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<mods:title id="1E3B10D100C5EBC2427FF2A430D5C1E5">A revision of the Nearctic species of the genus Halobrecta Thomson, 1858 (Coleoptera: Staphylinidae: Aleocharinae) with notes on some Palaearctic species of the genus</mods:title>
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(
<figureCitation id="95FC2A332F0222205666BA07F394FADC" box="[273,398,1353,1379]" captionStart-0="FIGURES 1 – 5" captionStart-1="FIGURES 6 – 12" captionStart-10="FIGURES 52 – 55" captionStart-11="FIGURES 56 – 61" captionStart-12="FIGURES 62 – 65" captionStart-2="FIGURES 13 – 17" captionStart-3="FIGURES 18 – 21" captionStart-4="FIGURES 22 – 29" captionStart-5="FIGURES 30 – 36" captionStart-6="FIGURES 37 – 40" captionStart-7="FIGURES 41 – 44" captionStart-8="FIGURES 45 – 48" captionStart-9="FIGURES 49 – 51" captionStartId-0="4.[264,383,1512,1536]" captionStartId-1="5.[264,383,1714,1738]" captionStartId-10="18.[264,382,1806,1830]" captionStartId-11="19.[264,382,1740,1764]" captionStartId-12="20.[264,383,1746,1770]" captionStartId-2="6.[264,382,1493,1517]" captionStartId-3="9.[264,382,1204,1228]" captionStartId-4="10.[264,382,1447,1471]" captionStartId-5="11.[264,382,1692,1716]" captionStartId-6="13.[264,382,1414,1438]" captionStartId-7="14.[264,382,1781,1805]" captionStartId-8="15.[264,382,1419,1443]" captionStartId-9="16.[264,382,1105,1129]" captionTargetBox-0="[294,1291,289,1499]" captionTargetBox-1="[312,1278,649,1705]" captionTargetBox-10="[268,1316,933,1806]" captionTargetBox-11="[304,1276,668,1702]" captionTargetBox-12="[328,1216,1040,1731]" captionTargetBox-2="[453,1127,301,1448]" captionTargetBox-3="[292,1295,296,1174]" captionTargetBox-4="[297,1292,297,1431]" captionTargetBox-5="[304,1285,459,1679]" captionTargetBox-6="[292,1310,552,1389]" captionTargetBox-7="[346,1257,1081,1771]" captionTargetBox-8="[298,1297,593,1401]" captionTargetBox-9="[346,1246,419,1083]" captionTargetId-0="figure@4.[291,1296,283,1505]" captionTargetId-1="figure@5.[299,1288,635,1710]" captionTargetId-10="figure@18.[268,1318,922,1806]" captionTargetId-11="figure@19.[287,1300,657,1716]" captionTargetId-12="figure@20.[320,1227,1026,1736]" captionTargetId-2="figure@6.[441,1145,290,1457]" captionTargetId-3="figure@9.[283,1304,284,1181]" captionTargetId-4="figure@10.[285,1301,286,1440]" captionTargetId-5="figure@11.[295,1292,446,1690]" captionTargetId-6="figure@13.[280,1321,538,1399]" captionTargetId-7="figure@14.[339,1264,1071,1778]" captionTargetId-8="figure@15.[281,1306,582,1409]" captionTargetId-9="figure@16.[331,1257,409,1090]" captionTargetPageId-0="4" captionTargetPageId-1="5" captionTargetPageId-10="18" captionTargetPageId-11="19" captionTargetPageId-12="20" captionTargetPageId-2="6" captionTargetPageId-3="9" captionTargetPageId-4="10" captionTargetPageId-5="11" captionTargetPageId-6="13" captionTargetPageId-7="14" captionTargetPageId-8="15" captionTargetPageId-9="16" captionText-0="FIGURES 1 – 5. Mouthparts of Halobrecta sp. aff. algae (Hardy) (female, Grado, Italy). 1 — labrum; 2 — epipharynx; 3 — left mandible, dorsal view; 4 — left mandible, ventral view; 5 — right mandible, dorsal view. Scale bar 0.1 mm." captionText-1="FIGURES 6 – 12. Mouthparts of Halobrecta sp. aff. algae (Hardy) (female, Grado, Italy (6 – 11 )) and Ha. sp. cf. halensis Mulsant &amp; Rey, 1873 (female, Lignano, Italy (12 )). 6 — prementum; 7 — hypopharynx; 8 — right maxilla, ventral view; 9 — right galea, ventral view; 10 — right lacinia, ventral view; 11 — right lacinia, dorsal view; 12 — mentum. Scale bar 0.1 mm (6 – 7, 9 – 12), 0.2 mm (8)." captionText-10="FIGURES 52 – 55. Abdominal segment 8 of Halobrecta algae (Hardy) (male lectotype (52 – 53) and female paralectotype of Homalota puncticeps Thomson (54 – 55 )). 52 — male tergum 8; 53 — male sternum 8; 54 — female tergum 8; 55 — female sternum 8. Scale bar 0.2 mm." captionText-11="FIGURES 56 – 61. Genitalia of Halobrecta algae (Hardy) (male lectotype (56 – 60) and female paralectotype of Homalota puncticeps Thomson (61 )). 56 — median lobe, parameral view; 57 — apex of median lobe, parameral view; 58 — median lobe, lateral view; 59 — apex of median lobe, lateral view, 60 — apex of left paramere, marginal view; 61 — spermatheca. Scale bar 0.1 mm (57, 59 – 61), 0.2 mm (56, 58)." captionText-12="FIGURES 62 – 65. Details of Halobrecta sp. aff. algae (Hardy) (male, Grado, Italy (62 – 63 )) and Ha. sp. cf. halensis Mulsant &amp; Rey, 1873 (male, Lignano, Italy (64 – 65 )). 62 — median lobe with everted internal sac, parameral view; 63 — median lobe with everted internal sac, lateral view; 64 — median lobe with partially everted internal sac, parameral view; 65 — median lobe with partially everted internal sac, lateral view. Scale bar 0.2 mm." captionText-2="FIGURES 13 – 17. Details of Halobrecta sp. aff. algae (Hardy) (female, Grado, Italy). 13 — right antenna; 14 — pronotum; 15 — meso­ and metathorax; 16 — left metatarsal segment 5 and metapretarsus, anterior view; 17 — right metatarsus, posterior view. Scale bar 0.1 mm (16), 0.2 mm (13, 17), 0.4 mm (14 – 15)." captionText-3="FIGURES 18 – 21. Abdominal segment 8 of Halobrecta algophila (Fenyes) (male lectotype (18 – 19) and female paralectotype of At. algophila (20 – 21 )). 18 — male tergum 8; 19 — male sternum 8; 20 — female tergum 8; 21 — female sternum 8. Scale bar 0.2 mm." captionText-4="FIGURES 22 – 29. Aedeagus of Halobrecta algophila (Fenyes) (male lectotype of At. algophila (22 – 25), and male from the environs of Bolinas, California (26 – 29 )). 22, 27 — apex of median lobe, parameral view; 23, 26 — apex of median lobe, parameral view; 25, 28 — median lobe, lateral view; 24, 29 — apex of median lobe, lateral view. Scale bar 0.1 mm (22, 24, 27, 29), 0.2 mm (23, 25, 26, 28)." captionText-5="FIGURES 30 – 36. Genitalia of Halobrecta algophila (Fenyes) (male lectotype (30 – 31, 33 – 36) and female paralectotype of At. algophila (32 )). 30 — details of retracted internal sac, abparameral view; 31 — details of retracted internal sac, lateral view; 32 — spermatheca; 33 — left paramere, view from the side facing the median lobe; 34 — left paramere, marginal view; 35 — left paramere, view from the side opposite to the median lobe; 36 — apex of left paramere, marginal view. Scale bar 0.1 mm (32, 36), 0.2 mm (30 – 31, 33 – 35)." captionText-6="FIGURES 37 – 40. Abdominal segment 8 of Halobrecta flavipes Thomson (male (37 – 38) and female paralectotypes Atheta pocahontas Casey (39 – 40 )). 37 — male tergum 8; 38 — male sternum 8; 39 — female tergum 8; 40 — female sternum 8. Scale bar 0.2 mm." captionText-7="FIGURES 41 – 44. Aedeagus of Halobrecta flavipes Thomson (male from Oskarshamn, Sweden (41 – 42), and male paralectotype of Atheta pocahontas Casey (43 – 44 )). 41, 43 — median lobe, parameral view; 42, 44 — apex of median lobe, parameral view. Scale bar 0.1 mm (42, 44), 0.2 mm (41, 43)." captionText-8="FIGURES 45 – 48. Aedeagus of Halobrecta flavipes Thomson (males, from Oskarshamn, Sweden (45 – 46), and lectotype of Atheta pocahontas Casey (47 – 48 )). 45, 47 — median lobe, lateral view; 46, 48 — apex of median lobe, lateral view. Scale bar 0.1 mm (46, 48), 0.2 mm (45, 47)." captionText-9="FIGURES 49 – 51. Genitalia of Halobrecta flavipes Thomson (male (49 – 50) and female paralectotypes of Atheta pocahontas (51 )). 49 — details of retracted internal sac, lateral view; 50 — apex of left paramere, marginal view; 51 — spermatheca. Scale bar 0.1 mm (50 – 51), 0.2 mm (49)." httpUri-0="https://zenodo.org/record/158214/files/figure.png" httpUri-1="https://zenodo.org/record/158215/files/figure.png" httpUri-10="https://zenodo.org/record/158224/files/figure.png" httpUri-11="https://zenodo.org/record/158225/files/figure.png" httpUri-12="https://zenodo.org/record/158226/files/figure.png" httpUri-2="https://zenodo.org/record/158216/files/figure.png" httpUri-3="https://zenodo.org/record/158217/files/figure.png" httpUri-4="https://zenodo.org/record/158218/files/figure.png" httpUri-5="https://zenodo.org/record/158219/files/figure.png" httpUri-6="https://zenodo.org/record/158220/files/figure.png" httpUri-7="https://zenodo.org/record/158221/files/figure.png" httpUri-8="https://zenodo.org/record/158222/files/figure.png" httpUri-9="https://zenodo.org/record/158223/files/figure.png" pageId="1" pageNumber="2">Figs. 1–65</figureCitation>
)
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<emphasis id="3FB3EAA42F022220567FBAD4F39AFA0E" box="[264,384,1434,1457]" italics="true" pageId="1" pageNumber="2">Halobrecta</emphasis>
<treatmentCitation id="8C6610A72F02222056F0BAD7F046FA0F" author="Thomson" box="[391,604,1433,1456]" page="35" pageId="1" pageNumber="2" year="1858">
<bibRefCitation id="69564B472F02222056F0BAD7F02BFA0F" author="Thomson" box="[391,561,1433,1456]" pageId="1" pageNumber="25" refString="Thomson, C G. (1858) Forsok till uppstallning af Sveriges Staphyliner. Ofversigt af Kongl. Vetenskaps-Akademiens Forkhandlingar, 15, 27 - 40." type="journal article" year="1858">Thomson, 1858</bibRefCitation>
: 35
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(
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species 
<taxonomicName id="CAC74D352F0222205585BAD4F69EFA0F" ID-CoL="6MC29" authority="Thomson, 1852" authorityName="Thomson" authorityYear="1852" box="[754,1156,1433,1457]" class="Insecta" family="Staphylinidae" genus="Homalota" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="2" phylum="Arthropoda" rank="species" species="puncticeps">
<emphasis id="3FB3EAA42F0222205585BAD4F1CEFA0E" box="[754,980,1434,1457]" italics="true" pageId="1" pageNumber="2">Homalota puncticeps</emphasis>
<bibRefCitation id="69564B472F02222054ACBAD7F69EFA0F" author="Thomson" box="[987,1156,1433,1456]" pageId="1" pageNumber="25" refString="Thomson, C. G. (1852) Insekt-slagtet Homalota. Ofversigt af Kongl. Vetenskaps-Akademiens Forkhandlingar, 9 (6), 131 - 146." type="journal article" year="1852">Thomson, 1852</bibRefCitation>
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, by monotypy).
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<taxonomicName id="CAC74D352F022220567FBAF4F060FA6F" authority="Thomson, 1861: 49" authorityName="Thomson" authorityPageNumber="49" authorityYear="1861" box="[264,634,1465,1489]" class="Insecta" family="Staphylinidae" genus="Halobrectha" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="2" phylum="Arthropoda" rank="genus">
<emphasis id="3FB3EAA42F022220567FBAF4F394FA6E" box="[264,398,1466,1489]" italics="true" pageId="1" pageNumber="2">Halobrectha</emphasis>
: 
<treatmentCitation id="8C6610A72F02222056D5BAF7F060FA6F" author="Thomson" box="[418,634,1465,1488]" page="49" pageId="1" pageNumber="2" year="1861">
<bibRefCitation id="69564B472F02222056D5BAF7F057FA6F" author="Thomson" box="[418,589,1465,1488]" pageId="1" pageNumber="25" refString="Thomson, C. G. (1861) Skandinaviens Coleoptera, synoptiskt bearbetade. Tom III. Berlingska Boktryckeriet, Lund, 278 pp." type="book" year="1861">Thomson, 1861</bibRefCitation>
: 49
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(as valid genus; incorrect subsequent spelling).
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<emphasis id="3FB3EAA42F022220567FBA94F379FA4E" box="[264,355,1498,1521]" italics="true" pageId="1" pageNumber="2">Glaphya</emphasis>
Mulsant &amp; 
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<bibRefCitation id="69564B472F0222205684BA97F072FA4F" author="Rey" box="[499,616,1497,1520]" pageId="1" pageNumber="24" refString="Muslant, E. &amp; Rey, C. (1873) Description de divers Coleopteres Brevipennes nouveaux ou peu connus. Opuscules Entomologiques, 15, 147 - 189." type="journal article" year="1873">Rey, 1873</bibRefCitation>
: 172
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</taxonomicName>
(as subgenus of 
<taxonomicName id="CAC74D352F0222205405BA94F689FA4F" ID-CoL="8VRML" authority="Thomson, 1858" authorityName="Thomson" authorityYear="1858" box="[882,1171,1497,1521]" class="Insecta" family="Staphylinidae" genus="Dinaraea" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="2" phylum="Arthropoda" rank="genus">
<emphasis id="3FB3EAA42F0222205405BA94F1C2FA4E" box="[882,984,1498,1521]" italics="true" pageId="1" pageNumber="2">Dinaraea</emphasis>
<bibRefCitation id="69564B472F0222205493BA97F689FA4F" author="Thomson" box="[996,1171,1497,1520]" pageId="1" pageNumber="25" refString="Thomson, C G. (1858) Forsok till uppstallning af Sveriges Staphyliner. Ofversigt af Kongl. Vetenskaps-Akademiens Forkhandlingar, 15, 27 - 40." type="journal article" year="1858">Thomson, 1858</bibRefCitation>
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; 
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species 
<taxonomicName id="CAC74D352F022220564FBAB4F0C8F9AF" authority="Mulsant &amp; Rey, 1873" authorityName="Mulsant &amp; Rey" authorityYear="1873" box="[312,722,1529,1553]" class="Insecta" family="Staphylinidae" genus="Dinaraea" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="2" phylum="Arthropoda" rank="species" species="pubes">
<emphasis id="3FB3EAA42F022220564FBAB4F3F9F9AE" box="[312,483,1530,1553]" italics="true" pageId="1" pageNumber="2">Dinaraea pubes</emphasis>
Mulsant &amp; 
<bibRefCitation id="69564B472F0222205514BAB7F0C8F9AF" author="Rey" box="[611,722,1529,1552]" pageId="1" pageNumber="24" refString="Muslant, E. &amp; Rey, C. (1873) Description de divers Coleopteres Brevipennes nouveaux ou peu connus. Opuscules Entomologiques, 15, 147 - 189." type="journal article" year="1873">Rey, 1873</bibRefCitation>
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, by monotypy).
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<treatmentCitationGroup id="2DD711982F022220567FB954F6A9F98F" box="[264,1203,1561,1585]" pageId="1" pageNumber="2">
<taxonomicName id="CAC74D352F022220567FB954F0ADF98F" authority="Mulsant &amp; Rey, 1875: 35" authorityName="Mulsant &amp; Rey" authorityPageNumber="35" authorityYear="1875" box="[264,695,1561,1585]" class="Insecta" family="Staphylinidae" genus="Halobrechta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="2" phylum="Arthropoda" rank="genus">
<emphasis id="3FB3EAA42F022220567FB954F394F98E" box="[264,398,1562,1585]" italics="true" pageId="1" pageNumber="2">Halobrechta</emphasis>
: Mulsant &amp; 
<treatmentCitation id="8C6610A72F022220556BB957F0ADF98F" author="Rey" box="[540,695,1561,1584]" page="35" pageId="1" pageNumber="2" year="1875">
<bibRefCitation id="69564B472F022220556BB957F091F98F" author="Rey" box="[540,651,1561,1584]" pageId="1" pageNumber="24" refString="Muslant, E. &amp; Rey, C. (1875) Tribu des Brevipennes. Famille des Aleochariens (suite). Septieme branche: Myrmedoniaires. 2 e partie. Annales de la Societe d'Agriculture, Histoire Naturelle et Arts Utiles de Lyon, serie 4, 7 [1874], 27 - 496, Pl. vi - ix." type="journal article" year="1875">Rey, 1875</bibRefCitation>
: 35
</treatmentCitation>
</taxonomicName>
(as valid genus; incorrect subsequent spelling).
</treatmentCitationGroup>
</paragraph>
<paragraph id="0D7836B62F022220567FB974F180F9EF" blockId="1.[264,1323,1433,1905]" box="[264,922,1593,1617]" pageId="1" pageNumber="2">
<treatmentCitationGroup id="2DD711982F022220567FB974F180F9EF" box="[264,922,1593,1617]" pageId="1" pageNumber="2">
<taxonomicName id="CAC74D352F022220567FB974F3FBF9EF" ID-CoL="8VRBL" authority="(Halobrecta)" baseAuthorityName="Halobrecta" box="[264,481,1593,1617]" class="Insecta" family="Staphylinidae" genus="Atheta" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="2" phylum="Arthropoda" rank="genus">
<emphasis id="3FB3EAA42F022220567FB974F357F9EE" box="[264,333,1594,1617]" italics="true" pageId="1" pageNumber="2">Atheta</emphasis>
(
<emphasis id="3FB3EAA42F022220562AB974F3CFF9EE" box="[349,469,1594,1617]" italics="true" pageId="1" pageNumber="2">Halobrecta</emphasis>
)
</taxonomicName>
: 
<treatmentCitation id="8C6610A72F0222205684B977F0A4F9EF" author="Fenyes" box="[499,702,1593,1616]" page="185" pageId="1" pageNumber="2" year="1920">
<bibRefCitation id="69564B472F0222205684B977F09FF9EF" author="Fenyes" box="[499,645,1593,1616]" pageId="1" pageNumber="23" refString="Fenyes, A. (1920) Coleoptera. Fam. Staphylinidae, subfam. Aleocharinae. In: Wytsman, P. (Ed.), Genera Insectorum, Fasc. 173 B. L. Desmet-Verteneuil, Bruxelles, pp. 111 - 414." type="book chapter" year="1920">Fenyes, 1920</bibRefCitation>
: 185
</treatmentCitation>
(as valid subgenus).
