750 lines
109 KiB
XML
750 lines
109 KiB
XML
<document id="99B1694E4BB76505103A2E6CC8693A4F" ID-CLB-Dataset="2777" ID-DOI="10.1206/0003-0090.452.1.1" ID-GBIF-Dataset="01c03222-2a9d-47ec-9f63-776920b6a540" ID-ISSN="0003-0090" ID-Zenodo-Dep="5825896" checkinTime="1638304321381" checkinUser="carolina" docAuthor="Goodman, Aaron M., Prendini, Lorenzo, Francke, Oscar F. & Esposito, Lauren A." docDate="2021" docId="510E87A56D5AFF9E2436FBA1FF74FB62" docLanguage="en" docName="BulAmeMusNatHis.2021.452.1-92.pdf" docOrigin="Bulletin of the American Museum of Natural History 2021 (452)" docStyle="DocumentStyle:915933466F796C9C739DF4DB6B8DCFA6.9:BulAmeMusNatHis.2011-.journal_article.1cover.type1" docStyleId="915933466F796C9C739DF4DB6B8DCFA6" docStyleName="BulAmeMusNatHis.2011-.journal_article.1cover.type1" docStyleVersion="9" docTitle="Centruroides berstoni Goodman & Prendini & Francke & Esposito 2021, sp. nov." docType="treatment" docVersion="6" lastPageNumber="13" masterDocId="AD37FFDD6D52FF91273DFFC9FFB8FFF2" masterDocTitle="Systematic Revision Of The Arboreal Neotropical “ Thorellii ” Clade Of Centruroides Marx, 1890, Bark Scorpions (Buthidae C. L. Koch, 1837) With Descriptions Of Six New Species" masterLastPageNumber="93" masterPageNumber="1" pageNumber="8" updateTime="1698480871315" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
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<mods:title id="DDD591BD318FDB86C23E14E6FD34126F">Systematic Revision Of The Arboreal Neotropical “ Thorellii ” Clade Of Centruroides Marx, 1890, Bark Scorpions (Buthidae C. L. Koch, 1837) With Descriptions Of Six New Species</mods:title>
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<mods:namePart id="DC095996CB124FF2274B1F692BC10585">Goodman, Aaron M.</mods:namePart>
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<mods:affiliation id="1995D2599373739C11D0151B8ED9D77E">Graduate School and University Center, City University of New York; Division of Invertebrate Zoology, American Museum of Natural History; Institute for Biodiversity Science and Sustainability, California Academy of Sciences</mods:affiliation>
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<mods:namePart id="14B3E06E68BE8621101EE06A489030B2">Prendini, Lorenzo</mods:namePart>
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<mods:affiliation id="31B58B7072646CA70935913F096EA732">Arachnology Lab and Scorpion Systematics Research Group, Division of Invertebrate Zoology, American Museum of Natural History</mods:affiliation>
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<mods:namePart id="3D62724C883116B50258BA7E5EA85493">Francke, Oscar F.</mods:namePart>
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<mods:affiliation id="84DB5C0BFF5C4A86F2F466F55DEB2BAA">Colección Nacional de Arácnidos, Departamento de Zoología, Instituto de Biología, Universidad Nacional Autónoma de México</mods:affiliation>
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<mods:namePart id="ABA337E7E169373130BDBB131E7E7C92">Esposito, Lauren A.</mods:namePart>
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<mods:affiliation id="B9210CBF157AAC146BAD9307BAB4299D">Graduate School and University Center, City University of New York; Division of Invertebrate Zoology, American Museum of Natural History; Institute for Biodiversity Science and Sustainability, California Academy of Sciences</mods:affiliation>
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<treatment id="510E87A56D5AFF9E2436FBA1FF74FB62" ID-DOI="http://doi.org/10.5281/zenodo.5825906" ID-GBIF-Taxon="191919053" ID-Zenodo-Dep="5825906" LSID="urn:lsid:plazi:treatment:510E87A56D5AFF9E2436FBA1FF74FB62" httpUri="http://treatment.plazi.org/id/510E87A56D5AFF9E2436FBA1FF74FB62" lastPageId="15" lastPageNumber="13" pageId="8" pageNumber="8">
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<heading id="825081DF6D5AFF992436FBA1FBE6FB72" box="[779,1118,1128,1153]" centered="true" fontSize="9" level="2" pageId="8" pageNumber="8" reason="2">
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<taxonomicName id="1EA74D306D5AFF992436FBA1FC46FB72" authority="Goodman & Prendini & Francke & Esposito, 2021" authorityName="Goodman & Prendini & Francke & Esposito" authorityYear="2021" box="[779,1022,1128,1152]" class="Insecta" family="Cerambycidae" genus="Centruroides" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="8" phylum="Arthropoda" rank="species" species="berstoni" status="sp. nov.">
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<emphasis id="EBD3EAA16D5AFF992436FBA1FC46FB72" bold="true" box="[779,1022,1128,1152]" italics="true" pageId="8" pageNumber="8">Centruroides berstoni</emphasis>
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<taxonomicNameLabel id="F0E057DA6D5AFF992336FBA0FBE6FB72" box="[1035,1118,1129,1152]" pageId="8" pageNumber="8" rank="species">sp. nov.</taxonomicNameLabel>
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<figureCitation id="419C2A366D5AFF99258DFB53FCFBFB40" box="[688,835,1178,1203]" captionStart="FIGURE 1" captionStartId="9.[109,195,1628,1649]" captionTargetBox="[166,1154,224,1599]" captionTargetId="figure-13@9.[166,1154,224,1599]" captionTargetPageId="9" captionText="FIGURE 1. Representative species of the arboreal Neotropical “thorellii” clade of Centruroides Marx, 1890, bark scorpions (Buthidae C.L. Koch, 1837), habitus in life. A, C. C. schmidti Sissom, 1995, A. ♀, El Progreso, Finca El Olvido, Las Minas, Guatemala, C. ♂, Zacapa, Guatemala. B, D. C. berstoni, sp. nov., B. ♀, D. ♂, Izabal, Rio Dulce, Hotel Tijax, Guatemala. Note elongated metasoma in males. Both species were collected in vegetation, 3–4 m above ground level." figureDoi="http://doi.org/10.5281/zenodo.5825898" httpUri="https://zenodo.org/record/5825898/files/figure.png" pageId="8" pageNumber="8">Figures 1B, D</figureCitation>
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,
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<figureCitation id="419C2A366D5AFF992473FB52FCE4FB41" box="[846,860,1179,1203]" captionStart="FIGURE 2" captionStartId="10.[108,194,978,999]" captionTargetBox="[109,1184,232,938]" captionTargetId="figure-310@10.[108,1212,224,947]" captionTargetPageId="10" captionText="FIGURE 2. Phylogeny of the arboreal Neotropical “thorellii” clade of Centruroides Marx, 1890, bark scorpions (Buthidae C.L. Koch, 1837), obtained by simultaneous phylogenetic analysis of 112 morphological characters and DNA sequences of the mitochondrial cytochrome oxidase c subunit I gene. Posterior probabilities, obtained with Bayesian inference, and nonparametric bootstraps, obtained with maximum likelihood, indicated above and below branches for corresponding nodes." figureDoi="http://doi.org/10.5281/zenodo.5825902" httpUri="https://zenodo.org/record/5825902/files/figure.png" pageId="8" pageNumber="8">2</figureCitation>
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<figureCitation id="419C2A366D5AFF992455FB52FCCFFB41" box="[872,887,1179,1203]" captionStart="FIGURE 4" captionStartId="13.[109,195,1048,1069]" captionTargetBox="[111,1203,241,1014]" captionTargetId="figure-13@13.[107,1211,224,1022]" captionTargetPageId="13" captionText="FIGURE 4. Map of southeastern Mexico and northwestern Central America, plotting known locality records of five species of the arboreal Neotropical “thorellii” clade of Centruroides Marx, 1890, bark scorpions (Buthidae C.L. Koch, 1837): C. berstoni, sp. nov. (circles); C. hamadryas, sp. nov. (diamonds); C. hoffmanni Armas, 1996 (triangles); C. schmidti Sissom, 1995 (squares); C. yucatanensis, sp. nov. (crosses)." figureDoi="http://doi.org/10.5281/zenodo.5825908" httpUri="https://zenodo.org/record/5825908/files/figure.png" pageId="8" pageNumber="8">4</figureCitation>
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<figureCitation id="419C2A366D5AFF9924BCFB53FC7AFB40" box="[897,962,1178,1202]" captionStart="FIGURE 6" captionStartId="18.[107,193,1647,1668]" captionTargetBox="[158,1159,223,1618]" captionTargetId="figure-10@18.[158,1159,223,1618]" captionTargetPageId="18" captionText="FIGURE 6. Centruroides Marx, 1890, carapace, dorsal aspect. A, B. C. hamadryas, sp. nov., A. holotype ♂ (CNAN T01408), B. paratype ♀ (CNAN T01415). C, D. C. berstoni, sp. nov., C. holotype ♂ (CASENT 9073325), D. paratype ♀ (CASENT 9073313). E, F. C. schmidti Sissom, 1995, E. ♂ (CASENT 9073316), F. ♀ (CASENT 9073317). Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5825914" httpUri="https://zenodo.org/record/5825914/files/figure.png" pageId="8" pageNumber="8">6C, D</figureCitation>
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<figureCitation id="419C2A366D5AFF9924F0FB52FBB5FB40" box="[973,1037,1179,1203]" captionStart="FIGURE 9" captionStartId="24.[107,193,1676,1697]" captionTargetBox="[146,1171,224,1647]" captionTargetId="figure-9@24.[146,1171,224,1647]" captionTargetPageId="24" captionText="FIGURE 9. Centruroides Marx, 1890, pectines and sternum, ventral aspect. A, B. C. hamadryas, sp. nov., A. holotype ♂ (CNAN T01408), B. paratype ♀ (CNAN T01415). C, D. C. berstoni, sp. nov., C. holotype ♂ (CASENT 9073325), D. paratype ♀ (CASENT 9073313). E, F. C. schmidti Sissom, 1995, E. ♂ (CASENT 9073316), F. ♀ (CASENT 9073317). Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5825920" httpUri="https://zenodo.org/record/5825920/files/figure.png" pageId="8" pageNumber="8">9C, D</figureCitation>
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<figureCitation id="419C2A366D5AFF992325FB52FBFBFB41" box="[1048,1091,1179,1203]" captionStart="FIGURE 13" captionStartId="30.[108,194,1021,1042]" captionTargetBox="[116,1203,224,993]" captionTargetId="figure-247@30.[116,1203,224,993]" captionTargetPageId="30" captionText="FIGURE 13. Centruroides Marx, 1890, pedipalp chela, retrolateral aspect. A. C. hamadryas, sp. nov., holotype ♂ (CNAN T01408). B. C. berstoni, sp. nov., holotype ♂ (CASENT 9073325). C. C. schmidti Sissom, 1995, ♂ (CASENT 9073316). Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5825928" httpUri="https://zenodo.org/record/5825928/files/figure.png" pageId="8" pageNumber="8">13B</figureCitation>
