httpUri="https://zenodo.org/record/1052305/files/figure.png" pageId="1" pageNumber="496">6A</figureCitation>
); postfrontal, postorbital cristae long, sharp, almost straight or gently sinuous (
<figureCitation id="76DA2A389372365AFCCDFC9235EAFC93" box="[879,979,823,848]" captionStart="FIGURE 1" captionStartId="2.[151,250,1966,1988]" captionTargetBox="[235,1330,193,1888]" captionTargetId="figure@2.[235,1330,193,1888]" captionTargetPageId="2" captionText="FIGURE 1. Parathelphusa pardus sp. nov., holotype male (37.6 × 28.2 mm) (MZB Cru 4388), Kampar River, Sumatra. A, overall dorsal view; B, anterior thoraCiC sternum and male abdominal somites 5, 6 and telson; C, outer view of Chelae." httpUri="https://zenodo.org/record/1052295/files/figure.png" pageId="1" pageNumber="496">Figs. 1A</figureCitation>
, 
<figureCitation id="76DA2A389372365AFC7DFC9D321DFC8C" box="[991,1060,824,848]" captionStart="FIGURE 2" captionStartId="3.[151,250,1942,1964]" captionTargetBox="[236,1352,193,1921]" captionTargetId="figure@3.[236,1352,193,1921]" captionTargetPageId="3" captionText="FIGURE 2. Parathelphusa pardus sp. nov., paratype male (34.9 × 26.6 mm) (ZRC 2015.0452), Kampar River, Sumatra. A, overall dorsal view; B, dorsal view of CarapaCe; C, frontal view of CarapaCe." httpUri="https://zenodo.org/record/1052297/files/figure.png" pageId="1" pageNumber="496">2A, B</figureCitation>
, 
<figureCitation id="76DA2A389372365AFB92FC92326FFC93" box="[1072,1110,823,848]" captionStart="FIGURE 6" captionStartId="7.[151,250,1796,1818]" captionTargetBox="[151,1436,213,1748]" captionTargetId="figure@7.[151,1436,213,1748]" captionTargetPageId="7" captionText="FIGURE 6. Parathelphusa pardus sp. nov. A, paratype female (34.7 × 26.9 mm) (ZRC 2015.0452), Kampar River, Sumatra; B, paratype female (46.1 × 34.7 mm) (ZRC 2015.0451), Kampar River, Sumatra. A, overall dorsal view; B, thoraCiC sternum showing vulvae." httpUri="https://zenodo.org/record/1052305/files/figure.png" pageId="1" pageNumber="496">6A</figureCitation>
); branchial regions distinctly convex; appears gently inflated from lateral view (
<figureCitation id="76DA2A389372365AFD77FCF93503FCB7" box="[725,826,860,885]" captionStart="FIGURE 1" captionStartId="2.[151,250,1966,1988]" captionTargetBox="[235,1330,193,1888]" captionTargetId="figure@2.[235,1330,193,1888]" captionTargetPageId="2" captionText="FIGURE 1. Parathelphusa pardus sp. nov., holotype male (37.6 × 28.2 mm) (MZB Cru 4388), Kampar River, Sumatra. A, overall dorsal view; B, anterior thoraCiC sternum and male abdominal somites 5, 6 and telson; C, outer view of Chelae." httpUri="https://zenodo.org/record/1052295/files/figure.png" pageId="1" pageNumber="496">Figs. 1A</figureCitation>
, 
<figureCitation id="76DA2A389372365AFCE5FCF935B7FCB7" box="[839,910,860,884]" captionStart="FIGURE 2" captionStartId="3.[151,250,1942,1964]" captionTargetBox="[236,1352,193,1921]" captionTargetId="figure@3.[236,1352,193,1921]" captionTargetPageId="3" captionText="FIGURE 2. Parathelphusa pardus sp. nov., paratype male (34.9 × 26.6 mm) (ZRC 2015.0452), Kampar River, Sumatra. A, overall dorsal view; B, dorsal view of CarapaCe; C, frontal view of CarapaCe." httpUri="https://zenodo.org/record/1052297/files/figure.png" pageId="1" pageNumber="496">2A, B</figureCitation>
, 
<figureCitation id="76DA2A389372365AFC38FCF935F9FCB6" box="[922,960,860,885]" captionStart="FIGURE 6" captionStartId="7.[151,250,1796,1818]" captionTargetBox="[151,1436,213,1748]" captionTargetId="figure@7.[151,1436,213,1748]" captionTargetPageId="7" captionText="FIGURE 6. Parathelphusa pardus sp. nov. A, paratype female (34.7 × 26.9 mm) (ZRC 2015.0452), Kampar River, Sumatra; B, paratype female (46.1 × 34.7 mm) (ZRC 2015.0451), Kampar River, Sumatra. A, overall dorsal view; B, thoraCiC sternum showing vulvae." httpUri="https://zenodo.org/record/1052305/files/figure.png" pageId="1" pageNumber="496">6A</figureCitation>
); external orbital tooth broadly triangular, external margin gently curved, with distal two-thirds almost straight, separated from first epibranchial tooth by Ushaped notch; first epibranchial tooth acutely triangular, directed anteriorly; second epibranchial tooth acutely triangular, directed obliquely outwards (
<figureCitation id="76DA2A389372365AFDFAFC623485FC23" box="[600,700,967,992]" captionStart="FIGURE 1" captionStartId="2.[151,250,1966,1988]" captionTargetBox="[235,1330,193,1888]" captionTargetId="figure@2.[235,1330,193,1888]" captionTargetPageId="2" captionText="FIGURE 1. Parathelphusa pardus sp. nov., holotype male (37.6 × 28.2 mm) (MZB Cru 4388), Kampar River, Sumatra. A, overall dorsal view; B, anterior thoraCiC sternum and male abdominal somites 5, 6 and telson; C, outer view of Chelae." httpUri="https://zenodo.org/record/1052295/files/figure.png" pageId="1" pageNumber="496">Figs. 1A</figureCitation>
, 
<figureCitation id="76DA2A389372365AFD65FC6D3534FC1C" box="[711,781,968,992]" captionStart="FIGURE 2" captionStartId="3.[151,250,1942,1964]" captionTargetBox="[236,1352,193,1921]" captionTargetId="figure@3.[236,1352,193,1921]" captionTargetPageId="3" captionText="FIGURE 2. Parathelphusa pardus sp. nov., paratype male (34.9 × 26.6 mm) (ZRC 2015.0452), Kampar River, Sumatra. A, overall dorsal view; B, dorsal view of CarapaCe; C, frontal view of CarapaCe." httpUri="https://zenodo.org/record/1052297/files/figure.png" pageId="1" pageNumber="496">2A, B</figureCitation>
, 
<figureCitation id="76DA2A389372365AFCBAFC623507FC23" box="[792,830,967,992]" captionStart="FIGURE 6" captionStartId="7.[151,250,1796,1818]" captionTargetBox="[151,1436,213,1748]" captionTargetId="figure@7.[151,1436,213,1748]" captionTargetPageId="7" captionText="FIGURE 6. Parathelphusa pardus sp. nov. A, paratype female (34.7 × 26.9 mm) (ZRC 2015.0452), Kampar River, Sumatra; B, paratype female (46.1 × 34.7 mm) (ZRC 2015.0451), Kampar River, Sumatra. A, overall dorsal view; B, thoraCiC sternum showing vulvae." httpUri="https://zenodo.org/record/1052305/files/figure.png" pageId="1" pageNumber="496">6A</figureCitation>
); surfaces adjacent to posterolateral margin with short striae, otherwise almost smooth (
<figureCitation id="76DA2A389372365AFDAEFC493448FBC7" box="[524,625,1004,1029]" captionStart="FIGURE 1" captionStartId="2.[151,250,1966,1988]" captionTargetBox="[235,1330,193,1888]" captionTargetId="figure@2.[235,1330,193,1888]" captionTargetPageId="2" captionText="FIGURE 1. Parathelphusa pardus sp. nov., holotype male (37.6 × 28.2 mm) (MZB Cru 4388), Kampar River, Sumatra. A, overall dorsal view; B, anterior thoraCiC sternum and male abdominal somites 5, 6 and telson; C, outer view of Chelae." httpUri="https://zenodo.org/record/1052295/files/figure.png" pageId="1" pageNumber="496">Figs. 1A</figureCitation>
, 
<figureCitation id="76DA2A389372365AFDDFFC4934FDFBC7" box="[637,708,1004,1028]" captionStart="FIGURE 2" captionStartId="3.[151,250,1942,1964]" captionTargetBox="[236,1352,193,1921]" captionTargetId="figure@3.[236,1352,193,1921]" captionTargetPageId="3" captionText="FIGURE 2. Parathelphusa pardus sp. nov., paratype male (34.9 × 26.6 mm) (ZRC 2015.0452), Kampar River, Sumatra. A, overall dorsal view; B, dorsal view of CarapaCe; C, frontal view of CarapaCe." httpUri="https://zenodo.org/record/1052297/files/figure.png" pageId="1" pageNumber="496">2A, B</figureCitation>
, 
<figureCitation id="76DA2A389372365AFD72FC4934CFFBC6" box="[720,758,1004,1029]" captionStart="FIGURE 6" captionStartId="7.[151,250,1796,1818]" captionTargetBox="[151,1436,213,1748]" captionTargetId="figure@7.[151,1436,213,1748]" captionTargetPageId="7" captionText="FIGURE 6. Parathelphusa pardus sp. nov. A, paratype female (34.7 × 26.9 mm) (ZRC 2015.0452), Kampar River, Sumatra; B, paratype female (46.1 × 34.7 mm) (ZRC 2015.0451), Kampar River, Sumatra. A, overall dorsal view; B, thoraCiC sternum showing vulvae." httpUri="https://zenodo.org/record/1052305/files/figure.png" pageId="1" pageNumber="496">6A</figureCitation>
); chelae with fingers not pigmented black (
<figureCitation id="76DA2A389372365AFB42FC49335EFBC7" box="[1248,1383,1004,1029]" captionStart="FIGURE 1" captionStartId="2.[151,250,1966,1988]" captionTargetBox="[235,1330,193,1888]" captionTargetId="figure@2.[235,1330,193,1888]" captionTargetPageId="2" captionText="FIGURE 1. Parathelphusa pardus sp. nov., holotype male (37.6 × 28.2 mm) (MZB Cru 4388), Kampar River, Sumatra. A, overall dorsal view; B, anterior thoraCiC sternum and male abdominal somites 5, 6 and telson; C, outer view of Chelae." httpUri="https://zenodo.org/record/1052295/files/figure.png" pageId="1" pageNumber="496">Figs. 1A, C</figureCitation>
, 
<figureCitation id="76DA2A389372365AFAD1FC4933AEFBC7" box="[1395,1431,1004,1028]" captionStart="FIGURE 2" captionStartId="3.[151,250,1942,1964]" captionTargetBox="[236,1352,193,1921]" captionTargetId="figure@3.[236,1352,193,1921]" captionTargetPageId="3" captionText="FIGURE 2. Parathelphusa pardus sp. nov., paratype male (34.9 × 26.6 mm) (ZRC 2015.0452), Kampar River, Sumatra. A, overall dorsal view; B, dorsal view of CarapaCe; C, frontal view of CarapaCe." httpUri="https://zenodo.org/record/1052297/files/figure.png" pageId="1" pageNumber="496">2A</figureCitation>
, 
<figureCitation id="76DA2A389372365AFF35FBB536EFFBEB" box="[151,214,1040,1064]" captionStart="FIGURE 3" captionStartId="4.[151,250,1626,1648]" captionTargetBox="[151,1436,193,1605]" captionTargetId="figure@4.[151,1436,193,1605]" captionTargetPageId="4" captionText="FIGURE 3. Parathelphusa pardus sp. nov., paratype male (34.9 × 26.6 mm) (ZRC 2015.0452), Kampar River, Sumatra. A, frontal view showing frontal median triangle of front; B, left third maxilliped; C, thoraCiC sternites 1 – 4 and telson; D, anterior thoraCiC sternum and male abdominal somites 5, 6 and telson; E, posterior thoraCiC sternums and male abdominal somites 1 – 5; F, right minor Chela; G, left major Chela." httpUri="https://zenodo.org/record/1052299/files/figure.png" pageId="1" pageNumber="496">3F, G</figureCitation>
, 
<figureCitation id="76DA2A389372365AFF40FBAA3731FBEB" box="[226,264,1039,1064]" captionStart="FIGURE 6" captionStartId="7.[151,250,1796,1818]" captionTargetBox="[151,1436,213,1748]" captionTargetId="figure@7.[151,1436,213,1748]" captionTargetPageId="7" captionText="FIGURE 6. Parathelphusa pardus sp. nov. A, paratype female (34.7 × 26.9 mm) (ZRC 2015.0452), Kampar River, Sumatra; B, paratype female (46.1 × 34.7 mm) (ZRC 2015.0451), Kampar River, Sumatra. A, overall dorsal view; B, thoraCiC sternum showing vulvae." httpUri="https://zenodo.org/record/1052305/files/figure.png" pageId="1" pageNumber="496">6A</figureCitation>
); ambulatory legs relatively short, merus 1–3 with sharp subdistal spine, merus 4 with sharp angle, not spiniform (
<figureCitation id="76DA2A389372365AFEB7FB913745FB8F" box="[277,380,1076,1101]" captionStart="FIGURE 1" captionStartId="2.[151,250,1966,1988]" captionTargetBox="[235,1330,193,1888]" captionTargetId="figure@2.[235,1330,193,1888]" captionTargetPageId="2" captionText="FIGURE 1. Parathelphusa pardus sp. nov., holotype male (37.6 × 28.2 mm) (MZB Cru 4388), Kampar River, Sumatra. A, overall dorsal view; B, anterior thoraCiC sternum and male abdominal somites 5, 6 and telson; C, outer view of Chelae." httpUri="https://zenodo.org/record/1052295/files/figure.png" pageId="1" pageNumber="496">Figs. 1A</figureCitation>
, 
<figureCitation id="76DA2A389372365AFE28FB913796FB8F" box="[394,431,1076,1100]" captionStart="FIGURE 2" captionStartId="3.[151,250,1942,1964]" captionTargetBox="[236,1352,193,1921]" captionTargetId="figure@3.[236,1352,193,1921]" captionTargetPageId="3" captionText="FIGURE 2. Parathelphusa pardus sp. nov., paratype male (34.9 × 26.6 mm) (ZRC 2015.0452), Kampar River, Sumatra. A, overall dorsal view; B, dorsal view of CarapaCe; C, frontal view of CarapaCe." httpUri="https://zenodo.org/record/1052297/files/figure.png" pageId="1" pageNumber="496">2A</figureCitation>
, 
<figureCitation id="76DA2A389372365AFE1FFB9137F4FB8F" box="[445,461,1076,1100]" captionStart="FIGURE 4" captionStartId="5.[151,250,1933,1955]" captionTargetBox="[163,1423,503,1907]" captionTargetId="figure@5.[163,1423,503,1907]" captionTargetPageId="5" captionText="FIGURE 4. Parathelphusa pardus sp. nov. A – C, paratype male (34.9 × 26.6 mm) (ZRC 2015.0452), Kampar River, Sumatra; D, paratype female (34.7 × 26.9 mm) (ZRC 2015.0452), Kampar River, Sumatra. A, right fourth ambulatory leg; B, right third ambulatory leg; C, D, right seCond ambulatory legs." httpUri="https://zenodo.org/record/1052301/files/figure.png" pageId="1" pageNumber="496">4</figureCitation>
, 
<figureCitation id="76DA2A389372365AFE7EFB913438FB8E" box="[476,513,1076,1101]" captionStart="FIGURE 6" captionStartId="7.[151,250,1796,1818]" captionTargetBox="[151,1436,213,1748]" captionTargetId="figure@7.[151,1436,213,1748]" captionTargetPageId="7" captionText="FIGURE 6. Parathelphusa pardus sp. nov. A, paratype female (34.7 × 26.9 mm) (ZRC 2015.0452), Kampar River, Sumatra; B, paratype female (46.1 × 34.7 mm) (ZRC 2015.0451), Kampar River, Sumatra. A, overall dorsal view; B, thoraCiC sternum showing vulvae." httpUri="https://zenodo.org/record/1052305/files/figure.png" pageId="1" pageNumber="496">6A</figureCitation>
); suture separating thoracic sternites 2, 3 sinuous, medially concave with lateral parts convex (towards buccal cavity); abdomen with telson slightly longer than broad, somite 6 1.5–2 times as long as broad, lateral margins gently concave (
<figureCitation id="76DA2A389372365AFDC4FBD934FEFB57" box="[614,711,1148,1173]" captionStart="FIGURE 1" captionStartId="2.[151,250,1966,1988]" captionTargetBox="[235,1330,193,1888]" captionTargetId="figure@2.[235,1330,193,1888]" captionTargetPageId="2" captionText="FIGURE 1. Parathelphusa pardus sp. nov., holotype male (37.6 × 28.2 mm) (MZB Cru 4388), Kampar River, Sumatra. A, overall dorsal view; B, anterior thoraCiC sternum and male abdominal somites 5, 6 and telson; C, outer view of Chelae." httpUri="https://zenodo.org/record/1052295/files/figure.png" pageId="1" pageNumber="496">Figs. 1B</figureCitation>
, 
<figureCitation id="76DA2A389372365AFD71FBD9352FFB57" box="[723,790,1148,1172]" captionStart="FIGURE 3" captionStartId="4.[151,250,1626,1648]" captionTargetBox="[151,1436,193,1605]" captionTargetId="figure@4.[151,1436,193,1605]" captionTargetPageId="4" captionText="FIGURE 3. Parathelphusa pardus sp. nov., paratype male (34.9 × 26.6 mm) (ZRC 2015.0452), Kampar River, Sumatra. A, frontal view showing frontal median triangle of front; B, left third maxilliped; C, thoraCiC sternites 1 – 4 and telson; D, anterior thoraCiC sternum and male abdominal somites 5, 6 and telson; E, posterior thoraCiC sternums and male abdominal somites 1 – 5; F, right minor Chela; G, left major Chela." httpUri="https://zenodo.org/record/1052299/files/figure.png" pageId="1" pageNumber="496">3C–E</figureCitation>
); G1 relatively slender, almost straight, distal part slightly curved laterally, outer margin with distinct cleft on proximal part; distal part tapering to sharp tip, opening distinct, laterally positioned (
<figureCitation id="76DA2A389372365AFEDCFB6137C2FB1E" box="[382,507,1220,1245]" captionStart="FIGURE 5" captionStartId="6.[151,250,1907,1929]" captionTargetBox="[224,1324,206,1867]" captionTargetId="figure@6.[206,1359,193,1881]" captionTargetPageId="6" captionText="FIGURE 5. Parathelphusa pardus sp. nov. A – D, holotype male (37.6 × 28.2 mm) (MZB Cru 4388), Kampar River, Sumatra; E – H, paratype male (34.9 × 26.6 mm) (ZRC 2015.0452), Kampar River, Sumatra. A, E, ventral view of left G 1; B, F, ventral view of distal part of left G 1; C, G, dorsal view of distal part of left G 1; D, H, left G 2 s. SCales: A, D = 1.0 mm; B, C = 0.5 mm." httpUri="https://zenodo.org/record/1052303/files/figure.png" pageId="1" pageNumber="496">Fig. 5A–G</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="A6FB653693723653FF65FB4237D4FBF6" lastPageId="8" lastPageNumber="503" pageId="1" pageNumber="496" type="description">
<paragraph id="EE5E36BD9372365AFF65FB423535F8A7" blockId="1.[151,1437,607,2037]" pageId="1" pageNumber="496">
<emphasis id="DC95EAAF9372365AFF65FB423781FAC3" bold="true" box="[199,440,1255,1280]" pageId="1" pageNumber="496">Description of male</emphasis>
. Carapace hexagonal; transversely wider than long; dorsal carapace surface distinctly convex, smooth; regions poorly demarcated; cervical groove broad, shallow; median H-shaped gastro-cardiac depression distinct (
<figureCitation id="76DA2A389372365AFEDDFA8A37DFFA8B" box="[383,486,1327,1352]" captionStart="FIGURE 1" captionStartId="2.[151,250,1966,1988]" captionTargetBox="[235,1330,193,1888]" captionTargetId="figure@2.[235,1330,193,1888]" captionTargetPageId="2" captionText="FIGURE 1. Parathelphusa pardus sp. nov., holotype male (37.6 × 28.2 mm) (MZB Cru 4388), Kampar River, Sumatra. A, overall dorsal view; B, anterior thoraCiC sternum and male abdominal somites 5, 6 and telson; C, outer view of Chelae." httpUri="https://zenodo.org/record/1052295/files/figure.png" pageId="1" pageNumber="496">Figs. 1A</figureCitation>
, 
<figureCitation id="76DA2A389372365AFE54FA953407FA8B" box="[502,574,1328,1352]" captionStart="FIGURE 2" captionStartId="3.[151,250,1942,1964]" captionTargetBox="[236,1352,193,1921]" captionTargetId="figure@3.[236,1352,193,1921]" captionTargetPageId="3" captionText="FIGURE 2. Parathelphusa pardus sp. nov., paratype male (34.9 × 26.6 mm) (ZRC 2015.0452), Kampar River, Sumatra. A, overall dorsal view; B, dorsal view of CarapaCe; C, frontal view of CarapaCe." httpUri="https://zenodo.org/record/1052297/files/figure.png" pageId="1" pageNumber="496">2A, B</figureCitation>
, 
