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<mods:title id="738A7442AA6A30293E5CC6F57D6B8239">A revision of the Nearctic species of the genus Halobrecta Thomson, 1858 (Coleoptera: Staphylinidae: Aleocharinae) with notes on some Palaearctic species of the genus</mods:title>
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<paragraph id="0D7836B62F112233567FBD2AF090FDC2" blockId="18.[264,650,611,673]" box="[264,650,611,638]" pageId="18" pageNumber="19">
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<taxonomicName id="CAC74D352F112233567FBD2AF090FDC2" ID-CoL="6KZTW" authority="Hardy, 1851" authorityName="Hardy" authorityYear="1851" box="[264,650,611,638]" class="Insecta" family="Staphylinidae" genus="Halobrecta" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="algae">
<emphasis id="3FB3EAA42F112233567FBD2AF090FDC2" bold="true" box="[264,650,611,638]" pageId="18" pageNumber="19">
<emphasis id="3FB3EAA42F112233567FBD2AF3CDFDC1" bold="true" box="[264,471,612,638]" italics="true" pageId="18" pageNumber="19">Halobrecta algae</emphasis>
(
<bibRefCitation id="69564B472F1122335691BD2DF098FDC2" author="Hardy" box="[486,642,611,637]" pageId="18" pageNumber="24" refString="Hardy, J. (1851) Family 1. Staphylinidae, Leach. In: Hardy, J. &amp; Bold, T. J. A Catalogue of the Insects of Northumberland and Durham (Part ii). Transactions of the Tyneside Naturalists' Field Club, 2, 21 - 97." type="journal article" year="1851">Hardy, 1851</bibRefCitation>
)
</emphasis>
</taxonomicName>
</heading>
</paragraph>
<paragraph id="0D7836B62F112233567FBDC9F3BFFD1E" blockId="18.[264,650,611,673]" box="[264,421,647,673]" pageId="18" pageNumber="19">
(
<figureCitation id="95FC2A332F1122335666BDC9F384FD1E" box="[273,414,647,673]" captionStart-0="FIGURES 52 – 55" captionStart-1="FIGURES 56 – 61" captionStartId-0="18.[264,382,1806,1830]" captionStartId-1="19.[264,382,1740,1764]" captionTargetBox-0="[268,1316,933,1806]" captionTargetBox-1="[304,1276,668,1702]" captionTargetId-0="figure@18.[268,1318,922,1806]" captionTargetId-1="figure@19.[287,1300,657,1716]" captionTargetPageId-0="18" captionTargetPageId-1="19" captionText-0="FIGURES 52 – 55. Abdominal segment 8 of Halobrecta algae (Hardy) (male lectotype (52 – 53) and female paralectotype of Homalota puncticeps Thomson (54 – 55 )). 52 — male tergum 8; 53 — male sternum 8; 54 — female tergum 8; 55 — female sternum 8. Scale bar 0.2 mm." captionText-1="FIGURES 56 – 61. Genitalia of Halobrecta algae (Hardy) (male lectotype (56 – 60) and female paralectotype of Homalota puncticeps Thomson (61 )). 56 — median lobe, parameral view; 57 — apex of median lobe, parameral view; 58 — median lobe, lateral view; 59 — apex of median lobe, lateral view, 60 — apex of left paramere, marginal view; 61 — spermatheca. Scale bar 0.1 mm (57, 59 – 61), 0.2 mm (56, 58)." httpUri-0="https://zenodo.org/record/158224/files/figure.png" httpUri-1="https://zenodo.org/record/158225/files/figure.png" pageId="18" pageNumber="19">Figs. 52–61</figureCitation>
)
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<subSubSection id="45DD653D2F112235567FBD99F38DFE39" lastPageId="20" lastPageNumber="21" pageId="18" pageNumber="19" type="reference_group">
<paragraph id="0D7836B62F112233567FBD99F06FFD52" blockId="18.[264,1235,726,880]" box="[264,629,726,750]" pageId="18" pageNumber="19">
<treatmentCitationGroup id="2DD711982F112233567FBD99F06FFD52" box="[264,629,726,750]" pageId="18" pageNumber="19">
<taxonomicName id="CAC74D352F112233567FBD99F06BFD52" ID-CoL="6LZZ7" authority="Hardy, 1851: 78" authorityName="Hardy" authorityPageNumber="78" authorityYear="1851" box="[264,625,726,750]" class="Insecta" family="Staphylinidae" genus="Homalota" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="algae">
<emphasis id="3FB3EAA42F112233567FBD99F3AEFD51" box="[264,436,727,750]" italics="true" pageId="18" pageNumber="19">Homalota algae</emphasis>
<treatmentCitation id="8C6610A72F11223356CCBD98F06BFD52" author="Hardy" box="[443,625,726,749]" page="78" pageId="18" pageNumber="19" year="1851">
<bibRefCitation id="69564B472F11223356CCBD98F059FD52" author="Hardy" box="[443,579,726,749]" pageId="18" pageNumber="24" refString="Hardy, J. (1851) Family 1. Staphylinidae, Leach. In: Hardy, J. &amp; Bold, T. J. A Catalogue of the Insects of Northumberland and Durham (Part ii). Transactions of the Tyneside Naturalists' Field Club, 2, 21 - 97." type="journal article" year="1851">Hardy, 1851</bibRefCitation>
: 78
</treatmentCitation>
</taxonomicName>
.