</treatmentCitationGroup>
</paragraph>
<paragraph id="0D7836B62F022220567FB914F1C1F9CF" blockId="1.[264,1323,1433,1905]" box="[264,987,1625,1649]" pageId="1" pageNumber="2">
<treatmentCitationGroup id="2DD711982F022220567FB914F1C1F9CF" box="[264,987,1625,1649]" pageId="1" pageNumber="2">
<taxonomicName id="CAC74D352F022220567FB914F059F9CF" authority="Fenyes, 1920: 185" authorityName="Fenyes" authorityPageNumber="185" authorityYear="1920" box="[264,579,1625,1649]" class="Insecta" family="Staphylinidae" genus="Glaphya" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="2" phylum="Arthropoda" rank="genus">
<emphasis id="3FB3EAA42F022220567FB914F37EF9CE" box="[264,356,1626,1649]" italics="true" pageId="1" pageNumber="2">Glaphya</emphasis>
: 
<treatmentCitation id="8C6610A72F022220560EB917F059F9CF" author="Fenyes" box="[377,579,1625,1648]" page="185" pageId="1" pageNumber="2" year="1920">
<bibRefCitation id="69564B472F022220560EB917F011F9CF" author="Fenyes" box="[377,523,1625,1648]" pageId="1" pageNumber="23" refString="Fenyes, A. (1920) Coleoptera. Fam. Staphylinidae, subfam. Aleocharinae. In: Wytsman, P. (Ed.), Genera Insectorum, Fasc. 173 B. L. Desmet-Verteneuil, Bruxelles, pp. 111 - 414." type="book chapter" year="1920">Fenyes, 1920</bibRefCitation>
: 185
</treatmentCitation>
</taxonomicName>
(as synonym of 
<taxonomicName id="CAC74D352F0222205582B914F121F9CE" ID-CoL="8VRBL" box="[757,827,1626,1649]" class="Insecta" family="Staphylinidae" genus="Atheta" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="2" phylum="Arthropoda" rank="genus">
<emphasis id="3FB3EAA42F0222205582B914F121F9CE" box="[757,827,1626,1649]" italics="true" pageId="1" pageNumber="2">Atheta</emphasis>
</taxonomicName>
(
<taxonomicName id="CAC74D352F022220543DB914F1D9F9CE" ID-CoL="8VRQB" box="[842,963,1626,1649]" class="Insecta" family="Staphylinidae" genus="Halobrecta" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="2" phylum="Arthropoda" rank="genus">
<emphasis id="3FB3EAA42F022220543DB914F1D9F9CE" box="[842,963,1626,1649]" italics="true" pageId="1" pageNumber="2">Halobrecta</emphasis>
</taxonomicName>
)).
</treatmentCitationGroup>
</paragraph>
<paragraph id="0D7836B62F022220567FB934F084F90F" blockId="1.[264,1323,1433,1905]" pageId="1" pageNumber="2">
<treatmentCitationGroup id="2DD711982F022220567FB934F084F90F" pageId="1" pageNumber="2">
<taxonomicName id="CAC74D352F022220567FB934F046F92F" ID-CoL="8W25D" authority="Cameron, 1920: 265" authorityName="Cameron" authorityPageNumber="265" authorityYear="1920" box="[264,604,1657,1681]" class="Insecta" family="Staphylinidae" genus="Exatheta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="2" phylum="Arthropoda" rank="genus">
<emphasis id="3FB3EAA42F022220567FB934F37DF92E" box="[264,359,1658,1681]" italics="true" pageId="1" pageNumber="2">Exatheta</emphasis>
<treatmentCitation id="8C6610A72F0222205606B937F046F92F" author="Cameron" box="[369,604,1657,1680]" page="265" pageId="1" pageNumber="2" year="1920">
<bibRefCitation id="69564B472F0222205606B937F006F92F" author="Cameron" box="[369,540,1657,1680]" pageId="1" pageNumber="23" refString="Cameron, M. (1920) New species of Staphylinidae from Singapore. Part III. Transactions of the Entomological Society of London, 1920, 212 - 284." type="journal article" year="1920">
<collectingCountry id="75D076262F0222205606B937F3CCF92F" box="[369,470,1657,1680]" name="Cameroon" pageId="1" pageNumber="2">Cameron</collectingCountry>
, 1920
</bibRefCitation>
: 265
</treatmentCitation>
</taxonomicName>
(
<typeStatus id="D27C88142F0222205519B937F087F92F" box="[622,669,1657,1680]" pageId="1" pageNumber="2">type</typeStatus>
species 
<taxonomicName id="CAC74D352F0222205588B934F69BF92F" ID-CoL="3DJSQ" authority="Cameron, 1920" authorityName="Cameron" authorityYear="1920" box="[767,1153,1657,1681]" class="Insecta" family="Staphylinidae" genus="Exatheta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="2" phylum="Arthropoda" rank="species" species="cingulata">
<emphasis id="3FB3EAA42F0222205588B934F1D7F92E" box="[767,973,1658,1681]" italics="true" pageId="1" pageNumber="2">Exatheta cingulata</emphasis>
<bibRefCitation id="69564B472F02222054A1B937F69BF92F" author="Cameron" box="[982,1153,1657,1680]" pageId="1" pageNumber="23" refString="Cameron, M. (1920) New species of Staphylinidae from Singapore. Part III. Transactions of the Entomological Society of London, 1920, 212 - 284." type="journal article" year="1920">
<collectingCountry id="75D076262F02222054A1B937F620F92F" box="[982,1082,1657,1680]" name="Cameroon" pageId="1" pageNumber="2">Cameron</collectingCountry>
, 1920
</bibRefCitation>
</taxonomicName>
, by subsequent designation 
<bibRefCitation id="69564B472F02222056CEB9D7F080F90F" author="Blackwelder" box="[441,666,1689,1712]" pageId="1" pageNumber="23" refString="Blackwelder, R. E. (1952) The generic names of the beetle family Staphylinidae with an essay on genotypy. United States National Museum Bulletin, 200, i - iv + 483 pp." type="book" year="1952">(Blackwelder 1952))</bibRefCitation>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="0D7836B62F022220567FB9F4F3D0F94F" blockId="1.[264,1323,1433,1905]" pageId="1" pageNumber="2">
<treatmentCitationGroup id="2DD711982F022220567FB9F4F3D0F94F" pageId="1" pageNumber="2">
<taxonomicName id="CAC74D352F022220567FB9F4F3FBF96F" ID-CoL="8VRBL" authority="(Halobrecta)" baseAuthorityName="Halobrecta" box="[264,481,1721,1745]" class="Insecta" family="Staphylinidae" genus="Atheta" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="2" phylum="Arthropoda" rank="genus">
<emphasis id="3FB3EAA42F022220567FB9F4F357F96E" box="[264,333,1722,1745]" italics="true" pageId="1" pageNumber="2">Atheta</emphasis>
(
<emphasis id="3FB3EAA42F022220562AB9F4F3CFF96E" box="[349,469,1722,1745]" italics="true" pageId="1" pageNumber="2">Halobrecta</emphasis>
)
</taxonomicName>
: 
<treatmentCitation id="8C6610A72F0222205683B9F7F199F96F" author="Bernhauer" box="[500,899,1721,1744]" page="621" pageId="1" pageNumber="2" year="1926">
<bibRefCitation id="69564B472F0222205683B9F7F153F96F" author="Bernhauer" box="[500,841,1721,1744]" pageId="1" pageNumber="23" refString="Bernhauer, M. &amp; Scheerpeltz, O. (1926) Staphylinidae VI. In: Junk, W. &amp; Schenkling, S. (Eds.), Coleopterorum Catalogus, Pars 82. W. Junk, Berlin, pp. 499 - 988." type="journal article" year="1926">Bernhauer &amp; Scheerpeltz, 1926</bibRefCitation>
: 621
</treatmentCitation>
(as valid subgenus in subtribe 
<taxonomicName id="CAC74D352F02222053B8B9F7F3A4F94F" ID-CoL="8VRC4" authority="Casey, 1910" authorityName="Casey" authorityYear="1910" class="Insecta" family="Staphylinidae" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="2" phylum="Arthropoda" rank="subTribe" subTribe="Athetina">
Athetina 
<bibRefCitation id="69564B472F022220564FB997F3A4F94F" author="Casey" box="[312,446,1753,1776]" pageId="1" pageNumber="23" refString="Casey, T. L. (1910) New Species of the Staphylinid Tribe Myrmedoniini. Memoirs on the Coleoptera I. The New Era Printing Company, Lancaster, pp. 1 - 183." type="book chapter" year="1910">Casey, 1910</bibRefCitation>
</taxonomicName>
).
</treatmentCitationGroup>
</paragraph>
<paragraph id="0D7836B62F022220567FB9B4F687F8AF" blockId="1.[264,1323,1433,1905]" box="[264,1181,1785,1809]" pageId="1" pageNumber="2">
<treatmentCitationGroup id="2DD711982F022220567FB9B4F687F8AF" box="[264,1181,1785,1809]" pageId="1" pageNumber="2">
<taxonomicName id="CAC74D352F022220567FB9B4F11CF8AF" authority="Bernhauer &amp; Scheerpeltz, 1926: 621" authorityName="Bernhauer &amp; Scheerpeltz" authorityPageNumber="621" authorityYear="1926" box="[264,774,1785,1809]" class="Insecta" family="Staphylinidae" genus="Glaphya" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="2" phylum="Arthropoda" rank="genus">
<emphasis id="3FB3EAA42F022220567FB9B4F37EF8AE" box="[264,356,1786,1809]" italics="true" pageId="1" pageNumber="2">Glaphya</emphasis>
: 
<treatmentCitation id="8C6610A72F022220560FB9B7F11CF8AF" author="Bernhauer" box="[376,774,1785,1808]" page="621" pageId="1" pageNumber="2" year="1926">
<bibRefCitation id="69564B472F022220560FB9B7F0D7F8AF" author="Bernhauer" box="[376,717,1785,1808]" pageId="1" pageNumber="23" refString="Bernhauer, M. &amp; Scheerpeltz, O. (1926) Staphylinidae VI. In: Junk, W. &amp; Schenkling, S. (Eds.), Coleopterorum Catalogus, Pars 82. W. Junk, Berlin, pp. 499 - 988." type="journal article" year="1926">Bernhauer &amp; Scheerpeltz, 1926</bibRefCitation>
: 621
</treatmentCitation>
</taxonomicName>
(as synonym of 
<taxonomicName id="CAC74D352F02222054C0B9B4F1E7F8AE" ID-CoL="8VRBL" box="[951,1021,1786,1809]" class="Insecta" family="Staphylinidae" genus="Atheta" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="2" phylum="Arthropoda" rank="genus">
<emphasis id="3FB3EAA42F02222054C0B9B4F1E7F8AE" box="[951,1021,1786,1809]" italics="true" pageId="1" pageNumber="2">Atheta</emphasis>
</taxonomicName>
(
<taxonomicName id="CAC74D352F022220537BB9B4F69EF8AE" ID-CoL="8VRQB" box="[1036,1156,1786,1809]" class="Insecta" family="Staphylinidae" genus="Halobrecta" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="2" phylum="Arthropoda" rank="genus">
<emphasis id="3FB3EAA42F022220537BB9B4F69EF8AE" box="[1036,1156,1786,1809]" italics="true" pageId="1" pageNumber="2">Halobrecta</emphasis>
</taxonomicName>
)).
</treatmentCitationGroup>
</paragraph>
<paragraph id="0D7836B62F022220567FB854F367F8EF" blockId="1.[264,1323,1433,1905]" pageId="1" pageNumber="2">
<treatmentCitationGroup id="2DD711982F022220567FB854F367F8EF" pageId="1" pageNumber="2">
<taxonomicName id="CAC74D352F022220567FB854F116F88F" ID-CoL="8W25D" authority="Bernhauer &amp; Scheerpeltz, 1926: 681" authorityName="Bernhauer &amp; Scheerpeltz" authorityPageNumber="681" authorityYear="1926" box="[264,780,1817,1841]" class="Insecta" family="Staphylinidae" genus="Exatheta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="2" phylum="Arthropoda" rank="genus">
<emphasis id="3FB3EAA42F022220567FB854F37DF88E" box="[264,359,1818,1841]" italics="true" pageId="1" pageNumber="2">Exatheta</emphasis>
: 
<treatmentCitation id="8C6610A72F022220560BB857F116F88F" author="Bernhauer" box="[380,780,1817,1840]" page="681" pageId="1" pageNumber="2" year="1926">
<bibRefCitation id="69564B472F022220560BB857F0C8F88F" author="Bernhauer" box="[380,722,1817,1840]" pageId="1" pageNumber="23" refString="Bernhauer, M. &amp; Scheerpeltz, O. (1926) Staphylinidae VI. In: Junk, W. &amp; Schenkling, S. (Eds.), Coleopterorum Catalogus, Pars 82. W. Junk, Berlin, pp. 499 - 988." type="journal article" year="1926">Bernhauer &amp; Scheerpeltz, 1926</bibRefCitation>
: 681
</treatmentCitation>
</taxonomicName>
(as valid genus in subtribe Schistogeniina 
<bibRefCitation id="69564B472F02222053AFB857F36BF8EF" author="Fenyes" pageId="1" pageNumber="23" refString="Fenyes, A. (1918) Coleoptera. Fam. Staphylinidae, subfam. Aleocharinae. In: Wytsman, P. (Ed.), Genera Insectorum, Fasc. 173 A. L. Desmet-Verteneuil, Bruxelles, pp. 1 - 110." type="book chapter" year="1918">Fenyes, 1918</bibRefCitation>
).
</treatmentCitationGroup>
</paragraph>
<paragraph id="0D7836B62F022220567FB814F1C3F8CF" blockId="1.[264,1323,1433,1905]" box="[264,985,1881,1905]" pageId="1" pageNumber="2">
<taxonomicName id="CAC74D352F022220567FB814F3FBF8CF" ID-CoL="8VRBL" authority="(Halobrecta)" baseAuthorityName="Halobrecta" box="[264,481,1881,1905]" class="Insecta" family="Staphylinidae" genus="Atheta" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="2" phylum="Arthropoda" rank="genus">
<emphasis id="3FB3EAA42F022220567FB814F357F8CE" box="[264,333,1882,1905]" italics="true" pageId="1" pageNumber="2">Atheta</emphasis>
(
<emphasis id="3FB3EAA42F022220562AB814F3CFF8CE" box="[349,469,1882,1905]" italics="true" pageId="1" pageNumber="2">Halobrecta</emphasis>
)
</taxonomicName>
: Scheerpeltz, 1934: 1600 (as valid subgenus).
</paragraph>
<paragraph id="0D7836B62F012223567FBE52F14BFE8C" blockId="2.[264,1323,284,659]" box="[264,849,284,307]" pageId="2" pageNumber="3">
<treatmentCitationGroup id="2DD711982F012223567FBE52F14BFE8C" box="[264,849,284,307]" pageId="2" pageNumber="3">
<taxonomicName id="CAC74D352F012223567FBE52F080FE8C" ID-CoL="8VRQB" authority="Blackwelder, 1952: 179" authorityName="Blackwelder" authorityPageNumber="179" authorityYear="1952" box="[264,666,284,307]" class="Insecta" family="Staphylinidae" genus="Halobrecta" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="genus">
<emphasis id="3FB3EAA42F012223567FBE52F39AFE8C" box="[264,384,284,307]" italics="true" pageId="2" pageNumber="3">Halobrecta</emphasis>
: 
<treatmentCitation id="8C6610A72F01222356E2BE52F080FE8C" author="Blackwelder" box="[405,666,284,307]" page="179" pageId="2" pageNumber="3" year="1952">
<bibRefCitation id="69564B472F01222356E2BE52F078FE8C" author="Blackwelder" box="[405,610,284,307]" pageId="2" pageNumber="23" refString="Blackwelder, R. E. (1952) The generic names of the beetle family Staphylinidae with an essay on genotypy. United States National Museum Bulletin, 200, i - iv + 483 pp." type="book" year="1952">Blackwelder, 1952</bibRefCitation>
: 179
</treatmentCitation>
</taxonomicName>
(as valid genus).
</treatmentCitationGroup>
</paragraph>
<paragraph id="0D7836B62F012223567FBE72F1A2FEEC" blockId="2.[264,1323,284,659]" box="[264,952,316,339]" pageId="2" pageNumber="3">
<treatmentCitationGroup id="2DD711982F012223567FBE72F1A2FEEC" box="[264,952,316,339]" pageId="2" pageNumber="3">
<taxonomicName id="CAC74D352F012223567FBE72F064FEEC" authority="Blackwelder, 1952: 170" authorityName="Blackwelder" authorityPageNumber="170" authorityYear="1952" box="[264,638,316,339]" class="Insecta" family="Staphylinidae" genus="Glaphya" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="genus">
<emphasis id="3FB3EAA42F012223567FBE72F37EFEEC" box="[264,356,316,339]" italics="true" pageId="2" pageNumber="3">Glaphya</emphasis>
: 
<treatmentCitation id="8C6610A72F012223560FBE72F064FEEC" author="Blackwelder" box="[376,638,316,339]" page="170" pageId="2" pageNumber="3" year="1952">
<bibRefCitation id="69564B472F012223560FBE72F05FFEEC" author="Blackwelder" box="[376,581,316,339]" pageId="2" pageNumber="23" refString="Blackwelder, R. E. (1952) The generic names of the beetle family Staphylinidae with an essay on genotypy. United States National Museum Bulletin, 200, i - iv + 483 pp." type="book" year="1952">Blackwelder, 1952</bibRefCitation>
: 170
</treatmentCitation>
</taxonomicName>
(as synonym of 
<taxonomicName id="CAC74D352F0122235458BE72F1BDFEEC" ID-CoL="8VRQB" box="[815,935,316,339]" class="Insecta" family="Staphylinidae" genus="Halobrecta" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="genus">
<emphasis id="3FB3EAA42F0122235458BE72F1BDFEEC" box="[815,935,316,339]" italics="true" pageId="2" pageNumber="3">Halobrecta</emphasis>
</taxonomicName>
).