<figureCitation id="76DA2A389372365AFDECFA8A344AFA8B" box="[590,627,1327,1352]" captionStart="FIGURE 6" captionStartId="7.[151,250,1796,1818]" captionTargetBox="[151,1436,213,1748]" captionTargetId="figure@7.[151,1436,213,1748]" captionTargetPageId="7" captionText="FIGURE 6. Parathelphusa pardus sp. nov. A, paratype female (34.7 × 26.9 mm) (ZRC 2015.0452), Kampar River, Sumatra; B, paratype female (46.1 × 34.7 mm) (ZRC 2015.0451), Kampar River, Sumatra. A, overall dorsal view; B, thoraCiC sternum showing vulvae." httpUri="https://zenodo.org/record/1052305/files/figure.png" pageId="1" pageNumber="496">6A</figureCitation>
). Frontal margin broad, contiguous with supraorbital margin, cristate, appearing almost straight from dorsal view, sinuous from frontal view; frontal median triangle distinct, broad (
<figureCitation id="76DA2A389372365AFAC6FAF13682FA53" captionStart="FIGURE 2" captionStartId="3.[151,250,1942,1964]" captionTargetBox="[236,1352,193,1921]" captionTargetId="figure@3.[236,1352,193,1921]" captionTargetPageId="3" captionText="FIGURE 2. Parathelphusa pardus sp. nov., paratype male (34.9 × 26.6 mm) (ZRC 2015.0452), Kampar River, Sumatra. A, overall dorsal view; B, dorsal view of CarapaCe; C, frontal view of CarapaCe." httpUri="https://zenodo.org/record/1052297/files/figure.png" pageId="1" pageNumber="496">Figs. 2A</figureCitation>
, 
<figureCitation id="76DA2A389372365AFF6BFADD36D7FA53" box="[201,238,1400,1424]" captionStart="FIGURE 3" captionStartId="4.[151,250,1626,1648]" captionTargetBox="[151,1436,193,1605]" captionTargetId="figure@4.[151,1436,193,1605]" captionTargetPageId="4" captionText="FIGURE 3. Parathelphusa pardus sp. nov., paratype male (34.9 × 26.6 mm) (ZRC 2015.0452), Kampar River, Sumatra. A, frontal view showing frontal median triangle of front; B, left third maxilliped; C, thoraCiC sternites 1 – 4 and telson; D, anterior thoraCiC sternum and male abdominal somites 5, 6 and telson; E, posterior thoraCiC sternums and male abdominal somites 1 – 5; F, right minor Chela; G, left major Chela." httpUri="https://zenodo.org/record/1052299/files/figure.png" pageId="1" pageNumber="496">3A</figureCitation>
). Postfrontal cristae sharp, almost straight separated from each other medially by short, narrow Y-shaped fissure; postorbital cristae long, sharp, almost straight or gently sinuous, contiguous with postfrontal cristae, cristae ending before of base of first epibranchial tooth (
<figureCitation id="76DA2A389372365AFD15FA1A3523FA1B" box="[695,794,1471,1496]" captionStart="FIGURE 1" captionStartId="2.[151,250,1966,1988]" captionTargetBox="[235,1330,193,1888]" captionTargetId="figure@2.[235,1330,193,1888]" captionTargetPageId="2" captionText="FIGURE 1. Parathelphusa pardus sp. nov., holotype male (37.6 × 28.2 mm) (MZB Cru 4388), Kampar River, Sumatra. A, overall dorsal view; B, anterior thoraCiC sternum and male abdominal somites 5, 6 and telson; C, outer view of Chelae." httpUri="https://zenodo.org/record/1052295/files/figure.png" pageId="1" pageNumber="496">Figs. 1A</figureCitation>
, 
<figureCitation id="76DA2A389372365AFC87FA653550FA1B" box="[805,873,1472,1496]" captionStart="FIGURE 2" captionStartId="3.[151,250,1942,1964]" captionTargetBox="[236,1352,193,1921]" captionTargetId="figure@3.[236,1352,193,1921]" captionTargetPageId="3" captionText="FIGURE 2. Parathelphusa pardus sp. nov., paratype male (34.9 × 26.6 mm) (ZRC 2015.0452), Kampar River, Sumatra. A, overall dorsal view; B, dorsal view of CarapaCe; C, frontal view of CarapaCe." httpUri="https://zenodo.org/record/1052297/files/figure.png" pageId="1" pageNumber="496">2A, B</figureCitation>
, 
<figureCitation id="76DA2A389372365AFCD6FA1A35A3FA1B" box="[884,922,1471,1496]" captionStart="FIGURE 6" captionStartId="7.[151,250,1796,1818]" captionTargetBox="[151,1436,213,1748]" captionTargetId="figure@7.[151,1436,213,1748]" captionTargetPageId="7" captionText="FIGURE 6. Parathelphusa pardus sp. nov. A, paratype female (34.7 × 26.9 mm) (ZRC 2015.0452), Kampar River, Sumatra; B, paratype female (46.1 × 34.7 mm) (ZRC 2015.0451), Kampar River, Sumatra. A, overall dorsal view; B, thoraCiC sternum showing vulvae." httpUri="https://zenodo.org/record/1052305/files/figure.png" pageId="1" pageNumber="496">6A</figureCitation>
). Branchial regions distinctly convex; appears gently inflated from lateral view (
<figureCitation id="76DA2A389372365AFDBAFA413447FA3F" box="[536,638,1508,1533]" captionStart="FIGURE 1" captionStartId="2.[151,250,1966,1988]" captionTargetBox="[235,1330,193,1888]" captionTargetId="figure@2.[235,1330,193,1888]" captionTargetPageId="2" captionText="FIGURE 1. Parathelphusa pardus sp. nov., holotype male (37.6 × 28.2 mm) (MZB Cru 4388), Kampar River, Sumatra. A, overall dorsal view; B, anterior thoraCiC sternum and male abdominal somites 5, 6 and telson; C, outer view of Chelae." httpUri="https://zenodo.org/record/1052295/files/figure.png" pageId="1" pageNumber="496">Figs. 1A</figureCitation>
, 
<figureCitation id="76DA2A389372365AFD29FA4134EBFA3F" box="[651,722,1508,1532]" captionStart="FIGURE 2" captionStartId="3.[151,250,1942,1964]" captionTargetBox="[236,1352,193,1921]" captionTargetId="figure@3.[236,1352,193,1921]" captionTargetPageId="3" captionText="FIGURE 2. Parathelphusa pardus sp. nov., paratype male (34.9 × 26.6 mm) (ZRC 2015.0452), Kampar River, Sumatra. A, overall dorsal view; B, dorsal view of CarapaCe; C, frontal view of CarapaCe." httpUri="https://zenodo.org/record/1052297/files/figure.png" pageId="1" pageNumber="496">2A, B</figureCitation>
, 
<figureCitation id="76DA2A389372365AFD7DFA41353CFA3E" box="[735,773,1508,1533]" captionStart="FIGURE 6" captionStartId="7.[151,250,1796,1818]" captionTargetBox="[151,1436,213,1748]" captionTargetId="figure@7.[151,1436,213,1748]" captionTargetPageId="7" captionText="FIGURE 6. Parathelphusa pardus sp. nov. A, paratype female (34.7 × 26.9 mm) (ZRC 2015.0452), Kampar River, Sumatra; B, paratype female (46.1 × 34.7 mm) (ZRC 2015.0451), Kampar River, Sumatra. A, overall dorsal view; B, thoraCiC sternum showing vulvae." httpUri="https://zenodo.org/record/1052305/files/figure.png" pageId="1" pageNumber="496">6A</figureCitation>
). Orbits large, ovate; eyes well developed with pigmented cornea, completely filling orbit (
<figureCitation id="76DA2A389372365AFDA5F9A23452F9E3" box="[519,619,1543,1568]" captionStart="FIGURE 1" captionStartId="2.[151,250,1966,1988]" captionTargetBox="[235,1330,193,1888]" captionTargetId="figure@2.[235,1330,193,1888]" captionTargetPageId="2" captionText="FIGURE 1. Parathelphusa pardus sp. nov., holotype male (37.6 × 28.2 mm) (MZB Cru 4388), Kampar River, Sumatra. A, overall dorsal view; B, anterior thoraCiC sternum and male abdominal somites 5, 6 and telson; C, outer view of Chelae." httpUri="https://zenodo.org/record/1052295/files/figure.png" pageId="1" pageNumber="496">Figs. 1A</figureCitation>
, 
<figureCitation id="76DA2A389372365AFDDBF9AD34B0F9E3" box="[633,649,1544,1568]" captionStart="FIGURE 2" captionStartId="3.[151,250,1942,1964]" captionTargetBox="[236,1352,193,1921]" captionTargetId="figure@3.[236,1352,193,1921]" captionTargetPageId="3" captionText="FIGURE 2. Parathelphusa pardus sp. nov., paratype male (34.9 × 26.6 mm) (ZRC 2015.0452), Kampar River, Sumatra. A, overall dorsal view; B, dorsal view of CarapaCe; C, frontal view of CarapaCe." httpUri="https://zenodo.org/record/1052297/files/figure.png" pageId="1" pageNumber="496">2</figureCitation>
, 
<figureCitation id="76DA2A389372365AFD34F9AD3483F9E3" box="[662,698,1544,1568]" captionStart="FIGURE 3" captionStartId="4.[151,250,1626,1648]" captionTargetBox="[151,1436,193,1605]" captionTargetId="figure@4.[151,1436,193,1605]" captionTargetPageId="4" captionText="FIGURE 3. Parathelphusa pardus sp. nov., paratype male (34.9 × 26.6 mm) (ZRC 2015.0452), Kampar River, Sumatra. A, frontal view showing frontal median triangle of front; B, left third maxilliped; C, thoraCiC sternites 1 – 4 and telson; D, anterior thoraCiC sternum and male abdominal somites 5, 6 and telson; E, posterior thoraCiC sternums and male abdominal somites 1 – 5; F, right minor Chela; G, left major Chela." httpUri="https://zenodo.org/record/1052299/files/figure.png" pageId="1" pageNumber="496">3A</figureCitation>
, 
<figureCitation id="76DA2A389372365AFD65F9A234D4F9E3" box="[711,749,1543,1568]" captionStart="FIGURE 6" captionStartId="7.[151,250,1796,1818]" captionTargetBox="[151,1436,213,1748]" captionTargetId="figure@7.[151,1436,213,1748]" captionTargetPageId="7" captionText="FIGURE 6. Parathelphusa pardus sp. nov. A, paratype female (34.7 × 26.9 mm) (ZRC 2015.0452), Kampar River, Sumatra; B, paratype female (46.1 × 34.7 mm) (ZRC 2015.0451), Kampar River, Sumatra. A, overall dorsal view; B, thoraCiC sternum showing vulvae." httpUri="https://zenodo.org/record/1052305/files/figure.png" pageId="1" pageNumber="496">6A</figureCitation>
). Supraorbital margin concave, entire; confluent with frontal margin; suborbital margin concave, entire (
<figureCitation id="76DA2A389372365AFDD5F98934E3F987" box="[631,730,1580,1605]" captionStart="FIGURE 1" captionStartId="2.[151,250,1966,1988]" captionTargetBox="[235,1330,193,1888]" captionTargetId="figure@2.[235,1330,193,1888]" captionTargetPageId="2" captionText="FIGURE 1. Parathelphusa pardus sp. nov., holotype male (37.6 × 28.2 mm) (MZB Cru 4388), Kampar River, Sumatra. A, overall dorsal view; B, anterior thoraCiC sternum and male abdominal somites 5, 6 and telson; C, outer view of Chelae." httpUri="https://zenodo.org/record/1052295/files/figure.png" pageId="1" pageNumber="496">Figs. 1A</figureCitation>
, 
<figureCitation id="76DA2A389372365AFD47F9893510F987" box="[741,809,1580,1604]" captionStart="FIGURE 2" captionStartId="3.[151,250,1942,1964]" captionTargetBox="[236,1352,193,1921]" captionTargetId="figure@3.[236,1352,193,1921]" captionTargetPageId="3" captionText="FIGURE 2. Parathelphusa pardus sp. nov., paratype male (34.9 × 26.6 mm) (ZRC 2015.0452), Kampar River, Sumatra. A, overall dorsal view; B, dorsal view of CarapaCe; C, frontal view of CarapaCe." httpUri="https://zenodo.org/record/1052297/files/figure.png" pageId="1" pageNumber="496">2A, B</figureCitation>
, 
<figureCitation id="76DA2A389372365AFC96F9893563F986" box="[820,858,1580,1605]" captionStart="FIGURE 6" captionStartId="7.[151,250,1796,1818]" captionTargetBox="[151,1436,213,1748]" captionTargetId="figure@7.[151,1436,213,1748]" captionTargetPageId="7" captionText="FIGURE 6. Parathelphusa pardus sp. nov. A, paratype female (34.7 × 26.9 mm) (ZRC 2015.0452), Kampar River, Sumatra; B, paratype female (46.1 × 34.7 mm) (ZRC 2015.0451), Kampar River, Sumatra. A, overall dorsal view; B, thoraCiC sternum showing vulvae." httpUri="https://zenodo.org/record/1052305/files/figure.png" pageId="1" pageNumber="496">6A</figureCitation>
). Pterygostomian region covered with scattered low granules, short striae; suborbital, subhepatic regions with low striae, granules (
<figureCitation id="76DA2A389372365AFBA4F9EA3251F9AB" box="[1030,1128,1615,1640]" captionStart="FIGURE 2" captionStartId="3.[151,250,1942,1964]" captionTargetBox="[236,1352,193,1921]" captionTargetId="figure@3.[236,1352,193,1921]" captionTargetPageId="3" captionText="FIGURE 2. Parathelphusa pardus sp. nov., paratype male (34.9 × 26.6 mm) (ZRC 2015.0452), Kampar River, Sumatra. A, overall dorsal view; B, dorsal view of CarapaCe; C, frontal view of CarapaCe." httpUri="https://zenodo.org/record/1052297/files/figure.png" pageId="1" pageNumber="496">Figs. 2C</figureCitation>
, 
<figureCitation id="76DA2A389372365AFBD1F9F532A0F9AB" box="[1139,1177,1616,1640]" captionStart="FIGURE 3" captionStartId="4.[151,250,1626,1648]" captionTargetBox="[151,1436,193,1605]" captionTargetId="figure@4.[151,1436,193,1605]" captionTargetPageId="4" captionText="FIGURE 3. Parathelphusa pardus sp. nov., paratype male (34.9 × 26.6 mm) (ZRC 2015.0452), Kampar River, Sumatra. A, frontal view showing frontal median triangle of front; B, left third maxilliped; C, thoraCiC sternites 1 – 4 and telson; D, anterior thoraCiC sternum and male abdominal somites 5, 6 and telson; E, posterior thoraCiC sternums and male abdominal somites 1 – 5; F, right minor Chela; G, left major Chela." httpUri="https://zenodo.org/record/1052299/files/figure.png" pageId="1" pageNumber="496">3A</figureCitation>
). External orbital tooth broadly triangular, external margin gently curved, with distal two-thirds almost straight; inner angle triangular, extending to level of frontal margin, separated from first epibranchial tooth by U-shaped notch; first epibranchial tooth sharp, acutely triangular, directed anteriorly; second epibranchial tooth sharp, acutely triangular, directed obliquely outwards (
<figureCitation id="76DA2A389372365AFE27F97A37D4F93B" box="[389,493,1759,1784]" captionStart="FIGURE 1" captionStartId="2.[151,250,1966,1988]" captionTargetBox="[235,1330,193,1888]" captionTargetId="figure@2.[235,1330,193,1888]" captionTargetPageId="2" captionText="FIGURE 1. Parathelphusa pardus sp. nov., holotype male (37.6 × 28.2 mm) (MZB Cru 4388), Kampar River, Sumatra. A, overall dorsal view; B, anterior thoraCiC sternum and male abdominal somites 5, 6 and telson; C, outer view of Chelae." httpUri="https://zenodo.org/record/1052295/files/figure.png" pageId="1" pageNumber="496">Figs. 1A</figureCitation>
, 
<figureCitation id="76DA2A389372365AFE5FF945347FF93B" box="[509,582,1760,1784]" captionStart="FIGURE 2" captionStartId="3.[151,250,1942,1964]" captionTargetBox="[236,1352,193,1921]" captionTargetId="figure@3.[236,1352,193,1921]" captionTargetPageId="3" captionText="FIGURE 2. Parathelphusa pardus sp. nov., paratype male (34.9 × 26.6 mm) (ZRC 2015.0452), Kampar River, Sumatra. A, overall dorsal view; B, dorsal view of CarapaCe; C, frontal view of CarapaCe." httpUri="https://zenodo.org/record/1052297/files/figure.png" pageId="1" pageNumber="496">2A, B</figureCitation>
, 
<figureCitation id="76DA2A389372365AFDF4F97A3445F93B" box="[598,636,1759,1784]" captionStart="FIGURE 6" captionStartId="7.[151,250,1796,1818]" captionTargetBox="[151,1436,213,1748]" captionTargetId="figure@7.[151,1436,213,1748]" captionTargetPageId="7" captionText="FIGURE 6. Parathelphusa pardus sp. nov. A, paratype female (34.7 × 26.9 mm) (ZRC 2015.0452), Kampar River, Sumatra; B, paratype female (46.1 × 34.7 mm) (ZRC 2015.0451), Kampar River, Sumatra. A, overall dorsal view; B, thoraCiC sternum showing vulvae." httpUri="https://zenodo.org/record/1052305/files/figure.png" pageId="1" pageNumber="496">6A</figureCitation>
). Posterolateral margin gently concave, without visible crista, lateral surfaces with short striae, otherwise almost smooth; posterolateral margins converging towards almost straight posterior carapace margin (
<figureCitation id="76DA2A389372365AFE6DF882340CF883" box="[463,565,1831,1856]" captionStart="FIGURE 1" captionStartId="2.[151,250,1966,1988]" captionTargetBox="[235,1330,193,1888]" captionTargetId="figure@2.[235,1330,193,1888]" captionTargetPageId="2" captionText="FIGURE 1. Parathelphusa pardus sp. nov., holotype male (37.6 × 28.2 mm) (MZB Cru 4388), Kampar River, Sumatra. A, overall dorsal view; B, anterior thoraCiC sternum and male abdominal somites 5, 6 and telson; C, outer view of Chelae." httpUri="https://zenodo.org/record/1052295/files/figure.png" pageId="1" pageNumber="496">Figs. 1A</figureCitation>
, 
<figureCitation id="76DA2A389372365AFDE1F88D34B2F883" box="[579,651,1832,1856]" captionStart="FIGURE 2" captionStartId="3.[151,250,1942,1964]" captionTargetBox="[236,1352,193,1921]" captionTargetId="figure@3.[236,1352,193,1921]" captionTargetPageId="3" captionText="FIGURE 2. Parathelphusa pardus sp. nov., paratype male (34.9 × 26.6 mm) (ZRC 2015.0452), Kampar River, Sumatra. A, overall dorsal view; B, dorsal view of CarapaCe; C, frontal view of CarapaCe." httpUri="https://zenodo.org/record/1052297/files/figure.png" pageId="1" pageNumber="496">2A, B</figureCitation>
, 
<figureCitation id="76DA2A389372365AFD3BF8823486F883" box="[665,703,1831,1856]" captionStart="FIGURE 6" captionStartId="7.[151,250,1796,1818]" captionTargetBox="[151,1436,213,1748]" captionTargetId="figure@7.[151,1436,213,1748]" captionTargetPageId="7" captionText="FIGURE 6. Parathelphusa pardus sp. nov. A, paratype female (34.7 × 26.9 mm) (ZRC 2015.0452), Kampar River, Sumatra; B, paratype female (46.1 × 34.7 mm) (ZRC 2015.0451), Kampar River, Sumatra. A, overall dorsal view; B, thoraCiC sternum showing vulvae." httpUri="https://zenodo.org/record/1052305/files/figure.png" pageId="1" pageNumber="496">6A</figureCitation>
). Posterior margin of epistome sinuous; median tooth triangular with rounded tip, lateral margins convex (
<figureCitation id="76DA2A389372365AFDCEF8E934F7F8A7" box="[620,718,1868,1893]" captionStart="FIGURE 2" captionStartId="3.[151,250,1942,1964]" captionTargetBox="[236,1352,193,1921]" captionTargetId="figure@3.[236,1352,193,1921]" captionTargetPageId="3" captionText="FIGURE 2. Parathelphusa pardus sp. nov., paratype male (34.9 × 26.6 mm) (ZRC 2015.0452), Kampar River, Sumatra. A, overall dorsal view; B, dorsal view of CarapaCe; C, frontal view of CarapaCe." httpUri="https://zenodo.org/record/1052297/files/figure.png" pageId="1" pageNumber="496">Figs. 2C</figureCitation>
, 
<figureCitation id="76DA2A389372365AFD78F8E934C6F8A7" box="[730,767,1868,1892]" captionStart="FIGURE 3" captionStartId="4.[151,250,1626,1648]" captionTargetBox="[151,1436,193,1605]" captionTargetId="figure@4.[151,1436,193,1605]" captionTargetPageId="4" captionText="FIGURE 3. Parathelphusa pardus sp. nov., paratype male (34.9 × 26.6 mm) (ZRC 2015.0452), Kampar River, Sumatra. A, frontal view showing frontal median triangle of front; B, left third maxilliped; C, thoraCiC sternites 1 – 4 and telson; D, anterior thoraCiC sternum and male abdominal somites 5, 6 and telson; E, posterior thoraCiC sternums and male abdominal somites 1 – 5; F, right minor Chela; G, left major Chela." httpUri="https://zenodo.org/record/1052299/files/figure.png" pageId="1" pageNumber="496">3A</figureCitation>
).
