treatments-xml/data/5A/35/41/5A35417E7C64355FAF7BEE43FCF15DA2.xml
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<document id="B079E21DF0B951510431836DB40CD7B8" ID-CLB-Dataset="295745" ID-DOI="10.1590/1982-0224-2022-0075" ID-GBIF-Dataset="bdd12854-e0c4-400c-bd81-fbd691e9c073" ID-ISSN="1982-0224" ID-Zenodo-Dep="11124758" ID-ZooBank="4FD4576C-30E9-49A0-B668-9D2C01BE312A" IM.bibliography_approvedBy="tatiana" IM.illustrations_approvedBy="tatiana" IM.materialsCitations_approvedBy="tatiana" IM.metadata_approvedBy="tatiana" IM.tables_approvedBy="tatiana" IM.treatments_approvedBy="tatiana" checkinTime="1712845666065" checkinUser="felipe" docAuthor="Shibatta, Oscar Akio &amp; Souza-Shibatta, Lenice" docDate="2023" docId="5A35417E7C64355FAF7BEE43FCF15DA2" docLanguage="en" docName="NeotropIchthyol.2023.21.1.e220075.pdf" docOrigin="Neotropical Ichthyology (e 220075) 21 (1)" docSource="http://dx.doi.org/10.1590/1982-0224-2022-0075" docStyle="DocumentStyle:FC0C3FD10650AF21F8DA93D98D275823.3:NeotropIchthyol.2020-.journal_article" docStyleId="FC0C3FD10650AF21F8DA93D98D275823" docStyleName="NeotropIchthyol.2020-.journal_article" docStyleVersion="3" docTitle="Rhyacoglanis varii Shibatta &amp; Souza-Shibatta 2023, new species" docType="treatment" docVersion="3" lastPageNumber="16" masterDocId="A60C39067C673550AC69ED7BFFC35E62" masterDocTitle="New species of Rhyacoglanis (Siluriformes: Pseudopimelodidae) from the upper rio Tocantins basin" masterLastPageNumber="20" masterPageNumber="1" pageNumber="4" updateTime="1715068936461" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
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<mods:title id="C1FFE2BFC0FEB1D850F534F57783F9F5">New species of Rhyacoglanis (Siluriformes: Pseudopimelodidae) from the upper rio Tocantins basin</mods:title>
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<mods:namePart id="72EB9424E0B11F3BCCC53AA99F4F8C1F">Shibatta, Oscar Akio</mods:namePart>
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<mods:namePart id="AC221B246AED81B7FDE4A68EFFB544F6">Souza-Shibatta, Lenice</mods:namePart>
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<mods:title id="05649E462DEFB01A5080D1D33A9112A3">Neotropical Ichthyology</mods:title>
</mods:titleInfo>
<mods:part id="B3C052D10A9D78E15D04E2A6DCD725CE">
<mods:date id="2BB8FFEC2242F56EA6C4EF93E0A40D34">2023</mods:date>
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<mods:classification id="A30BDAA0D3E35B8CDEB4D33D8E48CC6A">journal article</mods:classification>
<mods:identifier id="764D8251FDE5D114D62C22E839905AEE" type="CLB-Dataset">295745</mods:identifier>
<mods:identifier id="0E51DD786DC57A682B447529283AFBDE" type="DOI">10.1590/1982-0224-2022-0075</mods:identifier>
<mods:identifier id="5DA6FB1C0403DA6F00F12361D9D40B49" type="GBIF-Dataset">bdd12854-e0c4-400c-bd81-fbd691e9c073</mods:identifier>
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<treatment id="5A35417E7C64355FAF7BEE43FCF15DA2" ID-DOI="http://doi.org/10.5281/zenodo.11124776" ID-Zenodo-Dep="11124776" LSID="urn:lsid:plazi:treatment:5A35417E7C64355FAF7BEE43FCF15DA2" httpUri="http://treatment.plazi.org/id/5A35417E7C64355FAF7BEE43FCF15DA2" lastPageId="15" lastPageNumber="16" pageId="3" pageNumber="4">
<subSubSection id="9A86A3E37C643553AF7BEE43FAD55DF8" pageId="3" pageNumber="4" type="nomenclature">
<paragraph id="D223F0687C643553AF7BEE43FB4B5D32" blockId="3.[786,1160,824,848]" box="[786,1160,824,848]" pageId="3" pageNumber="4">
<heading id="896B47047C643553AF7BEE43FB4B5D32" box="[786,1160,824,848]" centered="true" fontSize="10" level="2" pageId="3" pageNumber="4" reason="2">
<taxonomicName id="159C8BEB7C643553AF7BEE43FC345D32" authority="Shibatta &amp; Souza-Shibatta, 2023" authorityName="Shibatta &amp; Souza-Shibatta" authorityYear="2023" box="[786,1015,824,848]" family="Pseudopimelodidae" genus="Rhyacoglanis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Siluriformes" pageId="3" pageNumber="4" phylum="Chordata" rank="species" species="varii" status="sp. nov.">
<emphasis id="E0E82C7A7C643553AF7BEE43FC345D32" bold="true" box="[786,1015,824,848]" pageId="3" pageNumber="4">Rhyacoglanis varii</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="FBDB91017C643553A86CEE42FB4B5D32" box="[1029,1160,824,848]" pageId="3" pageNumber="4" rank="species">new species</taxonomicNameLabel>
</heading>
</paragraph>
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<uri id="A60DFC6A7C643553AEEDEEFCFAD55DF8" box="[644,1302,903,922]" pageId="3" pageNumber="4">
urn:lsid:zoobank.org:act:
<uuid id="A63ACABD7C643553AF08EEFCFAD55DF8" box="[865,1302,903,922]" pageId="3" pageNumber="4">2A1AD513-0AB2-4D10-91AA-9C3F865A7152</uuid>
</uri>
</paragraph>
</subSubSection>
<subSubSection id="9A86A3E37C643553AF0FEEB5FBA35A4D" pageId="3" pageNumber="4" type="description">
<paragraph id="D223F0687C643553AF0FEEB5FBF65D84" blockId="3.[870,1077,974,998]" box="[870,1077,974,998]" pageId="3" pageNumber="4">
(
<figureCitation id="4AA7ECED7C643553AF07EEB5FC155D84" box="[878,982,974,998]" captionStart-0="FIGURE 1" captionStart-1="FIGURE 2" captionStart-2="FIGURE 3" captionStartId-0="4.[488,561,1761,1781]" captionStartId-1="5.[136,209,1873,1893]" captionStartId-2="6.[136,209,1686,1706]" captionTargetBox-0="[488,1459,432,1737]" captionTargetBox-1="[147,1436,1497,1804]" captionTargetBox-2="[178,1437,733,1622]" captionTargetId-0="figure-20@4.[488,1459,432,1737]" captionTargetId-1="figure-405@5.[136,1459,1471,1821]" captionTargetId-2="figure-21@6.[138,1459,733,1664]" captionTargetPageId-0="4" captionTargetPageId-1="5" captionTargetPageId-2="6" captionText-0="FIGURE 1 | Holotype of Rhyacoglanis varii, MZUEL 22317, 67.1 mm SL, rio do Peixe, upper rio Tocantins basin, Goianésia, state of Goiás, Brazil." captionText-1="FIGURE 2 | Rhyacoglanis varii pectoral-fin spine, paratype, MZUEL 6039, 55.5 mm SL. Scale bar = 1.0 mm." captionText-2="FIGURE 3 | Rhyacoglanis varii caudal-fin skeleton, paratype, MZUEL 6039, 53.5 mm SL. Bones in yellow ocher and cartilage in blue. H1 to H5 = hypurals 1 to 5. Scale bar = 1.0 mm." figureDoi-0="http://doi.org/10.5281/zenodo.11124760" figureDoi-1="http://doi.org/10.5281/zenodo.11124762" figureDoi-2="http://doi.org/10.5281/zenodo.11124764" httpUri-0="https://zenodo.org/record/11124760/files/figure.png" httpUri-1="https://zenodo.org/record/11124762/files/figure.png" httpUri-2="https://zenodo.org/record/11124764/files/figure.png" pageId="3" pageNumber="4">Figs. 13</figureCitation>
;
<tableCitation id="9F1EC5D37C643553AF8AEEB5FBE95D84" box="[995,1066,974,998]" captionStart="TABLE 1" captionStartId="7.[488,550,193,213]" captionTargetId="graphics-25@7.[488,1459,389,1328]" captionTargetPageId="7" captionText="TABLE 1 | Morphometry of Rhyacoglanis varii from upper rio Tocantins basin (n = 42). Statistics of minimum (Min), maximum (Max), mean, and standard deviation (SD) include holotype." httpUri="http://table.plazi.org/id/86E3A0E07C603557AD81EDBAFADD5E99" pageId="3" pageNumber="4" tableUuid="86E3A0E07C603557AD81EDBAFADD5E99">Tab. 1</tableCitation>
)
</paragraph>
<paragraph id="D223F0687C643553AE6FE967FBA35A4D" blockId="3.[518,1120,1052,1072]" box="[518,1120,1052,1072]" pageId="3" pageNumber="4">
<taxonomicName id="159C8BEB7C643553AE6FE967FD505A52" box="[518,659,1052,1072]" family="Pseudopimelodidae" genus="Rhyacoglanis" kingdom="Animalia" order="Siluriformes" pageId="3" pageNumber="4" phylum="Chordata" rank="species" species="undetermined">
<emphasis id="E0E82C7A7C643553AE6FE967FDB65A4D" box="[518,629,1052,1071]" italics="true" pageId="3" pageNumber="4">Rhyacoglanis</emphasis>
sp.
</taxonomicName>
―Abrahão
<emphasis id="E0E82C7A7C643553AE95E966FCDC5A4D" box="[764,799,1052,1071]" italics="true" pageId="3" pageNumber="4">et al</emphasis>
., 2018:3 [brain gross morphology].
</paragraph>
</subSubSection>
<subSubSection id="9A86A3E37C643553AE6FE918FAF55BE2" pageId="3" pageNumber="4" type="materials_examined">
<paragraph id="D223F0687C643553AE6FE918FC705AA4" blockId="3.[488,1459,1123,1222]" pageId="3" pageNumber="4">
<materialsCitation id="62F4FA357C643553AE6FE918FC705AA4" collectingDate="2010-07-14" collectionCode="R" collectorName="O. A. Shibatta &amp; E. Santana &amp; A. Claro-Garcia &amp; Jarduli." country="Brazil" latitude="-15.473277" location="Goianesia" longLatPrecision="1" longitude="-49.224693" municipality="Peixe" pageId="3" pageNumber="4" specimenCode="MZUEL 22317" specimenCount="1" stateProvince="Goias" typeStatus="holotype">
<emphasis id="E0E82C7A7C643553AE6FE918FDBE5A19" bold="true" box="[518,637,1123,1147]" pageId="3" pageNumber="4">
<typeStatus id="0D274ECA7C643553AE6FE918FDBA5A19" box="[518,633,1123,1147]" pageId="3" pageNumber="4" type="holotype">Holotype</typeStatus>
.
</emphasis>
<specimenCode id="823A58137C643553AEEEE91FFCF95A19" box="[647,826,1123,1147]" collectionCode="MZUEL" pageId="3" pageNumber="4">MZUEL 22317</specimenCode>
,
<quantity id="15645D8D7C643553AF2EE918FC6E5A19" box="[839,941,1123,1147]" metricMagnitude="-2" metricUnit="m" metricValue="6.709999999999999" pageId="3" pageNumber="4" unit="mm" value="67.1">67.1 mm</quantity>
SL,
<collectingCountry id="AA8BB0F87C643553AF8DE918FBEB5A19" box="[996,1064,1123,1147]" name="Brazil" pageId="3" pageNumber="4">Brazil</collectingCountry>
,
<collectingRegion id="10583E8A7C643553A85FE918FBBB5A19" box="[1078,1144,1123,1147]" country="Brazil" name="Goias" pageId="3" pageNumber="4">Goiás</collectingRegion>
, upper rio Tocantins basin, rio do
<collectingMunicipality id="32476A127C643553AE50E9F3FDBB5AC2" box="[569,632,1160,1184]" pageId="3" pageNumber="4">Peixe</collectingMunicipality>
,
<location id="D743A6B37C643553AEEEE9F3FD345AC2" LSID="urn:lsid:plazi:treatment:5A35417E7C64355FAF7BEE43FCF15DA2:D743A6B37C643553AEEEE9F3FD345AC2" box="[647,759,1160,1184]" country="Brazil" latitude="-15.473277" longLatPrecision="1" longitude="-49.224693" municipality="Peixe" name="Goianesia" pageId="3" pageNumber="4" stateProvince="Goias">Goianésia</location>
,
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15
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2823.8”S
</geoCoordinate>
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49
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1328.9”W
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,
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<collectingDate id="B6662F407C643553A825E9F2FB175AC3" box="[1100,1236,1160,1185]" pageId="3" pageNumber="4" value="2010-07-14">14 Jul 2010</collectingDate>
</date>
,
<collectorName id="7F6995BE7C643553A88AE9F2FA4F5AC2" box="[1251,1420,1160,1185]" pageId="3" pageNumber="4">O. A. Shibatta</collectorName>
,
<collectorName id="7F6995BE7C643553A9F5E9F2FD825AA4" pageId="3" pageNumber="4">E. Santana</collectorName>
,
<collectorName id="7F6995BE7C643553AE25E9D5FCC25AA4" box="[588,769,1198,1222]" pageId="3" pageNumber="4">A. Claro-García</collectorName>
&amp; L.
<collectionCode id="B48D68AD7C643553AF2BE9D5FC9B5AA7" box="[834,856,1198,1221]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="3" pageNumber="4">R</collectionCode>
.
<collectorName id="7F6995BE7C643553AF0BE9D5FC705AA4" box="[866,947,1198,1222]" pageId="3" pageNumber="4">Jarduli.</collectorName>
</materialsCitation>
</paragraph>
<paragraph id="D223F0687C643553AE6FE982FAF55BE2" blockId="3.[488,1459,1272,1409]" pageId="3" pageNumber="4">
<materialsCitation id="62F4FA357C643553AE6FE982FDCF5B39" collectingDate="2010-07-14" collectingDateMax="1996-11-04" collectingDateMin="1996-10-28" collectionCode="MZUEL" collectorName="O. A. Shibatta &amp; E. Santana &amp; A. Claro-Garcia &amp; Jarduli." country="Brazil" county="rio Tocantinzinho" latitude="-15.473277" location="Goianesia" longLatPrecision="1" longitude="-49.224693" municipality="Peixe" pageId="3" pageNumber="4" specimenCode="MZUEL 6039, 29.6, 69.1" specimenCount="1" stateProvince="Goias" typeStatus="paratype">
<emphasis id="E0E82C7A7C643553AE6FE982FC805B72" bold="true" box="[518,835,1272,1297]" pageId="3" pageNumber="4">
<typeStatus id="0D274ECA7C643553AE6FE982FDBA5B73" box="[518,633,1273,1297]" pageId="3" pageNumber="4" type="paratype">Paratypes</typeStatus>
(
<specimenCount id="C49A3BE17C643553AEE7E983FCF55B72" box="[654,822,1272,1296]" count="20" pageId="3" pageNumber="4" type="generic">20 specimens</specimenCount>
).
</emphasis>
<specimenCode id="823A58137C643553AF39E982FC3A5B72" box="[848,1017,1272,1296]" collectionCode="MZUEL" pageId="3" pageNumber="4">MZUEL 6039</specimenCode>
,
<specimenCode id="823A58137C643553A862E983FBFF5B72" box="[1035,1084,1272,1296]" collectionCode="MZUEL" pageId="3" pageNumber="4">29.6</specimenCode>
<specimenCode id="823A58137C643553A823E983FBBB5B72" box="[1098,1144,1272,1296]" collectionCode="MZUEL" pageId="3" pageNumber="4">69.1</specimenCode>
(10 eth, 5 C&amp;S; 4 COI),
<quantity id="15645D8D7C643553AD81E865FD4F5B57" box="[488,652,1310,1334]" metricMagnitude="-2" metricUnit="m" metricValue="4.734999999999999" metricValueMax="6.909999999999999" metricValueMin="2.56" pageId="3" pageNumber="4" unit="mm" value="47.349999999999994" valueMax="69.1" valueMin="25.6">25.669.1 mm</quantity>
SL, same data as the holotype.
<specimenCode id="823A58137C643553A86BE865FB715B54" box="[1026,1202,1310,1334]" collectionCode="MZUSP" country="Brazil" httpUri="http://grbio.org/cool/6yx0-3nm7" name="Museu de Zoologia da Universidade de Sao Paulo" pageId="3" pageNumber="4">MZUSP 54082</specimenCode>
,
<specimenCode id="823A58137C643553A8A8E865FB0C5B54" box="[1217,1231,1310,1334]" collectionCode="MZUSP" country="Brazil" httpUri="http://grbio.org/cool/6yx0-3nm7" name="Museu de Zoologia da Universidade de Sao Paulo" pageId="3" pageNumber="4">5</specimenCode>
eth, 37.0
<quantity id="15645D8D7C643553A922E865FA705B57" box="[1355,1459,1310,1334]" metricMagnitude="-2" metricUnit="m" metricValue="4.76" pageId="3" pageNumber="4" unit="mm" value="47.6">47.6 mm</quantity>
SL,
</materialsCitation>
<materialsCitation id="62F4FA357C643553AE7EE838FAF55BE2" collectingDate="1996-10-28" collectingDateMax="1996-11-04" collectingDateMin="1996-10-28" country="Brazil" county="rio Tocantinzinho" latitude="-14.208895" location="Minacu" longLatPrecision="1" longitude="-48.07088" municipality="Serra da Mesa region" pageId="3" pageNumber="4" specimenCount="1" stateProvince="Goias" typeStatus="paratype">
<collectingCountry id="AA8BB0F87C643553AE7EE838FD985B39" box="[535,603,1347,1371]" name="Brazil" pageId="3" pageNumber="4">Brazil</collectingCountry>
,
<collectingRegion id="10583E8A7C643553AE03E838FD6F5B39" box="[618,684,1347,1371]" country="Brazil" name="Goias" pageId="3" pageNumber="4">Goiás</collectingRegion>
,
<collectingCounty id="3B4288E47C643553AED2E838FC4D5B39" box="[699,910,1347,1371]" pageId="3" pageNumber="4">rio Tocantinzinho</collectingCounty>
and tributaries,
<collectingMunicipality id="32476A127C643553A827E838FA845B39" box="[1102,1351,1347,1371]" pageId="3" pageNumber="4">Serra da Mesa region</collectingMunicipality>
,
<location id="D743A6B37C643553A93FE838FA6C5B39" LSID="urn:lsid:plazi:treatment:5A35417E7C64355FAF7BEE43FCF15DA2:D743A6B37C643553A93FE838FA6C5B39" box="[1366,1455,1347,1371]" country="Brazil" county="rio Tocantinzinho" latitude="-14.208895" longLatPrecision="1" longitude="-48.07088" municipality="Serra da Mesa region" name="Minacu" pageId="3" pageNumber="4" stateProvince="Goias">Minaçu</location>
,
<geoCoordinate id="B7A896AF7C643553AD81E812FD425BE3" box="[488,641,1383,1409]" degrees="14" direction="south" minutes="12" orientation="latitude" pageId="3" pageNumber="4" precision="1" seconds="32.02" value="-14.208895">
14
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1232.02”S
</geoCoordinate>
<geoCoordinate id="B7A896AF7C643553AEE0E812FCF55BE2" box="[649,822,1383,1409]" degrees="48" direction="west" minutes="04" orientation="longitude" pageId="3" pageNumber="4" precision="1" seconds="15.17" value="-48.07088">
48
<superScript id="25E95D207C643553AECDE81CFD6F5B17" attach="none" box="[676,684,1383,1397]" fontSize="6" pageId="3" pageNumber="4">o</superScript>
0415.17”W
</geoCoordinate>
,
<date id="A622D6A87C643553AF28E812FBFF5BE2" box="[833,1084,1384,1409]" pageId="3" pageNumber="4" value="1996-10-28" valueMax="1996-11-04" valueMin="1996-10-28">
<collectingDate id="B6662F407C643553AF28E812FBFF5BE2" box="[833,1084,1384,1409]" pageId="3" pageNumber="4" value="1996-10-28" valueMax="1996-11-04" valueMin="1996-10-28">28 Oct to 4 Nov 1996</collectingDate>
</date>
,
<collectionCode id="B48D68AD7C643553A821E812FB645BE3" box="[1096,1191,1385,1409]" country="Brazil" httpUri="http://grbio.org/cool/6yx0-3nm7" name="Museu de Zoologia da Universidade de Sao Paulo" pageId="3" pageNumber="4">MZUSP</collectionCode>
/
<collectionCode id="B48D68AD7C643553A8C6E812FB385BE3" box="[1199,1275,1385,1409]" country="Brazil" httpUri="http://grbio.org/cool/zi1i-a0b5" name="Museu Nacional/Universidade Federal de Rio de Janeiro" pageId="3" pageNumber="4">MNRJ</collectionCode>
staff.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="9A86A3E37C643554AE6FE8C8FC805F09" lastPageId="4" lastPageNumber="5" pageId="3" pageNumber="4" type="diagnosis">
<paragraph id="D223F0687C643554AE6FE8C8FC805F09" blockId="3.[488,1459,1459,1894]" lastBlockId="4.[488,1459,190,363]" lastPageId="4" lastPageNumber="5" pageId="3" pageNumber="4">
<emphasis id="E0E82C7A7C643553AE6FE8C8FD455BA9" bold="true" box="[518,646,1459,1483]" pageId="3" pageNumber="4">Diagnosis.</emphasis>
<taxonomicName id="159C8BEB7C643553AEF9E8C8FC915BA9" authorityName="Shibatta &amp; Souza-Shibatta" authorityYear="2023" box="[656,850,1459,1483]" family="Pseudopimelodidae" genus="Rhyacoglanis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Siluriformes" pageId="3" pageNumber="4" phylum="Chordata" rank="species" species="varii">
<emphasis id="E0E82C7A7C643553AEF9E8C8FC915BA9" box="[656,850,1459,1483]" italics="true" pageId="3" pageNumber="4">Rhyacoglanis varii</emphasis>
</taxonomicName>
is easily distinguished from congeners (except for
<taxonomicName id="159C8BEB7C643553A9F2E8CFFDF15B92" baseAuthorityName="Boulenger" baseAuthorityYear="1887" family="Pseudopimelodidae" genus="Rhyacoglanis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Siluriformes" pageId="3" pageNumber="4" phylum="Chordata" rank="species" species="pulcher">
<emphasis id="E0E82C7A7C643553A9F2E8CFFDF15B92" italics="true" pageId="3" pageNumber="4">R. pulcher</emphasis>
</taxonomicName>
) by the color pattern, with the dark brown vertical bar below the dorsal-fin base not confluent with a dark brown bar below the adipose-fin base (
<emphasis id="E0E82C7A7C643553A8A1E884FB1E5874" box="[1224,1245,1535,1558]" italics="true" pageId="3" pageNumber="4">vs</emphasis>
. broadly confluent in
<taxonomicName id="159C8BEB7C643553AE6EEB5FFD495859" authorityName="Shibatta &amp; Vari" authorityYear="2017" box="[519,650,1572,1595]" family="Pseudopimelodidae" genus="Rhyacoglanis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Siluriformes" pageId="3" pageNumber="4" phylum="Chordata" rank="species" species="seminiger">
<emphasis id="E0E82C7A7C643553AE6EEB5FFD495859" box="[519,650,1572,1595]" italics="true" pageId="3" pageNumber="4">R. seminiger</emphasis>
</taxonomicName>
), the subdorsal dark brown bar becomes abruptly thinner ventrally (
<emphasis id="E0E82C7A7C643553A9FEEB5FFA6F5859" box="[1431,1452,1572,1595]" italics="true" pageId="3" pageNumber="4">vs</emphasis>
. broader in
<taxonomicName id="159C8BEB7C643553AE08EB32FD295802" authorityName="Shibatta &amp; Vari" authorityYear="2017" box="[609,746,1609,1632]" family="Pseudopimelodidae" genus="Rhyacoglanis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Siluriformes" pageId="3" pageNumber="4" phylum="Chordata" rank="species" species="paranensis">
<emphasis id="E0E82C7A7C643553AE08EB32FD295802" box="[609,746,1609,1632]" italics="true" pageId="3" pageNumber="4">R. paranensis</emphasis>
</taxonomicName>
), dark brown spots sparsely distributed on the body (
<emphasis id="E0E82C7A7C643553A95FEB32FA885802" box="[1334,1355,1609,1632]" italics="true" pageId="3" pageNumber="4">vs</emphasis>
. dense in
<taxonomicName id="159C8BEB7C643553AD81EB14FDAA58E4" authorityName="Shibatta &amp; Vari" authorityYear="2017" box="[488,617,1646,1670]" family="Pseudopimelodidae" genus="Rhyacoglanis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Siluriformes" pageId="3" pageNumber="4" phylum="Chordata" rank="species" species="epiblepsis">
<emphasis id="E0E82C7A7C643553AD81EB14FDAA58E4" box="[488,617,1646,1670]" italics="true" pageId="3" pageNumber="4">R. epiblepsis</emphasis>
</taxonomicName>
and
<taxonomicName id="159C8BEB7C643553AECBEB14FC9D58E4" authorityName="Shibatta, Rocha &amp; Oliveira" authorityYear="2021" box="[674,862,1646,1670]" family="Pseudopimelodidae" genus="Rhyacoglanis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Siluriformes" pageId="3" pageNumber="4" phylum="Chordata" rank="species" species="rapppydanielae">
<emphasis id="E0E82C7A7C643553AECBEB14FC9D58E4" box="[674,862,1646,1670]" italics="true" pageId="3" pageNumber="4">R. rapppydanielae</emphasis>
</taxonomicName>
), and a solid dark brown bar on caudal peduncle (
<emphasis id="E0E82C7A7C643553A9FEEB14FA6F58E4" box="[1431,1452,1647,1670]" italics="true" pageId="3" pageNumber="4">vs</emphasis>
. becoming light brown inside, resembling a circle, in
<taxonomicName id="159C8BEB7C643553A839EBEFFB1458C9" authorityName="Shibatta &amp; Vari" authorityYear="2017" box="[1104,1239,1683,1707]" family="Pseudopimelodidae" genus="Rhyacoglanis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Siluriformes" pageId="3" pageNumber="4" phylum="Chordata" rank="species" species="annulatus">
<emphasis id="E0E82C7A7C643553A839EBEFFB1458C9" box="[1104,1239,1683,1707]" italics="true" pageId="3" pageNumber="4">R. annulatus</emphasis>
</taxonomicName>
).
