105 lines
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105 lines
12 KiB
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<document id="D1E36E4D911220F57AAD7AF16E604325" ID-CLB-Dataset="35790" ID-DOI="http://dx.doi.org/10.3897/dez.63.10748" ID-GBIF-Dataset="d0ba9001-850f-4877-ac8e-b62dd23bff50" ID-Pensoft-Pub="1860-1324-2-287" ID-ZooBank="C09ADD81A914416A93388525B0E35BB0" ModsDocAuthor="" ModsDocDate="2016" ModsDocID="1860-1324-2-287" ModsDocOrigin="Deutsche Entomologische Zeitschrift 63 (2)" ModsDocTitle="Hidden species within the genus Ocys Stephens: the widespread species O.harpaloides (Audinet-Serville) and O.tachysoides (Antoine) (Coleoptera, Carabidae, Bembidiini)" checkinTime="1480571263651" checkinUser="pensoft" docAuthor="Maddison, David R. & Anderson, Roy" docDate="2016" docId="8BB993C0597B586F4951C97A083A6C57" docLanguage="en" docName="DeutEntomolZeit 63(2): 287-301" docOrigin="Deutsche Entomologische Zeitschrift 63 (2)" docSource="http://dx.doi.org/10.3897/dez.63.10748" docTitle="Ocys tachysoides Antoine 1933" docType="treatment" docVersion="6" lastPageNumber="297" masterDocId="FF8FFFA2FFA7B31D52192C54FF9CFFD7" masterDocTitle="Hidden species within the genus Ocys Stephens: the widespread species O. harpaloides (Audinet-Serville) and O. tachysoides (Antoine) (Coleoptera, Carabidae, Bembidiini)" masterLastPageNumber="301" masterPageNumber="287" pageNumber="296" updateTime="1701381165205" updateUser="ExternalLinkService">
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<mods:title id="F676CE3B4D9361ABBA20BB4A4A8C9ECA">Hidden species within the genus Ocys Stephens: the widespread species O. harpaloides (Audinet-Serville) and O. tachysoides (Antoine) (Coleoptera, Carabidae, Bembidiini)</mods:title>
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<mods:namePart id="E7F80D10DB92853CA2EC122C1590A005">Maddison, David R.</mods:namePart>
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<mods:namePart id="FAD77B7D1C4B34626FE27F277BCCE940">Anderson, Roy</mods:namePart>
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<mods:typeOfResource id="B4720F6337D402EBAECDD2FF64533036">text</mods:typeOfResource>
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<mods:title id="91ED494780F66C9B0DFB22D47295765D">Deutsche Entomologische Zeitschrift</mods:title>
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<mods:date id="E5B399500D3BB23F6AB7445157977469">2016</mods:date>
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<mods:number id="4022B6EE34E4F4D2374EA7F2B7810A2D">63</mods:number>
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<mods:number id="405FF336850E2CC18A7D55422E8809F1">2</mods:number>
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<mods:url id="C666AA3C2EE2BA6F2FCFE483E82CECF2">http://dx.doi.org/10.3897/dez.63.10748</mods:url>
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<mods:classification id="167630D190CF81BC6CFAF5541EC2B32F">journal article</mods:classification>
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<mods:identifier id="59AC6BFB88C1A7C41D453F04C10CA6EA" type="DOI">http://dx.doi.org/10.3897/dez.63.10748</mods:identifier>
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<treatment id="8BB993C0597B586F4951C97A083A6C57" ID-GBIF-Taxon="127860011" LSID="urn:lsid:plazi:treatment:8BB993C0597B586F4951C97A083A6C57" httpUri="http://treatment.plazi.org/id/8BB993C0597B586F4951C97A083A6C57" lastPageId="10" lastPageNumber="297" pageId="9" pageNumber="296">
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<subSubSection id="3E71753D34CE5AED7C2C6CD4BFB3CC93" pageId="9" pageNumber="296" type="nomenclature">
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<paragraph id="DC0D20AE7520E062851C7DCCA9EB4C81" pageId="9" pageNumber="296">
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<taxonomicName id="FC5F847556655EBD2C6EF2C1A48F5AA9" ID-CoL="48MN4" authority="Antoine, 1933" authorityName="Antoine" authorityYear="1933" class="Insecta" family="Carabidae" genus="Ocys" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ocys tachysoides" order="Coleoptera" pageId="9" pageNumber="296" phylum="Arthropoda" rank="species" species="tachysoides">
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<pageBreakToken id="6773D350BB65445DA9217160FD195456" pageId="9" pageNumber="296" start="start">Ocys</pageBreakToken>