</treatmentCitationGroup>
</paragraph>
<paragraph id="0D7836B62F012223567FBE12F12DFECC" blockId="2.[264,1323,284,659]" box="[264,823,348,371]" pageId="2" pageNumber="3">
<treatmentCitationGroup id="2DD711982F012223567FBE12F12DFECC" box="[264,823,348,371]" pageId="2" pageNumber="3">
<taxonomicName id="CAC74D352F012223567FBE12F09BFECC" ID-CoL="8W25D" authority="Blackwelder, 1952: 163" authorityName="Blackwelder" authorityPageNumber="163" authorityYear="1952" box="[264,641,348,371]" class="Insecta" family="Staphylinidae" genus="Exatheta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="genus">
<emphasis id="3FB3EAA42F012223567FBE12F37DFECC" box="[264,359,348,371]" italics="true" pageId="2" pageNumber="3">Exatheta</emphasis>
: 
<treatmentCitation id="8C6610A72F012223560CBE12F09BFECC" author="Blackwelder" box="[379,641,348,371]" page="163" pageId="2" pageNumber="3" year="1952">
<bibRefCitation id="69564B472F012223560CBE12F052FECC" author="Blackwelder" box="[379,584,348,371]" pageId="2" pageNumber="23" refString="Blackwelder, R. E. (1952) The generic names of the beetle family Staphylinidae with an essay on genotypy. United States National Museum Bulletin, 200, i - iv + 483 pp." type="book" year="1952">Blackwelder, 1952</bibRefCitation>
: 163
</treatmentCitation>
</taxonomicName>
(as valid genus).
</treatmentCitationGroup>
</paragraph>
<paragraph id="0D7836B62F012223567FBE32F6D5FE2C" blockId="2.[264,1323,284,659]" box="[264,1231,380,403]" pageId="2" pageNumber="3">
<treatmentCitationGroup id="2DD711982F012223567FBE32F6D5FE2C" box="[264,1231,380,403]" pageId="2" pageNumber="3">
<taxonomicName id="CAC74D352F012223567FBE32F0DEFE2C" ID-CoL="8VRQB" authority="Benick &amp; Lohse, 1974: 219" authorityName="Benick &amp; Lohse" authorityPageNumber="219" authorityYear="1974" box="[264,708,380,403]" class="Insecta" family="Staphylinidae" genus="Halobrecta" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="genus">
<emphasis id="3FB3EAA42F012223567FBE32F39AFE2C" box="[264,384,380,403]" italics="true" pageId="2" pageNumber="3">Halobrecta</emphasis>
: 
<treatmentCitation id="8C6610A72F01222356E2BE32F0DEFE2C" author="Benick" box="[405,708,380,403]" page="219" pageId="2" pageNumber="3" year="1974">
<bibRefCitation id="69564B472F01222356E2BE32F090FE2C" author="Benick" box="[405,650,380,403]" pageId="2" pageNumber="23" refString="Benick, G. &amp; Lohse, G. A. (1974) 14. Tribus: Callicerini (Athetae). In: Freude, H., Harde, K. W. &amp; Lohse, G. A. (Eds.), Die Kafer Mitteleuropas. Band 5, Staphylinidae II (Hypocyphtinae und Aleocharinae). Pselaphidae. Goecke &amp; Evers Verlag, Krefeld, pp. 72 - 220." type="book chapter" year="1974">Benick &amp; Lohse, 1974</bibRefCitation>
: 219
</treatmentCitation>
</taxonomicName>
(as valid genus in tribe 
<taxonomicName id="CAC74D352F01222354B4BE32F6D8FE2C" ID-CoL="8VSTP" authority="Lohse, 1969" authorityName="Lohse" authorityYear="1969" box="[963,1218,380,403]" class="Insecta" family="Staphylinidae" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="tribe" tribe="Callicerini">
Callicerini 
<bibRefCitation id="69564B472F012223534CBE32F6D8FE2C" author="Lohse" box="[1083,1218,380,403]" pageId="2" pageNumber="24" refString="Lohse, G. A. (1969) Vorschlage zur Anderung der Aleocharinensystematik (Coleoptera: Staphylinidae). In: Bericht uber die 10. Wanderversammlung deutscher Entomologen, 15. Bis 19. September 1965 in Dresden. Deutsche Akademie der Landwirtschaftswissenschaften zu Berlin, Berlin, pp. 169 - 175." type="journal article" year="1969">Lohse, 1969</bibRefCitation>
</taxonomicName>
).
</treatmentCitationGroup>
</paragraph>
<paragraph id="0D7836B62F012223567FBED2F165FE0C" blockId="2.[264,1323,284,659]" box="[264,895,412,435]" pageId="2" pageNumber="3">
<treatmentCitationGroup id="2DD711982F012223567FBED2F165FE0C" box="[264,895,412,435]" pageId="2" pageNumber="3">
<taxonomicName id="CAC74D352F012223567FBED2F0D2FE0C" ID-CoL="8VRQB" authority="Moore &amp; Legner, 1975: 434" authorityName="Moore &amp; Legner" authorityPageNumber="434" authorityYear="1975" box="[264,712,412,435]" class="Insecta" family="Staphylinidae" genus="Halobrecta" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="genus">
<emphasis id="3FB3EAA42F012223567FBED2F39AFE0C" box="[264,384,412,435]" italics="true" pageId="2" pageNumber="3">Halobrecta</emphasis>
: 
<treatmentCitation id="8C6610A72F01222356E2BED2F0D2FE0C" author="Moore" box="[405,712,412,435]" page="434" pageId="2" pageNumber="3" year="1975">
<bibRefCitation id="69564B472F01222356E2BED2F095FE0C" author="Moore" box="[405,655,412,435]" pageId="2" pageNumber="24" refString="Moore, I. &amp; Legner, E. F. (1975) A catalogue of the Staphylinidae of America North of Mexico (Coleoptera). Special publication 3015, Division of Agricultural Sciences, University of California, 514 pp." type="book" year="1975">Moore &amp; Legner, 1975</bibRefCitation>
: 434
</treatmentCitation>
</taxonomicName>
(as valid genus).
</treatmentCitationGroup>
</paragraph>
<paragraph id="0D7836B62F012223567FBEF2F383FE4C" blockId="2.[264,1323,284,659]" pageId="2" pageNumber="3">
<treatmentCitationGroup id="2DD711982F012223567FBEF2F383FE4C" pageId="2" pageNumber="3">
<taxonomicName id="CAC74D352F012223567FBEF2F09FFE6C" ID-CoL="8VRQB" authority="Seevers, 1978: 121" authorityName="Seevers" authorityPageNumber="121" authorityYear="1978" box="[264,645,444,467]" class="Insecta" family="Staphylinidae" genus="Halobrecta" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="genus">
<emphasis id="3FB3EAA42F012223567FBEF2F39AFE6C" box="[264,384,444,467]" italics="true" pageId="2" pageNumber="3">Halobrecta</emphasis>
: 
<treatmentCitation id="8C6610A72F01222356D6BEF2F09FFE6C" author="Seevers" box="[417,645,444,467]" page="121" pageId="2" pageNumber="3" year="1978">
<bibRefCitation id="69564B472F01222356D6BEF2F05AFE6C" author="Seevers" box="[417,576,444,467]" pageId="2" pageNumber="25" refString="Seevers, C. H. (1978) A generic and tribal revision of the North American Aleocharinae (Coleoptera: Staphylinidae). Fieldiana: Zoology, 71, vi + 275 pp." type="book" year="1978">Seevers, 1978</bibRefCitation>
: 121
</treatmentCitation>
</taxonomicName>
(as valid genus in subtribe Xenotae 
<bibRefCitation id="69564B472F0122235344BEF2F6D4FE6C" author="Seevers" box="[1075,1230,444,467]" pageId="2" pageNumber="25" refString="Seevers, C. H. (1978) A generic and tribal revision of the North American Aleocharinae (Coleoptera: Staphylinidae). Fieldiana: Zoology, 71, vi + 275 pp." type="book" year="1978">Seevers, 1978</bibRefCitation>
(
<emphasis id="3FB3EAA42F0122235393BEF2F39AFE4C" italics="true" pageId="2" pageNumber="3">nomen nudum</emphasis>
)).
</treatmentCitationGroup>
</paragraph>
<paragraph id="0D7836B62F012223567FBEB2F611FDAC" blockId="2.[264,1323,284,659]" box="[264,1035,508,531]" pageId="2" pageNumber="3">
<treatmentCitationGroup id="2DD711982F012223567FBEB2F611FDAC" box="[264,1035,508,531]" pageId="2" pageNumber="3">
<taxonomicName id="CAC74D352F012223567FBEB2F072FDAC" ID-CoL="8VRQB" authority="Sawada, 1985: 108" authorityName="Sawada" authorityPageNumber="108" authorityYear="1985" box="[264,616,508,531]" class="Insecta" family="Staphylinidae" genus="Halobrecta" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="genus">
<emphasis id="3FB3EAA42F012223567FBEB2F39AFDAC" box="[264,384,508,531]" italics="true" pageId="2" pageNumber="3">Halobrecta</emphasis>
: 
<treatmentCitation id="8C6610A72F01222356E2BEB2F072FDAC" author="Sawada" box="[405,616,508,531]" page="108" pageId="2" pageNumber="3" year="1985">
<bibRefCitation id="69564B472F01222356E2BEB2F035FDAC" author="Sawada" box="[405,559,508,531]" pageId="2" pageNumber="24" refString="Sawada, K. (1985) Atheta and its allies of Southeast Asia (Coleoptera: Staphylinidae). IV. Ten oriental species described by M. Bernhauer and M. Cameron. Contributions from the Biological Laboratory, Kyoto University, 27 (1), 91 - 110." type="journal article" year="1985">Sawada, 1985</bibRefCitation>
: 108
</treatmentCitation>
</taxonomicName>
(as valid genus in 
<taxonomicName id="CAC74D352F0122235447BEB2F1ADFDAC" box="[816,951,508,531]" class="Insecta" family="Staphylinidae" genus="Coprothassa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="genus">
<emphasis id="3FB3EAA42F0122235447BEB2F1ADFDAC" box="[816,951,508,531]" italics="true" pageId="2" pageNumber="3">Coprothassa</emphasis>
</taxonomicName>
series).
</treatmentCitationGroup>
</paragraph>
<paragraph id="0D7836B62F012223567FBD52F192FD8C" blockId="2.[264,1323,284,659]" box="[264,904,540,563]" pageId="2" pageNumber="3">
<treatmentCitationGroup id="2DD711982F012223567FBD52F192FD8C" box="[264,904,540,563]" pageId="2" pageNumber="3">
<taxonomicName id="CAC74D352F012223567FBD52F054FD8C" ID-CoL="8W25D" authority="Sawada, 1985: 108" authorityName="Sawada" authorityPageNumber="108" authorityYear="1985" box="[264,590,540,563]" class="Insecta" family="Staphylinidae" genus="Exatheta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="genus">
<emphasis id="3FB3EAA42F012223567FBD52F37DFD8C" box="[264,359,540,563]" italics="true" pageId="2" pageNumber="3">Exatheta</emphasis>
: 
<treatmentCitation id="8C6610A72F012223560CBD52F054FD8C" author="Sawada" box="[379,590,540,563]" page="108" pageId="2" pageNumber="3" year="1985">
<bibRefCitation id="69564B472F012223560CBD52F00FFD8C" author="Sawada" box="[379,533,540,563]" pageId="2" pageNumber="24" refString="Sawada, K. (1985) Atheta and its allies of Southeast Asia (Coleoptera: Staphylinidae). IV. Ten oriental species described by M. Bernhauer and M. Cameron. Contributions from the Biological Laboratory, Kyoto University, 27 (1), 91 - 110." type="journal article" year="1985">Sawada, 1985</bibRefCitation>
: 108
</treatmentCitation>
</taxonomicName>
(as synonym of 
<taxonomicName id="CAC74D352F0122235477BD52F162FD8C" ID-CoL="8VRQB" box="[768,888,540,563]" class="Insecta" family="Staphylinidae" genus="Halobrecta" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="genus">
<emphasis id="3FB3EAA42F0122235477BD52F162FD8C" box="[768,888,540,563]" italics="true" pageId="2" pageNumber="3">Halobrecta</emphasis>
</taxonomicName>
).
</treatmentCitationGroup>
</paragraph>
<paragraph id="0D7836B62F012223567FBD72F1A4FDEC" blockId="2.[264,1323,284,659]" box="[264,958,572,595]" pageId="2" pageNumber="3">
<treatmentCitationGroup id="2DD711982F012223567FBD72F1A4FDEC" box="[264,958,572,595]" pageId="2" pageNumber="3">
<taxonomicName id="CAC74D352F012223567FBD72F04CFDEC" ID-CoL="8VRQB" authority="Lohse, 1989: 219" authorityName="Lohse" authorityPageNumber="219" authorityYear="1989" box="[264,598,572,595]" class="Insecta" family="Staphylinidae" genus="Halobrecta" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="genus">
<emphasis id="3FB3EAA42F012223567FBD72F39AFDEC" box="[264,384,572,595]" italics="true" pageId="2" pageNumber="3">Halobrecta</emphasis>
: 
<treatmentCitation id="8C6610A72F01222356E2BD72F04CFDEC" author="Lohse" box="[405,598,572,595]" page="219" pageId="2" pageNumber="3" year="1989">
<bibRefCitation id="69564B472F01222356E2BD72F007FDEC" author="Lohse" box="[405,541,572,595]" pageId="2" pageNumber="24" refString="Lohse, G. A. (1989) Erganzungen und Berichtigungen zu Freude - Harde - Lohse &quot; Die Kafer Mitteleuropas &quot; Band 5 (1974) In: Lohse, G. A. &amp; Lucht, W. H. (Eds.), Die Kafer Mitteleuropas. 1. Supplementband mit Katalogteil. Goecke &amp; Evers Verlag, Krefeld, pp. 185 - 243." type="journal article" year="1989">Lohse, 1989</bibRefCitation>
: 219
</treatmentCitation>
</taxonomicName>
(as valid genus in tribe 
<taxonomicName id="CAC74D352F0122235421BD72F1A8FDEC" ID-CoL="8VR6Z" box="[854,946,572,595]" class="Insecta" family="Staphylinidae" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="tribe" tribe="Athetini">Athetini</taxonomicName>
).
</treatmentCitationGroup>
</paragraph>
<paragraph id="0D7836B62F012223567FBD12F036FD2C" blockId="2.[264,1323,284,659]" pageId="2" pageNumber="3">
<treatmentCitationGroup id="2DD711982F012223567FBD12F036FD2C" pageId="2" pageNumber="3">
<taxonomicName id="CAC74D352F012223567FBD12F3C8FDCC" ID-CoL="8VRQB" authority="Ashe" authorityName="Ashe" box="[264,466,604,627]" class="Insecta" family="Staphylinidae" genus="Halobrecta" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="genus">
<emphasis id="3FB3EAA42F012223567FBD12F39AFDCC" box="[264,384,604,627]" italics="true" pageId="2" pageNumber="3">Halobrecta</emphasis>
: Ashe
</taxonomicName>
<emphasis id="3FB3EAA42F01222356ACBD12F3EAFDCC" box="[475,496,604,627]" italics="true" pageId="2" pageNumber="3">in</emphasis>
<treatmentCitation id="8C6610A72F012223568EBD12F1E5FDCC" author="Newton" box="[505,1023,604,627]" page="369" pageId="2" pageNumber="3" year="2000">
<bibRefCitation id="69564B472F012223568EBD12F1DBFDCC" author="Newton" box="[505,961,604,627]" pageId="2" pageNumber="24" refString="Newton, A. F., Thayer, M. K., Ashe, J. S. &amp; Chandler, D. S. (2000) Staphylinidae Latreille, 1802. In: Arnett, R. H., Thomas, M. C. (Eds.), American Beetles. Vol. 1. Archostemata, Myxophaga, Adephaga, Polyphaga: Staphyliniformia. CRC Press, Boca Raton, Florida, pp. 272 - 418." type="book chapter" year="2000">Newton, Thayer, Ashe &amp; Chandler, 2000</bibRefCitation>
: 369
</treatmentCitation>
(as valid genus in subtribe 
<taxonomicName id="CAC74D352F012223564FBD32F03AFD2C" ID-CoL="8VRC4" authority="Casey, 1910" authorityName="Casey" authorityYear="1910" box="[312,544,636,659]" class="Insecta" family="Staphylinidae" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="subTribe" subTribe="Athetina">
Athetina 
<bibRefCitation id="69564B472F01222356EDBD32F03AFD2C" author="Casey" box="[410,544,636,659]" pageId="2" pageNumber="23" refString="Casey, T. L. (1910) New Species of the Staphylinid Tribe Myrmedoniini. Memoirs on the Coleoptera I. The New Era Printing Company, Lancaster, pp. 1 - 183." type="book chapter" year="1910">Casey, 1910</bibRefCitation>
</taxonomicName>
).