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<figureCitation id="419C2A366D5AFF992373FB52FBC1FB41" box="[1102,1145,1179,1203]" captionStart="FIGURE 14" captionStartId="31.[109,195,1174,1195]" captionTargetBox="[116,1204,224,1147]" captionTargetId="figure-218@31.[116,1204,222,1147]" captionTargetPageId="31" captionText="FIGURE 14. Centruroides Marx, 1890, pedipalp chela, retrolateral aspect. A. C. hamadryas, sp. nov., paratype ♀ (CNAN T01415). B. C. berstoni, sp. nov., paratype ♀ (CASENT 9073313). C. C. schmidti Sissom, 1995, ♀ (CASENT 9073317). Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5825930" httpUri="https://zenodo.org/record/5825930/files/figure.png" pageId="8" pageNumber="8">14B</figureCitation>
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<figureCitation id="419C2A366D5AFF9923B9FB52FD62FB21" captionStart="FIGURE 17" captionStartId="38.[107,193,1462,1483]" captionTargetBox="[224,1093,224,1434]" captionTargetId="figure-9@38.[224,1093,224,1434]" captionTargetPageId="38" captionText="FIGURE 17. Centruroides Marx, 1890, metasomal segments I and II, dorsal aspect. A, D. C. rileyi Sissom, 1995, A. ♂ (CNAN SC4002), D. ♀ (CNAN SC4003). B, E. C. hamadryas, sp. nov., B. holotype ♂ (CNAN T01408), E. paratype ♀ (CNAN T01415). C, F. C. hoffmanni Armas, 1996, C. ♂, F. ♀ (CNAN SC3996). G, J. C. cuauhmapan, sp. nov., G. holotype ♂ (CNAN T01396), J. paratype ♀ (CNAN T01399). H, K. C. berstoni, sp. nov., H. holotype ♂ (CASENT 9073325), K. paratype ♀ (CASENT 9073313). I, L. C. chanae, sp. nov., I. holotype ♂ (CNAN T01403), L. paratype ♀ (CNAN T01405). M, P. C. catemacoensis, sp. nov., M. holotype ♂ (CNAN T01424), P.paratype ♀ (CNAN T01423). N, Q. C. schmidti Sissom, 1995, N. ♂ (CASENT 9073316), Q. ♀ (CASENT 9073317). O, R. C. yucatanensis, sp. nov., O. holotype ♂ (CNAN T01416), R. paratype ♀ (CNAN T01417. Scale bars = 2 mm." figureDoi="http://doi.org/10.5281/zenodo.5825936" httpUri="https://zenodo.org/record/5825936/files/figure.png" pageId="8" pageNumber="8">17H, K</figureCitation>
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<figureCitation id="419C2A366D5AFF9925D8FB75FC8CFB21" box="[741,820,1212,1236]" captionStart="FIGURE 18" captionStartId="39.[109,195,1462,1483]" captionTargetBox="[225,1095,224,1434]" captionTargetId="figure-13@39.[225,1095,224,1434]" captionTargetPageId="39" captionText="FIGURE 18. Centruroides Marx, 1890, metasomal segments I and II, ventral aspect. A, D. C. rileyi Sissom, 1995, A. ♂ (CNAN SC4002), D. ♀ (CNAN SC4003). B, E. C. hamadryas, sp. nov., B. holotype ♂ (CNAN T01408), E. paratype ♀ (CNAN T01415). C, F. C. hoffmanni Armas, 1996, C. ♂, F. ♀ (CNAN SC3996). G, J. C. cuauhmapan, sp. nov., G. holotype ♂ (CNAN T01396), J. paratype ♀ (CNAN T01399). H, K. C. berstoni, sp. nov., H. holotype ♂ (CASENT 9073325), K. paratype ♀ (CASENT 9073313). I, L. C. chanae, sp. nov., I. holotype ♂ (CNAN T01403), L. paratype ♀ (CNAN T01405). M, P. C. catemacoensis, sp. nov., M. holotype ♂ (CNAN T01424), P.paratype ♀ (CNAN T01423). N, Q. C. schmidti Sissom, 1995, N. ♂ (CASENT 9073316), Q. ♀ (CASENT 9073317). O, R. C. yucatanensis, sp. nov., O. holotype ♂ (CNAN T01416), R. paratype ♀ (CNAN T01417. Scale bars = 2 mm." figureDoi="http://doi.org/10.5281/zenodo.5825938" httpUri="https://zenodo.org/record/5825938/files/figure.png" pageId="8" pageNumber="8">18H, K</figureCitation>
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<figureCitation id="419C2A366D5AFF992402FB75FC37FB21" box="[831,911,1212,1236]" captionStart="FIGURE 19" captionStartId="40.[108,193,1460,1481]" captionTargetBox="[229,1098,224,1434]" captionTargetId="figure-172@40.[229,1098,224,1434]" captionTargetPageId="40" captionText="FIGURE 19. Centruroides Marx, 1890, metasomal segments I and II, lateral aspect. A, D. C. rileyi Sissom, 1995, A. ♂ (CNAN SC4002), D. ♀ (CNAN SC4003). B, E. C. hamadryas, sp. nov., B. holotype ♂ (CNAN T01408), E. paratype ♀ (CNAN T01415). C, F. C. hoffmanni Armas, 1996, C. ♂, F. ♀ (CNAN SC3996). G, J. C. cuauhmapan, sp. nov., G. holotype ♂ (CNAN T01396), J. paratype ♀ (CNAN T01399). H, K. C. berstoni, sp. nov., H. holotype ♂ (CASENT 9073325), K. paratype ♀ (CASENT 9073313). I, L. C. chanae, sp. nov., I. holotype ♂ (CNAN T01403), L. paratype ♀ (CNAN T01405). M, P. C. catemacoensis, sp. nov., M. holotype ♂ (CNAN T01424), P. paratype ♀ (CNAN T01423). N, Q. C. schmidti Sissom, 1995, N. ♂ (CASENT 9073316), Q. ♀ (CASENT 9073317). O, R. C. yucatanensis, sp. nov., O. holotype ♂ (CNAN T01416), R. paratype ♀ (CNAN T01417). Scale bars = 2 mm." figureDoi="http://doi.org/10.5281/zenodo.5825942" httpUri="https://zenodo.org/record/5825942/files/figure.png" pageId="8" pageNumber="8">19H, K</figureCitation>
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|
||
<figureCitation id="419C2A366D5AFF9924A7FB75FC51FB21" box="[922,1001,1212,1236]" captionStart="FIGURE 20" captionStartId="44.[108,194,1460,1481]" captionTargetBox="[224,1093,224,1434]" captionTargetId="figure-9@44.[224,1093,224,1434]" captionTargetPageId="44" captionText="FIGURE 20. Centruroides Marx, 1890, metasomal segment V, dorsal aspect. A, D. C. rileyi Sissom, 1995, A. ♂ (CNAN SC4002), D. ♀ (CNAN SC4003). B, E. C. hamadryas, sp. nov., B. holotype ♂ (CNAN T01408), E. paratype ♀ (CNAN T01415). C, F. C. hoffmanni Armas, 1996, C. ♂, F. ♀ (CNAN SC3996). G, J. C. cuauhmapan, sp. nov., G. holotype ♂ (CNAN T01396), J. paratype ♀ (CNAN T01399). H, K. C. berstoni, sp. nov., H. holotype ♂ (CASENT 9073325), K. paratype ♀ (CASENT 9073313). I, L. C. chanae, sp. nov., I. holotype ♂ (CNAN T01403), L. paratype ♀ (CNAN T01405). M, P. C. catemacoensis, sp. nov., M. holotype ♂ (CNAN T01424), P. paratype ♀ (CNAN T01423). N, Q. C. schmidti Sissom, 1995, N. ♂ (CASENT 9073316), Q. ♀ (CASENT 9073317). O, R. C. yucatanensis, sp. nov., O. holotype ♂ (CNAN T01416), R. paratype ♀ (CNAN T01417). Scale bars = 2 mm." figureDoi="http://doi.org/10.5281/zenodo.5825944" httpUri="https://zenodo.org/record/5825944/files/figure.png" pageId="8" pageNumber="8">20H, K</figureCitation>
|
||
,
|
||
<figureCitation id="419C2A366D5AFF9924C9FB75FBFBFB21" box="[1012,1091,1212,1236]" captionStart="FIGURE 21" captionStartId="45.[107,193,1461,1482]" captionTargetBox="[226,1094,224,1434]" captionTargetId="figure-14@45.[226,1094,224,1434]" captionTargetPageId="45" captionText="FIGURE 21. Centruroides Marx, 1890, metasomal segment V, ventral aspect. A, D. C. rileyi Sissom, 1995, A. ♂ (CNAN SC4002), D. ♀ (CNAN SC4003). B, E. C. hamadryas, sp. nov., B. holotype ♂ (CNAN T01408), E. paratype ♀ (CNAN T01415). C, F. C. hoffmanni Armas, 1996, C. ♂, F. ♀ (CNAN SC3996). G, J. C. cuauhmapan, sp. nov., G. holotype ♂ (CNAN T01396), J. paratype ♀ (CNAN T01399). H, K. C berstoni, sp. nov., H. holotype ♂ (CASENT 9073325), K. paratype ♀ (CASENT 9073313). I, L. C. chanae, sp. nov., I. holotype ♂ (CNAN T01403), L. paratype ♀ (CNAN T01405). M, P. C. catemacoensis, sp. nov., M. holotype ♂ (CNAN T01424), P. paratype ♀ (CNAN T01423). N, Q. C. schmidti Sissom, 1995, N. ♂ (CASENT 9073316), Q. ♀ (CASENT 9073317). O, R. C. yucatanensis, sp. nov., O. holotype ♂ (CNAN T01416), R. paratype ♀ (CNAN T01417). Scale bars = 2 mm." figureDoi="http://doi.org/10.5281/zenodo.5825946" httpUri="https://zenodo.org/record/5825946/files/figure.png" pageId="8" pageNumber="8">21H, K</figureCitation>
|
||
,
|
||
<figureCitation id="419C2A366D5AFF992373FB75FB26FB21" box="[1102,1182,1212,1236]" captionStart="FIGURE 22" captionStartId="46.[109,195,1462,1483]" captionTargetBox="[223,1094,224,1434]" captionTargetId="figure-9@46.[223,1094,224,1434]" captionTargetPageId="46" captionText="FIGURE 22. Centruroides Marx, 1890, metasomal segment V, lateral aspect. A, D. C. rileyi Sissom, 1995, A. ♂ (CNAN SC4002), D. ♀ (CNAN SC4003). B, E. C. hamadryas, sp. nov., B. holotype ♂ (CNAN T01408), E. paratype ♀ (CNAN T01415). C, F. C. hoffmanni Armas, 1996, C. ♂, F. ♀ (CNAN SC3996). G, J. C. cuauhmapan, sp. nov., G. holotype ♂ (CNAN T01396), J. paratype ♀ (CNAN T01399). H, K. C. berstoni, sp. nov., H. holotype ♂ (CASENT 9073325), K. paratype ♀ (CASENT 9073313). I, L. C. chanae, sp. nov., I. holotype ♂ (CNAN T01403), L. paratype ♀ (CNAN T01405). M, P. C. catemacoensis, sp. nov., M. holotype ♂ (CNAN T01424), P.paratype ♀ (CNAN T01423). N, Q. C. schmidti Sissom, 1995, N. ♂ (CASENT 9073316), Q. ♀ (CASENT 9073317). O, R. C. yucatanensis, sp. nov., O. holotype ♂ (CNAN T01416), R. paratype ♀ (CNAN T01417). Scale bars = 2 mm." figureDoi="http://doi.org/10.5281/zenodo.5825948" httpUri="https://zenodo.org/record/5825948/files/figure.png" pageId="8" pageNumber="8">22H, K</figureCitation>
|
||
,
|
||
<figureCitation id="419C2A366D5AFF992580FB14FCB4FB06" box="[701,780,1245,1269]" captionStart="FIGURE 23" captionStartId="50.[108,195,1463,1484]" captionTargetBox="[224,1093,224,1434]" captionTargetId="figure-166@50.[224,1093,224,1434]" captionTargetPageId="50" captionText="FIGURE 23. Centruroides Marx, 1890, telson, dorsal aspect. A, D. C. rileyi Sissom, 1995, A. ♂ (CNAN SC4002), D. ♀ (CNAN SC4003). B, E. C. hamadryas, sp. nov., B. holotype ♂ (CNAN T01408), E. paratype ♀ (CNAN T01415). C, F. C. hoffmanni Armas, 1996, C. ♂, F. ♀ (CNAN SC3996). G, J. C. cuauhmapan, sp. nov., G. holotype ♂ (CNAN T01396), J. paratype ♀ (CNAN T01399). H, K. C. berstoni, sp. nov., H. holotype ♂ (CASENT 9073325), K. paratype ♀ (CASENT 9073313). I, L. C. chanae, sp. nov., I. holotype ♂ (CNAN T01403), L. paratype ♀ (CNAN T01405). M, P. C. catemacoensis, sp. nov., M. holotype ♂ (CNAN T01424), P.paratype ♀ (CNAN T01423). N, Q. C. schmidti Sissom, 1995, N. ♂ (CASENT 9073316), Q. ♀ (CASENT 9073317). O, R. C. yucatanensis, sp. nov., O. holotype ♂ (CNAN T01416), R. paratype ♀ (CNAN T01418). Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5825950" httpUri="https://zenodo.org/record/5825950/files/figure.png" pageId="8" pageNumber="8">23H, K</figureCitation>
|
||
,
|
||
