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<mods:title id="1F66C4AC987EFDE8C79C96C6E699385C">Taxonomic Revision of the Spider Genus Actinopus Perty, 1833 (Araneae, Mygalomorphae, Actinopodidae)</mods:title>
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<mods:namePart id="AE0ECB785B10AA754CA40802DFA4ACEC">Miglio, Laura Tavares</mods:namePart>
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<mods:affiliation id="54754CE1321F36C0A434866610ABE954">Museu Paraense Emílio Goeldi, Coordenação de Zoologia, Laboratório de Aracnologia, Av. Perimetral, 1901, Terra Firme, Cep: 66077 - 830, Belém, Pará, Brazil. https: // orcid. org / 0000 - 0002 - 9101 - 8657,</mods:affiliation>
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<mods:affiliation id="E031F90AA7D14A8546EA8F0739D6B4D9">Facultad de Ciencias, Sección Entomología, Iguá 4225, 11400, Montevideo, Uruguay. ® myga @ fcien. edu. uy, https: // orcid. org / 0000 - 0002 - 4367 - 6276</mods:affiliation>
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<mods:namePart id="3E80E5B39F686BC206C6E87D79E9451A">Bonaldo, Alexandre B.</mods:namePart>
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<emphasis id="B3A87E6EFF69CEB9FCD5FE82DF503D43" bold="true" box="[810,1027,338,365]" italics="true" pageId="228" pageNumber="229">Actinopus clavero</emphasis>
<bibRefCitation id="E54DDF8DFF69CEB9FBF4FE83DEDC3D43" author="Rios-Tamayo, D. &amp; Goloboff, P. A." box="[1035,1423,338,365]" pageId="228" pageNumber="229" pagination="1 - 83" refId="ref140496" refString="Rios-Tamayo, D. &amp; Goloboff, P. A. (2018) Taxonomic revision and morphology of the trapdoor spider genus Actinopus (Mygalomorphae: Actinopodidae) in Argentina. Bulletin of the American Museum of Natural History, 419, 1 - 83. https: // doi. org / 10.1206 / 0003 - 0090 - 419.1.1" type="journal article" year="2018">Ríos-Tamayo &amp; Goloboff, 2018</bibRefCitation>
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<figureCitation id="19E7BEF9FF69CEB9FCD5FEA6D8953DBF" box="[810,966,375,401]" captionStart-0="FIGURE 207" captionStart-1="FIGURE 208" captionStart-2="FIGURE 209" captionStartId-0="229.[136,235,1645,1669]" captionStartId-1="230.[136,235,1155,1179]" captionStartId-2="231.[136,235,1452,1476]" captionTargetBox-0="[171,1407,205,1604]" captionTargetBox-1="[180,1407,206,1115]" captionTargetBox-2="[175,1434,193,1409]" captionTargetId-0="figure-134@229.[151,1435,189,1621]" captionTargetId-1="figure-393@230.[151,1435,189,1132]" captionTargetId-2="figure-186@231.[151,1435,189,1429]" captionTargetPageId-0="229" captionTargetPageId-1="230" captionTargetPageId-2="231" captionText-0="FIGURE 207. Actinopus clavero Ríos-Tamayo &amp; Goloboff, 2018, male:AF MACN-Ar 19827:A.Abdomen, dorsal; B. Carapace, dorsal; C. Sternum, ventral; D. Cheliceral teeth, ventral; E. Rastellum, dorsal; F. Rastellum, ventral." captionText-1="FIGURE 208. Actinopus clavero Ríos-Tamayo &amp; Goloboff, 2018, male:AD MACN-Ar 19827:A. Patella III, dorsal; B. Tibia III, dorsal; C. Patella IV, dorsal; D. Tibia IV, dorsal." captionText-2="FIGURE 209. Actinopus clavero Ríos-Tamayo &amp; Goloboff, 2018, male:AC MACN-Ar 19827:A. Copulatory bulb, prolateral; B. Copulatory bulb, dorsal; C. Copulatory bulb, retrolateral. PA-Paraembolic apophysis; BTA-Basal Tegular Apophysis; PI-Prolateral Inferior keel; PS-Prolateral Superior keel. Scale line: 1mm." figureDoi-0="http://doi.org/10.5281/zenodo.5597689" figureDoi-1="http://doi.org/10.5281/zenodo.5597691" figureDoi-2="http://doi.org/10.5281/zenodo.5597693" httpUri-0="https://zenodo.org/record/5597689/files/figure.png" httpUri-1="https://zenodo.org/record/5597691/files/figure.png" httpUri-2="https://zenodo.org/record/5597693/files/figure.png" pageId="228" pageNumber="229">Figs 207209</figureCitation>