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="45DD653D2F012223567FBD8AF032FAE1" pageId="2" pageNumber="3" type="diagnosis">
<paragraph id="0D7836B62F012223567FBD8AF032FAE1" blockId="2.[264,1323,708,1894]" pageId="2" pageNumber="3">
<emphasis id="3FB3EAA42F012223567FBD8AF392FD61" bold="true" box="[264,392,708,734]" pageId="2" pageNumber="3">Diagnosis.</emphasis>
<taxonomicName id="CAC74D352F01222356EFBD8AF007FD62" box="[408,541,708,733]" class="Insecta" family="Staphylinidae" genus="Halobrecta" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="genus">
<emphasis id="3FB3EAA42F01222356EFBD8AF007FD62" box="[408,541,708,733]" italics="true" pageId="2" pageNumber="3">Halobrecta</emphasis>
</taxonomicName>
is distinguished from other athetine genera by the combination of the following characters: body parallel­sided; anterior margin of labrum straight; sensilla 
<emphasis id="3FB3EAA42F012223567FBC5AF30DFC92" box="[264,279,788,813]" italics="true" pageId="2" pageNumber="3">a</emphasis>
of epipharynx long; antennal article 2 longer than article 3, articles 7–10 transverse; ligula long, parallel­sided, with narrow base and slightly split apically (
<figureCitation id="95FC2A332F0122235324BC72F686FCE9" box="[1107,1180,828,854]" captionStart="FIGURES 6 – 12" captionStartId="5.[264,383,1714,1738]" captionTargetBox="[312,1278,649,1705]" captionTargetId="figure@5.[299,1288,635,1710]" captionTargetPageId="5" captionText="FIGURES 6 – 12. Mouthparts of Halobrecta sp. aff. algae (Hardy) (female, Grado, Italy (6 – 11 )) and Ha. sp. cf. halensis Mulsant &amp; Rey, 1873 (female, Lignano, Italy (12 )). 6 — prementum; 7 — hypopharynx; 8 — right maxilla, ventral view; 9 — right galea, ventral view; 10 — right lacinia, ventral view; 11 — right lacinia, dorsal view; 12 — mentum. Scale bar 0.1 mm (6 – 7, 9 – 12), 0.2 mm (8)." httpUri="https://zenodo.org/record/158215/files/figure.png" pageId="2" pageNumber="3">Fig. 6</figureCitation>
); labial palpus with setae and present (
<figureCitation id="95FC2A332F0122235503BC2AF0A6FCC1" box="[628,700,868,894]" captionStart="FIGURES 6 – 12" captionStartId="5.[264,383,1714,1738]" captionTargetBox="[312,1278,649,1705]" captionTargetId="figure@5.[299,1288,635,1710]" captionTargetPageId="5" captionText="FIGURES 6 – 12. Mouthparts of Halobrecta sp. aff. algae (Hardy) (female, Grado, Italy (6 – 11 )) and Ha. sp. cf. halensis Mulsant &amp; Rey, 1873 (female, Lignano, Italy (12 )). 6 — prementum; 7 — hypopharynx; 8 — right maxilla, ventral view; 9 — right galea, ventral view; 10 — right lacinia, ventral view; 11 — right lacinia, dorsal view; 12 — mentum. Scale bar 0.1 mm (6 – 7, 9 – 12), 0.2 mm (8)." httpUri="https://zenodo.org/record/158215/files/figure.png" pageId="2" pageNumber="3">Fig. 6</figureCitation>
); pronotum slightly transverse, 1.1–1.2 times as wide as long, with microsetae directed anteriorly along anterior half of midline, and posteriorly along posterior half of midline; in lateral portions of the disc microsetae directed laterally (
<typeStatus id="D27C88142F0122235667BC92F356FC49" box="[272,332,988,1014]" pageId="2" pageNumber="3">Type</typeStatus>
III, 
<bibRefCitation id="69564B472F01222356F6BC92F092FC49" author="Benick" box="[385,648,988,1014]" pageId="2" pageNumber="23" refString="Benick, G. &amp; Lohse, G. A. (1974) 14. Tribus: Callicerini (Athetae). In: Freude, H., Harde, K. W. &amp; Lohse, G. A. (Eds.), Die Kafer Mitteleuropas. Band 5, Staphylinidae II (Hypocyphtinae und Aleocharinae). Pselaphidae. Goecke &amp; Evers Verlag, Krefeld, pp. 72 - 220." type="book chapter" year="1974">Benick &amp; Lohse 1974</bibRefCitation>
) (
<figureCitation id="95FC2A332F01222355D6BC92F0E3FC49" box="[673,761,988,1014]" captionStart="FIGURES 13 – 17" captionStartId="6.[264,382,1493,1517]" captionTargetBox="[453,1127,301,1448]" captionTargetId="figure@6.[441,1145,290,1457]" captionTargetPageId="6" captionText="FIGURES 13 – 17. Details of Halobrecta sp. aff. algae (Hardy) (female, Grado, Italy). 13 — right antenna; 14 — pronotum; 15 — meso­ and metathorax; 16 — left metatarsal segment 5 and metapretarsus, anterior view; 17 — right metatarsus, posterior view. Scale bar 0.1 mm (16), 0.2 mm (13, 17), 0.4 mm (14 – 15)." httpUri="https://zenodo.org/record/158216/files/figure.png" pageId="2" pageNumber="3">Fig. 14</figureCitation>
); pronotal macrosetae long; pronotal hypomera fully visible in lateral view; medial macroseta of mesotibia thin, as long as tibial width; mesothoracic process narrow (
<figureCitation id="95FC2A332F0122235502BB62F0D7FBF9" box="[629,717,1068,1094]" captionStart="FIGURES 13 – 17" captionStartId="6.[264,382,1493,1517]" captionTargetBox="[453,1127,301,1448]" captionTargetId="figure@6.[441,1145,290,1457]" captionTargetPageId="6" captionText="FIGURES 13 – 17. Details of Halobrecta sp. aff. algae (Hardy) (female, Grado, Italy). 13 — right antenna; 14 — pronotum; 15 — meso­ and metathorax; 16 — left metatarsal segment 5 and metapretarsus, anterior view; 17 — right metatarsus, posterior view. Scale bar 0.1 mm (16), 0.2 mm (13, 17), 0.4 mm (14 – 15)." httpUri="https://zenodo.org/record/158216/files/figure.png" pageId="2" pageNumber="3">Fig. 15</figureCitation>
); posterior margin of elytra emarginate; tarsal formula 4­5­5; metatarsal segment 1 longer than segment 2; single empodial seta longer than claws; abdominal terga 3–5 with transverse basal impression; posterior margin of female tergum 8 with comb of scattered tiny projections (
<figureCitation id="95FC2A332F012223542BBBEAF1A5FB01" box="[860,959,1188,1214]" captionStart="FIGURES 18 – 21" captionStartId="9.[264,382,1204,1228]" captionTargetBox="[292,1295,296,1174]" captionTargetId="figure@9.[283,1304,284,1181]" captionTargetPageId="9" captionText="FIGURES 18 – 21. Abdominal segment 8 of Halobrecta algophila (Fenyes) (male lectotype (18 – 19) and female paralectotype of At. algophila (20 – 21 )). 18 — male tergum 8; 19 — male sternum 8; 20 — female tergum 8; 21 — female sternum 8. Scale bar 0.2 mm." httpUri="https://zenodo.org/record/158217/files/figure.png" pageId="2" pageNumber="3">Figs. 20</figureCitation>
, 
<figureCitation id="95FC2A332F01222354BABBEAF1F4FB01" box="[973,1006,1188,1214]" captionStart="FIGURES 37 – 40" captionStartId="13.[264,382,1414,1438]" captionTargetBox="[292,1310,552,1389]" captionTargetId="figure@13.[280,1321,538,1399]" captionTargetPageId="13" captionText="FIGURES 37 – 40. Abdominal segment 8 of Halobrecta flavipes Thomson (male (37 – 38) and female paralectotypes Atheta pocahontas Casey (39 – 40 )). 37 — male tergum 8; 38 — male sternum 8; 39 — female tergum 8; 40 — female sternum 8. Scale bar 0.2 mm." httpUri="https://zenodo.org/record/158220/files/figure.png" pageId="2" pageNumber="3">39</figureCitation>
); median lobe of aedeagus without athetine bridge, apex of paramere long and narrow, medial lamellae of internal sac absent; copulatory piece with pointed apex (
<figureCitation id="95FC2A332F0122235464BBBAF16DFAB1" box="[787,887,1268,1294]" captionStart="FIGURES 30 – 36" captionStartId="11.[264,382,1692,1716]" captionTargetBox="[304,1285,459,1679]" captionTargetId="figure@11.[295,1292,446,1690]" captionTargetPageId="11" captionText="FIGURES 30 – 36. Genitalia of Halobrecta algophila (Fenyes) (male lectotype (30 – 31, 33 – 36) and female paralectotype of At. algophila (32 )). 30 — details of retracted internal sac, abparameral view; 31 — details of retracted internal sac, lateral view; 32 — spermatheca; 33 — left paramere, view from the side facing the median lobe; 34 — left paramere, marginal view; 35 — left paramere, view from the side opposite to the median lobe; 36 — apex of left paramere, marginal view. Scale bar 0.1 mm (32, 36), 0.2 mm (30 – 31, 33 – 35)." httpUri="https://zenodo.org/record/158219/files/figure.png" pageId="2" pageNumber="3">Figs. 30</figureCitation>
, 
<figureCitation id="95FC2A332F01222354F3BBBAF1BEFAB1" box="[900,932,1268,1294]" captionStart="FIGURES 41 – 44" captionStartId="14.[264,382,1781,1805]" captionTargetBox="[346,1257,1081,1771]" captionTargetId="figure@14.[339,1264,1071,1778]" captionTargetPageId="14" captionText="FIGURES 41 – 44. Aedeagus of Halobrecta flavipes Thomson (male from Oskarshamn, Sweden (41 – 42), and male paralectotype of Atheta pocahontas Casey (43 – 44 )). 41, 43 — median lobe, parameral view; 42, 44 — apex of median lobe, parameral view. Scale bar 0.1 mm (42, 44), 0.2 mm (41, 43)." httpUri="https://zenodo.org/record/158221/files/figure.png" pageId="2" pageNumber="3">41</figureCitation>
, 
<figureCitation id="95FC2A332F01222354C6BBBAF1CAFAB1" box="[945,976,1268,1294]" captionStart="FIGURES 62 – 65" captionStartId="20.[264,383,1746,1770]" captionTargetBox="[328,1216,1040,1731]" captionTargetId="figure@20.[320,1227,1026,1736]" captionTargetPageId="20" captionText="FIGURES 62 – 65. Details of Halobrecta sp. aff. algae (Hardy) (male, Grado, Italy (62 – 63 )) and Ha. sp. cf. halensis Mulsant &amp; Rey, 1873 (male, Lignano, Italy (64 – 65 )). 62 — median lobe with everted internal sac, parameral view; 63 — median lobe with everted internal sac, lateral view; 64 — median lobe with partially everted internal sac, parameral view; 65 — median lobe with partially everted internal sac, lateral view. Scale bar 0.2 mm." httpUri="https://zenodo.org/record/158226/files/figure.png" pageId="2" pageNumber="3">62</figureCitation>
) and without sclerotized suspensoria; spermatheca short, not divided into distal and proximal portions, with large umbilicus (
<figureCitation id="95FC2A332F01222356FBBA0AF3F4FAE1" box="[396,494,1348,1374]" captionStart="FIGURES 30 – 36" captionStartId="11.[264,382,1692,1716]" captionTargetBox="[304,1285,459,1679]" captionTargetId="figure@11.[295,1292,446,1690]" captionTargetPageId="11" captionText="FIGURES 30 – 36. Genitalia of Halobrecta algophila (Fenyes) (male lectotype (30 – 31, 33 – 36) and female paralectotype of At. algophila (32 )). 30 — details of retracted internal sac, abparameral view; 31 — details of retracted internal sac, lateral view; 32 — spermatheca; 33 — left paramere, view from the side facing the median lobe; 34 — left paramere, marginal view; 35 — left paramere, view from the side opposite to the median lobe; 36 — apex of left paramere, marginal view. Scale bar 0.1 mm (32, 36), 0.2 mm (30 – 31, 33 – 35)." httpUri="https://zenodo.org/record/158219/files/figure.png" pageId="2" pageNumber="3">Figs. 32</figureCitation>
, 
<figureCitation id="95FC2A332F012223568DBA0AF000FAE1" box="[506,538,1348,1374]" captionStart="FIGURES 49 – 51" captionStartId="16.[264,382,1105,1129]" captionTargetBox="[346,1246,419,1083]" captionTargetId="figure@16.[331,1257,409,1090]" captionTargetPageId="16" captionText="FIGURES 49 – 51. Genitalia of Halobrecta flavipes Thomson (male (49 – 50) and female paralectotypes of Atheta pocahontas (51 )). 49 — details of retracted internal sac, lateral view; 50 — apex of left paramere, marginal view; 51 — spermatheca. Scale bar 0.1 mm (50 – 51), 0.2 mm (49)." httpUri="https://zenodo.org/record/158223/files/figure.png" pageId="2" pageNumber="3">51</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="45DD653D2F012223564FBA22F073F9C9" pageId="2" pageNumber="3" type="reference_group">
<paragraph id="0D7836B62F012223564FBA22F073F9C9" blockId="2.[264,1323,708,1894]" pageId="2" pageNumber="3">
<taxonomicName id="CAC74D352F012223564FBA22F3A7FA3A" ID-CoL="8VRQB" box="[312,445,1388,1413]" class="Insecta" family="Staphylinidae" genus="Halobrecta" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="genus">
<emphasis id="3FB3EAA42F012223564FBA22F3A7FA3A" box="[312,445,1388,1413]" italics="true" pageId="2" pageNumber="3">Halobrecta</emphasis>
</taxonomicName>
differs from other littoral aleocharines with the same tarsal formula (
<taxonomicName id="CAC74D352F0122235393BA22F387FA11" ID-CoL="8VRFR" authority="Casey, 1910" authorityName="Casey" authorityYear="1910" class="Insecta" family="Staphylinidae" genus="Adota" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="genus">
<emphasis id="3FB3EAA42F0122235393BA22F730FA3A" box="[1252,1322,1388,1413]" italics="true" pageId="2" pageNumber="3">Adota</emphasis>
<bibRefCitation id="69564B472F012223567FBADAF387FA11" author="Casey" box="[264,413,1428,1454]" pageId="2" pageNumber="23" refString="Casey, T. L. (1910) New Species of the Staphylinid Tribe Myrmedoniini. Memoirs on the Coleoptera I. The New Era Printing Company, Lancaster, pp. 1 - 183." type="book chapter" year="1910">Casey, 1910</bibRefCitation>
</taxonomicName>
, 
<taxonomicName id="CAC74D352F01222356DCBADAF17FFA11" ID-CoL="8VRZX" authority="Yosii &amp; Sawada, 1976" authorityName="Yosii &amp; Sawada" authorityYear="1976" box="[427,869,1428,1454]" class="Insecta" family="Staphylinidae" genus="Psammostiba" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="genus">
<emphasis id="3FB3EAA42F01222356DCBADAF053FA12" box="[427,585,1428,1453]" italics="true" pageId="2" pageNumber="3">Psammostiba</emphasis>
<bibRefCitation id="69564B472F0122235524BADAF17FFA11" author="Yosii" box="[595,869,1428,1454]" pageId="2" pageNumber="25" refString="Yosii, R. &amp; Sawada, K. (1976) Studies on the genus Atheta Thomson and its allies (Coleoptera, Staphylinidae). II: Diagnostic characters of genera and subgenera with description of representative species. Contributions from the Biological Laboratory, Kyoto University, 25 (1), 11 - 140." type="journal article" year="1976">Yosii &amp; Sawada, 1976</bibRefCitation>
</taxonomicName>
, 
<taxonomicName id="CAC74D352F0122235404BADAF6B3FA11" ID-CoL="8VRZP" authority="Casey, 1885" authorityName="Casey" authorityYear="1885" box="[883,1193,1428,1454]" class="Insecta" family="Staphylinidae" genus="Pontomalota" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="genus">
<emphasis id="3FB3EAA42F0122235404BADAF611FA12" box="[883,1035,1428,1453]" italics="true" pageId="2" pageNumber="3">Pontomalota</emphasis>
<bibRefCitation id="69564B472F0122235362BADAF6B3FA11" author="Casey" box="[1045,1193,1428,1454]" pageId="2" pageNumber="23" refString="Casey, T. L. (1885) New genera and species of Californian Coleoptera. Bulletin of the California Academy of Sciences, 1 (4), 283 - 336." type="journal article" year="1885">Casey, 1885</bibRefCitation>
</taxonomicName>
, 
<taxonomicName id="CAC74D352F01222353CFBADAF386FA69" ID-CoL="8VS4Z" authority="Casey, 1894" authorityName="Casey" authorityYear="1894" class="Insecta" family="Staphylinidae" genus="Tarphiota" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="genus">
<emphasis id="3FB3EAA42F01222353CFBADAF732FA12" box="[1208,1320,1428,1453]" italics="true" pageId="2" pageNumber="3">Tarphiota</emphasis>
<bibRefCitation id="69564B472F012223567FBAF2F386FA69" author="Casey" box="[264,412,1468,1494]" pageId="2" pageNumber="23" refString="Casey, T. L. (1894) Coleopterological notices. V. Annals of the New York Academy of Sciences, 7 [1893], 281 - 606." type="journal article" year="1894">Casey, 1894</bibRefCitation>
</taxonomicName>
) in having a different 
<typeStatus id="D27C88142F01222355C2BAF2F0F2FA69" box="[693,744,1468,1494]" pageId="2" pageNumber="3">type</typeStatus>
of pronotal pubescence (
<figureCitation id="95FC2A332F0122235369BAF2F663FA69" box="[1054,1145,1468,1494]" captionStart="FIGURES 13 – 17" captionStartId="6.[264,382,1493,1517]" captionTargetBox="[453,1127,301,1448]" captionTargetId="figure@6.[441,1145,290,1457]" captionTargetPageId="6" captionText="FIGURES 13 – 17. Details of Halobrecta sp. aff. algae (Hardy) (female, Grado, Italy). 13 — right antenna; 14 — pronotum; 15 — meso­ and metathorax; 16 — left metatarsal segment 5 and metapretarsus, anterior view; 17 — right metatarsus, posterior view. Scale bar 0.1 mm (16), 0.2 mm (13, 17), 0.4 mm (14 – 15)." httpUri="https://zenodo.org/record/158216/files/figure.png" pageId="2" pageNumber="3">Fig. 14</figureCitation>
); the posterior margin of the elytra emarginate; the posterior margin of the female tergum 8 with a comb of scattered tiny projections (
<figureCitation id="95FC2A332F012223551CB942F0CBF999" box="[619,721,1548,1574]" captionStart="FIGURES 18 – 21" captionStartId="9.[264,382,1204,1228]" captionTargetBox="[292,1295,296,1174]" captionTargetId="figure@9.[283,1304,284,1181]" captionTargetPageId="9" captionText="FIGURES 18 – 21. Abdominal segment 8 of Halobrecta algophila (Fenyes) (male lectotype (18 – 19) and female paralectotype of At. algophila (20 – 21 )). 18 — male tergum 8; 19 — male sternum 8; 20 — female tergum 8; 21 — female sternum 8. Scale bar 0.2 mm." httpUri="https://zenodo.org/record/158217/files/figure.png" pageId="2" pageNumber="3">Figs. 20</figureCitation>
, 
<figureCitation id="95FC2A332F0122235597B942F11BF999" box="[736,769,1548,1574]" captionStart="FIGURES 37 – 40" captionStartId="13.[264,382,1414,1438]" captionTargetBox="[292,1310,552,1389]" captionTargetId="figure@13.[280,1321,538,1399]" captionTargetPageId="13" captionText="FIGURES 37 – 40. Abdominal segment 8 of Halobrecta flavipes Thomson (male (37 – 38) and female paralectotypes Atheta pocahontas Casey (39 – 40 )). 37 — male tergum 8; 38 — male sternum 8; 39 — female tergum 8; 40 — female sternum 8. Scale bar 0.2 mm." httpUri="https://zenodo.org/record/158220/files/figure.png" pageId="2" pageNumber="3">39</figureCitation>
); the median lobe without the athetine bridge (
<figureCitation id="95FC2A332F0122235666B97AF3BAF9F1" box="[273,416,1588,1614]" captionStart="FIGURES 22 – 29" captionStartId="10.[264,382,1447,1471]" captionTargetBox="[297,1292,297,1431]" captionTargetId="figure@10.[285,1301,286,1440]" captionTargetPageId="10" captionText="FIGURES 22 – 29. Aedeagus of Halobrecta algophila (Fenyes) (male lectotype of At. algophila (22 – 25), and male from the environs of Bolinas, California (26 – 29 )). 22, 27 — apex of median lobe, parameral view; 23, 26 — apex of median lobe, parameral view; 25, 28 — median lobe, lateral view; 24, 29 — apex of median lobe, lateral view. Scale bar 0.1 mm (22, 24, 27, 29), 0.2 mm (23, 25, 26, 28)." httpUri="https://zenodo.org/record/158218/files/figure.png" pageId="2" pageNumber="3">Figs. 25, 28</figureCitation>