<figureCitation id="419C2A366D5AFF992425FB14FCDFFB06" box="[792,871,1245,1269]" captionStart="FIGURE 24" captionStartId="51.[108,195,1463,1484]" captionTargetBox="[225,1095,224,1434]" captionTargetId="figure-169@51.[225,1095,224,1434]" captionTargetPageId="51" captionText="FIGURE 24. Centruroides Marx, 1890, telson, ventral aspect. A, D. C. rileyi Sissom, 1995, A. ♂ (CNAN SC4002), D. ♀ (CNAN SC4003). B, E. C. hamadryas, sp. nov., B. holotype ♂ (CNAN T01408), E. paratype ♀ (CNAN T01415). C, F. C. hoffmanni Armas, 1996, C. ♂, F. ♀ (CNAN SC3996). G, J. C. cuauhmapan, sp. nov., G. holotype ♂ (CNAN T01396), J. paratype ♀ (CNAN T01399). H, K. C. berstoni, sp. nov., H. holotype ♂ (CASENT 9073325), K. paratype ♀ (CASENT 9073313). I, L. C. chanae, sp. nov., I. holotype ♂ (CNAN T01403), L. paratype ♀ (CNAN T01405). M, P. C. catemacoensis, sp. nov., M. holotype ♂ (CNAN T01424), P.paratype ♀ (CNAN T01423). N, Q. C. schmidti Sissom, 1995, N. ♂ (CASENT 9073316), Q. ♀ (CASENT 9073317). O, R. C. yucatanensis, sp. nov., O. holotype ♂ (CNAN T01416), R. paratype ♀ (CNAN T01418). Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5825952" httpUri="https://zenodo.org/record/5825952/files/figure.png" pageId="8" pageNumber="8">24H, K</figureCitation>
|
||
,
|
||
<figureCitation id="419C2A366D5AFF99244FFB15FC79FB06" box="[882,961,1244,1268]" captionStart="FIGURE 25" captionStartId="52.[108,195,1463,1484]" captionTargetBox="[224,1093,224,1434]" captionTargetId="figure-9@52.[224,1093,224,1434]" captionTargetPageId="52" captionText="FIGURE 25. Centruroides Marx, 1890, telson, lateral aspect. A, D. C. rileyi Sissom, 1995, A. ♂ (CNAN SC4002), D. ♀ (CNAN SC4003). B, E. C. hamadryas, sp. nov., B. holotype ♂ (CNAN T01408), E. paratype ♀ (CNAN T01415). C, F. C. hoffmanni Armas, 1996, C. ♂, F. ♀ (CNAN SC3996). G, J. C. cuauhmapan, sp. nov., G. holotype ♂ (CNAN T01396), J. paratype ♀ (CNAN T01399). H, K. C. berstoni, sp. nov., H. holotype ♂ (CASENT 9073325), K. paratype ♀ (CASENT 9073313). I, L. C. chanae, sp. nov., I. holotype ♂ (CNAN T01403), L. paratype ♀ (CNAN T01405). M, P. C. catemacoensis, sp. nov., M. holotype ♂ (CNAN T01424), P.paratype ♀ (CNAN T01423). N, Q. C. schmidti Sissom, 1995, N. ♂ (CASENT 9073316), Q. ♀ (CASENT 9073317). O, R. C. yucatanensis, sp. nov., O. holotype ♂ (CNAN T01416), R. paratype ♀ (CNAN T01418). Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5825954" httpUri="https://zenodo.org/record/5825954/files/figure.png" pageId="8" pageNumber="8">25H, K</figureCitation>
|
||
, 34, 35,
|
||
<tableCitation id="942503086D5AFF992324FB14FBD2FB07" box="[1049,1130,1245,1269]" captionStart="TABLE 1" captionStartId="5.[610,690,226,250]" captionTargetBox="[119,1126,358,746]" captionTargetPageId="5" captionText="TABLE 1 Species of the arboreal Neotropical scorpion “thorellii” clade of Centruroides Marx, 1890, bark scorpions, with countries, and departments or states in which recorded" httpUri="http://table.plazi.org/id/8DD8663B6D57FF94255FFF2BFC7DFEB6" pageId="8" pageNumber="8" tableUuid="8DD8663B6D57FF94255FFF2BFC7DFEB6">tables 1</tableCitation>
|
||
,
|
||
<tableCitation id="942503086D5AFF99234AFB15FB3DFB06" box="[1143,1157,1244,1268]" captionStart="TABLE 5" captionStartId="32.[610,690,226,250]" captionTargetBox="[119,1193,387,1697]" captionTargetPageId="32" captionText="TABLE 5 Meristic data for type material of Centruroides berstoni, sp. nov. Material deposited in the California Academy of Sciences (CASENT), San Francisco. Measurements follow Stahnke (1970), Lamoral (1979), and Prendini (2001b)." httpUri="http://table.plazi.org/id/8DD8663B6D72FFB1255FFF2BFC18FE90" pageId="8" pageNumber="8" tableUuid="8DD8663B6D72FFB1255FFF2BFC18FE90">5</tableCitation>
|
||
,
|
||
<tableCitation id="942503086D5AFF9923ACFB14FB13FB07" box="[1169,1195,1245,1269]" captionStart="TABLE 10" captionStartId="71.[603,683,226,250]" captionTargetBox="[119,1152,358,736]" captionTargetPageId="71" captionText="TABLE 10 Diagnostic ratios (mean/median/mode) for species of the arboreal Neotropical “thorelli” clade of Centruroides Marx, 1890, bark scorpions" httpUri="http://table.plazi.org/id/8DD8663B6D15FFD62566FF2BFCD9FEB6" pageId="8" pageNumber="8" tableUuid="8DD8663B6D15FFD62566FF2BFCD9FEB6">10</tableCitation>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="91BD65386D5AFF992591FADBFBF3FA9D" pageId="8" pageNumber="8" type="reference_group">
|
||
<paragraph id="D91836B36D5AFF992591FADBFBF3FA9D" blockId="8.[684,1202,1298,1391]" pageId="8" pageNumber="8">
|
||
<taxonomicName id="1EA74D306D5AFF992591FADBFBC3FAD8" authority=": Sissom, 1995: 94 - 96" authorityName=": Sissom" authorityPageNumber="94 - 96" authorityYear="1995" box="[684,1147,1298,1322]" class="Insecta" family="Cerambycidae" genus="Centruroides" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="8" phylum="Arthropoda" rank="species" species="schmidti">
|
||
<emphasis id="EBD3EAA16D5AFF992591FADBFC2AFAD8" box="[684,914,1298,1322]" italics="true" pageId="8" pageNumber="8">Centruroides schmidti</emphasis>
|
||
:
|
||
<bibRefCitation id="BD364B426D5AFF99249DFADBFBC3FAD8" author="Sissom, W. D." box="[928,1147,1298,1322]" journalOrPublisher="Journal of Arachnology" pageId="8" pageNumber="8" pagination="91 - 99" part="23" refId="ref40441" refString="Sissom, W. D. 1995. Redescription of the scorpion Centruroides thorelli Kraepelin (Buthidae) and description of two new species. Journal of Arachnology 23: 91 - 99." title="Redescription of the scorpion Centruroides thorelli Kraepelin (Buthidae) and description of two new species" type="journal article" year="1995">Sissom, 1995: 94–96</bibRefCitation>
|
||
</taxonomicName>
|
||
; figs. 10–18,
|
||
<tableCitation id="942503086D5AFF992417FAFCFCCAFABF" box="[810,882,1333,1357]" captionStart="TABLE 1" captionStartId="5.[610,690,226,250]" captionTargetBox="[119,1126,358,746]" captionTargetPageId="5" captionText="TABLE 1 Species of the arboreal Neotropical scorpion “thorellii” clade of Centruroides Marx, 1890, bark scorpions, with countries, and departments or states in which recorded" httpUri="http://table.plazi.org/id/8DD8663B6D57FF94255FFF2BFC7DFEB6" pageId="8" pageNumber="8" tableUuid="8DD8663B6D57FF94255FFF2BFC7DFEB6">table 1</tableCitation>
|
||
(misidentification:
|
||
<typeStatus id="061C88116D5AFF992379FAFCFB12FABF" box="[1092,1194,1333,1357]" pageId="8" pageNumber="8">paratypes</typeStatus>
|
||
from Escobas,
|
||
<collectingRegion id="1B63F8516D5AFF992446FA9EFC05FA9D" box="[891,957,1367,1391]" country="Guatemala" name="Izabal" pageId="8" pageNumber="8">Izabal</collectingRegion>
|
||
,
|
||
<collectingCountry id="A1B076236D5AFF9924FAFA9EFB87FA9D" box="[967,1087,1367,1391]" name="Guatemala" pageId="8" pageNumber="8">Guatemala</collectingCountry>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="91BD65386D5AFF9B25F0FA42FE25FB4A" lastPageId="10" lastPageNumber="10" pageId="8" pageNumber="8" type="materials_examined">
|
||
<paragraph id="D91836B36D5AFF9B25F0FA42FE25FB4A" blockId="8.[684,1212,1418,1740]" lastBlockId="10.[108,637,1150,1737]" lastPageId="10" lastPageNumber="10" pageId="8" pageNumber="8">
|
||
<collectionCode id="BFB6AE766D5AFF9925F0FA42FC30FA51" box="[717,904,1419,1443]" pageId="8" pageNumber="8">TYPE MATERIAL</collectionCode>
|
||
:
|
||
<materialsCitation id="69CF3CEE6D5AFF9924A6FA43FC26FA16" ID-GBIF-Occurrence="3427257344" collectingDate="2019-09-24" collectingDateMax="2019-09-25" collectingDateMin="2019-09-24" collectionCode="TYPE, MATERIAL" country="Guatemala" elevation="17" latitude="15.734806" longLatPrecision="1" longitude="-88.91589" municipality="Municipio Livingston" pageId="8" pageNumber="8" specimenCode="CASENT 9073325" specimenCount="1" specimenCount-female="4" specimenCount-male="1" stateProvince="Departamento Izabal" typeStatus="holotype">
|
||
<emphasis id="EBD3EAA16D5AFF9924A6FA43FBF2FA50" bold="true" box="[923,1098,1418,1442]" pageId="8" pageNumber="8">
|
||
<collectingCountry id="A1B076236D5AFF9924A6FA43FBFFFA50" box="[923,1095,1418,1442]" name="Guatemala" pageId="8" pageNumber="8">GUATEMALA</collectingCountry>
|
||
:
|
||
</emphasis>
|
||
<emphasis id="EBD3EAA16D5AFF992365FA42FC8FFA31" italics="true" pageId="8" pageNumber="8">
|
||
<collectingRegion id="1B63F8516D5AFF992365FA42FC8FFA31" country="Guatemala" name="Izabal" pageId="8" pageNumber="8">Departamento Izabal</collectingRegion>
|
||
</emphasis>
|
||
:
|
||
<collectingCounty id="30794E3F6D5AFF99247EFA62FB89FA31" box="[835,1073,1451,1475]" pageId="8" pageNumber="8">Município Livingston</collectingCounty>
|
||
:
|
||
<typeStatus id="061C88116D5AFF992307FA62FB27FA31" box="[1082,1183,1451,1475]" pageId="8" pageNumber="8">Holotype</typeStatus>
|
||
<specimenCount id="CFA1FD3A6D5AFF992398FA63FB04FA31" box="[1189,1212,1450,1475]" pageId="8" pageNumber="8" type="male">♂</specimenCount>
|
||
(
|
||
<specimenCode id="89019EC86D5AFF99258EFA04FC2BFA16" box="[691,915,1484,1509]" pageId="8" pageNumber="8">CASENT 9073325</specimenCode>
|
||
),
|
||
</materialsCitation>
|
||
<materialsCitation id="69CF3CEE6D5AFF992492FA04FCBEF978" ID-GBIF-Occurrence="3427257337" collectingDate="2019-09-24" collectingDateMax="2019-09-25" collectingDateMin="2019-09-24" collectionCode="TYPE, MATERIAL" collectorName="A. M. Goodman" country="Guatemala" elevation="17" latitude="15.734806" location="Hotel Tijax" longLatPrecision="1" longitude="-88.91589" municipality="Municipio Livingston" pageId="8" pageNumber="8" specimenCode="CASENT 9073298" specimenCount="1" specimenCount-female="4" specimenCount-male="1" stateProvince="Izabal" typeStatus="paratype">
|
||
<typeStatus id="061C88116D5AFF992492FA04FBA9FA17" box="[943,1041,1485,1509]" pageId="8" pageNumber="8">paratype</typeStatus>
|
||
<specimenCount id="CFA1FD3A6D5AFF99231FFA02FB81FA16" box="[1058,1081,1483,1508]" pageId="8" pageNumber="8" type="male">♂</specimenCount>
|
||
(
|
||
<specimenCode id="89019EC86D5AFF99236CFA04FCB4F9F4" pageId="8" pageNumber="8">CASENT 9073298</specimenCode>
|
||
),
|
||
<collectingMunicipality id="397CACC96D5AFF99241DFA27FC2AF9F4" box="[800,914,1518,1542]" pageId="8" pageNumber="8">Río Dulce</collectingMunicipality>
|
||
,
|
||
<location id="DC7860686D5AFF9924A2FA27FBA6F9F4" LSID="urn:lsid:plazi:treatment:510E87A56D5AFF9E2436FBA1FF74FB62:DC7860686D5AFF9924A2FA27FBA6F9F4" box="[927,1054,1518,1542]" country="Guatemala" county="Municipio Livingston" latitude="15.734806" longLatPrecision="1" longitude="-88.91589" municipality="Rio Dulce" name="Hotel Tijax" pageId="8" pageNumber="8" stateProvince="Izabal">Hotel Tijax</location>
|
||
,
|
||
<geoCoordinate id="BC9350746D5AFF992316FA24FB04F9F7" box="[1067,1212,1517,1542]" degrees="15" direction="north" minutes="39" orientation="latitude" pageId="8" pageNumber="8" precision="1" seconds="51.2" value="15.664222">15°39′51.2″N</geoCoordinate>