,
<figureCitation id="19E7BEF9FF69CEB9FC2EFEA9DF743DBC" box="[977,1063,375,402]" captionStart="MAP 15" captionStartId="228.[136,192,2053,2077]" captionTargetBox="[157,1428,1127,2021]" captionTargetId="figure-458@228.[151,1435,1121,2029]" captionText="MAP 15. Distribution map of Actinopus clavero Ríos-Tamayo &amp; Goloboff, 2018." figureDoi="http://doi.org/10.5281/zenodo.5655464" httpUri="https://zenodo.org/record/5655464/files/figure.png" pageId="228" pageNumber="229">Map 15</figureCitation>
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<emphasis id="B3A87E6EFF69CEB9FCD5FE6ED8843DFD" box="[810,983,446,468]" italics="true" pageId="228" pageNumber="229">Actinopus clavero</emphasis>
<bibRefCitation id="E54DDF8DFF69CEB9FC23FE6FDE653DFA" author="Rios-Tamayo, D. &amp; Goloboff, P. A." box="[988,1334,446,468]" pageId="228" pageNumber="229" pagination="1 - 83" refId="ref140496" refString="Rios-Tamayo, D. &amp; Goloboff, P. A. (2018) Taxonomic revision and morphology of the trapdoor spider genus Actinopus (Mygalomorphae: Actinopodidae) in Argentina. Bulletin of the American Museum of Natural History, 419, 1 - 83. https: // doi. org / 10.1206 / 0003 - 0090 - 419.1.1" type="journal article" year="2018">Ríos-Tamayo &amp; Goloboff, 2018: 14</bibRefCitation>
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, fig. 4 AH, 5 AD, 43 (
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♂, [
<geoCoordinate id="E4E8C4BBFF69CEB9FBB7FE0FDFCD3DDD" box="[1096,1182,478,500]" degrees="31" direction="south" minutes="46" orientation="latitude" pageId="228" pageNumber="229" precision="925" value="-31.766666">31º 46 S</geoCoordinate>
<geoCoordinate id="E4E8C4BBFF69CEB9FB5CFE0FDE573DDA" box="[1187,1284,478,500]" degrees="64" direction="west" minutes="59" orientation="longitude" pageId="228" pageNumber="229" precision="925" value="-64.98333">64º 59 W</geoCoordinate>
], Nono, Córdoba,
<collectingCountry id="F9CBE2ECFF69CEB9FCA0FE2FD8923E3A" box="[863,961,510,532]" name="Argentina" pageId="228" pageNumber="229">Argentina</collectingCountry>
,
<date id="F56284BCFF69CEB9FC36FE2FDF5A3E3A" box="[969,1033,510,532]" pageId="228" pageNumber="229" value="1998-01">i.1998</date>
, P. Goloboff and C. Szumik leg., MACN-Ar 36533;
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, [
<geoCoordinate id="E4E8C4BBFF69CEB9FBD1FDCFDFDE3E1D" box="[1070,1165,542,564]" degrees="31" direction="south" minutes="36" orientation="latitude" pageId="228" pageNumber="229" precision="925" value="-31.6">31º 36 S</geoCoordinate>
<geoCoordinate id="E4E8C4BBFF69CEB9FB68FDCFDE523E1D" box="[1175,1281,541,564]" degrees="64" direction="west" minutes="54" orientation="longitude" pageId="228" pageNumber="229" precision="925" value="-64.9">64º 54 W</geoCoordinate>
], Camino a Villa Carlos Paz,
<quantity id="46240F99FF69CEB9FC20FDEFDF453E7A" box="[991,1046,574,596]" metricMagnitude="4" metricUnit="m" metricValue="1.5" pageId="228" pageNumber="229" unit="km" value="15.0">15km</quantity>
de Mina Clavero, Córdoba,
<collectingCountry id="F9CBE2ECFF69CEB9FAB9FDEFDEFB3E7A" box="[1350,1448,574,596]" name="Argentina" pageId="228" pageNumber="229">Argentina</collectingCountry>
,
<date id="F56284BCFF69CEB9FCA0FD8FD8CE3E5A" box="[863,925,606,628]" pageId="228" pageNumber="229" value="1981-01">i.1981</date>
, P. Grosso, A. Zanetic and A. Goloboff leg., MACN-Ar 36533);
<bibRefCitation id="E54DDF8DFF69CEB9FC30FDAFDF8D3EBD" author="World Spider Catalog" box="[975,1246,638,660]" pageId="228" pageNumber="229" refId="ref140996" refString="World Spider Catalog (2020) World Spider Catalog. Natural History Museum Bern. Available from: http: // wsc. nmbe. ch (accessed on 22 July 2020)" type="url" year="2020">World Spider Catalog, 2020</bibRefCitation>
.