<taxonomicName id="159C8BEB7C643553A899EBE8FA7058C9" authorityName="Shibatta &amp; Souza-Shibatta" authorityYear="2023" box="[1264,1459,1683,1707]" family="Pseudopimelodidae" genus="Rhyacoglanis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Siluriformes" pageId="3" pageNumber="4" phylum="Chordata" rank="species" species="varii">
<emphasis id="E0E82C7A7C643553A899EBE8FA7058C9" box="[1264,1459,1683,1707]" italics="true" pageId="3" pageNumber="4">Rhyacoglanis varii</emphasis>
</taxonomicName>
has the posterior tip of the post-cleithral process reaching vertical through the base of the dorsal-fin spine (
<emphasis id="E0E82C7A7C643553AEB9EBA4FD265894" box="[720,741,1759,1782]" italics="true" pageId="3" pageNumber="4">vs</emphasis>
. not reaching in
<taxonomicName id="159C8BEB7C643553AFC0EBA4FBE95894" authorityName="Shibatta &amp; Vari" authorityYear="2017" box="[937,1066,1758,1782]" family="Pseudopimelodidae" genus="Rhyacoglanis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Siluriformes" pageId="3" pageNumber="4" phylum="Chordata" rank="species" species="epiblepsis">
<emphasis id="E0E82C7A7C643553AFC0EBA4FBE95894" box="[937,1066,1758,1782]" italics="true" pageId="3" pageNumber="4">R. epiblepsis</emphasis>
</taxonomicName>
and
<taxonomicName id="159C8BEB7C643553A80AEBA4FAD25894" box="[1123,1297,1758,1782]" family="Pseudopimelodidae" genus="Rhyacoglanis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Siluriformes" pageId="3" pageNumber="4" phylum="Chordata" rank="species" species="rappydanielae">
<emphasis id="E0E82C7A7C643553A80AEBA4FAD25894" box="[1123,1297,1758,1782]" italics="true" pageId="3" pageNumber="4">R. rappydanielae</emphasis>
</taxonomicName>
).
<taxonomicName id="159C8BEB7C643553A940EBA5FDD55979" authorityName="Shibatta &amp; Souza-Shibatta" authorityYear="2023" family="Pseudopimelodidae" genus="Rhyacoglanis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Siluriformes" pageId="3" pageNumber="4" phylum="Chordata" rank="species" species="varii">
<emphasis id="E0E82C7A7C643553A940EBA5FDD55979" italics="true" pageId="3" pageNumber="4">Rhyacoglanis varii</emphasis>
</taxonomicName>
differs from
<taxonomicName id="159C8BEB7C643553AECEEA7FFCF25979" authorityName="Shibatta &amp; Vari" authorityYear="2017" box="[679,817,1796,1819]" family="Pseudopimelodidae" genus="Rhyacoglanis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Siluriformes" pageId="3" pageNumber="4" phylum="Chordata" rank="species" species="paranensis">
<emphasis id="E0E82C7A7C643553AECEEA7FFCF25979" box="[679,817,1796,1819]" italics="true" pageId="3" pageNumber="4">R. paranensis</emphasis>
</taxonomicName>
by the longer distance of anus to anal-fin origin (mean = 11.4±0.8%
<emphasis id="E0E82C7A7C643553AE0DEA52FDBA5922" box="[612,633,1833,1856]" italics="true" pageId="3" pageNumber="4">vs</emphasis>
. 10.0±1.4% SL), maxillary barbel length (mean = 79.6±5.8%
<emphasis id="E0E82C7A7C643553A972EA52FAF35922" box="[1307,1328,1833,1856]" italics="true" pageId="3" pageNumber="4">vs</emphasis>
. 74.8±8.7% SL), and caudal peduncle depth (mean = 9.5±0.5%
<emphasis id="E0E82C7A7C643553A85AEA34FB8B5904" box="[1075,1096,1871,1894]" italics="true" pageId="3" pageNumber="4">vs</emphasis>
. 8.8±0.5% SL); minor posterior nostril to eye distance (mean = 4.8±0.9%
<emphasis id="E0E82C7A7C633554AF8AEDC4FC3B5EB4" box="[995,1016,191,214]" italics="true" pageId="4" pageNumber="5">vs</emphasis>
. 6.7±1.3% SL), body depth (mean = 18.2±1.5%
<emphasis id="E0E82C7A7C633554AE0EED9FFDBF5E99" box="[615,636,228,251]" italics="true" pageId="4" pageNumber="5">vs</emphasis>
. 21.4±2.6% SL), and postcleithral process length (mean = 12.9±1.1%
<emphasis id="E0E82C7A7C633554A9FEED9FFA6F5E99" box="[1431,1452,228,251]" italics="true" pageId="4" pageNumber="5">vs</emphasis>
. 13.9±1.2% SL).
<taxonomicName id="159C8BEB7C633554AEF5EC73FC9D5F42" authorityName="Shibatta &amp; Souza-Shibatta" authorityYear="2023" box="[668,862,264,288]" family="Pseudopimelodidae" genus="Rhyacoglanis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Siluriformes" pageId="4" pageNumber="5" phylum="Chordata" rank="species" species="varii">
<emphasis id="E0E82C7A7C633554AEF5EC73FC9D5F42" box="[668,862,264,288]" italics="true" pageId="4" pageNumber="5">Rhyacoglanis varii</emphasis>
</taxonomicName>
differs from
<taxonomicName id="159C8BEB7C633554AF9AEC72FB9C5F42" baseAuthorityName="Boulenger" baseAuthorityYear="1887" box="[1011,1119,264,288]" family="Pseudopimelodidae" genus="Rhyacoglanis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Siluriformes" pageId="4" pageNumber="5" phylum="Chordata" rank="species" species="pulcher">
<emphasis id="E0E82C7A7C633554AF9AEC72FB9C5F42" box="[1011,1119,264,288]" italics="true" pageId="4" pageNumber="5">R. pulcher</emphasis>
</taxonomicName>
and
<taxonomicName id="159C8BEB7C633554A8F0EC72FADF5F42" authorityName="Shibatta &amp; Vari" authorityYear="2017" box="[1177,1308,265,288]" family="Pseudopimelodidae" genus="Rhyacoglanis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Siluriformes" pageId="4" pageNumber="5" phylum="Chordata" rank="species" species="seminiger">
<emphasis id="E0E82C7A7C633554A8F0EC72FADF5F42" box="[1177,1308,265,288]" italics="true" pageId="4" pageNumber="5">R. seminiger</emphasis>
</taxonomicName>
by the shape of the pectoral-fin spine, with anterior serrae distributed along the entire margin (
<emphasis id="E0E82C7A7C633554A9FEEC54FA6F5F24" box="[1431,1452,303,326]" italics="true" pageId="4" pageNumber="5">vs</emphasis>
. restricted to the proximal half).
</paragraph>
</subSubSection>
<caption id="86E3A0E07C633554AD81EB9AFC965979" ID-DOI="http://doi.org/10.5281/zenodo.11124760" ID-Zenodo-Dep="11124760" httpUri="https://zenodo.org/record/11124760/files/figure.png" pageId="4" pageNumber="5" startId="4.[488,561,1761,1781]" targetBox="[488,1459,432,1737]" targetPageId="4" targetType="figure">
<paragraph id="D223F0687C633554AD81EB9AFC965979" blockId="4.[488,1459,1761,1819]" pageId="4" pageNumber="5">
<emphasis id="E0E82C7A7C633554AD81EB9AFD965897" bold="true" box="[488,597,1761,1781]" pageId="4" pageNumber="5">FIGURE 1 |</emphasis>
Holotype of
<taxonomicName id="159C8BEB7C633554AEA6EB9AFC065897" authority=", MZUEL" authorityName="MZUEL" box="[719,965,1761,1781]" family="Pseudopimelodidae" genus="Rhyacoglanis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Siluriformes" pageId="4" pageNumber="5" phylum="Chordata" rank="species" species="varii">
<emphasis id="E0E82C7A7C633554AEA6EB9AFCB25897" bold="true" box="[719,881,1761,1781]" italics="true" pageId="4" pageNumber="5">Rhyacoglanis varii</emphasis>
, MZUEL
</taxonomicName>
22317, 67.1 mm SL, rio do Peixe, upper rio Tocantins basin, Goianésia, state of Goiás, Brazil.
</paragraph>
</caption>
<subSubSection id="9A86A3E37C62355AAE6FEDC5FC365CD9" lastPageId="10" lastPageNumber="11" pageId="5" pageNumber="6" type="description">
<paragraph id="D223F0687C623556AE6FEDC5FCD35C13" blockId="5.[488,1459,190,1372]" lastBlockId="6.[488,1459,190,625]" lastPageId="6" lastPageNumber="7" pageId="5" pageNumber="6">
<emphasis id="E0E82C7A7C623555AE6FEDC5FD5F5EB4" bold="true" box="[518,668,190,214]" pageId="5" pageNumber="6">Description.</emphasis>
Body proportions are given in
<tableCitation id="9F1EC5D37C623555A86BEDC5FB8E5EB4" box="[1026,1101,190,214]" captionStart="TABLE 1" captionStartId="7.[488,550,193,213]" captionTargetId="graphics-25@7.[488,1459,389,1328]" captionTargetPageId="7" captionText="TABLE 1 | Morphometry of Rhyacoglanis varii from upper rio Tocantins basin (n = 42). Statistics of minimum (Min), maximum (Max), mean, and standard deviation (SD) include holotype." httpUri="http://table.plazi.org/id/86E3A0E07C603557AD81EDBAFADD5E99" pageId="5" pageNumber="6" tableUuid="86E3A0E07C603557AD81EDBAFADD5E99">Tab. 1</tableCitation>
. Body depressed from snout tip to dorsal-fin origin; progressively compressed from that point to caudal-fin base. Dorsal profile of head and anterior body scarcely convex from snout tip to dorsal-fin origin, then nearly straight, scarcely descending to adipose-fin origin. Ventral head convex, then almost straight from pectoral-fin origin to posterior limit of anal-fin base. Caudal peduncle profile slightly concave along dorsal and ventral margins. Head depressed; slightly longer than wide; anterior margin convex (semicircular) in dorsal view. Head with weakly developed unculiferous tubercles scattered laterally and dorsally. Mouth terminal and wide; width more than one-half of head length (HL). Upper jaw slightly longer than, or same length as, lower jaw. Lips thick and well developed, more so proximate to rictus. Premaxillary tooth plate posterolaterally pointed. Anterior nostril immediately posterior to vertical through rictus. Eye small, superior, covered by skin, slightly posterior to anterior one-third of HL. Opercular membrane well developed; margin falling short of pectoral-fin origin. Maxillary barbel base enlarged. Tip of adpressed maxillary barbel falling short of opercular margin. Adpressed inner mental barbel tip surpassing outer mental barbel base but falling short of its tip. Adpressed outer mental barbel tip reaching opercular membrane margin. Dorsal-fin trapezoidal, distal margin rounded, and first branched ray longer than dorsal-fin base. Dorsal-fin origin immediately posterior to anterior one-third of body but anterior to one-half of SL. Adpressed dorsal-fin tip reaching slightly beyond midpoint between dorsal-fin base terminus and adipose-fin origin. First dorsal-fin ray (spinelet) small, rigid, forming dorsal-fin spine-locking mechanism. Second dorsal-fin ray forming strong, rigid, pointed spine; anterior margin smooth, posterior margin with retrorse serrations. Dorsal-fin rays II,6* (5 C&amp;S, 16 eth). Adipose fin long; base longer than anal-fin base; posterior margin free and rounded. Pectoral-fin margin somewhat triangular, posterior margin rounded. Adpressed pectoral fin tip falling short pelvic-fin origin. First pectoral-fin ray forming strong, rigid, pointed spine, serrated along anterior and posterior margins; posterior serrae retrorse, distinctly larger than anterior serrae (approximately twice as long); pectoral-fin spine notched distally (
<figureCitation id="4AA7ECED7C623555AFD2E9A8FC3D5A89" box="[955,1022,1235,1259]" captionStart="FIGURE 2" captionStartId="5.[136,209,1873,1893]" captionTargetBox="[147,1436,1497,1804]" captionTargetId="figure-405@5.[136,1459,1471,1821]" captionTargetPageId="5" captionText="FIGURE 2 | Rhyacoglanis varii pectoral-fin spine, paratype, MZUEL 6039, 55.5 mm SL. Scale bar = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.11124762" httpUri="https://zenodo.org/record/11124762/files/figure.png" pageId="5" pageNumber="6">Fig. 2</figureCitation>
). Pectoral-fin rays I,6* (5 C&amp;S, 15 eth), I,5 (1 eth). Pelvic-fin profile almost triangular; posterior margin rounded. Pelvic-fin origin immediately posterior to vertical through dorsal-fin base terminus. Adpressed pelvic fin tip reaching vertical through adipose fin origin. Pelvic-fin rays i,5* (5 C&amp;S, 16 eth). Anal-fin margin rounded; anal-fin rays iii,5 (1 eth), ii,6* (5 eth), iii,6 (9 eth), iv,6 (1 C&amp;S, 1 eth), v,5 (1 C&amp;S), or v,6 (3 C&amp;S). Caudal fin forked; lobes pointed; ventral lobe usually slightly longer than dorsal lobe. Caudal-fin principal rays i,6,5,i (1 eth), i,6,7,i (1 C&amp;S, 1 eth), i,6,8,i* (4 C&amp;S, 13 eth) or i,7,8,i (1 eth). Dorsal procurrent rays 17 (1 C&amp;S), 18 (1 C&amp;S), or 19 (3 C&amp;S); ventral procurrent rays 15 (3 C&amp;S), 16 (1 C&amp;S), or 17 (1 C&amp;S). Skeletal elements supporting the caudal fin as follow: parhypural, hypurals 1 and 2 fused; hypural 3 and 4 fused; hypural 5 free (
<figureCitation id="4AA7ECED7C613556A852ECE5FBBD5FD4" box="[1083,1150,414,438]" captionStart="FIGURE 3" captionStartId="6.[136,209,1686,1706]" captionTargetBox="[178,1437,733,1622]" captionTargetId="figure-21@6.[138,1459,733,1664]" captionTargetPageId="6" captionText="FIGURE 3 | Rhyacoglanis varii caudal-fin skeleton, paratype, MZUEL 6039, 53.5 mm SL. Bones in yellow ocher and cartilage in blue. H1 to H5 = hypurals 1 to 5. Scale bar = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.11124764" httpUri="https://zenodo.org/record/11124764/files/figure.png" pageId="6" pageNumber="7">Fig. 3</figureCitation>
). Posterior cleithral process well developed, pointed, tip reaching vertical through anterior dorsal-fin base. Axillary pore present. Lateral line complete. Total vertebrae 33 (2 x-r), 35 (3 C&amp;S), 36 (2 C&amp;S). Ribs 8 (1 X-r), 9 (3 C&amp;S, 1 X-r), 10 (1 C&amp;S), or 11 (1 C&amp;S). Gill rakers 1,0,5* (1 C&amp;S, 5 eth), 1,0,6 (2 C&amp;S, 2 eth), 1,0,7 (6 eth), 1,0,8 (2 eth), 1,1,5 (2 C&amp;S). Branchiostegal rays 7 (1 C&amp;S), 8 (4 C&amp;S).
</paragraph>
<caption id="86E3A0E07C623555ACE1EA2AFB9A5907" ID-DOI="http://doi.org/10.5281/zenodo.11124762" ID-Zenodo-Dep="11124762" box="[136,1113,1873,1893]" httpUri="https://zenodo.org/record/11124762/files/figure.png" pageId="5" pageNumber="6" startId="5.[136,209,1873,1893]" targetBox="[147,1436,1497,1804]" targetPageId="5" targetType="figure">
<paragraph id="D223F0687C623555ACE1EA2AFB9A5907" blockId="5.[136,1113,1873,1893]" box="[136,1113,1873,1893]" pageId="5" pageNumber="6">
<emphasis id="E0E82C7A7C623555ACE1EA2AFE595907" bold="true" box="[136,410,1873,1893]" pageId="5" pageNumber="6">
FIGURE 2 |
<taxonomicName id="159C8BEB7C623555AC91EA2AFE595907" box="[248,410,1873,1893]" family="Pseudopimelodidae" genus="Rhyacoglanis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Siluriformes" pageId="5" pageNumber="6" phylum="Chordata" rank="species" species="varii">
<emphasis id="E0E82C7A7C623555AC91EA2AFE595907" bold="true" box="[248,410,1873,1893]" italics="true" pageId="5" pageNumber="6">Rhyacoglanis varii</emphasis>
</taxonomicName>
</emphasis>
pectoral-fin spine, paratype, MZUEL 6039, 55.5 mm SL. Scale bar = 1.0 mm.
</paragraph>
</caption>
<caption id="86E3A0E07C613556ACE1EBEDFE0E58B2" ID-DOI="http://doi.org/10.5281/zenodo.11124764" ID-Zenodo-Dep="11124764" httpUri="https://zenodo.org/record/11124764/files/figure.png" pageId="6" pageNumber="7" startId="6.[136,209,1686,1706]" targetBox="[178,1437,733,1622]" targetPageId="6" targetType="figure">
<paragraph id="D223F0687C613556ACE1EBEDFE0E58B2" blockId="6.[136,1459,1686,1744]" pageId="6" pageNumber="7">
<emphasis id="E0E82C7A7C613556ACE1EBEDFE5958C9" bold="true" box="[136,410,1686,1707]" pageId="6" pageNumber="7">
FIGURE 3 |
<taxonomicName id="159C8BEB7C613556AC91EBECFE5958C9" box="[248,410,1687,1707]" family="Pseudopimelodidae" genus="Rhyacoglanis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Siluriformes" pageId="6" pageNumber="7" phylum="Chordata" rank="species" species="varii">
<emphasis id="E0E82C7A7C613556AC91EBECFE5958C9" bold="true" box="[248,410,1687,1707]" italics="true" pageId="6" pageNumber="7">Rhyacoglanis varii</emphasis>
</taxonomicName>
</emphasis>
caudal-fin skeleton, paratype, MZUEL 6039, 53.5 mm SL. Bones in yellow ocher and cartilage in blue. H1 to H5 = hypurals 1 to 5. Scale bar = 1.0 mm.
</paragraph>
</caption>
<caption id="86E3A0E07C603557AD81EDBAFADD5E99" ID-Table-UUID="86E3A0E07C603557AD81EDBAFADD5E99" httpUri="http://table.plazi.org/id/86E3A0E07C603557AD81EDBAFADD5E99" pageId="7" pageNumber="8" startId="7.[488,550,193,213]" targetBox="[502,1412,308,1718]" targetIsTable="true" targetPageId="7" targetType="table">
<paragraph id="D223F0687C603557AD81EDBAFADD5E99" blockId="7.[488,1459,193,251]" pageId="7" pageNumber="8">
<emphasis id="E0E82C7A7C603557AD81EDBAFD935EB7" bold="true" box="[488,592,193,213]" pageId="7" pageNumber="8">TABLE 1 |</emphasis>
Morphometry of
<taxonomicName id="159C8BEB7C603557AF69EDBAFC665EB7" box="[768,933,193,213]" family="Pseudopimelodidae" genus="Rhyacoglanis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Siluriformes" pageId="7" pageNumber="8" phylum="Chordata" rank="species" species="varii">
<emphasis id="E0E82C7A7C603557AF69EDBAFC665EB7" bold="true" box="[768,933,193,213]" italics="true" pageId="7" pageNumber="8">Rhyacoglanis varii</emphasis>
</taxonomicName>
from upper rio Tocantins basin (n = 42). Statistics of minimum (Min), maximum (Max), mean, and standard deviation (SD) include holotype.