|
||
<geoCoordinate id="BC9350746D5AFF992591F9C6FCFAF9D4" box="[684,834,1550,1575]" degrees="89" direction="west" minutes="00" orientation="longitude" pageId="8" pageNumber="8" precision="1" seconds="14.6" value="-89.00406">89°00′14.6″W</geoCoordinate>
|
||
,
|
||
<quantity id="1E5F9B566D5AFF992471F9C6FC3CF9D4" box="[844,900,1551,1575]" metricMagnitude="1" metricUnit="m" metricValue="1.7" pageId="8" pageNumber="8" unit="m" value="17.0">
|
||
<elevation id="528AD1806D5AFF992471F9C6FC3CF9D4" box="[844,900,1551,1575]" metricMagnitude="1" metricUnit="m" metricValue="1.7" pageId="8" pageNumber="8" unit="m" value="17.0">17 m</elevation>
|
||
</quantity>
|
||
,
|
||
<date id="AD1910736D5AFF9924B3F9C6FC47F9D5" box="[910,1023,1551,1575]" pageId="8" pageNumber="8" value="2019-09-24">
|
||
<collectingDate id="BD5DE99B6D5AFF9924B3F9C6FC47F9D5" box="[910,1023,1551,1575]" pageId="8" pageNumber="8" value="2019-09-24">24.ix.2019</collectingDate>
|
||
</date>
|
||
,
|
||
<collectorName id="745253656D5AFF992334F9C6FB00F9D5" box="[1033,1208,1551,1575]" pageId="8" pageNumber="8">A.M. Goodman</collectorName>
|
||
, collected along gravel road of Hacienda Tijax Parking Lot, flanked by bamboo groves, and live fencing;
|
||
</materialsCitation>
|
||
<materialsCitation id="69CF3CEE6D5AFF99242BF9BBFCA1F959" ID-GBIF-Occurrence="3427257332" collectingDate="2019-09-24" collectingDateMax="2019-09-25" collectingDateMin="2019-09-24" collectionCode="TYPE, MATERIAL" country="Guatemala" county="Municipio Livingston" elevation="17" latitude="15.734806" location="Hotel Tijax" longLatPrecision="1" longitude="-88.91589" municipality="Rio Dulce" pageId="8" pageNumber="8" specimenCode="CASENT 9073312, 9073326" specimenCount="2" specimenCount-female="4" specimenCount-male="2" stateProvince="Izabal" typeStatus="paratype">
|
||
<specimenCount id="CFA1FD3A6D5AFF99242BF9BBFCF1F978" box="[790,841,1649,1674]" pageId="8" pageNumber="8" type="male">2 ♂</specimenCount>
|
||
<typeStatus id="061C88116D5AFF992464F9BBFC7DF978" box="[857,965,1650,1674]" pageId="8" pageNumber="8">paratypes</typeStatus>
|
||
(
|
||
<specimenCode id="89019EC86D5AFF9924E1F9BBFB00F978" box="[988,1208,1650,1674]" pageId="8" pageNumber="8">CASENT 9073312</specimenCode>
|
||
,
|
||
<specimenCode id="89019EC86D5AFF992591F95AFCB6F959" box="[684,782,1683,1707]" pageId="8" pageNumber="8">9073326</specimenCode>
|
||
),
|
||
</materialsCitation>
|
||
<materialsCitation id="69CF3CEE6D5AFF9B241BF95AFE25FB4A" ID-GBIF-Occurrence="3427257314" collectingDate="2019-09-25" collectingDateMax="2019-09-25" collectingDateMin="2019-09-24" collectionCode="TYPE, MATERIAL" collectorName="A. M. Goodman." country="Guatemala" elevation="15" lastPageId="10" lastPageNumber="10" latitude="15.734806" location="Chocon Machacas" longLatPrecision="1" longitude="-88.91589" municipality="Livingston" pageId="8" pageNumber="8" specimenCode="CASENT 9073297, 9073313, 9073324, 9073368" specimenCount="4" specimenCount-female="4" specimenCount-male="4" stateProvince="Izabal" typeStatus="paratype">
|
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<specimenCount id="CFA1FD3A6D5AFF99241BF95AFCEFF959" box="[806,855,1682,1707]" pageId="8" pageNumber="8" type="female">4 ♀</specimenCount>
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<typeStatus id="061C88116D5AFF99245EF95DFC76F95E" box="[867,974,1684,1708]" pageId="8" pageNumber="8">paratypes</typeStatus>
|
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(
|
||
<specimenCode id="89019EC86D5AFF9924DFF95AFB00F959" box="[994,1208,1683,1707]" pageId="8" pageNumber="8">CASENT 9073297</specimenCode>
|
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,
|
||
<specimenCode id="89019EC86D5AFF992591F97DFCB3F93E" box="[684,779,1716,1740]" pageId="8" pageNumber="8">9073313</specimenCode>
|
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,
|
||
<specimenCode id="89019EC86D5AFF992427F97DFCC1F93E" box="[794,889,1716,1740]" pageId="8" pageNumber="8">9073324</specimenCode>
|
||
,
|
||
<specimenCode id="89019EC86D5AFF9924B4F97DFC52F93E" box="[905,1002,1716,1740]" pageId="8" pageNumber="8">9073368</specimenCode>
|
||
), Biotopo
|
||
<location id="DC7860686D5AFF9B2359F97DFF64FB65" LSID="urn:lsid:plazi:treatment:510E87A56D5AFF9E2436FBA1FF74FB62:DC7860686D5AFF9B2359F97DFF64FB65" country="Guatemala" county="Municipio Livingston" lastPageId="10" lastPageNumber="10" latitude="15.734806" longLatPrecision="1" longitude="-88.91589" municipality="Rio Dulce" name="Chocon Machacas" pageId="8" pageNumber="8" stateProvince="Izabal">Chocón Machacas</location>
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||
,
|
||
<geoCoordinate id="BC9350746D58FF9B27D1FBB6FE39FB64" box="[236,385,1150,1175]" degrees="15" direction="north" minutes="44" orientation="latitude" pageId="10" pageNumber="10" precision="1" seconds="05.3" value="15.734806">15°44′05.3″N</geoCoordinate>
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<geoCoordinate id="BC9350746D58FF9B26B3FBB6FD9FFB64" box="[398,551,1150,1175]" degrees="88" direction="west" minutes="54" orientation="longitude" pageId="10" pageNumber="10" precision="1" seconds="57.2" value="-88.91589">88°54′57.2″W</geoCoordinate>
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,
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||
<quantity id="1E5F9B566D58FF9B2505FBB6FDCFFB64" box="[568,631,1151,1175]" metricMagnitude="1" metricUnit="m" metricValue="1.5" pageId="10" pageNumber="10" unit="m" value="15.0">
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<elevation id="528AD1806D58FF9B2505FBB6FDCFFB64" box="[568,631,1151,1175]" metricMagnitude="1" metricUnit="m" metricValue="1.5" pageId="10" pageNumber="10" unit="m" value="15.0">15 m</elevation>
|
||
</quantity>
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,
|
||
<date id="AD1910736D58FF9B2751FB69FF64FB4A" box="[108,220,1184,1208]" pageId="10" pageNumber="10" value="2019-09-25">
|
||
<collectingDate id="BD5DE99B6D58FF9B2751FB69FF64FB4A" box="[108,220,1184,1208]" pageId="10" pageNumber="10" value="2019-09-25">25.ix.2019</collectingDate>
|
||
</date>
|
||
,
|
||
<collectorName id="745253656D58FF9B27D5FB69FE25FB4A" box="[232,413,1184,1208]" pageId="10" pageNumber="10">A.M. Goodman.</collectorName>
|
||
</materialsCitation>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<caption id="8DD8663B6D5BFF982750F995FE49F915" ID-DOI="http://doi.org/10.5281/zenodo.5825898" ID-Zenodo-Dep="5825898" httpUri="https://zenodo.org/record/5825898/files/figure.png" pageId="9" pageNumber="9" startId="9.[109,195,1628,1649]" targetBox="[166,1154,224,1599]" targetPageId="9">
|
||
<paragraph id="D91836B36D5BFF982750F995FE49F915" blockId="9.[109,1212,1626,1767]" pageId="9" pageNumber="9">
|
||
FIGURE 1. Representative species of the arboreal Neotropical
|
||
<emphasis id="EBD3EAA16D5BFF9825C8F993FCEDF980" box="[757,853,1626,1650]" italics="true" pageId="9" pageNumber="9">
|
||
“
|
||
<taxonomicName id="1EA74D306D5BFF9825C1F993FCF6F980" baseAuthorityName="Kraepelin" baseAuthorityYear="1891" box="[764,846,1626,1650]" class="Insecta" family="Cerambycidae" genus="Centruroides" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="9" phylum="Arthropoda" rank="species" species="thorellii">thorellii</taxonomicName>
|
||
”
|
||
</emphasis>
|
||
clade of
|
||
<taxonomicName id="1EA74D306D5BFF98248AF993FB00F980" authority="Marx, 1890" authorityName="MARX" authorityYear="1890" box="[951,1208,1626,1650]" class="Insecta" family="Cerambycidae" genus="Centruroides" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="9" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="EBD3EAA16D5BFF98248AF993FB8FF980" box="[951,1079,1626,1650]" italics="true" pageId="9" pageNumber="9">Centruroides</emphasis>
|
||
<bibRefCitation id="BD364B426D5BFF98237DF993FB00F980" author="Marx" box="[1088,1208,1626,1650]" firstAuthor="Marx" pageId="9" pageNumber="9" pagination="207 - 211" refId="ref39358" refString="Marx, G. 1890 (" 1889 "). Arachnida. In L. O. Howard (editor), Scientific results of the explorations by the U. S. Fish Commission Steamer Albatross, No. V. Annotated catalogue of the insects collected in 1887 - ' 88. Proceedings of the United States National Museum 12: 207 - 211." type="journal article" year="1890">Marx, 1890</bibRefCitation>
|
||
</taxonomicName>
|
||
, bark scorpions (
|
||
<taxonomicName id="1EA74D306D5BFF982632F9BEFED4F97D" box="[271,364,1655,1679]" class="Arachnida" family="Buthidae" kingdom="Animalia" order="Scorpiones" pageId="9" pageNumber="9" phylum="Arthropoda" rank="family">Buthidae</taxonomicName>
|
||
C.L. Koch, 1837), habitus in life.
|
||
<emphasis id="EBD3EAA16D5BFF9825FDF9BEFD4EF97D" bold="true" box="[704,758,1655,1679]" pageId="9" pageNumber="9">A, C.</emphasis>
|
||
<taxonomicName id="1EA74D306D5BFF9825C1F9BEFC40F97D" authority="Sissom, 1995" authorityName=": Sissom" authorityYear="1995" box="[764,1016,1655,1679]" class="Insecta" family="Cerambycidae" genus="Centruroides" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="9" phylum="Arthropoda" rank="species" species="schmidti">
|
||
<emphasis id="EBD3EAA16D5BFF9825C1F9BEFCD4F97D" box="[764,876,1655,1679]" italics="true" pageId="9" pageNumber="9">C. schmidti</emphasis>
|
||
<bibRefCitation id="BD364B426D5BFF98244FF9BEFC40F97D" author="Sissom, W. D." box="[882,1016,1655,1679]" journalOrPublisher="Journal of Arachnology" pageId="9" pageNumber="9" pagination="91 - 99" part="23" refId="ref40441" refString="Sissom, W. D. 1995. Redescription of the scorpion Centruroides thorelli Kraepelin (Buthidae) and description of two new species. Journal of Arachnology 23: 91 - 99." title="Redescription of the scorpion Centruroides thorelli Kraepelin (Buthidae) and description of two new species" type="journal article" year="1995">Sissom, 1995</bibRefCitation>
|
||
</taxonomicName>
|
||
,
|
||
<emphasis id="EBD3EAA16D5BFF98233FF9BEFBA2F97D" bold="true" box="[1026,1050,1655,1679]" pageId="9" pageNumber="9">A.</emphasis>
|
||
♀, El Progreso, Finca El Olvido, Las Minas, Guatemala,
|
||
<emphasis id="EBD3EAA16D5BFF98252CF95DFD91F95E" bold="true" box="[529,553,1684,1708]" pageId="9" pageNumber="9">C.</emphasis>
|
||
♂, Zacapa, Guatemala.