</paragraph>
<paragraph id="EE5E36BD9372365AFF65F8CA3505F837" blockId="1.[151,1437,607,2037]" pageId="1" pageNumber="496">
Third maxilliped elongated, completely covering buccal cavity when closed; merus subquadrate, as wide as long, median part gently depressed, anterolateral margin slightly auriculiform, rounded; ischium rectangular, with deep submedian oblique sulcus; exopod relatively stout, reaching to median part of merus, with distinct subterminal tooth on inner margin, flagellum long (
<figureCitation id="76DA2A389372365AFD75F8793517F837" box="[727,814,2012,2037]" captionStart="FIGURE 3" captionStartId="4.[151,250,1626,1648]" captionTargetBox="[151,1436,193,1605]" captionTargetId="figure@4.[151,1436,193,1605]" captionTargetPageId="4" captionText="FIGURE 3. Parathelphusa pardus sp. nov., paratype male (34.9 × 26.6 mm) (ZRC 2015.0452), Kampar River, Sumatra. A, frontal view showing frontal median triangle of front; B, left third maxilliped; C, thoraCiC sternites 1 – 4 and telson; D, anterior thoraCiC sternum and male abdominal somites 5, 6 and telson; E, posterior thoraCiC sternums and male abdominal somites 1 – 5; F, right minor Chela; G, left major Chela." httpUri="https://zenodo.org/record/1052299/files/figure.png" pageId="1" pageNumber="496">Fig. 3B</figureCitation>
).
</paragraph>
<caption id="BA9E663593713659FF35F80B337AF820" httpUri="https://zenodo.org/record/1052295/files/figure.png" pageId="2" pageNumber="497" targetBox="[235,1330,193,1888]" targetPageId="2">
<paragraph id="EE5E36BD93713659FF35F80B337AF820" blockId="2.[151,1435,1966,2019]" pageId="2" pageNumber="497">
<emphasis id="DC95EAAF93713659FF35F80B3737F807" bold="true" box="[151,270,1966,1988]" pageId="2" pageNumber="497">FIGURE 1</emphasis>
. 
<taxonomicName id="29E14D3E93713659FEBEF80B37C6F807" authority="Ng &amp; Riady, 2016" authorityName="Ng &amp; Riady" authorityYear="2016" box="[284,511,1966,1988]" class="Malacostraca" family="Gecarcinucidae" genus="Parathelphusa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="497" phylum="Arthropoda" rank="species" species="pardus" status="sp. nov.">
<emphasis id="DC95EAAF93713659FEBEF80B37C6F807" box="[284,511,1966,1988]" italics="true" pageId="2" pageNumber="497">Parathelphusa pardus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="C7A657D493713659FDA5F80A3460F807" box="[519,601,1967,1988]" pageId="2" pageNumber="497" rank="species">
<emphasis id="DC95EAAF93713659FDA5F80A3460F807" bold="true" box="[519,601,1967,1988]" pageId="2" pageNumber="497">sp. nov.</emphasis>
</taxonomicNameLabel>
, holotype male (37.6 × 28.2 mm) (MZB Cru 4388), Kampar River, Sumatra. A, overall dorsal view; B, anterior thoraCiC sternum and male abdominal somites 5, 6 and telson; C, outer view of Chelae.
</paragraph>
</caption>
<caption id="BA9E663593703658FF35F83335A2F809" httpUri="https://zenodo.org/record/1052297/files/figure.png" pageId="3" pageNumber="498" targetBox="[236,1352,193,1921]" targetPageId="3">
<paragraph id="EE5E36BD93703658FF35F83335A2F809" blockId="3.[151,1435,1942,1994]" pageId="3" pageNumber="498">
<emphasis id="DC95EAAF93703658FF35F8333737F868" bold="true" box="[151,270,1942,1964]" pageId="3" pageNumber="498">FIGURE 2</emphasis>
. 
<taxonomicName id="29E14D3E93703658FEBEF83337C7F86F" authority="Ng &amp; Riady, 2016" authorityName="Ng &amp; Riady" authorityYear="2016" box="[284,510,1942,1964]" class="Malacostraca" family="Gecarcinucidae" genus="Parathelphusa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="498" phylum="Arthropoda" rank="species" species="pardus" status="sp. nov.">
<emphasis id="DC95EAAF93703658FEBEF83337C7F86F" box="[284,510,1942,1964]" italics="true" pageId="3" pageNumber="498">Parathelphusa pardus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="C7A657D493703658FDA4F8323461F86F" box="[518,600,1943,1964]" pageId="3" pageNumber="498" rank="species">
<emphasis id="DC95EAAF93703658FDA4F8323461F86F" bold="true" box="[518,600,1943,1964]" pageId="3" pageNumber="498">sp. nov.</emphasis>
</taxonomicNameLabel>
, paratype male (34.9 × 26.6 mm) (ZRC 2015.0452), Kampar River, Sumatra. A, overall dorsal view; B, dorsal view of CarapaCe; C, frontal view of CarapaCe.
</paragraph>
</caption>
<caption id="BA9E66359377365FFF35F9FF3417F90F" httpUri="https://zenodo.org/record/1052299/files/figure.png" pageId="4" pageNumber="499" targetBox="[151,1436,193,1605]" targetPageId="4">
<paragraph id="EE5E36BD9377365FFF35F9FF3417F90F" blockId="4.[151,1436,1626,1740]" pageId="4" pageNumber="499">
<emphasis id="DC95EAAF9377365FFF35F9FF3737F9AC" bold="true" box="[151,270,1626,1648]" pageId="4" pageNumber="499">FIGURE 3</emphasis>
. 
<taxonomicName id="29E14D3E9377365FFEBEF9FF37C7F9B3" authority="Ng &amp; Riady, 2016" authorityName="Ng &amp; Riady" authorityYear="2016" box="[284,510,1626,1648]" class="Malacostraca" family="Gecarcinucidae" genus="Parathelphusa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="499" phylum="Arthropoda" rank="species" species="pardus" status="sp. nov.">
<emphasis id="DC95EAAF9377365FFEBEF9FF37C7F9B3" box="[284,510,1626,1648]" italics="true" pageId="4" pageNumber="499">Parathelphusa pardus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="C7A657D49377365FFDA4F9FE3461F9B3" box="[518,600,1627,1648]" pageId="4" pageNumber="499" rank="species">
<emphasis id="DC95EAAF9377365FFDA4F9FE3461F9B3" bold="true" box="[518,600,1627,1648]" pageId="4" pageNumber="499">sp. nov.</emphasis>
</taxonomicNameLabel>
, paratype male (34.9 × 26.6 mm) (ZRC 2015.0452), Kampar River, Sumatra. A, frontal view showing frontal median triangle of front; B, left third maxilliped; C, thoraCiC sternites 1–4 and telson; D, anterior thoraCiC sternum and male abdominal somites 5, 6 and telson; E, posterior thoraCiC sternums and male abdominal somites 1–5; F, right minor Chela; G, left major Chela.
</paragraph>
</caption>
<paragraph id="EE5E36BD9377365FFF65F95C3512F861" blockId="4.[151,1436,1785,2026]" pageId="4" pageNumber="499">
Chelipeds asymmetrical in adult males; merus relatively short; with distinct subdistal spine; carpus subovate, outer surface rugose, inner distal angle with prominent, elongated spine, inner margin gently serrated; chelae enlarged, outer surface smooth or almost so; fingers not pigmented black, those on major chela of larger males gently curved, subequal or slightly longer than palm, forming basal gap when closed; cutting edges with small and large rounded teeth, denticles (
<figureCitation id="76DA2A389377365FFE4DF82C344DF861" box="[495,628,1929,1954]" captionStart="FIGURE 1" captionStartId="2.[151,250,1966,1988]" captionTargetBox="[235,1330,193,1888]" captionTargetId="figure@2.[235,1330,193,1888]" captionTargetPageId="2" captionText="FIGURE 1. Parathelphusa pardus sp. nov., holotype male (37.6 × 28.2 mm) (MZB Cru 4388), Kampar River, Sumatra. A, overall dorsal view; B, anterior thoraCiC sternum and male abdominal somites 5, 6 and telson; C, outer view of Chelae." httpUri="https://zenodo.org/record/1052295/files/figure.png" pageId="4" pageNumber="499">Figs. 1A, C</figureCitation>
, 
<figureCitation id="76DA2A389377365FFD22F82F349DF861" box="[640,676,1930,1954]" captionStart="FIGURE 2" captionStartId="3.[151,250,1942,1964]" captionTargetBox="[236,1352,193,1921]" captionTargetId="figure@3.[236,1352,193,1921]" captionTargetPageId="3" captionText="FIGURE 2. Parathelphusa pardus sp. nov., paratype male (34.9 × 26.6 mm) (ZRC 2015.0452), Kampar River, Sumatra. A, overall dorsal view; B, dorsal view of CarapaCe; C, frontal view of CarapaCe." httpUri="https://zenodo.org/record/1052297/files/figure.png" pageId="4" pageNumber="499">2A</figureCitation>
, 
<figureCitation id="76DA2A389377365FFD12F82F34D4F861" box="[688,749,1930,1954]" captionStart="FIGURE 3" captionStartId="4.[151,250,1626,1648]" captionTargetBox="[151,1436,193,1605]" captionTargetId="figure@4.[151,1436,193,1605]" captionTargetPageId="4" captionText="FIGURE 3. Parathelphusa pardus sp. nov., paratype male (34.9 × 26.6 mm) (ZRC 2015.0452), Kampar River, Sumatra. A, frontal view showing frontal median triangle of front; B, left third maxilliped; C, thoraCiC sternites 1 – 4 and telson; D, anterior thoraCiC sternum and male abdominal somites 5, 6 and telson; E, posterior thoraCiC sternums and male abdominal somites 1 – 5; F, right minor Chela; G, left major Chela." httpUri="https://zenodo.org/record/1052299/files/figure.png" pageId="4" pageNumber="499">3F, G</figureCitation>
, 
<figureCitation id="76DA2A389377365FFD5AF82C3527F861" box="[760,798,1929,1954]" captionStart="FIGURE 6" captionStartId="7.[151,250,1796,1818]" captionTargetBox="[151,1436,213,1748]" captionTargetId="figure@7.[151,1436,213,1748]" captionTargetPageId="7" captionText="FIGURE 6. Parathelphusa pardus sp. nov. A, paratype female (34.7 × 26.9 mm) (ZRC 2015.0452), Kampar River, Sumatra; B, paratype female (46.1 × 34.7 mm) (ZRC 2015.0451), Kampar River, Sumatra. A, overall dorsal view; B, thoraCiC sternum showing vulvae." httpUri="https://zenodo.org/record/1052305/files/figure.png" pageId="4" pageNumber="499">6A</figureCitation>
).
</paragraph>
<paragraph id="EE5E36BD9377365EFF65F808372FFEDE" blockId="4.[151,1436,1785,2026]" lastBlockId="5.[151,1436,151,464]" lastPageId="5" lastPageNumber="500" pageId="4" pageNumber="499">
Ambulatory legs relatively short, stout; second leg longest; merus subrectangular, margins cristate, dorsal margins of meri 1–3 with sharp subdistal spine or angle, that of merus 4 with sharp angle, not spiniform; carpus unarmed, with cristate dorsal margin, subparallel crista on outer median surface; propodus with dorsal margin cristate, ventral margin gently serrated; outer surface with shallow submedian longitudinal depression; dactylus gently curved with corneous tip, quadrate in cross-section, margins with 4 rows of strong sharp spines (
<figureCitation id="76DA2A389376365EFA90FF7A33AEFF3B" box="[1330,1431,223,248]" captionStart="FIGURE 1" captionStartId="2.[151,250,1966,1988]" captionTargetBox="[235,1330,193,1888]" captionTargetId="figure@2.[235,1330,193,1888]" captionTargetPageId="2" captionText="FIGURE 1. Parathelphusa pardus sp. nov., holotype male (37.6 × 28.2 mm) (MZB Cru 4388), Kampar River, Sumatra. A, overall dorsal view; B, anterior thoraCiC sternum and male abdominal somites 5, 6 and telson; C, outer view of Chelae." httpUri="https://zenodo.org/record/1052295/files/figure.png" pageId="5" pageNumber="500">Figs. 1A</figureCitation>
, 
<figureCitation id="76DA2A389376365EFF35FEA13682FEDF" box="[151,187,260,284]" captionStart="FIGURE 2" captionStartId="3.[151,250,1942,1964]" captionTargetBox="[236,1352,193,1921]" captionTargetId="figure@3.[236,1352,193,1921]" captionTargetPageId="3" captionText="FIGURE 2. Parathelphusa pardus sp. nov., paratype male (34.9 × 26.6 mm) (ZRC 2015.0452), Kampar River, Sumatra. A, overall dorsal view; B, dorsal view of CarapaCe; C, frontal view of CarapaCe." httpUri="https://zenodo.org/record/1052297/files/figure.png" pageId="5" pageNumber="500">2A</figureCitation>
, 
<figureCitation id="76DA2A389376365EFF65FEA136EEFEDF" box="[199,215,260,284]" captionStart="FIGURE 4" captionStartId="5.[151,250,1933,1955]" captionTargetBox="[163,1423,503,1907]" captionTargetId="figure@5.[163,1423,503,1907]" captionTargetPageId="5" captionText="FIGURE 4. Parathelphusa pardus sp. nov. A – C, paratype male (34.9 × 26.6 mm) (ZRC 2015.0452), Kampar River, Sumatra; D, paratype female (34.7 × 26.9 mm) (ZRC 2015.0452), Kampar River, Sumatra. A, right fourth ambulatory leg; B, right third ambulatory leg; C, D, right seCond ambulatory legs." httpUri="https://zenodo.org/record/1052301/files/figure.png" pageId="5" pageNumber="500">4</figureCitation>
, 
<figureCitation id="76DA2A389376365EFF41FEA13730FEDE" box="[227,265,260,285]" captionStart="FIGURE 6" captionStartId="7.[151,250,1796,1818]" captionTargetBox="[151,1436,213,1748]" captionTargetId="figure@7.[151,1436,213,1748]" captionTargetPageId="7" captionText="FIGURE 6. Parathelphusa pardus sp. nov. A, paratype female (34.7 × 26.9 mm) (ZRC 2015.0452), Kampar River, Sumatra; B, paratype female (46.1 × 34.7 mm) (ZRC 2015.0451), Kampar River, Sumatra. A, overall dorsal view; B, thoraCiC sternum showing vulvae." httpUri="https://zenodo.org/record/1052305/files/figure.png" pageId="5" pageNumber="500">6A</figureCitation>
).
