360 lines
38 KiB
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360 lines
38 KiB
XML
<document ID-DOI="http://doi.org/10.5281/zenodo.4657185" ID-GBIF-Dataset="acf2b7a6-7f3c-443b-a3fe-307fda431e51" ID-GBIF-Taxon="180784025" ID-ISSN="0067-1975" ID-Zenodo-Dep="4657185" checkinTime="1617214739793" checkinUser="felipe" docAuthor="Just, Jean" docDate="1998" docId="B128F8650E24E827FE29F8E7F71DD235" docLanguage="en" docName="RecAustMus.50.1.27-54.pdf.imf" docOrigin="Records of the Australian Museum 50 (1)" docSource="https://journals.australian.museum/just-1998-rec-aust-mus-501-2754/" docStyle="DocumentStyle:CF64E561BE6B94FA5F31B9132C6D4316.1:RecAustMus.1998.journal_article" docStyleId="CF64E561BE6B94FA5F31B9132C6D4316" docStyleName="RecAustMus.1998.journal_article" docStyleVersion="1" docTitle="Caribboecetes Just 1983" docType="treatment" docVersion="5" lastPageId="3" lastPageNumber="30" masterDocId="4D11801D0E26E824FFF1FFC6FFD4D472" masterDocTitle="Siphonoecetinae (Crustacea: Amphipoda: Ischyroceridae) 7: Australian Concholestids, Ambicholestes n. gen., with a Description of Six New Species, and a New, Restricted Diagnosis for Caribboecetes Just, 1983" masterLastPageNumber="54" masterPageNumber="27" pageId="2" pageNumber="29" updateTime="1632023270205" updateUser="ExternalLinkService">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>Siphonoecetinae (Crustacea: Amphipoda: Ischyroceridae) 7: Australian Concholestids, Ambicholestes n. gen., with a Description of Six New Species, and a New, Restricted Diagnosis for Caribboecetes Just, 1983</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Just, Jean</mods:namePart>
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<mods:affiliation>Australian Biological Resources Study, GPO Box 636, Canberra ACT 2601, Australia</mods:affiliation>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:titleInfo>
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<mods:title>Records of the Australian Museum</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>1998</mods:date>
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<mods:detail type="pubDate">
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<mods:number>1998-05-13</mods:number>
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</mods:detail>
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<mods:detail type="volume">
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<mods:number>50</mods:number>
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</mods:detail>
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<mods:detail type="issue">
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<mods:number>1</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>27</mods:start>
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<mods:end>54</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:location>
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<mods:url>https://journals.australian.museum/just-1998-rec-aust-mus-501-2754/</mods:url>
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</mods:location>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">10.3853/j.0067-1975.50.1998.1273</mods:identifier>
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<mods:identifier type="GBIF-Dataset">acf2b7a6-7f3c-443b-a3fe-307fda431e51</mods:identifier>
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<mods:identifier type="ISSN">0067-1975</mods:identifier>
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<mods:identifier type="Zenodo-Dep">4656898</mods:identifier>
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</mods:mods>
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<treatment ID-DOI="http://doi.org/10.5281/zenodo.4657185" ID-GBIF-Taxon="180784025" ID-Zenodo-Dep="4657185" LSID="urn:lsid:plazi:treatment:B128F8650E24E827FE29F8E7F71DD235" httpUri="http://treatment.plazi.org/id/B128F8650E24E827FE29F8E7F71DD235" lastPageId="3" lastPageNumber="30" pageId="2" pageNumber="29">
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<subSubSection box="[472,920,1825,1864]" pageId="2" pageNumber="29" type="nomenclature">
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<paragraph blockId="2.[472,920,1825,1864]" box="[472,920,1825,1864]" pageId="2" pageNumber="29">
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<heading box="[472,920,1825,1864]" centered="true" fontSize="10" level="2" pageId="2" pageNumber="29" reason="2">
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<taxonomicName authority="Just, 1983" authorityName="Just" authorityYear="1983" box="[472,920,1825,1864]" class="Malacostraca" family="Ischyroceridae" genus="Caribboecetes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Amphipoda" pageId="2" pageNumber="29" phylum="Arthropoda" rank="genus">
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<emphasis box="[472,721,1825,1864]" italics="true" pageId="2" pageNumber="29">Caribboecetes</emphasis>
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<bibRefCitation author="Just, J." box="[741,920,1825,1864]" pageId="2" pageNumber="29" pagination="35" refId="ref11748" refString="Just, J., 1983. Siphonoecetinae subfam. n. (Crustacea, Amphipoda, Corophiidae) 1: Classification. Steenstrupia 9 (6): 117 - l 35." type="journal article" year="1983">Just, 1983</bibRefCitation>
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</taxonomicName>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection pageId="2" pageNumber="29" type="diagnosis">
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<paragraph blockId="2.[203,1188,1907,2315]" pageId="2" pageNumber="29">
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New restricted diagnosis. Concholestid
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<taxonomicName authority="(sensu Just, 1988)" baseAuthorityName="sensu Just" baseAuthorityYear="1988" class="Malacostraca" family="Ischyroceridae" kingdom="Animalia" order="Amphipoda" pageId="2" pageNumber="29" phylum="Arthropoda" rank="subFamily" subFamily="Siphonoecetinae">
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Siphonoecetinae
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<emphasis box="[207,310,1953,1992]" italics="true" pageId="2" pageNumber="29">(sensu</emphasis>
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<bibRefCitation author="Just, J." box="[326,498,1953,1992]" pageId="2" pageNumber="29" pagination="193 - 208" refId="ref11940" refString="Just, J., 1988. Siphonoecetinae (Crustacea, Amphipoda, Corophiidae) 6: A survey of phylogeny, distribution, and biology. Crustaceana, Supplement l 3: 193 - 208." type="journal article" year="1988">Just, 1988</bibRefCitation>
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)
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</taxonomicName>
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with inner ramus of uropod 1 at most 1/2 length of outer ramus, flattened, with robust setae on margin. Uropod 3 without ramus. Mandibular palp 2- articulate, blunt, reaching to about apex of incisor. Inner lobes of lower lip simple. Ocular field of eye lobes not inflated. Article 2 of pereopod 7 without single midlateral row of setae. Oostegites on pereopods 2-5. Gills on pereopods 4-6.