</paragraph>
</caption>
<paragraph id="D223F0687C603557AFB2EC4EFABE58D4" pageId="7" pageNumber="8">
<table id="A09C02C87C60CAAFAD9FEC4FFA4758D4" box="[502,1412,308,1718]" gridcols="4" gridrows="32" pageId="7" pageNumber="8">
<tr id="6CACF22A7C60CAAFAD9FEC4FFA475F24" box="[502,1412,308,326]" gridrow="0" pageId="7" pageNumber="8" rowspan-0="1">
<th id="2F7D9B567C60CAAFAFB2EC4FFBF25F24" box="[987,1073,308,326]" gridcol="1" gridrow="0" pageId="7" pageNumber="8">
<emphasis id="E0E82C7A7C603557AFB2EC4EFBF25F27" bold="true" box="[987,1073,309,325]" pageId="7" pageNumber="8">Holotype</emphasis>
</th>
<th id="2F7D9B567C60CAAFA8EBEC4FFB155F24" box="[1154,1238,308,326]" gridcol="2" gridrow="0" pageId="7" pageNumber="8">
<emphasis id="E0E82C7A7C603557A8EBEC4FFB155F24" bold="true" box="[1154,1238,308,326]" pageId="7" pageNumber="8">Min-Max</emphasis>
</th>
<th id="2F7D9B567C60CAAFA945EC4FFA475F24" box="[1324,1412,308,326]" gridcol="3" gridrow="0" pageId="7" pageNumber="8">
<emphasis id="E0E82C7A7C603557A945EC4FFA475F24" bold="true" box="[1324,1412,308,326]" pageId="7" pageNumber="8">Mean±SD</emphasis>
</th>
</tr>
<tr id="6CACF22A7C60CAAFAD9FEC11FA475F1E" box="[502,1412,362,380]" gridrow="1" pageId="7" pageNumber="8">
<th id="2F7D9B567C60CAAFAD9FEC11FC865F1E" box="[502,837,362,380]" gridcol="0" gridrow="1" pageId="7" pageNumber="8">Standard length (mm)</th>
<td id="2F7D9B567C60CAAFAFB2EC11FBF25F1E" box="[987,1073,362,380]" gridcol="1" gridrow="1" pageId="7" pageNumber="8">67.1</td>
<td id="2F7D9B567C60CAAFA8EBEC11FB155F1E" box="[1154,1238,362,380]" gridcol="2" gridrow="1" pageId="7" pageNumber="8">29.669.1</td>
<td id="2F7D9B567C60CAAFA945EC11FA475F1E" box="[1324,1412,362,380]" gridcol="3" gridrow="1" pageId="7" pageNumber="8">44.3±11.1</td>
</tr>
<tr id="6CACF22A7C60CAAFAD9FECEDFA475FCA" box="[502,1412,406,424]" gridrow="2" pageId="7" pageNumber="8">
<th id="2F7D9B567C60CAAFAD9FECEDFA475FCA" box="[502,1412,406,424]" colspan="4" colspanRight="3" gridcol="0" gridrow="2" pageId="7" pageNumber="8">
<emphasis id="E0E82C7A7C603557AD9EECECFD305FC5" bold="true" box="[503,755,406,424]" pageId="7" pageNumber="8">Percent of standard length</emphasis>
</th>
</tr>
<tr id="6CACF22A7C60CAAFAD9FECB8FA475FB7" box="[502,1412,451,469]" gridrow="3" pageId="7" pageNumber="8">
<th id="2F7D9B567C60CAAFAD9FECB8FC865FB7" box="[502,837,451,469]" gridcol="0" gridrow="3" pageId="7" pageNumber="8">Head length</th>
<td id="2F7D9B567C60CAAFAFB2ECB8FBF25FB7" box="[987,1073,451,469]" gridcol="1" gridrow="3" pageId="7" pageNumber="8">30.6</td>
<td id="2F7D9B567C60CAAFA8EBECB8FB155FB7" box="[1154,1238,451,469]" gridcol="2" gridrow="3" pageId="7" pageNumber="8">27.030.6</td>
<td id="2F7D9B567C60CAAFA945ECB8FA475FB7" box="[1324,1412,451,469]" gridcol="3" gridrow="3" pageId="7" pageNumber="8">28.8±1.0</td>
</tr>
<tr id="6CACF22A7C60CAAFAD9FEC8BFA475C60" box="[502,1412,496,514]" gridrow="4" pageId="7" pageNumber="8">
<th id="2F7D9B567C60CAAFAD9FEC8BFC865C60" box="[502,837,496,514]" gridcol="0" gridrow="4" pageId="7" pageNumber="8">Pectoral girdle width</th>
<td id="2F7D9B567C60CAAFAFB2EC8BFBF25C60" box="[987,1073,496,514]" gridcol="1" gridrow="4" pageId="7" pageNumber="8">27.6</td>
<td id="2F7D9B567C60CAAFA8EBEC8BFB155C60" box="[1154,1238,496,514]" gridcol="2" gridrow="4" pageId="7" pageNumber="8">23.128.4</td>
<td id="2F7D9B567C60CAAFA945EC8BFA475C60" box="[1324,1412,496,514]" gridcol="3" gridrow="4" pageId="7" pageNumber="8">26.5±1.3</td>
</tr>
<tr id="6CACF22A7C60CAAFAD9FEF67FA475C4C" box="[502,1412,540,558]" gridrow="5" pageId="7" pageNumber="8">
<th id="2F7D9B567C60CAAFAD9FEF67FC865C4C" box="[502,837,540,558]" gridcol="0" gridrow="5" pageId="7" pageNumber="8">Predorsal distance</th>
<td id="2F7D9B567C60CAAFAFB2EF67FBF25C4C" box="[987,1073,540,558]" gridcol="1" gridrow="5" pageId="7" pageNumber="8">37.3</td>
<td id="2F7D9B567C60CAAFA8EBEF67FB155C4C" box="[1154,1238,540,558]" gridcol="2" gridrow="5" pageId="7" pageNumber="8">35.137.5</td>
<td id="2F7D9B567C60CAAFA945EF67FA475C4C" box="[1324,1412,540,558]" gridcol="3" gridrow="5" pageId="7" pageNumber="8">36.4±0.8</td>
</tr>
<tr id="6CACF22A7C60CAAFAD9FEF32FA475C39" box="[502,1412,585,603]" gridrow="6" pageId="7" pageNumber="8">
<th id="2F7D9B567C60CAAFAD9FEF32FC865C39" box="[502,837,585,603]" gridcol="0" gridrow="6" pageId="7" pageNumber="8">Dorsal-fin base length</th>
<td id="2F7D9B567C60CAAFAFB2EF32FBF25C39" box="[987,1073,585,603]" gridcol="1" gridrow="6" pageId="7" pageNumber="8">17.2</td>
<td id="2F7D9B567C60CAAFA8EBEF32FB155C39" box="[1154,1238,585,603]" gridcol="2" gridrow="6" pageId="7" pageNumber="8">15.017.2</td>
<td id="2F7D9B567C60CAAFA945EF32FA475C39" box="[1324,1412,585,603]" gridcol="3" gridrow="6" pageId="7" pageNumber="8">16.3±0.6</td>
</tr>
<tr id="6CACF22A7C60CAAFAD9FEF0EFA475CE5" box="[502,1412,629,647]" gridrow="7" pageId="7" pageNumber="8">
<th id="2F7D9B567C60CAAFAD9FEF0EFC865CE5" box="[502,837,629,647]" gridcol="0" gridrow="7" pageId="7" pageNumber="8">Adipose-fin base length</th>
<td id="2F7D9B567C60CAAFAFB2EF0EFBF25CE5" box="[987,1073,629,647]" gridcol="1" gridrow="7" pageId="7" pageNumber="8">18.4</td>
<td id="2F7D9B567C60CAAFA8EBEF0EFB155CE5" box="[1154,1238,629,647]" gridcol="2" gridrow="7" pageId="7" pageNumber="8">16.221.5</td>
<td id="2F7D9B567C60CAAFA945EF0EFA475CE5" box="[1324,1412,629,647]" gridcol="3" gridrow="7" pageId="7" pageNumber="8">18.3±1.5</td>
</tr>
<tr id="6CACF22A7C60CAAFAD9FEFD9FA475CD6" box="[502,1412,674,692]" gridrow="8" pageId="7" pageNumber="8">
<th id="2F7D9B567C60CAAFAD9FEFD9FC865CD6" box="[502,837,674,692]" gridcol="0" gridrow="8" pageId="7" pageNumber="8">Prepelvic length</th>
<td id="2F7D9B567C60CAAFAFB2EFD9FBF25CD6" box="[987,1073,674,692]" gridcol="1" gridrow="8" pageId="7" pageNumber="8">50.5</td>
<td id="2F7D9B567C60CAAFA8EBEFD9FB155CD6" box="[1154,1238,674,692]" gridcol="2" gridrow="8" pageId="7" pageNumber="8">46.851.7</td>
<td id="2F7D9B567C60CAAFA945EFD9FA475CD6" box="[1324,1412,674,692]" gridcol="3" gridrow="8" pageId="7" pageNumber="8">49.5±1.6</td>
</tr>
<tr id="6CACF22A7C60CAAFAD9FEFB4FA475C83" box="[502,1412,719,737]" gridrow="9" pageId="7" pageNumber="8">
<th id="2F7D9B567C60CAAFAD9FEFB4FC865C83" box="[502,837,719,737]" gridcol="0" gridrow="9" pageId="7" pageNumber="8">Pelvic fin to anal fin distance</th>
<td id="2F7D9B567C60CAAFAFB2EFB4FBF25C83" box="[987,1073,719,737]" gridcol="1" gridrow="9" pageId="7" pageNumber="8">24.7</td>
<td id="2F7D9B567C60CAAFA8EBEFB4FB155C83" box="[1154,1238,719,737]" gridcol="2" gridrow="9" pageId="7" pageNumber="8">22.525.7</td>
<td id="2F7D9B567C60CAAFA945EFB4FA475C83" box="[1324,1412,719,737]" gridcol="3" gridrow="9" pageId="7" pageNumber="8">24.1±0.9</td>
</tr>
<tr id="6CACF22A7C60CAAFAD9FEF80FA475D6F" box="[502,1412,763,781]" gridrow="10" pageId="7" pageNumber="8">
<th id="2F7D9B567C60CAAFAD9FEF80FC865D6F" box="[502,837,763,781]" gridcol="0" gridrow="10" pageId="7" pageNumber="8">Anal-fin base length</th>
<td id="2F7D9B567C60CAAFAFB2EF80FBF25D6F" box="[987,1073,763,781]" gridcol="1" gridrow="10" pageId="7" pageNumber="8">10.3</td>
<td id="2F7D9B567C60CAAFA8EBEF80FB155D6F" box="[1154,1238,763,781]" gridcol="2" gridrow="10" pageId="7" pageNumber="8">9.213.1</td>
<td id="2F7D9B567C60CAAFA945EF80FA475D6F" box="[1324,1412,763,781]" gridcol="3" gridrow="10" pageId="7" pageNumber="8">10.5±0.9</td>
</tr>
<tr id="6CACF22A7C60CAAFAD9FEE53FA475D58" box="[502,1412,808,826]" gridrow="11" pageId="7" pageNumber="8">
<th id="2F7D9B567C60CAAFAD9FEE53FC865D58" box="[502,837,808,826]" gridcol="0" gridrow="11" pageId="7" pageNumber="8">Caudal peduncle length</th>
<td id="2F7D9B567C60CAAFAFB2EE53FBF25D58" box="[987,1073,808,826]" gridcol="1" gridrow="11" pageId="7" pageNumber="8">15.1</td>
<td id="2F7D9B567C60CAAFA8EBEE53FB155D58" box="[1154,1238,808,826]" gridcol="2" gridrow="11" pageId="7" pageNumber="8">15.119.9</td>
<td id="2F7D9B567C60CAAFA945EE53FA475D58" box="[1324,1412,808,826]" gridcol="3" gridrow="11" pageId="7" pageNumber="8">17.7±1.3</td>
</tr>
<tr id="6CACF22A7C60CAAFAD9FEE2EFA475D05" box="[502,1412,853,871]" gridrow="12" pageId="7" pageNumber="8">
<th id="2F7D9B567C60CAAFAD9FEE2EFC865D05" box="[502,837,853,871]" gridcol="0" gridrow="12" pageId="7" pageNumber="8">Body depth</th>
<td id="2F7D9B567C60CAAFAFB2EE2EFBF25D05" box="[987,1073,853,871]" gridcol="1" gridrow="12" pageId="7" pageNumber="8">20.7</td>
<td id="2F7D9B567C60CAAFA8EBEE2EFB155D05" box="[1154,1238,853,871]" gridcol="2" gridrow="12" pageId="7" pageNumber="8">16.121.2</td>
<td id="2F7D9B567C60CAAFA945EE2EFA475D05" box="[1324,1412,853,871]" gridcol="3" gridrow="12" pageId="7" pageNumber="8">18.2±1.5</td>
</tr>
<tr id="6CACF22A7C60CAAFAD9FEEFAFA475DF1" box="[502,1412,897,915]" gridrow="13" pageId="7" pageNumber="8">
<th id="2F7D9B567C60CAAFAD9FEEFAFC865DF1" box="[502,837,897,915]" gridcol="0" gridrow="13" pageId="7" pageNumber="8">Caudal peduncle depth</th>
<td id="2F7D9B567C60CAAFAFB2EEFAFBF25DF1" box="[987,1073,897,915]" gridcol="1" gridrow="13" pageId="7" pageNumber="8">9.4</td>
<td id="2F7D9B567C60CAAFA8EBEEFAFB155DF1" box="[1154,1238,897,915]" gridcol="2" gridrow="13" pageId="7" pageNumber="8">8.510.5</td>
<td id="2F7D9B567C60CAAFA945EEFAFA475DF1" box="[1324,1412,897,915]" gridcol="3" gridrow="13" pageId="7" pageNumber="8">9.5±0.5</td>
</tr>
<tr id="6CACF22A7C60CAAFAD9FEED5FA475DA2" box="[502,1412,942,960]" gridrow="14" pageId="7" pageNumber="8">
<th id="2F7D9B567C60CAAFAD9FEED5FC865DA2" box="[502,837,942,960]" gridcol="0" gridrow="14" pageId="7" pageNumber="8">Pectoral-fin spine length</th>
<td id="2F7D9B567C60CAAFAFB2EED5FBF25DA2" box="[987,1073,942,960]" gridcol="1" gridrow="14" pageId="7" pageNumber="8">17.5</td>
<td id="2F7D9B567C60CAAFA8EBEED5FB155DA2" box="[1154,1238,942,960]" gridcol="2" gridrow="14" pageId="7" pageNumber="8">16.321.5</td>
<td id="2F7D9B567C60CAAFA945EED5FA475DA2" box="[1324,1412,942,960]" gridcol="3" gridrow="14" pageId="7" pageNumber="8">18.6±1.5</td>
</tr>
<tr id="6CACF22A7C60CAAFAD9FEEA1FA475D8E" box="[502,1412,986,1004]" gridrow="15" pageId="7" pageNumber="8">
<th id="2F7D9B567C60CAAFAD9FEEA1FC865D8E" box="[502,837,986,1004]" gridcol="0" gridrow="15" pageId="7" pageNumber="8">Dorsal-fin spine length</th>
<td id="2F7D9B567C60CAAFAFB2EEA1FBF25D8E" box="[987,1073,986,1004]" gridcol="1" gridrow="15" pageId="7" pageNumber="8">15.3</td>
<td id="2F7D9B567C60CAAFA8EBEEA1FB155D8E" box="[1154,1238,986,1004]" gridcol="2" gridrow="15" pageId="7" pageNumber="8">14.119.7</td>
<td id="2F7D9B567C60CAAFA945EEA1FA475D8E" box="[1324,1412,986,1004]" gridcol="3" gridrow="15" pageId="7" pageNumber="8">17.7±1.6</td>
</tr>
<tr id="6CACF22A7C60CAAFAD9FE97CFA475A7B" box="[502,1412,1031,1049]" gridrow="16" pageId="7" pageNumber="8">
<th id="2F7D9B567C60CAAFAD9FE97CFC865A7B" box="[502,837,1031,1049]" gridcol="0" gridrow="16" pageId="7" pageNumber="8">Pelvic fin length</th>
<td id="2F7D9B567C60CAAFAFB2E97CFBF25A7B" box="[987,1073,1031,1049]" gridcol="1" gridrow="16" pageId="7" pageNumber="8">19.6</td>
<td id="2F7D9B567C60CAAFA8EBE97CFB155A7B" box="[1154,1238,1031,1049]" gridcol="2" gridrow="16" pageId="7" pageNumber="8">17.419.8</td>
<td id="2F7D9B567C60CAAFA945E97CFA475A7B" box="[1324,1412,1031,1049]" gridcol="3" gridrow="16" pageId="7" pageNumber="8">18.8±0.7</td>
</tr>
<tr id="6CACF22A7C60CAAFAD9FE94FFA475A24" box="[502,1412,1076,1094]" gridrow="17" pageId="7" pageNumber="8">
<th id="2F7D9B567C60CAAFAD9FE94FFC865A24" box="[502,837,1076,1094]" gridcol="0" gridrow="17" pageId="7" pageNumber="8">Postcleithral process length</th>
<td id="2F7D9B567C60CAAFAFB2E94FFBF25A24" box="[987,1073,1076,1094]" gridcol="1" gridrow="17" pageId="7" pageNumber="8">12.6</td>
<td id="2F7D9B567C60CAAFA8EBE94FFB155A24" box="[1154,1238,1076,1094]" gridcol="2" gridrow="17" pageId="7" pageNumber="8">10.114.7</td>
<td id="2F7D9B567C60CAAFA945E94FFA475A24" box="[1324,1412,1076,1094]" gridcol="3" gridrow="17" pageId="7" pageNumber="8">12.9±1.1</td>
</tr>
<tr id="6CACF22A7C60CAAFAD9FE91BFA475A10" box="[502,1412,1120,1138]" gridrow="18" pageId="7" pageNumber="8">
<th id="2F7D9B567C60CAAFAD9FE91BFC865A10" box="[502,837,1120,1138]" gridcol="0" gridrow="18" pageId="7" pageNumber="8">Dorsal fin to pelvic fin distance</th>
<td id="2F7D9B567C60CAAFAFB2E91BFBF25A10" box="[987,1073,1120,1138]" gridcol="1" gridrow="18" pageId="7" pageNumber="8">26.2</td>
<td id="2F7D9B567C60CAAFA8EBE91BFB155A10" box="[1154,1238,1120,1138]" gridcol="2" gridrow="18" pageId="7" pageNumber="8">22.426.2</td>
<td id="2F7D9B567C60CAAFA945E91BFA475A10" box="[1324,1412,1120,1138]" gridcol="3" gridrow="18" pageId="7" pageNumber="8">23.8±1.2</td>
</tr>
<tr id="6CACF22A7C60CAAFAD9FE9F6FA475AFD" box="[502,1412,1165,1183]" gridrow="19" pageId="7" pageNumber="8">
<th id="2F7D9B567C60CAAFAD9FE9F6FC865AFD" box="[502,837,1165,1183]" gridcol="0" gridrow="19" pageId="7" pageNumber="8">Pelvic fins distance</th>
<td id="2F7D9B567C60CAAFAFB2E9F6FBF25AFD" box="[987,1073,1165,1183]" gridcol="1" gridrow="19" pageId="7" pageNumber="8">13.7</td>
<td id="2F7D9B567C60CAAFA8EBE9F6FB155AFD" box="[1154,1238,1165,1183]" gridcol="2" gridrow="19" pageId="7" pageNumber="8">10.913.7</td>
<td id="2F7D9B567C60CAAFA945E9F6FA475AFD" box="[1324,1412,1165,1183]" gridcol="3" gridrow="19" pageId="7" pageNumber="8">12.3±0.7</td>
</tr>
<tr id="6CACF22A7C60CAAFAD9FE9C2FA475AA9" box="[502,1412,1209,1227]" gridrow="20" pageId="7" pageNumber="8">
<th id="2F7D9B567C60CAAFAD9FE9C2FC865AA9" box="[502,837,1209,1227]" gridcol="0" gridrow="20" pageId="7" pageNumber="8">Pelvic fin to anus distance</th>
<td id="2F7D9B567C60CAAFAFB2E9C2FBF25AA9" box="[987,1073,1209,1227]" gridcol="1" gridrow="20" pageId="7" pageNumber="8">14.5</td>
<td id="2F7D9B567C60CAAFA8EBE9C2FB155AA9" box="[1154,1238,1209,1227]" gridcol="2" gridrow="20" pageId="7" pageNumber="8">11.914.5</td>
<td id="2F7D9B567C60CAAFA945E9C2FA475AA9" box="[1324,1412,1209,1227]" gridcol="3" gridrow="20" pageId="7" pageNumber="8">13.1±0.8</td>
</tr>
<tr id="6CACF22A7C60CAAFAD9FE99DFA475A9A" box="[502,1412,1254,1272]" gridrow="21" pageId="7" pageNumber="8">
<th id="2F7D9B567C60CAAFAD9FE99DFC865A9A" box="[502,837,1254,1272]" gridcol="0" gridrow="21" pageId="7" pageNumber="8">Anus to anal fin distance</th>
<td id="2F7D9B567C60CAAFAFB2E99DFBF25A9A" box="[987,1073,1254,1272]" gridcol="1" gridrow="21" pageId="7" pageNumber="8">11.9</td>
<td id="2F7D9B567C60CAAFA8EBE99DFB155A9A" box="[1154,1238,1254,1272]" gridcol="2" gridrow="21" pageId="7" pageNumber="8">9.812.5</td>
<td id="2F7D9B567C60CAAFA945E99DFA475A9A" box="[1324,1412,1254,1272]" gridcol="3" gridrow="21" pageId="7" pageNumber="8">11.4±0.8</td>
</tr>
<tr id="6CACF22A7C60CAAFAD9FE868FA475B47" box="[502,1412,1299,1317]" gridrow="22" pageId="7" pageNumber="8">
<th id="2F7D9B567C60CAAFAD9FE868FA475B47" box="[502,1412,1299,1317]" colspan="4" colspanRight="3" gridcol="0" gridrow="22" pageId="7" pageNumber="8">
<emphasis id="E0E82C7A7C603557AD9FE868FD365B41" bold="true" box="[502,757,1299,1317]" pageId="7" pageNumber="8">Percentages of head length</emphasis>
</th>
</tr>
<tr id="6CACF22A7C60CAAFAD9FE844FA475B33" box="[502,1412,1343,1361]" gridrow="23" pageId="7" pageNumber="8">
<th id="2F7D9B567C60CAAFAD9FE844FC865B33" box="[502,837,1343,1361]" gridcol="0" gridrow="23" pageId="7" pageNumber="8">Eye diameter</th>
<td id="2F7D9B567C60CAAFAFB2E844FBF25B33" box="[987,1073,1343,1361]" gridcol="1" gridrow="23" pageId="7" pageNumber="8">9.4</td>
<td id="2F7D9B567C60CAAFA8EBE844FB155B33" box="[1154,1238,1343,1361]" gridcol="2" gridrow="23" pageId="7" pageNumber="8">8.911.8</td>
<td id="2F7D9B567C60CAAFA945E844FA475B33" box="[1324,1412,1343,1361]" gridcol="3" gridrow="23" pageId="7" pageNumber="8">10.4±0.8</td>
</tr>
<tr id="6CACF22A7C60CAAFAD9FE817FA475B1C" box="[502,1412,1388,1406]" gridrow="24" pageId="7" pageNumber="8">
<th id="2F7D9B567C60CAAFAD9FE817FC865B1C" box="[502,837,1388,1406]" gridcol="0" gridrow="24" pageId="7" pageNumber="8">Interorbital distance</th>
<td id="2F7D9B567C60CAAFAFB2E817FBF25B1C" box="[987,1073,1388,1406]" gridcol="1" gridrow="24" pageId="7" pageNumber="8">31.1</td>
<td id="2F7D9B567C60CAAFA8EBE817FB155B1C" box="[1154,1238,1388,1406]" gridcol="2" gridrow="24" pageId="7" pageNumber="8">26.334.5</td>
<td id="2F7D9B567C60CAAFA945E817FA475B1C" box="[1324,1412,1388,1406]" gridcol="3" gridrow="24" pageId="7" pageNumber="8">31.7±1.9</td>
</tr>
<tr id="6CACF22A7C60CAAFAD9FE8E2FA475BC9" box="[502,1412,1433,1451]" gridrow="25" pageId="7" pageNumber="8">
<th id="2F7D9B567C60CAAFAD9FE8E2FC865BC9" box="[502,837,1433,1451]" gridcol="0" gridrow="25" pageId="7" pageNumber="8">Snout length</th>
<td id="2F7D9B567C60CAAFAFB2E8E2FBF25BC9" box="[987,1073,1433,1451]" gridcol="1" gridrow="25" pageId="7" pageNumber="8">39.5</td>
<td id="2F7D9B567C60CAAFA8EBE8E2FB155BC9" box="[1154,1238,1433,1451]" gridcol="2" gridrow="25" pageId="7" pageNumber="8">34.239.8</td>
<td id="2F7D9B567C60CAAFA945E8E2FA475BC9" box="[1324,1412,1433,1451]" gridcol="3" gridrow="25" pageId="7" pageNumber="8">37.0±1.6</td>
</tr>
<tr id="6CACF22A7C60CAAFAD9FE8BEFA475BB5" box="[502,1412,1477,1495]" gridrow="26" pageId="7" pageNumber="8">