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</paragraph>
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<paragraph id="8163A27CFF69CEB9FCD5FD12DFFC380B" blockId="228.[810,1452,707,1098]" pageId="228" pageNumber="229">
<emphasis id="B3A87E6EFF69CEB9FCD5FD12D8F73EF3" bold="true" box="[810,932,707,733]" pageId="228" pageNumber="229">Diagnosis.</emphasis>
Males of
<taxonomicName id="46DCD9FFFF69CEB9FBD4FD12DFAF3EF3" authorityName="Rios-Tamayo &amp; Goloboff" authorityYear="2018" box="[1067,1276,707,733]" class="Arachnida" family="Actinopodidae" genus="Actinopus" kingdom="Animalia" order="Araneae" pageId="228" pageNumber="229" phylum="Arthropoda" rank="species" species="clavero">
<emphasis id="B3A87E6EFF69CEB9FBD4FD12DFAF3EF3" box="[1067,1276,707,733]" italics="true" pageId="228" pageNumber="229">Actinopus clavero</emphasis>
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are unique in
<taxonomicName id="46DCD9FFFF69CEB9FCD5FD39D8C93F2C" authorityName="Perty" authorityYear="1833" box="[810,922,744,770]" class="Arachnida" family="Actinopodidae" genus="Actinopus" kingdom="Animalia" order="Araneae" pageId="228" pageNumber="229" phylum="Arthropoda" rank="genus">
<emphasis id="B3A87E6EFF69CEB9FCD5FD39D8C93F2C" box="[810,922,744,770]" italics="true" pageId="228" pageNumber="229">Actinopus</emphasis>
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by the shape of the copulatory bulb, with embolus elongated and winding along longitudinal line of tegulum, resembling in that regard the shape of the copulatory bulb in
<taxonomicName id="46DCD9FFFF69CEB9FBFBFC84DF243F41" authorityName="Walckenaer" authorityYear="1805" box="[1028,1143,853,879]" class="Arachnida" family="Actinopodidae" genus="Missulena" kingdom="Animalia" order="Araneae" pageId="228" pageNumber="229" phylum="Arthropoda" rank="genus">
<emphasis id="B3A87E6EFF69CEB9FBFBFC84DF243F41" box="[1028,1143,853,879]" italics="true" pageId="228" pageNumber="229">Missulena</emphasis>
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. (
<figureCitation id="19E7BEF9FF69CEB9FB70FC84DFBC3F41" box="[1167,1263,853,879]" captionStart="FIGURE 209" captionStartId="231.[136,235,1452,1476]" captionTargetBox="[175,1434,193,1409]" captionTargetId="figure-186@231.[151,1435,189,1429]" captionTargetPageId="231" captionText="FIGURE 209. Actinopus clavero Ríos-Tamayo &amp; Goloboff, 2018, male:AC MACN-Ar 19827:A. Copulatory bulb, prolateral; B. Copulatory bulb, dorsal; C. Copulatory bulb, retrolateral. PA-Paraembolic apophysis; BTA-Basal Tegular Apophysis; PI-Prolateral Inferior keel; PS-Prolateral Superior keel. Scale line: 1mm." figureDoi="http://doi.org/10.5281/zenodo.5597693" httpUri="https://zenodo.org/record/5597693/files/figure.png" pageId="228" pageNumber="229">Fig. 209</figureCitation>
AC); they also present the dorsal surface of ocular area and chelicerae slightly granulated (
<figureCitation id="19E7BEF9FF69CEB9FBF5FC4CDFD03F96" box="[1034,1155,925,952]" captionStart="FIGURE 207" captionStartId="229.[136,235,1645,1669]" captionTargetBox="[171,1407,205,1604]" captionTargetId="figure-134@229.[151,1435,189,1621]" captionTargetPageId="229" captionText="FIGURE 207. Actinopus clavero Ríos-Tamayo &amp; Goloboff, 2018, male:AF MACN-Ar 19827:A.Abdomen, dorsal; B. Carapace, dorsal; C. Sternum, ventral; D. Cheliceral teeth, ventral; E. Rastellum, dorsal; F. Rastellum, ventral." figureDoi="http://doi.org/10.5281/zenodo.5597689" httpUri="https://zenodo.org/record/5597689/files/figure.png" pageId="228" pageNumber="229">Fig. 207 B</figureCitation>
) and three spines on basal third on dorsal surface of tibia III (
<figureCitation id="19E7BEF9FF69CEB9FB45FC13DE6B3FF2" box="[1210,1336,962,988]" captionStart="FIGURE 208" captionStartId="230.[136,235,1155,1179]" captionTargetBox="[180,1407,206,1115]" captionTargetId="figure-393@230.[151,1435,189,1132]" captionTargetPageId="230" captionText="FIGURE 208. Actinopus clavero Ríos-Tamayo &amp; Goloboff, 2018, male:AD MACN-Ar 19827:A. Patella III, dorsal; B. Tibia III, dorsal; C. Patella IV, dorsal; D. Tibia IV, dorsal." figureDoi="http://doi.org/10.5281/zenodo.5597691" httpUri="https://zenodo.org/record/5597691/files/figure.png" pageId="228" pageNumber="229">Fig. 208 C</figureCitation>
). Females are unique in
<taxonomicName id="46DCD9FFFF69CEB9FC3BFC36DF67382F" authorityName="Perty" authorityYear="1833" box="[964,1076,999,1025]" class="Arachnida" family="Actinopodidae" genus="Actinopus" kingdom="Animalia" order="Araneae" pageId="228" pageNumber="229" phylum="Arthropoda" rank="genus">
<emphasis id="B3A87E6EFF69CEB9FC3BFC36DF67382F" box="[964,1076,999,1025]" italics="true" pageId="228" pageNumber="229">Actinopus</emphasis>
</taxonomicName>
by the shape of receptacles, very elongated and with a tapered apex.