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tachysoides (Antoine, 1933)
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</taxonomicName>
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Figs 2B; 3B; 6C, D; 7C, D; 8B, 9C, D; 10C, D; 11C, D
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</paragraph>
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</subSubSection>
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<subSubSection id="CDFB1B69439A815D54B67782C88C253D" pageId="10" pageNumber="297" type="reference_group">
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<paragraph id="8E5319025437EF82EFC238D83CBC793A" pageId="10" pageNumber="297">
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<taxonomicName id="A33275CB57604F041200FDA8DC2976F7" class="Insecta" family="Carabidae" genus="Bembidium" higherTaxonomySource="CoL" infraspecific-rank="v." kingdom="Animalia" lsidName="Bembidium harpaloides var. tachysoides" order="Coleoptera" pageId="10" pageNumber="297" phylum="Arthropoda" rank="variety" species="harpaloides" variety="tachysoides">
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<pageBreakToken id="ABBA7BB1841F14C9E2427541C8D95A5A" pageId="10" pageNumber="297" start="start">Bembidium</pageBreakToken>
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harpaloides v. tachysoides
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</taxonomicName>
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Antoine, 1933:79. Holotype female, in the MNHN, examined, with three labels: "O. Nefifik (Maroc) Antoine I.1928" [partly handwritten],
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<normalizedToken id="863FB6C33BC5ED06EF9ACDFFDECB9CE0" originalValue="“Holotype”">"Holotype"</normalizedToken>
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[handwritten on red paper], "
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<taxonomicName id="0A66EDBB9D1AF20E1D61015BEF6D9141" family="Carabidae" genus="Bembidiina" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="Bembidiina tachysoides" order="Coleoptera" pageId="10" pageNumber="297" rank="species" species="tachysoides">tachysoides</taxonomicName>
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Antoine det. m." [partly handwritten]. Type locality: Mouth of the Oued Nefifikh, east of Casablanca, Morocco (in the vicinity of
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<geoCoordinate id="E1CCB5DD04CFBE84C58BCEAE02B25678" direction="north" orientation="latitude" precision="555" value="33.72">33.72°N</geoCoordinate>
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,
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<geoCoordinate id="E3E2F3963574CA3EA93FBAEFC90B0756" direction="west" orientation="longitude" precision="555" value="-7.34">7.34°W</geoCoordinate>
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).
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</paragraph>
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</subSubSection>
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<subSubSection id="F7BCB7FAD8E2DD9664917D99A6389C64" pageId="10" pageNumber="297" type="notes">
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<paragraph id="A97B59BF43F49FD162E4C81A084D9392" pageId="10" pageNumber="297">Nomenclatural notes.</paragraph>
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<paragraph id="5CCF12364355220193A5BAC56129150E" pageId="10" pageNumber="297">
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The type of
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<taxonomicName id="8697FAFE94EAECC06FB8BB5679386E20" class="Insecta" family="Carabidae" genus="Bembidium" higherTaxonomySource="CoL" infraspecific-rank="v." kingdom="Animalia" lsidName="Bembidium harpaloides var. tachysoides" order="Coleoptera" pageId="10" pageNumber="297" phylum="Arthropoda" rank="variety" species="harpaloides" variety="tachysoides">Bembidium harpaloides v. tachysoides</taxonomicName>
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Antoine belongs to this species. We have examined external features of the holotype, including elytral microsculpture (which has a density of 21 lines per 0.1 mm), as well as shape of the spermatheca. Both second gonocoxites are broken in the holotype, and thus we were not able to measure their lengths.