<th id="2F7D9B567C60CAAFAD9FE8BEFC865BB5" box="[502,837,1477,1495]" gridcol="0" gridrow="26" pageId="7" pageNumber="8">Mouth width</th>
<td id="2F7D9B567C60CAAFAFB2E8BEFBF25BB5" box="[987,1073,1477,1495]" gridcol="1" gridrow="26" pageId="7" pageNumber="8">57.5</td>
<td id="2F7D9B567C60CAAFA8EBE8BEFB155BB5" box="[1154,1238,1477,1495]" gridcol="2" gridrow="26" pageId="7" pageNumber="8">45.960.2</td>
<td id="2F7D9B567C60CAAFA945E8BEFA475BB5" box="[1324,1412,1477,1495]" gridcol="3" gridrow="26" pageId="7" pageNumber="8">52.5±4.8</td>
</tr>
<tr id="6CACF22A7C60CAAFAD9FE889FA475866" box="[502,1412,1522,1540]" gridrow="27" pageId="7" pageNumber="8">
<th id="2F7D9B567C60CAAFAD9FE889FC865866" box="[502,837,1522,1540]" gridcol="0" gridrow="27" pageId="7" pageNumber="8">Head depth</th>
<td id="2F7D9B567C60CAAFAFB2E889FBF25866" box="[987,1073,1522,1540]" gridcol="1" gridrow="27" pageId="7" pageNumber="8">39.4</td>
<td id="2F7D9B567C60CAAFA8EBE889FB155866" box="[1154,1238,1522,1540]" gridcol="2" gridrow="27" pageId="7" pageNumber="8">35.050.9</td>
<td id="2F7D9B567C60CAAFA945E889FA475866" box="[1324,1412,1522,1540]" gridcol="3" gridrow="27" pageId="7" pageNumber="8">41.0±4.5</td>
</tr>
<tr id="6CACF22A7C60CAAFAD9FEB65FA475852" box="[502,1412,1566,1584]" gridrow="28" pageId="7" pageNumber="8">
<th id="2F7D9B567C60CAAFAD9FEB65FC865852" box="[502,837,1566,1584]" gridcol="0" gridrow="28" pageId="7" pageNumber="8">Maxillary barbel length</th>
<td id="2F7D9B567C60CAAFAFB2EB65FBF25852" box="[987,1073,1566,1584]" gridcol="1" gridrow="28" pageId="7" pageNumber="8">75.7</td>
<td id="2F7D9B567C60CAAFA8EBEB65FB155852" box="[1154,1238,1566,1584]" gridcol="2" gridrow="28" pageId="7" pageNumber="8">72.092.9</td>
<td id="2F7D9B567C60CAAFA945EB65FA475852" box="[1324,1412,1566,1584]" gridcol="3" gridrow="28" pageId="7" pageNumber="8">79.6±5.8</td>
</tr>
<tr id="6CACF22A7C60CAAFAD9FEB30FA47583F" box="[502,1412,1611,1629]" gridrow="29" pageId="7" pageNumber="8">
<th id="2F7D9B567C60CAAFAD9FEB30FC86583F" box="[502,837,1611,1629]" gridcol="0" gridrow="29" pageId="7" pageNumber="8">Anterior to posterior nostrils distance</th>
<td id="2F7D9B567C60CAAFAFB2EB30FBF2583F" box="[987,1073,1611,1629]" gridcol="1" gridrow="29" pageId="7" pageNumber="8">14.3</td>
<td id="2F7D9B567C60CAAFA8EBEB30FB15583F" box="[1154,1238,1611,1629]" gridcol="2" gridrow="29" pageId="7" pageNumber="8">14.319.8</td>
<td id="2F7D9B567C60CAAFA945EB30FA47583F" box="[1324,1412,1611,1629]" gridcol="3" gridrow="29" pageId="7" pageNumber="8">17.0±1.6</td>
</tr>
<tr id="6CACF22A7C60CAAFAD9FEB03FA4758E8" box="[502,1412,1656,1674]" gridrow="30" pageId="7" pageNumber="8">
<th id="2F7D9B567C60CAAFAD9FEB03FC8658E8" box="[502,837,1656,1674]" gridcol="0" gridrow="30" pageId="7" pageNumber="8">Posterior nostril to eye distance</th>
<td id="2F7D9B567C60CAAFAFB2EB03FBF258E8" box="[987,1073,1656,1674]" gridcol="1" gridrow="30" pageId="7" pageNumber="8">5.5</td>
<td id="2F7D9B567C60CAAFA8EBEB03FB1558E8" box="[1154,1238,1656,1674]" gridcol="2" gridrow="30" pageId="7" pageNumber="8">3.46.2</td>
<td id="2F7D9B567C60CAAFA945EB03FA4758E8" box="[1324,1412,1656,1674]" gridcol="3" gridrow="30" pageId="7" pageNumber="8">4.8±0.9</td>
</tr>
<tr id="6CACF22A7C60CAAFAD9FEBDFFA4758D4" box="[502,1412,1700,1718]" gridrow="31" pageId="7" pageNumber="8">
<th id="2F7D9B567C60CAAFAD9FEBDFFC8658D4" box="[502,837,1700,1718]" gridcol="0" gridrow="31" pageId="7" pageNumber="8">Posterior nostrils distance</th>
<td id="2F7D9B567C60CAAFAFB2EBDFFBF258D4" box="[987,1073,1700,1718]" gridcol="1" gridrow="31" pageId="7" pageNumber="8">20.9</td>
<td id="2F7D9B567C60CAAFA8EBEBDFFB1558D4" box="[1154,1238,1700,1718]" gridcol="2" gridrow="31" pageId="7" pageNumber="8">18.723.6</td>
<td id="2F7D9B567C60CAAFA945EBDFFA4758D4" box="[1324,1412,1700,1718]" gridcol="3" gridrow="31" pageId="7" pageNumber="8">20.4±1.2</td>
</tr>
</table>
</paragraph>
<paragraph id="D223F0687C6F3558AE6FEDC5FA825FB9" blockId="8.[488,1459,190,475]" pageId="8" pageNumber="9">
<emphasis id="E0E82C7A7C6F3558AE6FEDC5FC025EB4" bold="true" box="[518,961,190,214]" pageId="8" pageNumber="9">Head laterosensory canal and pores.</emphasis>
Supraorbital(s) canal begins anteromedially to anterior nostril just before vertical through nostril, with five branches (s1, s2, s3, s6 (epiphyseal branch), and s8 (parietal branch)). Infraorbital canal with six branches (i1 to i6), bifurcated anteriorly; i1 lateral to anterior nostril; i2 between anterior and posterior nostrils (
<figureCitation id="4AA7ECED7C6F3558AE2FEC28FD4A5F09" box="[582,649,339,363]" captionStart="FIGURE 4" captionStartId="8.[487,560,1798,1818]" captionTargetBox="[505,1428,1270,1756]" captionTargetId="figure-321@8.[488,1459,1247,1777]" captionTargetPageId="8" captionText="FIGURE 4 | Rhyacoglanis varii laterosensory canal pores, MZUSP 54082, 47.6 mm SL, paratype, in dorsal (A) and ventral views (B). i1 to i6 = infraorbital pores 1 to 6; pm1 to pm11 = preoperculomandubular pores 1 to 11; po1 and po2 = postotic pores 1 and 2; s1 to s8 = supraorbital pores 1 to 8. Scale bar = 5 mm." figureDoi="http://doi.org/10.5281/zenodo.11124766" httpUri="https://zenodo.org/record/11124766/files/figure.png" pageId="8" pageNumber="9">Fig. 4</figureCitation>
). Preoperculomandibular (pm) canal with ten branches; arched in ventral view; pm1 medio-ventral on head; pm9 at base of opercular bone; canal continuing dorsally, finishing with pm11 fused with first postotic branch (po1). Postotic canal with two branches (po1 and po2), followed posteriorly by trunk lateral-line canal.
</paragraph>
<paragraph id="D223F0687C6F3559AE6FEF75FA635F09" blockId="8.[488,1460,526,1147]" lastBlockId="9.[488,1460,190,363]" lastPageId="9" lastPageNumber="10" pageId="8" pageNumber="9">
<emphasis id="E0E82C7A7C6F3558AE6FEF75FCCC5C44" bold="true" box="[518,783,526,550]" pageId="8" pageNumber="9">Coloration in alcohol.</emphasis>
Lateral surfaces of body with yellowish brown ground color. Dorsal surface of head light gray with scattered dark brown specks; pale yellow blotch on adductor muscle region. Maxillary barbel light brown with dark brown specks; mentonian barbel light yellow. Trunk and caudal regions with three dark brown bars. Subdorsal trunk bar approximately triangular, attenuated ventrally, lowermost portion slightly entering belly. Subadipose bar broad, spanning distance between bases of adipose and anal fins, and confluent with contralateral bar, shape roughly rectangular with anterior and posterior margins irregularly concave; rarely in extreme dorsal and ventral contact with dark brown bar on caudal peduncle. Caudal peduncle bar slightly narrower, approximately rectangular, anterior, and posterior margins irregularly concave, dorsally, and ventrally confluent with contralateral bar. Dark brown spots scattered throughout lateral region of body. Ventral region predominantly light yellow, without dark spots. Area between pelvic fins sometimes with tiny spots. Dorsal fin dark brown, distal third with hyaline band; occasionally, band of hyaline elliptical spots on membranes between rays in second lower quarter. Adipose fin with central dark brown vertical band completely separating light yellow spots on anterior and posterior regions, respectively. Pectoral fin hyaline with narrow dark brown band on median region. Pelvic fin hyaline with irregular or inconspicuous dark brown band across middle of rays. Anal fin hyaline; dark brown, approximately triangular spot near origin, confluent with subadipose band; dark brown oval spots forming irregular band across middle rays. Caudal fin with 3-shaped dark brown band through posterior region; median tip usually confluent with the dark brown bar on caudal peduncle; distal margin hyaline (
<figureCitation id="4AA7ECED7C6E3559A926EC28FA4C5F09" box="[1359,1423,339,363]" captionStart="FIGURE 1" captionStartId="4.[488,561,1761,1781]" captionTargetBox="[488,1459,432,1737]" captionTargetId="figure-20@4.[488,1459,432,1737]" captionTargetPageId="4" captionText="FIGURE 1 | Holotype of Rhyacoglanis varii, MZUEL 22317, 67.1 mm SL, rio do Peixe, upper rio Tocantins basin, Goianésia, state of Goiás, Brazil." figureDoi="http://doi.org/10.5281/zenodo.11124760" httpUri="https://zenodo.org/record/11124760/files/figure.png" pageId="9" pageNumber="10">Fig. 1</figureCitation>
).
</paragraph>
<caption id="86E3A0E07C6F3558AD8EEA7DFAA65907" ID-DOI="http://doi.org/10.5281/zenodo.11124766" ID-Zenodo-Dep="11124766" httpUri="https://zenodo.org/record/11124766/files/figure.png" pageId="8" pageNumber="9" startId="8.[487,560,1798,1818]" targetBox="[505,1428,1270,1756]" targetPageId="8" targetType="figure">
<paragraph id="D223F0687C6F3558AD8EEA7DFAA65907" blockId="8.[487,1464,1798,1893]" pageId="8" pageNumber="9">
<emphasis id="E0E82C7A7C6F3558AD8EEA7DFD385979" bold="true" box="[487,763,1798,1819]" pageId="8" pageNumber="9">
FIGURE 4 |
<taxonomicName id="159C8BEB7C6F3558AE30EA7CFD385979" box="[601,763,1799,1819]" family="Pseudopimelodidae" genus="Rhyacoglanis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Siluriformes" pageId="8" pageNumber="9" phylum="Chordata" rank="species" species="varii">
<emphasis id="E0E82C7A7C6F3558AE30EA7CFD385979" bold="true" box="[601,763,1799,1819]" italics="true" pageId="8" pageNumber="9">Rhyacoglanis varii</emphasis>
</taxonomicName>
</emphasis>
laterosensory canal pores, MZUSP 54082, 47.6 mm SL, paratype, in dorsal (
<emphasis id="E0E82C7A7C6F3558AD87EA57FE3D595D" bold="true" box="[494,510,1836,1855]" pageId="8" pageNumber="9">A</emphasis>
) and ventral views (
<emphasis id="E0E82C7A7C6F3558AED2EA57FD0A595D" bold="true" box="[699,713,1836,1855]" pageId="8" pageNumber="9">B</emphasis>
). i1 to i6 = infraorbital pores 1 to 6; pm1 to pm11 = preoperculomandubular pores 1 to 11; po1 and po2 = postotic pores 1 and 2; s1 to s8 = supraorbital pores 1 to 8. Scale bar = 5 mm.
</paragraph>
</caption>
<paragraph id="D223F0687C6E3559AE6FECE5FD505D14" blockId="9.[488,1459,414,886]" pageId="9" pageNumber="10">
<emphasis id="E0E82C7A7C6E3559AE6FECE5FD195FD4" bold="true" box="[518,730,414,438]" pageId="9" pageNumber="10">Coloration in life.</emphasis>
Head dorsal region reddish-brown, lateral region with yellowish- brown spot, peppered with dark brown spots of about pupil size. Maxillary barbel base light-brown, grayish posteriorly. Eye gray. Trunk and caudal peduncle ground color light yellowish-brown, with three large black vertical bars (subdorsal, subadipose, and caudal peduncle), peppered with dark brown spots of eye diameter size. Subdorsal bar triangular; attenuated ventrally; subadipose bar trapezoidal; caudal peduncle bar vertically rectangular. Pectoral fin hyaline, reddish-brown, with dark brown band crossing middle of rays. Dorsal fin about 3/4 black, with distinct reddish-brown distal margin. Pelvic fin hyaline, reddish-brown, with dark brown spots crossing middle of rays. Adipose fin black with light brown spots on anterior and posterior regions, respectively. Anal fin light brown with dark brown band across middle of rays. Caudal fin hyaline, light brown, with 3-shaped black band on posterior half; distal margin hyaline (
<figureCitation id="4AA7ECED7C6E3559AE20EE25FD485D14" box="[585,651,862,886]" captionStart="FIGURE 5" captionStartId="9.[136,209,1649,1669]" captionTargetBox="[136,1459,966,1627]" captionTargetId="figure-235@9.[136,1459,966,1627]" captionTargetPageId="9" captionText="FIGURE 5 | Rhyacoglanis varii coloration in life, MZUEL 6039, paratype, 46.1 mm SL, rio do Peixe, upper rio Tocantins basin, Goianésia, state of Goiás, Brazil." figureDoi="http://doi.org/10.5281/zenodo.11124768" httpUri="https://zenodo.org/record/11124768/files/figure.png" pageId="9" pageNumber="10">Fig. 5</figureCitation>
)
</paragraph>
<caption id="86E3A0E07C6E3559ACE1EB0AFEDE58C9" ID-DOI="http://doi.org/10.5281/zenodo.11124768" ID-Zenodo-Dep="11124768" httpUri="https://zenodo.org/record/11124768/files/figure.png" pageId="9" pageNumber="10" startId="9.[136,209,1649,1669]" targetBox="[136,1459,966,1627]" targetPageId="9" targetType="figure">
<paragraph id="D223F0687C6E3559ACE1EB0AFEDE58C9" blockId="9.[136,1459,1649,1707]" pageId="9" pageNumber="10">
<emphasis id="E0E82C7A7C6E3559ACE1EB0AFE5F58E7" bold="true" box="[136,412,1649,1669]" pageId="9" pageNumber="10">
FIGURE 5 |
<taxonomicName id="159C8BEB7C6E3559AC93EB0AFE5F58E7" box="[250,412,1649,1669]" family="Pseudopimelodidae" genus="Rhyacoglanis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Siluriformes" pageId="9" pageNumber="10" phylum="Chordata" rank="species" species="varii">
<emphasis id="E0E82C7A7C6E3559AC93EB0AFE5F58E7" bold="true" box="[250,412,1649,1669]" italics="true" pageId="9" pageNumber="10">Rhyacoglanis varii</emphasis>
</taxonomicName>
</emphasis>
coloration in life, MZUEL 6039, paratype, 46.1 mm SL, rio do Peixe, upper rio Tocantins basin, Goianésia, state of Goiás, Brazil.
</paragraph>
</caption>
<paragraph id="D223F0687C6D355AAE6FEDC5FC395E99" blockId="10.[488,1459,190,251]" pageId="10" pageNumber="11">
<emphasis id="E0E82C7A7C6D355AAE6FEDC5FC895EB4" bold="true" box="[518,842,190,214]" pageId="10" pageNumber="11">Geographical distribution.</emphasis>
<taxonomicName id="159C8BEB7C6D355AAF3CEDC5FBDA5EB4" box="[853,1049,190,214]" family="Pseudopimelodidae" genus="Rhyacoglanis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Siluriformes" pageId="10" pageNumber="11" phylum="Chordata" rank="species" species="varii">
<emphasis id="E0E82C7A7C6D355AAF3CEDC5FBDA5EB4" box="[853,1049,190,214]" italics="true" pageId="10" pageNumber="11">Rhyacoglanis varii</emphasis>
</taxonomicName>
is known only from the upper rio
<collectingRegion id="10583E8A7C6D355AAD81ED98FD9A5E99" box="[488,601,227,251]" country="Brazil" name="Tocantins" pageId="10" pageNumber="11">Tocantins</collectingRegion>
basin on the Brazilian shield (
<figureCitation id="4AA7ECED7C6D355AAFC3ED98FC2E5E99" box="[938,1005,227,251]" captionStart="FIGURE 6" captionStartId="10.[136,209,1798,1818]" captionTargetBox="[149,1430,917,1733]" captionTargetId="figure-208@10.[136,1459,880,1754]" captionTargetPageId="10" captionText="FIGURE 6 | Map of Rhyacoglanis varii distribution through upper rio Tocantins basin (red dot = rio Tocantinzinho tributary; red star = type locality, rio do Peixe), state of Goiás (GO), Brazil. DF = Federal District of Brazil." figureDoi="http://doi.org/10.5281/zenodo.11124772" httpUri="https://zenodo.org/record/11124772/files/figure.png" pageId="10" pageNumber="11">Fig. 6</figureCitation>
).
</paragraph>
<paragraph id="D223F0687C6D355AAE6FEC55FC365CD9" blockId="10.[488,1460,302,699]" pageId="10" pageNumber="11">
<emphasis id="E0E82C7A7C6D355AAE6FEC55FD145F24" bold="true" box="[518,727,302,326]" pageId="10" pageNumber="11">Ecological notes.</emphasis>
The
<typeStatus id="0D274ECA7C6D355AAF75EC55FC8E5F24" box="[796,845,302,326]" pageId="10" pageNumber="11">type</typeStatus>
locality in the rio do Peixe presented clear running waters, mostly shallow, with deeper pools approximately knee height. The bottom is predominantly pebbles, with rocks, sand, and organic litter (
<figureCitation id="4AA7ECED7C6D355AA8C2EC03FAEB5FF3" box="[1195,1320,376,401]" captionStart="FIGURE 7" captionStartId="11.[488,561,753,773]" captionTargetBox="[488,1459,176,730]" captionTargetId="figure-306@11.[488,1459,176,731]" captionTargetPageId="11" captionText="FIGURE 7 | A. Panoramic view of Rhyacoglanis varii type locality, rio do Peixe, Goianésia, state of Goiás, Brazil; B. Bottom with rocks, pebbles, and sand where specimens were collected; C. Blue arrow indicating water flow and pointing to a specimen beneath rocks." figureDoi="http://doi.org/10.5281/zenodo.11124774" httpUri="https://zenodo.org/record/11124774/files/figure.png" pageId="10" pageNumber="11">Figs. 7A,B</figureCitation>
). The color pattern of the species, yellowish or orange-brown background with alternating dark brown bars, allows for perfect camouflage amid the rocks and litter (
<figureCitation id="4AA7ECED7C6D355AA88DECB8FAFF5FB9" box="[1252,1340,451,475]" captionStart="FIGURE 7" captionStartId="11.[488,561,753,773]" captionTargetBox="[488,1459,176,730]" captionTargetId="figure-306@11.[488,1459,176,731]" captionTargetPageId="11" captionText="FIGURE 7 | A. Panoramic view of Rhyacoglanis varii type locality, rio do Peixe, Goianésia, state of Goiás, Brazil; B. Bottom with rocks, pebbles, and sand where specimens were collected; C. Blue arrow indicating water flow and pointing to a specimen beneath rocks." figureDoi="http://doi.org/10.5281/zenodo.11124774" httpUri="https://zenodo.org/record/11124774/files/figure.png" pageId="10" pageNumber="11">Fig. 7C</figureCitation>
). The area is in a rural and pristine zone, with marginal vegetation well preserved, which is the source of the organic litter. Dissolved oxygen
<quantity id="15645D8D7C6D355AAF8CEF75FBF75C44" box="[997,1076,526,550]" metricMagnitude="-6" metricUnit="kg" metricValue="8.3" pageId="10" pageNumber="11" unit="mg" value="8.3">8.3 mg</quantity>
.L-
<superScript id="25E95D207C6D355AA839EF77FB9B5C78" attach="right" box="[1104,1112,524,538]" fontSize="6" pageId="10" pageNumber="11">1</superScript>
, pH 8.06, conductivity 83.4μS.