</paragraph>
<paragraph id="EE5E36BD9376365EFF65FE823339FE13" blockId="5.[151,1436,151,464]" pageId="5" pageNumber="500">
Thoracic sternum evenly pitted to smooth; sternites 1, 2 completely fused forming triangular structure, relatively narrow longitudinally; suture separating sternites 2, 3 sinuous, medially concave with lateral parts convex (towards buccal cavity); sternites 3, 4 completely fused; sternoabdominal cavity extending to just before suture between sternites 2, 3, reaching imaginary line jointing anterior edges of coxae of chelipeds (
<figureCitation id="76DA2A389376365EFAB3FE3133A9FE6F" box="[1297,1424,404,429]" captionStart="FIGURE 3" captionStartId="4.[151,250,1626,1648]" captionTargetBox="[151,1436,193,1605]" captionTargetId="figure@4.[151,1436,193,1605]" captionTargetPageId="4" captionText="FIGURE 3. Parathelphusa pardus sp. nov., paratype male (34.9 × 26.6 mm) (ZRC 2015.0452), Kampar River, Sumatra. A, frontal view showing frontal median triangle of front; B, left third maxilliped; C, thoraCiC sternites 1 – 4 and telson; D, anterior thoraCiC sternum and male abdominal somites 5, 6 and telson; E, posterior thoraCiC sternums and male abdominal somites 1 – 5; F, right minor Chela; G, left major Chela." httpUri="https://zenodo.org/record/1052299/files/figure.png" pageId="5" pageNumber="500">Fig. 3C, D</figureCitation>
). Tubercle of press-button male abdominal locking mechanism rounded, on anterior edge of sternite 5.
</paragraph>
<caption id="BA9E66359376365EFF35F828349CF823" httpUri="https://zenodo.org/record/1052301/files/figure.png" pageId="5" pageNumber="500" targetBox="[163,1423,503,1907]" targetPageId="5">
<paragraph id="EE5E36BD9376365EFF35F828349CF823" blockId="5.[151,1436,1933,2016]" pageId="5" pageNumber="500">
<emphasis id="DC95EAAF9376365EFF35F8283734F861" bold="true" box="[151,269,1933,1955]" pageId="5" pageNumber="500">FIGURE 4</emphasis>
. 
<taxonomicName id="29E14D3E9376365EFEBBF82837C3F860" authority="Ng &amp; Riady, 2016" authorityName="Ng &amp; Riady" authorityYear="2016" box="[281,506,1933,1955]" class="Malacostraca" family="Gecarcinucidae" genus="Parathelphusa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="500" phylum="Arthropoda" rank="species" species="pardus" status="sp. nov.">
<emphasis id="DC95EAAF9376365EFEBBF82837C3F860" box="[281,506,1933,1955]" italics="true" pageId="5" pageNumber="500">Parathelphusa pardus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="C7A657D49376365EFDA3F8283468F861" box="[513,593,1933,1954]" pageId="5" pageNumber="500" rank="species">
<emphasis id="DC95EAAF9376365EFDA3F8283468F861" bold="true" box="[513,593,1933,1954]" pageId="5" pageNumber="500">sp. nov.</emphasis>
</taxonomicNameLabel>
A–C, paratype male (34.9 × 26.6 mm) (ZRC 2015.0452), Kampar River, Sumatra; D, paratype female (34.7 × 26.9 mm) (ZRC 2015.0452), Kampar River, Sumatra. A, right fourth ambulatory leg; B, right third ambulatory leg; C, D, right seCond ambulatory legs.
</paragraph>
</caption>
<caption id="BA9E66359375365DFF35F8D633A4F805" httpUri="https://zenodo.org/record/1052303/files/figure.png" pageId="6" pageNumber="501" targetBox="[224,1324,206,1867]" targetPageId="6">
<paragraph id="EE5E36BD9375365DFF35F8D633A4F805" blockId="6.[151,1437,1907,1990]" pageId="6" pageNumber="501">
<emphasis id="DC95EAAF9375365DFF35F8D63734F84B" bold="true" box="[151,269,1907,1929]" pageId="6" pageNumber="501">FIGURE 5</emphasis>
. 
<taxonomicName id="29E14D3E9375365DFEB8F8D637C3F84A" authority="Ng &amp; Riady, 2016" authorityName="Ng &amp; Riady" authorityYear="2016" box="[282,506,1907,1929]" class="Malacostraca" family="Gecarcinucidae" genus="Parathelphusa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="501" phylum="Arthropoda" rank="species" species="pardus" status="sp. nov.">
<emphasis id="DC95EAAF9375365DFEB8F8D637C3F84A" box="[282,506,1907,1929]" italics="true" pageId="6" pageNumber="501">Parathelphusa pardus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="C7A657D49375365DFDA0F8D6346BF84B" box="[514,594,1907,1928]" pageId="6" pageNumber="501" rank="species">
<emphasis id="DC95EAAF9375365DFDA0F8D6346BF84B" bold="true" box="[514,594,1907,1928]" pageId="6" pageNumber="501">sp. nov.</emphasis>
</taxonomicNameLabel>
A–D, holotype male (37.6 × 28.2 mm) (MZB Cru 4388), Kampar River, Sumatra; E–H, paratype male (34.9 × 26.6 mm) (ZRC 2015.0452), Kampar River, Sumatra. A, E, ventral view of left G1; B, F, ventral view of distal part of left G1; C, G, dorsal view of distal part of left G1; D, H, left G2s. SCales: A, D = 1.0 mm; B, C = 0.5 mm.
</paragraph>
</caption>
<caption id="BA9E66359374365CFF35F8A13707F894" httpUri="https://zenodo.org/record/1052305/files/figure.png" pageId="7" pageNumber="502" targetBox="[151,1436,213,1748]" targetPageId="7">
<paragraph id="EE5E36BD9374365CFF35F8A13707F894" blockId="7.[151,1436,1796,1879]" pageId="7" pageNumber="502">
<emphasis id="DC95EAAF9374365CFF35F8A13734F8DA" bold="true" box="[151,269,1796,1818]" pageId="7" pageNumber="502">FIGURE 6</emphasis>
. 
<taxonomicName id="29E14D3E9374365CFEB8F8A137C2F8D9" authority="Ng &amp; Riady, 2016" authorityName="Ng &amp; Riady" authorityYear="2016" box="[282,507,1796,1818]" class="Malacostraca" family="Gecarcinucidae" genus="Parathelphusa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="502" phylum="Arthropoda" rank="species" species="pardus" status="sp. nov.">
<emphasis id="DC95EAAF9374365CFEB8F8A137C2F8D9" box="[282,507,1796,1818]" italics="true" pageId="7" pageNumber="502">Parathelphusa pardus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="C7A657D49374365CFDA1F8A1346DF8DA" box="[515,596,1796,1817]" pageId="7" pageNumber="502" rank="species">
<emphasis id="DC95EAAF9374365CFDA1F8A1346DF8DA" bold="true" box="[515,596,1796,1817]" pageId="7" pageNumber="502">sp. nov.</emphasis>
</taxonomicNameLabel>
A, paratype female (34.7 × 26.9 mm) (ZRC 2015.0452), Kampar River, Sumatra; B, paratype female (46.1 × 34.7 mm) (ZRC 2015.0451), Kampar River, Sumatra. A, overall dorsal view; B, thoraCiC sternum showing vulvae.
</paragraph>
</caption>
<paragraph id="EE5E36BD93743653FF65F82134AEFF17" blockId="7.[151,1436,1924,2021]" lastBlockId="8.[151,1437,151,1976]" lastPageId="8" lastPageNumber="503" pageId="7" pageNumber="502">
Abdomen distinctly T-shaped; telson triangular, slightly longer than broad, lateral margins gently sinuous, tip rounded; somite 6 rectangular, 1.5–2 times as long as broad, lateral margins gently concave; somites 3–5 trapezoidal; lateral margins of somites 4, 5 concave; somite 3 broad, proximal edge reaching to coxae of last ambulatory legs, lateral margins convex; somites 1, 2 longitudinally narrow, wide, reaching to bases of coxae of last pair of ambulatory legs (
<figureCitation id="76DA2A38937B3653FE7AFF193403FF17" box="[472,570,188,213]" captionStart="FIGURE 1" captionStartId="2.[151,250,1966,1988]" captionTargetBox="[235,1330,193,1888]" captionTargetId="figure@2.[235,1330,193,1888]" captionTargetPageId="2" captionText="FIGURE 1. Parathelphusa pardus sp. nov., holotype male (37.6 × 28.2 mm) (MZB Cru 4388), Kampar River, Sumatra. A, overall dorsal view; B, anterior thoraCiC sternum and male abdominal somites 5, 6 and telson; C, outer view of Chelae." httpUri="https://zenodo.org/record/1052295/files/figure.png" pageId="8" pageNumber="503">Figs. 1B</figureCitation>
, 
<figureCitation id="76DA2A38937B3653FDE4FF1934B3FF17" box="[582,650,188,212]" captionStart="FIGURE 3" captionStartId="4.[151,250,1626,1648]" captionTargetBox="[151,1436,193,1605]" captionTargetId="figure@4.[151,1436,193,1605]" captionTargetPageId="4" captionText="FIGURE 3. Parathelphusa pardus sp. nov., paratype male (34.9 × 26.6 mm) (ZRC 2015.0452), Kampar River, Sumatra. A, frontal view showing frontal median triangle of front; B, left third maxilliped; C, thoraCiC sternites 1 – 4 and telson; D, anterior thoraCiC sternum and male abdominal somites 5, 6 and telson; E, posterior thoraCiC sternums and male abdominal somites 1 – 5; F, right minor Chela; G, left major Chela." httpUri="https://zenodo.org/record/1052299/files/figure.png" pageId="8" pageNumber="503">3C–E</figureCitation>
).
</paragraph>
<paragraph id="EE5E36BD937B3653FF65FF4534B4FE6F" blockId="8.[151,1437,151,1976]" pageId="8" pageNumber="503">
G1 relatively slender, terminal, subterminal segments not differentiated; entire structure almost straight, distal part slightly curved laterally; outer margin almost straight, entire along distal two-thirds, with distinct cleft on proximal part; lateral margins of proximal part with dense, long plumose setae; outer margin of distal quarter with long and short simple setae; subdistal surface with very short simple setae, scale-like setae; distal part tapering to sharp tip, opening distinct, laterally positioned (
<figureCitation id="76DA2A38937B3653FD18FECA3500FE4B" box="[698,825,367,392]" captionStart="FIGURE 5" captionStartId="6.[151,250,1907,1929]" captionTargetBox="[224,1324,206,1867]" captionTargetId="figure@6.[206,1359,193,1881]" captionTargetPageId="6" captionText="FIGURE 5. Parathelphusa pardus sp. nov. A – D, holotype male (37.6 × 28.2 mm) (MZB Cru 4388), Kampar River, Sumatra; E – H, paratype male (34.9 × 26.6 mm) (ZRC 2015.0452), Kampar River, Sumatra. A, E, ventral view of left G 1; B, F, ventral view of distal part of left G 1; C, G, dorsal view of distal part of left G 1; D, H, left G 2 s. SCales: A, D = 1.0 mm; B, C = 0.5 mm." httpUri="https://zenodo.org/record/1052303/files/figure.png" pageId="8" pageNumber="503">Fig. 5A–G</figureCitation>
). G2 much longer than G1; distal segment elongated, almost as long as basal segment (
<figureCitation id="76DA2A38937B3653FDA9FE3134BCFE6F" box="[523,645,404,429]" captionStart="FIGURE 5" captionStartId="6.[151,250,1907,1929]" captionTargetBox="[224,1324,206,1867]" captionTargetId="figure@6.[206,1359,193,1881]" captionTargetPageId="6" captionText="FIGURE 5. Parathelphusa pardus sp. nov. A – D, holotype male (37.6 × 28.2 mm) (MZB Cru 4388), Kampar River, Sumatra; E – H, paratype male (34.9 × 26.6 mm) (ZRC 2015.0452), Kampar River, Sumatra. A, E, ventral view of left G 1; B, F, ventral view of distal part of left G 1; C, G, dorsal view of distal part of left G 1; D, H, left G 2 s. SCales: A, D = 1.0 mm; B, C = 0.5 mm." httpUri="https://zenodo.org/record/1052303/files/figure.png" pageId="8" pageNumber="503">Fig. 5D, H</figureCitation>
)
</paragraph>
<paragraph id="EE5E36BD937B3653FF65FE12357EFD33" blockId="8.[151,1437,151,1976]" pageId="8" pageNumber="503">
<emphasis id="DC95EAAF937B3653FF65FE123799FE13" bold="true" box="[199,416,439,464]" pageId="8" pageNumber="503">Variation in males</emphasis>
. The male chelipeds are asymmetrical in large males in excess of about 
<quantity id="29199B58937B3653FB6AFE12332EFE0C" box="[1224,1303,439,464]" metricMagnitude="-2" metricUnit="m" metricValue="3.5" pageId="8" pageNumber="503" unit="mm" value="35.0">35 mm</quantity>
in carapace width. In smaller adult males, the major chela is relatively less inflated with the fingers less elongated and curved (
<figureCitation id="76DA2A38937B3653FF3DFE5A36CEFDDB" box="[159,247,511,536]" captionStart="FIGURE 3" captionStartId="4.[151,250,1626,1648]" captionTargetBox="[151,1436,193,1605]" captionTargetId="figure@4.[151,1436,193,1605]" captionTargetPageId="4" captionText="FIGURE 3. Parathelphusa pardus sp. nov., paratype male (34.9 × 26.6 mm) (ZRC 2015.0452), Kampar River, Sumatra. A, frontal view showing frontal median triangle of front; B, left third maxilliped; C, thoraCiC sternites 1 – 4 and telson; D, anterior thoraCiC sternum and male abdominal somites 5, 6 and telson; E, posterior thoraCiC sternums and male abdominal somites 1 – 5; F, right minor Chela; G, left major Chela." httpUri="https://zenodo.org/record/1052299/files/figure.png" pageId="8" pageNumber="503">Fig. 3G</figureCitation>
) when compared to those of larger males (e.g., 
<figureCitation id="76DA2A38937B3653FCB3FE5A3551FDDB" box="[785,872,511,536]" captionStart="FIGURE 1" captionStartId="2.[151,250,1966,1988]" captionTargetBox="[235,1330,193,1888]" captionTargetId="figure@2.[235,1330,193,1888]" captionTargetPageId="2" captionText="FIGURE 1. Parathelphusa pardus sp. nov., holotype male (37.6 × 28.2 mm) (MZB Cru 4388), Kampar River, Sumatra. A, overall dorsal view; B, anterior thoraCiC sternum and male abdominal somites 5, 6 and telson; C, outer view of Chelae." httpUri="https://zenodo.org/record/1052295/files/figure.png" pageId="8" pageNumber="503">Fig. 1C</figureCitation>
). The suture between thoracic sternites 2, 3 varies slightly in males less than 
<quantity id="29199B58937B3653FE6DFD813418FDFF" box="[463,545,548,572]" metricMagnitude="-2" metricUnit="m" metricValue="3.0" pageId="8" pageNumber="503" unit="mm" value="30.0">30 mm</quantity>
in carapace width, from sinuous to almost straight or gently evenly concave towards the buccal cavity. In adult males larger than 
<quantity id="29199B58937B3653FD55FDE2357EFD9C" box="[759,839,583,608]" metricMagnitude="-2" metricUnit="m" metricValue="3.5" pageId="8" pageNumber="503" unit="mm" value="35.0">35 mm</quantity>
carapace width, it is constant (see description). The male abdominal somite 6 varies in shape slightly. In smaller males (e.g., 34.9 × 
<quantity id="29199B58937B3653FBB6FDC93243FD47" box="[1044,1146,620,645]" metricMagnitude="-2" metricUnit="m" metricValue="2.66" pageId="8" pageNumber="503" unit="mm" value="26.6">26.6 mm</quantity>
, ZRC 2015.0452), somite 6 is more quadrate with the length only about 1.5 times the length (
<figureCitation id="76DA2A38937B3653FC2EFD2A35DEFD6B" box="[908,999,655,680]" captionStart="FIGURE 3" captionStartId="4.[151,250,1626,1648]" captionTargetBox="[151,1436,193,1605]" captionTargetId="figure@4.[151,1436,193,1605]" captionTargetPageId="4" captionText="FIGURE 3. Parathelphusa pardus sp. nov., paratype male (34.9 × 26.6 mm) (ZRC 2015.0452), Kampar River, Sumatra. A, frontal view showing frontal median triangle of front; B, left third maxilliped; C, thoraCiC sternites 1 – 4 and telson; D, anterior thoraCiC sternum and male abdominal somites 5, 6 and telson; E, posterior thoraCiC sternums and male abdominal somites 1 – 5; F, right minor Chela; G, left major Chela." httpUri="https://zenodo.org/record/1052299/files/figure.png" pageId="8" pageNumber="503">Fig. 3D</figureCitation>
). In larger males (e.g., 
<typeStatus id="315A881F937B3653FB48FD2A3375FD6B" box="[1258,1356,655,680]" pageId="8" pageNumber="503" type="holotype">holotype</typeStatus>
37.6 × 
<quantity id="29199B58937B3653FF35FD1136C4FD0F" box="[151,253,692,716]" metricMagnitude="-2" metricUnit="m" metricValue="2.82" pageId="8" pageNumber="503" unit="mm" value="28.2">28.2 mm</quantity>
, MZB Cru 4388), this ratio is about two times (
<figureCitation id="76DA2A38937B3653FCB5FD113557FD0F" box="[791,878,692,717]" captionStart="FIGURE 1" captionStartId="2.[151,250,1966,1988]" captionTargetBox="[235,1330,193,1888]" captionTargetId="figure@2.[235,1330,193,1888]" captionTargetPageId="2" captionText="FIGURE 1. Parathelphusa pardus sp. nov., holotype male (37.6 × 28.2 mm) (MZB Cru 4388), Kampar River, Sumatra. A, overall dorsal view; B, anterior thoraCiC sternum and male abdominal somites 5, 6 and telson; C, outer view of Chelae." httpUri="https://zenodo.org/record/1052295/files/figure.png" pageId="8" pageNumber="503">Fig. 1B</figureCitation>
). This is typical for many gecarcinucids in which somite 6, the telson are proportionately longer in large males.
</paragraph>
<paragraph id="EE5E36BD937B3653FF65FD5937BAFC2E" blockId="8.[151,1437,151,1976]" pageId="8" pageNumber="503">
<emphasis id="DC95EAAF937B3653FF65FD593727FCD6" bold="true" box="[199,286,764,789]" pageId="8" pageNumber="503">Female</emphasis>
. Females are similar to males in most non-sexual features. The chelipeds are symmetrical, not inflated and relatively slender, even in large females (
<figureCitation id="76DA2A38937B3653FD2FFCBA34DEFCFB" box="[653,743,799,824]" captionStart="FIGURE 6" captionStartId="7.[151,250,1796,1818]" captionTargetBox="[151,1436,213,1748]" captionTargetId="figure@7.[151,1436,213,1748]" captionTargetPageId="7" captionText="FIGURE 6. Parathelphusa pardus sp. nov. A, paratype female (34.7 × 26.9 mm) (ZRC 2015.0452), Kampar River, Sumatra; B, paratype female (46.1 × 34.7 mm) (ZRC 2015.0451), Kampar River, Sumatra. A, overall dorsal view; B, thoraCiC sternum showing vulvae." httpUri="https://zenodo.org/record/1052305/files/figure.png" pageId="8" pageNumber="503">Fig. 6A</figureCitation>
). The abdomen is almost round and covers most of the surface of the thoracic sternum; with all the somites and telson free; telson broadly triangular with gently convex lateral margins. Female specimens less than 
<quantity id="29199B58937B3653FDEBFCC234A2FCBC" box="[585,667,871,896]" metricMagnitude="-2" metricUnit="m" metricValue="3.5" pageId="8" pageNumber="503" unit="mm" value="35.0">35 mm</quantity>
in carapace width (e.g., female 30.2 × 
<quantity id="29199B58937B3653FBC7FCCD32F7FCBC" box="[1125,1230,872,896]" metricMagnitude="-2" metricUnit="m" metricValue="2.38" pageId="8" pageNumber="503" unit="mm" value="23.8">23.8 mm</quantity>
, ZRC 2015.0451) have more subovate abdomens and are not mature. The vulvae are relatively large, on surface of thoracic sternite 6 closer to suture between sternites 5 and 6; posterior part of vulva convex, anterior part with a soft, semilunate operculum (
<figureCitation id="76DA2A38937B3653FEBCFC71374FFC2E" box="[286,374,980,1005]" captionStart="FIGURE 6" captionStartId="7.[151,250,1796,1818]" captionTargetBox="[151,1436,213,1748]" captionTargetId="figure@7.[151,1436,213,1748]" captionTargetPageId="7" captionText="FIGURE 6. Parathelphusa pardus sp. nov. A, paratype female (34.7 × 26.9 mm) (ZRC 2015.0452), Kampar River, Sumatra; B, paratype female (46.1 × 34.7 mm) (ZRC 2015.0451), Kampar River, Sumatra. A, overall dorsal view; B, thoraCiC sternum showing vulvae." httpUri="https://zenodo.org/record/1052305/files/figure.png" pageId="8" pageNumber="503">Fig. 6B</figureCitation>
).