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</paragraph>
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</subSubSection>
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<subSubSection id="577F4CA1FF8BDEFA905B993595E2176C" pageId="10" pageNumber="297" type="diagnosis">
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<paragraph id="500607E634E2D10F874F4502D73B1A27" pageId="10" pageNumber="297">Diagnosis.</paragraph>
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<paragraph id="F5ED5E2B924BCA9AB3FDFE96FED07024" pageId="10" pageNumber="297">
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Body length 4.0-5.8 mm (
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<bibRefCitation id="95B406FC9CFAF3C471607E8C54A24F24" author="Toribio, M" journalOrPublisher="Revista gaditana de Entomologi ́ a" pageId="12" pageNumber="299" pagination="1 - 5" title="Datos sobre algunos Cara ́ bidos de la Peni ́ nsula Ibe ́ rica (Coleoptera)." volume="4" year="2013">Toribio 2013</bibRefCitation>
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and our observations; average length of males 4.79 mm (n=7) of females 5.16 mm (n=7)). Head and pronotum a deep red-brown contrasting with the dark brown to blackish elytra (Fig. 3), only the epipleural gutters and sometimes sutures paler, light reddish-brown. Occasionally with a bluish lustre. Microsculpture of elytral disc less transverse, with more of tendency to form sculpticells (Fig. 7C, D); density of microsculpture lines 20-21 per 0.1 mm (n=4 males). Hind margin of prothorax with a slight emargination laterally, such that the hind margin at the hind angles is directed posteriorly (Fig. 10C, D). Elytra less parallel-sided, greatest width around middle. Basal margin at shoulder in most specimens more or less straight (Fig. 11C), relatively few specimens slightly arcuate with a forward-directed concavity (Fig. 11D). Elytral striae 2 through 4 more marked in the apical third. Aedeagus with ventral margin bend more or less straight (Fig. 6C, D); apex less rounded, blunt. Anterior edge of central sclerite complex of internal sac angulate; brush sclerite smaller; dorsal membranes of internal sac paler, less evident. Gonocoxite relatively short (Fig. 8B), GCR=0.55-0.60 (average 0.58, n=5); spermathecal margin opposite the efferent duct of the spermathecal gland more or less straight (Fig. 8C,D; n=5).
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</paragraph>
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</subSubSection>
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<subSubSection id="6B378743C1B5EA37AD8D15793414624D" pageId="10" pageNumber="297" type="distribution">
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<paragraph id="5F68A78629808D423E901FD38F4D84A2" pageId="10" pageNumber="297">Geographic distribution</paragraph>
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<paragraph id="FD8E30B548F565D969407EE84FCB3DC8" pageId="10" pageNumber="297">
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(Fig. 12). In Africa, known from Morocco (
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<bibRefCitation id="64EDBC8D3F594762FF0383F049343E1E" author="Antoine, M" journalOrPublisher="Memoires de la Societe des sciences naturelles et physiques du Maroc (Nouvelle serie)" pageId="12" pageNumber="299" pagination="1 - 77" title="Coleopteres carabiques du Maroc. Premiere partie." volume="1" year="1955">Antoine 1955</bibRefCitation>
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). In Europe, from Portugal (
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<bibRefCitation id="3BE6A6925A5A065FB784FA2AB34D572F" author="Sciaky, R" journalOrPublisher="Fragmenta Entomologica, Roma" pageId="12" pageNumber="299" pagination="243 - 251" title="Trechusjeannei n. sp. della Spagna settentrionale e note su altre specie di carabidi della Penisola Iberica (Coleoptera, Carabidae)." volume="30" year="1998">Sciaky 1998</bibRefCitation>
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), Spain (
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<bibRefCitation id="E9886A9B538E61E0AB663D176031AD74" author="Toribio, M" journalOrPublisher="Revista gaditana de Entomologi ́ a" pageId="12" pageNumber="299" pagination="1 - 5" title="Datos sobre algunos Cara ́ bidos de la Peni ́ nsula Ibe ́ rica (Coleoptera)." volume="4" year="2013">Toribio 2013</bibRefCitation>
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), France, Belgium, Germany, and the United Kingdom. Examination of specimens in additional collections will likely show it to be more widely distributed.
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</paragraph>
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</subSubSection>
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<subSubSection id="9A9CFDEE1C9B7CD914E187BE23B5206A" pageId="10" pageNumber="297" type="materials_examined">
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<paragraph id="D71CEFF94F198438BB59EA0E210525FF" pageId="10" pageNumber="297">Specimens examined.</paragraph>
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<paragraph id="AD245F9695255EE4AEE89151F36F964E" pageId="10" pageNumber="297">In addition to the type specimen, and those listed in Table 1, we examined specimens from France: Vaires (MNHN).</paragraph>
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</subSubSection>
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</treatment>
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</document> |