|
||
<emphasis id="EBD3EAA16D5BFF98241AF95DFCE6F95E" bold="true" box="[807,862,1684,1708]" pageId="9" pageNumber="9">B, D.</emphasis>
|
||
<taxonomicName id="1EA74D306D5BFF98245AF95CFC6CF95E" authority="Goodman & Prendini & Francke & Esposito, 2021" authorityName="Goodman & Prendini & Francke & Esposito" authorityYear="2021" box="[871,980,1684,1708]" class="Insecta" family="Cerambycidae" genus="Centruroides" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="9" phylum="Arthropoda" rank="species" species="berstoni" status="sp. nov.">
|
||
<emphasis id="EBD3EAA16D5BFF98245AF95CFC6CF95E" box="[871,980,1684,1708]" italics="true" pageId="9" pageNumber="9">C. berstoni</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicNameLabel id="F0E057DA6D5BFF9824DFF95CFB8AF95E" box="[994,1074,1685,1708]" pageId="9" pageNumber="9" rank="species">sp. nov.</taxonomicNameLabel>
|
||
,
|
||
<emphasis id="EBD3EAA16D5BFF982303F95DFBECF95E" bold="true" box="[1086,1108,1684,1708]" pageId="9" pageNumber="9">B.</emphasis>
|
||
♀,
|
||
<emphasis id="EBD3EAA16D5BFF982342F95DFB20F95E" bold="true" box="[1151,1176,1684,1708]" pageId="9" pageNumber="9">D.</emphasis>
|
||
♂, Izabal, Rio Dulce, Hotel Tijax, Guatemala. Note elongated metasoma in males. Both species were collected in vegetation, 3–4 m above ground level.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="8DD8663B6D58FF9B2751FC1BFD05FBAC" ID-DOI="http://doi.org/10.5281/zenodo.5825902" ID-Zenodo-Dep="5825902" httpUri="https://zenodo.org/record/5825902/files/figure.png" pageId="10" pageNumber="10" startId="10.[108,194,978,999]" targetBox="[109,1184,232,938]" targetPageId="10">
|
||
<paragraph id="D91836B36D58FF9B2751FC1BFD05FBAC" blockId="10.[108,1212,976,1118]" pageId="10" pageNumber="10">
|
||
FIGURE 2. Phylogeny of the arboreal Neotropical
|
||
<emphasis id="EBD3EAA16D58FF9B2555FC19FD7FFC1A" box="[616,711,976,1000]" italics="true" pageId="10" pageNumber="10">
|
||
“
|
||
<taxonomicName id="1EA74D306D58FF9B2553FC19FD78FC1A" baseAuthorityName="Kraepelin" baseAuthorityYear="1891" box="[622,704,976,1000]" class="Insecta" family="Cerambycidae" genus="Centruroides" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="10" phylum="Arthropoda" rank="species" species="thorellii">thorellii</taxonomicName>
|
||
”
|
||
</emphasis>
|
||
clade of
|
||
<taxonomicName id="1EA74D306D58FF9B241CFC19FBA5FC1A" authority="Marx, 1890" authorityName="MARX" authorityYear="1890" box="[801,1053,976,1000]" class="Insecta" family="Cerambycidae" genus="Centruroides" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="10" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="EBD3EAA16D58FF9B241CFC19FC19FC1A" box="[801,929,976,1000]" italics="true" pageId="10" pageNumber="10">Centruroides</emphasis>
|
||
<bibRefCitation id="BD364B426D58FF9B249AFC18FBA5FC1A" author="Marx" box="[935,1053,976,1000]" firstAuthor="Marx" pageId="10" pageNumber="10" pagination="207 - 211" refId="ref39358" refString="Marx, G. 1890 (" 1889 "). Arachnida. In L. O. Howard (editor), Scientific results of the explorations by the U. S. Fish Commission Steamer Albatross, No. V. Annotated catalogue of the insects collected in 1887 - ' 88. Proceedings of the United States National Museum 12: 207 - 211." type="journal article" year="1890">Marx, 1890</bibRefCitation>
|
||
</taxonomicName>
|
||
, bark scorpions (
|
||
<taxonomicName id="1EA74D306D58FF9B274EFC27FF68FBF4" box="[115,208,1006,1030]" class="Arachnida" family="Buthidae" kingdom="Animalia" order="Scorpiones" pageId="10" pageNumber="10" phylum="Arthropoda" rank="family">Buthidae</taxonomicName>
|
||
C.L. Koch, 1837), obtained by simultaneous phylogenetic analysis of 112 morphological characters and DNA sequences of the mitochondrial cytochrome oxidase
|
||
<emphasis id="EBD3EAA16D58FF9B242DFBC5FCA2FBD1" box="[784,794,1036,1059]" italics="true" pageId="10" pageNumber="10">c</emphasis>
|
||
subunit I gene. Posterior probabilities, obtained with Bayesian inference, and nonparametric bootstraps, obtained with maximum likelihood, indicated above and below branches for corresponding nodes.
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection id="91BD65386D58FF9B27B1FB08FDBAFAAF" pageId="10" pageNumber="10" type="etymology">
|
||
<paragraph id="D91836B36D58FF9B27B1FB08FDBAFAAF" blockId="10.[108,637,1150,1737]" pageId="10" pageNumber="10">
|
||
ETYMOLOGY: The species name is a patronym honoring the late Hyman Maxwell Berston, the first author’s grandfather (1930–2021), who inspired his interest in natural history during childhood visits to the
|
||
<collectingRegion id="1B63F8516D58FF9B2792FA8CFEA0FAAF" box="[175,280,1349,1373]" country="United States of America" name="California" pageId="10" pageNumber="10">California</collectingRegion>
|
||
Academy of Sciences.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="91BD65386D58FF9A27B1FAAFFF77FB84" lastPageId="11" lastPageNumber="11" pageId="10" pageNumber="10" type="diagnosis">
|
||
<paragraph id="D91836B36D58FF9A27B1FAAFFEB7FE6D" blockId="10.[108,637,1150,1737]" lastBlockId="11.[108,638,226,1738]" lastPageId="11" lastPageNumber="11" pageId="10" pageNumber="10">
|
||
DIAGNOSIS:
|
||
<taxonomicName id="1EA74D306D58FF9B2637FAAFFE5BFA8C" authorityName="Goodman & Prendini & Francke & Esposito" authorityYear="2021" box="[266,483,1382,1406]" class="Insecta" family="Cerambycidae" genus="Centruroides" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="10" phylum="Arthropoda" rank="species" species="berstoni">
|
||
<emphasis id="EBD3EAA16D58FF9B2637FAAFFE5BFA8C" box="[266,483,1382,1406]" italics="true" pageId="10" pageNumber="10">Centruroides berstoni</emphasis>
|
||
</taxonomicName>
|
||
is most closely related to
|
||
<taxonomicName id="1EA74D306D58FF9B27E7FA41FED6FA6D" authorityName="Goodman & Prendini & Francke & Esposito" authorityYear="2021" box="[218,366,1415,1439]" class="Insecta" family="Cerambycidae" genus="Centruroides" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="10" phylum="Arthropoda" rank="species" species="hamadryas">
|
||
<emphasis id="EBD3EAA16D58FF9B27E7FA41FED6FA6D" box="[218,366,1415,1439]" italics="true" pageId="10" pageNumber="10">C. hamadryas</emphasis>
|
||
</taxonomicName>
|
||
, from which it differs as follows. The carapace is sparsely granular, more densely so on the interocular triangle, in the female of
|
||
<taxonomicName id="1EA74D306D58FF9B27B6FA22FF45F9F0" authorityName="Goodman & Prendini & Francke & Esposito" authorityYear="2021" box="[139,253,1514,1538]" class="Insecta" family="Cerambycidae" genus="Centruroides" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="10" phylum="Arthropoda" rank="species" species="berstoni">
|
||
<emphasis id="EBD3EAA16D58FF9B27B6FA22FF45F9F0" box="[139,253,1514,1538]" italics="true" pageId="10" pageNumber="10">C. berstoni</emphasis>
|
||
</taxonomicName>
|
||
but densely granular, with distinct lateral ocular carinae, in the female of
|
||
<taxonomicName id="1EA74D306D58FF9B252BF9C5FF1CF9B7" authorityName="Goodman & Prendini & Francke & Esposito" authorityYear="2021" class="Insecta" family="Cerambycidae" genus="Centruroides" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="10" phylum="Arthropoda" rank="species" species="hamadryas">
|
||
<emphasis id="EBD3EAA16D58FF9B252BF9C5FF1CF9B7" italics="true" pageId="10" pageNumber="10">C. hamadryas</emphasis>
|
||
</taxonomicName>
|
||
(fig. 6B, D). The pedipalp chela manus of the male is proportionally less incrassate in
|
||
<taxonomicName id="1EA74D306D58FF9B2530F987FDC4F994" authorityName="Goodman & Prendini & Francke & Esposito" authorityYear="2021" box="[525,636,1614,1638]" class="Insecta" family="Cerambycidae" genus="Centruroides" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="10" phylum="Arthropoda" rank="species" species="berstoni">
|
||
<emphasis id="EBD3EAA16D58FF9B2530F987FDC4F994" box="[525,636,1614,1638]" italics="true" pageId="10" pageNumber="10">C. berstoni</emphasis>
|
||
</taxonomicName>
|
||
(figs. 13–15B) than
|
||
<taxonomicName id="1EA74D306D58FF9B2676F9A6FE5AF975" authorityName="Goodman & Prendini & Francke & Esposito" authorityYear="2021" box="[331,482,1647,1671]" class="Insecta" family="Cerambycidae" genus="Centruroides" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="10" phylum="Arthropoda" rank="species" species="hamadryas">
|
||
<emphasis id="EBD3EAA16D58FF9B2676F9A6FE5AF975" box="[331,482,1647,1671]" italics="true" pageId="10" pageNumber="10">C. hamadryas</emphasis>
|
||
</taxonomicName>
|
||
(fig. 13–15A,
|
||
<tableCitation id="942503086D58FF9B2751F959FF7BF95A" box="[108,195,1680,1704]" captionStart="TABLE 4" captionStartId="26.[610,690,226,250]" captionTargetBox="[118,1191,387,1697]" captionTargetPageId="26" captionText="TABLE 4 Meristic data for type material of Centruroides hamadryas, sp. nov. Material deposited in the Colección Nacional de Arácnidos (CNAN), Universidad Nacional Autónoma de México, Mexico City. Measurements follow Stahnke (1970), Lamoral (1979), and Prendini (2001b)." httpUri="http://table.plazi.org/id/8DD8663B6D48FF8B255FFF2BFBCCFE90" pageId="10" pageNumber="10" tableUuid="8DD8663B6D48FF8B255FFF2BFBCCFE90">tables 4</tableCitation>
|
||
,
|
||
<tableCitation id="942503086D58FF9B27ECF959FF58F95A" box="[209,224,1680,1704]" captionStart="TABLE 5" captionStartId="32.[610,690,226,250]" captionTargetBox="[119,1193,387,1697]" captionTargetPageId="32" captionText="TABLE 5 Meristic data for type material of Centruroides berstoni, sp. nov. Material deposited in the California Academy of Sciences (CASENT), San Francisco. Measurements follow Stahnke (1970), Lamoral (1979), and Prendini (2001b)." httpUri="http://table.plazi.org/id/8DD8663B6D72FFB1255FFF2BFC18FE90" pageId="10" pageNumber="10" tableUuid="8DD8663B6D72FFB1255FFF2BFC18FE90">5</tableCitation>
|
||
). The retrodorsal carina of the chela manus is weakly granular and the dorsomedian carina absent in the male of
|
||
<taxonomicName id="1EA74D306D58FF9B24E1FBB7FBF4FB64" authorityName="Goodman & Prendini & Francke & Esposito" authorityYear="2021" box="[988,1100,1150,1174]" class="Insecta" family="Cerambycidae" genus="Centruroides" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="10" phylum="Arthropoda" rank="species" species="berstoni">
|
||
<emphasis id="EBD3EAA16D58FF9B24E1FBB7FBF4FB64" box="[988,1100,1150,1174]" italics="true" pageId="10" pageNumber="10">C. berstoni</emphasis>
|
||
</taxonomicName>
|
||
(fig. 13B), whereas the retrodorsal carina is complete and the dorsomedian carina weakly developed and restricted to the distal half in the male of
|
||
<taxonomicName id="1EA74D306D58FF9B2366FB2BFD5EFAE8" authorityName="Goodman & Prendini & Francke & Esposito" authorityYear="2021" class="Insecta" family="Cerambycidae" genus="Centruroides" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="10" phylum="Arthropoda" rank="species" species="hamadryas">
|
||
<emphasis id="EBD3EAA16D58FF9B2366FB2BFD5EFAE8" italics="true" pageId="10" pageNumber="10">C. hamadryas</emphasis>
|
||
</taxonomicName>
|
||
(fig. 13A). The retrodorsal carina of the manus is finely granular and the prodorsal carina absent in the female of
|
||
<taxonomicName id="1EA74D306D58FF9B24FBFA8CFB87FAAE" authorityName="Goodman & Prendini & Francke & Esposito" authorityYear="2021" box="[966,1087,1348,1372]" class="Insecta" family="Cerambycidae" genus="Centruroides" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="10" phylum="Arthropoda" rank="species" species="berstoni">