</paragraph>
</subSubSection>
<subSubSection id="C9C6F1F7FF69CEBBFCA0FBFEDE323B7E" lastPageId="230" lastPageNumber="231" pageId="228" pageNumber="229" type="description">
<paragraph id="8163A27CFF69CEBBFCA0FBFEDE323B7E" blockId="228.[810,1452,707,1098]" lastBlockId="230.[810,1452,1330,2056]" lastPageId="230" lastPageNumber="231" pageId="228" pageNumber="229">
MALE (MACN-Ar 19827): Total length 12.25; Carapace, long 5.37; wide 5. Carapace rounded. Anterior eye row slightly procurved, posterior row slightly recurved (
<figureCitation id="19E7BEF9FF68CEB8FF6FF8A3DA5A3BA3" box="[144,265,1906,1933]" captionStart="FIGURE 207" captionStartId="229.[136,235,1645,1669]" captionTargetBox="[171,1407,205,1604]" captionTargetId="figure-134@229.[151,1435,189,1621]" captionTargetPageId="229" captionText="FIGURE 207. Actinopus clavero Ríos-Tamayo &amp; Goloboff, 2018, male:AF MACN-Ar 19827:A.Abdomen, dorsal; B. Carapace, dorsal; C. Sternum, ventral; D. Cheliceral teeth, ventral; E. Rastellum, dorsal; F. Rastellum, ventral." figureDoi="http://doi.org/10.5281/zenodo.5597689" httpUri="https://zenodo.org/record/5597689/files/figure.png" pageId="229" pageNumber="230">Fig. 207 B</figureCitation>
). With short and weak bristles between AMEclypeus, betweenALEALP,between posterior eyes-fovea and lateral eyes-edge of carapace. Sternum with eight sigilla converging to the middle but separeted (
<figureCitation id="19E7BEF9FF68CEB8FD56F83BDBCD3402" captionStart="FIGURE 207" captionStartId="229.[136,235,1645,1669]" captionTargetBox="[171,1407,205,1604]" captionTargetId="figure-134@229.[151,1435,189,1621]" captionTargetPageId="229" captionText="FIGURE 207. Actinopus clavero Ríos-Tamayo &amp; Goloboff, 2018, male:AF MACN-Ar 19827:A.Abdomen, dorsal; B. Carapace, dorsal; C. Sternum, ventral; D. Cheliceral teeth, ventral; E. Rastellum, dorsal; F. Rastellum, ventral." figureDoi="http://doi.org/10.5281/zenodo.5597689" httpUri="https://zenodo.org/record/5597689/files/figure.png" pageId="229" pageNumber="230">Fig. 207 C</figureCitation>
). Rastellum protuberant, digitiform, hirsute, with many short spines covering apex (
<figureCitation id="19E7BEF9FF68CEB8FB87F8C9DF8F3B1C" box="[1144,1244,1816,1842]" captionStart="FIGURE 207" captionStartId="229.[136,235,1645,1669]" captionTargetBox="[171,1407,205,1604]" captionTargetId="figure-134@229.[151,1435,189,1621]" captionTargetPageId="229" captionText="FIGURE 207. Actinopus clavero Ríos-Tamayo &amp; Goloboff, 2018, male:AF MACN-Ar 19827:A.Abdomen, dorsal; B. Carapace, dorsal; C. Sternum, ventral; D. Cheliceral teeth, ventral; E. Rastellum, dorsal; F. Rastellum, ventral." figureDoi="http://doi.org/10.5281/zenodo.5597689" httpUri="https://zenodo.org/record/5597689/files/figure.png" pageId="229" pageNumber="230">Fig. 207</figureCitation>
EF). Chelicerae with twelve denticles along prolateral row of teeth. Prolateral row with six megateeth. Retrolateral row with five megateeth and one tooth between two basal megateeth (
<figureCitation id="19E7BEF9FF68CEB8FCCDF86ED8EB3BF7" box="[818,952,1983,2009]" captionStart="FIGURE 207" captionStartId="229.[136,235,1645,1669]" captionTargetBox="[171,1407,205,1604]" captionTargetId="figure-134@229.[151,1435,189,1621]" captionTargetPageId="229" captionText="FIGURE 207. Actinopus clavero Ríos-Tamayo &amp; Goloboff, 2018, male:AF MACN-Ar 19827:A.Abdomen, dorsal; B. Carapace, dorsal; C. Sternum, ventral; D. Cheliceral teeth, ventral; E. Rastellum, dorsal; F. Rastellum, ventral." figureDoi="http://doi.org/10.5281/zenodo.5597689" httpUri="https://zenodo.org/record/5597689/files/figure.png" pageId="229" pageNumber="230">Fig. 207 D</figureCitation>
). Patella and tibia III with distal crown of well-developed thorns, not interrupted in middle (