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</paragraph>
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</subSubSection>
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<caption ID-DOI="http://doi.org/10.5281/zenodo.4656904" ID-Zenodo-Dep="4656904" httpUri="https://zenodo.org/record/4656904/files/figure.png" pageId="2" pageNumber="29" startId="2.[1258,1313,2258,2294]" targetBox="[1258,2244,853,2219]" targetPageId="2">
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<paragraph blockId="2.[1253,2244,2258,2544]" pageId="2" pageNumber="29">
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Fig. 2. Lower lip of selected concholestid
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<taxonomicName baseAuthorityName="sensu Just" baseAuthorityYear="1988" box="[1932,2185,2258,2294]" class="Malacostraca" family="Ischyroceridae" kingdom="Animalia" order="Amphipoda" pageId="2" pageNumber="29" phylum="Arthropoda" rank="subFamily" subFamily="Siphonoecetinae">Siphonoecetinae</taxonomicName>
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. A,
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<taxonomicName authority="Just, 1987" authorityName="Just" authorityYear="1987" box="[1261,1800,2300,2335]" class="Malacostraca" family="Corophiidae" genus="Concholestes" kingdom="Animalia" order="Amphipoda" pageId="2" pageNumber="29" phylum="Arthropoda" rank="species" species="omani">
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<emphasis box="[1261,1601,2300,2335]" italics="true" pageId="2" pageNumber="29">Concholestes omani</emphasis>
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<bibRefCitation author="Just, J." box="[1621,1800,2300,2335]" pageId="2" pageNumber="29" pagination="93 - 99" refId="ref11900" refString="Just, J., 1987. Siphonoecetinae (Crustacea, Amphipoda, Corophiidae) 5: Concholestes omani n. sp. from the coast of Oman. Steenstrupia l 3 (2): 93 - 99." type="journal article" year="1987">Just, 1987</bibRefCitation>
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</taxonomicName>
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; B,
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<taxonomicName authority="(Griffiths, 1974)" baseAuthorityName="Griffiths" baseAuthorityYear="1974" class="Malacostraca" family="Corophiidae" genus="Africoecetes" kingdom="Animalia" order="Amphipoda" pageId="2" pageNumber="29" phylum="Arthropoda" rank="species" species="armatus">
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<emphasis box="[1879,2244,2300,2335]" italics="true" pageId="2" pageNumber="29">Africoecetes armatus</emphasis>
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(
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<bibRefCitation author="Griffiths" box="[1270,1506,2341,2377]" pageId="2" pageNumber="29" pagination="251 - 336" refId="ref11704" refString="Griffiths, c. L., 1974. The Amphipoda of southern Africa 4: The Gammaridea and Caprellidea of the Cape Province east of Cape Agulhas. Annals of the South African Museum 65 (9): 251 - 336." type="journal article" year="1974">Griffiths, 1974</bibRefCitation>
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)
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</taxonomicName>
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; C,
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<taxonomicName authority="Just, 1983" authorityName="Just" authorityYear="1983" box="[1592,2184,2341,2377]" class="Malacostraca" family="Ischyroceridae" genus="Caribboecetes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Amphipoda" pageId="2" pageNumber="29" phylum="Arthropoda" rank="species" species="barbadensis">
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<emphasis box="[1592,2006,2341,2377]" italics="true" pageId="2" pageNumber="29">Caribboecetes barbadensis</emphasis>
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<bibRefCitation author="Just, J." box="[2022,2184,2341,2377]" pageId="2" pageNumber="29" pagination="35" refId="ref11748" refString="Just, J., 1983. Siphonoecetinae subfam. n. (Crustacea, Amphipoda, Corophiidae) 1: Classification. Steenstrupia 9 (6): 117 - l 35." type="journal article" year="1983">Just, 1983</bibRefCitation>
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</taxonomicName>
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; D,
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<taxonomicName authority="Just, 1984" authorityName="Just" authorityYear="1984" box="[1261,1918,2382,2418]" class="Malacostraca" family="Ischyroceridae" genus="Caribboecetes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Amphipoda" pageId="2" pageNumber="29" phylum="Arthropoda" rank="species" species="intermedius">
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<emphasis box="[1261,1719,2382,2418]" italics="true" pageId="2" pageNumber="29">Caribboecetes intermedius</emphasis>
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Just, 1984
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</taxonomicName>
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; E,
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<taxonomicName authority="Just, 1984" authorityName="Just" authorityYear="1984" class="Malacostraca" family="Ischyroceridae" genus="Caribboecetes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Amphipoda" pageId="2" pageNumber="29" phylum="Arthropoda" rank="species" species="jenikarpae">
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<emphasis italics="true" pageId="2" pageNumber="29">Caribboecetes jenikarpae</emphasis>
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Just, 1984
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</taxonomicName>
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; F,
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<taxonomicName baseAuthorityName="Just" baseAuthorityYear="1984" box="[1645,2057,2424,2460]" class="Malacostraca" family="Ischyroceridae" genus="Ambicholestes" kingdom="Animalia" order="Amphipoda" pageId="2" pageNumber="29" phylum="Arthropoda" rank="species" species="crassicornis">
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<emphasis box="[1645,2057,2424,2460]" italics="true" pageId="2" pageNumber="29">Ambicholestes crassicornis</emphasis>
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</taxonomicName>
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(Just, 1984, n. comb.); G,
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<taxonomicName baseAuthorityName="Just" baseAuthorityYear="1984" box="[1473,1888,2466,2502]" class="Malacostraca" family="Ischyroceridae" genus="Ambicholestes" kingdom="Animalia" order="Amphipoda" pageId="2" pageNumber="29" phylum="Arthropoda" rank="species" species="squamiferus">