|
||
<emphasis id="EBD3EAA16D58FF9B24FBFA8CFB87FAAE" box="[966,1087,1348,1372]" italics="true" pageId="10" pageNumber="10">C. berstoni</emphasis>
|
||
</taxonomicName>
|
||
(fig. 14B), whereas the retrodorsal carina is complete and the prodorsal carina restricted to the distal third in the female of
|
||
<taxonomicName id="1EA74D306D58FF9B2428FA61FC1FFA4D" authorityName="Goodman & Prendini & Francke & Esposito" authorityYear="2021" box="[789,935,1447,1471]" class="Insecta" family="Cerambycidae" genus="Centruroides" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="10" phylum="Arthropoda" rank="species" species="hamadryas">
|
||
<emphasis id="EBD3EAA16D58FF9B2428FA61FC1FFA4D" box="[789,935,1447,1471]" italics="true" pageId="10" pageNumber="10">C. hamadryas</emphasis>
|
||
</taxonomicName>
|
||
(fig. 14A). The first leg of the male is greater than 2× the length of the carapace in
|
||
<taxonomicName id="1EA74D306D58FF9B25C3FA23FCD5F9F0" authorityName="Goodman & Prendini & Francke & Esposito" authorityYear="2021" box="[766,877,1514,1538]" class="Insecta" family="Cerambycidae" genus="Centruroides" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="10" phylum="Arthropoda" rank="species" species="berstoni">
|
||
<emphasis id="EBD3EAA16D58FF9B25C3FA23FCD5F9F0" box="[766,877,1514,1538]" italics="true" pageId="10" pageNumber="10">C. berstoni</emphasis>
|
||
</taxonomicName>
|
||
but less than 2× its length in
|
||
<taxonomicName id="1EA74D306D58FF9B2398FA23FCA7F9D1" authorityName="Goodman & Prendini & Francke & Esposito" authorityYear="2021" class="Insecta" family="Cerambycidae" genus="Centruroides" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="10" phylum="Arthropoda" rank="species" species="hamadryas">
|
||
<emphasis id="EBD3EAA16D58FF9B2398FA23FCA7F9D1" italics="true" pageId="10" pageNumber="10">C. hamadryas</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<tableCitation id="942503086D58FF9B2410F9C2FC3BF9D1" box="[813,899,1547,1571]" captionStart="TABLE 10" captionStartId="71.[603,683,226,250]" captionTargetBox="[119,1152,358,736]" captionTargetPageId="71" captionText="TABLE 10 Diagnostic ratios (mean/median/mode) for species of the arboreal Neotropical “thorelli” clade of Centruroides Marx, 1890, bark scorpions" httpUri="http://table.plazi.org/id/8DD8663B6D15FFD62566FF2BFCD9FEB6" pageId="10" pageNumber="10" tableUuid="8DD8663B6D15FFD62566FF2BFCD9FEB6">table 10</tableCitation>
|
||
). The dorsomedian carinae of the mesosomal tergites are weakly developed and restricted to the posterior half in
|
||
<taxonomicName id="1EA74D306D58FF9B2320F984FB37F997" authorityName="Goodman & Prendini & Francke & Esposito" authorityYear="2021" box="[1053,1167,1613,1637]" class="Insecta" family="Cerambycidae" genus="Centruroides" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="10" phylum="Arthropoda" rank="species" species="berstoni">
|
||
<emphasis id="EBD3EAA16D58FF9B2320F984FB37F997" box="[1053,1167,1613,1637]" italics="true" pageId="10" pageNumber="10">C. berstoni</emphasis>
|
||
</taxonomicName>
|
||
but distinct and complete in
|
||
<taxonomicName id="1EA74D306D58FF9B2497F9A7FB80F974" authorityName="Goodman & Prendini & Francke & Esposito" authorityYear="2021" box="[938,1080,1646,1670]" class="Insecta" family="Cerambycidae" genus="Centruroides" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="10" phylum="Arthropoda" rank="species" species="hamadryas">
|
||
<emphasis id="EBD3EAA16D58FF9B2497F9A7FB80F974" box="[938,1080,1646,1670]" italics="true" pageId="10" pageNumber="10">C. hamadryas</emphasis>
|
||
</taxonomicName>
|
||
. The ventrolateral carinae of mesosomal sternite VII are weakly developed and the ventrosubmedian carinae absent in
|
||
<taxonomicName id="1EA74D306D59FF9A27BBFF2BFF4AFF08" authorityName="Goodman & Prendini & Francke & Esposito" authorityYear="2021" box="[134,242,226,250]" class="Insecta" family="Cerambycidae" genus="Centruroides" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="11" phylum="Arthropoda" rank="species" species="berstoni">
|
||
<emphasis id="EBD3EAA16D59FF9A27BBFF2BFF4AFF08" box="[134,242,226,250]" italics="true" pageId="11" pageNumber="11">C. berstoni</emphasis>
|
||
</taxonomicName>
|
||
, whereas the ventrolateral and ventrosubmedian carinae are weakly developed to absent in
|
||
<taxonomicName id="1EA74D306D59FF9A27B3FEEDFE9EFECE" authorityName="Goodman & Prendini & Francke & Esposito" authorityYear="2021" box="[142,294,292,316]" class="Insecta" family="Cerambycidae" genus="Centruroides" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="11" phylum="Arthropoda" rank="species" species="hamadryas">
|
||
<emphasis id="EBD3EAA16D59FF9A27B3FEEDFE9EFECE" box="[142,294,292,316]" italics="true" pageId="11" pageNumber="11">C. hamadryas</emphasis>
|
||
</taxonomicName>
|
||
. The telson vesicle is sparsely setose in
|
||
<taxonomicName id="1EA74D306D59FF9A27F6FE8CFE81FEAF" authorityName="Goodman & Prendini & Francke & Esposito" authorityYear="2021" box="[203,313,325,349]" class="Insecta" family="Cerambycidae" genus="Centruroides" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="11" phylum="Arthropoda" rank="species" species="berstoni">
|
||
<emphasis id="EBD3EAA16D59FF9A27F6FE8CFE81FEAF" box="[203,313,325,349]" italics="true" pageId="11" pageNumber="11">C. berstoni</emphasis>
|
||
</taxonomicName>
|
||
(figs. 23H, K, 24H, K, 25H, K), but densely setose in
|
||
<taxonomicName id="1EA74D306D59FF9A266BFEAFFE52FE8C" authorityName="Goodman & Prendini & Francke & Esposito" authorityYear="2021" box="[342,490,358,382]" class="Insecta" family="Cerambycidae" genus="Centruroides" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="11" phylum="Arthropoda" rank="species" species="hamadryas">
|
||
<emphasis id="EBD3EAA16D59FF9A266BFEAFFE52FE8C" box="[342,490,358,382]" italics="true" pageId="11" pageNumber="11">C. hamadryas</emphasis>
|
||
</taxonomicName>
|
||
(figs. 23B, E, 24B, E, 25B, E).
|
||
</paragraph>
|
||
<paragraph id="D91836B36D59FF9A27B1FE61FF77FB84" blockId="11.[108,638,226,1738]" pageId="11" pageNumber="11">
|
||
Additional differences between
|
||
<taxonomicName id="1EA74D306D59FF9A26E1FE61FDF4FE32" authorityName="Goodman & Prendini & Francke & Esposito" authorityYear="2021" box="[476,588,424,448]" class="Insecta" family="Cerambycidae" genus="Centruroides" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="11" phylum="Arthropoda" rank="species" species="berstoni">
|
||
<emphasis id="EBD3EAA16D59FF9A26E1FE61FDF4FE32" box="[476,588,424,448]" italics="true" pageId="11" pageNumber="11">C. berstoni</emphasis>
|
||
</taxonomicName>
|
||
and other species of the clade are as follows. The ventrosubmedian and ventrolateral carinae of metasomal segments I–IV in the male are weakly developed, smooth in
|
||
<taxonomicName id="1EA74D306D59FF9A27E6FDE4FEFFFDB6" authorityName="Goodman & Prendini & Francke & Esposito" authorityYear="2021" box="[219,327,556,580]" class="Insecta" family="Cerambycidae" genus="Centruroides" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="11" phylum="Arthropoda" rank="species" species="berstoni">
|
||
<emphasis id="EBD3EAA16D59FF9A27E6FDE4FEFFFDB6" box="[219,327,556,580]" italics="true" pageId="11" pageNumber="11">C. berstoni</emphasis>
|
||
</taxonomicName>
|
||
(figs. 18H, 19H) but absent in
|
||
<taxonomicName id="1EA74D306D59FF9A2751FD87FEA2FD97" authorityName="Goodman & Prendini & Francke & Esposito" authorityYear="2021" box="[108,282,590,613]" class="Insecta" family="Cerambycidae" genus="Centruroides" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="11" phylum="Arthropoda" rank="species" species="catemacoensis">
|
||
<emphasis id="EBD3EAA16D59FF9A2751FD87FEA2FD97" box="[108,282,590,613]" italics="true" pageId="11" pageNumber="11">C. catemacoensis</emphasis>
|
||
</taxonomicName>
|
||
(figs. 18, 19M), well developed in
|
||
<taxonomicName id="1EA74D306D59FF9A2751FDA6FEAAFD74" authorityName="Goodman & Prendini & Francke & Esposito" authorityYear="2021" box="[108,274,622,646]" class="Insecta" family="Cerambycidae" genus="Centruroides" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="11" phylum="Arthropoda" rank="species" species="cuauhmapan">
|
||
<emphasis id="EBD3EAA16D59FF9A2751FDA6FEAAFD74" box="[108,274,622,646]" italics="true" pageId="11" pageNumber="11">C. cuauhmapan</emphasis>
|
||
</taxonomicName>
|
||
(figs. 18, 19G), weakly developed on segments I–III, absent on IV in
|
||
<taxonomicName id="1EA74D306D59FF9A26D7FD59FDC5FD55" authorityName="Goodman & Prendini & Francke & Esposito" authorityYear="2021" box="[490,637,655,679]" class="Insecta" family="Cerambycidae" genus="Centruroides" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="11" phylum="Arthropoda" rank="species" species="hamadryas">
|
||
<emphasis id="EBD3EAA16D59FF9A26D7FD59FDC5FD55" box="[490,637,655,679]" italics="true" pageId="11" pageNumber="11">C. hamadryas</emphasis>
|
||
</taxonomicName>
|
||
(figs. 18–19B, 21–22B), and vestigial, smooth in
|
||
<taxonomicName id="1EA74D306D59FF9A2558FD78FF18FD18" authorityName="Sissom" authorityYear="1995" class="Insecta" family="Cerambycidae" genus="Centruroides" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="11" phylum="Arthropoda" rank="species" species="rileyi">
|
||
<emphasis id="EBD3EAA16D59FF9A2558FD78FF18FD18" italics="true" pageId="11" pageNumber="11">C. rileyi</emphasis>
|
||
</taxonomicName>
|
||
(figs. 18, 19A, 21–22A). The metasomal carinae of the female are finely granular in
|
||
<taxonomicName id="1EA74D306D59FF9A2530FD3AFDC4FCF9" authorityName="Goodman & Prendini & Francke & Esposito" authorityYear="2021" box="[525,636,755,779]" class="Insecta" family="Cerambycidae" genus="Centruroides" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="11" phylum="Arthropoda" rank="species" species="berstoni">
|
||
<emphasis id="EBD3EAA16D59FF9A2530FD3AFDC4FCF9" box="[525,636,755,779]" italics="true" pageId="11" pageNumber="11">C. berstoni</emphasis>
|
||
</taxonomicName>
|
||
(figs. 17–22K) but well developed, granular in
|
||
<taxonomicName id="1EA74D306D59FF9A2558FCDDFF45FCBF" authorityName="Goodman & Prendini & Francke & Esposito" authorityYear="2021" class="Insecta" family="Cerambycidae" genus="Centruroides" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="11" phylum="Arthropoda" rank="species" species="catemacoensis">
|
||
<emphasis id="EBD3EAA16D59FF9A2558FCDDFF45FCBF" italics="true" pageId="11" pageNumber="11">C. catemacoensis</emphasis>
|
||
</taxonomicName>
|
||
(figs. 17–22P). The telson vesicle is short and not posteriorly bilobed in the male of
|
||
<taxonomicName id="1EA74D306D59FF9A2558FC9FFF06FC7D" authorityName="Goodman & Prendini & Francke & Esposito" authorityYear="2021" class="Insecta" family="Cerambycidae" genus="Centruroides" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="11" phylum="Arthropoda" rank="species" species="berstoni">
|
||
<emphasis id="EBD3EAA16D59FF9A2558FC9FFF06FC7D" italics="true" pageId="11" pageNumber="11">C. berstoni</emphasis>
|
||
</taxonomicName>
|
||
(figs. 23–25H), unlike in
|
||
<taxonomicName id="1EA74D306D59FF9A26E8FCBEFDC4FC7D" authorityName="Goodman & Prendini & Francke & Esposito" authorityYear="2021" box="[469,636,887,911]" class="Insecta" family="Cerambycidae" genus="Centruroides" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="11" phylum="Arthropoda" rank="species" species="cuauhmapan">