<superScript id="25E95D207C6D355AAD81EF4FFDDA5C22" attach="left" box="[488,537,562,587]" fontSize="6" pageId="10" pageNumber="11">cm-1</superScript>
, and temperature 22.5° C (at 12:54h Brasília time). Specimens were found under rocks, protected from the water current (
<figureCitation id="4AA7ECED7C6D355AAFAAEF23FBDD5C13" box="[963,1054,600,625]" captionStart="FIGURE 7" captionStartId="11.[488,561,753,773]" captionTargetBox="[488,1459,176,730]" captionTargetId="figure-306@11.[488,1459,176,731]" captionTargetPageId="11" captionText="FIGURE 7 | A. Panoramic view of Rhyacoglanis varii type locality, rio do Peixe, Goianésia, state of Goiás, Brazil; B. Bottom with rocks, pebbles, and sand where specimens were collected; C. Blue arrow indicating water flow and pointing to a specimen beneath rocks." figureDoi="http://doi.org/10.5281/zenodo.11124774" httpUri="https://zenodo.org/record/11124774/files/figure.png" pageId="10" pageNumber="11">Fig. 7C</figureCitation>
), and collected with a sieve with a high-density polyethylene mesh of
<quantity id="15645D8D7C6D355AAF13EF05FC7E5CF7" box="[890,957,638,662]" metricMagnitude="-3" metricUnit="m" metricValue="2.0" pageId="10" pageNumber="11" unit="mm" value="2.0">2 mm</quantity>
size as rocks were turned. The use of trawls or fishing nets was inefficient for their capture.
</paragraph>
</subSubSection>
<subSubSection id="9A86A3E37C6D355FAE6FEF95FCF15DA2" lastPageId="15" lastPageNumber="16" pageId="10" pageNumber="11" type="etymology">
<paragraph id="D223F0687C6D355AAE6FEF95FA9C5D49" blockId="10.[488,1459,750,811]" pageId="10" pageNumber="11">
<emphasis id="E0E82C7A7C6D355AAE6FEF95FD525D64" bold="true" box="[518,657,750,774]" pageId="10" pageNumber="11">Etymology.</emphasis>
The species name
<taxonomicName id="159C8BEB7C6D355AAF0DEF94FC515D64" box="[868,914,751,774]" family="Pseudopimelodidae" genus="Rhyacoglanis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Siluriformes" pageId="10" pageNumber="11" phylum="Chordata" rank="species" species="varii">
<emphasis id="E0E82C7A7C6D355AAF0DEF94FC515D64" box="[868,914,751,774]" italics="true" pageId="10" pageNumber="11">varii</emphasis>
</taxonomicName>
honors the ichthyologist Richard P. Vari (1949 2016) for his outstanding contributions to the systematics of Neotropical fishes.
</paragraph>
<caption id="86E3A0E07C6D355AACE1EA7DFCB15922" ID-DOI="http://doi.org/10.5281/zenodo.11124772" ID-Zenodo-Dep="11124772" httpUri="https://zenodo.org/record/11124772/files/figure.png" pageId="10" pageNumber="11" startId="10.[136,209,1798,1818]" targetBox="[149,1430,917,1733]" targetPageId="10" targetType="figure">
<paragraph id="D223F0687C6D355AACE1EA7DFCB15922" blockId="10.[136,1459,1798,1856]" pageId="10" pageNumber="11">
<emphasis id="E0E82C7A7C6D355AACE1EA7DFF345978" bold="true" box="[136,247,1798,1818]" pageId="10" pageNumber="11">FIGURE 6 |</emphasis>
Map of
<taxonomicName id="159C8BEB7C6D355AAD2FEA7CFE2A5979" box="[326,489,1799,1819]" family="Pseudopimelodidae" genus="Rhyacoglanis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Siluriformes" pageId="10" pageNumber="11" phylum="Chordata" rank="species" species="varii">
<emphasis id="E0E82C7A7C6D355AAD2FEA7CFE2A5979" bold="true" box="[326,489,1799,1819]" italics="true" pageId="10" pageNumber="11">Rhyacoglanis varii</emphasis>
</taxonomicName>
distribution through upper rio Tocantins basin (red dot = rio Tocantinzinho tributary; red star = type locality, rio do Peixe), state of Goiás (GO), Brazil. DF = Federal District of Brazil.
</paragraph>
</caption>
<caption id="86E3A0E07C6C355BAD81EF8AFB8A5D32" ID-DOI="http://doi.org/10.5281/zenodo.11124774" ID-Zenodo-Dep="11124774" httpUri="https://zenodo.org/record/11124774/files/figure.png" pageId="11" pageNumber="12" startId="11.[488,561,753,773]" targetBox="[488,1459,176,730]" targetPageId="11" targetType="figure">
<paragraph id="D223F0687C6C355BAD81EF8AFB8A5D32" blockId="11.[488,1459,753,848]" pageId="11" pageNumber="12">
<emphasis id="E0E82C7A7C6C355BAD81EF8AFDBE5D66" bold="true" box="[488,637,753,773]" pageId="11" pageNumber="12">FIGURE 7 | A.</emphasis>
Panoramic view of
<taxonomicName id="159C8BEB7C6C355BAF2CEF8AFC295D67" box="[837,1002,753,773]" family="Pseudopimelodidae" genus="Rhyacoglanis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Siluriformes" pageId="11" pageNumber="12" phylum="Chordata" rank="species" species="varii">
<emphasis id="E0E82C7A7C6C355BAF2CEF8AFC295D67" bold="true" box="[837,1002,753,773]" italics="true" pageId="11" pageNumber="12">Rhyacoglanis varii</emphasis>
</taxonomicName>
type locality, rio do Peixe, Goianésia, state of Goiás, Brazil;
<emphasis id="E0E82C7A7C6C355BAE02EE6CFD435D48" bold="true" box="[619,640,791,810]" pageId="11" pageNumber="12">B.</emphasis>
Bottom with rocks, pebbles, and sand where specimens were collected;
<emphasis id="E0E82C7A7C6C355BA946EE6CFA875D48" bold="true" box="[1327,1348,791,810]" pageId="11" pageNumber="12">C.</emphasis>
Blue arrow indicating water flow and pointing to a specimen beneath rocks.
</paragraph>
</caption>
<paragraph id="D223F0687C6C355BAE6FEED3FD3A5BE2" blockId="11.[488,1459,936,1408]" pageId="11" pageNumber="12">
<emphasis id="E0E82C7A7C6C355BAE6FEED3FD3A5DA3" bold="true" box="[518,761,936,961]" pageId="11" pageNumber="12">Conservation status.</emphasis>
Considering the criterion established by the
<collectionCode id="B48D68AD7C6C355BA89FEED2FA835DA3" box="[1270,1344,937,961]" pageId="11" pageNumber="12">IUCN</collectionCode>
Standards and Petitions Committee (
<collectionCode id="B48D68AD7C6C355BAF73EEB5FCA45D84" box="[794,871,974,998]" pageId="11" pageNumber="12">IUCN</collectionCode>
, 2022), the species is Data Deficient (DD) because “very little information is known about a taxon, but the available information indicates that the taxon may be threatened”. The species is apparently restricted to the upper
<collectingRegion id="10583E8A7C6C355BAD81E945FD9A5A34" box="[488,601,1086,1110]" country="Brazil" name="Tocantins" pageId="11" pageNumber="12">Tocantins</collectingRegion>
basin, but the absence of the species in several collections points out its low occurrence in the basin. For instance, in the same expedition when the species was collected, the effort failed to capture it in 18 other localities of Barro Alto, Goianésia, and Pirenópolis municipalities (
<collectionCode id="B48D68AD7C6C355BAF73E9D5FC975AA4" box="[794,852,1198,1222]" pageId="11" pageNumber="12">OAS</collectionCode>
, pers. obs.). Furthermore, it is worth mentioning that the rapids environment where the species occurs often lends itself to the construction of hydroelectric plants. Dams entirely alter the rivers flow and seasonality (
<bibRefCitation id="B60D8D997C6C355BA959E983FD9C5B54" author="Winemiller KO &amp; Mcintyre PB &amp; Castello L &amp; Fluet-Chouinard E &amp; Giarrizzo T &amp; Nam S" pageId="11" pageNumber="12" pagination="128 - 29" refId="ref9064" refString="• Winemiller KO, Mcintyre PB, Castello L, Fluet-Chouinard E, Giarrizzo T, Nam S et al. Balancing hydropower and biodiversity in the Amazon, Congo, and Mekong. Science. 2016; 351 (6269): 128 - 29. https: // doi. org / 10.1126 / science. aac 7082" type="journal article" year="2016">
Winemiller
<emphasis id="E0E82C7A7C6C355BAD81E864FDD95B54" box="[488,538,1310,1334]" italics="true" pageId="11" pageNumber="12">et al.</emphasis>
, 2016
</bibRefCitation>
) and can thereby jeopardize the local population. Also, the dam of the Serra da Mesa power plant may obstruct movement between populations upstream and downstream of the dam.
</paragraph>
<paragraph id="D223F0687C6C355BAE6FE8C8FAF35979" blockId="11.[488,1459,1459,1819]" pageId="11" pageNumber="12">
<emphasis id="E0E82C7A7C6C355BAE6FE8C8FC085BA9" bold="true" box="[518,971,1459,1483]" pageId="11" pageNumber="12">Multivariate morphometric analysis.</emphasis>
Differences in the shape of
<taxonomicName id="159C8BEB7C6C355BA943E8C8FDD55B92" family="Pseudopimelodidae" genus="Rhyacoglanis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Siluriformes" pageId="11" pageNumber="12" phylum="Chordata" rank="species" species="varii">
<emphasis id="E0E82C7A7C6C355BA943E8C8FDD55B92" italics="true" pageId="11" pageNumber="12">Rhyacoglanis varii</emphasis>
</taxonomicName>
,
<taxonomicName id="159C8BEB7C6C355BAE4CE8A2FD735B92" authorityName="Shibatta &amp; Vari" authorityYear="2017" box="[549,688,1497,1520]" family="Pseudopimelodidae" genus="Rhyacoglanis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Siluriformes" pageId="11" pageNumber="12" phylum="Chordata" rank="species" species="paranensis">
<emphasis id="E0E82C7A7C6C355BAE4CE8A2FD735B92" box="[549,688,1497,1520]" italics="true" pageId="11" pageNumber="12">R. paranensis</emphasis>
</taxonomicName>
, and
<taxonomicName id="159C8BEB7C6C355BAE9AE8A2FC6D5B92" authorityName="Shibatta, Rocha &amp; Oliveira" authorityYear="2021" box="[755,942,1496,1520]" family="Pseudopimelodidae" genus="Rhyacoglanis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Siluriformes" pageId="11" pageNumber="12" phylum="Chordata" rank="species" species="rapppydanielae">
<emphasis id="E0E82C7A7C6C355BAE9AE8A2FC6D5B92" box="[755,942,1496,1520]" italics="true" pageId="11" pageNumber="12">R. rapppydanielae</emphasis>
</taxonomicName>
are represented by the second and third axes of the principal components analysis (
<figureCitation id="4AA7ECED7C6C355BAFF8E885FC165874" box="[913,981,1534,1558]" captionStart="FIGURE 8" captionStartId="13.[488,561,1052,1072]" captionTargetBox="[504,1453,177,1014]" captionTargetId="figure-23@13.[487,1459,171,1023]" captionTargetPageId="13" captionText="FIGURE 8 | Second and third principal components axes of combined samples of Rhyacoglanis varii (red cross, n = 16), R. paranensis (blue square; n = 31), and R. rapppydanielae (pink square; n = 13)." figureDoi="http://doi.org/10.5281/zenodo.11124778" httpUri="https://zenodo.org/record/11124778/files/figure.png" pageId="11" pageNumber="12">Fig. 8</figureCitation>
). The character loadings, eigenvalues, and percentage of variation are presented in
<tableCitation id="9F1EC5D37C6C355BAFF1EB58FC235859" box="[920,992,1571,1595]" captionStart="TABLE 2" captionStartId="12.[488,550,193,213]" captionTargetPageId="12" captionText="TABLE 2 | Character loadings of the first three principal components axes of combined analysis of Rhyacoglanis varii (N = 16), R. paranensis (N = 31), and R. rapppydanielae (N = 13)." httpUri="http://table.plazi.org/id/86E3A0E07C6B355CAD81EDBAFB785E99" pageId="11" pageNumber="12" tableUuid="86E3A0E07C6B355CAD81EDBAFB785E99">Tab. 2</tableCitation>
.
<taxonomicName id="159C8BEB7C6C355BAF80EB58FB665859" box="[1001,1189,1571,1595]" family="Pseudopimelodidae" genus="Rhyacoglanis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Siluriformes" pageId="11" pageNumber="12" phylum="Chordata" rank="species" species="varii">
<emphasis id="E0E82C7A7C6C355BAF80EB58FB665859" box="[1001,1189,1571,1595]" italics="true" pageId="11" pageNumber="12">Rhyacoglanis varii</emphasis>
</taxonomicName>
differs from
<taxonomicName id="159C8BEB7C6C355BA944EB5FFA705859" authorityName="Shibatta &amp; Vari" authorityYear="2017" box="[1325,1459,1572,1595]" family="Pseudopimelodidae" genus="Rhyacoglanis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Siluriformes" pageId="11" pageNumber="12" phylum="Chordata" rank="species" species="paranensis">
<emphasis id="E0E82C7A7C6C355BA944EB5FFA705859" box="[1325,1459,1572,1595]" italics="true" pageId="11" pageNumber="12">R. paranensis</emphasis>
</taxonomicName>
by the longer anus-to-anal-fin origin distance, maxillary barbel length, caudal peduncle depth (positive loadings), and minor posterior nostril-to-eye distance, body depth, and postcleithral process length (negative loadings).
<taxonomicName id="159C8BEB7C6C355BAF97EBE8FAEB58C9" authorityName="Shibatta, Rocha &amp; Oliveira" authorityYear="2021" box="[1022,1320,1683,1707]" family="Pseudopimelodidae" genus="Rhyacoglanis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Siluriformes" pageId="11" pageNumber="12" phylum="Chordata" rank="species" species="rapppydanielae">
<emphasis id="E0E82C7A7C6C355BAF97EBE8FAEB58C9" box="[1022,1320,1683,1707]" italics="true" pageId="11" pageNumber="12">Rhyacoglanis rapppydanielae</emphasis>
</taxonomicName>
differs from
<taxonomicName id="159C8BEB7C6C355BAD81EBC2FDF758B2" box="[488,564,1721,1744]" family="Pseudopimelodidae" genus="Rhyacoglanis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Siluriformes" pageId="11" pageNumber="12" phylum="Chordata" rank="species" species="varii">
<emphasis id="E0E82C7A7C6C355BAD81EBC2FDF758B2" box="[488,564,1721,1744]" italics="true" pageId="11" pageNumber="12">R. varii</emphasis>
</taxonomicName>
and
<taxonomicName id="159C8BEB7C6C355BAE00EBC2FD3358B2" authorityName="Shibatta &amp; Vari" authorityYear="2017" box="[617,752,1721,1744]" family="Pseudopimelodidae" genus="Rhyacoglanis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Siluriformes" pageId="11" pageNumber="12" phylum="Chordata" rank="species" species="paranensis">
<emphasis id="E0E82C7A7C6C355BAE00EBC2FD3358B2" box="[617,752,1721,1744]" italics="true" pageId="11" pageNumber="12">R. paranensis</emphasis>
</taxonomicName>
by the longer distance between posterior nostril to eye distance, anus to anal-fin origin distance, and mouth width (positive loadings), and by the smaller dorsal-fin spine length and post-cleithral process length (negative loadings).
</paragraph>
<caption id="86E3A0E07C6B355CAD81EDBAFB785E99" ID-Table-UUID="86E3A0E07C6B355CAD81EDBAFB785E99" httpUri="http://table.plazi.org/id/86E3A0E07C6B355CAD81EDBAFB785E99" pageId="12" pageNumber="13" startId="12.[488,550,193,213]" targetBox="[503,1414,298,1661]" targetIsTable="true" targetPageId="12" targetType="table">
<paragraph id="D223F0687C6B355CAD81EDBAFB785E99" blockId="12.[488,1459,193,251]" pageId="12" pageNumber="13">
<emphasis id="E0E82C7A7C6B355CAD81EDBAFD8C5EB7" bold="true" box="[488,591,193,213]" pageId="12" pageNumber="13">TABLE 2 |</emphasis>
Character loadings of the first three principal components axes of combined analysis of
<taxonomicName id="159C8BEB7C6B355CAD81ED9CFD4A5E99" box="[488,649,231,251]" family="Pseudopimelodidae" genus="Rhyacoglanis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Siluriformes" pageId="12" pageNumber="13" phylum="Chordata" rank="species" species="varii">
<emphasis id="E0E82C7A7C6B355CAD81ED9CFD4A5E99" bold="true" box="[488,649,231,251]" italics="true" pageId="12" pageNumber="13">Rhyacoglanis varii</emphasis>
</taxonomicName>
(N = 16),
<taxonomicName id="159C8BEB7C6B355CAEB4ED9CFC915E99" authorityName="Shibatta &amp; Vari" authorityYear="2017" box="[733,850,231,251]" family="Pseudopimelodidae" genus="Rhyacoglanis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Siluriformes" pageId="12" pageNumber="13" phylum="Chordata" rank="species" species="paranensis">
<emphasis id="E0E82C7A7C6B355CAEB4ED9CFC915E99" bold="true" box="[733,850,231,251]" italics="true" pageId="12" pageNumber="13">R. paranensis</emphasis>
</taxonomicName>
(N = 31), and
<taxonomicName id="159C8BEB7C6B355CAFA7ED9CFBAF5E99" authorityName="Shibatta, Rocha &amp; Oliveira" authorityYear="2021" box="[974,1132,231,251]" family="Pseudopimelodidae" genus="Rhyacoglanis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Siluriformes" pageId="12" pageNumber="13" phylum="Chordata" rank="species" species="rapppydanielae">
<emphasis id="E0E82C7A7C6B355CAFA7ED9CFBAF5E99" bold="true" box="[974,1132,231,251]" italics="true" pageId="12" pageNumber="13">R. rapppydanielae</emphasis>
</taxonomicName>
(N = 13).