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<emphasis box="[1473,1888,2466,2502]" italics="true" pageId="2" pageNumber="29">Ambicholestes squamiferus</emphasis>
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</taxonomicName>
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(Just, 1984, n. comb.);
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<emphasis bold="true" box="[1259,1295,2508,2544]" pageId="2" pageNumber="29">H,</emphasis>
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<taxonomicName authorityName="Just" authorityYear="1998" box="[1309,1691,2508,2544]" class="Malacostraca" family="Ischyroceridae" genus="Ambicholestes" kingdom="Animalia" order="Amphipoda" pageId="2" pageNumber="29" phylum="Arthropoda" rank="species" species="berentsae" status="n.sp.">
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<emphasis box="[1309,1691,2508,2544]" italics="true" pageId="2" pageNumber="29">Ambicholestes berentsae</emphasis>
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</taxonomicName>
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<taxonomicNameLabel box="[1707,1778,2508,2544]" pageId="2" pageNumber="29" rank="species">n.sp.</taxonomicNameLabel>
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, lateral view. Not to scale
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</paragraph>
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</caption>
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<subSubSection pageId="2" pageNumber="29" type="materials_examined">
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<paragraph blockId="2.[205,1185,2356,2441]" pageId="2" pageNumber="29">
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<typeStatus box="[205,295,2356,2395]" pageId="2" pageNumber="29">Type</typeStatus>
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species.
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<taxonomicName authority="Just, 1983" authorityName="Just" authorityYear="1983" box="[480,1185,2356,2395]" class="Malacostraca" family="Ischyroceridae" genus="Caribboecetes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Amphipoda" pageId="2" pageNumber="29" phylum="Arthropoda" rank="species" species="barbadensis">
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<emphasis box="[480,978,2356,2395]" italics="true" pageId="2" pageNumber="29">Caribboecetes barbadensis</emphasis>
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<bibRefCitation author="Just, J." box="[998,1185,2356,2395]" pageId="2" pageNumber="29" pagination="35" refId="ref11748" refString="Just, J., 1983. Siphonoecetinae subfam. n. (Crustacea, Amphipoda, Corophiidae) 1: Classification. Steenstrupia 9 (6): 117 - l 35." type="journal article" year="1983">Just, 1983</bibRefCitation>
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</taxonomicName>
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(original designation).
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</paragraph>
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<paragraph blockId="2.[203,1185,2483,2615]" pageId="2" pageNumber="29">
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Additional species.
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<taxonomicName authority="Just, 1984 a" authorityName="Just" authorityYear="1984" class="Malacostraca" family="Ischyroceridae" genus="Caribboecetes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Amphipoda" pageId="2" pageNumber="29" phylum="Arthropoda" rank="species" species="intermedius">
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<emphasis box="[593,1085,2483,2523]" italics="true" pageId="2" pageNumber="29">Caribboecetes intermedius</emphasis>
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<bibRefCitation author="Just, J." pageId="2" pageNumber="29" pagination="37 - 64" refId="ref11782" refString="Just, J., 1984 a. Siphonoecetinae (Crustacea, Amphipoda, Corophiidae) 2: Caribboecetes Just, 1983, with description of six new species. Steenstrupia 10 (2): 37 - 64." type="journal article" year="1984">Just, 1984a</bibRefCitation>
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</taxonomicName>
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, C.
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<taxonomicName authority="Just, 1984 a" authorityName="Just" authorityYear="1984" box="[388,806,2530,2569]" class="Malacostraca" family="Ischyroceridae" genus="Caribboecetes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Amphipoda" pageId="2" pageNumber="29" phylum="Arthropoda" rank="species" species="jenikarpae">
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<emphasis box="[388,580,2530,2569]" italics="true" pageId="2" pageNumber="29">jenikarpae</emphasis>
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<bibRefCitation author="Just, J." box="[600,806,2530,2569]" pageId="2" pageNumber="29" pagination="37 - 64" refId="ref11782" refString="Just, J., 1984 a. Siphonoecetinae (Crustacea, Amphipoda, Corophiidae) 2: Caribboecetes Just, 1983, with description of six new species. Steenstrupia 10 (2): 37 - 64." type="journal article" year="1984">Just, 1984a</bibRefCitation>
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</taxonomicName>
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, C.
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<taxonomicName authority="Just, 1984 a" authorityName="Just" authorityYear="1984" class="Malacostraca" family="Ischyroceridae" genus="Caribboecetes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Amphipoda" pageId="2" pageNumber="29" phylum="Arthropoda" rank="species" species="pterycornis">
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<emphasis box="[886,1089,2530,2569]" italics="true" pageId="2" pageNumber="29">pterycornis</emphasis>
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<bibRefCitation author="Just, J." pageId="2" pageNumber="29" pagination="37 - 64" refId="ref11782" refString="Just, J., 1984 a. Siphonoecetinae (Crustacea, Amphipoda, Corophiidae) 2: Caribboecetes Just, 1983, with description of six new species. Steenstrupia 10 (2): 37 - 64." type="journal article" year="1984">Just, 1984a</bibRefCitation>
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</taxonomicName>
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(tentatively).
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</paragraph>
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</subSubSection>
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<subSubSection pageId="2" pageNumber="29" type="description">
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<paragraph blockId="2.[199,1188,2657,3202]" pageId="2" pageNumber="29">
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Further description. Rostrum present or absent. Coxal plates 5-6 antero- as well as posterolobate (except C.