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</paragraph>
<paragraph id="0D7836B62F112233567FBDB9F0C7FCB2" blockId="18.[264,1235,726,880]" box="[264,733,758,782]" pageId="18" pageNumber="19">
<treatmentCitationGroup id="2DD711982F112233567FBDB9F0C7FCB2" box="[264,733,758,782]" pageId="18" pageNumber="19">
<taxonomicName id="CAC74D352F112233567FBDB9F0C3FCB2" ID-CoL="6MC29" authority="Thomson, 1852: 133" authorityName="Thomson" authorityPageNumber="133" authorityYear="1852" box="[264,729,758,782]" class="Insecta" family="Staphylinidae" genus="Homalota" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="puncticeps">
<emphasis id="3FB3EAA42F112233567FBDB9F3F1FCB1" box="[264,491,759,782]" italics="true" pageId="18" pageNumber="19">Homalota puncticeps</emphasis>
<treatmentCitation id="8C6610A72F1122335685BDB8F0C3FCB2" author="Thomson" box="[498,729,758,781]" page="133" pageId="18" pageNumber="19" year="1852">
<bibRefCitation id="69564B472F1122335685BDB8F087FCB2" author="Thomson" box="[498,669,758,781]" pageId="18" pageNumber="25" refString="Thomson, C. G. (1852) Insekt-slagtet Homalota. Ofversigt af Kongl. Vetenskaps-Akademiens Forkhandlingar, 9 (6), 131 - 146." type="journal article" year="1852">Thomson, 1852</bibRefCitation>
: 133
</treatmentCitation>
</taxonomicName>
.
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</paragraph>
<paragraph id="0D7836B62F112233567FBC59F0F6FC92" blockId="18.[264,1235,726,880]" box="[264,748,790,814]" pageId="18" pageNumber="19">
<treatmentCitationGroup id="2DD711982F112233567FBC59F0F6FC92" box="[264,748,790,814]" pageId="18" pageNumber="19">
<taxonomicName id="CAC74D352F112233567FBC59F0F2FC92" ID-CoL="8Y4LS" authority="Thomson, 1858: 35" authorityName="Thomson" authorityPageNumber="35" authorityYear="1858" box="[264,744,790,814]" class="Insecta" family="Staphylinidae" genus="Halobrecta" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="puncticeps">
<emphasis id="3FB3EAA42F112233567FBC59F3E3FC91" box="[264,505,791,814]" italics="true" pageId="18" pageNumber="19">Halobrecta puncticeps</emphasis>
: 
<treatmentCitation id="8C6610A72F1122335579BC58F0F2FC92" author="Thomson" box="[526,744,790,813]" page="35" pageId="18" pageNumber="19" year="1858">
<bibRefCitation id="69564B472F1122335579BC58F0A3FC92" author="Thomson" box="[526,697,790,813]" pageId="18" pageNumber="25" refString="Thomson, C G. (1858) Forsok till uppstallning af Sveriges Staphyliner. Ofversigt af Kongl. Vetenskaps-Akademiens Forkhandlingar, 15, 27 - 40." type="journal article" year="1858">Thomson, 1858</bibRefCitation>
: 35
</treatmentCitation>
</taxonomicName>
.
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</paragraph>
<paragraph id="0D7836B62F112233567FBC79F6C9FCD0" blockId="18.[264,1235,726,880]" pageId="18" pageNumber="19">
<treatmentCitationGroup id="2DD711982F112233567FBC79F6C9FCD0" pageId="18" pageNumber="19">
<taxonomicName id="CAC74D352F112233567FBC79F0FFFCF2" ID-CoL="8Y4LS" authority="Thomson, 1859: 39" authorityName="Thomson" authorityPageNumber="39" authorityYear="1859" box="[264,741,822,846]" class="Insecta" family="Staphylinidae" genus="Halobrecta" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="puncticeps">
<emphasis id="3FB3EAA42F112233567FBC79F3E3FCF1" box="[264,505,823,846]" italics="true" pageId="18" pageNumber="19">Halobrecta puncticeps</emphasis>
: 
<treatmentCitation id="8C6610A72F1122335579BC78F0FFFCF2" author="Thomson" box="[526,741,822,845]" page="39" pageId="18" pageNumber="19" year="1859">
<bibRefCitation id="69564B472F1122335579BC78F0A3FCF2" author="Thomson" box="[526,697,822,845]" pageId="18" pageNumber="25" refString="Thomson, C. G. (1859) Skandinaviens Coleoptera, synoptiskt bearbetade. Tom 1. Berlingska Boktryckeriet, Lund, v + 290 pp." type="book" year="1859">Thomson, 1859</bibRefCitation>
: 39
</treatmentCitation>
</taxonomicName>
(misspelled as 
<taxonomicName id="CAC74D352F11223354FDBC79F60AFCF1" box="[906,1040,823,846]" class="Insecta" family="Staphylinidae" genus="Halobrectha" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="genus">
<emphasis id="3FB3EAA42F11223354FDBC79F60AFCF1" box="[906,1040,823,846]" italics="true" pageId="18" pageNumber="19">Halobrectha</emphasis>
</taxonomicName>
; as valid species). 
<taxonomicName id="CAC74D352F112233567FBC17F0FFFCD0" ID-CoL="8Y4LS" authority="Thomson, 1861: 49" authorityName="Thomson" authorityPageNumber="49" authorityYear="1861" box="[264,741,856,880]" class="Insecta" family="Staphylinidae" genus="Halobrecta" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="puncticeps">
<emphasis id="3FB3EAA42F112233567FBC17F3E3FCCF" box="[264,505,857,880]" italics="true" pageId="18" pageNumber="19">Halobrecta puncticeps</emphasis>
: 
<treatmentCitation id="8C6610A72F1122335579BC16F0FFFCD0" author="Thomson" box="[526,741,856,879]" page="49" pageId="18" pageNumber="19" year="1861">
<bibRefCitation id="69564B472F1122335579BC16F0A3FCD0" author="Thomson" box="[526,697,856,879]" pageId="18" pageNumber="25" refString="Thomson, C. G. (1861) Skandinaviens Coleoptera, synoptiskt bearbetade. Tom III. Berlingska Boktryckeriet, Lund, 278 pp." type="book" year="1861">Thomson, 1861</bibRefCitation>
: 49
</treatmentCitation>
</taxonomicName>
(misspelled as 
<taxonomicName id="CAC74D352F11223354FDBC17F60AFCCF" box="[906,1040,857,880]" class="Insecta" family="Staphylinidae" genus="Halobrectha" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="genus">
<emphasis id="3FB3EAA42F11223354FDBC17F60AFCCF" box="[906,1040,857,880]" italics="true" pageId="18" pageNumber="19">Halobrectha</emphasis>
</taxonomicName>
; as valid species).