</paragraph>
</caption>
<paragraph id="D223F0687C6B355CAD9EEC51FAB4581F" pageId="12" pageNumber="13">
<table id="A09C02C87C6BCAAFAD9EEC51FA45581F" box="[503,1414,298,1661]" gridcols="4" gridrows="31" pageId="12" pageNumber="13">
<tr id="6CACF22A7C6BCAAFAD9EEC51FA455F5E" box="[503,1414,298,316]" gridrow="0" pageId="12" pageNumber="13">
<th id="2F7D9B567C6BCAAFAD9EEC51FCFD5F5E" box="[503,830,298,316]" gridcol="0" gridrow="0" pageId="12" pageNumber="13">
<emphasis id="E0E82C7A7C6B355CAD9EEC51FD9C5F5E" bold="true" box="[503,607,298,316]" pageId="12" pageNumber="13">Characters</emphasis>
</th>
<th id="2F7D9B567C6BCAAFAFB2EC51FBEA5F5E" box="[987,1065,298,316]" gridcol="1" gridrow="0" pageId="12" pageNumber="13">
<emphasis id="E0E82C7A7C6B355CAF87EC51FBD65F5E" bold="true" box="[1006,1045,298,316]" pageId="12" pageNumber="13">PC 1</emphasis>
</th>
<th id="2F7D9B567C6BCAAFA8E8EC51FB195F5E" box="[1153,1242,298,316]" gridcol="2" gridrow="0" pageId="12" pageNumber="13">
<emphasis id="E0E82C7A7C6B355CA8F3EC51FB025F5E" bold="true" box="[1178,1217,298,316]" pageId="12" pageNumber="13">PC 2</emphasis>
</th>
<th id="2F7D9B567C6BCAAFA944EC51FA455F5E" box="[1325,1414,298,316]" gridcol="3" gridrow="0" pageId="12" pageNumber="13">
<emphasis id="E0E82C7A7C6B355CA92FEC51FAAE5F5E" bold="true" box="[1350,1389,298,316]" pageId="12" pageNumber="13">PC 3</emphasis>
</th>
</tr>
<tr id="6CACF22A7C6BCAAFAD9EEC26FA455F0D" box="[503,1414,349,367]" gridrow="1" pageId="12" pageNumber="13">
<th id="2F7D9B567C6BCAAFAD9EEC26FCFD5F0D" box="[503,830,349,367]" gridcol="0" gridrow="1" pageId="12" pageNumber="13">Head length</th>
<td id="2F7D9B567C6BCAAFAFB2EC26FBEA5F0D" box="[987,1065,349,367]" gridcol="1" gridrow="1" pageId="12" pageNumber="13">0.1915</td>
<td id="2F7D9B567C6BCAAFA8E8EC26FB195F0D" box="[1153,1242,349,367]" gridcol="2" gridrow="1" pageId="12" pageNumber="13">0.015</td>
<td id="2F7D9B567C6BCAAFA944EC26FA455F0D" box="[1325,1414,349,367]" gridcol="3" gridrow="1" pageId="12" pageNumber="13">0.05995</td>
</tr>
<tr id="6CACF22A7C6BCAAFAD9EECF1FA455FFE" box="[503,1414,394,412]" gridrow="2" pageId="12" pageNumber="13">
<th id="2F7D9B567C6BCAAFAD9EECF1FCFD5FFE" box="[503,830,394,412]" gridcol="0" gridrow="2" pageId="12" pageNumber="13">Eye diameter</th>
<td id="2F7D9B567C6BCAAFAFB2ECF1FBEA5FFE" box="[987,1065,394,412]" gridcol="1" gridrow="2" pageId="12" pageNumber="13">0.1345</td>
<td id="2F7D9B567C6BCAAFA8E8ECF1FB195FFE" box="[1153,1242,394,412]" gridcol="2" gridrow="2" pageId="12" pageNumber="13">0.09743</td>
<td id="2F7D9B567C6BCAAFA944ECF1FA455FFE" box="[1325,1414,394,412]" gridcol="3" gridrow="2" pageId="12" pageNumber="13">0.1875</td>
</tr>
<tr id="6CACF22A7C6BCAAFAD9EECCDFA455FAA" box="[503,1414,438,456]" gridrow="3" pageId="12" pageNumber="13">
<th id="2F7D9B567C6BCAAFAD9EECCDFCFD5FAA" box="[503,830,438,456]" gridcol="0" gridrow="3" pageId="12" pageNumber="13">Interorbital distance</th>
<td id="2F7D9B567C6BCAAFAFB2ECCDFBEA5FAA" box="[987,1065,438,456]" gridcol="1" gridrow="3" pageId="12" pageNumber="13">0.1988</td>
<td id="2F7D9B567C6BCAAFA8E8ECCDFB195FAA" box="[1153,1242,438,456]" gridcol="2" gridrow="3" pageId="12" pageNumber="13">0.01607</td>
<td id="2F7D9B567C6BCAAFA944ECCDFA455FAA" box="[1325,1414,438,456]" gridcol="3" gridrow="3" pageId="12" pageNumber="13">-0.04845</td>
</tr>
<tr id="6CACF22A7C6BCAAFAD9EEC98FA455F97" box="[503,1414,483,501]" gridrow="4" pageId="12" pageNumber="13">
<th id="2F7D9B567C6BCAAFAD9EEC98FCFD5F97" box="[503,830,483,501]" gridcol="0" gridrow="4" pageId="12" pageNumber="13">Snout length</th>
<td id="2F7D9B567C6BCAAFAFB2EC98FBEA5F97" box="[987,1065,483,501]" gridcol="1" gridrow="4" pageId="12" pageNumber="13">0.1942</td>
<td id="2F7D9B567C6BCAAFA8E8EC98FB195F97" box="[1153,1242,483,501]" gridcol="2" gridrow="4" pageId="12" pageNumber="13">-0.009734</td>
<td id="2F7D9B567C6BCAAFA944EC98FA455F97" box="[1325,1414,483,501]" gridcol="3" gridrow="4" pageId="12" pageNumber="13">0.02416</td>
</tr>
<tr id="6CACF22A7C6BCAAFAD9EEF6BFA455C40" box="[503,1414,528,546]" gridrow="5" pageId="12" pageNumber="13">
<th id="2F7D9B567C6BCAAFAD9EEF6BFCFD5C40" box="[503,830,528,546]" gridcol="0" gridrow="5" pageId="12" pageNumber="13">Mouth width</th>
<td id="2F7D9B567C6BCAAFAFB2EF6BFBEA5C40" box="[987,1065,528,546]" gridcol="1" gridrow="5" pageId="12" pageNumber="13">0.2077</td>
<td id="2F7D9B567C6BCAAFA8E8EF6BFB195C40" box="[1153,1242,528,546]" gridcol="2" gridrow="5" pageId="12" pageNumber="13">0.1981</td>
<td id="2F7D9B567C6BCAAFA944EF6BFA455C40" box="[1325,1414,528,546]" gridcol="3" gridrow="5" pageId="12" pageNumber="13">0.1694</td>
</tr>
<tr id="6CACF22A7C6BCAAFAD9EEF47FA455C2C" box="[503,1414,572,590]" gridrow="6" pageId="12" pageNumber="13">
<th id="2F7D9B567C6BCAAFAD9EEF47FCFD5C2C" box="[503,830,572,590]" gridcol="0" gridrow="6" pageId="12" pageNumber="13">Pectoral girdle width</th>
<td id="2F7D9B567C6BCAAFAFB2EF47FBEA5C2C" box="[987,1065,572,590]" gridcol="1" gridrow="6" pageId="12" pageNumber="13">0.2087</td>
<td id="2F7D9B567C6BCAAFA8E8EF47FB195C2C" box="[1153,1242,572,590]" gridcol="2" gridrow="6" pageId="12" pageNumber="13">0.05512</td>
<td id="2F7D9B567C6BCAAFA944EF47FA455C2C" box="[1325,1414,572,590]" gridcol="3" gridrow="6" pageId="12" pageNumber="13">0.01591</td>
</tr>
<tr id="6CACF22A7C6BCAAFAD9EEF12FA455C19" box="[503,1414,617,635]" gridrow="7" pageId="12" pageNumber="13">
<th id="2F7D9B567C6BCAAFAD9EEF12FCFD5C19" box="[503,830,617,635]" gridcol="0" gridrow="7" pageId="12" pageNumber="13">Predorsal distance</th>
<td id="2F7D9B567C6BCAAFAFB2EF12FBEA5C19" box="[987,1065,617,635]" gridcol="1" gridrow="7" pageId="12" pageNumber="13">0.1722</td>
<td id="2F7D9B567C6BCAAFA8E8EF12FB195C19" box="[1153,1242,617,635]" gridcol="2" gridrow="7" pageId="12" pageNumber="13">0.01745</td>
<td id="2F7D9B567C6BCAAFA944EF12FA455C19" box="[1325,1414,617,635]" gridcol="3" gridrow="7" pageId="12" pageNumber="13">0.03117</td>
</tr>
<tr id="6CACF22A7C6BCAAFAD9EEFEDFA455CCA" box="[503,1414,662,680]" gridrow="8" pageId="12" pageNumber="13">
<th id="2F7D9B567C6BCAAFAD9EEFEDFCFD5CCA" box="[503,830,662,680]" gridcol="0" gridrow="8" pageId="12" pageNumber="13">Dorsal-fin base length</th>
<td id="2F7D9B567C6BCAAFAFB2EFEDFBEA5CCA" box="[987,1065,662,680]" gridcol="1" gridrow="8" pageId="12" pageNumber="13">0.2031</td>
<td id="2F7D9B567C6BCAAFA8E8EFEDFB195CCA" box="[1153,1242,662,680]" gridcol="2" gridrow="8" pageId="12" pageNumber="13">-0.08244</td>
<td id="2F7D9B567C6BCAAFA944EFEDFA455CCA" box="[1325,1414,662,680]" gridcol="3" gridrow="8" pageId="12" pageNumber="13">-0.06269</td>
</tr>
<tr id="6CACF22A7C6BCAAFAD9EEFB9FA455CB6" box="[503,1414,706,724]" gridrow="9" pageId="12" pageNumber="13">
<th id="2F7D9B567C6BCAAFAD9EEFB9FCFD5CB6" box="[503,830,706,724]" gridcol="0" gridrow="9" pageId="12" pageNumber="13">Adipose-fin base length</th>
<td id="2F7D9B567C6BCAAFAFB2EFB9FBEA5CB6" box="[987,1065,706,724]" gridcol="1" gridrow="9" pageId="12" pageNumber="13">0.1661</td>
<td id="2F7D9B567C6BCAAFA8E8EFB9FB195CB6" box="[1153,1242,706,724]" gridcol="2" gridrow="9" pageId="12" pageNumber="13">0.06053</td>
<td id="2F7D9B567C6BCAAFA944EFB9FA455CB6" box="[1325,1414,706,724]" gridcol="3" gridrow="9" pageId="12" pageNumber="13">0.1708</td>
</tr>
<tr id="6CACF22A7C6BCAAFAD9EEF94FA455D63" box="[503,1414,751,769]" gridrow="10" pageId="12" pageNumber="13">
<th id="2F7D9B567C6BCAAFAD9EEF94FCFD5D63" box="[503,830,751,769]" gridcol="0" gridrow="10" pageId="12" pageNumber="13">Prepelvic distance</th>
<td id="2F7D9B567C6BCAAFAFB2EF94FBEA5D63" box="[987,1065,751,769]" gridcol="1" gridrow="10" pageId="12" pageNumber="13">0.186</td>
<td id="2F7D9B567C6BCAAFA8E8EF94FB195D63" box="[1153,1242,751,769]" gridcol="2" gridrow="10" pageId="12" pageNumber="13">0.02634</td>
<td id="2F7D9B567C6BCAAFA944EF94FA455D63" box="[1325,1414,751,769]" gridcol="3" gridrow="10" pageId="12" pageNumber="13">0.01305</td>
</tr>
<tr id="6CACF22A7C6BCAAFAD9EEE60FA455D4F" box="[503,1414,795,813]" gridrow="11" pageId="12" pageNumber="13">
<th id="2F7D9B567C6BCAAFAD9EEE60FCFD5D4F" box="[503,830,795,813]" gridcol="0" gridrow="11" pageId="12" pageNumber="13">Pelvic fin to anal fin distance</th>
<td id="2F7D9B567C6BCAAFAFB2EE60FBEA5D4F" box="[987,1065,795,813]" gridcol="1" gridrow="11" pageId="12" pageNumber="13">0.1913</td>
<td id="2F7D9B567C6BCAAFA8E8EE60FB195D4F" box="[1153,1242,795,813]" gridcol="2" gridrow="11" pageId="12" pageNumber="13">0.1087</td>
<td id="2F7D9B567C6BCAAFA944EE60FA455D4F" box="[1325,1414,795,813]" gridcol="3" gridrow="11" pageId="12" pageNumber="13">0.1432</td>
</tr>
<tr id="6CACF22A7C6BCAAFAD9EEE33FA455D38" box="[503,1414,840,858]" gridrow="12" pageId="12" pageNumber="13">
<th id="2F7D9B567C6BCAAFAD9EEE33FCFD5D38" box="[503,830,840,858]" gridcol="0" gridrow="12" pageId="12" pageNumber="13">Anal-fin base length</th>
<td id="2F7D9B567C6BCAAFAFB2EE33FBEA5D38" box="[987,1065,840,858]" gridcol="1" gridrow="12" pageId="12" pageNumber="13">0.1823</td>
<td id="2F7D9B567C6BCAAFA8E8EE33FB195D38" box="[1153,1242,840,858]" gridcol="2" gridrow="12" pageId="12" pageNumber="13">-0.07779</td>
<td id="2F7D9B567C6BCAAFA944EE33FA455D38" box="[1325,1414,840,858]" gridcol="3" gridrow="12" pageId="12" pageNumber="13">0.001686</td>
</tr>
<tr id="6CACF22A7C6BCAAFAD9EEE0EFA455DE5" box="[503,1414,885,903]" gridrow="13" pageId="12" pageNumber="13">
<th id="2F7D9B567C6BCAAFAD9EEE0EFCFD5DE5" box="[503,830,885,903]" gridcol="0" gridrow="13" pageId="12" pageNumber="13">Caudal peduncle length</th>
<td id="2F7D9B567C6BCAAFAFB2EE0EFBEA5DE5" box="[987,1065,885,903]" gridcol="1" gridrow="13" pageId="12" pageNumber="13">0.1888</td>
<td id="2F7D9B567C6BCAAFA8E8EE0EFB195DE5" box="[1153,1242,885,903]" gridcol="2" gridrow="13" pageId="12" pageNumber="13">-0.1205</td>
<td id="2F7D9B567C6BCAAFA944EE0EFA455DE5" box="[1325,1414,885,903]" gridcol="3" gridrow="13" pageId="12" pageNumber="13">0.1033</td>
</tr>
<tr id="6CACF22A7C6BCAAFAD9EEEDAFA455DD1" box="[503,1414,929,947]" gridrow="14" pageId="12" pageNumber="13">
<th id="2F7D9B567C6BCAAFAD9EEEDAFCFD5DD1" box="[503,830,929,947]" gridcol="0" gridrow="14" pageId="12" pageNumber="13">Head depth</th>
<td id="2F7D9B567C6BCAAFAFB2EEDAFBEA5DD1" box="[987,1065,929,947]" gridcol="1" gridrow="14" pageId="12" pageNumber="13">0.2193</td>
<td id="2F7D9B567C6BCAAFA8E8EEDAFB195DD1" box="[1153,1242,929,947]" gridcol="2" gridrow="14" pageId="12" pageNumber="13">-0.09948</td>
<td id="2F7D9B567C6BCAAFA944EEDAFA455DD1" box="[1325,1414,929,947]" gridcol="3" gridrow="14" pageId="12" pageNumber="13">-0.003842</td>
</tr>
<tr id="6CACF22A7C6BCAAFAD9EEEB5FA455D82" box="[503,1414,974,992]" gridrow="15" pageId="12" pageNumber="13">
<th id="2F7D9B567C6BCAAFAD9EEEB5FCFD5D82" box="[503,830,974,992]" gridcol="0" gridrow="15" pageId="12" pageNumber="13">Body depth</th>
<td id="2F7D9B567C6BCAAFAFB2EEB5FBEA5D82" box="[987,1065,974,992]" gridcol="1" gridrow="15" pageId="12" pageNumber="13">0.2262</td>
<td id="2F7D9B567C6BCAAFA8E8EEB5FB195D82" box="[1153,1242,974,992]" gridcol="2" gridrow="15" pageId="12" pageNumber="13">0.0633</td>
<td id="2F7D9B567C6BCAAFA944EEB5FA455D82" box="[1325,1414,974,992]" gridcol="3" gridrow="15" pageId="12" pageNumber="13">-0.2802</td>
</tr>
<tr id="6CACF22A7C6BCAAFAD9EEE81FA455A6E" box="[503,1414,1018,1036]" gridrow="16" pageId="12" pageNumber="13">
<th id="2F7D9B567C6BCAAFAD9EEE81FCFD5A6E" box="[503,830,1018,1036]" gridcol="0" gridrow="16" pageId="12" pageNumber="13">Caudal peduncle depth</th>
<td id="2F7D9B567C6BCAAFAFB2EE81FBEA5A6E" box="[987,1065,1018,1036]" gridcol="1" gridrow="16" pageId="12" pageNumber="13">0.1791</td>
<td id="2F7D9B567C6BCAAFA8E8EE81FB195A6E" box="[1153,1242,1018,1036]" gridcol="2" gridrow="16" pageId="12" pageNumber="13">0.09839</td>
<td id="2F7D9B567C6BCAAFA944EE81FA455A6E" box="[1325,1414,1018,1036]" gridcol="3" gridrow="16" pageId="12" pageNumber="13">0.2062</td>
</tr>
<tr id="6CACF22A7C6BCAAFAD9EE95CFA455A5B" box="[503,1414,1063,1081]" gridrow="17" pageId="12" pageNumber="13">
<th id="2F7D9B567C6BCAAFAD9EE95CFCFD5A5B" box="[503,830,1063,1081]" gridcol="0" gridrow="17" pageId="12" pageNumber="13">Pectoral-fin spine length</th>
<td id="2F7D9B567C6BCAAFAFB2E95CFBEA5A5B" box="[987,1065,1063,1081]" gridcol="1" gridrow="17" pageId="12" pageNumber="13">0.1622</td>
<td id="2F7D9B567C6BCAAFA8E8E95CFB195A5B" box="[1153,1242,1063,1081]" gridcol="2" gridrow="17" pageId="12" pageNumber="13">-0.1404</td>
<td id="2F7D9B567C6BCAAFA944E95CFA455A5B" box="[1325,1414,1063,1081]" gridcol="3" gridrow="17" pageId="12" pageNumber="13">0.04719</td>
</tr>
<tr id="6CACF22A7C6BCAAFAD9EE92FFA455A04" box="[503,1414,1108,1126]" gridrow="18" pageId="12" pageNumber="13">
<th id="2F7D9B567C6BCAAFAD9EE92FFCFD5A04" box="[503,830,1108,1126]" gridcol="0" gridrow="18" pageId="12" pageNumber="13">Dorsal-fin spine length</th>
<td id="2F7D9B567C6BCAAFAFB2E92FFBEA5A04" box="[987,1065,1108,1126]" gridcol="1" gridrow="18" pageId="12" pageNumber="13">0.171</td>
<td id="2F7D9B567C6BCAAFA8E8E92FFB195A04" box="[1153,1242,1108,1126]" gridcol="2" gridrow="18" pageId="12" pageNumber="13">-0.4545</td>
<td id="2F7D9B567C6BCAAFA944E92FFA455A04" box="[1325,1414,1108,1126]" gridcol="3" gridrow="18" pageId="12" pageNumber="13">-0.0166</td>
</tr>
<tr id="6CACF22A7C6BCAAFAD9EE9FBFA455AF0" box="[503,1414,1152,1170]" gridrow="19" pageId="12" pageNumber="13">
<th id="2F7D9B567C6BCAAFAD9EE9FBFCFD5AF0" box="[503,830,1152,1170]" gridcol="0" gridrow="19" pageId="12" pageNumber="13">Maxillary barbel length</th>
<td id="2F7D9B567C6BCAAFAFB2E9FBFBEA5AF0" box="[987,1065,1152,1170]" gridcol="1" gridrow="19" pageId="12" pageNumber="13">0.1625</td>
<td id="2F7D9B567C6BCAAFA8E8E9FBFB195AF0" box="[1153,1242,1152,1170]" gridcol="2" gridrow="19" pageId="12" pageNumber="13">-0.03909</td>
<td id="2F7D9B567C6BCAAFA944E9FBFA455AF0" box="[1325,1414,1152,1170]" gridcol="3" gridrow="19" pageId="12" pageNumber="13">0.2676</td>
</tr>
<tr id="6CACF22A7C6BCAAFAD9EE9D6FA455ADD" box="[503,1414,1197,1215]" gridrow="20" pageId="12" pageNumber="13">
<th id="2F7D9B567C6BCAAFAD9EE9D6FCFD5ADD" box="[503,830,1197,1215]" gridcol="0" gridrow="20" pageId="12" pageNumber="13">Pelvic length</th>
<td id="2F7D9B567C6BCAAFAFB2E9D6FBEA5ADD" box="[987,1065,1197,1215]" gridcol="1" gridrow="20" pageId="12" pageNumber="13">0.179</td>
<td id="2F7D9B567C6BCAAFA8E8E9D6FB195ADD" box="[1153,1242,1197,1215]" gridcol="2" gridrow="20" pageId="12" pageNumber="13">-0.1042</td>
<td id="2F7D9B567C6BCAAFA944E9D6FA455ADD" box="[1325,1414,1197,1215]" gridcol="3" gridrow="20" pageId="12" pageNumber="13">-0.05121</td>
</tr>
<tr id="6CACF22A7C6BCAAFAD9EE9A2FA455A89" box="[503,1414,1241,1259]" gridrow="21" pageId="12" pageNumber="13">
<th id="2F7D9B567C6BCAAFAD9EE9A2FCFD5A89" box="[503,830,1241,1259]" gridcol="0" gridrow="21" pageId="12" pageNumber="13">Anterior to posterior nostril distance</th>
<td id="2F7D9B567C6BCAAFAFB2E9A2FBEA5A89" box="[987,1065,1241,1259]" gridcol="1" gridrow="21" pageId="12" pageNumber="13">0.1462</td>
<td id="2F7D9B567C6BCAAFA8E8E9A2FB195A89" box="[1153,1242,1241,1259]" gridcol="2" gridrow="21" pageId="12" pageNumber="13">0.05437</td>
<td id="2F7D9B567C6BCAAFA944E9A2FA455A89" box="[1325,1414,1241,1259]" gridcol="3" gridrow="21" pageId="12" pageNumber="13">0.006021</td>
</tr>
<tr id="6CACF22A7C6BCAAFAD9EE87DFA455B7A" box="[503,1414,1286,1304]" gridrow="22" pageId="12" pageNumber="13">
<th id="2F7D9B567C6BCAAFAD9EE87DFCFD5B7A" box="[503,830,1286,1304]" gridcol="0" gridrow="22" pageId="12" pageNumber="13">Posterior nostril to eye distance</th>
<td id="2F7D9B567C6BCAAFAFB2E87DFBEA5B7A" box="[987,1065,1286,1304]" gridcol="1" gridrow="22" pageId="12" pageNumber="13">0.1529</td>
<td id="2F7D9B567C6BCAAFA8E8E87DFB195B7A" box="[1153,1242,1286,1304]" gridcol="2" gridrow="22" pageId="12" pageNumber="13">0.5834</td>
<td id="2F7D9B567C6BCAAFA944E87DFA455B7A" box="[1325,1414,1286,1304]" gridcol="3" gridrow="22" pageId="12" pageNumber="13">-0.5648</td>
</tr>
<tr id="6CACF22A7C6BCAAFAD9EE848FA455B27" box="[503,1414,1331,1349]" gridrow="23" pageId="12" pageNumber="13">
<th id="2F7D9B567C6BCAAFAD9EE848FCFD5B27" box="[503,830,1331,1349]" gridcol="0" gridrow="23" pageId="12" pageNumber="13">Posterior nostrils distance</th>
<td id="2F7D9B567C6BCAAFAFB2E848FBEA5B27" box="[987,1065,1331,1349]" gridcol="1" gridrow="23" pageId="12" pageNumber="13">0.2059</td>
<td id="2F7D9B567C6BCAAFA8E8E848FB195B27" box="[1153,1242,1331,1349]" gridcol="2" gridrow="23" pageId="12" pageNumber="13">-0.04857</td>
<td id="2F7D9B567C6BCAAFA944E848FA455B27" box="[1325,1414,1331,1349]" gridcol="3" gridrow="23" pageId="12" pageNumber="13">-0.002237</td>
</tr>
<tr id="6CACF22A7C6BCAAFAD9EE824FA455B13" box="[503,1414,1375,1393]" gridrow="24" pageId="12" pageNumber="13">
<th id="2F7D9B567C6BCAAFAD9EE824FCFD5B13" box="[503,830,1375,1393]" gridcol="0" gridrow="24" pageId="12" pageNumber="13">Postcleithral process length</th>
<td id="2F7D9B567C6BCAAFAFB2E824FBEA5B13" box="[987,1065,1375,1393]" gridcol="1" gridrow="24" pageId="12" pageNumber="13">0.2379</td>
<td id="2F7D9B567C6BCAAFA8E8E824FB195B13" box="[1153,1242,1375,1393]" gridcol="2" gridrow="24" pageId="12" pageNumber="13">-0.4045</td>
<td id="2F7D9B567C6BCAAFA944E824FA455B13" box="[1325,1414,1375,1393]" gridcol="3" gridrow="24" pageId="12" pageNumber="13">-0.273</td>
</tr>
<tr id="6CACF22A7C6BCAAFAD9EE8F7FA455BFC" box="[503,1414,1420,1438]" gridrow="25" pageId="12" pageNumber="13">
<th id="2F7D9B567C6BCAAFAD9EE8F7FCFD5BFC" box="[503,830,1420,1438]" gridcol="0" gridrow="25" pageId="12" pageNumber="13">Dorsal fin to pelvic fin distance</th>
<td id="2F7D9B567C6BCAAFAFB2E8F7FBEA5BFC" box="[987,1065,1420,1438]" gridcol="1" gridrow="25" pageId="12" pageNumber="13">0.215</td>
<td id="2F7D9B567C6BCAAFA8E8E8F7FB195BFC" box="[1153,1242,1420,1438]" gridcol="2" gridrow="25" pageId="12" pageNumber="13">0.09364</td>
<td id="2F7D9B567C6BCAAFA944E8F7FA455BFC" box="[1325,1414,1420,1438]" gridcol="3" gridrow="25" pageId="12" pageNumber="13">-0.02694</td>
</tr>
<tr id="6CACF22A7C6BCAAFAD9EE8C2FA455BA9" box="[503,1414,1465,1483]" gridrow="26" pageId="12" pageNumber="13">
<th id="2F7D9B567C6BCAAFAD9EE8C2FCFD5BA9" box="[503,830,1465,1483]" gridcol="0" gridrow="26" pageId="12" pageNumber="13">Pelvic fin distance</th>