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<emphasis box="[199,409,2749,2788]" italics="true" pageId="2" pageNumber="29">
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<taxonomicName authorityName="Just" authorityYear="1984" box="[199,403,2749,2788]" class="Malacostraca" family="Ischyroceridae" genus="Caribboecetes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Amphipoda" pageId="2" pageNumber="29" phylum="Arthropoda" rank="species" species="pterycornis">pterycornis</taxonomicName>
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,
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</emphasis>
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posterolobate only). Upper lip with apical notch. Antenna 1 article 3 50-60% (c.
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<emphasis box="[887,1186,2795,2834]" italics="true" pageId="2" pageNumber="29">
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<taxonomicName authorityName="Just" authorityYear="1984" box="[887,1081,2795,2834]" class="Malacostraca" family="Ischyroceridae" genus="Caribboecetes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Amphipoda" pageId="2" pageNumber="29" phylum="Arthropoda" rank="species" species="pterycornis">pterycornis</taxonomicName>
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70%)
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</emphasis>
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length of article 1, article 2 80-95% length of article 1. Antenna 2 article 4 75-90% length of article 5. Antenna 1 reaching from just beyond peduncle of antenna 2 to about tip of flagellum of antenna 2. Pereopod 1 article 6 80- 90% length of article 5, without posteroapical tooth, with posteroapical robust seta and 1-2 robust setae on posterior margin; article 5 with 1 posteroapical robust seat. Pereopod 2 article 6 10-45% longer than article 5 (except c.
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</paragraph>
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<paragraph blockId="2.[1253,2244,2258,2544]" box="[2198,2203,2508,2544]" pageId="2" pageNumber="29">.</paragraph>
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<paragraph blockId="2.[1255,2246,2756,3072]" pageId="2" pageNumber="29">
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<emphasis box="[1255,1459,2756,2796]" italics="true" pageId="2" pageNumber="29">
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<taxonomicName authorityName="Just" authorityYear="1984" box="[1255,1453,2756,2796]" class="Malacostraca" family="Ischyroceridae" genus="Caribboecetes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Amphipoda" pageId="2" pageNumber="29" phylum="Arthropoda" rank="species" species="pterycornis">pterycornis</taxonomicName>
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,
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</emphasis>
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70%), with posteroapical triangular tooth and varying number of posterior robust setae. Pereopods 3-4 with posteriorrobust setae on article 5. Pereopods 5-6 with crescent of equal-sized short, robust setae on article 5. Pleopods with peduncle broader than long; rami as long as or longer than width of peduncle; inner ramus slightly longer than outer ramus.
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</paragraph>
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</subSubSection>
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<subSubSection lastPageId="3" lastPageNumber="30" pageId="2" pageNumber="29" type="distribution">
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<paragraph blockId="2.[1259,2247,3113,3199]" pageId="2" pageNumber="29">
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Distribution. Caribbean, Pacific
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<collectingRegion box="[1830,1960,3113,3153]" country="Mexico" name="Mexico" pageId="2" pageNumber="29">Mexico</collectingRegion>
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; from low water mark to
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<specimenCount box="[1405,1500,3161,3199]" pageId="2" pageNumber="29" type="generic">
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<quantity box="[1405,1495,3161,3199]" metricMagnitude="1" metricUnit="m" metricValue="5.4" pageId="2" pageNumber="29" unit="m" value="54.0">54 m</quantity>
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.
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</specimenCount>
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</paragraph>
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<paragraph blockId="3.[471,944,404,443]" box="[471,944,404,443]" pageId="3" pageNumber="30">Australian Concholestids</paragraph>
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</subSubSection>
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<subSubSection pageId="3" pageNumber="30" type="discussion">
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<paragraph blockId="3.[216,1199,486,1537]" pageId="3" pageNumber="30">
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A full cladistic analysis of the
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<taxonomicName baseAuthorityName="sensu Just" baseAuthorityYear="1988" box="[766,1046,486,524]" class="Malacostraca" family="Ischyroceridae" kingdom="Animalia" order="Amphipoda" pageId="3" pageNumber="30" phylum="Arthropoda" rank="subFamily" subFamily="Siphonoecetinae">Siphonoecetinae</taxonomicName>
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, and the concholestid component in particular, is not feasible at the present time. Many siphonoecetines remain to be described, including groups of species exhibiting characters which will influence the outcome of such an analysis. The problem is compounded by the present uncertainties, at the outgroup level, of where the
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<taxonomicName baseAuthorityName="sensu Just" baseAuthorityYear="1988" box="[794,1068,762,801]" class="Malacostraca" family="Ischyroceridae" kingdom="Animalia" order="Amphipoda" pageId="3" pageNumber="30" phylum="Arthropoda" rank="subFamily" subFamily="Siphonoecetinae">Siphonoecetinae</taxonomicName>