</treatmentCitationGroup>
</paragraph>
<caption id="59B8663E2F112233567FB840F691F8D4" httpUri="https://zenodo.org/record/158224/files/figure.png" pageId="18" pageNumber="19" targetBox="[268,1316,933,1806]" targetPageId="18">
<paragraph id="0D7836B62F112233567FB840F691F8D4" blockId="18.[264,1323,1806,1899]" pageId="18" pageNumber="19">
<emphasis id="3FB3EAA42F112233567FB840F3C8F899" bold="true" box="[264,466,1806,1830]" pageId="18" pageNumber="19">FIGURES 52–55.</emphasis>
Abdominal segment 8 of 
<taxonomicName id="CAC74D352F1122335477B841F601F899" ID-CoL="6KZTW" authority="Hardy" authorityName="Hardy" box="[768,1051,1807,1830]" class="Insecta" family="Staphylinidae" genus="Halobrecta" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="algae">
<emphasis id="3FB3EAA42F1122335477B841F1A7F899" box="[768,957,1807,1830]" italics="true" pageId="18" pageNumber="19">Halobrecta algae</emphasis>
(Hardy)
</taxonomicName>
(male lectotype (52–53) and female paralectotype of 
<taxonomicName id="CAC74D352F112233554EB87CF192F8F7" ID-CoL="6MC29" authority="Thomson" authorityName="Thomson" box="[569,904,1841,1865]" class="Insecta" family="Staphylinidae" genus="Homalota" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="puncticeps">
<emphasis id="3FB3EAA42F112233554EB87CF107F8F6" box="[569,797,1842,1865]" italics="true" pageId="18" pageNumber="19">Homalota puncticeps</emphasis>
Thomson
</taxonomicName>
(54–55)). 52 — male tergum 8; 53 — male sternum 8; 54 — female tergum 8; 55 — female sternum 8. Scale bar 0.2 mm.
</paragraph>
</caption>
<paragraph id="0D7836B62F102232567FBE52F36DFEEC" blockId="19.[264,1323,284,531]" pageId="19" pageNumber="20">
<treatmentCitationGroup id="2DD711982F102232567FBE52F36DFEEC" pageId="19" pageNumber="20">
<taxonomicName id="CAC74D352F102232567FBE52F110FE8C" authority="Fenyes, 1920: 185" authorityName="Fenyes" authorityPageNumber="185" authorityYear="1920" box="[264,778,284,307]" class="Insecta" family="Staphylinidae" genus="Atheta" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="algae" subGenus="Halobrecta">
<emphasis id="3FB3EAA42F102232567FBE52F357FE8C" box="[264,333,284,307]" italics="true" pageId="19" pageNumber="20">Atheta</emphasis>
(
<emphasis id="3FB3EAA42F1022325628BE52F3CDFE8C" box="[351,471,284,307]" italics="true" pageId="19" pageNumber="20">Halobrecta</emphasis>
) 
<emphasis id="3FB3EAA42F102232569FBE52F039FE8C" box="[488,547,284,307]" italics="true" pageId="19" pageNumber="20">algae</emphasis>
: 
<treatmentCitation id="8C6610A72F102232554CBE52F110FE8C" author="Fenyes" box="[571,778,284,307]" page="185" pageId="19" pageNumber="20" year="1920">
<bibRefCitation id="69564B472F102232554CBE52F0D4FE8C" author="Fenyes" box="[571,718,284,307]" pageId="19" pageNumber="23" refString="Fenyes, A. (1920) Coleoptera. Fam. Staphylinidae, subfam. Aleocharinae. In: Wytsman, P. (Ed.), Genera Insectorum, Fasc. 173 B. L. Desmet-Verteneuil, Bruxelles, pp. 111 - 414." type="book chapter" year="1920">Fenyes, 1920</bibRefCitation>
: 185
</treatmentCitation>
</taxonomicName>
(as synonym both of 
<taxonomicName id="CAC74D352F102232548FBE52F69DFE8C" box="[1016,1159,284,307]" class="Insecta" family="Staphylinidae" genus="Homalota" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="flaviceps">
<emphasis id="3FB3EAA42F102232548FBE52F69DFE8C" box="[1016,1159,284,307]" italics="true" pageId="19" pageNumber="20">Ha. flaviceps</emphasis>
</taxonomicName>
and 
<taxonomicName id="CAC74D352F10223253CABE52F37DFEEC" class="Insecta" family="Staphylinidae" genus="Homalota" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="pucticeps">
<emphasis id="3FB3EAA42F10223253CABE52F37DFEEC" italics="true" pageId="19" pageNumber="20">Ha. pucticeps</emphasis>
</taxonomicName>
).