<td id="2F7D9B567C6BCAAFAFB2E8C2FBEA5BA9" box="[987,1065,1465,1483]" gridcol="1" gridrow="26" pageId="12" pageNumber="13">0.1971</td>
<td id="2F7D9B567C6BCAAFA8E8E8C2FB195BA9" box="[1153,1242,1465,1483]" gridcol="2" gridrow="26" pageId="12" pageNumber="13">0.08527</td>
<td id="2F7D9B567C6BCAAFA944E8C2FA455BA9" box="[1325,1414,1465,1483]" gridcol="3" gridrow="26" pageId="12" pageNumber="13">-0.1839</td>
</tr>
<tr id="6CACF22A7C6BCAAFAD9EE89EFA455B95" box="[503,1414,1509,1527]" gridrow="27" pageId="12" pageNumber="13">
<th id="2F7D9B567C6BCAAFAD9EE89EFCFD5B95" box="[503,830,1509,1527]" gridcol="0" gridrow="27" pageId="12" pageNumber="13">Pelvic fin to anus distance</th>
<td id="2F7D9B567C6BCAAFAFB2E89EFBEA5B95" box="[987,1065,1509,1527]" gridcol="1" gridrow="27" pageId="12" pageNumber="13">0.2148</td>
<td id="2F7D9B567C6BCAAFA8E8E89EFB195B95" box="[1153,1242,1509,1527]" gridcol="2" gridrow="27" pageId="12" pageNumber="13">-0.08011</td>
<td id="2F7D9B567C6BCAAFA944E89EFA455B95" box="[1325,1414,1509,1527]" gridcol="3" gridrow="27" pageId="12" pageNumber="13">-0.1393</td>
</tr>
<tr id="6CACF22A7C6BCAAFAD9EEB69FA455846" box="[503,1414,1554,1572]" gridrow="28" pageId="12" pageNumber="13">
<th id="2F7D9B567C6BCAAFAD9EEB69FCFD5846" box="[503,830,1554,1572]" gridcol="0" gridrow="28" pageId="12" pageNumber="13">Anus to anal fin distance</th>
<td id="2F7D9B567C6BCAAFAFB2EB69FBEA5846" box="[987,1065,1554,1572]" gridcol="1" gridrow="28" pageId="12" pageNumber="13">0.1499</td>
<td id="2F7D9B567C6BCAAFA8E8EB69FB195846" box="[1153,1242,1554,1572]" gridcol="2" gridrow="28" pageId="12" pageNumber="13">0.3313</td>
<td id="2F7D9B567C6BCAAFA944EB69FA455846" box="[1325,1414,1554,1572]" gridcol="3" gridrow="28" pageId="12" pageNumber="13">0.4677</td>
</tr>
<tr id="6CACF22A7C6BCAAFAD9EEB45FA455832" box="[503,1414,1598,1616]" gridrow="29" pageId="12" pageNumber="13">
<th id="2F7D9B567C6BCAAFAD9EEB45FCFD5832" box="[503,830,1598,1616]" gridcol="0" gridrow="29" pageId="12" pageNumber="13">Eigenvalue</th>
<td id="2F7D9B567C6BCAAFAFB2EB45FBEA5832" box="[987,1065,1598,1616]" gridcol="1" gridrow="29" pageId="12" pageNumber="13">0.333129</td>
<td id="2F7D9B567C6BCAAFA8E8EB45FB195832" box="[1153,1242,1598,1616]" gridcol="2" gridrow="29" pageId="12" pageNumber="13">0.0280641</td>
<td id="2F7D9B567C6BCAAFA944EB45FA455832" box="[1325,1414,1598,1616]" gridcol="3" gridrow="29" pageId="12" pageNumber="13">0.0112928</td>
</tr>
<tr id="6CACF22A7C6BCAAFAD9EEB10FA45581F" box="[503,1414,1643,1661]" gridrow="30" pageId="12" pageNumber="13">
<th id="2F7D9B567C6BCAAFAD9EEB10FCFD581F" box="[503,830,1643,1661]" gridcol="0" gridrow="30" pageId="12" pageNumber="13">% variance</th>
<td id="2F7D9B567C6BCAAFAFB2EB10FBEA581F" box="[987,1065,1643,1661]" gridcol="1" gridrow="30" pageId="12" pageNumber="13">83.209</td>
<td id="2F7D9B567C6BCAAFA8E8EB10FB19581F" box="[1153,1242,1643,1661]" gridcol="2" gridrow="30" pageId="12" pageNumber="13">7.0099</td>
<td id="2F7D9B567C6BCAAFA944EB10FA45581F" box="[1325,1414,1643,1661]" gridcol="3" gridrow="30" pageId="12" pageNumber="13">2.8207</td>
</tr>
</table>
</paragraph>
<caption id="86E3A0E07C6A355DAD81E967FAF25A37" ID-DOI="http://doi.org/10.5281/zenodo.11124778" ID-Zenodo-Dep="11124778" httpUri="https://zenodo.org/record/11124778/files/figure.png" pageId="13" pageNumber="14" startId="13.[488,561,1052,1072]" targetBox="[504,1453,177,1014]" targetPageId="13" targetType="figure">
<paragraph id="D223F0687C6A355DAD81E967FAF25A37" blockId="13.[488,1458,1052,1109]" pageId="13" pageNumber="14">
<emphasis id="E0E82C7A7C6A355DAD81E967FD905A52" bold="true" box="[488,595,1052,1072]" pageId="13" pageNumber="14">FIGURE 8 |</emphasis>
Second and third principal components axes of combined samples of
<taxonomicName id="159C8BEB7C6A355DA88FE967FA4B5A52" box="[1254,1416,1052,1072]" family="Pseudopimelodidae" genus="Rhyacoglanis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Siluriformes" pageId="13" pageNumber="14" phylum="Chordata" rank="species" species="varii">
<emphasis id="E0E82C7A7C6A355DA88FE967FA4B5A52" bold="true" box="[1254,1416,1052,1072]" italics="true" pageId="13" pageNumber="14">Rhyacoglanis varii</emphasis>
</taxonomicName>
(red cross, n = 16),
<taxonomicName id="159C8BEB7C6A355DAE01E93AFD1E5A37" authorityName="Shibatta &amp; Vari" authorityYear="2017" box="[616,733,1089,1109]" family="Pseudopimelodidae" genus="Rhyacoglanis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Siluriformes" pageId="13" pageNumber="14" phylum="Chordata" rank="species" species="paranensis">
<emphasis id="E0E82C7A7C6A355DAE01E93AFD1E5A37" bold="true" box="[616,733,1089,1109]" italics="true" pageId="13" pageNumber="14">R. paranensis</emphasis>
</taxonomicName>
(blue square; n = 31), and
<taxonomicName id="159C8BEB7C6A355DAFA4E93AFBA85A37" authorityName="Shibatta, Rocha &amp; Oliveira" authorityYear="2021" box="[973,1131,1089,1109]" family="Pseudopimelodidae" genus="Rhyacoglanis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Siluriformes" pageId="13" pageNumber="14" phylum="Chordata" rank="species" species="rapppydanielae">
<emphasis id="E0E82C7A7C6A355DAFA4E93AFBA85A37" bold="true" box="[973,1131,1089,1109]" italics="true" pageId="13" pageNumber="14">R. rapppydanielae</emphasis>
</taxonomicName>
(pink square; n = 13).
</paragraph>
</caption>
<paragraph id="D223F0687C6A355DAE6FE9D5FB935979" blockId="13.[488,1459,1198,1819]" pageId="13" pageNumber="14">
<emphasis id="E0E82C7A7C6A355DAE6FE9D5FCD85AA4" bold="true" box="[518,795,1198,1222]" pageId="13" pageNumber="14">DNA barcode analysis.</emphasis>
The Kimura 2-parameters genetic distance (K2P) between
<taxonomicName id="159C8BEB7C6A355DAD81E9A8FD6E5A89" box="[488,685,1235,1259]" family="Pseudopimelodidae" genus="Rhyacoglanis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Siluriformes" pageId="13" pageNumber="14" phylum="Chordata" rank="species" species="varii">
<emphasis id="E0E82C7A7C6A355DAD81E9A8FD6E5A89" box="[488,685,1235,1259]" italics="true" pageId="13" pageNumber="14">Rhyacoglanis varii</emphasis>
</taxonomicName>
and its congeners ranged from 1.9% (
<taxonomicName id="159C8BEB7C6A355DA8EAE9AFFAD05A89" authorityName="Shibatta &amp; Vari" authorityYear="2017" box="[1155,1299,1236,1259]" family="Pseudopimelodidae" genus="Rhyacoglanis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Siluriformes" pageId="13" pageNumber="14" phylum="Chordata" rank="species" species="paranensis">
<emphasis id="E0E82C7A7C6A355DA8EAE9AFFAD05A89" box="[1155,1299,1236,1259]" italics="true" pageId="13" pageNumber="14">R. paranensis</emphasis>
</taxonomicName>
) to 3.4% (
<taxonomicName id="159C8BEB7C6A355DA9F2E9AFFD8E5B72" authorityName="Shibatta &amp; Vari" authorityYear="2017" family="Pseudopimelodidae" genus="Rhyacoglanis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Siluriformes" pageId="13" pageNumber="14" phylum="Chordata" rank="species" species="annulatus">
<emphasis id="E0E82C7A7C6A355DA9F2E9AFFD8E5B72" italics="true" pageId="13" pageNumber="14">R. annulatus</emphasis>
</taxonomicName>
), with low intraspecific variation (
<tableCitation id="9F1EC5D37C6A355DAF8BE983FBF25B73" box="[994,1073,1272,1297]" captionStart="TABLE 3" captionStartId="15.[488,550,193,213]" captionTargetPageId="15" captionText="TABLE 3 | Interspecific and intraspecific genetic distances are based on the mitochondrial gene cytochrome c oxidase I (COI). Values are shown as proportions. Intraspecific distances are in bold." httpUri="http://table.plazi.org/id/86E3A0E07C68355FAD81EDBAFA465E99" pageId="13" pageNumber="14" tableUuid="86E3A0E07C68355FAD81EDBAFA465E99">Tab. 3</tableCitation>
). Based on COI gene sequences, species delimitation methods identified 7 MOTUs, corroborating
<taxonomicName id="159C8BEB7C6A355DA885E864FAFD5B54" box="[1260,1342,1311,1334]" family="Pseudopimelodidae" genus="Rhyacoglanis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Siluriformes" pageId="13" pageNumber="14" phylum="Chordata" rank="species" species="varii">
<emphasis id="E0E82C7A7C6A355DA885E864FAFD5B54" box="[1260,1342,1311,1334]" italics="true" pageId="13" pageNumber="14">R. varii</emphasis>
</taxonomicName>
as a new species.
<collectionCode id="B48D68AD7C6A355DAE2FE838FD5E5B39" box="[582,669,1347,1371]" pageId="13" pageNumber="14">GMYC</collectionCode>
results indicated seven species with a confidence interval of 5 to 8 species; the maximum likelihood of the null model = 169.5058, the maximum likelihood of the
<collectionCode id="B48D68AD7C6A355DAE5FE8F5FD4E5BC4" box="[566,653,1422,1446]" pageId="13" pageNumber="14">GMYC</collectionCode>
model = 175.6366, and the threshold time = -0.0034.
<collectionCode id="B48D68AD7C6A355DA96DE8F5FA975BC4" box="[1284,1364,1422,1446]" pageId="13" pageNumber="14">ABGD</collectionCode>
analysis resulted in 7 partitions where partition one (maximum previous distance p = 0.001) indicated 7 MOTUs.
<collectionCode id="B48D68AD7C6A355DAEB6E8A2FCE15B93" box="[735,802,1497,1521]" pageId="13" pageNumber="14">ASAP</collectionCode>
analysis retrieved nine partitions, with the lowest scoring partition (2.5) finding 7 MOTUs. The maximum likelihood solution of bPTP delimited 7 MOTUs and suggested a clear separation between the analyzed species. In the Bayesian inference analysis,
<taxonomicName id="159C8BEB7C6A355DAF4BEB32FCAC5802" box="[802,879,1609,1632]" family="Pseudopimelodidae" genus="Rhyacoglanis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Siluriformes" pageId="13" pageNumber="14" phylum="Chordata" rank="species" species="varii">
<emphasis id="E0E82C7A7C6A355DAF4BEB32FCAC5802" box="[802,879,1609,1632]" italics="true" pageId="13" pageNumber="14">R. varii</emphasis>
</taxonomicName>
was closely related to
<taxonomicName id="159C8BEB7C6A355DA801EB32FB315802" authorityName="Shibatta &amp; Vari" authorityYear="2017" box="[1128,1266,1609,1632]" family="Pseudopimelodidae" genus="Rhyacoglanis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Siluriformes" pageId="13" pageNumber="14" phylum="Chordata" rank="species" species="paranensis">
<emphasis id="E0E82C7A7C6A355DA801EB32FB315802" box="[1128,1266,1609,1632]" italics="true" pageId="13" pageNumber="14">R. paranensis</emphasis>
</taxonomicName>
, and
<taxonomicName id="159C8BEB7C6A355DA947EB32FA705802" authorityName="Shibatta &amp; Vari" authorityYear="2017" box="[1326,1459,1608,1632]" family="Pseudopimelodidae" genus="Rhyacoglanis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Siluriformes" pageId="13" pageNumber="14" phylum="Chordata" rank="species" species="annulatus">
<emphasis id="E0E82C7A7C6A355DA947EB32FA705802" box="[1326,1459,1608,1632]" italics="true" pageId="13" pageNumber="14">R. annulatus</emphasis>
</taxonomicName>
was the sister group of these two species.
<taxonomicName id="159C8BEB7C6A355DAFD4EB15FBAE58E7" box="[957,1133,1646,1670]" family="Pseudopimelodidae" genus="Rhyacoglanis" kingdom="Animalia" order="Siluriformes" pageId="13" pageNumber="14" phylum="Chordata" rank="species" species="undetermined">
<emphasis id="E0E82C7A7C6A355DAFD4EB15FB8458E4" box="[957,1095,1646,1670]" italics="true" pageId="13" pageNumber="14">Rhyacoglanis</emphasis>
sp.
</taxonomicName>
from the upper rio Araguaia basin (
<figureCitation id="4AA7ECED7C6A355DAE59EBE8FDB258C9" box="[560,625,1683,1707]" captionStart="FIGURE 9" captionStartId="14.[488,561,454,474]" captionTargetBox="[488,1459,185,432]" captionTargetId="figure-82@14.[488,1459,185,432]" captionTargetPageId="14" captionText="FIGURE 9 | Lateral view of sequenced Rhyacoglanis sp., LBP 1685, 21.6 mm SL, from Barra do Garças, state of Goiás, upper rio Araguaia basin." figureDoi="http://doi.org/10.5281/zenodo.11124780" httpUri="https://zenodo.org/record/11124780/files/figure.png" pageId="13" pageNumber="14">Fig. 9</figureCitation>
) was the basal species of the clade and was not closely related to
<taxonomicName id="159C8BEB7C6A355DA951EBEFFA4658C9" box="[1336,1413,1684,1707]" family="Pseudopimelodidae" genus="Rhyacoglanis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Siluriformes" pageId="13" pageNumber="14" phylum="Chordata" rank="species" species="varii">
<emphasis id="E0E82C7A7C6A355DA951EBEFFA4658C9" box="[1336,1413,1684,1707]" italics="true" pageId="13" pageNumber="14">R. varii</emphasis>
</taxonomicName>
. All
<taxonomicName id="159C8BEB7C6A355DAD81EBC3FD0458B2" box="[488,711,1720,1744]" pageId="13" pageNumber="14">
<emphasis id="E0E82C7A7C6A355DAD81EBC3FDB158B2" box="[488,626,1720,1744]" italics="true" pageId="13" pageNumber="14">Rhyacoglanis</emphasis>
species
</taxonomicName>
clustered in a clade with high posterior probability support (
<figureCitation id="4AA7ECED7C6A355DA9EFEBC3FDC65894" captionStart="FIGURE 10" captionStartId="14.[136,209,1724,1744]" captionTargetBox="[153,1438,659,1652]" captionTargetId="figure-21@14.[136,1459,619,1697]" captionTargetPageId="14" captionText="FIGURE 10 | Pseudopimelodidae species delimitation with Bayesian GMYC analysis. The cluster of supported species is indicated in red, and the Rhyacoglanis cluster is highlighted in yellow ocher. The numbers on the nodes represent the posterior probability, and the branch length represents the time in My. Genbank accession number in parenthesis; * = specimens identified as Pseudopimelodus mangurus in Genbank." figureDoi="http://doi.org/10.5281/zenodo.11124782" httpUri="https://zenodo.org/record/11124782/files/figure.png" pageId="13" pageNumber="14">Fig. 10</figureCitation>
), corroborating the monophyly of the genus.
<taxonomicName id="159C8BEB7C6A355DA878EBA5FB005894" authorityName="Bleeker" authorityYear="1858" box="[1041,1219,1758,1782]" family="Pseudopimelodidae" genus="Pseudopimelodus" kingdom="Animalia" order="Siluriformes" pageId="13" pageNumber="14" phylum="Chordata" rank="genus">
<emphasis id="E0E82C7A7C6A355DA878EBA5FB005894" box="[1041,1219,1758,1782]" italics="true" pageId="13" pageNumber="14">Pseudopimelodus</emphasis>
</taxonomicName>
and
<taxonomicName id="159C8BEB7C6A355DA895EBA5FAB05894" authorityName="Ortega-Lara &amp; Lehmann" authorityYear="2006" box="[1276,1395,1758,1782]" family="Pseudopimelodidae" genus="Cruciglanis" kingdom="Animalia" order="Siluriformes" pageId="13" pageNumber="14" phylum="Chordata" rank="genus">
<emphasis id="E0E82C7A7C6A355DA895EBA5FAB05894" box="[1276,1395,1758,1782]" italics="true" pageId="13" pageNumber="14">Cruciglanis</emphasis>
</taxonomicName>
were more closely related to each other than to
<taxonomicName id="159C8BEB7C6A355DAFD6EA78FB8A5979" authorityName="Shibatta &amp; Vari" authorityYear="2017" box="[959,1097,1795,1819]" family="Pseudopimelodidae" genus="Rhyacoglanis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Siluriformes" pageId="13" pageNumber="14" phylum="Chordata" rank="genus">
<emphasis id="E0E82C7A7C6A355DAFD6EA78FB8A5979" box="[959,1097,1795,1819]" italics="true" pageId="13" pageNumber="14">Rhyacoglanis</emphasis>
</taxonomicName>
.
</paragraph>
<caption id="86E3A0E07C69355EAD81ECBDFCF25C62" ID-DOI="http://doi.org/10.5281/zenodo.11124780" ID-Zenodo-Dep="11124780" httpUri="https://zenodo.org/record/11124780/files/figure.png" pageId="14" pageNumber="15" startId="14.[488,561,454,474]" targetBox="[488,1459,185,432]" targetPageId="14" targetType="figure">
<paragraph id="D223F0687C69355EAD81ECBDFCF25C62" blockId="14.[488,1455,454,512]" pageId="14" pageNumber="15">
<emphasis id="E0E82C7A7C69355EAD81ECBDFD935FB8" bold="true" box="[488,592,454,474]" pageId="14" pageNumber="15">FIGURE 9 |</emphasis>
Lateral view of sequenced
<taxonomicName id="159C8BEB7C69355EAF20ECBCFC1E5FB9" box="[841,989,455,475]" family="Pseudopimelodidae" genus="Rhyacoglanis" kingdom="Animalia" order="Siluriformes" pageId="14" pageNumber="15" phylum="Chordata" rank="species" species="undetermined">
<emphasis id="E0E82C7A7C69355EAF20ECBCFC7F5FB9" bold="true" box="[841,956,455,475]" italics="true" pageId="14" pageNumber="15">Rhyacoglanis</emphasis>
sp.
</taxonomicName>
, LBP 1685, 21.6 mm SL, from Barra do Garças, state of Goiás, upper rio Araguaia basin.
</paragraph>
</caption>
<caption id="86E3A0E07C69355EACE1EBC7FA485979" ID-DOI="http://doi.org/10.5281/zenodo.11124782" ID-Zenodo-Dep="11124782" httpUri="https://zenodo.org/record/11124782/files/figure.png" pageId="14" pageNumber="15" startId="14.[136,209,1724,1744]" targetBox="[153,1438,659,1652]" targetPageId="14" targetType="figure">
<paragraph id="D223F0687C69355EACE1EBC7FA485979" blockId="14.[136,1459,1724,1819]" pageId="14" pageNumber="15">
<emphasis id="E0E82C7A7C69355EACE1EBC7FF3D58B2" bold="true" box="[136,254,1724,1744]" pageId="14" pageNumber="15">FIGURE 10 |</emphasis>
<taxonomicName id="159C8BEB7C69355EAD6AEBC7FE7858B2" authorityName="Fernandez-Yepez &amp; Anton" authorityYear="1966" box="[259,443,1724,1744]" family="Pseudopimelodidae" kingdom="Animalia" order="Siluriformes" pageId="14" pageNumber="15" phylum="Chordata" rank="family">Pseudopimelodidae</taxonomicName>
species delimitation with Bayesian GMYC analysis. The cluster of supported species is indicated in red, and the
<taxonomicName id="159C8BEB7C69355EACC5EB9AFEDC5897" authorityName="Shibatta &amp; Vari" authorityYear="2017" box="[172,287,1761,1781]" family="Pseudopimelodidae" genus="Rhyacoglanis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Siluriformes" pageId="14" pageNumber="15" phylum="Chordata" rank="genus">
<emphasis id="E0E82C7A7C69355EACC5EB9AFEDC5897" bold="true" box="[172,287,1761,1781]" italics="true" pageId="14" pageNumber="15">Rhyacoglanis</emphasis>
</taxonomicName>
cluster is highlighted in yellow ocher. The numbers on the nodes represent the posterior probability, and the branch length represents the time in My. Genbank accession number in parenthesis; * = specimens identified as
<taxonomicName id="159C8BEB7C69355EA849EA7CFAD15979" authorityName="Shibatta et al." authorityYear="2021" baseAuthorityName="Valenciennes" baseAuthorityYear="1835" box="[1056,1298,1799,1819]" family="Pseudopimelodidae" genus="Pseudopimelodus" kingdom="Animalia" order="Siluriformes" pageId="14" pageNumber="15" phylum="Chordata" rank="species" species="mangurus">
<emphasis id="E0E82C7A7C69355EA849EA7CFAD15979" bold="true" box="[1056,1298,1799,1819]" italics="true" pageId="14" pageNumber="15">Pseudopimelodus mangurus</emphasis>
</taxonomicName>
in Genbank.