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belong in the
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<taxonomicName authorityName="Leach" authorityYear="1814" box="[339,576,808,847]" pageId="3" pageNumber="54" rank="superFamily" superFamily="Corophioidea">Corophioidea</taxonomicName>
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, and by difficulties in delimiting families with which
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<taxonomicName baseAuthorityName="sensu Just" baseAuthorityYear="1988" box="[583,868,854,893]" class="Malacostraca" family="Ischyroceridae" kingdom="Animalia" order="Amphipoda" pageId="3" pageNumber="30" phylum="Arthropoda" rank="subFamily" subFamily="Siphonoecetinae">Siphonoecetinae</taxonomicName>
|
|
are believed to be related. The proposed classification of the Australian concholestids is based on immediately obvious synapomorphies, such as a complex lower lip versus a simple one, and an elongated, I-articulate, pointed mandibular palp versus the otherwise ubiquitous short, club shaped palp with a tiny second article (in a single known species the first article is broadly expanded). The use of subgeneric classification is a subjective convenience to keep together species sharing a major synapomorphy, e.g., the complex lower lip in
|
|
<emphasis box="[647,894,1314,1353]" italics="true" pageId="3" pageNumber="30">
|
|
<taxonomicName authorityName="Just" authorityYear="1998" box="[647,888,1314,1353]" class="Malacostraca" family="Ischyroceridae" genus="Ambicholestes" kingdom="Animalia" order="Amphipoda" pageId="3" pageNumber="30" phylum="Arthropoda" rank="genus">Ambicholestes</taxonomicName>
|
|
,
|
|
</emphasis>
|
|
but differing at a perceived subordinate sister group level, here
|
|
<taxonomicName authorityName="Just" authorityYear="1998" box="[965,1196,1360,1400]" class="Malacostraca" family="Ischyroceridae" genus="Ambicholestes" kingdom="Animalia" order="Amphipoda" pageId="3" pageNumber="30" phylum="Arthropoda" rank="genus" sensu="stricto">
|
|
<emphasis box="[965,1196,1360,1400]" italics="true" pageId="3" pageNumber="30">Ambicholestes</emphasis>
|
|
</taxonomicName>
|
|
<taxonomicNameLabel box="[218,415,1406,1445]" pageId="3" pageNumber="30" sensu="stricto">sensu stricto</taxonomicNameLabel>
|
|
versus
|
|
<emphasis box="[536,1005,1406,1445]" italics="true" pageId="3" pageNumber="30">
|
|
<taxonomicName authority="" box="[536,1000,1406,1445]" class="Malacostraca" family="Ischyroceridae" genus="Ambicholestes" kingdom="Animalia" order="Amphipoda" pageId="3" pageNumber="30" phylum="Arthropoda" rank="subGenus" subGenus="Austrolestes">Ambicholestes (Austrolestes)</taxonomicName>
|
|
.
|
|
</emphasis>
|
|
This use of the subgenus category is consistent with
|
|
<bibRefCitation author="Just, J." box="[968,1183,1452,1491]" pageId="3" pageNumber="30" pagination="35" refId="ref11748" refString="Just, J., 1983. Siphonoecetinae subfam. n. (Crustacea, Amphipoda, Corophiidae) 1: Classification. Steenstrupia 9 (6): 117 - l 35." type="journal article" year="1983">Just (1983</bibRefCitation>
|
|
-
|
|
<emphasis box="[216,467,1498,1537]" italics="true" pageId="3" pageNumber="30">
|
|
<taxonomicName authorityName="Kroyer" authorityYear="1845" box="[216,461,1498,1537]" class="Malacostraca" family="Ischyroceridae" genus="Siphonoecetes" kingdom="Animalia" order="Amphipoda" pageId="3" pageNumber="30" phylum="Arthropoda" rank="genus">Siphonoecetes</taxonomicName>
|
|
,
|
|
</emphasis>
|
|
and
|
|
<emphasis box="[566,960,1498,1537]" italics="true" pageId="3" pageNumber="30">
|
|
I985-
|
|
<taxonomicName authorityName="Just" authorityYear="1983" box="[661,942,1498,1537]" class="Malacostraca" family="Ischyroceridae" genus="Australoecetes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Amphipoda" pageId="3" pageNumber="30" phylum="Arthropoda" rank="genus">Australoecetes</taxonomicName>
|
|
).
|
|
</emphasis>
|
|
</paragraph>
|
|
<paragraph blockId="3.[464,951,1616,1654]" box="[464,951,1616,1654]" pageId="3" pageNumber="30">Urosomites and the telson</paragraph>
|
|
<paragraph blockId="3.[214,1197,1697,2471]" pageId="3" pageNumber="30">
|
|
In my first contribution to the taxonomy of the
|
|
<taxonomicName authority="(Just, 1983)" baseAuthorityName="Just" baseAuthorityYear="1983" box="[217,729,1743,1782]" class="Malacostraca" family="Ischyroceridae" kingdom="Animalia" order="Amphipoda" pageId="3" pageNumber="30" phylum="Arthropoda" rank="subFamily" subFamily="Siphonoecetinae">
|
|
Siphonoecetinae (
|
|
<bibRefCitation author="Just, J." box="[534,717,1743,1782]" pageId="3" pageNumber="30" pagination="35" refId="ref11748" refString="Just, J., 1983. Siphonoecetinae subfam. n. (Crustacea, Amphipoda, Corophiidae) 1: Classification. Steenstrupia 9 (6): 117 - l 35." type="journal article" year="1983">Just, 1983</bibRefCitation>
|
|
)
|
|
</taxonomicName>
|
|
an important character in the diagnosis of the new genus
|
|
<taxonomicName authorityName="Just" authorityYear="1983" box="[799,1045,1789,1828]" class="Malacostraca" family="Ischyroceridae" genus="Caribboecetes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Amphipoda" pageId="3" pageNumber="30" phylum="Arthropoda" rank="genus">
|
|
<emphasis box="[799,1045,1789,1828]" italics="true" pageId="3" pageNumber="30">Caribboecetes</emphasis>
|
|
</taxonomicName>
|
|
was urosomite 3 and telson fused dorsally". This still holds good for the four species retained in that genus. For the three species of
|
|
<taxonomicName authorityName="Just" authorityYear="1983" box="[503,741,1927,1966]" class="Malacostraca" family="Ischyroceridae" genus="Caribboecetes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Amphipoda" pageId="3" pageNumber="30" phylum="Arthropoda" rank="genus">
|
|
<emphasis box="[503,741,1927,1966]" italics="true" pageId="3" pageNumber="30">Caribboecetes</emphasis>
|
|
</taxonomicName>
|
|
described by
|
|
<bibRefCitation author="Just, J." box="[991,1189,1927,1966]" pageId="3" pageNumber="30" pagination="37 - 64" refId="ref11782" refString="Just, J., 1984 a. Siphonoecetinae (Crustacea, Amphipoda, Corophiidae) 2: Caribboecetes Just, 1983, with description of six new species. Steenstrupia 10 (2): 37 - 64." type="journal article" year="1984">Just (1984a</bibRefCitation>
|
|
, C.