</paragraph>
<paragraph id="EE5E36BD937B3653FF65FC5237D4FBF6" blockId="8.[151,1437,151,1976]" pageId="8" pageNumber="503">
<emphasis id="DC95EAAF937B3653FF65FC52375CFBD3" bold="true" box="[199,357,1015,1040]" pageId="8" pageNumber="503">Colour in life</emphasis>
. The carapace and pereopods have a spotted and reticulated pattern in live and freshly preserved specimens (
<figureCitation id="76DA2A38937B3653FEB9FBB93747FBF7" box="[283,382,1052,1077]" captionStart="FIGURE 1" captionStartId="2.[151,250,1966,1988]" captionTargetBox="[235,1330,193,1888]" captionTargetId="figure@2.[235,1330,193,1888]" captionTargetPageId="2" captionText="FIGURE 1. Parathelphusa pardus sp. nov., holotype male (37.6 × 28.2 mm) (MZB Cru 4388), Kampar River, Sumatra. A, overall dorsal view; B, anterior thoraCiC sternum and male abdominal somites 5, 6 and telson; C, outer view of Chelae." httpUri="https://zenodo.org/record/1052295/files/figure.png" pageId="8" pageNumber="503">Figs. 1A</figureCitation>
, 
<figureCitation id="76DA2A38937B3653FE28FBB93796FBF7" box="[394,431,1052,1076]" captionStart="FIGURE 2" captionStartId="3.[151,250,1942,1964]" captionTargetBox="[236,1352,193,1921]" captionTargetId="figure@3.[236,1352,193,1921]" captionTargetPageId="3" captionText="FIGURE 2. Parathelphusa pardus sp. nov., paratype male (34.9 × 26.6 mm) (ZRC 2015.0452), Kampar River, Sumatra. A, overall dorsal view; B, dorsal view of CarapaCe; C, frontal view of CarapaCe." httpUri="https://zenodo.org/record/1052297/files/figure.png" pageId="8" pageNumber="503">2A</figureCitation>
, 
<figureCitation id="76DA2A38937B3653FE18FBB937D9FBF6" box="[442,480,1052,1077]" captionStart="FIGURE 6" captionStartId="7.[151,250,1796,1818]" captionTargetBox="[151,1436,213,1748]" captionTargetId="figure@7.[151,1436,213,1748]" captionTargetPageId="7" captionText="FIGURE 6. Parathelphusa pardus sp. nov. A, paratype female (34.7 × 26.9 mm) (ZRC 2015.0452), Kampar River, Sumatra; B, paratype female (46.1 × 34.7 mm) (ZRC 2015.0451), Kampar River, Sumatra. A, overall dorsal view; B, thoraCiC sternum showing vulvae." httpUri="https://zenodo.org/record/1052305/files/figure.png" pageId="8" pageNumber="503">6A</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="A6FB6536937B3653FF65FB9A34E2FBBE" pageId="8" pageNumber="503" type="etymology">
<paragraph id="EE5E36BD937B3653FF65FB9A34E2FBBE" blockId="8.[151,1437,151,1976]" pageId="8" pageNumber="503">
<emphasis id="DC95EAAF937B3653FF65FB9A377FFB9B" bold="true" box="[199,326,1087,1112]" pageId="8" pageNumber="503">Etymology</emphasis>
. The name is derived from the Latin 
<taxonomicName id="29E14D3E937B3653FD56FBE53578FB9B" authorityName="Ng &amp; Riady" authorityYear="2016" box="[756,833,1088,1112]" class="Malacostraca" family="Gecarcinucidae" genus="Parathelphusa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="503" phylum="Arthropoda" rank="species" species="pardus">
<emphasis id="DC95EAAF937B3653FD56FBE53578FB9B" box="[756,833,1088,1112]" italics="true" pageId="8" pageNumber="503">pardus</emphasis>
</taxonomicName>
for “leopard”, alluding to the pattern of spots on the carapace. The name is used as a noun in apposition.
</paragraph>
</subSubSection>
<subSubSection id="A6FB6536937B3651FF65FB22370EFCFB" lastPageId="10" lastPageNumber="505" pageId="8" pageNumber="503" type="discussion">
<paragraph id="EE5E36BD937B3653FF65FB22323CFABB" blockId="8.[151,1437,151,1976]" pageId="8" pageNumber="503">
<emphasis id="DC95EAAF937B3653FF65FB22370DFB63" bold="true" box="[199,308,1159,1184]" pageId="8" pageNumber="503">Remarks</emphasis>
. The colour pattern of 
<taxonomicName id="29E14D3E937B3653FD96FB2D3515FB63" authority="Ng &amp; Riady, 2016" authorityName="Ng &amp; Riady" authorityYear="2016" box="[564,812,1160,1184]" class="Malacostraca" family="Gecarcinucidae" genus="Parathelphusa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="503" phylum="Arthropoda" rank="species" species="pardus" status="sp. nov.">
<emphasis id="DC95EAAF937B3653FD96FB2D3515FB63" box="[564,812,1160,1184]" italics="true" pageId="8" pageNumber="503">Parathelphusa pardus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="C7A657D4937B3653FC90FB2235B2FB63" box="[818,907,1159,1184]" pageId="8" pageNumber="503" rank="species">
<emphasis id="DC95EAAF937B3653FC90FB2235B2FB63" bold="true" box="[818,907,1159,1184]" pageId="8" pageNumber="503">sp. nov.</emphasis>
</taxonomicNameLabel>
is distinctive, and the large series of specimens examined shows it is relatively constant character, even in small specimens. In 
<collectingRegion id="2C25F85F937B3653FB89FB0932B2FB07" box="[1067,1163,1196,1220]" country="Indonesia" name="Sumatera Utara" pageId="8" pageNumber="503">Sumatra</collectingRegion>
, this is a colour pattern shared with 
<taxonomicName id="29E14D3E937B3653FE89FB7437F9FB2B" baseAuthorityName="Rathbun" baseAuthorityYear="1902" box="[299,448,1232,1256]" class="Malacostraca" family="Gecarcinucidae" genus="Parathelphusa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="503" phylum="Arthropoda" rank="species" species="maindroni">
<emphasis id="DC95EAAF937B3653FE89FB7437F9FB2B" box="[299,448,1232,1256]" italics="true" pageId="8" pageNumber="503">P. maindroni</emphasis>
</taxonomicName>
and 
<taxonomicName id="29E14D3E937B3653FDA3FB7434A5FB2B" authorityName="Ng" authorityYear="1992" box="[513,668,1232,1256]" class="Malacostraca" family="Gecarcinucidae" genus="Parathelphusa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="503" phylum="Arthropoda" rank="species" species="batamensis">
<emphasis id="DC95EAAF937B3653FDA3FB7434A5FB2B" box="[513,668,1232,1256]" italics="true" pageId="8" pageNumber="503">P. batamensis</emphasis>
</taxonomicName>
(cf. 
<bibRefCitation id="8A704B4C937B3653FD7DFB743570FB2B" author="Ng" box="[735,841,1232,1257]" pageId="8" pageNumber="503" refString="Ng, P. K. L. (1992) The freshwater Crabs and palaemonid prawns (CrustaCea: DeCapoda) of Batam island, Riau ArChipelago, Indonesia, inCluding desCriptions of two new speCies. Proceedings of the Biological Society of Washington, 105 (4), 788 - 794." type="journal article" year="1992">Ng 1992</bibRefCitation>
, 
<bibRefCitation id="8A704B4C937B3653FCFBFB7535ACFB2B" author="Ng" box="[857,917,1232,1256]" pageId="8" pageNumber="503" refString="Ng, P. K. L. (1993) Parathelphusa maindroni (Rathbun, 1902), a peat swamp Crab from Peninsular Malaysia and Sumatra (CrustaCea: DeCapoda: BraChyura: Parathelphusidae). Malayan Nature Journal, 46, 189 - 200." type="journal article" year="1993">1993</bibRefCitation>
). This colour pattern is also observed in 
<taxonomicName id="29E14D3E937B3653FA25FB74374DFACF" authority="Ng, 1990" authorityName="Ng" authorityYear="1990" class="Malacostraca" family="Gecarcinucidae" genus="Parathelphusa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="503" phylum="Arthropoda" rank="species" species="reticulata">
<emphasis id="DC95EAAF937B3653FA25FB74373AFACF" italics="true" pageId="8" pageNumber="503">P. reticulata</emphasis>
Ng, 1990
</taxonomicName>
(from 
<collectingCountry id="96F6762D937B3653FE67FB513401FACE" box="[453,568,1268,1293]" name="Singapore" pageId="8" pageNumber="503">Singapore</collectingCountry>
) and 
<taxonomicName id="29E14D3E937B3653FDDBFB5035B5FACF" authority="Nobili, 1901" authorityName="Nobili" authorityYear="1901" box="[633,908,1268,1293]" class="Malacostraca" family="Gecarcinucidae" genus="Parathelphusa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="503" phylum="Arthropoda" rank="species" species="oxygona">
<emphasis id="DC95EAAF937B3653FDDBFB5034CCFACF" box="[633,757,1269,1292]" italics="true" pageId="8" pageNumber="503">P. oxygona</emphasis>
Nobili, 1901
</taxonomicName>
(from western 
<collectingRegion id="2C25F85F937B3653FB9EFB5132A7FACE" box="[1084,1182,1268,1293]" country="Malaysia" name="Sarawak" pageId="8" pageNumber="503">
<collectingCountry id="96F6762D937B3653FB9EFB5132A7FACE" box="[1084,1182,1268,1293]" name="Malaysia" pageId="8" pageNumber="503">Sarawak</collectingCountry>
</collectingRegion>
) (cf. 
<bibRefCitation id="8A704B4C937B3653FB7FFB503369FACE" author="Ng" box="[1245,1360,1268,1293]" pageId="8" pageNumber="503" refString="Ng, P. K. L. (1990 b) Parathelphusa reticulata speC. nov., a new speCies of freshwater Crab from blaCkwater swamps in Singapore (CrustaCea: DeCapoda: BraChyura: GeCarCinuCoidea). Zoologische Mededelingen, 63 (18), 241 - 254." type="journal article" year="1990" yearSuffix="b">Ng 1990b</bibRefCitation>
, 
<bibRefCitation id="8A704B4C937B3653FAFFFB5133A1FACF" author="Ng" box="[1373,1432,1268,1292]" pageId="8" pageNumber="503" refString="Ng, P. K. L. (2004) CrustaCea: DeCapoda, BraChyura. In: Yule, C. &amp; Yong, H. S. (Eds.), Freshwater Invertebrates of the Malaysian Region. Malaysian ACademy of SCienCes, pp. 311 - 336." type="book chapter" year="2004">2004</bibRefCitation>
, 
<bibRefCitation id="8A704B4C937B3653FF35FABD36EDFAF3" author="Ng" box="[151,212,1304,1328]" pageId="8" pageNumber="503" refString="Ng, P. K. L. (2014) The identity of the Sarawak freshwater Crab Parathelphusa oxygona Nobili, 1901, with desCription of a new speCies, Parathelphusa nobilii, from Western Kalimantan, Indonesia, Borneo (CrustaCea: BraChyura: GeCarCinuCidae). Zootaxa, 3774 (1), 31 - 44. http: // dx. doi. org / 10.11646 / zootaxa. 3774.1.2" type="journal article" year="2014">2014</bibRefCitation>
). Like 
<taxonomicName id="29E14D3E937B3653FE81FABC37B5FAF3" authority="Ng &amp; Riady, 2016" authorityName="Ng &amp; Riady" authorityYear="2016" box="[291,396,1304,1328]" class="Malacostraca" family="Gecarcinucidae" genus="Parathelphusa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="503" phylum="Arthropoda" rank="species" species="pardus" status="sp. nov.">
<emphasis id="DC95EAAF937B3653FE81FABC37B5FAF3" box="[291,396,1304,1328]" italics="true" pageId="8" pageNumber="503">P. pardus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="C7A657D4937B3653FE36FAB237D6FAF3" box="[404,495,1303,1328]" pageId="8" pageNumber="503" rank="species">
<emphasis id="DC95EAAF937B3653FE36FAB237D6FAF3" bold="true" box="[404,495,1303,1328]" pageId="8" pageNumber="503">sp. nov.</emphasis>
</taxonomicNameLabel>
these species are also usually associated with freshwater and peat-swamp habitats. Non swamp dwelling species like 
<taxonomicName id="29E14D3E937B3653FDB4FA9834B4FA97" authorityName="De Man" authorityYear="1879" box="[534,653,1341,1364]" class="Malacostraca" family="Gecarcinucidae" genus="Parathelphusa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="503" phylum="Arthropoda" rank="species" species="convexa">
<emphasis id="DC95EAAF937B3653FDB4FA9834B4FA97" box="[534,653,1341,1364]" italics="true" pageId="8" pageNumber="503">P. convexa</emphasis>
</taxonomicName>
and 
<taxonomicName id="29E14D3E937B3653FD61FA98357FFA97" authorityName="De Man" authorityYear="1879" box="[707,838,1340,1364]" class="Malacostraca" family="Gecarcinucidae" genus="Parathelphusa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="503" phylum="Arthropoda" rank="species" species="maculata">
<emphasis id="DC95EAAF937B3653FD61FA98357FFA97" box="[707,838,1340,1364]" italics="true" pageId="8" pageNumber="503">P. maculata</emphasis>
</taxonomicName>
sometimes also have an unevenly patterned carapace but it is never as distinctly patterned or brightly coloured as the above species.
</paragraph>
<paragraph id="EE5E36BD937B3653FF65FA21326CF95B" blockId="8.[151,1437,151,1976]" pageId="8" pageNumber="503">
The swelling of the dorsal carapace surface of 
<taxonomicName id="29E14D3E937B3653FD6EFA20350DFA5F" authority="Ng &amp; Riady, 2016" authorityName="Ng &amp; Riady" authorityYear="2016" box="[716,820,1412,1436]" class="Malacostraca" family="Gecarcinucidae" genus="Parathelphusa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="503" phylum="Arthropoda" rank="species" species="pardus" status="sp. nov.">
<emphasis id="DC95EAAF937B3653FD6EFA20350DFA5F" box="[716,820,1412,1436]" italics="true" pageId="8" pageNumber="503">P. pardus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="C7A657D4937B3653FC98FA2135AAFA5E" box="[826,915,1412,1437]" pageId="8" pageNumber="503" rank="species">
<emphasis id="DC95EAAF937B3653FC98FA2135AAFA5E" bold="true" box="[826,915,1412,1437]" pageId="8" pageNumber="503">sp. nov.</emphasis>
</taxonomicNameLabel>
is most like 
<taxonomicName id="29E14D3E937B3653FBBDFA203297FA5F" baseAuthorityName="Rathbun" baseAuthorityYear="1902" box="[1055,1198,1412,1436]" class="Malacostraca" family="Gecarcinucidae" genus="Parathelphusa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="503" phylum="Arthropoda" rank="species" species="maindroni">
<emphasis id="DC95EAAF937B3653FBBDFA203297FA5F" box="[1055,1198,1412,1436]" italics="true" pageId="8" pageNumber="503">P. maindroni</emphasis>
</taxonomicName>
(cf. 
<bibRefCitation id="8A704B4C937B3653FB43FA20337CFA5F" author="Ng" box="[1249,1349,1412,1437]" pageId="8" pageNumber="503" refString="Ng, P. K. L. (1993) Parathelphusa maindroni (Rathbun, 1902), a peat swamp Crab from Peninsular Malaysia and Sumatra (CrustaCea: DeCapoda: BraChyura: Parathelphusidae). Malayan Nature Journal, 46, 189 - 200." type="journal article" year="1993">Ng 1993</bibRefCitation>
: pl. 1A, B, pl. 2A, pl. 3A, C, pl. 4; 
<bibRefCitation id="8A704B4C937B3653FE67FA0C3400FA03" author="Ng" box="[453,569,1447,1473]" pageId="8" pageNumber="503" refString="Ng, P. K. L. (1990 b) Parathelphusa reticulata speC. nov., a new speCies of freshwater Crab from blaCkwater swamps in Singapore (CrustaCea: DeCapoda: BraChyura: GeCarCinuCoidea). Zoologische Mededelingen, 63 (18), 241 - 254." type="journal article" year="1990" yearSuffix="b">Ng 1990b</bibRefCitation>
: fig. 6A), but the suture between sternites 2 and 3 are more sinuous, with the median part gently concave and the lateral parts convex towards the buccal cavity (
<figureCitation id="76DA2A38937B3653FBD1FA6932F4FA27" box="[1139,1229,1484,1509]" captionStart="FIGURE 3" captionStartId="4.[151,250,1626,1648]" captionTargetBox="[151,1436,193,1605]" captionTargetId="figure@4.[151,1436,193,1605]" captionTargetPageId="4" captionText="FIGURE 3. Parathelphusa pardus sp. nov., paratype male (34.9 × 26.6 mm) (ZRC 2015.0452), Kampar River, Sumatra. A, frontal view showing frontal median triangle of front; B, left third maxilliped; C, thoraCiC sternites 1 – 4 and telson; D, anterior thoraCiC sternum and male abdominal somites 5, 6 and telson; E, posterior thoraCiC sternums and male abdominal somites 1 – 5; F, right minor Chela; G, left major Chela." httpUri="https://zenodo.org/record/1052299/files/figure.png" pageId="8" pageNumber="503">Fig. 3C</figureCitation>
) (evenly concave toward the buccal cavity in 
<taxonomicName id="29E14D3E937B3653FE68FA543460F9CB" baseAuthorityName="Rathbun" baseAuthorityYear="1902" box="[458,601,1520,1544]" class="Malacostraca" family="Gecarcinucidae" genus="Parathelphusa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="503" phylum="Arthropoda" rank="species" species="maindroni">
<emphasis id="DC95EAAF937B3653FE68FA543460F9CB" box="[458,601,1520,1544]" italics="true" pageId="8" pageNumber="503">P. maindroni</emphasis>
</taxonomicName>
, cf. 