); the paramere with a long apex (
<figureCitation id="95FC2A332F0122235452B97AF192F9F1" box="[805,904,1588,1614]" captionStart="FIGURES 30 – 36" captionStartId="11.[264,382,1692,1716]" captionTargetBox="[304,1285,459,1679]" captionTargetId="figure@11.[295,1292,446,1690]" captionTargetPageId="11" captionText="FIGURES 30 – 36. Genitalia of Halobrecta algophila (Fenyes) (male lectotype (30 – 31, 33 – 36) and female paralectotype of At. algophila (32 )). 30 — details of retracted internal sac, abparameral view; 31 — details of retracted internal sac, lateral view; 32 — spermatheca; 33 — left paramere, view from the side facing the median lobe; 34 — left paramere, marginal view; 35 — left paramere, view from the side opposite to the median lobe; 36 — apex of left paramere, marginal view. Scale bar 0.1 mm (32, 36), 0.2 mm (30 – 31, 33 – 35)." httpUri="https://zenodo.org/record/158219/files/figure.png" pageId="2" pageNumber="3">Figs. 36</figureCitation>
, 
<figureCitation id="95FC2A332F01222354E4B97AF1A8F9F1" box="[915,946,1588,1614]" captionStart="FIGURES 49 – 51" captionStartId="16.[264,382,1105,1129]" captionTargetBox="[346,1246,419,1083]" captionTargetId="figure@16.[331,1257,409,1090]" captionTargetPageId="16" captionText="FIGURES 49 – 51. Genitalia of Halobrecta flavipes Thomson (male (49 – 50) and female paralectotypes of Atheta pocahontas (51 )). 49 — details of retracted internal sac, lateral view; 50 — apex of left paramere, marginal view; 51 — spermatheca. Scale bar 0.1 mm (50 – 51), 0.2 mm (49)." httpUri="https://zenodo.org/record/158223/files/figure.png" pageId="2" pageNumber="3">50</figureCitation>
) and a short spermatheca with a large umbilicus (
<figureCitation id="95FC2A332F01222356BAB912F035F9C9" box="[461,559,1628,1654]" captionStart="FIGURES 30 – 36" captionStartId="11.[264,382,1692,1716]" captionTargetBox="[304,1285,459,1679]" captionTargetId="figure@11.[295,1292,446,1690]" captionTargetPageId="11" captionText="FIGURES 30 – 36. Genitalia of Halobrecta algophila (Fenyes) (male lectotype (30 – 31, 33 – 36) and female paralectotype of At. algophila (32 )). 30 — details of retracted internal sac, abparameral view; 31 — details of retracted internal sac, lateral view; 32 — spermatheca; 33 — left paramere, view from the side facing the median lobe; 34 — left paramere, marginal view; 35 — left paramere, view from the side opposite to the median lobe; 36 — apex of left paramere, marginal view. Scale bar 0.1 mm (32, 36), 0.2 mm (30 – 31, 33 – 35)." httpUri="https://zenodo.org/record/158219/files/figure.png" pageId="2" pageNumber="3">Figs. 32</figureCitation>
, 
<figureCitation id="95FC2A332F012223554CB912F046F9C9" box="[571,604,1628,1654]" captionStart="FIGURES 49 – 51" captionStartId="16.[264,382,1105,1129]" captionTargetBox="[346,1246,419,1083]" captionTargetId="figure@16.[331,1257,409,1090]" captionTargetPageId="16" captionText="FIGURES 49 – 51. Genitalia of Halobrecta flavipes Thomson (male (49 – 50) and female paralectotypes of Atheta pocahontas (51 )). 49 — details of retracted internal sac, lateral view; 50 — apex of left paramere, marginal view; 51 — spermatheca. Scale bar 0.1 mm (50 – 51), 0.2 mm (49)." httpUri="https://zenodo.org/record/158223/files/figure.png" pageId="2" pageNumber="3">51</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="45DD653D2F012222564FB9CAF378F899" lastPageId="3" lastPageNumber="4" pageId="2" pageNumber="3" type="description">
<paragraph id="0D7836B62F012223564FB9CAF3D4F951" blockId="2.[264,1323,708,1894]" pageId="2" pageNumber="3">
<emphasis id="3FB3EAA42F012223564FB9CAF3D5F921" bold="true" box="[312,463,1668,1694]" pageId="2" pageNumber="3">Description.</emphasis>
Length 
<quantity id="CA3F9B532F012223553EB9CAF0C5F921" box="[585,735,1668,1694]" metricMagnitude="-3" metricUnit="m" metricValue="3.4" metricValueMax="3.9" metricValueMin="2.9" pageId="2" pageNumber="3" unit="mm" value="3.4" valueMax="3.9" valueMin="2.9">2.9–3.9 mm</quantity>
, pronotal width 
<quantity id="CA3F9B532F01222354DAB9CAF67BF921" box="[941,1121,1668,1694]" metricMagnitude="-4" metricUnit="m" metricValue="5.949999999999999" metricValueMax="6.6000000000000005" metricValueMin="5.300000000000001" pageId="2" pageNumber="3" unit="mm" value="0.595" valueMax="0.66" valueMin="0.53">0.53–0.66 mm</quantity>
. Body parallelsided, dark brown, with legs, basal or all antennal articles, and mouthparts yellowish brown to yellow.
</paragraph>
<paragraph id="0D7836B62F012222564FB9B2F610FC09" blockId="2.[264,1323,708,1894]" lastBlockId="3.[264,1325,284,1870]" lastPageId="3" lastPageNumber="4" pageId="2" pageNumber="3">
Head transverse; eye length to temple length ratio 0.7–1.2; infraorbital carina complete. Antennal article 2 longer than article 3, article 4 elongate, 5 elongate or subquadrate, 6 subquadrate or transverse, 7–10 transverse, apical article without coeloconic sensilla, as long as articles 9 and 10 combined. Labrum (
<figureCitation id="95FC2A332F00222254C1BE52F1E1FE89" box="[950,1019,284,310]" captionStart="FIGURES 1 – 5" captionStartId="4.[264,383,1512,1536]" captionTargetBox="[294,1291,289,1499]" captionTargetId="figure@4.[291,1296,283,1505]" captionTargetPageId="4" captionText="FIGURES 1 – 5. Mouthparts of Halobrecta sp. aff. algae (Hardy) (female, Grado, Italy). 1 — labrum; 2 — epipharynx; 3 — left mandible, dorsal view; 4 — left mandible, ventral view; 5 — right mandible, dorsal view. Scale bar 0.1 mm." httpUri="https://zenodo.org/record/158214/files/figure.png" pageId="3" pageNumber="4">Fig. 1</figureCitation>
) transverse, with straight anterior margin. Epipharynx (
<figureCitation id="95FC2A332F0022225505BE0AF0A0FEE1" box="[626,698,324,350]" captionStart="FIGURES 1 – 5" captionStartId="4.[264,383,1512,1536]" captionTargetBox="[294,1291,289,1499]" captionTargetId="figure@4.[291,1296,283,1505]" captionTargetPageId="4" captionText="FIGURES 1 – 5. Mouthparts of Halobrecta sp. aff. algae (Hardy) (female, Grado, Italy). 1 — labrum; 2 — epipharynx; 3 — left mandible, dorsal view; 4 — left mandible, ventral view; 5 — right mandible, dorsal view. Scale bar 0.1 mm." httpUri="https://zenodo.org/record/158214/files/figure.png" pageId="3" pageNumber="4">Fig. 2</figureCitation>
) with long sensilla 
<emphasis id="3FB3EAA42F00222254DFBE0AF1ACFEE2" box="[936,950,324,349]" italics="true" pageId="3" pageNumber="4">a</emphasis>
, with three pairs of small marginal setae, medial field with 32 pores, lateral rows with two pores each, anterolateral groups with three pores each, transverse row with six pores, posterolateral groups with three–four pores each, with one medial proximal pore on each side and two lateral proximal pores. Mandibles (
<figureCitation id="95FC2A332F002222556CBEAAF090FE41" box="[539,650,484,510]" captionStart="FIGURES 1 – 5" captionStartId="4.[264,383,1512,1536]" captionTargetBox="[294,1291,289,1499]" captionTargetId="figure@4.[291,1296,283,1505]" captionTargetPageId="4" captionText="FIGURES 1 – 5. Mouthparts of Halobrecta sp. aff. algae (Hardy) (female, Grado, Italy). 1 — labrum; 2 — epipharynx; 3 — left mandible, dorsal view; 4 — left mandible, ventral view; 5 — right mandible, dorsal view. Scale bar 0.1 mm." httpUri="https://zenodo.org/record/158214/files/figure.png" pageId="3" pageNumber="4">Figs. 3–5</figureCitation>
) broad, right mandible with a small medial tooth; velvety patch of dorsal molar area not visible at 400x. Maxilla (
<figureCitation id="95FC2A332F00222254D3BD42F63BFD99" box="[932,1057,524,550]" captionStart="FIGURES 6 – 12" captionStartId="5.[264,383,1714,1738]" captionTargetBox="[312,1278,649,1705]" captionTargetId="figure@5.[299,1288,635,1710]" captionTargetPageId="5" captionText="FIGURES 6 – 12. Mouthparts of Halobrecta sp. aff. algae (Hardy) (female, Grado, Italy (6 – 11 )) and Ha. sp. cf. halensis Mulsant &amp; Rey, 1873 (female, Lignano, Italy (12 )). 6 — prementum; 7 — hypopharynx; 8 — right maxilla, ventral view; 9 — right galea, ventral view; 10 — right lacinia, ventral view; 11 — right lacinia, dorsal view; 12 — mentum. Scale bar 0.1 mm (6 – 7, 9 – 12), 0.2 mm (8)." httpUri="https://zenodo.org/record/158215/files/figure.png" pageId="3" pageNumber="4">Figs. 8–11</figureCitation>
) with galea projecting slightly beyond apex of lacinia; apical lobe of galea covered with numerous fine and short setae; internal margin of galea with long subapical setae (
<figureCitation id="95FC2A332F00222254D8BD12F1E2FDC9" box="[943,1016,604,630]" captionStart="FIGURES 6 – 12" captionStartId="5.[264,383,1714,1738]" captionTargetBox="[312,1278,649,1705]" captionTargetId="figure@5.[299,1288,635,1710]" captionTargetPageId="5" captionText="FIGURES 6 – 12. Mouthparts of Halobrecta sp. aff. algae (Hardy) (female, Grado, Italy (6 – 11 )) and Ha. sp. cf. halensis Mulsant &amp; Rey, 1873 (female, Lignano, Italy (12 )). 6 — prementum; 7 — hypopharynx; 8 — right maxilla, ventral view; 9 — right galea, ventral view; 10 — right lacinia, ventral view; 11 — right lacinia, dorsal view; 12 — mentum. Scale bar 0.1 mm (6 – 7, 9 – 12), 0.2 mm (8)." httpUri="https://zenodo.org/record/158215/files/figure.png" pageId="3" pageNumber="4">Fig. 9</figureCitation>
); distal comb of lacinia is divided into isolated groups of 5 and 2 closely placed spines (
<figureCitation id="95FC2A332F00222254ADBDCAF672FD21" box="[986,1128,644,670]" captionStart="FIGURES 6 – 12" captionStartId="5.[264,383,1714,1738]" captionTargetBox="[312,1278,649,1705]" captionTargetId="figure@5.[299,1288,635,1710]" captionTargetPageId="5" captionText="FIGURES 6 – 12. Mouthparts of Halobrecta sp. aff. algae (Hardy) (female, Grado, Italy (6 – 11 )) and Ha. sp. cf. halensis Mulsant &amp; Rey, 1873 (female, Lignano, Italy (12 )). 6 — prementum; 7 — hypopharynx; 8 — right maxilla, ventral view; 9 — right galea, ventral view; 10 — right lacinia, ventral view; 11 — right lacinia, dorsal view; 12 — mentum. Scale bar 0.1 mm (6 – 7, 9 – 12), 0.2 mm (8)." httpUri="https://zenodo.org/record/158215/files/figure.png" pageId="3" pageNumber="4">Figs. 10–11</figureCitation>
), middle portion produced medially and covered with numerous fine setae (
<figureCitation id="95FC2A332F00222254CEBDE2F65DFD79" box="[953,1095,684,710]" captionStart="FIGURES 6 – 12" captionStartId="5.[264,383,1714,1738]" captionTargetBox="[312,1278,649,1705]" captionTargetId="figure@5.[299,1288,635,1710]" captionTargetPageId="5" captionText="FIGURES 6 – 12. Mouthparts of Halobrecta sp. aff. algae (Hardy) (female, Grado, Italy (6 – 11 )) and Ha. sp. cf. halensis Mulsant &amp; Rey, 1873 (female, Lignano, Italy (12 )). 6 — prementum; 7 — hypopharynx; 8 — right maxilla, ventral view; 9 — right galea, ventral view; 10 — right lacinia, ventral view; 11 — right lacinia, dorsal view; 12 — mentum. Scale bar 0.1 mm (6 – 7, 9 – 12), 0.2 mm (8)." httpUri="https://zenodo.org/record/158215/files/figure.png" pageId="3" pageNumber="4">Figs. 10–11</figureCitation>
), ventral surface of lacinia with a marginal group of 4 strong setae (
<figureCitation id="95FC2A332F002222543EBD9AF1B8FD51" box="[841,930,724,750]" captionStart="FIGURES 6 – 12" captionStartId="5.[264,383,1714,1738]" captionTargetBox="[312,1278,649,1705]" captionTargetId="figure@5.[299,1288,635,1710]" captionTargetPageId="5" captionText="FIGURES 6 – 12. Mouthparts of Halobrecta sp. aff. algae (Hardy) (female, Grado, Italy (6 – 11 )) and Ha. sp. cf. halensis Mulsant &amp; Rey, 1873 (female, Lignano, Italy (12 )). 6 — prementum; 7 — hypopharynx; 8 — right maxilla, ventral view; 9 — right galea, ventral view; 10 — right lacinia, ventral view; 11 — right lacinia, dorsal view; 12 — mentum. Scale bar 0.1 mm (6 – 7, 9 – 12), 0.2 mm (8)." httpUri="https://zenodo.org/record/158215/files/figure.png" pageId="3" pageNumber="4">Fig. 10</figureCitation>
), dorsal surface of lacinia with a row of 18 weak setae (
<figureCitation id="95FC2A332F0022225566BDB2F07DFCA9" box="[529,615,764,790]" captionStart="FIGURES 6 – 12" captionStartId="5.[264,383,1714,1738]" captionTargetBox="[312,1278,649,1705]" captionTargetId="figure@5.[299,1288,635,1710]" captionTargetPageId="5" captionText="FIGURES 6 – 12. Mouthparts of Halobrecta sp. aff. algae (Hardy) (female, Grado, Italy (6 – 11 )) and Ha. sp. cf. halensis Mulsant &amp; Rey, 1873 (female, Lignano, Italy (12 )). 6 — prementum; 7 — hypopharynx; 8 — right maxilla, ventral view; 9 — right galea, ventral view; 10 — right lacinia, ventral view; 11 — right lacinia, dorsal view; 12 — mentum. Scale bar 0.1 mm (6 – 7, 9 – 12), 0.2 mm (8)." httpUri="https://zenodo.org/record/158215/files/figure.png" pageId="3" pageNumber="4">Fig. 11</figureCitation>
). Labium as in 
<figureCitation id="95FC2A332F0022225452BDB2F1DBFCA9" box="[805,961,764,790]" captionStart="FIGURES 6 – 12" captionStartId="5.[264,383,1714,1738]" captionTargetBox="[312,1278,649,1705]" captionTargetId="figure@5.[299,1288,635,1710]" captionTargetPageId="5" captionText="FIGURES 6 – 12. Mouthparts of Halobrecta sp. aff. algae (Hardy) (female, Grado, Italy (6 – 11 )) and Ha. sp. cf. halensis Mulsant &amp; Rey, 1873 (female, Lignano, Italy (12 )). 6 — prementum; 7 — hypopharynx; 8 — right maxilla, ventral view; 9 — right galea, ventral view; 10 — right lacinia, ventral view; 11 — right lacinia, dorsal view; 12 — mentum. Scale bar 0.1 mm (6 – 7, 9 – 12), 0.2 mm (8)." httpUri="https://zenodo.org/record/158215/files/figure.png" pageId="3" pageNumber="4">Figs. 6–7, 12</figureCitation>
; ligula long and parallel­sided, with narrow base, slightly split apically (
<figureCitation id="95FC2A332F0022225586BC6AF121FC81" box="[753,827,804,830]" captionStart="FIGURES 6 – 12" captionStartId="5.[264,383,1714,1738]" captionTargetBox="[312,1278,649,1705]" captionTargetId="figure@5.[299,1288,635,1710]" captionTargetPageId="5" captionText="FIGURES 6 – 12. Mouthparts of Halobrecta sp. aff. algae (Hardy) (female, Grado, Italy (6 – 11 )) and Ha. sp. cf. halensis Mulsant &amp; Rey, 1873 (female, Lignano, Italy (12 )). 6 — prementum; 7 — hypopharynx; 8 — right maxilla, ventral view; 9 — right galea, ventral view; 10 — right lacinia, ventral view; 11 — right lacinia, dorsal view; 12 — mentum. Scale bar 0.1 mm (6 – 7, 9 – 12), 0.2 mm (8)." httpUri="https://zenodo.org/record/158215/files/figure.png" pageId="3" pageNumber="4">Fig. 6</figureCitation>
); medial area of prementum with 2 pores and 12 pseudopores, lateral areas each with two asetose pores, single setose pore and 9–10 pseudopores (
<figureCitation id="95FC2A332F00222256DDBC3AF3EBFC31" box="[426,497,884,910]" captionStart="FIGURES 6 – 12" captionStartId="5.[264,383,1714,1738]" captionTargetBox="[312,1278,649,1705]" captionTargetId="figure@5.[299,1288,635,1710]" captionTargetPageId="5" captionText="FIGURES 6 – 12. Mouthparts of Halobrecta sp. aff. algae (Hardy) (female, Grado, Italy (6 – 11 )) and Ha. sp. cf. halensis Mulsant &amp; Rey, 1873 (female, Lignano, Italy (12 )). 6 — prementum; 7 — hypopharynx; 8 — right maxilla, ventral view; 9 — right galea, ventral view; 10 — right lacinia, ventral view; 11 — right lacinia, dorsal view; 12 — mentum. Scale bar 0.1 mm (6 – 7, 9 – 12), 0.2 mm (8)." httpUri="https://zenodo.org/record/158215/files/figure.png" pageId="3" pageNumber="4">Fig. 6</figureCitation>
). Hypopharyngeal lobes as in 
<figureCitation id="95FC2A332F002222542DBC3AF1BAFC31" box="[858,928,884,910]" captionStart="FIGURES 6 – 12" captionStartId="5.[264,383,1714,1738]" captionTargetBox="[312,1278,649,1705]" captionTargetId="figure@5.[299,1288,635,1710]" captionTargetPageId="5" captionText="FIGURES 6 – 12. Mouthparts of Halobrecta sp. aff. algae (Hardy) (female, Grado, Italy (6 – 11 )) and Ha. sp. cf. halensis Mulsant &amp; Rey, 1873 (female, Lignano, Italy (12 )). 6 — prementum; 7 — hypopharynx; 8 — right maxilla, ventral view; 9 — right galea, ventral view; 10 — right lacinia, ventral view; 11 — right lacinia, dorsal view; 12 — mentum. Scale bar 0.1 mm (6 – 7, 9 – 12), 0.2 mm (8)." httpUri="https://zenodo.org/record/158215/files/figure.png" pageId="3" pageNumber="4">Fig. 7</figureCitation>
. Labial palpus with setae and present (
<figureCitation id="95FC2A332F002222561ABCD2F3AEFC09" box="[365,436,924,950]" captionStart="FIGURES 6 – 12" captionStartId="5.[264,383,1714,1738]" captionTargetBox="[312,1278,649,1705]" captionTargetId="figure@5.[299,1288,635,1710]" captionTargetPageId="5" captionText="FIGURES 6 – 12. Mouthparts of Halobrecta sp. aff. algae (Hardy) (female, Grado, Italy (6 – 11 )) and Ha. sp. cf. halensis Mulsant &amp; Rey, 1873 (female, Lignano, Italy (12 )). 6 — prementum; 7 — hypopharynx; 8 — right maxilla, ventral view; 9 — right galea, ventral view; 10 — right lacinia, ventral view; 11 — right lacinia, dorsal view; 12 — mentum. Scale bar 0.1 mm (6 – 7, 9 – 12), 0.2 mm (8)." httpUri="https://zenodo.org/record/158215/files/figure.png" pageId="3" pageNumber="4">Fig. 6</figureCitation>
). Mentum (
<figureCitation id="95FC2A332F0022225532BCD2F080FC09" box="[581,666,924,950]" captionStart="FIGURES 6 – 12" captionStartId="5.[264,383,1714,1738]" captionTargetBox="[312,1278,649,1705]" captionTargetId="figure@5.[299,1288,635,1710]" captionTargetPageId="5" captionText="FIGURES 6 – 12. Mouthparts of Halobrecta sp. aff. algae (Hardy) (female, Grado, Italy (6 – 11 )) and Ha. sp. cf. halensis Mulsant &amp; Rey, 1873 (female, Lignano, Italy (12 )). 6 — prementum; 7 — hypopharynx; 8 — right maxilla, ventral view; 9 — right galea, ventral view; 10 — right lacinia, ventral view; 11 — right lacinia, dorsal view; 12 — mentum. Scale bar 0.1 mm (6 – 7, 9 – 12), 0.2 mm (8)." httpUri="https://zenodo.org/record/158215/files/figure.png" pageId="3" pageNumber="4">Fig. 12</figureCitation>
) with concave anterior margin.