|
|
<emphasis box="[271,486,1973,2012]" italics="true" pageId="3" pageNumber="30">
|
|
<taxonomicName baseAuthorityName="Just" baseAuthorityYear="1984" box="[271,480,1973,2012]" class="Malacostraca" family="Ischyroceridae" genus="Ambicholestes" kingdom="Animalia" order="Amphipoda" pageId="3" pageNumber="30" phylum="Arthropoda" rank="species" species="crassicornis">crassicornis</taxonomicName>
|
|
,
|
|
</emphasis>
|
|
C.
|
|
<emphasis box="[563,741,1973,2012]" italics="true" pageId="3" pageNumber="30">
|
|
<taxonomicName authorityName="Just, I" authorityYear="1984" box="[563,735,1973,2012]" class="Malacostraca" family="Ischyroceridae" genus="Caribboecetes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Amphipoda" pageId="3" pageNumber="30" phylum="Arthropoda" rank="species" species="magellani">magellani</taxonomicName>
|
|
,
|
|
</emphasis>
|
|
and C.
|
|
<emphasis box="[894,1125,1973,2012]" italics="true" pageId="3" pageNumber="30">
|
|
<taxonomicName baseAuthorityName="Just" baseAuthorityYear="1984" box="[894,1107,1973,2012]" class="Malacostraca" family="Ischyroceridae" genus="Ambicholestes" kingdom="Animalia" order="Amphipoda" pageId="3" pageNumber="30" phylum="Arthropoda" rank="species" species="squamiferus">squamiferus</taxonomicName>
|
|
),
|
|
</emphasis>
|
|
but here transferred to
|
|
<taxonomicName authorityName="Just" authorityYear="1998" box="[526,768,2019,2058]" class="Malacostraca" family="Ischyroceridae" genus="Ambicholestes" kingdom="Animalia" order="Amphipoda" pageId="3" pageNumber="30" phylum="Arthropoda" rank="genus" status="n.gen.">
|
|
<emphasis box="[526,768,2019,2058]" italics="true" pageId="3" pageNumber="30">Ambicholestes</emphasis>
|
|
</taxonomicName>
|
|
<taxonomicNameLabel box="[779,882,2019,2058]" pageId="3" pageNumber="30" rank="genus">n.gen.</taxonomicNameLabel>
|
|
, it was noted (Just, I984a: 38) that the urosomite-telson configuration is not intraspecifically stable (occurrence of incomplete fusion between urosomite 3 and the telson, more or less complete fusion between urosomites 1 and 2 and/or 2 and 3). The same lability is found in the Australian concholestids and will be commented upon where relevant. For that reason the urosomite-telson configuration has not been included in the diagnoses of
|
|
<taxonomicName authorityName="Just" authorityYear="1983" box="[724,973,2386,2425]" class="Malacostraca" family="Ischyroceridae" genus="Caribboecetes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Amphipoda" pageId="3" pageNumber="30" phylum="Arthropoda" rank="genus">
|
|
<emphasis box="[724,973,2386,2425]" italics="true" pageId="3" pageNumber="30">Caribboecetes</emphasis>
|
|
</taxonomicName>
|
|
and the new genus group taxa below.
|
|
</paragraph>
|
|
<paragraph blockId="3.[523,890,2550,2589]" box="[523,890,2550,2589]" pageId="3" pageNumber="30">Stridulation organs</paragraph>
|
|
<paragraph blockId="3.[212,1197,2631,3175]" lastBlockId="3.[1265,2251,403,810]" pageId="3" pageNumber="30">
|
|
Three species in
|
|
<taxonomicName authorityName="Just" authorityYear="1998" box="[508,750,2631,2670]" class="Malacostraca" family="Ischyroceridae" genus="Ambicholestes" kingdom="Animalia" order="Amphipoda" pageId="3" pageNumber="30" phylum="Arthropoda" rank="genus" sensu="stricto">
|
|
<emphasis box="[508,750,2631,2670]" italics="true" pageId="3" pageNumber="30">Ambicholestes</emphasis>
|
|
</taxonomicName>
|
|
<taxonomicNameLabel box="[768,821,2631,2670]" pageId="3" pageNumber="30" sensu="stricto">S.S.</taxonomicNameLabel>
|
|
, viz.,
|
|
<emphasis italics="true" pageId="3" pageNumber="30">
|
|
<taxonomicName baseAuthorityName="Just" baseAuthorityYear="1984" box="[930,1187,2631,2670]" class="Malacostraca" family="Ischyroceridae" genus="Ambicholestes" kingdom="Animalia" order="Amphipoda" pageId="3" pageNumber="30" phylum="Arthropoda" rank="species" species="crassicornis">A. crassicornis</taxonomicName>
|
|
,
|
|
<taxonomicName baseAuthorityName="Just" baseAuthorityYear="1984" box="[212,431,2677,2716]" class="Malacostraca" family="Ischyroceridae" genus="Ambicholestes" kingdom="Animalia" order="Amphipoda" pageId="3" pageNumber="30" phylum="Arthropoda" rank="species" species="magellani">A. magellani</taxonomicName>
|
|
,
|
|
</emphasis>
|
|
and
|
|
<taxonomicName authorityName="Just" authorityYear="1998" box="[522,734,2677,2716]" class="Malacostraca" family="Ischyroceridae" genus="Ambicholestes" kingdom="Animalia" order="Amphipoda" pageId="3" pageNumber="30" phylum="Arthropoda" rank="species" species="trilobatus" status="n.sp.">
|
|
<emphasis box="[522,734,2677,2716]" italics="true" pageId="3" pageNumber="30">A. trilobatus</emphasis>
|
|
</taxonomicName>
|
|
<taxonomicNameLabel box="[748,824,2677,2716]" pageId="3" pageNumber="30" rank="species">n.sp.</taxonomicNameLabel>
|
|