<bibRefCitation id="8A704B4C937B3653FD2BFA5434C5F9CB" author="Ng" box="[649,764,1519,1545]" pageId="8" pageNumber="503" refString="Ng, P. K. L. (1990 b) Parathelphusa reticulata speC. nov., a new speCies of freshwater Crab from blaCkwater swamps in Singapore (CrustaCea: DeCapoda: BraChyura: GeCarCinuCoidea). Zoologische Mededelingen, 63 (18), 241 - 254." type="journal article" year="1990" yearSuffix="b">Ng 1990b</bibRefCitation>
: fig. 7C; 
<bibRefCitation id="8A704B4C937B3653FCC3FA5435FFF9CB" author="Ng" box="[865,966,1520,1545]" pageId="8" pageNumber="503" refString="Ng, P. K. L. (1993) Parathelphusa maindroni (Rathbun, 1902), a peat swamp Crab from Peninsular Malaysia and Sumatra (CrustaCea: DeCapoda: BraChyura: Parathelphusidae). Malayan Nature Journal, 46, 189 - 200." type="journal article" year="1993">Ng 1993</bibRefCitation>
: fig. 1G); and most significantly, the outer margin of the proximal part of the G1 has a distinct cleft (
<figureCitation id="76DA2A38937B3653FC89F9B135D1F9EF" box="[811,1000,1556,1581]" captionStart="FIGURE 5" captionStartId="6.[151,250,1907,1929]" captionTargetBox="[224,1324,206,1867]" captionTargetId="figure@6.[206,1359,193,1881]" captionTargetPageId="6" captionText="FIGURE 5. Parathelphusa pardus sp. nov. A – D, holotype male (37.6 × 28.2 mm) (MZB Cru 4388), Kampar River, Sumatra; E – H, paratype male (34.9 × 26.6 mm) (ZRC 2015.0452), Kampar River, Sumatra. A, E, ventral view of left G 1; B, F, ventral view of distal part of left G 1; C, G, dorsal view of distal part of left G 1; D, H, left G 2 s. SCales: A, D = 1.0 mm; B, C = 0.5 mm." httpUri="https://zenodo.org/record/1052303/files/figure.png" pageId="8" pageNumber="503">Fig. 5A–D, E–G</figureCitation>
) (outer margin of the proximal part of the G1 is entire in 
<taxonomicName id="29E14D3E937B3653FECDF99C3439F993" baseAuthorityName="Rathbun" baseAuthorityYear="1902" box="[367,512,1592,1616]" class="Malacostraca" family="Gecarcinucidae" genus="Parathelphusa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="503" phylum="Arthropoda" rank="species" species="maindroni">
<emphasis id="DC95EAAF937B3653FECDF99C3439F993" box="[367,512,1592,1616]" italics="true" pageId="8" pageNumber="503">P. maindroni</emphasis>
</taxonomicName>
, cf. 
<bibRefCitation id="8A704B4C937B3653FD96F99C3493F993" author="Ng" box="[564,682,1591,1617]" pageId="8" pageNumber="503" refString="Ng, P. K. L. (1990 b) Parathelphusa reticulata speC. nov., a new speCies of freshwater Crab from blaCkwater swamps in Singapore (CrustaCea: DeCapoda: BraChyura: GeCarCinuCoidea). Zoologische Mededelingen, 63 (18), 241 - 254." type="journal article" year="1990" yearSuffix="b">Ng 1990b</bibRefCitation>
: fig. 8A–D; 
<bibRefCitation id="8A704B4C937B3653FC9BF99C3599F993" author="Ng" box="[825,928,1592,1617]" pageId="8" pageNumber="503" refString="Ng, P. K. L. (1993) Parathelphusa maindroni (Rathbun, 1902), a peat swamp Crab from Peninsular Malaysia and Sumatra (CrustaCea: DeCapoda: BraChyura: Parathelphusidae). Malayan Nature Journal, 46, 189 - 200." type="journal article" year="1993">Ng 1993</bibRefCitation>
: figs. 1B, C, E, F, 2). The 
<typeStatus id="315A881F937B3653FB70F99C333BF992" box="[1234,1282,1593,1617]" pageId="8" pageNumber="503">type</typeStatus>
locality of 
<taxonomicName id="29E14D3E937B3653FA25F99C36DDF9B7" authority="Ng &amp; Riady, 2016" authorityName="Ng &amp; Riady" authorityYear="2016" class="Malacostraca" family="Gecarcinucidae" genus="Parathelphusa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="503" phylum="Arthropoda" rank="species" species="pardus" status="sp. nov.">
<emphasis id="DC95EAAF937B3653FA25F99C36DDF9B7" italics="true" pageId="8" pageNumber="503">P. pardus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="C7A657D4937B3653FF4FF9F93770F9B6" box="[237,329,1628,1653]" pageId="8" pageNumber="503" rank="species">
<emphasis id="DC95EAAF937B3653FF4FF9F93770F9B6" bold="true" box="[237,329,1628,1653]" pageId="8" pageNumber="503">sp. nov.</emphasis>
</taxonomicNameLabel>
, Pekanbaru, is more than 
<quantity id="29199B58937B3653FDD8F9F934EBF9B6" box="[634,722,1628,1653]" metricMagnitude="5" metricUnit="m" metricValue="1.0" pageId="8" pageNumber="503" unit="km" value="100.0">100 km</quantity>
southwest of Bengkalis, the 
<typeStatus id="315A881F937B3653FB80F9F8326BF9B6" box="[1058,1106,1629,1653]" pageId="8" pageNumber="503">type</typeStatus>
locality of 
<taxonomicName id="29E14D3E937B3653FB78F9F83355F9B7" baseAuthorityName="Rathbun" baseAuthorityYear="1902" box="[1242,1388,1628,1652]" class="Malacostraca" family="Gecarcinucidae" genus="Parathelphusa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="503" phylum="Arthropoda" rank="species" species="maindroni">
<emphasis id="DC95EAAF937B3653FB78F9F83355F9B7" box="[1242,1388,1628,1652]" italics="true" pageId="8" pageNumber="503">P. maindroni</emphasis>
</taxonomicName>
(cf. 
<bibRefCitation id="8A704B4C937B3653FF35F9DA3700F95B" author="Rathbun" box="[151,313,1663,1688]" pageId="8" pageNumber="503" refString="Rathbun, M. J. (1902) DesCription des nouvelles espECes de Parathelphusa appartenant au Museum de Paris. Bulletin du Museum d'Histoire naturelle, 8 (3), 184 - 187." type="journal article" year="1902">Rathbun 1902</bibRefCitation>
), with the latter species also present in Peninsular 
<collectingCountry id="96F6762D937B3653FCCEF9DA35EAF95B" box="[876,979,1663,1688]" name="Malaysia" pageId="8" pageNumber="503">Malaysia</collectingCountry>
(
<bibRefCitation id="8A704B4C937B3653FC40F9253270F95B" author="Ng" box="[994,1097,1664,1688]" pageId="8" pageNumber="503" refString="Ng, P. K. L. (1993) Parathelphusa maindroni (Rathbun, 1902), a peat swamp Crab from Peninsular Malaysia and Sumatra (CrustaCea: DeCapoda: BraChyura: Parathelphusidae). Malayan Nature Journal, 46, 189 - 200." type="journal article" year="1993">Ng 1993</bibRefCitation>
).
</paragraph>
<paragraph id="EE5E36BD937B3653FF65F901371CF87B" blockId="8.[151,1437,151,1976]" pageId="8" pageNumber="503">
The dorsal surface of the carapace of 
<taxonomicName id="29E14D3E937B3653FDD2F90034E0F97F" authority="Ng &amp; Riady, 2016" authorityName="Ng &amp; Riady" authorityYear="2016" box="[624,729,1700,1724]" class="Malacostraca" family="Gecarcinucidae" genus="Parathelphusa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="503" phylum="Arthropoda" rank="species" species="pardus" status="sp. nov.">
<emphasis id="DC95EAAF937B3653FDD2F90034E0F97F" box="[624,729,1700,1724]" italics="true" pageId="8" pageNumber="503">P. pardus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="C7A657D4937B3653FD43F9013502F97E" box="[737,827,1700,1725]" pageId="8" pageNumber="503" rank="species">
<emphasis id="DC95EAAF937B3653FD43F9013502F97E" bold="true" box="[737,827,1700,1725]" pageId="8" pageNumber="503">sp. nov.</emphasis>
</taxonomicNameLabel>
is more convex and swollen (
<figureCitation id="76DA2A38937B3653FB2CF90132CAF97F" box="[1166,1267,1700,1725]" captionStart="FIGURE 1" captionStartId="2.[151,250,1966,1988]" captionTargetBox="[235,1330,193,1888]" captionTargetId="figure@2.[235,1330,193,1888]" captionTargetPageId="2" captionText="FIGURE 1. Parathelphusa pardus sp. nov., holotype male (37.6 × 28.2 mm) (MZB Cru 4388), Kampar River, Sumatra. A, overall dorsal view; B, anterior thoraCiC sternum and male abdominal somites 5, 6 and telson; C, outer view of Chelae." httpUri="https://zenodo.org/record/1052295/files/figure.png" pageId="8" pageNumber="503">Figs. 1A</figureCitation>
, 
<figureCitation id="76DA2A38937B3653FB5DF9013336F97F" box="[1279,1295,1700,1724]" captionStart="FIGURE 2" captionStartId="3.[151,250,1942,1964]" captionTargetBox="[236,1352,193,1921]" captionTargetId="figure@3.[236,1352,193,1921]" captionTargetPageId="3" captionText="FIGURE 2. Parathelphusa pardus sp. nov., paratype male (34.9 × 26.6 mm) (ZRC 2015.0452), Kampar River, Sumatra. A, overall dorsal view; B, dorsal view of CarapaCe; C, frontal view of CarapaCe." httpUri="https://zenodo.org/record/1052297/files/figure.png" pageId="8" pageNumber="503">2</figureCitation>
, 
<figureCitation id="76DA2A38937B3653FABEF9013379F97E" box="[1308,1344,1700,1725]" captionStart="FIGURE 6" captionStartId="7.[151,250,1796,1818]" captionTargetBox="[151,1436,213,1748]" captionTargetId="figure@7.[151,1436,213,1748]" captionTargetPageId="7" captionText="FIGURE 6. Parathelphusa pardus sp. nov. A, paratype female (34.7 × 26.9 mm) (ZRC 2015.0452), Kampar River, Sumatra; B, paratype female (46.1 × 34.7 mm) (ZRC 2015.0451), Kampar River, Sumatra. A, overall dorsal view; B, thoraCiC sternum showing vulvae." httpUri="https://zenodo.org/record/1052305/files/figure.png" pageId="8" pageNumber="503">6A</figureCitation>
) than 
<taxonomicName id="29E14D3E937B3653FA25F900372CF923" authorityName="Ng" authorityYear="1992" class="Malacostraca" family="Gecarcinucidae" genus="Parathelphusa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="503" phylum="Arthropoda" rank="species" species="batamensis">
<emphasis id="DC95EAAF937B3653FA25F900372CF923" italics="true" pageId="8" pageNumber="503">P. batamensis</emphasis>
</taxonomicName>
(cf. 
<bibRefCitation id="8A704B4C937B3653FEECF96C378FF923" author="Ng" box="[334,438,1736,1761]" pageId="8" pageNumber="503" refString="Ng, P. K. L. (1992) The freshwater Crabs and palaemonid prawns (CrustaCea: DeCapoda) of Batam island, Riau ArChipelago, Indonesia, inCluding desCriptions of two new speCies. Proceedings of the Biological Society of Washington, 105 (4), 788 - 794." type="journal article" year="1992">Ng 1992</bibRefCitation>
: fig. 4A); the suture between sternites 2 and 3 are more sinuous, with the median part gently concave and the lateral parts convex towards the buccal cavity (
<figureCitation id="76DA2A38937B3653FC13F949323FF8C7" box="[945,1030,1772,1797]" captionStart="FIGURE 3" captionStartId="4.[151,250,1626,1648]" captionTargetBox="[151,1436,193,1605]" captionTargetId="figure@4.[151,1436,193,1605]" captionTargetPageId="4" captionText="FIGURE 3. Parathelphusa pardus sp. nov., paratype male (34.9 × 26.6 mm) (ZRC 2015.0452), Kampar River, Sumatra. A, frontal view showing frontal median triangle of front; B, left third maxilliped; C, thoraCiC sternites 1 – 4 and telson; D, anterior thoraCiC sternum and male abdominal somites 5, 6 and telson; E, posterior thoraCiC sternums and male abdominal somites 1 – 5; F, right minor Chela; G, left major Chela." httpUri="https://zenodo.org/record/1052299/files/figure.png" pageId="8" pageNumber="503">Fig. 3C</figureCitation>
) (evenly concave toward the buccal cavity in 
<taxonomicName id="29E14D3E937B3653FEA5F8B4379DF8EB" authorityName="Ng" authorityYear="1992" box="[263,420,1808,1832]" class="Malacostraca" family="Gecarcinucidae" genus="Parathelphusa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="503" phylum="Arthropoda" rank="species" species="batamensis">
<emphasis id="DC95EAAF937B3653FEA5F8B4379DF8EB" box="[263,420,1808,1832]" italics="true" pageId="8" pageNumber="503">P. batamensis</emphasis>
</taxonomicName>
, cf. 
<bibRefCitation id="8A704B4C937B3653FE7CF8B4347EF8EB" author="Ng" box="[478,583,1808,1833]" pageId="8" pageNumber="503" refString="Ng, P. K. L. (1992) The freshwater Crabs and palaemonid prawns (CrustaCea: DeCapoda) of Batam island, Riau ArChipelago, Indonesia, inCluding desCriptions of two new speCies. Proceedings of the Biological Society of Washington, 105 (4), 788 - 794." type="journal article" year="1992">Ng 1992</bibRefCitation>
: fig. 2H); the lateral margins of the male abdominal somite 6 are gently concave (
<figureCitation id="76DA2A38937B3653FEA4F8913758F88F" box="[262,353,1844,1869]" captionStart="FIGURE 3" captionStartId="4.[151,250,1626,1648]" captionTargetBox="[151,1436,193,1605]" captionTargetId="figure@4.[151,1436,193,1605]" captionTargetPageId="4" captionText="FIGURE 3. Parathelphusa pardus sp. nov., paratype male (34.9 × 26.6 mm) (ZRC 2015.0452), Kampar River, Sumatra. A, frontal view showing frontal median triangle of front; B, left third maxilliped; C, thoraCiC sternites 1 – 4 and telson; D, anterior thoraCiC sternum and male abdominal somites 5, 6 and telson; E, posterior thoraCiC sternums and male abdominal somites 1 – 5; F, right minor Chela; G, left major Chela." httpUri="https://zenodo.org/record/1052299/files/figure.png" pageId="8" pageNumber="503">Fig. 3D</figureCitation>
) (gently convex in 
<taxonomicName id="29E14D3E937B3653FDE9F89034D1F88F" authorityName="Ng" authorityYear="1992" box="[587,744,1844,1868]" class="Malacostraca" family="Gecarcinucidae" genus="Parathelphusa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="503" phylum="Arthropoda" rank="species" species="batamensis">
<emphasis id="DC95EAAF937B3653FDE9F89034D1F88F" box="[587,744,1844,1868]" italics="true" pageId="8" pageNumber="503">P. batamensis</emphasis>
</taxonomicName>
, cf. 
<bibRefCitation id="8A704B4C937B3653FC83F89035B3F88F" author="Ng" box="[801,906,1844,1869]" pageId="8" pageNumber="503" refString="Ng, P. K. L. (1992) The freshwater Crabs and palaemonid prawns (CrustaCea: DeCapoda) of Batam island, Riau ArChipelago, Indonesia, inCluding desCriptions of two new speCies. Proceedings of the Biological Society of Washington, 105 (4), 788 - 794." type="journal article" year="1992">Ng 1992</bibRefCitation>
: fig. 2G); and the G1 has the distalmost part slightly curved and the outer margin of the proximal part has a distinct cleft (
<figureCitation id="76DA2A38937B3653FBADF8F232F7F8B3" box="[1039,1230,1879,1904]" captionStart="FIGURE 5" captionStartId="6.[151,250,1907,1929]" captionTargetBox="[224,1324,206,1867]" captionTargetId="figure@6.[206,1359,193,1881]" captionTargetPageId="6" captionText="FIGURE 5. Parathelphusa pardus sp. nov. A – D, holotype male (37.6 × 28.2 mm) (MZB Cru 4388), Kampar River, Sumatra; E – H, paratype male (34.9 × 26.6 mm) (ZRC 2015.0452), Kampar River, Sumatra. A, E, ventral view of left G 1; B, F, ventral view of distal part of left G 1; C, G, dorsal view of distal part of left G 1; D, H, left G 2 s. SCales: A, D = 1.0 mm; B, C = 0.5 mm." httpUri="https://zenodo.org/record/1052303/files/figure.png" pageId="8" pageNumber="503">Fig. 5A–D, E–G</figureCitation>
) (distal half more prominently curved laterally and the outer margin of the proximal part is entire in 
<taxonomicName id="29E14D3E937B3653FB95F8D832EBF857" authorityName="Ng" authorityYear="1992" box="[1079,1234,1916,1940]" class="Malacostraca" family="Gecarcinucidae" genus="Parathelphusa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="503" phylum="Arthropoda" rank="species" species="batamensis">
<emphasis id="DC95EAAF937B3653FB95F8D832EBF857" box="[1079,1234,1916,1940]" italics="true" pageId="8" pageNumber="503">P. batamensis</emphasis>
</taxonomicName>
, cf. 
<bibRefCitation id="8A704B4C937B3653FAA6F8D83353F857" author="Ng" box="[1284,1386,1916,1941]" pageId="8" pageNumber="503" refString="Ng, P. K. L. (1992) The freshwater Crabs and palaemonid prawns (CrustaCea: DeCapoda) of Batam island, Riau ArChipelago, Indonesia, inCluding desCriptions of two new speCies. Proceedings of the Biological Society of Washington, 105 (4), 788 - 794." type="journal article" year="1992">Ng 1992</bibRefCitation>
: fig. 4B, C, E, F).
</paragraph>
<caption id="BA9E6635937A3652FF35F8F83599F852" httpUri="https://zenodo.org/record/1052307/files/figure.png" pageId="9" pageNumber="504" targetBox="[152,1438,193,1857]" targetPageId="9">
<paragraph id="EE5E36BD937A3652FF35F8F83599F852" blockId="9.[151,1436,1884,1937]" pageId="9" pageNumber="504">
<emphasis id="DC95EAAF937A3652FF35F8F83736F8B1" bold="true" box="[151,271,1885,1907]" pageId="9" pageNumber="504">FIGURE 7</emphasis>
. The habitat of 
<taxonomicName id="29E14D3E937A3652FE1AF8F934A2F8B1" authority="Ng &amp; Riady, 2016" authorityName="Ng &amp; Riady" authorityYear="2016" box="[440,667,1884,1906]" class="Malacostraca" family="Gecarcinucidae" genus="Parathelphusa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="504" phylum="Arthropoda" rank="species" species="pardus" status="sp. nov.">
<emphasis id="DC95EAAF937A3652FE1AF8F934A2F8B1" box="[440,667,1884,1906]" italics="true" pageId="9" pageNumber="504">Parathelphusa pardus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="C7A657D4937A3652FD01F8F834CFF8B1" box="[675,758,1885,1906]" pageId="9" pageNumber="504" rank="species">
<emphasis id="DC95EAAF937A3652FD01F8F834CFF8B1" bold="true" box="[675,758,1885,1906]" pageId="9" pageNumber="504">sp. nov.</emphasis>
</taxonomicNameLabel>
A, one of the tributaries flowing through the swamp forest; B, a shallow seCtion of the swamp with 
<taxonomicName id="29E14D3E937A3652FE5EF8DE3461F852" authorityName="Wallich" authorityYear="1827" box="[508,600,1915,1937]" class="Magnoliopsida" family="Nymphaeaceae" genus="Barclaya" higherTaxonomySource="GBIF" kingdom="Plantae" order="Nymphaeales" pageId="9" pageNumber="504" phylum="Tracheophyta" rank="genus">
<emphasis id="DC95EAAF937A3652FE5EF8DE3461F852" box="[508,600,1915,1937]" italics="true" pageId="9" pageNumber="504">Barclaya</emphasis>
</taxonomicName>
sp. and a Crab foraging at night.