</paragraph>
<paragraph id="0D7836B62F002222564FBC8AF16CFAD1" blockId="3.[264,1325,284,1870]" pageId="3" pageNumber="4">
Pronotum (
<figureCitation id="95FC2A332F00222256CBBC8AF00BFC61" box="[444,529,964,990]" captionStart="FIGURES 13 – 17" captionStartId="6.[264,382,1493,1517]" captionTargetBox="[453,1127,301,1448]" captionTargetId="figure@6.[441,1145,290,1457]" captionTargetPageId="6" captionText="FIGURES 13 – 17. Details of Halobrecta sp. aff. algae (Hardy) (female, Grado, Italy). 13 — right antenna; 14 — pronotum; 15 — meso­ and metathorax; 16 — left metatarsal segment 5 and metapretarsus, anterior view; 17 — right metatarsus, posterior view. Scale bar 0.1 mm (16), 0.2 mm (13, 17), 0.4 mm (14 – 15)." httpUri="https://zenodo.org/record/158216/files/figure.png" pageId="3" pageNumber="4">Fig. 14</figureCitation>
) slightly transverse, 1.1–1.2 times as wide as long, with microsetae directed anteriorly along anterior half of midline, posteriorly along posterior half of midline, and laterally in lateral portions of the disc (
<typeStatus id="D27C88142F002222544DBB5AF16CFB91" box="[826,886,1044,1070]" pageId="3" pageNumber="4">Type</typeStatus>
III, 
<bibRefCitation id="69564B472F00222254DEBB5AF6B6FB91" author="Benick" box="[937,1196,1044,1070]" pageId="3" pageNumber="23" refString="Benick, G. &amp; Lohse, G. A. (1974) 14. Tribus: Callicerini (Athetae). In: Freude, H., Harde, K. W. &amp; Lohse, G. A. (Eds.), Die Kafer Mitteleuropas. Band 5, Staphylinidae II (Hypocyphtinae und Aleocharinae). Pselaphidae. Goecke &amp; Evers Verlag, Krefeld, pp. 72 - 220." type="book chapter" year="1974">Benick &amp; Lohse 1974</bibRefCitation>
) (
<figureCitation id="95FC2A332F00222253B2BB5AF701FB91" box="[1221,1307,1044,1070]" captionStart="FIGURES 13 – 17" captionStartId="6.[264,382,1493,1517]" captionTargetBox="[453,1127,301,1448]" captionTargetId="figure@6.[441,1145,290,1457]" captionTargetPageId="6" captionText="FIGURES 13 – 17. Details of Halobrecta sp. aff. algae (Hardy) (female, Grado, Italy). 13 — right antenna; 14 — pronotum; 15 — meso­ and metathorax; 16 — left metatarsal segment 5 and metapretarsus, anterior view; 17 — right metatarsus, posterior view. Scale bar 0.1 mm (16), 0.2 mm (13, 17), 0.4 mm (14 – 15)." httpUri="https://zenodo.org/record/158216/files/figure.png" pageId="3" pageNumber="4">Fig. 14</figureCitation>
); macrosetae long; hypomera fully visible in lateral view. Meso­ and metasternum as in 
<figureCitation id="95FC2A332F002222538BBB72F33DFBC1" captionStart="FIGURES 13 – 17" captionStartId="6.[264,382,1493,1517]" captionTargetBox="[453,1127,301,1448]" captionTargetId="figure@6.[441,1145,290,1457]" captionTargetPageId="6" captionText="FIGURES 13 – 17. Details of Halobrecta sp. aff. algae (Hardy) (female, Grado, Italy). 13 — right antenna; 14 — pronotum; 15 — meso­ and metathorax; 16 — left metatarsal segment 5 and metapretarsus, anterior view; 17 — right metatarsus, posterior view. Scale bar 0.1 mm (16), 0.2 mm (13, 17), 0.4 mm (14 – 15)." httpUri="https://zenodo.org/record/158216/files/figure.png" pageId="3" pageNumber="4">Fig. 15</figureCitation>
, mesosternal process narrow, extending about 3/5 length of mesocoxal cavities, metasternal process short, mesosternum and mesosternal process not carinate medially; relative lengths of mesosternal process: isthmus: metasternal process in ratio of about 3:1:1; mesocoxal cavities margined posteriorly; mesocoxae contiguous. Medial macroseta of mesotibia thin, as long as tibial width. Tarsal segmentation 4­5­5, metatarsal segment 1 longer than segment 2 (
<figureCitation id="95FC2A332F002222556EBA62F075FAF9" box="[537,623,1324,1350]" captionStart="FIGURES 13 – 17" captionStartId="6.[264,382,1493,1517]" captionTargetBox="[453,1127,301,1448]" captionTargetId="figure@6.[441,1145,290,1457]" captionTargetPageId="6" captionText="FIGURES 13 – 17. Details of Halobrecta sp. aff. algae (Hardy) (female, Grado, Italy). 13 — right antenna; 14 — pronotum; 15 — meso­ and metathorax; 16 — left metatarsal segment 5 and metapretarsus, anterior view; 17 — right metatarsus, posterior view. Scale bar 0.1 mm (16), 0.2 mm (13, 17), 0.4 mm (14 – 15)." httpUri="https://zenodo.org/record/158216/files/figure.png" pageId="3" pageNumber="4">Fig. 17</figureCitation>
). One empodial seta, longer than claws (
<figureCitation id="95FC2A332F0022225338BA62F6BCFAF9" box="[1103,1190,1324,1350]" captionStart="FIGURES 13 – 17" captionStartId="6.[264,382,1493,1517]" captionTargetBox="[453,1127,301,1448]" captionTargetId="figure@6.[441,1145,290,1457]" captionTargetPageId="6" captionText="FIGURES 13 – 17. Details of Halobrecta sp. aff. algae (Hardy) (female, Grado, Italy). 13 — right antenna; 14 — pronotum; 15 — meso­ and metathorax; 16 — left metatarsal segment 5 and metapretarsus, anterior view; 17 — right metatarsus, posterior view. Scale bar 0.1 mm (16), 0.2 mm (13, 17), 0.4 mm (14 – 15)." httpUri="https://zenodo.org/record/158216/files/figure.png" pageId="3" pageNumber="4">Fig. 16</figureCitation>
). Posterior margin of elytra emarginate. Wings fully developed.
</paragraph>
<paragraph id="0D7836B62F002222564FBA32F02FF9E1" blockId="3.[264,1325,284,1870]" pageId="3" pageNumber="4">
Abdominal terga 3–5 with moderate basal impressions. Tergum 7 is as long as tergum 6. Punctation on terga 6–7 finer and slightly sparser than on terga 3–5. Tergum 7 with wide white palisade fringe. Posterior margin of female sternum 8 with comb consisting of tiny projections (
<figureCitation id="95FC2A332F00222256A1BABAF02AF9B1" box="[470,560,1524,1550]" captionStart="FIGURES 18 – 21" captionStartId="9.[264,382,1204,1228]" captionTargetBox="[292,1295,296,1174]" captionTargetId="figure@9.[283,1304,284,1181]" captionTargetPageId="9" captionText="FIGURES 18 – 21. Abdominal segment 8 of Halobrecta algophila (Fenyes) (male lectotype (18 – 19) and female paralectotype of At. algophila (20 – 21 )). 18 — male tergum 8; 19 — male sternum 8; 20 — female tergum 8; 21 — female sternum 8. Scale bar 0.2 mm." httpUri="https://zenodo.org/record/158217/files/figure.png" pageId="3" pageNumber="4">Fig. 21</figureCitation>
). Posterior margin of female tergum 8 with irregular comb of scattered projections (
<figureCitation id="95FC2A332F002222557EB952F045F989" box="[521,607,1564,1590]" captionStart="FIGURES 18 – 21" captionStartId="9.[264,382,1204,1228]" captionTargetBox="[292,1295,296,1174]" captionTargetId="figure@9.[283,1304,284,1181]" captionTargetPageId="9" captionText="FIGURES 18 – 21. Abdominal segment 8 of Halobrecta algophila (Fenyes) (male lectotype (18 – 19) and female paralectotype of At. algophila (20 – 21 )). 18 — male tergum 8; 19 — male sternum 8; 20 — female tergum 8; 21 — female sternum 8. Scale bar 0.2 mm." httpUri="https://zenodo.org/record/158217/files/figure.png" pageId="3" pageNumber="4">Fig. 20</figureCitation>
, 
<figureCitation id="95FC2A332F002222551DB952F093F989" box="[618,649,1564,1590]" captionStart="FIGURES 37 – 40" captionStartId="13.[264,382,1414,1438]" captionTargetBox="[292,1310,552,1389]" captionTargetId="figure@13.[280,1321,538,1399]" captionTargetPageId="13" captionText="FIGURES 37 – 40. Abdominal segment 8 of Halobrecta flavipes Thomson (male (37 – 38) and female paralectotypes Atheta pocahontas Casey (39 – 40 )). 37 — male tergum 8; 38 — male sternum 8; 39 — female tergum 8; 40 — female sternum 8. Scale bar 0.2 mm." httpUri="https://zenodo.org/record/158220/files/figure.png" pageId="3" pageNumber="4">39</figureCitation>
) (in some males this comb is also present but may consist of just a few projections).
</paragraph>
<paragraph id="0D7836B62F002222564FB922F378F899" blockId="3.[264,1325,284,1870]" pageId="3" pageNumber="4">
Median lobe of aedeagus without athetine bridge (
<figureCitation id="95FC2A332F00222254E3B922F63DF939" box="[916,1063,1644,1670]" captionStart="FIGURES 22 – 29" captionStartId="10.[264,382,1447,1471]" captionTargetBox="[297,1292,297,1431]" captionTargetId="figure@10.[285,1301,286,1440]" captionTargetPageId="10" captionText="FIGURES 22 – 29. Aedeagus of Halobrecta algophila (Fenyes) (male lectotype of At. algophila (22 – 25), and male from the environs of Bolinas, California (26 – 29 )). 22, 27 — apex of median lobe, parameral view; 23, 26 — apex of median lobe, parameral view; 25, 28 — median lobe, lateral view; 24, 29 — apex of median lobe, lateral view. Scale bar 0.1 mm (22, 24, 27, 29), 0.2 mm (23, 25, 26, 28)." httpUri="https://zenodo.org/record/158218/files/figure.png" pageId="3" pageNumber="4">Figs. 25, 28</figureCitation>
), internal sac without medial lamellae (
<figureCitation id="95FC2A332F00222256A1B9DAF072F911" box="[470,616,1684,1710]" captionStart="FIGURES 62 – 65" captionStartId="20.[264,383,1746,1770]" captionTargetBox="[328,1216,1040,1731]" captionTargetId="figure@20.[320,1227,1026,1736]" captionTargetPageId="20" captionText="FIGURES 62 – 65. Details of Halobrecta sp. aff. algae (Hardy) (male, Grado, Italy (62 – 63 )) and Ha. sp. cf. halensis Mulsant &amp; Rey, 1873 (male, Lignano, Italy (64 – 65 )). 62 — median lobe with everted internal sac, parameral view; 63 — median lobe with everted internal sac, lateral view; 64 — median lobe with partially everted internal sac, parameral view; 65 — median lobe with partially everted internal sac, lateral view. Scale bar 0.2 mm." httpUri="https://zenodo.org/record/158226/files/figure.png" pageId="3" pageNumber="4">Figs. 62–65</figureCitation>
); copulatory piece with pointed apex (
<figureCitation id="95FC2A332F0022225346B9DAF68FF911" box="[1073,1173,1684,1710]" captionStart="FIGURES 30 – 36" captionStartId="11.[264,382,1692,1716]" captionTargetBox="[304,1285,459,1679]" captionTargetId="figure@11.[295,1292,446,1690]" captionTargetPageId="11" captionText="FIGURES 30 – 36. Genitalia of Halobrecta algophila (Fenyes) (male lectotype (30 – 31, 33 – 36) and female paralectotype of At. algophila (32 )). 30 — details of retracted internal sac, abparameral view; 31 — details of retracted internal sac, lateral view; 32 — spermatheca; 33 — left paramere, view from the side facing the median lobe; 34 — left paramere, marginal view; 35 — left paramere, view from the side opposite to the median lobe; 36 — apex of left paramere, marginal view. Scale bar 0.1 mm (32, 36), 0.2 mm (30 – 31, 33 – 35)." httpUri="https://zenodo.org/record/158219/files/figure.png" pageId="3" pageNumber="4">Figs. 30</figureCitation>
, 
<figureCitation id="95FC2A332F00222253D4B9DAF6F5F911" box="[1187,1263,1684,1710]" captionStart="FIGURES 62 – 65" captionStartId="20.[264,383,1746,1770]" captionTargetBox="[328,1216,1040,1731]" captionTargetId="figure@20.[320,1227,1026,1736]" captionTargetPageId="20" captionText="FIGURES 62 – 65. Details of Halobrecta sp. aff. algae (Hardy) (male, Grado, Italy (62 – 63 )) and Ha. sp. cf. halensis Mulsant &amp; Rey, 1873 (male, Lignano, Italy (64 – 65 )). 62 — median lobe with everted internal sac, parameral view; 63 — median lobe with everted internal sac, lateral view; 64 — median lobe with partially everted internal sac, parameral view; 65 — median lobe with partially everted internal sac, lateral view. Scale bar 0.2 mm." httpUri="https://zenodo.org/record/158226/files/figure.png" pageId="3" pageNumber="4">62, 64</figureCitation>
) and without sclerotized suspensoria; paramere with long and narrow apex (
<figureCitation id="95FC2A332F002222533DB9F2F6C0F969" box="[1098,1242,1724,1750]" captionStart="FIGURES 30 – 36" captionStartId="11.[264,382,1692,1716]" captionTargetBox="[304,1285,459,1679]" captionTargetId="figure@11.[295,1292,446,1690]" captionTargetPageId="11" captionText="FIGURES 30 – 36. Genitalia of Halobrecta algophila (Fenyes) (male lectotype (30 – 31, 33 – 36) and female paralectotype of At. algophila (32 )). 30 — details of retracted internal sac, abparameral view; 31 — details of retracted internal sac, lateral view; 32 — spermatheca; 33 — left paramere, view from the side facing the median lobe; 34 — left paramere, marginal view; 35 — left paramere, view from the side opposite to the median lobe; 36 — apex of left paramere, marginal view. Scale bar 0.1 mm (32, 36), 0.2 mm (30 – 31, 33 – 35)." httpUri="https://zenodo.org/record/158219/files/figure.png" pageId="3" pageNumber="4">Figs. 33–36</figureCitation>
); spermatheca short, not divided into distal and proximal portions, with large umbilicus (
<figureCitation id="95FC2A332F0022225387B9AAF332F899" captionStart="FIGURES 30 – 36" captionStartId="11.[264,382,1692,1716]" captionTargetBox="[304,1285,459,1679]" captionTargetId="figure@11.[295,1292,446,1690]" captionTargetPageId="11" captionText="FIGURES 30 – 36. Genitalia of Halobrecta algophila (Fenyes) (male lectotype (30 – 31, 33 – 36) and female paralectotype of At. algophila (32 )). 30 — details of retracted internal sac, abparameral view; 31 — details of retracted internal sac, lateral view; 32 — spermatheca; 33 — left paramere, view from the side facing the median lobe; 34 — left paramere, marginal view; 35 — left paramere, view from the side opposite to the median lobe; 36 — apex of left paramere, marginal view. Scale bar 0.1 mm (32, 36), 0.2 mm (30 – 31, 33 – 35)." httpUri="https://zenodo.org/record/158219/files/figure.png" pageId="3" pageNumber="4">Figs. 32</figureCitation>
, 
<figureCitation id="95FC2A332F0022225643B842F34EF899" box="[308,340,1804,1830]" captionStart="FIGURES 49 – 51" captionStartId="16.[264,382,1105,1129]" captionTargetBox="[346,1246,419,1083]" captionTargetId="figure@16.[331,1257,409,1090]" captionTargetPageId="16" captionText="FIGURES 49 – 51. Genitalia of Halobrecta flavipes Thomson (male (49 – 50) and female paralectotypes of Atheta pocahontas (51 )). 49 — details of retracted internal sac, lateral view; 50 — apex of left paramere, marginal view; 51 — spermatheca. Scale bar 0.1 mm (50 – 51), 0.2 mm (49)." httpUri="https://zenodo.org/record/158223/files/figure.png" pageId="3" pageNumber="4">51</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="45DD653D2F002222564FB87AF704F8F1" box="[312,1310,1844,1870]" pageId="3" pageNumber="4" type="materials_examined">
<paragraph id="0D7836B62F002222564FB87AF704F8F1" blockId="3.[264,1325,284,1870]" box="[312,1310,1844,1870]" pageId="3" pageNumber="4">
<emphasis id="3FB3EAA42F002222564FB87AF3C0F8F1" bold="true" box="[312,474,1844,1870]" pageId="3" pageNumber="4">
<typeStatus id="D27C88142F002222564FB87AF36FF8F1" box="[312,373,1844,1870]" pageId="3" pageNumber="4">Type</typeStatus>
species.