</treatmentCitationGroup>
</paragraph>
<paragraph id="0D7836B62F102232567FBE12F1E7FECC" blockId="19.[264,1323,284,531]" box="[264,1021,348,371]" pageId="19" pageNumber="20">
<treatmentCitationGroup id="2DD711982F102232567FBE12F1E7FECC" box="[264,1021,348,371]" pageId="19" pageNumber="20">
<taxonomicName id="CAC74D352F102232567FBE12F12DFECC" authority="Fenyes, 1920: 185" authorityName="Fenyes" authorityPageNumber="185" authorityYear="1920" box="[264,823,348,371]" class="Insecta" family="Staphylinidae" genus="Atheta" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="puncticeps" subGenus="Halobrecta">
<emphasis id="3FB3EAA42F102232567FBE12F357FECC" box="[264,333,348,371]" italics="true" pageId="19" pageNumber="20">Atheta</emphasis>
(
<emphasis id="3FB3EAA42F102232562ABE12F3CFFECC" box="[349,469,348,371]" italics="true" pageId="19" pageNumber="20">Halobrecta</emphasis>
) 
<emphasis id="3FB3EAA42F1022325692BE12F04DFECC" box="[485,599,348,371]" italics="true" pageId="19" pageNumber="20">puncticeps</emphasis>
: 
<treatmentCitation id="8C6610A72F102232551BBE12F12DFECC" author="Fenyes" box="[620,823,348,371]" page="185" pageId="19" pageNumber="20" year="1920">
<bibRefCitation id="69564B472F102232551BBE12F0E4FECC" author="Fenyes" box="[620,766,348,371]" pageId="19" pageNumber="23" refString="Fenyes, A. (1920) Coleoptera. Fam. Staphylinidae, subfam. Aleocharinae. In: Wytsman, P. (Ed.), Genera Insectorum, Fasc. 173 B. L. Desmet-Verteneuil, Bruxelles, pp. 111 - 414." type="book chapter" year="1920">Fenyes, 1920</bibRefCitation>
: 185
</treatmentCitation>
</taxonomicName>
(as valid species).
</treatmentCitationGroup>
</paragraph>
<paragraph id="0D7836B62F102232567FBE32F00FFE0C" blockId="19.[264,1323,284,531]" pageId="19" pageNumber="20">
<treatmentCitationGroup id="2DD711982F102232567FBE32F00FFE0C" pageId="19" pageNumber="20">
<taxonomicName id="CAC74D352F102232567FBE32F1D9FE2C" authority="Bernhauer &amp; Scheerpeltz, 1926: 621" authorityName="Bernhauer &amp; Scheerpeltz" authorityPageNumber="621" authorityYear="1926" box="[264,963,380,403]" class="Insecta" family="Staphylinidae" genus="Atheta" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="algae" subGenus="Halobrecta">
<emphasis id="3FB3EAA42F102232567FBE32F357FE2C" box="[264,333,380,403]" italics="true" pageId="19" pageNumber="20">Atheta</emphasis>
(
<emphasis id="3FB3EAA42F102232562ABE32F3CFFE2C" box="[349,469,380,403]" italics="true" pageId="19" pageNumber="20">Halobrecta</emphasis>
) 
<emphasis id="3FB3EAA42F1022325692BE32F03AFE2C" box="[485,544,380,403]" italics="true" pageId="19" pageNumber="20">algae</emphasis>
: 
<treatmentCitation id="8C6610A72F1022325542BE32F1D9FE2C" author="Bernhauer" box="[565,963,380,403]" page="621" pageId="19" pageNumber="20" year="1926">
<bibRefCitation id="69564B472F1022325542BE32F190FE2C" author="Bernhauer" box="[565,906,380,403]" pageId="19" pageNumber="23" refString="Bernhauer, M. &amp; Scheerpeltz, O. (1926) Staphylinidae VI. In: Junk, W. &amp; Schenkling, S. (Eds.), Coleopterorum Catalogus, Pars 82. W. Junk, Berlin, pp. 499 - 988." type="journal article" year="1926">Bernhauer &amp; Scheerpeltz, 1926</bibRefCitation>
: 621
</treatmentCitation>
</taxonomicName>
(as synonym both of 
<taxonomicName id="CAC74D352F10223253DDBE32F731FE2C" ID-CoL="6KZTS" box="[1194,1323,380,403]" class="Insecta" family="Staphylinidae" genus="Halobrecta" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="flavipes">
<emphasis id="3FB3EAA42F10223253DDBE32F731FE2C" box="[1194,1323,380,403]" italics="true" pageId="19" pageNumber="20">Ha. flavipes</emphasis>
</taxonomicName>
and 
<taxonomicName id="CAC74D352F1022325612BED2F01FFE0C" box="[357,517,412,435]" class="Insecta" family="Staphylinidae" genus="Atheta" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="puncticeps">
<emphasis id="3FB3EAA42F1022325612BED2F01FFE0C" box="[357,517,412,435]" italics="true" pageId="19" pageNumber="20">Ha. puncticeps</emphasis>
</taxonomicName>
).
</treatmentCitationGroup>
</paragraph>
<paragraph id="0D7836B62F102232567FBEF2F6A4FE6C" blockId="19.[264,1323,284,531]" box="[264,1214,444,467]" pageId="19" pageNumber="20">
<treatmentCitationGroup id="2DD711982F102232567FBEF2F6A4FE6C" box="[264,1214,444,467]" pageId="19" pageNumber="20">
<taxonomicName id="CAC74D352F102232567FBEF2F1E2FE6C" authority="Bernhauer &amp; Scheerpeltz, 1926: 621" authorityName="Bernhauer &amp; Scheerpeltz" authorityPageNumber="621" authorityYear="1926" box="[264,1016,444,467]" class="Insecta" family="Staphylinidae" genus="Atheta" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="puncticeps" subGenus="Halobrecta">
<emphasis id="3FB3EAA42F102232567FBEF2F357FE6C" box="[264,333,444,467]" italics="true" pageId="19" pageNumber="20">Atheta</emphasis>
(
<emphasis id="3FB3EAA42F102232562ABEF2F3CFFE6C" box="[349,469,444,467]" italics="true" pageId="19" pageNumber="20">Halobrecta</emphasis>
) 
<emphasis id="3FB3EAA42F1022325692BEF2F04DFE6C" box="[485,599,444,467]" italics="true" pageId="19" pageNumber="20">puncticeps</emphasis>
: 
<treatmentCitation id="8C6610A72F102232551BBEF2F1E2FE6C" author="Bernhauer" box="[620,1016,444,467]" page="621" pageId="19" pageNumber="20" year="1926">
<bibRefCitation id="69564B472F102232551BBEF2F1DAFE6C" author="Bernhauer" box="[620,960,444,467]" pageId="19" pageNumber="23" refString="Bernhauer, M. &amp; Scheerpeltz, O. (1926) Staphylinidae VI. In: Junk, W. &amp; Schenkling, S. (Eds.), Coleopterorum Catalogus, Pars 82. W. Junk, Berlin, pp. 499 - 988." type="journal article" year="1926">Bernhauer &amp; Scheerpeltz, 1926</bibRefCitation>
: 621
</treatmentCitation>
</taxonomicName>
(as valid species).