</paragraph>
</caption>
<paragraph id="EE5E36BD93793651FF65FF32341FFE6F" blockId="10.[151,1437,151,1040]" pageId="10" pageNumber="505">
The dorsal surface of the carapace is relatively less swollen in 
<taxonomicName id="29E14D3E93793651FC26FF3C35D5FF73" authority="Ng &amp; Riady, 2016" authorityName="Ng &amp; Riady" authorityYear="2016" box="[900,1004,152,176]" class="Malacostraca" family="Gecarcinucidae" genus="Parathelphusa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="505" phylum="Arthropoda" rank="species" species="pardus" status="sp. nov.">
<emphasis id="DC95EAAF93793651FC26FF3C35D5FF73" box="[900,1004,152,176]" italics="true" pageId="10" pageNumber="505">P. pardus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="C7A657D493793651FC51FF323274FF73" box="[1011,1101,151,176]" pageId="10" pageNumber="505" rank="species">
<emphasis id="DC95EAAF93793651FC51FF323274FF73" bold="true" box="[1011,1101,151,176]" pageId="10" pageNumber="505">sp. nov.</emphasis>
</taxonomicNameLabel>
(
<figureCitation id="76DA2A3893793651FBFEFF323286FF73" box="[1116,1215,151,176]" captionStart="FIGURE 1" captionStartId="2.[151,250,1966,1988]" captionTargetBox="[235,1330,193,1888]" captionTargetId="figure@2.[235,1330,193,1888]" captionTargetPageId="2" captionText="FIGURE 1. Parathelphusa pardus sp. nov., holotype male (37.6 × 28.2 mm) (MZB Cru 4388), Kampar River, Sumatra. A, overall dorsal view; B, anterior thoraCiC sternum and male abdominal somites 5, 6 and telson; C, outer view of Chelae." httpUri="https://zenodo.org/record/1052295/files/figure.png" pageId="10" pageNumber="505">Figs. 1A</figureCitation>
, 
<figureCitation id="76DA2A3893793651FB69FF3D3328FF6C" box="[1227,1297,152,176]" captionStart="FIGURE 2" captionStartId="3.[151,250,1942,1964]" captionTargetBox="[236,1352,193,1921]" captionTargetId="figure@3.[236,1352,193,1921]" captionTargetPageId="3" captionText="FIGURE 2. Parathelphusa pardus sp. nov., paratype male (34.9 × 26.6 mm) (ZRC 2015.0452), Kampar River, Sumatra. A, overall dorsal view; B, dorsal view of CarapaCe; C, frontal view of CarapaCe." httpUri="https://zenodo.org/record/1052297/files/figure.png" pageId="10" pageNumber="505">2A, B</figureCitation>
, 
<figureCitation id="76DA2A3893793651FABEFF323379FF73" box="[1308,1344,151,176]" captionStart="FIGURE 6" captionStartId="7.[151,250,1796,1818]" captionTargetBox="[151,1436,213,1748]" captionTargetId="figure@7.[151,1436,213,1748]" captionTargetPageId="7" captionText="FIGURE 6. Parathelphusa pardus sp. nov. A, paratype female (34.7 × 26.9 mm) (ZRC 2015.0452), Kampar River, Sumatra; B, paratype female (46.1 × 34.7 mm) (ZRC 2015.0451), Kampar River, Sumatra. A, overall dorsal view; B, thoraCiC sternum showing vulvae." httpUri="https://zenodo.org/record/1052305/files/figure.png" pageId="10" pageNumber="505">6A</figureCitation>
) than in 
<taxonomicName id="29E14D3E93793651FF35FF183726FF17" authorityName="Ng" authorityYear="1990" box="[151,287,188,212]" class="Malacostraca" family="Gecarcinucidae" genus="Parathelphusa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="505" phylum="Arthropoda" rank="species" species="reticulata">
<emphasis id="DC95EAAF93793651FF35FF183726FF17" box="[151,287,188,212]" italics="true" pageId="10" pageNumber="505">P. reticulata</emphasis>
</taxonomicName>
from 
<collectingCountry id="96F6762D93793651FEC6FF1937EFFF16" box="[356,470,188,213]" name="Singapore" pageId="10" pageNumber="505">Singapore</collectingCountry>
(cf. 
<bibRefCitation id="8A704B4C93793651FDA8FF183447FF16" author="Ng" box="[522,638,188,213]" pageId="10" pageNumber="505" refString="Ng, P. K. L. (1990 b) Parathelphusa reticulata speC. nov., a new speCies of freshwater Crab from blaCkwater swamps in Singapore (CrustaCea: DeCapoda: BraChyura: GeCarCinuCoidea). Zoologische Mededelingen, 63 (18), 241 - 254." type="journal article" year="1990" yearSuffix="b">Ng 1990b</bibRefCitation>
: figs. 1, 2); the suture between sternites 2 and 3 are more sinuous, with the median part gently concave and the lateral parts convex towards the buccal cavity (
<figureCitation id="76DA2A3893793651FBD9FF7A32EBFF3B" box="[1147,1234,223,248]" captionStart="FIGURE 3" captionStartId="4.[151,250,1626,1648]" captionTargetBox="[151,1436,193,1605]" captionTargetId="figure@4.[151,1436,193,1605]" captionTargetPageId="4" captionText="FIGURE 3. Parathelphusa pardus sp. nov., paratype male (34.9 × 26.6 mm) (ZRC 2015.0452), Kampar River, Sumatra. A, frontal view showing frontal median triangle of front; B, left third maxilliped; C, thoraCiC sternites 1 – 4 and telson; D, anterior thoraCiC sternum and male abdominal somites 5, 6 and telson; E, posterior thoraCiC sternums and male abdominal somites 1 – 5; F, right minor Chela; G, left major Chela." httpUri="https://zenodo.org/record/1052299/files/figure.png" pageId="10" pageNumber="505">Fig. 3C</figureCitation>
) (evenly concave toward the buccal cavity in 
<taxonomicName id="29E14D3E93793651FE7DFEA03453FEDF" authorityName="Ng" authorityYear="1990" box="[479,618,260,284]" class="Malacostraca" family="Gecarcinucidae" genus="Parathelphusa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="505" phylum="Arthropoda" rank="species" species="reticulata">
<emphasis id="DC95EAAF93793651FE7DFEA03453FEDF" box="[479,618,260,284]" italics="true" pageId="10" pageNumber="505">P. reticulata</emphasis>
</taxonomicName>
, cf. 
<bibRefCitation id="8A704B4C93793651FD00FEA03520FEDE" author="Ng" box="[674,793,260,285]" pageId="10" pageNumber="505" refString="Ng, P. K. L. (1990 b) Parathelphusa reticulata speC. nov., a new speCies of freshwater Crab from blaCkwater swamps in Singapore (CrustaCea: DeCapoda: BraChyura: GeCarCinuCoidea). Zoologische Mededelingen, 63 (18), 241 - 254." type="journal article" year="1990" yearSuffix="b">Ng 1990b</bibRefCitation>
: fig. 
<date id="9A5F107D93793651FCFAFEA1354AFEDF" box="[856,883,260,284]" pageId="10" pageNumber="505">7I</date>
); the ambulatory legs are relatively shorter and stouter (
<figureCitation id="76DA2A3893793651FF51FE82376FFE83" box="[243,342,295,320]" captionStart="FIGURE 1" captionStartId="2.[151,250,1966,1988]" captionTargetBox="[235,1330,193,1888]" captionTargetId="figure@2.[235,1330,193,1888]" captionTargetPageId="2" captionText="FIGURE 1. Parathelphusa pardus sp. nov., holotype male (37.6 × 28.2 mm) (MZB Cru 4388), Kampar River, Sumatra. A, overall dorsal view; B, anterior thoraCiC sternum and male abdominal somites 5, 6 and telson; C, outer view of Chelae." httpUri="https://zenodo.org/record/1052295/files/figure.png" pageId="10" pageNumber="505">Figs. 1A</figureCitation>
, 
<figureCitation id="76DA2A3893793651FEC6FE8D37B1FE83" box="[356,392,296,320]" captionStart="FIGURE 2" captionStartId="3.[151,250,1942,1964]" captionTargetBox="[236,1352,193,1921]" captionTargetId="figure@3.[236,1352,193,1921]" captionTargetPageId="3" captionText="FIGURE 2. Parathelphusa pardus sp. nov., paratype male (34.9 × 26.6 mm) (ZRC 2015.0452), Kampar River, Sumatra. A, overall dorsal view; B, dorsal view of CarapaCe; C, frontal view of CarapaCe." httpUri="https://zenodo.org/record/1052297/files/figure.png" pageId="10" pageNumber="505">2A</figureCitation>
, 
<figureCitation id="76DA2A3893793651FE37FE8D379CFE83" box="[405,421,296,320]" captionStart="FIGURE 4" captionStartId="5.[151,250,1933,1955]" captionTargetBox="[163,1423,503,1907]" captionTargetId="figure@5.[163,1423,503,1907]" captionTargetPageId="5" captionText="FIGURE 4. Parathelphusa pardus sp. nov. A – C, paratype male (34.9 × 26.6 mm) (ZRC 2015.0452), Kampar River, Sumatra; D, paratype female (34.7 × 26.9 mm) (ZRC 2015.0452), Kampar River, Sumatra. A, right fourth ambulatory leg; B, right third ambulatory leg; C, D, right seCond ambulatory legs." httpUri="https://zenodo.org/record/1052301/files/figure.png" pageId="10" pageNumber="505">4</figureCitation>
, 
<figureCitation id="76DA2A3893793651FE13FE8237ECFE83" box="[433,469,295,320]" captionStart="FIGURE 6" captionStartId="7.[151,250,1796,1818]" captionTargetBox="[151,1436,213,1748]" captionTargetId="figure@7.[151,1436,213,1748]" captionTargetPageId="7" captionText="FIGURE 6. Parathelphusa pardus sp. nov. A, paratype female (34.7 × 26.9 mm) (ZRC 2015.0452), Kampar River, Sumatra; B, paratype female (46.1 × 34.7 mm) (ZRC 2015.0451), Kampar River, Sumatra. A, overall dorsal view; B, thoraCiC sternum showing vulvae." httpUri="https://zenodo.org/record/1052305/files/figure.png" pageId="10" pageNumber="505">6A</figureCitation>
) than in 
<taxonomicName id="29E14D3E93793651FD9BFE8C34F8FE83" authorityName="Ng" authorityYear="1990" box="[569,705,296,320]" class="Malacostraca" family="Gecarcinucidae" genus="Parathelphusa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="505" phylum="Arthropoda" rank="species" species="reticulata">
<emphasis id="DC95EAAF93793651FD9BFE8C34F8FE83" box="[569,705,296,320]" italics="true" pageId="10" pageNumber="505">P. reticulata</emphasis>
</taxonomicName>
(cf. 
<bibRefCitation id="8A704B4C93793651FD54FE8D3553FE83" author="Ng" box="[758,874,295,320]" pageId="10" pageNumber="505" refString="Ng, P. K. L. (1990 b) Parathelphusa reticulata speC. nov., a new speCies of freshwater Crab from blaCkwater swamps in Singapore (CrustaCea: DeCapoda: BraChyura: GeCarCinuCoidea). Zoologische Mededelingen, 63 (18), 241 - 254." type="journal article" year="1990" yearSuffix="b">Ng 1990b</bibRefCitation>
: figs. 1, 2); the distalmost part of the G1 is gently curved and relatively stouter, with the outer margin of the proximal part possessing a distinct cleft (
<figureCitation id="76DA2A3893793651FB57FEE93694FE4B" captionStart="FIGURE 5" captionStartId="6.[151,250,1907,1929]" captionTargetBox="[224,1324,206,1867]" captionTargetId="figure@6.[206,1359,193,1881]" captionTargetPageId="6" captionText="FIGURE 5. Parathelphusa pardus sp. nov. A – D, holotype male (37.6 × 28.2 mm) (MZB Cru 4388), Kampar River, Sumatra; E – H, paratype male (34.9 × 26.6 mm) (ZRC 2015.0452), Kampar River, Sumatra. A, E, ventral view of left G 1; B, F, ventral view of distal part of left G 1; C, G, dorsal view of distal part of left G 1; D, H, left G 2 s. SCales: A, D = 1.0 mm; B, C = 0.5 mm." httpUri="https://zenodo.org/record/1052303/files/figure.png" pageId="10" pageNumber="505">Fig. 5A–D, E– G</figureCitation>
) (almost straight and slender, with the outer margin of the proximal part possessing a more shallow cleft in 
<taxonomicName id="29E14D3E93793651FA25FED4373AFE6F" authorityName="Ng" authorityYear="1990" class="Malacostraca" family="Gecarcinucidae" genus="Parathelphusa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="505" phylum="Arthropoda" rank="species" species="reticulata">
<emphasis id="DC95EAAF93793651FA25FED4373AFE6F" italics="true" pageId="10" pageNumber="505">P. reticulata</emphasis>
</taxonomicName>
, cf. 
<bibRefCitation id="8A704B4C93793651FE96FE303791FE6E" author="Ng" box="[308,424,404,429]" pageId="10" pageNumber="505" refString="Ng, P. K. L. (1990 b) Parathelphusa reticulata speC. nov., a new speCies of freshwater Crab from blaCkwater swamps in Singapore (CrustaCea: DeCapoda: BraChyura: GeCarCinuCoidea). Zoologische Mededelingen, 63 (18), 241 - 254." type="journal article" year="1990" yearSuffix="b">Ng 1990b</bibRefCitation>
: fig. 8F–I);
</paragraph>
<paragraph id="EE5E36BD93793651FF65FE123369FDA3" blockId="10.[151,1437,151,1040]" pageId="10" pageNumber="505">
The degree of swelling and spotted carapace pattern of 
<taxonomicName id="29E14D3E93793651FC9AFE1C3598FE13" authority="Ng &amp; Riady, 2016" authorityName="Ng &amp; Riady" authorityYear="2016" box="[824,929,440,464]" class="Malacostraca" family="Gecarcinucidae" genus="Parathelphusa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="505" phylum="Arthropoda" rank="species" species="pardus" status="sp. nov.">
<emphasis id="DC95EAAF93793651FC9AFE1C3598FE13" box="[824,929,440,464]" italics="true" pageId="10" pageNumber="505">P. pardus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="C7A657D493793651FC0BFE12323AFE13" box="[937,1027,439,464]" pageId="10" pageNumber="505" rank="species">
<emphasis id="DC95EAAF93793651FC0BFE12323AFE13" bold="true" box="[937,1027,439,464]" pageId="10" pageNumber="505">sp. nov.</emphasis>
</taxonomicNameLabel>
is also similar to another species, 
<taxonomicName id="29E14D3E93793651FA25FE1C36CFFE37" authorityName="Nobili" authorityYear="1901" class="Malacostraca" family="Gecarcinucidae" genus="Parathelphusa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="505" phylum="Arthropoda" rank="species" species="oxygona">
<emphasis id="DC95EAAF93793651FA25FE1C36CFFE37" italics="true" pageId="10" pageNumber="505">P. oxygona</emphasis>
</taxonomicName>
, from western 
<collectingRegion id="2C25F85F93793651FE06FE79343EFE36" box="[420,519,476,501]" country="Malaysia" name="Sarawak" pageId="10" pageNumber="505">
<collectingCountry id="96F6762D93793651FE06FE79343EFE36" box="[420,519,476,501]" name="Malaysia" pageId="10" pageNumber="505">Sarawak</collectingCountry>
</collectingRegion>
, Borneo. 
<taxonomicName id="29E14D3E93793651FDD4FE793548FE37" authority="Ng &amp; Riady, 2016" authorityName="Ng &amp; Riady" authorityYear="2016" box="[630,881,476,500]" class="Malacostraca" family="Gecarcinucidae" genus="Parathelphusa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="505" phylum="Arthropoda" rank="species" species="pardus" status="sp. nov.">
<emphasis id="DC95EAAF93793651FDD4FE793548FE37" box="[630,881,476,500]" italics="true" pageId="10" pageNumber="505">Parathelphusa pardus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="C7A657D493793651FCDBFE7935ECFE36" box="[889,981,476,501]" pageId="10" pageNumber="505" rank="species">
<emphasis id="DC95EAAF93793651FCDBFE7935ECFE36" bold="true" box="[889,981,476,501]" pageId="10" pageNumber="505">sp. nov.</emphasis>
</taxonomicNameLabel>
is easily distinguished from 
<taxonomicName id="29E14D3E93793651FA82FE7833A5FE37" authorityName="Nobili" authorityYear="1901" box="[1312,1436,477,500]" class="Malacostraca" family="Gecarcinucidae" genus="Parathelphusa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="505" phylum="Arthropoda" rank="species" species="oxygona">
<emphasis id="DC95EAAF93793651FA82FE7833A5FE37" box="[1312,1436,477,500]" italics="true" pageId="10" pageNumber="505">P. oxygona</emphasis>
</taxonomicName>
by having the outer margin of the external orbital almost straight (
<figureCitation id="76DA2A3893793651FC30FE5A35CEFDDB" box="[914,1015,511,536]" captionStart="FIGURE 1" captionStartId="2.[151,250,1966,1988]" captionTargetBox="[235,1330,193,1888]" captionTargetId="figure@2.[235,1330,193,1888]" captionTargetPageId="2" captionText="FIGURE 1. Parathelphusa pardus sp. nov., holotype male (37.6 × 28.2 mm) (MZB Cru 4388), Kampar River, Sumatra. A, overall dorsal view; B, anterior thoraCiC sternum and male abdominal somites 5, 6 and telson; C, outer view of Chelae." httpUri="https://zenodo.org/record/1052295/files/figure.png" pageId="10" pageNumber="505">Figs. 1A</figureCitation>
, 
<figureCitation id="76DA2A3893793651FBA4FDA53274FDD4" box="[1030,1101,512,536]" captionStart="FIGURE 2" captionStartId="3.[151,250,1942,1964]" captionTargetBox="[236,1352,193,1921]" captionTargetId="figure@3.[236,1352,193,1921]" captionTargetPageId="3" captionText="FIGURE 2. Parathelphusa pardus sp. nov., paratype male (34.9 × 26.6 mm) (ZRC 2015.0452), Kampar River, Sumatra. A, overall dorsal view; B, dorsal view of CarapaCe; C, frontal view of CarapaCe." httpUri="https://zenodo.org/record/1052297/files/figure.png" pageId="10" pageNumber="505">2A, B</figureCitation>
, 
<figureCitation id="76DA2A3893793651FBF9FE5A3247FDDB" box="[1115,1150,511,536]" captionStart="FIGURE 6" captionStartId="7.[151,250,1796,1818]" captionTargetBox="[151,1436,213,1748]" captionTargetId="figure@7.[151,1436,213,1748]" captionTargetPageId="7" captionText="FIGURE 6. Parathelphusa pardus sp. nov. A, paratype female (34.7 × 26.9 mm) (ZRC 2015.0452), Kampar River, Sumatra; B, paratype female (46.1 × 34.7 mm) (ZRC 2015.0451), Kampar River, Sumatra. A, overall dorsal view; B, thoraCiC sternum showing vulvae." httpUri="https://zenodo.org/record/1052305/files/figure.png" pageId="10" pageNumber="505">6A</figureCitation>
) (distinctly sinuous in 
<taxonomicName id="29E14D3E93793651FA25FDA436CFFDFF" authorityName="Nobili" authorityYear="1901" class="Malacostraca" family="Gecarcinucidae" genus="Parathelphusa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="505" phylum="Arthropoda" rank="species" species="oxygona">
<emphasis id="DC95EAAF93793651FA25FDA436CFFDFF" italics="true" pageId="10" pageNumber="505">P. oxygona</emphasis>
</taxonomicName>
, cf. 