|
||
<emphasis id="EBD3EAA16D59FF9A26E8FCBEFDC4FC7D" box="[469,636,887,911]" italics="true" pageId="11" pageNumber="11">C. cuauhmapan</emphasis>
|
||
</taxonomicName>
|
||
(figs. 23–25G) and
|
||
<taxonomicName id="1EA74D306D59FF9A260FFC51FE3AFC42" authorityName="Sissom" authorityYear="1995" box="[306,386,920,944]" class="Insecta" family="Cerambycidae" genus="Centruroides" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="11" phylum="Arthropoda" rank="species" species="rileyi">
|
||
<emphasis id="EBD3EAA16D59FF9A260FFC51FE3AFC42" box="[306,386,920,944]" italics="true" pageId="11" pageNumber="11">C. rileyi</emphasis>
|
||
</taxonomicName>
|
||
(figs 23–25A), and with surfaces sparsely granular in the female of
|
||
<taxonomicName id="1EA74D306D59FF9A2509FC73FF26FC00" authorityName="Goodman & Prendini & Francke & Esposito" authorityYear="2021" class="Insecta" family="Cerambycidae" genus="Centruroides" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="11" phylum="Arthropoda" rank="species" species="berstoni">
|
||
<emphasis id="EBD3EAA16D59FF9A2509FC73FF26FC00" italics="true" pageId="11" pageNumber="11">C. berstoni</emphasis>
|
||
</taxonomicName>
|
||
(figs. 23–25K), but smooth in the female of
|
||
<taxonomicName id="1EA74D306D59FF9A2751FC35FF46FBE1" authorityName="Goodman & Prendini & Francke & Esposito" authorityYear="2021" box="[108,254,1019,1043]" class="Insecta" family="Cerambycidae" genus="Centruroides" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="11" phylum="Arthropoda" rank="species" species="hamadryas">
|
||
<emphasis id="EBD3EAA16D59FF9A2751FC35FF46FBE1" box="[108,254,1019,1043]" italics="true" pageId="11" pageNumber="11">C. hamadryas</emphasis>
|
||
</taxonomicName>
|
||
(figs. 23–25E) and densely granular in the female of
|
||
<taxonomicName id="1EA74D306D59FF9A2618FBD4FE6FFBC6" authorityName="Goodman & Prendini & Francke & Esposito" authorityYear="2021" box="[293,471,1053,1076]" class="Insecta" family="Cerambycidae" genus="Centruroides" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="11" phylum="Arthropoda" rank="species" species="catemacoensis">
|
||
<emphasis id="EBD3EAA16D59FF9A2618FBD4FE6FFBC6" box="[293,471,1053,1076]" italics="true" pageId="11" pageNumber="11">C. catemacoensis</emphasis>
|
||
</taxonomicName>
|
||
(figs. 23–25P),
|
||
<taxonomicName id="1EA74D306D59FF9A2751FBF7FEADFBA7" authorityName="Goodman & Prendini & Francke & Esposito" authorityYear="2021" box="[108,277,1085,1109]" class="Insecta" family="Cerambycidae" genus="Centruroides" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="11" phylum="Arthropoda" rank="species" species="cuauhmapan">
|
||
<emphasis id="EBD3EAA16D59FF9A2751FBF7FEADFBA7" box="[108,277,1085,1109]" italics="true" pageId="11" pageNumber="11">C. cuauhmapan</emphasis>
|
||
</taxonomicName>
|
||
(figs. 23–25J), and
|
||
<taxonomicName id="1EA74D306D59FF9A26D6FBF7FDF8FBA7" authorityName="Sissom" authorityYear="1995" box="[491,576,1085,1109]" class="Insecta" family="Cerambycidae" genus="Centruroides" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="11" phylum="Arthropoda" rank="species" species="rileyi">
|
||
<emphasis id="EBD3EAA16D59FF9A26D6FBF7FDF8FBA7" box="[491,576,1085,1109]" italics="true" pageId="11" pageNumber="11">C. rileyi</emphasis>
|
||
</taxonomicName>
|
||
(figs. 23–25D).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="91BD65386D59FF9D27B1FB49FC59F9D4" lastPageId="12" lastPageNumber="12" pageId="11" pageNumber="11" type="description">
|
||
<paragraph id="D91836B36D59FF9A27B1FB49FF6CFB09" blockId="11.[108,638,226,1738]" pageId="11" pageNumber="11">
|
||
DESCRIPTION: The following description is based on the
|
||
<typeStatus id="061C88116D59FF9A262DFB68FECAFB4B" box="[272,370,1185,1209]" pageId="11" pageNumber="11">holotype</typeStatus>
|
||
male, with differences among other material noted in the section on variation.
|
||
</paragraph>
|
||
<paragraph id="D91836B36D59FF9A27B1FACDFBCCFECE" blockId="11.[108,638,226,1738]" lastBlockId="11.[684,1213,226,1738]" pageId="11" pageNumber="11">
|
||
<emphasis id="EBD3EAA16D59FF9A27B1FACDFEB8FAEE" box="[140,256,1284,1308]" italics="true" pageId="11" pageNumber="11">Coloration:</emphasis>
|
||
Base color pale yellow, with extensive infuscation, creating mottled or marbled pattern. Carapace with uniformly infuscate marbling, more densely infuscate medially. Pedipalp chela fingers and manus, dorsal and retrolateral intercarinal surfaces with moderately infuscate marbling; prolateral and ventral intercarinal surfaces mostly immaculate. Legs retrolateral surfaces with infuscate marbling; prolateral surfaces pale, immaculate. Tergites with unformly infuscate mottling, pale stripe medially, blackish spots submedially, and faint, narrow bands laterally. Sternites pale, with faintly infuscate triangular to trapezoidal marking at posterior margin of III, fading to infuscate mottling on VII. Metasomal segments uniformly, faintly marbled; segment V and telson markedly infuscate, noticeably darker than preceding segments.
|
||
</paragraph>
|
||
<paragraph id="D91836B36D59FF9A25F1FE8CFBADFDD1" blockId="11.[684,1213,226,1738]" pageId="11" pageNumber="11">
|
||
<emphasis id="EBD3EAA16D59FF9A25F1FE8CFC80FEAE" box="[716,824,325,348]" italics="true" pageId="11" pageNumber="11">Carapace:</emphasis>
|
||
Shape trapezoidal; anterior width four-fifths of posterior width (
|
||
<tableCitation id="942503086D59FF9A24D3FEAFFB80FE8C" box="[1006,1080,358,382]" captionStart="TABLE 5" captionStartId="32.[610,690,226,250]" captionTargetBox="[119,1193,387,1697]" captionTargetPageId="32" captionText="TABLE 5 Meristic data for type material of Centruroides berstoni, sp. nov. Material deposited in the California Academy of Sciences (CASENT), San Francisco. Measurements follow Stahnke (1970), Lamoral (1979), and Prendini (2001b)." httpUri="http://table.plazi.org/id/8DD8663B6D72FFB1255FFF2BFC18FE90" pageId="11" pageNumber="11" tableUuid="8DD8663B6D72FFB1255FFF2BFC18FE90">table 5</tableCitation>
|
||
); anteromedian sulcus moderately deep, oval; posteromedian sulcus shallow anteriorly, deeper posteriorly; median ocular tubercle weakly granular; carinae moderately developed, comprising small to medium-sized granules (fig. 6C).
|
||
</paragraph>
|
||
<paragraph id="D91836B36D59FF9A25F1FDE5FBCAFB65" blockId="11.[684,1213,226,1738]" pageId="11" pageNumber="11">
|
||
<emphasis id="EBD3EAA16D59FF9A25F1FDE5FC80FDB6" box="[716,824,556,580]" italics="true" pageId="11" pageNumber="11">Pedipalps:</emphasis>
|
||
Orthobothriotaxic,
|
||
<typeStatus id="061C88116D59FF9A2324FDE4FBF6FDB7" box="[1049,1102,557,581]" pageId="11" pageNumber="11">Type</typeStatus>
|
||
A; femur dorsal trichobothria with α configuration; pedipalp chela fixed finger, trichobothrium
|
||
<emphasis id="EBD3EAA16D59FF9A235AFDA7FB3AFD74" box="[1127,1154,622,646]" italics="true" pageId="11" pageNumber="11">db</emphasis>
|
||
situated slightly distal to
|
||
<emphasis id="EBD3EAA16D59FF9A24A8FD59FC17FD55" box="[917,943,656,679]" italics="true" pageId="11" pageNumber="11">et.</emphasis>
|
||
Femoral carinae serrate; retromedian carinae comprising spiniform granules; dorsal intercarinal surface moderately granular; prolateral intercarinal surface with series of large spiniform granules. Patella carinae strongly developed, granular; prolateral intercarinal surface with five or six large, subspiniform granules. Chela manus prodorsal and dorsomedian carinae absent; retrodorsal carina weakly developed, granular. Fixed finger, median denticle row comprising eight oblique subrows, each flanked by pro- and retrolateral supernumerary denticles. Movable finger, median denticle row with short terminal row comprising four denticles preceded by eight oblique subrows, each flanked by pro- and retrolateral supernumerary denticles.
|
||
</paragraph>
|
||
<paragraph id="D91836B36D59FF9A25F1FB68FBC5FB09" blockId="11.[684,1213,226,1738]" pageId="11" pageNumber="11">
|
||
<emphasis id="EBD3EAA16D59FF9A25F1FB68FCB9FB4A" box="[716,769,1185,1208]" italics="true" pageId="11" pageNumber="11">Legs:</emphasis>
|
||
Leg I length 2× carapace length (
|
||
<tableCitation id="942503086D59FF9A23B8FB68FD72FB28" captionStart="TABLE 10" captionStartId="71.[603,683,226,250]" captionTargetBox="[119,1152,358,736]" captionTargetPageId="71" captionText="TABLE 10 Diagnostic ratios (mean/median/mode) for species of the arboreal Neotropical “thorelli” clade of Centruroides Marx, 1890, bark scorpions" httpUri="http://table.plazi.org/id/8DD8663B6D15FFD62566FF2BFCD9FEB6" pageId="11" pageNumber="11" tableUuid="8DD8663B6D15FFD62566FF2BFCD9FEB6">table 10</tableCitation>
|
||
). Telotarsi ventral surfaces densely covered with short setae; ungues markedly curved.
|
||
</paragraph>
|
||
<paragraph id="D91836B36D59FF9A25F1FACCFBD9FAAC" blockId="11.[684,1213,226,1738]" pageId="11" pageNumber="11">
|
||
<emphasis id="EBD3EAA16D59FF9A25F1FACCFC90FAEE" box="[716,808,1285,1308]" italics="true" pageId="11" pageNumber="11">Pectines:</emphasis>
|
||
Pectinal plate 1.65× wider than long; posterior margin distinctly rounded; pectinal tooth count 16/15 (
|
||
<specimenCount id="CFA1FD3A6D59FF9A24BFFA8CFC21FAAC" box="[898,921,1349,1374]" pageId="11" pageNumber="11" type="male">♂</specimenCount>
|
||
) (fig. 9C,
|
||
<tableCitation id="942503086D59FF9A2335FA8FFBEBFAAC" box="[1032,1107,1350,1374]" captionStart="TABLE 5" captionStartId="32.[610,690,226,250]" captionTargetBox="[119,1193,387,1697]" captionTargetPageId="32" captionText="TABLE 5 Meristic data for type material of Centruroides berstoni, sp. nov. Material deposited in the California Academy of Sciences (CASENT), San Francisco. Measurements follow Stahnke (1970), Lamoral (1979), and Prendini (2001b)." httpUri="http://table.plazi.org/id/8DD8663B6D72FFB1255FFF2BFC18FE90" pageId="11" pageNumber="11" tableUuid="8DD8663B6D72FFB1255FFF2BFC18FE90">table 5</tableCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="D91836B36D59FF9A25F1FAA1FB18F938" blockId="11.[684,1213,226,1738]" pageId="11" pageNumber="11">
|
||
<emphasis id="EBD3EAA16D59FF9A25F1FAA1FCFAFA8D" box="[716,834,1384,1407]" italics="true" pageId="11" pageNumber="11">Mesosoma:</emphasis>
|
||
Tergites width similar to carapace posterior width; I and II slightly narrower (
|
||
<tableCitation id="942503086D59FF9A23BBFA41FD03FA33" captionStart="TABLE 5" captionStartId="32.[610,690,226,250]" captionTargetBox="[119,1193,387,1697]" captionTargetPageId="32" captionText="TABLE 5 Meristic data for type material of Centruroides berstoni, sp. nov. Material deposited in the California Academy of Sciences (CASENT), San Francisco. Measurements follow Stahnke (1970), Lamoral (1979), and Prendini (2001b)." httpUri="http://table.plazi.org/id/8DD8663B6D72FFB1255FFF2BFC18FE90" pageId="11" pageNumber="11" tableUuid="8DD8663B6D72FFB1255FFF2BFC18FE90">table 5</tableCitation>
|
||
). Pretergites surfaces smooth to finely granular. Posttergites surfaces weakly granular; I–VI with dorsomedian carinae weakly granular, restricted to posterior half of each segment; VII surface weakly granular, dorsomedian carinae moderately granular, dorsosubmedian carinae weakly granular, dorsolateral carinae absent. Sternites III–VI, surfaces smooth; VII surface smooth, ventrolateral carinae reduced to few granules.