</emphasis>
<taxonomicName id="CAC74D352F002222569FB87AF1BEF8F1" authority="Thomson, 1852" authorityName="Thomson" authorityYear="1852" box="[488,932,1844,1870]" class="Insecta" family="Staphylinidae" genus="Homalota" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="puncticeps">
<emphasis id="3FB3EAA42F002222569FB87AF0F8F8F2" box="[488,738,1844,1869]" italics="true" pageId="3" pageNumber="4">Homalota puncticeps</emphasis>
<bibRefCitation id="69564B472F002222559DB87AF1BEF8F1" author="Thomson" box="[746,932,1844,1870]" pageId="3" pageNumber="25" refString="Thomson, C. G. (1852) Insekt-slagtet Homalota. Ofversigt af Kongl. Vetenskaps-Akademiens Forkhandlingar, 9 (6), 131 - 146." type="journal article" year="1852">Thomson, 1852</bibRefCitation>
</taxonomicName>
, by monotypy (
<bibRefCitation id="69564B472F002222532DB87AF70BF8F1" author="Thomson" box="[1114,1297,1844,1870]" pageId="3" pageNumber="25" refString="Thomson, C G. (1858) Forsok till uppstallning af Sveriges Staphyliner. Ofversigt af Kongl. Vetenskaps-Akademiens Forkhandlingar, 15, 27 - 40." type="journal article" year="1858">Thomson 1858</bibRefCitation>
).
</paragraph>
</subSubSection>
<caption id="59B8663E2F072225567FBAA6F11BF9FA" httpUri="https://zenodo.org/record/158214/files/figure.png" pageId="4" pageNumber="5" targetBox="[294,1291,289,1499]" targetPageId="4">
<paragraph id="0D7836B62F072225567FBAA6F11BF9FA" blockId="4.[264,1323,1512,1605]" pageId="4" pageNumber="5">
<emphasis id="3FB3EAA42F072225567FBAA6F3ADF9BF" bold="true" box="[264,439,1512,1536]" pageId="4" pageNumber="5">FIGURES 1–5.</emphasis>
Mouthparts of 
<emphasis id="3FB3EAA42F0722255506BAA7F109F9BF" box="[625,787,1513,1536]" italics="true" pageId="4" pageNumber="5">
<taxonomicName id="CAC74D352F0722255506BAA7F0F3F9BF" box="[625,745,1513,1536]" class="Insecta" family="Staphylinidae" genus="Halobrecta" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="genus">Halobrecta</taxonomicName>
sp.
</emphasis>
<taxonomicName id="CAC74D352F072225546ABAA7F1F9F9BF" authority="(Hardy)" authorityName="Hardy" baseAuthorityName="Hardy" box="[797,995,1513,1536]" class="Insecta" family="Staphylinidae" genus="Halobrecta" isUncertain="true" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="algae">
<emphasis id="3FB3EAA42F072225546ABAA7F19EF9BF" box="[797,900,1513,1536]" italics="true" pageId="4" pageNumber="5">aff. algae</emphasis>
(Hardy)
</taxonomicName>
(female, Grado, Italy). 1 — labrum; 2 — epipharynx; 3 — left mandible, dorsal view; 4 — left mandible, ventral view; 5 — right mandible, dorsal view. Scale bar 0.1 mm.
</paragraph>
</caption>
<subSubSection id="45DD653D2F072226564FB9C6F07AFD79" lastPageId="7" lastPageNumber="8" pageId="4" pageNumber="5" type="discussion">
<paragraph id="0D7836B62F072224564FB9C6F149FDF1" blockId="4.[264,1323,1672,1899]" lastBlockId="5.[264,1323,284,590]" lastPageId="5" lastPageNumber="6" pageId="4" pageNumber="5">
<emphasis id="3FB3EAA42F072225564FB9C6F3D9F91D" bold="true" box="[312,451,1672,1698]" pageId="4" pageNumber="5">Discussion.</emphasis>
<taxonomicName id="CAC74D352F07222556A5B9C7F04DF91D" box="[466,599,1673,1698]" class="Insecta" family="Staphylinidae" genus="Halobrecta" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="genus">
<emphasis id="3FB3EAA42F07222556A5B9C7F04DF91D" box="[466,599,1673,1698]" italics="true" pageId="4" pageNumber="5">Halobrecta</emphasis>
</taxonomicName>
is usually placed in the tribe 
<taxonomicName id="CAC74D352F07222554C6B9C7F60BF91C" box="[945,1041,1673,1699]" class="Insecta" family="Staphylinidae" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="tribe" tribe="Athetini">Athetini</taxonomicName>
but it lacks the athetine bridge of the median lobe which is considered as an autapomorphy of 
<taxonomicName id="CAC74D352F072225532BB9FFF6A6F974" box="[1116,1212,1713,1739]" class="Insecta" family="Staphylinidae" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="tribe" tribe="Athetini">Athetini</taxonomicName>
(
<bibRefCitation id="69564B472F07222553B9B9FFF35CF94C" author="Seevers" pageId="4" pageNumber="25" refString="Seevers, C. H. (1978) A generic and tribal revision of the North American Aleocharinae (Coleoptera: Staphylinidae). Fieldiana: Zoology, 71, vi + 275 pp." type="book" year="1978">Seevers 1978</bibRefCitation>
; 
<bibRefCitation id="69564B472F0722255626B997F3F3F94C" author="Muona" box="[337,489,1753,1779]" pageId="4" pageNumber="24" refString="Muona, J. (1987) Some aspects of Aleocharinae systematics - a response to Dr. G. A. Lohse. Entomologische Blatter, 83 (1), 19 - 24." type="journal article" year="1987">Muona 1987</bibRefCitation>
; 
<bibRefCitation id="69564B472F0722255682B997F0CCF94C" author="Newton" box="[501,726,1753,1779]" pageId="4" pageNumber="24" refString="Newton, A. F., Thayer, M. K., Ashe, J. S. &amp; Chandler, D. S. (2000) Staphylinidae Latreille, 1802. In: Arnett, R. H., Thomas, M. C. (Eds.), American Beetles. Vol. 1. Archostemata, Myxophaga, Adephaga, Polyphaga: Staphyliniformia. CRC Press, Boca Raton, Florida, pp. 272 - 418." type="book chapter" year="2000">
Newton 
<emphasis id="3FB3EAA42F072225552EB997F089F94D" box="[601,659,1753,1778]" italics="true" pageId="4" pageNumber="5">et al.</emphasis>
2000
</bibRefCitation>
) although this character is also present in the tribes 
<taxonomicName id="CAC74D352F072225567FB84FF3BBF8A4" box="[264,417,1793,1819]" class="Insecta" family="Staphylinidae" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="tribe" tribe="Lomechusini">Lomechusini</taxonomicName>
<bibRefCitation id="69564B472F07222556DFB84FF045F8A4" author="Fleming" box="[424,607,1793,1819]" pageId="4" pageNumber="23" refString="Fleming, J. (1821) Insecta. In: Supplement to the fourth, fifth and sixth editions of the Encyclopaedia Britannica, with preliminary dissertations on the history of the sciences. Vol. 5. Archibald Constable and Company, Edinburgh, pp. 41 - 56." type="book" year="1821">Fleming (1821)</bibRefCitation>
and 
<taxonomicName id="CAC74D352F07222555EFB84FF110F8A4" box="[664,778,1793,1819]" class="Insecta" family="Staphylinidae" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="tribe" tribe="Falagriini">Falagriini</taxonomicName>
Mulsant &amp; 
<bibRefCitation id="69564B472F07222554E1B84FF605F8A4" box="[918,1055,1793,1819]" pageId="4" pageNumber="24" refString="Muslant, E. &amp; Rey, C. (1874) Tribu des Brevipennes. Famille des Aleochariens. Septieme branche: Myrmedoniaires. Annales de la Societe d'Agriculture, Histoire Naturelle et Arts Utiles de Lyon, serie 4. 6 [1873], 33 - 738." type="journal article">Rey (1874)</bibRefCitation>
. The lack of the athetine bridge in 
<taxonomicName id="CAC74D352F07222556C2B867F020F8FD" box="[437,570,1833,1858]" class="Insecta" family="Staphylinidae" genus="Halobrecta" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="genus">
<emphasis id="3FB3EAA42F07222556C2B867F020F8FD" box="[437,570,1833,1858]" italics="true" pageId="4" pageNumber="5">Halobrecta</emphasis>
</taxonomicName>
may suggest that it belongs to 
<taxonomicName id="CAC74D352F07222554B8B867F656F8FC" box="[975,1100,1833,1859]" class="Insecta" family="Staphylinidae" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="tribe" tribe="Oxypodini">Oxypodini</taxonomicName>
<bibRefCitation id="69564B472F072225532EB867F73DF8FC" author="Thomson" box="[1113,1319,1833,1859]" pageId="4" pageNumber="25" refString="Thomson, C. G. (1859) Skandinaviens Coleoptera, synoptiskt bearbetade. Tom 1. Berlingska Boktryckeriet, Lund, v + 290 pp." type="book" year="1859">Thomson (1859)</bibRefCitation>
. Another character traditionally used to distinguish between the aleocharine tribes is the tarsal formula (
<emphasis id="3FB3EAA42F06222456C9BE52F3F2FE8A" box="[446,488,284,309]" italics="true" pageId="5" pageNumber="6">e.g.</emphasis>
, Lohse 1974). In 
<taxonomicName id="CAC74D352F06222455B5BE52F15CFE8A" box="[706,838,284,309]" class="Insecta" family="Staphylinidae" genus="Halobrecta" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="genus">
<emphasis id="3FB3EAA42F06222455B5BE52F15CFE8A" box="[706,838,284,309]" italics="true" pageId="5" pageNumber="6">Halobrecta</emphasis>
</taxonomicName>
it is 4­5­5 and supports the placement of the genus in 
<taxonomicName id="CAC74D352F06222456DEBE0AF017FEE1" box="[425,525,324,350]" class="Insecta" family="Staphylinidae" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="tribe" tribe="Athetini">Athetini</taxonomicName>
. However there are examples of changes in tarsal formula even within a genus (
<taxonomicName id="CAC74D352F06222456BCBE22F1BCFE39" authority="Casey, 1894 (Seevers 1978)" authorityName="Casey, 1894 (Seevers" authorityYear="1978" box="[459,934,364,390]" class="Insecta" family="Staphylinidae" genus="Gyronycha" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="genus">
<emphasis id="3FB3EAA42F06222456BCBE22F056FE3A" box="[459,588,364,389]" italics="true" pageId="5" pageNumber="6">Gyronycha</emphasis>
<bibRefCitation id="69564B472F0622245522BE22F0FDFE39" author="Casey" box="[597,743,364,390]" pageId="5" pageNumber="23" refString="Casey, T. L. (1894) Coleopterological notices. V. Annals of the New York Academy of Sciences, 7 [1893], 281 - 606." type="journal article" year="1894">Casey, 1894</bibRefCitation>
(
<bibRefCitation id="69564B472F062224558FBE22F186FE39" author="Seevers" box="[760,924,364,390]" pageId="5" pageNumber="25" refString="Seevers, C. H. (1978) A generic and tribal revision of the North American Aleocharinae (Coleoptera: Staphylinidae). Fieldiana: Zoology, 71, vi + 275 pp." type="book" year="1978">Seevers 1978</bibRefCitation>
)
</taxonomicName>
, 
<taxonomicName id="CAC74D352F06222454C3BE22F6AEFE39" authority="Casey, 1910" authorityName="Casey" authorityYear="1910" box="[948,1204,364,390]" class="Insecta" family="Staphylinidae" genus="Microlia" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="genus">
<emphasis id="3FB3EAA42F06222454C3BE22F603FE3A" box="[948,1049,364,389]" italics="true" pageId="5" pageNumber="6">Microlia</emphasis>
<bibRefCitation id="69564B472F0622245354BE22F6AEFE39" author="Casey" box="[1059,1204,364,390]" pageId="5" pageNumber="23" refString="Casey, T. L. (1910) New Species of the Staphylinid Tribe Myrmedoniini. Memoirs on the Coleoptera I. The New Era Printing Company, Lancaster, pp. 1 - 183." type="book chapter" year="1910">Casey, 1910</bibRefCitation>
</taxonomicName>
<bibRefCitation id="69564B472F06222453CABE22F37DFE11" author="Gusarov" pageId="5" pageNumber="23" refString="Gusarov, V. I. (2002 a) A revision of the genus Microlia Casey, 1910 (Coleoptera: Staphylinidae: Aleocharinae: Hoplandriini). Zootaxa, 34, 1 - 24." type="book chapter" year="2002" yearSuffix="a">(Gusarov 2002a))</bibRefCitation>
. In addition to lacking the athetine bridge, 
<taxonomicName id="CAC74D352F0622245415BEDAF1FCFE12" box="[866,998,404,429]" class="Insecta" family="Staphylinidae" genus="Halobrecta" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="genus">
<emphasis id="3FB3EAA42F0622245415BEDAF1FCFE12" box="[866,998,404,429]" italics="true" pageId="5" pageNumber="6">Halobrecta</emphasis>
</taxonomicName>
is similar to some 
<taxonomicName id="CAC74D352F06222453B5BEDAF33DFE69" class="Insecta" family="Staphylinidae" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="tribe" tribe="Oxypodini">Oxypodini</taxonomicName>
in having the apex of the paramere long; the spermatheca short with a large umbilicus; the posterior margin of the elytra emarginate; the ligula bisetose, long and narrow, split only at the very apex. The tribal placement of 
<taxonomicName id="CAC74D352F062224543BBD42F1CAFD9A" box="[844,976,524,549]" class="Insecta" family="Staphylinidae" genus="Halobrecta" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="genus">
<emphasis id="3FB3EAA42F062224543BBD42F1CAFD9A" box="[844,976,524,549]" italics="true" pageId="5" pageNumber="6">Halobrecta</emphasis>
</taxonomicName>
cannot be resolved until the phylogeny of 
<taxonomicName id="CAC74D352F06222456DDBD7AF010FDF1" box="[426,522,564,590]" class="Insecta" family="Staphylinidae" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="tribe" tribe="Athetini">Athetini</taxonomicName>
and 
<taxonomicName id="CAC74D352F0622245534BD7AF0DAFDF1" box="[579,704,564,590]" class="Insecta" family="Staphylinidae" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="tribe" tribe="Oxypodini">Oxypodini</taxonomicName>
is analyzed.
</paragraph>
<caption id="59B8663E2F062224567FB9FCF344F8EC" httpUri="https://zenodo.org/record/158215/files/figure.png" pageId="5" pageNumber="6" targetBox="[312,1278,649,1705]" targetPageId="5">
<paragraph id="0D7836B62F062224567FB9FCF344F8EC" blockId="5.[264,1323,1714,1875]" pageId="5" pageNumber="6">
<emphasis id="3FB3EAA42F062224567FB9FCF3D9F975" bold="true" box="[264,451,1714,1738]" pageId="5" pageNumber="6">FIGURES 6–12.</emphasis>
Mouthparts of 
<emphasis id="3FB3EAA42F062224550FB9FCF102F976" box="[632,792,1714,1737]" italics="true" pageId="5" pageNumber="6">
<taxonomicName id="CAC74D352F062224550FB9FCF0EAF976" box="[632,752,1714,1737]" class="Insecta" family="Staphylinidae" genus="Halobrecta" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="genus">Halobrecta</taxonomicName>
sp.
</emphasis>
<taxonomicName id="CAC74D352F0622245456B9FCF1FFF976" authority="(Hardy)" authorityName="Hardy" baseAuthorityName="Hardy" box="[801,997,1714,1737]" class="Insecta" family="Staphylinidae" genus="Halobrecta" isUncertain="true" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="algae">
<emphasis id="3FB3EAA42F0622245456B9FCF192F976" box="[801,904,1714,1737]" italics="true" pageId="5" pageNumber="6">aff. algae</emphasis>
(Hardy)
</taxonomicName>
(female, Grado, Italy (6–11)) and 
<emphasis id="3FB3EAA42F0622245641B99BF01BF953" box="[310,513,1749,1772]" italics="true" pageId="5" pageNumber="6">
Ha. sp. 