</treatmentCitationGroup>
</paragraph>
<paragraph id="0D7836B62F102232567FBE92F618FE4C" blockId="19.[264,1323,284,531]" box="[264,1026,476,499]" pageId="19" pageNumber="20">
<treatmentCitationGroup id="2DD711982F102232567FBE92F618FE4C" box="[264,1026,476,499]" pageId="19" pageNumber="20">
<taxonomicName id="CAC74D352F102232567FBE92F126FE4C" ID-CoL="8Y4LS" authority="Benick &amp; Lohse, 1974: 219" authorityName="Benick &amp; Lohse" authorityPageNumber="219" authorityYear="1974" box="[264,828,476,499]" class="Insecta" family="Staphylinidae" genus="Halobrecta" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="puncticeps">
<emphasis id="3FB3EAA42F102232567FBE92F3E3FE4C" box="[264,505,476,499]" italics="true" pageId="19" pageNumber="20">Halobrecta puncticeps</emphasis>
: 
<treatmentCitation id="8C6610A72F1022325579BE92F126FE4C" author="Benick" box="[526,828,476,499]" page="219" pageId="19" pageNumber="20" year="1974">
<bibRefCitation id="69564B472F1022325579BE92F119FE4C" author="Benick" box="[526,771,476,499]" pageId="19" pageNumber="23" refString="Benick, G. &amp; Lohse, G. A. (1974) 14. Tribus: Callicerini (Athetae). In: Freude, H., Harde, K. W. &amp; Lohse, G. A. (Eds.), Die Kafer Mitteleuropas. Band 5, Staphylinidae II (Hypocyphtinae und Aleocharinae). Pselaphidae. Goecke &amp; Evers Verlag, Krefeld, pp. 72 - 220." type="book chapter" year="1974">Benick &amp; Lohse, 1974</bibRefCitation>
: 219
</treatmentCitation>
</taxonomicName>
(as valid species).
</treatmentCitationGroup>
</paragraph>
<paragraph id="0D7836B62F102232567FBEB2F197FDAC" blockId="19.[264,1323,284,531]" box="[264,909,508,531]" pageId="19" pageNumber="20">
<treatmentCitationGroup id="2DD711982F102232567FBEB2F197FDAC" box="[264,909,508,531]" pageId="19" pageNumber="20">
<taxonomicName id="CAC74D352F102232567FBEB2F0DDFDAC" ID-CoL="6KZTW" authority="Hammond, 2000: 275" authorityName="Hammond" authorityPageNumber="275" authorityYear="2000" box="[264,711,508,531]" class="Insecta" family="Staphylinidae" genus="Halobrecta" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="algae">
<emphasis id="3FB3EAA42F102232567FBEB2F3D8FDAC" box="[264,450,508,531]" italics="true" pageId="19" pageNumber="20">Halobrecta algae</emphasis>
: 
<treatmentCitation id="8C6610A72F10223256A0BEB2F0DDFDAC" author="Hammond" box="[471,711,508,531]" page="275" pageId="19" pageNumber="20" year="2000">
<bibRefCitation id="69564B472F10223256A0BEB2F094FDAC" author="Hammond" box="[471,654,508,531]" lastPageId="19" lastPageNumber="24" pageId="19" pageNumber="23" refString="Hammond, P. M. (2000) 16. Coastal Staphylinidae (rove beetles) in the British Isles, with special reference to saltmarshes. In: Sherwood, B. R., Gardiner, B. G. &amp; Harris T. (Eds.), British Saltmarshes. Joint Symposium on British Saltmarshes organized between the Linnean Society of London, the Royal Society for the Protection of Birds and English Nature (2000: London). Forrest Text, Cardingan, pp. 247 - 302." type="journal article" year="2000">Hammond, 2000</bibRefCitation>
: 275
</treatmentCitation>
</taxonomicName>
(as valid species).
</treatmentCitationGroup>
</paragraph>
<paragraph id="0D7836B62F102235567FBD0AF38DFE39" blockId="19.[264,580,580,603]" lastBlockId="20.[264,1323,283,990]" lastPageId="20" lastPageNumber="21" pageId="19" pageNumber="20">
(other references are omitted) 
<emphasis id="3FB3EAA42F172235567FBE55F3A4FE8A" bold="true" box="[264,446,283,309]" pageId="20" pageNumber="21">
<typeStatus id="D27C88142F172235567FBE55F359FE8A" box="[264,323,283,309]" pageId="20" pageNumber="21">Type</typeStatus>
material.
</emphasis>
<typeStatus id="D27C88142F17223556A3BE52F053FE89" box="[468,585,284,310]" pageId="20" pageNumber="21" type="lectotype">Lectotype</typeStatus>
of 
<taxonomicName id="CAC74D352F1722355502BE52F16BFE8A" ID-CoL="6MC29" box="[629,881,284,309]" class="Insecta" family="Staphylinidae" genus="Homalota" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="puncticeps">
<emphasis id="3FB3EAA42F1722355502BE52F16BFE8A" box="[629,881,284,309]" italics="true" pageId="20" pageNumber="21">Homalota puncticeps</emphasis>
</taxonomicName>
(here designated): ɗ, &quot;Lā [Lomma, 
<collectingCountry id="75D076262F172235567FBE0AF371FEE1" box="[264,363,324,350]" name="Sweden" pageId="20" pageNumber="21">Sweden</collectingCountry>
]&quot; (MZLU). 