</paragraph>
</caption>
<caption id="86E3A0E07C68355FAD81EDBAFA465E99" ID-Table-UUID="86E3A0E07C68355FAD81EDBAFA465E99" httpUri="http://table.plazi.org/id/86E3A0E07C68355FAD81EDBAFA465E99" pageId="15" pageNumber="16" startId="15.[488,550,193,213]" targetBox="[503,1434,291,627]" targetIsTable="true" targetPageId="15" targetType="table">
<paragraph id="D223F0687C68355FAD81EDBAFA465E99" blockId="15.[488,1459,193,251]" pageId="15" pageNumber="16">
<emphasis id="E0E82C7A7C68355FAD81EDBAFD965EB7" bold="true" box="[488,597,193,213]" pageId="15" pageNumber="16">TABLE 3 |</emphasis>
Interspecific and intraspecific genetic distances are based on the mitochondrial gene cytochrome c oxidase I (COI). Values are shown as proportions. Intraspecific distances are in bold.
</paragraph>
</caption>
<paragraph id="D223F0687C68355FAD9EEC58FA595C11" pageId="15" pageNumber="16">
<table id="A09C02C87C68CAAFAD9EEC58FA595C11" box="[503,1434,291,627]" gridcols="8" gridrows="8" pageId="15" pageNumber="16">
<tr id="6CACF22A7C68CAAFAD9EEC58FA595F57" box="[503,1434,291,309]" gridrow="0" pageId="15" pageNumber="16">
<th id="2F7D9B567C68CAAFAD9EEC58FCC25F57" box="[503,769,291,309]" gridcol="0" gridrow="0" pageId="15" pageNumber="16">
<emphasis id="E0E82C7A7C68355FAD9EEC58FDFE5F57" bold="true" box="[503,573,291,309]" pageId="15" pageNumber="16">Species</emphasis>
</th>
<th id="2F7D9B567C68CAAFAF42EC58FC995F57" box="[811,858,291,309]" gridcol="1" gridrow="0" pageId="15" pageNumber="16">
<emphasis id="E0E82C7A7C68355FAF54EC58FC845F57" bold="true" box="[829,839,291,309]" pageId="15" pageNumber="16">1</emphasis>
</th>
<th id="2F7D9B567C68CAAFAFE2EC58FC795F57" box="[907,954,291,309]" gridcol="2" gridrow="0" pageId="15" pageNumber="16">
<emphasis id="E0E82C7A7C68355FAFF4EC58FC645F57" bold="true" box="[925,935,291,309]" pageId="15" pageNumber="16">2</emphasis>
</th>
<th id="2F7D9B567C68CAAFAF82EC58FBD95F57" box="[1003,1050,291,309]" gridcol="3" gridrow="0" pageId="15" pageNumber="16">
<emphasis id="E0E82C7A7C68355FAF94EC58FBC45F57" bold="true" box="[1021,1031,291,309]" pageId="15" pageNumber="16">3</emphasis>
</th>
<th id="2F7D9B567C68CAAFA822EC58FBB95F57" box="[1099,1146,291,309]" gridcol="4" gridrow="0" pageId="15" pageNumber="16">
<emphasis id="E0E82C7A7C68355FA834EC58FBA45F57" bold="true" box="[1117,1127,291,309]" pageId="15" pageNumber="16">4</emphasis>
</th>
<th id="2F7D9B567C68CAAFA8C2EC58FB195F57" box="[1195,1242,291,309]" gridcol="5" gridrow="0" pageId="15" pageNumber="16">
<emphasis id="E0E82C7A7C68355FA8D4EC5FFB045F56" bold="true" box="[1213,1223,292,308]" pageId="15" pageNumber="16">5</emphasis>
</th>
<th id="2F7D9B567C68CAAFA962EC58FAF95F57" box="[1291,1338,291,309]" gridcol="6" gridrow="0" pageId="15" pageNumber="16">
<emphasis id="E0E82C7A7C68355FA977EC58FAEB5F57" bold="true" box="[1310,1320,291,309]" pageId="15" pageNumber="16">6</emphasis>
</th>
<th id="2F7D9B567C68CAAFA902EC58FA595F57" box="[1387,1434,291,309]" gridcol="7" gridrow="0" pageId="15" pageNumber="16">
<emphasis id="E0E82C7A7C68355FA917EC5FFA4B5F56" bold="true" box="[1406,1416,292,308]" pageId="15" pageNumber="16">7</emphasis>
</th>
</tr>
<tr id="6CACF22A7C68CAAFAD9EEC2EFA595F05" box="[503,1434,341,359]" gridrow="1" pageId="15" pageNumber="16" rowspan-2="1" rowspan-3="1" rowspan-4="1" rowspan-5="1" rowspan-6="1" rowspan-7="1">
<th id="2F7D9B567C68CAAFAD9EEC2EFCC25F05" box="[503,769,341,359]" gridcol="0" gridrow="1" pageId="15" pageNumber="16">
1.
<taxonomicName id="159C8BEB7C68355FAE62EC2EFD6E5F05" box="[523,685,341,359]" family="Pseudopimelodidae" genus="Rhyacoglanis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Siluriformes" pageId="15" pageNumber="16" phylum="Chordata" rank="species" species="varii">
<emphasis id="E0E82C7A7C68355FAE62EC2EFD6E5F05" box="[523,685,341,359]" italics="true" pageId="15" pageNumber="16">Rhyacoglanis varii</emphasis>
</taxonomicName>
</th>
<td id="2F7D9B567C68CAAFAF42EC2EFC995F05" box="[811,858,341,359]" gridcol="1" gridrow="1" pageId="15" pageNumber="16">
<emphasis id="E0E82C7A7C68355FAF42EC2EFC995F05" bold="true" box="[811,858,341,359]" pageId="15" pageNumber="16">0.000</emphasis>
</td>
</tr>
<tr id="6CACF22A7C68CAAFAD9EECF9FA595FF6" box="[503,1434,386,404]" gridrow="2" pageId="15" pageNumber="16" rowspan-3="1" rowspan-4="1" rowspan-5="1" rowspan-6="1" rowspan-7="1">
<th id="2F7D9B567C68CAAFAD9EECF9FCC25FF6" box="[503,769,386,404]" gridcol="0" gridrow="2" pageId="15" pageNumber="16">
2.
<taxonomicName id="159C8BEB7C68355FAE62ECF9FD5D5FF1" box="[523,670,386,404]" family="Pseudopimelodidae" genus="Rhyacoglanis" kingdom="Animalia" order="Siluriformes" pageId="15" pageNumber="16" phylum="Chordata" rank="species" species="undetermined">
<emphasis id="E0E82C7A7C68355FAE62ECF9FDBC5FF6" box="[523,639,386,404]" italics="true" pageId="15" pageNumber="16">Rhyacoglanis</emphasis>
sp.
</taxonomicName>
(Araguaia)
</th>
<td id="2F7D9B567C68CAAFAF42ECF9FC995FF6" box="[811,858,386,404]" gridcol="1" gridrow="2" pageId="15" pageNumber="16">0.029</td>
<td id="2F7D9B567C68CAAFAFE2ECF9FC795FF6" box="[907,954,386,404]" gridcol="2" gridrow="2" pageId="15" pageNumber="16">
<emphasis id="E0E82C7A7C68355FAFF6ECF8FC665FF1" bold="true" box="[927,933,387,403]" pageId="15" pageNumber="16">-</emphasis>
</td>
</tr>
<tr id="6CACF22A7C68CAAFAD9EECD4FA595FA3" box="[503,1434,431,449]" gridrow="3" pageId="15" pageNumber="16" rowspan-4="1" rowspan-5="1" rowspan-6="1" rowspan-7="1">
<th id="2F7D9B567C68CAAFAD9EECD4FCC25FA3" box="[503,769,431,449]" gridcol="0" gridrow="3" pageId="15" pageNumber="16">
3.
<taxonomicName id="159C8BEB7C68355FAE62ECD4FD205FA2" authorityName="Shibatta &amp; Vari" authorityYear="2017" box="[523,739,431,449]" family="Pseudopimelodidae" genus="Rhyacoglanis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Siluriformes" pageId="15" pageNumber="16" phylum="Chordata" rank="species" species="paranensis">
<emphasis id="E0E82C7A7C68355FAE62ECD4FD205FA2" box="[523,739,431,449]" italics="true" pageId="15" pageNumber="16">Rhyacoglanis paranensis</emphasis>
</taxonomicName>
</th>
<td id="2F7D9B567C68CAAFAF42ECD4FC995FA3" box="[811,858,431,449]" gridcol="1" gridrow="3" pageId="15" pageNumber="16">0.019</td>
<td id="2F7D9B567C68CAAFAFE2ECD4FC795FA3" box="[907,954,431,449]" gridcol="2" gridrow="3" pageId="15" pageNumber="16">0.041</td>
<td id="2F7D9B567C68CAAFAF82ECD4FBD95FA3" box="[1003,1050,431,449]" gridcol="3" gridrow="3" pageId="15" pageNumber="16">
<emphasis id="E0E82C7A7C68355FAF82ECD4FBD95FA3" bold="true" box="[1003,1050,431,449]" pageId="15" pageNumber="16">0.000</emphasis>
</td>
</tr>
<tr id="6CACF22A7C68CAAFAD9EECA0FA595F8F" box="[503,1434,475,493]" gridrow="4" pageId="15" pageNumber="16" rowspan-5="1" rowspan-6="1" rowspan-7="1">
<th id="2F7D9B567C68CAAFAD9EECA0FCC25F8F" box="[503,769,475,493]" gridcol="0" gridrow="4" pageId="15" pageNumber="16">
4.
<taxonomicName id="159C8BEB7C68355FAE62ECA0FD1F5F8F" authorityName="Shibatta &amp; Vari" authorityYear="2017" box="[523,732,475,493]" family="Pseudopimelodidae" genus="Rhyacoglanis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Siluriformes" pageId="15" pageNumber="16" phylum="Chordata" rank="species" species="annulatus">
<emphasis id="E0E82C7A7C68355FAE62ECA0FD1F5F8F" box="[523,732,475,493]" italics="true" pageId="15" pageNumber="16">Rhyacoglanis annulatus</emphasis>
</taxonomicName>
</th>
<td id="2F7D9B567C68CAAFAF42ECA0FC995F8F" box="[811,858,475,493]" gridcol="1" gridrow="4" pageId="15" pageNumber="16">0.034</td>
<td id="2F7D9B567C68CAAFAFE2ECA0FC795F8F" box="[907,954,475,493]" gridcol="2" gridrow="4" pageId="15" pageNumber="16">0.060</td>
<td id="2F7D9B567C68CAAFAF82ECA0FBD95F8F" box="[1003,1050,475,493]" gridcol="3" gridrow="4" pageId="15" pageNumber="16">0.048</td>
<td id="2F7D9B567C68CAAFA822ECA0FBB95F8F" box="[1099,1146,475,493]" gridcol="4" gridrow="4" pageId="15" pageNumber="16">
<emphasis id="E0E82C7A7C68355FA822ECA0FBB95F8F" bold="true" box="[1099,1146,475,493]" pageId="15" pageNumber="16">0.003</emphasis>
</td>
</tr>
<tr id="6CACF22A7C68CAAFAD9EEF73FA595C78" box="[503,1434,520,538]" gridrow="5" pageId="15" pageNumber="16" rowspan-6="1" rowspan-7="1">
<th id="2F7D9B567C68CAAFAD9EEF73FCC25C78" box="[503,769,520,538]" gridcol="0" gridrow="5" pageId="15" pageNumber="16">
5.
<taxonomicName id="159C8BEB7C68355FAE62EF73FD715C78" box="[523,690,520,538]" family="Pseudopimelodidae" genus="Cruciglanis" kingdom="Animalia" order="Siluriformes" pageId="15" pageNumber="16" phylum="Chordata" rank="species" species="pacifici">
<emphasis id="E0E82C7A7C68355FAE62EF73FD715C78" box="[523,690,520,538]" italics="true" pageId="15" pageNumber="16">Cruciglanis pacifici</emphasis>
</taxonomicName>
</th>
<td id="2F7D9B567C68CAAFAF42EF73FC995C78" box="[811,858,520,538]" gridcol="1" gridrow="5" pageId="15" pageNumber="16">0.138</td>
<td id="2F7D9B567C68CAAFAFE2EF73FC795C78" box="[907,954,520,538]" gridcol="2" gridrow="5" pageId="15" pageNumber="16">0.130</td>
<td id="2F7D9B567C68CAAFAF82EF73FBD95C78" box="[1003,1050,520,538]" gridcol="3" gridrow="5" pageId="15" pageNumber="16">0.149</td>
<td id="2F7D9B567C68CAAFA822EF73FBB95C78" box="[1099,1146,520,538]" gridcol="4" gridrow="5" pageId="15" pageNumber="16">0.152</td>
<td id="2F7D9B567C68CAAFA8C2EF73FB195C78" box="[1195,1242,520,538]" gridcol="5" gridrow="5" pageId="15" pageNumber="16">
<emphasis id="E0E82C7A7C68355FA8C2EF73FB195C78" bold="true" box="[1195,1242,520,538]" pageId="15" pageNumber="16">0.006</emphasis>
</td>
</tr>
<tr id="6CACF22A7C68CAAFAD9EEF4FFA595C24" box="[503,1434,564,582]" gridrow="6" pageId="15" pageNumber="16" rowspan-7="1">
<th id="2F7D9B567C68CAAFAD9EEF4FFCC25C24" box="[503,769,564,582]" gridcol="0" gridrow="6" pageId="15" pageNumber="16">
6.
<taxonomicName id="159C8BEB7C68355FAE62EF4FFD305C24" baseAuthorityName="Valenciennes" baseAuthorityYear="1840" box="[523,755,564,582]" family="Pseudopimelodidae" genus="Pseudopimelodus" kingdom="Animalia" order="Siluriformes" pageId="15" pageNumber="16" phylum="Chordata" rank="species" species="bufonius">
<emphasis id="E0E82C7A7C68355FAE62EF4FFD305C24" box="[523,755,564,582]" italics="true" pageId="15" pageNumber="16">Pseudopimelodus bufonius</emphasis>
</taxonomicName>
</th>
<td id="2F7D9B567C68CAAFAF42EF4FFC995C24" box="[811,858,564,582]" gridcol="1" gridrow="6" pageId="15" pageNumber="16">0.093</td>
<td id="2F7D9B567C68CAAFAFE2EF4FFC795C24" box="[907,954,564,582]" gridcol="2" gridrow="6" pageId="15" pageNumber="16">0.101</td>
<td id="2F7D9B567C68CAAFAF82EF4FFBD95C24" box="[1003,1050,564,582]" gridcol="3" gridrow="6" pageId="15" pageNumber="16">0.102</td>
<td id="2F7D9B567C68CAAFA822EF4FFBB95C24" box="[1099,1146,564,582]" gridcol="4" gridrow="6" pageId="15" pageNumber="16">0.116</td>
<td id="2F7D9B567C68CAAFA8C2EF4FFB195C24" box="[1195,1242,564,582]" gridcol="5" gridrow="6" pageId="15" pageNumber="16">0.117</td>
<td id="2F7D9B567C68CAAFA962EF4FFAF95C24" box="[1291,1338,564,582]" gridcol="6" gridrow="6" pageId="15" pageNumber="16">
<emphasis id="E0E82C7A7C68355FA962EF4FFAF95C24" bold="true" box="[1291,1338,564,582]" pageId="15" pageNumber="16">0.003</emphasis>
</td>
</tr>
<tr id="6CACF22A7C68CAAFAD9EEF1AFA595C11" box="[503,1434,609,627]" gridrow="7" pageId="15" pageNumber="16">
<th id="2F7D9B567C68CAAFAD9EEF1AFCC25C11" box="[503,769,609,627]" gridcol="0" gridrow="7" pageId="15" pageNumber="16">
7.
<taxonomicName id="159C8BEB7C68355FAE62EF1AFCC25C10" authorityName="Shibatta et al." authorityYear="2021" baseAuthorityName="Valenciennes" baseAuthorityYear="1835" box="[523,769,609,627]" family="Pseudopimelodidae" genus="Pseudopimelodus" kingdom="Animalia" order="Siluriformes" pageId="15" pageNumber="16" phylum="Chordata" rank="species" species="mangurus">
<emphasis id="E0E82C7A7C68355FAE62EF1AFCC25C10" box="[523,769,609,627]" italics="true" pageId="15" pageNumber="16">Pseudopimelodus mangurus</emphasis>
</taxonomicName>
</th>
<td id="2F7D9B567C68CAAFAF42EF1AFC995C11" box="[811,858,609,627]" gridcol="1" gridrow="7" pageId="15" pageNumber="16">0.107</td>
<td id="2F7D9B567C68CAAFAFE2EF1AFC795C11" box="[907,954,609,627]" gridcol="2" gridrow="7" pageId="15" pageNumber="16">0.109</td>
<td id="2F7D9B567C68CAAFAF82EF1AFBD95C11" box="[1003,1050,609,627]" gridcol="3" gridrow="7" pageId="15" pageNumber="16">0.109</td>
<td id="2F7D9B567C68CAAFA822EF1AFBB95C11" box="[1099,1146,609,627]" gridcol="4" gridrow="7" pageId="15" pageNumber="16">0.118</td>
<td id="2F7D9B567C68CAAFA8C2EF1AFB195C11" box="[1195,1242,609,627]" gridcol="5" gridrow="7" pageId="15" pageNumber="16">0.109</td>
<td id="2F7D9B567C68CAAFA962EF1AFAF95C11" box="[1291,1338,609,627]" gridcol="6" gridrow="7" pageId="15" pageNumber="16">0.027</td>
<td id="2F7D9B567C68CAAFA902EF1AFA595C11" box="[1387,1434,609,627]" gridcol="7" gridrow="7" pageId="15" pageNumber="16">0.003</td>
</tr>
</table>
</paragraph>
<paragraph id="D223F0687C68355FAE6FEF95FCF15DA2" blockId="15.[488,1459,750,960]" pageId="15" pageNumber="16">
<taxonomicName id="159C8BEB7C68355FAE6FEF95FD0F5D64" box="[518,716,750,774]" family="Pseudopimelodidae" genus="Rhyacoglanis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Siluriformes" pageId="15" pageNumber="16" phylum="Chordata" rank="species" species="varii">
<emphasis id="E0E82C7A7C68355FAE6FEF95FD0F5D64" box="[518,716,750,774]" italics="true" pageId="15" pageNumber="16">Rhyacoglanis varii</emphasis>
</taxonomicName>
was recovered from different methods of species delimitation using molecular analysis. The results of those methods are presented as supplementary material:
<figureCitation id="4AA7ECED7C68355FAE3AEE43FD655D32" box="[595,678,824,848]" captionStart="FIGURE 1" captionStartId="4.[488,561,1761,1781]" captionTargetBox="[488,1459,432,1737]" captionTargetId="figure-20@4.[488,1459,432,1737]" captionTargetPageId="4" captionText="FIGURE 1 | Holotype of Rhyacoglanis varii, MZUEL 22317, 67.1 mm SL, rio do Peixe, upper rio Tocantins basin, Goianésia, state of Goiás, Brazil." figureDoi="http://doi.org/10.5281/zenodo.11124760" httpUri="https://zenodo.org/record/11124760/files/figure.png" pageId="15" pageNumber="16">
Fig.
<emphasis id="E0E82C7A7C68355FAEE3EE43FD655D32" bold="true" box="[650,678,824,848]" pageId="15" pageNumber="16">S1</emphasis>
</figureCitation>
, Neighbor-joining tree;
<figureCitation id="4AA7ECED7C68355FAFA2EE43FBDD5D32" box="[971,1054,824,848]" captionStart="FIGURE 2" captionStartId="5.[136,209,1873,1893]" captionTargetBox="[147,1436,1497,1804]" captionTargetId="figure-405@5.[136,1459,1471,1821]" captionTargetPageId="5" captionText="FIGURE 2 | Rhyacoglanis varii pectoral-fin spine, paratype, MZUEL 6039, 55.5 mm SL. Scale bar = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.11124762" httpUri="https://zenodo.org/record/11124762/files/figure.png" pageId="15" pageNumber="16">
Fig.
<emphasis id="E0E82C7A7C68355FA86BEE43FBDD5D32" bold="true" box="[1026,1054,824,848]" pageId="15" pageNumber="16">S2</emphasis>
</figureCitation>
, Number of groups obtained with
<collectionCode id="B48D68AD7C68355FAD81EE25FDFB5D14" box="[488,568,862,886]" pageId="15" pageNumber="16">ABGD</collectionCode>
method;
<figureCitation id="4AA7ECED7C68355FAEC3EE25FD3E5D14" box="[682,765,862,886]" captionStart="FIGURE 3" captionStartId="6.[136,209,1686,1706]" captionTargetBox="[178,1437,733,1622]" captionTargetId="figure-21@6.[138,1459,733,1664]" captionTargetPageId="6" captionText="FIGURE 3 | Rhyacoglanis varii caudal-fin skeleton, paratype, MZUEL 6039, 53.5 mm SL. Bones in yellow ocher and cartilage in blue. H1 to H5 = hypurals 1 to 5. Scale bar = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.11124764" httpUri="https://zenodo.org/record/11124764/files/figure.png" pageId="15" pageNumber="16">
Fig.
<emphasis id="E0E82C7A7C68355FAE88EE25FD3E5D14" bold="true" box="[737,765,862,886]" pageId="15" pageNumber="16">S3</emphasis>
</figureCitation>
, Number of groups obtained with
<collectionCode id="B48D68AD7C68355FA8F5EE25FB1C5D14" box="[1180,1247,862,886]" pageId="15" pageNumber="16">ASAP</collectionCode>
method; Tab.
<emphasis id="E0E82C7A7C68355FA9F9EE25FA6E5D14" bold="true" box="[1424,1453,862,886]" pageId="15" pageNumber="16">S4</emphasis>
,
<collectionCode id="B48D68AD7C68355FAD81EEF8FDFC5DF9" box="[488,575,899,923]" pageId="15" pageNumber="16">GMYC</collectionCode>
method, and
<figureCitation id="4AA7ECED7C68355FAE9BEEF8FC8A5DF9" box="[754,841,899,923]" captionStart="FIGURE 5" captionStartId="9.[136,209,1649,1669]" captionTargetBox="[136,1459,966,1627]" captionTargetId="figure-235@9.[136,1459,966,1627]" captionTargetPageId="9" captionText="FIGURE 5 | Rhyacoglanis varii coloration in life, MZUEL 6039, paratype, 46.1 mm SL, rio do Peixe, upper rio Tocantins basin, Goianésia, state of Goiás, Brazil." figureDoi="http://doi.org/10.5281/zenodo.11124768" httpUri="https://zenodo.org/record/11124768/files/figure.png" pageId="15" pageNumber="16">
Fig.
<emphasis id="E0E82C7A7C68355FAF44EEF8FC8A5DF9" bold="true" box="[813,841,899,923]" pageId="15" pageNumber="16">S5</emphasis>
</figureCitation>
, Phylogeny of analyzed
<taxonomicName id="159C8BEB7C68355FA816EEF8FA9B5DF9" authorityName="Fernandez-Yepez &amp; Anton" authorityYear="1966" box="[1151,1368,899,923]" family="Pseudopimelodidae" kingdom="Animalia" order="Siluriformes" pageId="15" pageNumber="16" phylum="Chordata" rank="family">Pseudopimelodidae</taxonomicName>
species obtained with bPTP method.
</paragraph>
</subSubSection>
</treatment>
</document>