<figureCitation id="19E7BEF9FF68CEB8FAB0F838DEF8342D" box="[1359,1451,2025,2051]" captionStart="FIGURE 208" captionStartId="230.[136,235,1155,1179]" captionTargetBox="[180,1407,206,1115]" captionTargetId="figure-393@230.[151,1435,189,1132]" captionTargetPageId="230" captionText="FIGURE 208. Actinopus clavero Ríos-Tamayo &amp; Goloboff, 2018, male:AD MACN-Ar 19827:A. Patella III, dorsal; B. Tibia III, dorsal; C. Patella IV, dorsal; D. Tibia IV, dorsal." figureDoi="http://doi.org/10.5281/zenodo.5597691" httpUri="https://zenodo.org/record/5597691/files/figure.png" pageId="229" pageNumber="230">Fig. 208</figureCitation>
AB). Patellae III and IV with spines on prolaterodorsal face (
<figureCitation id="19E7BEF9FF6BCEBBFF36FAE3DA3D3962" box="[201,366,1330,1356]" captionStart="FIGURE 208" captionStartId="230.[136,235,1155,1179]" captionTargetBox="[180,1407,206,1115]" captionTargetId="figure-393@230.[151,1435,189,1132]" captionTargetPageId="230" captionText="FIGURE 208. Actinopus clavero Ríos-Tamayo &amp; Goloboff, 2018, male:AD MACN-Ar 19827:A. Patella III, dorsal; B. Tibia III, dorsal; C. Patella IV, dorsal; D. Tibia IV, dorsal." figureDoi="http://doi.org/10.5281/zenodo.5597691" httpUri="https://zenodo.org/record/5597691/files/figure.png" pageId="230" pageNumber="231">Fig. 208 A, C</figureCitation>
). Tibia IV without spines on dorsal surface (
<figureCitation id="19E7BEF9FF6BCEBBFF15FA86DA3F395F" box="[234,364,1367,1393]" captionStart="FIGURE 208" captionStartId="230.[136,235,1155,1179]" captionTargetBox="[180,1407,206,1115]" captionTargetId="figure-393@230.[151,1435,189,1132]" captionTargetPageId="230" captionText="FIGURE 208. Actinopus clavero Ríos-Tamayo &amp; Goloboff, 2018, male:AD MACN-Ar 19827:A. Patella III, dorsal; B. Tibia III, dorsal; C. Patella IV, dorsal; D. Tibia IV, dorsal." figureDoi="http://doi.org/10.5281/zenodo.5597691" httpUri="https://zenodo.org/record/5597691/files/figure.png" pageId="230" pageNumber="231">Fig. 208 D</figureCitation>
). Ventral pseudoscopulae occupying 50% of tarsus I, 70% of II, 20% of metatarsi IV and 100% of III and IV. Pseudoscopula of tarsi I, II and metatarsi IV with setae spaced, forming diffuse group of bristles; On tarsi III and IV, compact. Carapace, chelicerae, sternum, coxae, trochantera, femora, patellae, tibiae, metatarsi and tarsi reddish brown; Abdomen pale brown with a scutum covering 1/3 of anterior dorsal surface o abdomen (
<figureCitation id="19E7BEF9FF6BCEBBFD23F989DB853AB9" captionStart="FIGURE 207" captionStartId="229.[136,235,1645,1669]" captionTargetBox="[171,1407,205,1604]" captionTargetId="figure-134@229.[151,1435,189,1621]" captionTargetPageId="229" captionText="FIGURE 207. Actinopus clavero Ríos-Tamayo &amp; Goloboff, 2018, male:AF MACN-Ar 19827:A.Abdomen, dorsal; B. Carapace, dorsal; C. Sternum, ventral; D. Cheliceral teeth, ventral; E. Rastellum, dorsal; F. Rastellum, ventral." figureDoi="http://doi.org/10.5281/zenodo.5597689" httpUri="https://zenodo.org/record/5597689/files/figure.png" pageId="230" pageNumber="231">Fig. 207 A</figureCitation>
).
<emphasis id="B3A87E6EFF6BCEBBFF0FF9ACDA903AB9" bold="true" box="[240,451,1661,1687]" pageId="230" pageNumber="231">Eyes: Diameters:</emphasis>
PME 0.2, PLE 0.24, ALE 0.32, AME 0.2; MOQ: Length 0.8, front width 2.64, back width 2.56;
<emphasis id="B3A87E6EFF6BCEBBFEEAF916DA943ACF" bold="true" box="[277,455,1735,1761]" pageId="230" pageNumber="231">Interdistances:</emphasis>
PMEPME 1.4, PLEPME 0.2, AMEAME 0.28, ALEAME 0.8, ALEPLE 0.44, AMEPME 0.52.
<emphasis id="B3A87E6EFF6BCEBBFEA4F8C1DAA43B05" bold="true" box="[347,503,1808,1835]" pageId="230" pageNumber="231">Ocular area:</emphasis>
OAL 3.75, OAW 3.12, and IF 1.75.
<emphasis id="B3A87E6EFF6BCEBBFEE5F8E7DA323B7E" bold="true" box="[282,353,1846,1872]" pageId="230" pageNumber="231">Body:</emphasis>
Clypeus: 0.25; Fovea: 2.25; Labium: long 1.12; wide 1.12; Chelicerae: long 3.12; wide 1; Sternum: long 3.37; wide 3.12. Abdomen: long 6.5; wide 5.