<typeStatus id="D27C88142F1722355577BE0AF0B5FEE1" box="[512,687,324,350]" pageId="20" pageNumber="21" type="paralectotype">Paralectotypes</typeStatus>
: ɗ, 2ΨΨ, &quot;Lā [Lomma, 
<collectingCountry id="75D076262F17223554A4BE0AF62CFEE1" box="[979,1078,324,350]" name="Sweden" pageId="20" pageNumber="21">Sweden</collectingCountry>
]&quot; (MZLU (Thomson collection)).
</paragraph>
</subSubSection>
<caption id="59B8663E2F102232567FB982F00EF8D1" httpUri="https://zenodo.org/record/158225/files/figure.png" pageId="19" pageNumber="20" targetBox="[304,1276,668,1702]" targetPageId="19">
<paragraph id="0D7836B62F102232567FB982F00EF8D1" blockId="19.[264,1323,1740,1902]" pageId="19" pageNumber="20">
<emphasis id="3FB3EAA42F102232567FB982F3C8F95B" bold="true" box="[264,466,1740,1764]" pageId="19" pageNumber="20">FIGURES 56–61.</emphasis>
Genitalia of 
<taxonomicName id="CAC74D352F1022325502B982F189F95C" authority="Hardy" authorityName="Hardy" box="[629,915,1740,1763]" class="Insecta" family="Staphylinidae" genus="Halobrecta" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="algae">
<emphasis id="3FB3EAA42F1022325502B982F12EF95C" box="[629,820,1740,1763]" italics="true" pageId="19" pageNumber="20">Halobrecta algae</emphasis>
(Hardy)
</taxonomicName>
(male lectotype (56–60) and female paralectotype of 
<taxonomicName id="CAC74D352F10223256CAB9A1F114F8B9" authority="Thomson" authorityName="Thomson" box="[445,782,1775,1798]" class="Insecta" family="Staphylinidae" genus="Homalota" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="puncticeps">
<emphasis id="3FB3EAA42F10223256CAB9A1F0B8F8B9" box="[445,674,1775,1798]" italics="true" pageId="19" pageNumber="20">Homalota puncticeps</emphasis>
Thomson
</taxonomicName>
(61)). 56 — median lobe, parameral view; 57 — apex of median lobe, parameral view; 58 — median lobe, lateral view; 59 — apex of median lobe, lateral view, 60 — apex of left paramere, marginal view; 61 — spermatheca. Scale bar 0.1 mm (57, 59–61), 0.2 mm (56, 58).
</paragraph>
</caption>
<subSubSection id="45DD653D2F172235564FBEDDF36FFE69" pageId="20" pageNumber="21" type="materials_examined">
<paragraph id="0D7836B62F172235564FBEDDF36FFE69" blockId="20.[264,1323,283,990]" pageId="20" pageNumber="21">
<emphasis id="3FB3EAA42F172235564FBEDDF0FEFE12" bold="true" box="[312,740,403,429]" pageId="20" pageNumber="21">
Additional material. 
<collectingCountry id="75D076262F172235552CBEDDF0C5FE12" box="[603,735,403,429]" name="Sweden" pageId="20" pageNumber="21">SWEDEN</collectingCountry>
:
</emphasis>
ɗ, Ψ, Oskarshamn, 
<date id="797910762F1722355489BEDAF661FE11" box="[1022,1147,404,430]" pageId="20" pageNumber="21" value="1941-06-10">10.vi.1941</date>
(H.Bachlund) (
<collectionCode id="6BD6AE732F1722355667BEF2F372FE69" box="[272,360,444,470]" pageId="20" pageNumber="21">KSEM</collectionCode>
).
</paragraph>
</subSubSection>
<subSubSection id="45DD653D2F172235564FBEADF343FD99" pageId="20" pageNumber="21" type="diagnosis">
<paragraph id="0D7836B62F172235564FBEADF343FD99" blockId="20.[264,1323,283,990]" pageId="20" pageNumber="21">
<emphasis id="3FB3EAA42F172235564FBEADF3ADFE42" bold="true" box="[312,439,483,509]" pageId="20" pageNumber="21">Diagnosis.</emphasis>
See 
<bibRefCitation id="69564B472F1722355688BEAAF12EFE41" author="Benick" box="[511,820,484,510]" pageId="20" pageNumber="23" refString="Benick, G. &amp; Lohse, G. A. (1974) 14. Tribus: Callicerini (Athetae). In: Freude, H., Harde, K. W. &amp; Lohse, G. A. (Eds.), Die Kafer Mitteleuropas. Band 5, Staphylinidae II (Hypocyphtinae und Aleocharinae). Pselaphidae. Goecke &amp; Evers Verlag, Krefeld, pp. 72 - 220." type="book chapter" year="1974">Benick and Lohse (1974)</bibRefCitation>
. Male tergum 8 and aedeagus: 
<figureCitation id="95FC2A332F17223553B5BEAAF73CFE41" box="[1218,1318,484,510]" captionStart="FIGURES 52 – 55" captionStartId="18.[264,382,1806,1830]" captionTargetBox="[268,1316,933,1806]" captionTargetId="figure@18.[268,1318,922,1806]" captionTargetPageId="18" captionText="FIGURES 52 – 55. Abdominal segment 8 of Halobrecta algae (Hardy) (male lectotype (52 – 53) and female paralectotype of Homalota puncticeps Thomson (54 – 55 )). 52 — male tergum 8; 53 — male sternum 8; 54 — female tergum 8; 55 — female sternum 8. Scale bar 0.2 mm." httpUri="https://zenodo.org/record/158224/files/figure.png" pageId="20" pageNumber="21">Figs. 52</figureCitation>