possess what appears to be a sound producing organ. Coxal plate 2 carries on the posteroventral margin a row of backward and slightly outward pointing teeth; pereopod 2 article 2 carries medially along the anterior margin a number of short, pointed, proximally inflated, hollow setae, (Just, I984a,
|
|
<figureCitation box="[214,402,2952,2992]" captionStart-0="Fig" captionStart-1="Fig" captionStartId-0="16.[237,358,382,2728]" captionStartId-1="17.[376,431,2223,2259]" captionTargetBox-1="[241,2232,405,2091]" captionTargetPageId-1="17" captionText-0="Fig.13.Ambicholestes(Austrolestes)berentsaen.sp.,holotype." captionText-1="Fig. 14. Ambicholestes (Austrolestes) berentsae n.sp., holotype. co4, apex of coxal plate 4; p, pereopod (3 and 4); us, urosome, dorsal and ventral view." figureDoi-1="http://doi.org/10.5281/zenodo.4656927" httpUri-1="https://zenodo.org/record/4656927/files/figure.png" pageId="3" pageNumber="30">figs 13-14</figureCitation>
|
|
, 16; this paper
|
|
<figureCitation box="[691,800,2952,2992]" captionStart="Fig" captionStartId="11.[376,433,2640,2676]" captionTargetBox="[247,2247,408,2503]" captionTargetPageId="11" captionText="Fig. 8. Ambicholestes (Ambicholestes) trilobatus n.sp., holotype. col, ventral margin of coxal plate 1 with posterior, presumed stridulation teeth; p, pereopod (1, 2, 3 and 5); us, urosome, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.4656917" httpUri="https://zenodo.org/record/4656917/files/figure.png" pageId="3" pageNumber="30">Fig. 8</figureCitation>
|
|
). When pereopod 2 is moved forward the setae come into contact with the coxal plate teeth. This may result in the production of sound. The different arrangement of setae in the three species
|
|
<emphasis italics="true" pageId="3" pageNumber="30">
|
|
(
|
|
<taxonomicName baseAuthorityName="Just" baseAuthorityYear="1984" class="Malacostraca" family="Ischyroceridae" genus="Ambicholestes" kingdom="Animalia" order="Amphipoda" pageId="3" pageNumber="30" phylum="Arthropoda" rank="species" species="crassicornis">A. crassicornis</taxonomicName>
|
|
:
|
|
</emphasis>
|
|
nearly straight row,
|
|
<emphasis box="[817,1054,3136,3175]" italics="true" pageId="3" pageNumber="30">
|
|
<taxonomicName baseAuthorityName="Just" baseAuthorityYear="1984" box="[817,1049,3136,3175]" class="Malacostraca" family="Ischyroceridae" genus="Ambicholestes" kingdom="Animalia" order="Amphipoda" pageId="3" pageNumber="30" phylum="Arthropoda" rank="species" species="magellani">A. magellani</taxonomicName>
|
|
:
|
|
</emphasis>
|
|
on two projecting lobes,
|
|
<emphasis box="[1564,1788,403,442]" italics="true" pageId="3" pageNumber="30">
|
|
<taxonomicName authorityName="Just" authorityYear="1998" box="[1564,1783,403,442]" class="Malacostraca" family="Ischyroceridae" genus="Ambicholestes" kingdom="Animalia" order="Amphipoda" pageId="3" pageNumber="30" phylum="Arthropoda" rank="species" species="trilobatus">A. trilobatus</taxonomicName>
|
|
:
|
|
</emphasis>
|
|
on three projecting lobes) further suggests differences in the sound patterns produced. Stridulation in live animals has not been observed.
|
|
</paragraph>
|
|
<paragraph blockId="3.[1265,2251,403,810]" pageId="3" pageNumber="30">
|
|
In this paper the three species are placed
|
|
<emphasis box="[1974,2250,540,580]" italics="true" pageId="3" pageNumber="30">inAmbicholestes</emphasis>
|
|
s.s. together with
|
|
<emphasis box="[1566,2076,587,626]" italics="true" pageId="3" pageNumber="30">
|
|
<taxonomicName baseAuthorityName="Just" baseAuthorityYear="1984" box="[1566,1813,587,626]" class="Malacostraca" family="Ischyroceridae" genus="Ambicholestes" kingdom="Animalia" order="Amphipoda" pageId="3" pageNumber="30" phylum="Arthropoda" rank="species" species="squamiferus">A. squamiferus</taxonomicName>
|
|
,
|
|
<taxonomicName authorityName="Just" authorityYear="1998" box="[1835,2076,587,626]" class="Malacostraca" family="Ischyroceridae" genus="Ambicholestes" kingdom="Animalia" order="Amphipoda" pageId="3" pageNumber="30" phylum="Arthropoda" rank="species" species="cygnatratus" status="n.sp.">A. cygnatratus</taxonomicName>
|
|
</emphasis>
|
|
<taxonomicNameLabel box="[2093,2171,587,626]" pageId="3" pageNumber="30" rank="species">n.sp.</taxonomicNameLabel>
|
|
and
|
|
<taxonomicName authorityName="Just" authorityYear="1998" box="[1265,1421,633,672]" class="Malacostraca" family="Ischyroceridae" genus="Ambicholestes" kingdom="Animalia" order="Amphipoda" pageId="3" pageNumber="30" phylum="Arthropoda" rank="species" species="poorei" status="n.sp.">
|
|
<emphasis box="[1265,1421,633,672]" italics="true" pageId="3" pageNumber="30">A. poorei</emphasis>
|
|
</taxonomicName>
|
|
<taxonomicNameLabel box="[1434,1511,633,672]" pageId="3" pageNumber="30" rank="species">n.sp.</taxonomicNameLabel>
|
|
for other reasons, but their joint possession of a unique potential stridulation mechanism suggests they form a clade which may eventually warrant separate genus group recognition.