<emphasis id="B3A87E6EFF6BCEBBFF59F875DAC13B90" bold="true" box="[166,402,1956,1982]" pageId="230" pageNumber="231">Leg measurements:</emphasis>
I: Fe 5.62/ Pa 2.25/ Ti 3.12/ Me 3.87/ Ta 2/ total 16.87. II: 5.12/ 2.25/ 3/ 3.75/ 2/ 16.12. III: 3.87/ 2.25/ 2.37/ 3.75/ 2.12 /14.37. IV: 5/ 2.5/ 3.87/ 4.25/ 2.25/ 17.87. Formula 4123.
<emphasis id="B3A87E6EFF6BCEBBFE3BF7C3D9133402" bold="true" box="[452,576,2066,2092]" pageId="230" pageNumber="231">Spination:</emphasis>
I—Fe v0, d0, p0, r0; Pa v0, d0, p0, r0; ti v3-6-15, d0, p0-0-4, r2-2-4; Me v6-9-10, d0, p1-3-2, r1-3-3; ta v3-6-4, d0, p0-1-3, r1-2-3. II—Fe v0, d0, p0, r0; Pa v0, d0, p0, r0; ti v4-5-7, d0, p1- 3-3, r0-0-2; Me v4-5-1, d0, p1-2-2, r1-2-3; ta v0-4-4, d0, p1-1-2, r1-0-1. III—Fe v0, d0-4-8, p0, r0; Pa v0, d8-10- 29, p3-0-11, r0-0-4; ti v0, d3-0-11, p1-1-1, r0-2-4; Me v0- 0-2, d4-7-5, p1-1-2, r1-2-2; ta v0-0-2, d0, p1-2-3, r0-3-4; IV—Fe v0, d0, p0, r0; Pa v0, d21-23-8, p0, r0; ti v0, d0, p0, r0; Me v0, d0-0-1, p1-0-2, r0-0-1; ta v0, d0, p1-6-7, r1-4-3.
<emphasis id="B3A87E6EFF6BCEBBFC7FF9ACD8933AB9" bold="true" box="[896,960,1661,1687]" pageId="230" pageNumber="231">Palp:</emphasis>
PA conspicuous and continuous to surface between PS and PI. Embolus with two keels (PI and PS). BTA developed and displaced dorsally in relation to a middle longitudinal line along prolateral tegular surface; serrated area absent; embolus elongated, thin, winding along longitudinal line of tegulum (
<figureCitation id="19E7BEF9FF6BCEBBFB47F8E4DE453B7E" box="[1208,1302,1845,1872]" captionStart="FIGURE 209" captionStartId="231.[136,235,1452,1476]" captionTargetBox="[175,1434,193,1409]" captionTargetId="figure-186@231.[151,1435,189,1429]" captionTargetPageId="231" captionText="FIGURE 209. Actinopus clavero Ríos-Tamayo &amp; Goloboff, 2018, male:AC MACN-Ar 19827:A. Copulatory bulb, prolateral; B. Copulatory bulb, dorsal; C. Copulatory bulb, retrolateral. PA-Paraembolic apophysis; BTA-Basal Tegular Apophysis; PI-Prolateral Inferior keel; PS-Prolateral Superior keel. Scale line: 1mm." figureDoi="http://doi.org/10.5281/zenodo.5597693" httpUri="https://zenodo.org/record/5597693/files/figure.png" pageId="230" pageNumber="231">Fig. 209</figureCitation>
AC).
</paragraph>
</subSubSection>
<caption id="D5A3F2F4FF69CEB9FF77F7D4D8903430" ID-DOI="http://doi.org/10.5281/zenodo.5655464" ID-Zenodo-Dep="5655464" box="[136,963,2053,2078]" httpUri="https://zenodo.org/record/5655464/files/figure.png" pageId="228" pageNumber="229" startId="228.[136,192,2053,2077]" targetBox="[157,1428,1127,2021]" targetPageId="228">
<paragraph id="8163A27CFF69CEB9FF77F7D4D8903430" blockId="228.[136,963,2053,2078]" box="[136,963,2053,2078]" pageId="228" pageNumber="229">
<emphasis id="B3A87E6EFF69CEB9FF77F7D4DBB53430" bold="true" box="[136,230,2053,2078]" pageId="228" pageNumber="229">MAP 15.</emphasis>
Distribution map of
<taxonomicName id="46DCD9FFFF69CEB9FE44F7D7D8EC3430" authority="Rios-Tamayo &amp; Goloboff, 2018" authorityName="Rios-Tamayo &amp; Goloboff" authorityYear="2018" box="[443,959,2054,2078]" class="Arachnida" family="Actinopodidae" genus="Actinopus" kingdom="Animalia" order="Araneae" pageId="228" pageNumber="229" phylum="Arthropoda" rank="species" species="clavero">
<emphasis id="B3A87E6EFF69CEB9FE44F7D7D9213430" box="[443,626,2054,2078]" italics="true" pageId="228" pageNumber="229">Actinopus clavero</emphasis>
<bibRefCitation id="E54DDF8DFF69CEB9FD87F7D7D8EC3430" author="Rios-Tamayo, D. &amp; Goloboff, P. A." box="[632,959,2054,2078]" pageId="228" pageNumber="229" pagination="1 - 83" refId="ref140496" refString="Rios-Tamayo, D. &amp; Goloboff, P. A. (2018) Taxonomic revision and morphology of the trapdoor spider genus Actinopus (Mygalomorphae: Actinopodidae) in Argentina. Bulletin of the American Museum of Natural History, 419, 1 - 83. https: // doi. org / 10.1206 / 0003 - 0090 - 419.1.1" type="journal article" year="2018">Ríos-Tamayo &amp; Goloboff, 2018</bibRefCitation>
</taxonomicName>
.