, 
<figureCitation id="95FC2A332F172235567FBD42F34EFD99" box="[264,340,524,550]" captionStart="FIGURES 56 – 61" captionStartId="19.[264,382,1740,1764]" captionTargetBox="[304,1276,668,1702]" captionTargetId="figure@19.[287,1300,657,1716]" captionTargetPageId="19" captionText="FIGURES 56 – 61. Genitalia of Halobrecta algae (Hardy) (male lectotype (56 – 60) and female paralectotype of Homalota puncticeps Thomson (61 )). 56 — median lobe, parameral view; 57 — apex of median lobe, parameral view; 58 — median lobe, lateral view; 59 — apex of median lobe, lateral view, 60 — apex of left paramere, marginal view; 61 — spermatheca. Scale bar 0.1 mm (57, 59 – 61), 0.2 mm (56, 58)." httpUri="https://zenodo.org/record/158225/files/figure.png" pageId="20" pageNumber="21">56–60</figureCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="45DD653D2F172235564FBD7DF148FC61" pageId="20" pageNumber="21" type="discussion">
<paragraph id="0D7836B62F172235564FBD7DF148FC61" blockId="20.[264,1323,283,990]" pageId="20" pageNumber="21">
<emphasis id="3FB3EAA42F172235564FBD7DF3D9FDF2" bold="true" box="[312,451,563,589]" pageId="20" pageNumber="21">Discussion.</emphasis>
The 
<typeStatus id="D27C88142F1722355564BD7AF064FDF1" box="[531,638,564,590]" pageId="20" pageNumber="21" type="lectotype">lectotype</typeStatus>
of 
<taxonomicName id="CAC74D352F17223555DABD7AF17BFDF2" box="[685,865,564,589]" class="Insecta" family="Staphylinidae" genus="Homalota" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="puncticeps">
<emphasis id="3FB3EAA42F17223555DABD7AF17BFDF2" box="[685,865,564,589]" italics="true" pageId="20" pageNumber="21">Ho. puncticeps</emphasis>
</taxonomicName>
agrees with accepted concept of 
<taxonomicName id="CAC74D352F1722355388BD7AF353FDCA" class="Insecta" family="Staphylinidae" genus="Halobrecta" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="algae">
<emphasis id="3FB3EAA42F1722355388BD7AF353FDCA" italics="true" pageId="20" pageNumber="21">Ha. algae</emphasis>
</taxonomicName>
(see 
<bibRefCitation id="69564B472F17223556F0BD12F096FDC9" author="Benick" box="[391,652,604,630]" pageId="20" pageNumber="23" refString="Benick, G. &amp; Lohse, G. A. (1974) 14. Tribus: Callicerini (Athetae). In: Freude, H., Harde, K. W. &amp; Lohse, G. A. (Eds.), Die Kafer Mitteleuropas. Band 5, Staphylinidae II (Hypocyphtinae und Aleocharinae). Pselaphidae. Goecke &amp; Evers Verlag, Krefeld, pp. 72 - 220." type="book chapter" year="1974">Benick &amp; Lohse 1974</bibRefCitation>
; as 
<taxonomicName id="CAC74D352F17223555CFBD12F172FDCA" box="[696,872,604,629]" class="Insecta" family="Staphylinidae" genus="Homalota" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="puncticeps">
<emphasis id="3FB3EAA42F17223555CFBD12F172FDCA" box="[696,872,604,629]" italics="true" pageId="20" pageNumber="21">Ha. puncticeps</emphasis>
</taxonomicName>
). 
<taxonomicName id="CAC74D352F17223554FFBD12F68AFDCA" box="[904,1168,604,629]" class="Insecta" family="Staphylinidae" genus="Halobrecta" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="puncticeps">
<emphasis id="3FB3EAA42F17223554FFBD12F68AFDCA" box="[904,1168,604,629]" italics="true" pageId="20" pageNumber="21">Halobrecta puncticeps</emphasis>
</taxonomicName>
has not been recorded outside Europe, unlike 
<taxonomicName id="CAC74D352F17223555EABDCAF15FFD22" box="[669,837,644,669]" class="Insecta" family="Staphylinidae" genus="Halobrecta" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="algophila">
<emphasis id="3FB3EAA42F17223555EABDCAF15FFD22" box="[669,837,644,669]" italics="true" pageId="20" pageNumber="21">Ha. algophila</emphasis>
</taxonomicName>
, 
<taxonomicName id="CAC74D352F172235542DBDCAF1F4FD22" box="[858,1006,644,669]" class="Insecta" family="Staphylinidae" genus="Halobrecta" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="flavipes">
<emphasis id="3FB3EAA42F172235542DBDCAF1F4FD22" box="[858,1006,644,669]" italics="true" pageId="20" pageNumber="21">Ha. flavipes</emphasis>
</taxonomicName>
and 
<emphasis id="3FB3EAA42F1722355344BDCAF739FD22" box="[1075,1315,644,669]" italics="true" pageId="20" pageNumber="21">
Ha. sp. 