<taxonomicName id="CAC74D352F06222456FEB99BF01BF953" authority="Mulsant &amp; Rey, 1873" authorityName="Mulsant &amp; Rey" authorityYear="1873" box="[393,513,1749,1772]" class="Insecta" family="Staphylinidae" genus="Halobrecta" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="halensis">cf. halensis</taxonomicName>
</emphasis>
Mulsant &amp; Rey, 1873 (female, Lignano, Italy (12)). 6 — prementum; 7 — hypopharynx; 8 — right maxilla, ventral view; 9 — right galea, ventral view; 10 — right lacinia, ventral view; 11 — right lacinia, dorsal view; 12 — mentum. Scale bar 0.1 mm (6–7, 9–12), 0.2 mm (8).
</paragraph>
</caption>
<caption id="59B8663E2F052227567FBA9BF3F6F9EB" httpUri="https://zenodo.org/record/158216/files/figure.png" pageId="6" pageNumber="7" targetBox="[453,1127,301,1448]" targetPageId="6">
<paragraph id="0D7836B62F052227567FBA9BF3F6F9EB" blockId="6.[264,1323,1493,1620]" pageId="6" pageNumber="7">
<emphasis id="3FB3EAA42F052227567FBA9BF3D4FA52" bold="true" box="[264,462,1493,1517]" pageId="6" pageNumber="7">FIGURES 13–17.</emphasis>
Details of 
<emphasis id="3FB3EAA42F052227553BBA9BF0F0FA53" box="[588,746,1493,1516]" italics="true" pageId="6" pageNumber="7">
<taxonomicName id="CAC74D352F052227553BBA9BF0DEFA53" box="[588,708,1493,1516]" class="Insecta" family="Staphylinidae" genus="Halobrecta" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="genus">Halobrecta</taxonomicName>
sp.
</emphasis>
<taxonomicName id="CAC74D352F0522275586BA9BF1ABFA53" authority="(Hardy)" authorityName="Hardy" baseAuthorityName="Hardy" box="[753,945,1493,1516]" class="Insecta" family="Staphylinidae" genus="Halobrecta" isUncertain="true" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="algae">
<emphasis id="3FB3EAA42F0522275586BA9BF14CFA53" box="[753,854,1493,1516]" italics="true" pageId="6" pageNumber="7">aff. algae</emphasis>
(Hardy)
</taxonomicName>
(female, Grado, Italy). 13 — right antenna; 14 — pronotum; 15 — meso­ and metathorax; 16 — left metatarsal segment 5 and metapretarsus, anterior view; 17 — right metatarsus, posterior view. Scale bar 0.1 mm (16), 0.2 mm (13, 17), 0.4 mm (14–15).
</paragraph>
</caption>
<paragraph id="0D7836B62F052226564FB9FEF07AFD79" blockId="6.[264,1323,1712,1898]" lastBlockId="7.[264,1324,284,710]" lastPageId="7" lastPageNumber="8" pageId="6" pageNumber="7">
According to a recent review of the British intertidal beetles (
<bibRefCitation id="69564B472F0522275364B9FEF6CCF975" author="Hammond" box="[1043,1238,1712,1738]" lastPageId="6" lastPageNumber="24" pageId="6" pageNumber="23" refString="Hammond, P. M. (2000) 16. Coastal Staphylinidae (rove beetles) in the British Isles, with special reference to saltmarshes. In: Sherwood, B. R., Gardiner, B. G. &amp; Harris T. (Eds.), British Saltmarshes. Joint Symposium on British Saltmarshes organized between the Linnean Society of London, the Royal Society for the Protection of Birds and English Nature (2000: London). Forrest Text, Cardingan, pp. 247 - 302." type="journal article" year="2000">Hammond 2000</bibRefCitation>
) 
<taxonomicName id="CAC74D352F0522275391B9FEF34BF94E" class="Insecta" family="Staphylinidae" genus="Halobrecta" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="genus">
<emphasis id="3FB3EAA42F0522275391B9FEF34BF94E" italics="true" pageId="6" pageNumber="7">Halobrecta</emphasis>
</taxonomicName>
includes at least four valid species, all of them recorded from the British Isles: 
<taxonomicName id="CAC74D352F0522275388B996F3E6F8A5" authority="Hardy, 1851" authorityName="Hardy" authorityYear="1851" class="Insecta" family="Staphylinidae" genus="Halobrecta" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="algae">
<emphasis id="3FB3EAA42F0522275388B996F353F8A6" italics="true" pageId="6" pageNumber="7">Ha. algae</emphasis>
(
<bibRefCitation id="69564B472F052227562CB84EF3E9F8A5" author="Hardy" box="[347,499,1792,1818]" pageId="6" pageNumber="24" refString="Hardy, J. (1851) Family 1. Staphylinidae, Leach. In: Hardy, J. &amp; Bold, T. J. A Catalogue of the Insects of Northumberland and Durham (Part ii). Transactions of the Tyneside Naturalists' Field Club, 2, 21 - 97." type="journal article" year="1851">Hardy, 1851</bibRefCitation>
)
</taxonomicName>
, 
<taxonomicName id="CAC74D352F0522275563B84EF171F8A5" authority="Thomson, 1861" authorityName="Thomson" authorityYear="1861" box="[532,875,1792,1818]" class="Insecta" family="Staphylinidae" genus="Halobrecta" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="flavipes">
<emphasis id="3FB3EAA42F0522275563B84EF0BFF8A6" box="[532,677,1792,1817]" italics="true" pageId="6" pageNumber="7">Ha. flavipes</emphasis>
<bibRefCitation id="69564B472F05222755D9B84EF171F8A5" author="Thomson" box="[686,875,1792,1818]" pageId="6" pageNumber="25" refString="Thomson, C. G. (1861) Skandinaviens Coleoptera, synoptiskt bearbetade. Tom III. Berlingska Boktryckeriet, Lund, 278 pp." type="book" year="1861">Thomson, 1861</bibRefCitation>
</taxonomicName>
, 
<taxonomicName id="CAC74D352F052227540DB84EF6DBF8A5" authority="Sharp, 1869" authorityName="Sharp" authorityYear="1869" box="[890,1217,1792,1818]" class="Insecta" family="Staphylinidae" genus="Halobrecta" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="princeps">
<emphasis id="3FB3EAA42F052227540DB84EF60EF8A6" box="[890,1044,1792,1817]" italics="true" pageId="6" pageNumber="7">Ha. princeps</emphasis>
(
<bibRefCitation id="69564B472F0522275351B84EF6A3F8A5" author="Sharp" box="[1062,1209,1792,1818]" pageId="6" pageNumber="25" refString="Sharp, D. (1869) A revision of the British species of Homalota. Transactions of the Entomological Society of London, 1869 (2 - 3), 91 - 272." type="journal article" year="1869">Sharp, 1869</bibRefCitation>
)
</taxonomicName>
and 
<taxonomicName id="CAC74D352F0522275388B84EF05AF8FD" authority="Fenyes, 1909" authorityName="Fenyes" authorityYear="1909" class="Insecta" family="Staphylinidae" genus="Atheta" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="algophila">
<emphasis id="3FB3EAA42F0522275388B84EF362F8FE" italics="true" pageId="6" pageNumber="7">Ha. algophila</emphasis>
(
<bibRefCitation id="69564B472F05222756F8B866F02DF8FD" author="Fenyes" box="[399,567,1832,1858]" pageId="6" pageNumber="23" refString="Fenyes, A. (1909) New Aleocharinae (Staphylinidae, Col.) of the U. S. Entomological News, 20, 1909, 418 - 425." type="journal article" year="1909">Fenyes, 1909</bibRefCitation>
)
</taxonomicName>
. Two species from 
<collectingCountry id="75D076262F052227543DB866F1DAF8FD" box="[842,960,1832,1858]" name="Singapore" pageId="6" pageNumber="7">Singapore</collectingCountry>
were described by 
<bibRefCitation id="69564B472F05222753C9B866F34CF8D5" author="Cameron" pageId="6" pageNumber="23" refString="Cameron, M. (1920) New species of Staphylinidae from Singapore. Part III. Transactions of the Entomological Society of London, 1920, 212 - 284." type="journal article" year="1920">
<collectingCountry id="75D076262F05222753C9B866F733F8FD" box="[1214,1321,1832,1858]" name="Cameroon" pageId="6" pageNumber="7">Cameron</collectingCountry>
(1920)
</bibRefCitation>
in the genus 
<taxonomicName id="CAC74D352F0522275575B81EF134F8D5" authority="Cameron, 1920" authorityName="Cameron" authorityYear="1920" box="[514,814,1872,1898]" class="Insecta" family="Staphylinidae" genus="Exatheta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="genus">
<emphasis id="3FB3EAA42F0522275575B81EF070F8D6" box="[514,618,1872,1897]" italics="true" pageId="6" pageNumber="7">Exatheta</emphasis>
<bibRefCitation id="69564B472F0522275502B81EF134F8D5" author="Cameron" box="[629,814,1872,1898]" pageId="6" pageNumber="23" refString="Cameron, M. (1920) New species of Staphylinidae from Singapore. Part III. Transactions of the Entomological Society of London, 1920, 212 - 284." type="journal article" year="1920">
<collectingCountry id="75D076262F0522275502B81EF0F9F8D5" box="[629,739,1872,1898]" name="Cameroon" pageId="6" pageNumber="7">Cameron</collectingCountry>
, 1920
</bibRefCitation>
</taxonomicName>
(
<taxonomicName id="CAC74D352F0522275435B81EF1CCF8D6" box="[834,982,1872,1897]" class="Insecta" family="Staphylinidae" genus="Exatheta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="cingulata">
<emphasis id="3FB3EAA42F0522275435B81EF1CCF8D6" box="[834,982,1872,1897]" italics="true" pageId="6" pageNumber="7">E. cingulata</emphasis>
</taxonomicName>
and 
<taxonomicName id="CAC74D352F0522275361B81EF68FF8D6" box="[1046,1173,1872,1897]" class="Insecta" family="Staphylinidae" genus="Exatheta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="consors">
<emphasis id="3FB3EAA42F0522275361B81EF68FF8D6" box="[1046,1173,1872,1897]" italics="true" pageId="6" pageNumber="7">E. consors</emphasis>
</taxonomicName>
) and subsequently transferred to 
<taxonomicName id="CAC74D352F042226556EBE52F084FE8A" box="[537,670,284,309]" class="Insecta" family="Staphylinidae" genus="Halobrecta" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="genus">
<emphasis id="3FB3EAA42F042226556EBE52F084FE8A" box="[537,670,284,309]" italics="true" pageId="7" pageNumber="8">Halobrecta</emphasis>
</taxonomicName>
and synonymized with each other by 
<bibRefCitation id="69564B472F042226530FBE52F73CFE89" author="Sawada" box="[1144,1318,284,310]" pageId="7" pageNumber="24" refString="Sawada, K. (1985) Atheta and its allies of Southeast Asia (Coleoptera: Staphylinidae). IV. Ten oriental species described by M. Bernhauer and M. Cameron. Contributions from the Biological Laboratory, Kyoto University, 27 (1), 91 - 110." type="journal article" year="1985">Sawada (1985</bibRefCitation>
, 
<bibRefCitation id="69564B472F042226567FBE0AF35DFEE1" author="Sawada" box="[264,327,324,350]" lastPageId="7" lastPageNumber="25" pageId="7" pageNumber="24" refString="Sawada, K. (1987) Atheta and its allies of Southeast Asia (Coleoptera: Staphylinidae). V. Singaporean species described in Cameron, 1920. Contributions from the Biological Laboratory, Kyoto University, 27 (2), 137 - 150." type="journal article" year="1987">1987</bibRefCitation>
). 
<bibRefCitation id="69564B472F0422265610BE0AF3EFFEE1" author="Pace" box="[359,501,324,350]" pageId="7" pageNumber="24" refString="Pace, R. (1999) Aleocharinae del Cile (Coleoptera, Staphylinidae). Bollettino del Museo civico di Storia naturale di Verona, 23, 119 - 210." type="journal article" year="1999">Pace (1999)</bibRefCitation>
described 
<taxonomicName id="CAC74D352F042226550EBE0AF168FEE2" box="[633,882,324,349]" class="Insecta" family="Staphylinidae" genus="Halobrecta" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="discipula">
<emphasis id="3FB3EAA42F042226550EBE0AF168FEE2" box="[633,882,324,349]" italics="true" pageId="7" pageNumber="8">Halobrecta discipula</emphasis>
</taxonomicName>
from 
<collectingCountry id="75D076262F04222654C9BE0AF61BFEE1" box="[958,1025,324,350]" name="Chile" pageId="7" pageNumber="8">Chile</collectingCountry>
. I am aware of an additional species of 
<taxonomicName id="CAC74D352F04222656BBBE22F04BFE3A" box="[460,593,364,389]" class="Insecta" family="Staphylinidae" genus="Halobrecta" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="genus">
<emphasis id="3FB3EAA42F04222656BBBE22F04BFE3A" box="[460,593,364,389]" italics="true" pageId="7" pageNumber="8">Halobrecta</emphasis>
</taxonomicName>
(
<emphasis id="3FB3EAA42F0422265516BE25F10DFE3A" bold="true" box="[609,791,363,389]" pageId="7" pageNumber="8">
<collectingCountry id="75D076262F0422265516BE25F108FE3A" box="[609,786,363,389]" name="Australia" pageId="7" pageNumber="8">AUSTRALIA</collectingCountry>
:
</emphasis>
ɗ, Victoria (FMNH); 
<emphasis id="3FB3EAA42F0422265354BE25F6B2FE3A" bold="true" box="[1059,1192,363,389]" pageId="7" pageNumber="8">
<collectingCountry id="75D076262F0422265354BE25F6B9FE3A" box="[1059,1187,363,389]" name="Greece" pageId="7" pageNumber="8">GREECE</collectingCountry>
:
</emphasis>
ɗ, 
<specimenCount id="1BC1FD3F2F04222653A0BE22F354FE11" pageId="7" pageNumber="8" type="generic">5 specimens</specimenCount>
(sex undetermined), Kérkira (J.Sahlberg); ɗ, Elevsís (J.Sahlberg) (FMNH); 
<emphasis id="3FB3EAA42F04222653BBBEDDF731FE12" bold="true" box="[1228,1323,403,429]" pageId="7" pageNumber="8">
<collectingCountry id="75D076262F04222653BBBEDDF73CFE12" box="[1228,1318,403,429]" name="Italy" pageId="7" pageNumber="8">ITALY</collectingCountry>
:
</emphasis>
<specimenCount id="1BC1FD3F2F042226567FBEF2F3BCFE69" box="[264,422,444,470]" pageId="7" pageNumber="8" type="generic">18 specimens</specimenCount>
(sex undetermined), Friuli, Lignano, 
<date id="797910762F0422265429BEF2F1CBFE69" box="[862,977,444,470]" pageId="7" pageNumber="8" value="1929-05-25">25.v.1929</date>
(A.Gagliardi); ɗ, 
<specimenCount id="1BC1FD3F2F04222653D1BEF2F731FE69" box="[1190,1323,444,470]" pageId="7" pageNumber="8" type="generic">1 specimen</specimenCount>
(sex undetermined), Fiumicino, 
<date id="797910762F04222655F6BEAAF0F5FE41" box="[641,751,484,510]" pageId="7" pageNumber="8" value="1898-06-07">7.vi.1898</date>
(FMNH); 
<emphasis id="3FB3EAA42F0422265418BEADF629FE42" bold="true" box="[879,1075,483,509]" pageId="7" pageNumber="8">
<collectingCountry id="75D076262F0422265418BEADF635FE42" box="[879,1071,483,509]" name="Macedonia" pageId="7" pageNumber="8">MACEDONIA</collectingCountry>
:
</emphasis>
ɗ, Vardar (FMNH); 
<emphasis id="3FB3EAA42F042226567FBD45F3CEFD9A" bold="true" box="[264,468,523,549]" pageId="7" pageNumber="8">
<collectingCountry id="75D076262F042226567FBD45F3D5FD9A" box="[264,463,523,549]" name="Serbia" pageId="7" pageNumber="8">YUGOSLAVIA</collectingCountry>
:
</emphasis>
<specimenCount id="1BC1FD3F2F042226569EBD42F066FD99" box="[489,636,524,550]" pageId="7" pageNumber="8" type="generic">5 specimens</specimenCount>
(sex undetermined), Sutorina (Paganetti); ɗ, Ψ, Sutorina near Castelnuovo; ɗ, 
<specimenCount id="1BC1FD3F2F0422265572BD7AF08FFDF1" box="[517,661,564,590]" pageId="7" pageNumber="8" type="generic">2 specimens</specimenCount>
(sex undetermined), Castelnuovo (Hummler) (FMNH)). This species differs from the other four species in the shape of the median lobe (
<figureCitation id="95FC2A332F04222653CBBD12F33CFD21" captionStart="FIGURES 62 – 65" captionStartId="20.[264,383,1746,1770]" captionTargetBox="[328,1216,1040,1731]" captionTargetId="figure@20.[320,1227,1026,1736]" captionTargetPageId="20" captionText="FIGURES 62 – 65. Details of Halobrecta sp. aff. algae (Hardy) (male, Grado, Italy (62 – 63 )) and Ha. sp. cf. halensis Mulsant &amp; Rey, 1873 (male, Lignano, Italy (64 – 65 )). 62 — median lobe with everted internal sac, parameral view; 63 — median lobe with everted internal sac, lateral view; 64 — median lobe with partially everted internal sac, parameral view; 65 — median lobe with partially everted internal sac, lateral view. Scale bar 0.2 mm." httpUri="https://zenodo.org/record/158226/files/figure.png" pageId="7" pageNumber="8">Figs. 64– 65</figureCitation>
) and may be conspecific with 
<taxonomicName id="CAC74D352F04222655EABDCAF65BFD21" authority="Mulsant &amp; Rey, 1873" authorityName="Mulsant &amp; Rey" authorityYear="1873" box="[669,1089,644,670]" class="Insecta" family="Staphylinidae" genus="Halobrecta" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="halensis">
<emphasis id="3FB3EAA42F04222655EABDCAF128FD22" box="[669,818,644,669]" italics="true" pageId="7" pageNumber="8">Ha. halensis</emphasis>
Mulsant &amp; 
<bibRefCitation id="69564B472F04222654B0BDCAF65BFD21" author="Rey" box="[967,1089,644,670]" pageId="7" pageNumber="24" refString="Muslant, E. &amp; Rey, C. (1873) Description de divers Coleopteres Brevipennes nouveaux ou peu connus. Opuscules Entomologiques, 15, 147 - 189." type="journal article" year="1873">Rey, 1873</bibRefCitation>
</taxonomicName>
described from the coast of Languedoc (
<collectingCountry id="75D076262F042226568ABDE2F04EFD79" box="[509,596,684,710]" name="France" pageId="7" pageNumber="8">France</collectingCountry>
).
</paragraph>
</subSubSection>
</treatment>
</document>