<bibRefCitation id="8A704B4C93793651FE88FD8037A8FDFF" author="Ng" box="[298,401,548,573]" pageId="10" pageNumber="505" refString="Ng, P. K. L. (2014) The identity of the Sarawak freshwater Crab Parathelphusa oxygona Nobili, 1901, with desCription of a new speCies, Parathelphusa nobilii, from Western Kalimantan, Indonesia, Borneo (CrustaCea: BraChyura: GeCarCinuCidae). Zootaxa, 3774 (1), 31 - 44. http: // dx. doi. org / 10.11646 / zootaxa. 3774.1.2" type="journal article" year="2014">Ng 2014</bibRefCitation>
: figs. 1A, 2E), and the G1 is relatively stouter with only the distal part gently curved (
<figureCitation id="76DA2A3893793651FACCFD813727FDA3" captionStart="FIGURE 5" captionStartId="6.[151,250,1907,1929]" captionTargetBox="[224,1324,206,1867]" captionTargetId="figure@6.[206,1359,193,1881]" captionTargetPageId="6" captionText="FIGURE 5. Parathelphusa pardus sp. nov. A – D, holotype male (37.6 × 28.2 mm) (MZB Cru 4388), Kampar River, Sumatra; E – H, paratype male (34.9 × 26.6 mm) (ZRC 2015.0452), Kampar River, Sumatra. A, E, ventral view of left G 1; B, F, ventral view of distal part of left G 1; C, G, dorsal view of distal part of left G 1; D, H, left G 2 s. SCales: A, D = 1.0 mm; B, C = 0.5 mm." httpUri="https://zenodo.org/record/1052303/files/figure.png" pageId="10" pageNumber="505">Fig. 5A–D, E–G</figureCitation>
) (more slender with the distal part more strongly curved in 
<taxonomicName id="29E14D3E93793651FC18FDEC320DFDA3" authorityName="Nobili" authorityYear="1901" box="[954,1076,585,608]" class="Malacostraca" family="Gecarcinucidae" genus="Parathelphusa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="505" phylum="Arthropoda" rank="species" species="oxygona">
<emphasis id="DC95EAAF93793651FC18FDEC320DFDA3" box="[954,1076,585,608]" italics="true" pageId="10" pageNumber="505">P. oxygona</emphasis>
</taxonomicName>
, cf. 
<bibRefCitation id="8A704B4C93793651FBC4FDED32F2FDA3" author="Ng" box="[1126,1227,584,608]" pageId="10" pageNumber="505" refString="Ng, P. K. L. (2014) The identity of the Sarawak freshwater Crab Parathelphusa oxygona Nobili, 1901, with desCription of a new speCies, Parathelphusa nobilii, from Western Kalimantan, Indonesia, Borneo (CrustaCea: BraChyura: GeCarCinuCidae). Zootaxa, 3774 (1), 31 - 44. http: // dx. doi. org / 10.11646 / zootaxa. 3774.1.2" type="journal article" year="2014">Ng 2014</bibRefCitation>
: fig. 4A, B)
</paragraph>
<paragraph id="EE5E36BD93793651FF65FDC9370EFCFB" blockId="10.[151,1437,151,1040]" pageId="10" pageNumber="505">
The cleft on the outer margin of the proximal part of the G 
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<taxonomicName id="29E14D3E93793651FC2FFDC835CFFD47" authority="Ng &amp; Riady, 2016" authorityName="Ng &amp; Riady" authorityYear="2016" box="[909,1014,620,644]" class="Malacostraca" family="Gecarcinucidae" genus="Parathelphusa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="505" phylum="Arthropoda" rank="species" species="pardus" status="sp. nov.">
<emphasis id="DC95EAAF93793651FC2FFDC835CFFD47" box="[909,1014,620,644]" italics="true" pageId="10" pageNumber="505">P. pardus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="C7A657D493793651FC5FFDC9326FFD46" box="[1021,1110,620,645]" pageId="10" pageNumber="505" rank="species">
<emphasis id="DC95EAAF93793651FC5FFDC9326FFD46" bold="true" box="[1021,1110,620,645]" pageId="10" pageNumber="505">sp. nov.</emphasis>
</taxonomicNameLabel>
is a character also present in 
<taxonomicName id="29E14D3E93793651FF35FD343723FD6B" authorityName="De Man" authorityYear="1879" box="[151,282,656,680]" class="Malacostraca" family="Gecarcinucidae" genus="Parathelphusa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="505" phylum="Arthropoda" rank="species" species="maculata">
<emphasis id="DC95EAAF93793651FF35FD343723FD6B" box="[151,282,656,680]" italics="true" pageId="10" pageNumber="505">P. maculata</emphasis>
</taxonomicName>
from 
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and Peninsular 
<collectingCountry id="96F6762D93793651FDCEFD2A34EFFD6B" box="[620,726,655,680]" name="Malaysia" pageId="10" pageNumber="505">Malaysia</collectingCountry>
, and 
<taxonomicName id="29E14D3E93793651FCADFD3435BFFD6B" authorityName="De Man" authorityYear="1879" box="[783,902,657,680]" class="Malacostraca" family="Gecarcinucidae" genus="Parathelphusa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="505" phylum="Arthropoda" rank="species" species="convexa">
<emphasis id="DC95EAAF93793651FCADFD3435BFFD6B" box="[783,902,657,680]" italics="true" pageId="10" pageNumber="505">P. convexa</emphasis>
</taxonomicName>
from southern 
<collectingRegion id="2C25F85F93793651FB92FD3532B4FD6B" box="[1072,1165,656,680]" country="Indonesia" name="Sumatera Utara" pageId="10" pageNumber="505">Sumatra</collectingRegion>
and 
<collectingRegion id="2C25F85F93793651FB61FD3532CCFD64" box="[1219,1269,656,679]" country="Indonesia" name="Jawa Timur" pageId="10" pageNumber="505">Java</collectingRegion>
(cf. 
<bibRefCitation id="8A704B4C93793651FA85FD3533A1FD6B" author="Ng" box="[1319,1432,656,680]" pageId="10" pageNumber="505" refString="Ng, P. K. L. (1990 a) The taxonomy and biology of Parathelphusa maculata De Man, 1879 (CrustaCea: DeCapoda: BraChyura: Parathelphusidae). Malayan Nature Journal, 44 (1), 45 - 60." type="journal article" year="1990" yearSuffix="a">Ng 1990a</bibRefCitation>
: figs. 4A, B, F, I, 5A, B, E, F, H; 
<bibRefCitation id="8A704B4C93793651FDAFFD1034BBFD0E" author="Ng" box="[525,642,692,717]" pageId="10" pageNumber="505" refString="Ng, P. K. L. (1990 b) Parathelphusa reticulata speC. nov., a new speCies of freshwater Crab from blaCkwater swamps in Singapore (CrustaCea: DeCapoda: BraChyura: GeCarCinuCoidea). Zoologische Mededelingen, 63 (18), 241 - 254." type="journal article" year="1990" yearSuffix="b">Ng 1990b</bibRefCitation>
: fig. 8K, L). Adults of both 
<taxonomicName id="29E14D3E93793651FC66FD103270FD0F" authorityName="De Man" authorityYear="1879" box="[964,1097,692,716]" class="Malacostraca" family="Gecarcinucidae" genus="Parathelphusa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="505" phylum="Arthropoda" rank="species" species="maculata">
<emphasis id="DC95EAAF93793651FC66FD103270FD0F" box="[964,1097,692,716]" italics="true" pageId="10" pageNumber="505">P. maculata</emphasis>
</taxonomicName>
and 
<taxonomicName id="29E14D3E93793651FB23FD1032C3FD0F" authorityName="De Man" authorityYear="1879" box="[1153,1274,693,716]" class="Malacostraca" family="Gecarcinucidae" genus="Parathelphusa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="505" phylum="Arthropoda" rank="species" species="convexa">
<emphasis id="DC95EAAF93793651FB23FD1032C3FD0F" box="[1153,1274,693,716]" italics="true" pageId="10" pageNumber="505">P. convexa</emphasis>
</taxonomicName>
do not have a spotted or reticulated colour pattern (
<bibRefCitation id="8A704B4C93793651FD98FD7D3544FD33" author="Ng" box="[570,893,727,752]" pageId="10" pageNumber="505" refString="Ng, P. K. L. (1990 a) The taxonomy and biology of Parathelphusa maculata De Man, 1879 (CrustaCea: DeCapoda: BraChyura: Parathelphusidae). Malayan Nature Journal, 44 (1), 45 - 60." type="journal article" year="1990" yearSuffix="a">Ng 1990a; unpublished data</bibRefCitation>
). While the carapace of 
<taxonomicName id="29E14D3E93793651FB32FD7C332DFD33" authorityName="De Man" authorityYear="1879" box="[1168,1300,728,752]" class="Malacostraca" family="Gecarcinucidae" genus="Parathelphusa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="505" phylum="Arthropoda" rank="species" species="maculata">
<emphasis id="DC95EAAF93793651FB32FD7C332DFD33" box="[1168,1300,728,752]" italics="true" pageId="10" pageNumber="505">P. maculata</emphasis>
</taxonomicName>
has a much flatter carapace than 
<taxonomicName id="29E14D3E93793651FE23FD5837D0FCD7" authority="Ng &amp; Riady, 2016" authorityName="Ng &amp; Riady" authorityYear="2016" box="[385,489,764,788]" class="Malacostraca" family="Gecarcinucidae" genus="Parathelphusa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="505" phylum="Arthropoda" rank="species" species="pardus" status="sp. nov.">
<emphasis id="DC95EAAF93793651FE23FD5837D0FCD7" box="[385,489,764,788]" italics="true" pageId="10" pageNumber="505">P. pardus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="C7A657D493793651FE52FD593473FCD6" box="[496,586,764,789]" pageId="10" pageNumber="505" rank="species">
<emphasis id="DC95EAAF93793651FE52FD593473FCD6" bold="true" box="[496,586,764,789]" pageId="10" pageNumber="505">sp. nov.</emphasis>
</taxonomicNameLabel>
(cf. 
<bibRefCitation id="8A704B4C93793651FDDCFD5834C9FCD7" author="Ng" box="[638,752,764,789]" pageId="10" pageNumber="505" refString="Ng, P. K. L. (1990 a) The taxonomy and biology of Parathelphusa maculata De Man, 1879 (CrustaCea: DeCapoda: BraChyura: Parathelphusidae). Malayan Nature Journal, 44 (1), 45 - 60." type="journal article" year="1990" yearSuffix="a">Ng 1990a</bibRefCitation>
: figs. 1, 2), that of 
<taxonomicName id="29E14D3E93793651FC61FD583203FCD7" authorityName="De Man" authorityYear="1879" box="[963,1082,765,788]" class="Malacostraca" family="Gecarcinucidae" genus="Parathelphusa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="505" phylum="Arthropoda" rank="species" species="convexa">
<emphasis id="DC95EAAF93793651FC61FD583203FCD7" box="[963,1082,765,788]" italics="true" pageId="10" pageNumber="505">P. convexa</emphasis>
</taxonomicName>
is more inflated (cf. 
<bibRefCitation id="8A704B4C93793651FA87FD5833A1FCD7" author="Bott" box="[1317,1432,764,788]" pageId="10" pageNumber="505" refString="Bott, R. (1970) Die Susswasserkrabben von Europa, Asien, Australien und ihre StammesgesChiChte. Eine Revision der Potamoidea und der Parathelphusoidea (CrustaCea, DeCapoda). Abhandlungen der senckenbergischen naturforschenden Gesellschaft, 526, 1 - 338, figs. 1 - 8, pls. 1 - 58, map 1." type="journal article" year="1970">Bott 1970</bibRefCitation>
: pl. 23 fig. 69).
</paragraph>
</subSubSection>
<subSubSection id="A6FB653693793651FF65FCE1354EFBD3" pageId="10" pageNumber="505" type="biology_ecology">
<paragraph id="EE5E36BD93793651FF65FCE1354EFBD3" blockId="10.[151,1437,151,1040]" pageId="10" pageNumber="505">
<emphasis id="DC95EAAF93793651FF65FCE1371DFC9E" bold="true" box="[199,292,836,861]" pageId="10" pageNumber="505">Ecology</emphasis>
. All the specimens of 
<taxonomicName id="29E14D3E93793651FD86FCE034B4FC9F" authority="Ng &amp; Riady, 2016" authorityName="Ng &amp; Riady" authorityYear="2016" box="[548,653,836,860]" class="Malacostraca" family="Gecarcinucidae" genus="Parathelphusa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="505" phylum="Arthropoda" rank="species" species="pardus" status="sp. nov.">
<emphasis id="DC95EAAF93793651FD86FCE034B4FC9F" box="[548,653,836,860]" italics="true" pageId="10" pageNumber="505">P. pardus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="C7A657D493793651FD36FCE134D7FC9E" box="[660,750,836,861]" pageId="10" pageNumber="505" rank="species">
<emphasis id="DC95EAAF93793651FD36FCE134D7FC9E" bold="true" box="[660,750,836,861]" pageId="10" pageNumber="505">sp. nov.</emphasis>
</taxonomicNameLabel>
were collected from various tributaries of the Kampar River in 
<collectingRegion id="2C25F85F93793651FF14FCC23765FC43" box="[182,348,871,896]" country="Indonesia" name="Riau" pageId="10" pageNumber="505">Riau Province</collectingRegion>
, 
<collectingRegion id="2C25F85F93793651FEC8FCCD37F2FC43" box="[362,459,872,896]" country="Indonesia" name="Sumatera Utara" pageId="10" pageNumber="505">Sumatra</collectingRegion>
. Their preferred habitat is swampy areas along the river, often living in well-shaded areas under the vegetation (
<figureCitation id="76DA2A3893793651FE6EFC293428FC67" box="[460,529,908,933]" captionStart="FIGURE 7" captionStartId="9.[151,250,1885,1907]" captionTargetBox="[152,1438,193,1857]" captionTargetId="figure@9.[152,1438,193,1857]" captionTargetPageId="9" captionText="FIGURE 7. The habitat of Parathelphusa pardus sp. nov. A, one of the tributaries flowing through the swamp forest; B, a shallow seCtion of the swamp with Barclaya sp. and a Crab foraging at night." httpUri="https://zenodo.org/record/1052307/files/figure.png" pageId="10" pageNumber="505">Fig. 7</figureCitation>
). The main aquatic plant in these areas is a species of 
<taxonomicName id="29E14D3E93793651FBD0FC293345FC67" authority="Wallich, 1827" authorityName="Wallich" authorityYear="1827" box="[1138,1404,908,933]" class="Magnoliopsida" family="Nymphaeaceae" genus="Barclaya" higherTaxonomySource="GBIF" kingdom="Plantae" order="Nymphaeales" pageId="10" pageNumber="505" phylum="Tracheophyta" rank="genus">
<emphasis id="DC95EAAF93793651FBD0FC2932EEFC67" box="[1138,1239,908,932]" italics="true" pageId="10" pageNumber="505">Barclaya</emphasis>
Wallich, 1827
</taxonomicName>
(= 
<taxonomicName id="29E14D3E93793651FF35FC1537D8FC0B" authority="Wallich, 1827" authorityName="Wallich" authorityYear="1827" box="[151,481,943,968]" class="Magnoliopsida" family="Nymphaeaceae" genus="Hydrostemma" higherTaxonomySource="GBIF" kingdom="Plantae" order="Nymphaeales" pageId="10" pageNumber="505" phylum="Tracheophyta" rank="genus">
<emphasis id="DC95EAAF93793651FF35FC15370AFC0B" box="[151,307,944,968]" italics="true" pageId="10" pageNumber="505">Hydrostemma</emphasis>
Wallich, 1827
</taxonomicName>
) (family 
<taxonomicName id="29E14D3E93793651FDF6FC0A3538FC0B" box="[596,769,943,968]" family="Nymphaeaceae" higherTaxonomySource="GBIF" kingdom="Plantae" order="Nymphaeales" pageId="10" pageNumber="505" phylum="Magnoliopsida" rank="family">Nymphaeaceae</taxonomicName>
) (see 
<bibRefCitation id="8A704B4C93793651FCECFC0A3234FC0B" author="Lok" box="[846,1037,943,968]" pageId="10" pageNumber="505" refString="Lok, A. F. S. L., Ang, W. F., Lee, S. M. L., Tan, H. H. &amp; Tan, H. T. W. (2009) The status and distribution of Barclaya (NymphaeaCeae) in Singapore. Nature in Singapore, 2, 237 - 245." type="journal article" year="2009">
Lok 
<emphasis id="DC95EAAF93793651FC25FC1435FDFC0B" box="[903,964,944,968]" italics="true" pageId="10" pageNumber="505">et al.</emphasis>
2009
</bibRefCitation>
). The crabs are mainly nocturnal, coming out to forage only after dark, the same habit for swamp species like 
<taxonomicName id="29E14D3E93793651FBA4FC7032B6FC2F" authorityName="Ng" authorityYear="1990" box="[1030,1167,980,1004]" class="Malacostraca" family="Gecarcinucidae" genus="Parathelphusa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="505" phylum="Arthropoda" rank="species" species="reticulata">
<emphasis id="DC95EAAF93793651FBA4FC7032B6FC2F" box="[1030,1167,980,1004]" italics="true" pageId="10" pageNumber="505">P. reticulata</emphasis>
</taxonomicName>
(see 
<bibRefCitation id="8A704B4C93793651FB6CFC70337AFC2E" author="Ng" box="[1230,1347,980,1005]" pageId="10" pageNumber="505" refString="Ng, P. K. L. (1990 b) Parathelphusa reticulata speC. nov., a new speCies of freshwater Crab from blaCkwater swamps in Singapore (CrustaCea: DeCapoda: BraChyura: GeCarCinuCoidea). Zoologische Mededelingen, 63 (18), 241 - 254." type="journal article" year="1990" yearSuffix="b">Ng 1990b</bibRefCitation>
, c). For more details of the biology of the species, see 
<bibRefCitation id="8A704B4C93793651FD3FFC52354AFBD3" box="[669,883,1015,1040]" pageId="10" pageNumber="505" refString="Riady, R., Mahatma, R. &amp; Windarti (2014) Inventarisasi kepiting air tawar di KeCamatan Kampar Utara Kabupaten Kampar Provinsi Riau. JOM (Jurnal Online Mahasiswa) FMIPA (Fakultas Matematika dan Ilmu Pengetahuan Alam), 1 (2), 471 - 479. [Universitas Riau]" type="journal article">
Riady 
<emphasis id="DC95EAAF93793651FD44FC5C3526FBD3" box="[742,799,1016,1040]" italics="true" pageId="10" pageNumber="505">et al.</emphasis>
(2014)
</bibRefCitation>
.
</paragraph>
</subSubSection>
</treatment>
</document>