<taxonomicName id="CAC74D352F17223553EDBDCAF739FD22" box="[1178,1315,644,669]" class="Insecta" family="Staphylinidae" genus="Halobrecta" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="halensis">cf. halensis</taxonomicName>
</emphasis>
. However, considering the wide distribution ranges of the last three species, it is possible that 
<taxonomicName id="CAC74D352F172235564BBD9AF3F6FD52" box="[316,492,724,749]" class="Insecta" family="Staphylinidae" genus="Halobrecta" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="puncticeps">
<emphasis id="3FB3EAA42F172235564BBD9AF3F6FD52" box="[316,492,724,749]" italics="true" pageId="20" pageNumber="21">Ha. puncticeps</emphasis>
</taxonomicName>
has also been introduced to other parts of the world. Examined Mediterranean specimens, externally similar to 
<taxonomicName id="CAC74D352F1722355476BDB2F16DFCAA" box="[769,887,764,789]" class="Insecta" family="Staphylinidae" genus="Halobrecta" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="algae">
<emphasis id="3FB3EAA42F1722355476BDB2F16DFCAA" box="[769,887,764,789]" italics="true" pageId="20" pageNumber="21">Ha. algae</emphasis>
</taxonomicName>
, differ in the shape of the aedeagus, particularly in having a narrower median lobe (in parameral view: cf. 
<figureCitation id="95FC2A332F1722355336BC6AF68EFC81" box="[1089,1172,804,830]" captionStart="FIGURES 62 – 65" captionStartId="20.[264,383,1746,1770]" captionTargetBox="[328,1216,1040,1731]" captionTargetId="figure@20.[320,1227,1026,1736]" captionTargetPageId="20" captionText="FIGURES 62 – 65. Details of Halobrecta sp. aff. algae (Hardy) (male, Grado, Italy (62 – 63 )) and Ha. sp. cf. halensis Mulsant &amp; Rey, 1873 (male, Lignano, Italy (64 – 65 )). 62 — median lobe with everted internal sac, parameral view; 63 — median lobe with everted internal sac, lateral view; 64 — median lobe with partially everted internal sac, parameral view; 65 — median lobe with partially everted internal sac, lateral view. Scale bar 0.2 mm." httpUri="https://zenodo.org/record/158226/files/figure.png" pageId="20" pageNumber="21">Fig. 62</figureCitation>
and 
<figureCitation id="95FC2A332F17223553BABC6AF738FC81" box="[1229,1314,804,830]" captionStart="FIGURES 56 – 61" captionStartId="19.[264,382,1740,1764]" captionTargetBox="[304,1276,668,1702]" captionTargetId="figure@19.[287,1300,657,1716]" captionTargetPageId="19" captionText="FIGURES 56 – 61. Genitalia of Halobrecta algae (Hardy) (male lectotype (56 – 60) and female paralectotype of Homalota puncticeps Thomson (61 )). 56 — median lobe, parameral view; 57 — apex of median lobe, parameral view; 58 — median lobe, lateral view; 59 — apex of median lobe, lateral view, 60 — apex of left paramere, marginal view; 61 — spermatheca. Scale bar 0.1 mm (57, 59 – 61), 0.2 mm (56, 58)." httpUri="https://zenodo.org/record/158225/files/figure.png" pageId="20" pageNumber="21">Fig. 56</figureCitation>
) and in less bent median lobe (in lateral view: cf. 
<figureCitation id="95FC2A332F1722355421BC02F1B0FCD9" box="[854,938,844,870]" captionStart="FIGURES 62 – 65" captionStartId="20.[264,383,1746,1770]" captionTargetBox="[328,1216,1040,1731]" captionTargetId="figure@20.[320,1227,1026,1736]" captionTargetPageId="20" captionText="FIGURES 62 – 65. Details of Halobrecta sp. aff. algae (Hardy) (male, Grado, Italy (62 – 63 )) and Ha. sp. cf. halensis Mulsant &amp; Rey, 1873 (male, Lignano, Italy (64 – 65 )). 62 — median lobe with everted internal sac, parameral view; 63 — median lobe with everted internal sac, lateral view; 64 — median lobe with partially everted internal sac, parameral view; 65 — median lobe with partially everted internal sac, lateral view. Scale bar 0.2 mm." httpUri="https://zenodo.org/record/158226/files/figure.png" pageId="20" pageNumber="21">Fig. 63</figureCitation>
and 
<figureCitation id="95FC2A332F1722355491BC02F624FCD9" box="[998,1086,844,870]" captionStart="FIGURES 56 – 61" captionStartId="19.[264,382,1740,1764]" captionTargetBox="[304,1276,668,1702]" captionTargetId="figure@19.[287,1300,657,1716]" captionTargetPageId="19" captionText="FIGURES 56 – 61. Genitalia of Halobrecta algae (Hardy) (male lectotype (56 – 60) and female paralectotype of Homalota puncticeps Thomson (61 )). 56 — median lobe, parameral view; 57 — apex of median lobe, parameral view; 58 — median lobe, lateral view; 59 — apex of median lobe, lateral view, 60 — apex of left paramere, marginal view; 61 — spermatheca. Scale bar 0.1 mm (57, 59 – 61), 0.2 mm (56, 58)." httpUri="https://zenodo.org/record/158225/files/figure.png" pageId="20" pageNumber="21">Fig. 58</figureCitation>
). These Mediterranean specimens are likely to belong to an undecribed species. Revision of European species of 
<taxonomicName id="CAC74D352F1722355628BCD2F3FEFC0A" box="[351,484,924,949]" class="Insecta" family="Staphylinidae" genus="Halobrecta" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="21" phylum="Arthropoda" rank="genus">
<emphasis id="3FB3EAA42F1722355628BCD2F3FEFC0A" box="[351,484,924,949]" italics="true" pageId="20" pageNumber="21">Halobrecta</emphasis>
</taxonomicName>
is outside the scope of this paper. A revision of the British species of 
<taxonomicName id="CAC74D352F172235567FBC8AF396FC62" box="[264,396,964,989]" class="Insecta" family="Staphylinidae" genus="Halobrecta" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="21" phylum="Arthropoda" rank="genus">
<emphasis id="3FB3EAA42F172235567FBC8AF396FC62" box="[264,396,964,989]" italics="true" pageId="20" pageNumber="21">Halobrecta</emphasis>
</taxonomicName>
is being prepared by Peter Hammond.
</paragraph>
</subSubSection>
</treatment>
</document>