</paragraph>
</caption>
<caption id="D5A3F2F4FF68CEB8FF77F9BCDF283A87" ID-DOI="http://doi.org/10.5281/zenodo.5597689" ID-Zenodo-Dep="5597689" httpUri="https://zenodo.org/record/5597689/files/figure.png" pageId="229" pageNumber="230" startId="229.[136,235,1645,1669]" targetBox="[171,1407,205,1604]" targetPageId="229">
<paragraph id="8163A27CFF68CEB8FF77F9BCDF283A87" blockId="229.[136,1452,1645,1705]" pageId="229" pageNumber="230">
<emphasis id="B3A87E6EFF68CEB8FF77F9BCDA4F3AAB" bold="true" box="[136,284,1645,1669]" pageId="229" pageNumber="230">FIGURE 207.</emphasis>
<taxonomicName id="46DCD9FFFF68CEB8FEDEF9BCD84C3AAB" authority="Rios-Tamayo &amp; Goloboff, 2018" authorityName="Rios-Tamayo &amp; Goloboff" authorityYear="2018" box="[289,799,1645,1669]" class="Arachnida" family="Actinopodidae" genus="Actinopus" kingdom="Animalia" order="Araneae" pageId="229" pageNumber="230" phylum="Arthropoda" rank="species" species="clavero">
<emphasis id="B3A87E6EFF68CEB8FEDEF9BCDA843AAB" box="[289,471,1645,1669]" italics="true" pageId="229" pageNumber="230">Actinopus clavero</emphasis>
<bibRefCitation id="E54DDF8DFF68CEB8FE23F9BCD84C3AAB" author="Rios-Tamayo, D. &amp; Goloboff, P. A." box="[476,799,1645,1669]" pageId="229" pageNumber="230" pagination="1 - 83" refId="ref140496" refString="Rios-Tamayo, D. &amp; Goloboff, P. A. (2018) Taxonomic revision and morphology of the trapdoor spider genus Actinopus (Mygalomorphae: Actinopodidae) in Argentina. Bulletin of the American Museum of Natural History, 419, 1 - 83. https: // doi. org / 10.1206 / 0003 - 0090 - 419.1.1" type="journal article" year="2018">Ríos-Tamayo &amp; Goloboff, 2018</bibRefCitation>
</taxonomicName>
, male: AF MACN-Ar 19827: A. Abdomen, dorsal; B. Carapace, dorsal; C. Sternum, ventral; D. Cheliceral teeth, ventral; E. Rastellum, dorsal; F. Rastellum, ventral.
</paragraph>
</caption>
<caption id="D5A3F2F4FF6BCEBBFF77FB52D93D3891" ID-DOI="http://doi.org/10.5281/zenodo.5597691" ID-Zenodo-Dep="5597691" httpUri="https://zenodo.org/record/5597691/files/figure.png" pageId="230" pageNumber="231" startId="230.[136,235,1155,1179]" targetBox="[180,1407,206,1115]" targetPageId="230">
<paragraph id="8163A27CFF6BCEBBFF77FB52D93D3891" blockId="230.[136,1451,1155,1215]" pageId="230" pageNumber="231">
<emphasis id="B3A87E6EFF6BCEBBFF77FB52DA4E38B5" bold="true" box="[136,285,1155,1179]" pageId="230" pageNumber="231">FIGURE 208.</emphasis>
<taxonomicName id="46DCD9FFFF6BCEBBFEDDFB55D87238B5" authority="Rios-Tamayo &amp; Goloboff, 2018" authorityName="Rios-Tamayo &amp; Goloboff" authorityYear="2018" box="[290,801,1155,1179]" class="Arachnida" family="Actinopodidae" genus="Actinopus" kingdom="Animalia" order="Araneae" pageId="230" pageNumber="231" phylum="Arthropoda" rank="species" species="clavero">
<emphasis id="B3A87E6EFF6BCEBBFEDDFB55DA8B38B5" box="[290,472,1155,1179]" italics="true" pageId="230" pageNumber="231">Actinopus clavero</emphasis>
<bibRefCitation id="E54DDF8DFF6BCEBBFE22FB52D87238B5" author="Rios-Tamayo, D. &amp; Goloboff, P. A." box="[477,801,1155,1179]" pageId="230" pageNumber="231" pagination="1 - 83" refId="ref140496" refString="Rios-Tamayo, D. &amp; Goloboff, P. A. (2018) Taxonomic revision and morphology of the trapdoor spider genus Actinopus (Mygalomorphae: Actinopodidae) in Argentina. Bulletin of the American Museum of Natural History, 419, 1 - 83. https: // doi. org / 10.1206 / 0003 - 0090 - 419.1.1" type="journal article" year="2018">Ríos-Tamayo &amp; Goloboff, 2018</bibRefCitation>
</taxonomicName>
, male: AD MACN-Ar 19827: A. Patella III, dorsal; B. Tibia III, dorsal; C. Patella IV, dorsal; D. Tibia IV, dorsal.