|
|
</paragraph>
|
|
<paragraph blockId="3.[1578,1940,888,927]" box="[1578,1940,888,927]" pageId="3" pageNumber="30">Sexual dimorphism</paragraph>
|
|
<paragraph blockId="3.[1265,2253,969,1607]" pageId="3" pageNumber="30">
|
|
Some groups of siphonoecetines exhibit distinctive sexual dimorphism
|
|
<emphasis box="[1493,2060,1016,1055]" italics="true" pageId="3" pageNumber="30">
|
|
(
|
|
<taxonomicName authority="" box="[1504,2055,1016,1055]" class="Malacostraca" family="Ischyroceridae" genus="Siphonoecetes" kingdom="Animalia" order="Amphipoda" pageId="3" pageNumber="30" phylum="Arthropoda" rank="subGenus" subGenus="Centraloecetes">Siphonoecetes (Centraloecetes)</taxonomicName>
|
|
:
|
|
</emphasis>
|
|
uropod 1,
|
|
<bibRefCitation author="Just, J." box="[1268,1447,1062,1101]" pageId="3" pageNumber="30" pagination="35" refId="ref11748" refString="Just, J., 1983. Siphonoecetinae subfam. n. (Crustacea, Amphipoda, Corophiidae) 1: Classification. Steenstrupia 9 (6): 117 - l 35." type="journal article" year="1983">Just, 1983</bibRefCitation>
|
|
;
|
|
<emphasis box="[1469,2040,1062,1101]" italics="true" pageId="3" pageNumber="30">
|
|
<taxonomicName authority="" box="[1469,2035,1062,1101]" class="Malacostraca" family="Ischyroceridae" genus="Australoecetes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Amphipoda" pageId="3" pageNumber="30" phylum="Arthropoda" rank="subGenus" subGenus="Stebbingoecetes">Australoecetes (Stebbingoecetes)</taxonomicName>
|
|
:
|
|
</emphasis>
|
|
coxal plate 1,
|
|
<bibRefCitation author="Just, J." box="[1314,1492,1108,1147]" pageId="3" pageNumber="30" pagination="325 - 34" refId="ref11860" refString="Just, J., 1985. Siphonoecetinae (Crustacea, Amphipoda, Corophiidae) 4: Australoecetes Just, 1983, including Stebbingoecetes subgen. n. Records of the Australian Museum 37: 325 - 34 l." type="journal article" year="1985">Just, 1985</bibRefCitation>
|
|
;
|
|
<emphasis box="[1513,1731,1108,1147]" italics="true" pageId="3" pageNumber="30">
|
|
<taxonomicName authorityName="Just" authorityYear="1983" box="[1513,1726,1108,1147]" class="Malacostraca" family="Corophiidae" genus="Africoecetes" kingdom="Animalia" order="Amphipoda" pageId="3" pageNumber="30" phylum="Arthropoda" rank="genus">Africoecetes</taxonomicName>
|
|
:
|
|
</emphasis>
|
|
pereopod 2, Just, I984b). No such differences between the sexes have been observed in species of
|
|
<taxonomicName authorityName="Just" authorityYear="1983" box="[1457,1696,1200,1239]" class="Malacostraca" family="Ischyroceridae" genus="Caribboecetes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Amphipoda" pageId="3" pageNumber="30" phylum="Arthropoda" rank="genus">
|
|
<emphasis box="[1457,1696,1200,1239]" italics="true" pageId="3" pageNumber="30">Caribboecetes</emphasis>
|
|
</taxonomicName>
|
|
or
|
|
<emphasis box="[1761,2008,1200,1239]" italics="true" pageId="3" pageNumber="30">
|
|
<taxonomicName authorityName="Just" authorityYear="1998" box="[1761,2003,1200,1239]" class="Malacostraca" family="Ischyroceridae" genus="Ambicholestes" kingdom="Animalia" order="Amphipoda" pageId="3" pageNumber="30" phylum="Arthropoda" rank="genus">Ambicholestes</taxonomicName>
|
|
.
|
|
</emphasis>
|
|
In species of
|
|
<emphasis box="[1265,1539,1246,1285]" italics="true" pageId="3" pageNumber="30">
|
|
<taxonomicName authorityName="Just" authorityYear="1998" box="[1265,1533,1246,1285]" class="Malacostraca" family="Ischyroceridae" genus="Ambicholestes" kingdom="Animalia" order="Amphipoda" pageId="3" pageNumber="30" phylum="Arthropoda" rank="genus">Ambicholestes</taxonomicName>
|
|
,
|
|
</emphasis>
|
|
as in all other siphonoecetines, male antennae are longer, antenna 1 often with more flagellar articles, stouter, and more densely setose than in females; the body length of adult males is greater than in adult females; and females often have fewer robust setae on pereopods and uropods. The number of robust setae mentioned in descriptions below is the maximum number found in adult males, or otherwise in the material available.
|
|
</paragraph>
|
|
</subSubSection>
|
|
</treatment>
|
|
</document> |