</paragraph>
</caption>
<subSubSection id="C9C6F1F7FF6BCEBBFCA0F88BDF9F3B90" pageId="230" pageNumber="231" type="materials_examined">
<paragraph id="8163A27CFF6BCEBBFCA0F88BDF9F3B90" blockId="230.[810,1452,1330,2056]" pageId="230" pageNumber="231">
<emphasis id="B3A87E6EFF6BCEBBFCA0F88BDF1F3B5A" bold="true" box="[863,1100,1882,1908]" pageId="230" pageNumber="231">Material examined.</emphasis>
<materialsCitation id="31B4A821FF6BCEBBFBA9F88BDF9B3B90" ID-GBIF-Occurrence="3399910390" collectionCode="M" collectorName="Rumboll" country="Argentina" latitude="-30.923342" location="Cordoba" longLatPrecision="1" longitude="-64.499084" municipality="Los Cocos" pageId="230" pageNumber="231" specimenCount="1">
<collectingCountry id="F9CBE2ECFF6BCEBBFBA9F88BDFAD3B5A" box="[1110,1278,1882,1908]" name="Argentina" pageId="230" pageNumber="231">ARGENTINA</collectingCountry>
:
<emphasis id="B3A87E6EFF6BCEBBFAF4F88BDE243B5A" box="[1291,1399,1882,1908]" italics="true" pageId="230" pageNumber="231">
<location id="8403F4A7FF6BCEBBFAF4F88BDE223B5A" LSID="urn:lsid:plazi:treatment:0975136AFF69CEBBFCD5FE82D8C93426:8403F4A7FF6BCEBBFAF4F88BDE223B5A" box="[1291,1393,1882,1908]" country="Argentina" latitude="-30.923342" longLatPrecision="1" longitude="-64.499084" municipality="Los Cocos" name="Cordoba" pageId="230" pageNumber="231">Córdoba</location>
:
</emphasis>
<collectingMunicipality id="61073806FF6BCEBBFA7EF88AD8233BB7" pageId="230" pageNumber="231">Los Cocos</collectingMunicipality>
[
<geoCoordinate id="E4E8C4BBFF6BCEBBFC83F8AEDF7D3BB7" box="[892,1070,1919,1946]" degrees="30" direction="south" minutes="55" orientation="latitude" pageId="230" pageNumber="231" precision="1" seconds="24.03" value="-30.923342">30º 5524.03” S</geoCoordinate>
<geoCoordinate id="E4E8C4BBFF6BCEBBFBCDF8AEDFA73BB7" box="[1074,1268,1918,1946]" degrees="64" direction="west" minutes="29" orientation="longitude" pageId="230" pageNumber="231" precision="1" seconds="56.73" value="-64.499084">64º 29 56.73” W</geoCoordinate>
], 1♂, xxi.1995,
<collectionCode id="E7CD3AB9FF6BCEBBFCD5F875D8153B90" box="[810,838,1956,1982]" country="Germany" lsid="urn:lsid:biocol.org:col:15637" name="Botanische Staatssammlung München" pageId="230" pageNumber="231" type="Herbarium">M</collectionCode>
.
<collectorName id="2C29C7AAFF6BCEBBFCAEF875D8E03B90" box="[849,947,1956,1982]" pageId="230" pageNumber="231">Rumboll</collectorName>
leg. (MACN-Ar 19827)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C9C6F1F7FF6BCEBBFCA0F819D8C93426" pageId="230" pageNumber="231" type="distribution">
<paragraph id="8163A27CFF6BCEBBFCA0F819D8C93426" blockId="230.[810,1452,1330,2056]" pageId="230" pageNumber="231">
<emphasis id="B3A87E6EFF6BCEBBFCA0F819D8AA3BCC" bold="true" box="[863,1017,1992,2018]" pageId="230" pageNumber="231">Distribution.</emphasis>
<collectingCountry id="F9CBE2ECFF6BCEBBFBFCF818DFF83BCD" box="[1027,1195,1993,2019]" name="Argentina" pageId="230" pageNumber="231">ARGENTINA</collectingCountry>
: Córdoba: Los Cocos (
<figureCitation id="19E7BEF9FF6BCEBBFCCEF83FD8DF3426" box="[817,908,2029,2056]" captionStart="MAP 15" captionStartId="228.[136,192,2053,2077]" captionTargetBox="[157,1428,1127,2021]" captionTargetId="figure-458@228.[151,1435,1121,2029]" captionText="MAP 15. Distribution map of Actinopus clavero Ríos-Tamayo &amp; Goloboff, 2018." figureDoi="http://doi.org/10.5281/zenodo.5655464" httpUri="https://zenodo.org/record/5655464/files/figure.png" pageId="230" pageNumber="231">Map 15</figureCitation>
).
</paragraph>
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