|
||
</paragraph>
|
||
<caption id="8DD8663B6D5EFF9D2751FBDDFF68FB70" ID-DOI="http://doi.org/10.5281/zenodo.5825904" ID-Zenodo-Dep="5825904" httpUri="https://zenodo.org/record/5825904/files/figure.png" pageId="12" pageNumber="12" startId="12.[108,194,1044,1065]" targetBox="[111,1207,237,1014]" targetPageId="12">
|
||
<paragraph id="D91836B36D5EFF9D2751FBDDFF68FB70" blockId="12.[108,1213,1042,1154]" pageId="12" pageNumber="12">
|
||
FIGURE 3. Map of central Mexico, plotting known locality records of four species of the arboreal Neotropical
|
||
<emphasis id="EBD3EAA16D5EFF9D2751FBE6FF73FBB5" box="[108,203,1071,1095]" italics="true" pageId="12" pageNumber="12">
|
||
“
|
||
<taxonomicName id="1EA74D306D5EFF9D274EFBE6FF7DFBB5" baseAuthorityName="Kraepelin" baseAuthorityYear="1891" box="[115,197,1071,1095]" class="Insecta" family="Cerambycidae" genus="Centruroides" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="12" phylum="Arthropoda" rank="species" species="thorellii">thorellii</taxonomicName>
|
||
”
|
||
</emphasis>
|
||
clade of
|
||
<taxonomicName id="1EA74D306D5EFF9D261AFBE6FD9BFBB5" authority="Marx, 1890" authorityName="MARX" authorityYear="1890" box="[295,547,1071,1095]" class="Insecta" family="Cerambycidae" genus="Centruroides" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="12" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="EBD3EAA16D5EFF9D261AFBE6FE1FFBB5" box="[295,423,1071,1095]" italics="true" pageId="12" pageNumber="12">Centruroides</emphasis>
|
||
<bibRefCitation id="BD364B426D5EFF9D2690FBF9FD9BFBB5" author="Marx" box="[429,547,1071,1095]" firstAuthor="Marx" pageId="12" pageNumber="12" pagination="207 - 211" refId="ref39358" refString="Marx, G. 1890 (" 1889 "). Arachnida. In L. O. Howard (editor), Scientific results of the explorations by the U. S. Fish Commission Steamer Albatross, No. V. Annotated catalogue of the insects collected in 1887 - ' 88. Proceedings of the United States National Museum 12: 207 - 211." type="journal article" year="1890">Marx, 1890</bibRefCitation>
|
||
</taxonomicName>
|
||
, bark scorpions (
|
||
<taxonomicName id="1EA74D306D5EFF9D25EDFBE6FC95FBB5" box="[720,813,1071,1095]" class="Arachnida" family="Buthidae" kingdom="Animalia" order="Scorpiones" pageId="12" pageNumber="12" phylum="Arthropoda" rank="family">Buthidae</taxonomicName>
|
||
C.L. Koch, 1837):
|
||
<taxonomicName id="1EA74D306D5EFF9D24D1FBF9FB2CFBB5" authority="Goodman & Prendini & Francke & Esposito, 2021" authorityName="Goodman & Prendini & Francke & Esposito" authorityYear="2021" box="[1004,1172,1072,1095]" class="Insecta" family="Cerambycidae" genus="Centruroides" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="12" phylum="Arthropoda" rank="species" species="catemacoensis" status="sp. nov.">
|
||
<emphasis id="EBD3EAA16D5EFF9D24D1FBF9FB2CFBB5" box="[1004,1172,1072,1095]" italics="true" pageId="12" pageNumber="12">C. catemacoensis</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicNameLabel id="F0E057DA6D5EFF9D239DFBF9FF2FFB96" pageId="12" pageNumber="12" rank="species">sp. nov.</taxonomicNameLabel>
|
||
(triangles);
|
||
<taxonomicName id="1EA74D306D5EFF9D2624FB84FEC6FB97" authority="Goodman & Prendini & Francke & Esposito, 2021" authorityName="Goodman & Prendini & Francke & Esposito" authorityYear="2021" box="[281,382,1101,1125]" class="Insecta" family="Cerambycidae" genus="Centruroides" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="12" phylum="Arthropoda" rank="species" species="chanae" status="sp. nov.">
|
||
<emphasis id="EBD3EAA16D5EFF9D2624FB84FEC6FB97" box="[281,382,1101,1125]" italics="true" pageId="12" pageNumber="12">C. chanae</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicNameLabel id="F0E057DA6D5EFF9D26B3FB84FE66FB96" box="[398,478,1101,1124]" pageId="12" pageNumber="12" rank="species">sp. nov.</taxonomicNameLabel>
|
||
(circles);
|
||
<taxonomicName id="1EA74D306D5EFF9D257AFB84FD53FB97" authority="Goodman & Prendini & Francke & Esposito, 2021" authorityName="Goodman & Prendini & Francke & Esposito" authorityYear="2021" box="[583,747,1101,1125]" class="Insecta" family="Cerambycidae" genus="Centruroides" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="12" phylum="Arthropoda" rank="species" species="cuauhmapan" status="sp. nov.">
|
||
<emphasis id="EBD3EAA16D5EFF9D257AFB84FD53FB97" box="[583,747,1101,1125]" italics="true" pageId="12" pageNumber="12">C. cuauhmapan</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicNameLabel id="F0E057DA6D5EFF9D25C4FB84FCF2FB96" box="[761,842,1101,1124]" pageId="12" pageNumber="12" rank="species">sp. nov.</taxonomicNameLabel>
|
||
(diamonds);
|
||
<taxonomicName id="1EA74D306D5EFF9D24E5FB84FB04FB97" authority="Sissom, 1995" authorityName="Sissom" authorityYear="1995" box="[984,1212,1101,1125]" class="Insecta" family="Cerambycidae" genus="Centruroides" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="12" phylum="Arthropoda" rank="species" species="rileyi">
|
||
<emphasis id="EBD3EAA16D5EFF9D24E5FB84FB94FB97" box="[984,1068,1101,1125]" italics="true" pageId="12" pageNumber="12">C. rileyi</emphasis>
|
||
<bibRefCitation id="BD364B426D5EFF9D2308FB84FB04FB97" author="Sissom, W. D." box="[1077,1212,1101,1125]" journalOrPublisher="Journal of Arachnology" pageId="12" pageNumber="12" pagination="91 - 99" part="23" refId="ref40441" refString="Sissom, W. D. 1995. Redescription of the scorpion Centruroides thorelli Kraepelin (Buthidae) and description of two new species. Journal of Arachnology 23: 91 - 99." title="Redescription of the scorpion Centruroides thorelli Kraepelin (Buthidae) and description of two new species" type="journal article" year="1995">Sissom, 1995</bibRefCitation>
|
||
</taxonomicName>
|
||
(squares).
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="D91836B36D5EFF9D27B1FB6AFE94FA72" blockId="12.[108,637,1186,1706]" pageId="12" pageNumber="12">
|
||
<emphasis id="EBD3EAA16D5EFF9D27B1FB6AFEB9FB48" box="[140,257,1187,1210]" italics="true" pageId="12" pageNumber="12">Metasoma:</emphasis>
|
||
Metasoma length 3.23× mesosoma length (
|
||
<tableCitation id="942503086D5EFF9D27F8FB0AFE9DFB29" box="[197,293,1219,1243]" captionStart="TABLE 10" captionStartId="71.[603,683,226,250]" captionTargetBox="[119,1152,358,736]" captionTargetPageId="71" captionText="TABLE 10 Diagnostic ratios (mean/median/mode) for species of the arboreal Neotropical “thorelli” clade of Centruroides Marx, 1890, bark scorpions" httpUri="http://table.plazi.org/id/8DD8663B6D15FFD62566FF2BFCD9FEB6" pageId="12" pageNumber="12" tableUuid="8DD8663B6D15FFD62566FF2BFCD9FEB6">table 10</tableCitation>
|
||
). Segments longer than wide; increasing in length posteriorly, segment V 2× length of I; carinae weakly developed, smooth on segments I–IV (figs. 17–21H), absent or obsolete on V (figs. 20–22H); intercarinal surfaces sparsely granular.
|
||
</paragraph>
|
||
<paragraph id="D91836B36D5EFF9D27B1FA40FE4FF9D7" blockId="12.[108,637,1186,1706]" pageId="12" pageNumber="12">
|
||
<emphasis id="EBD3EAA16D5EFF9D27B1FA40FF6EFA53" box="[140,214,1417,1441]" italics="true" pageId="12" pageNumber="12">Telson:</emphasis>
|
||
Vesicle elongate, ovoid; ventral surface shallowly convex; subaculear tubercle narrow and angular in lateral aspect, directed toward midpoint of aculeus. Aculeus angled ventrally at slightly less than 90° (figs. 23–25H).
|
||
</paragraph>
|
||
<paragraph id="D91836B36D5EFF9D27B1F9E6FC59F9D4" blockId="12.[108,637,1186,1706]" lastBlockId="12.[684,1213,1186,1706]" pageId="12" pageNumber="12">
|
||
<emphasis id="EBD3EAA16D5EFF9D27B1F9E6FF4DF9B4" box="[140,245,1583,1606]" italics="true" pageId="12" pageNumber="12">Variation:</emphasis>
|
||
Base color varies from pale yellow to light orange with considerable variation in infuscation of the carapace and mesosoma, despite localities being less than
|
||
<quantity id="1E5F9B566D5EFF9D264CF95BFE0DF958" box="[369,437,1682,1706]" metricMagnitude="4" metricUnit="m" metricValue="5.0" pageId="12" pageNumber="12" unit="km" value="50.0">50 km</quantity>
|
||
apart. Adult males and females differ as follows. The mesosoma is proportionally longer and slenderer, the metasoma up to 3× longer, with segment V markedly longer, and the telson more elongate, with the vesicle more rounded and bilobed posteriorly, in males (figs. 20H, K, 21H, K, 22H, K,
|
||
<tableCitation id="942503086D5EFF9D230AFA8EFB38FAAD" box="[1079,1152,1351,1375]" captionStart="TABLE 5" captionStartId="32.[610,690,226,250]" captionTargetBox="[119,1193,387,1697]" captionTargetPageId="32" captionText="TABLE 5 Meristic data for type material of Centruroides berstoni, sp. nov. Material deposited in the California Academy of Sciences (CASENT), San Francisco. Measurements follow Stahnke (1970), Lamoral (1979), and Prendini (2001b)." httpUri="http://table.plazi.org/id/8DD8663B6D72FFB1255FFF2BFC18FE90" pageId="12" pageNumber="12" tableUuid="8DD8663B6D72FFB1255FFF2BFC18FE90">table 5</tableCitation>
|
||
). The tegument is more densely infuscate, the pectinal plate produced into a rounded lobe posteriorly, which is punctate and slightly infuscate, and the telson shorter and narrower with the vesicle surfaces less granular, in females (figs. 9C, D, 23H, K, 24H, K, 25H, K, 34–35A, B).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="91BD65386D5EFF9D25F1F9E6FB13F958" pageId="12" pageNumber="12" type="distribution">
|
||
<paragraph id="D91836B36D5EFF9D25F1F9E6FB13F958" blockId="12.[684,1213,1186,1706]" pageId="12" pageNumber="12">
|
||
DISTRIBUTION:
|
||
<taxonomicName id="1EA74D306D5EFF9D24ACF9E6FB2CF9B5" authorityName="Goodman & Prendini & Francke & Esposito" authorityYear="2021" box="[913,1172,1583,1607]" class="Insecta" family="Cerambycidae" genus="Centruroides" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="12" phylum="Arthropoda" rank="species" species="berstoni">
|
||
<emphasis id="EBD3EAA16D5EFF9D24ACF9E6FB2CF9B5" box="[913,1172,1583,1607]" italics="true" pageId="12" pageNumber="12">Centruroides berstoni</emphasis>
|
||
</taxonomicName>
|
||
is endemic to the
|
||
<collectingRegion id="1B63F8516D5EFF9D246BF999FB9BF995" box="[854,1059,1616,1640]" country="Guatemala" name="Izabal" pageId="12" pageNumber="12">Izabal Department</collectingRegion>
|
||
of
|
||
<collectingCountry id="A1B076236D5EFF9D237BF999FB04F99A" box="[1094,1212,1616,1640]" name="Guatemala" pageId="12" pageNumber="12">Guatemala</collectingCountry>
|
||
and has been recorded from several localities around Morales and Río Dulce (figs. 1B, D, 4).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<caption id="8DD8663B6D5FFF9C2750FBD1FC1FFB75" ID-DOI="http://doi.org/10.5281/zenodo.5825908" ID-Zenodo-Dep="5825908" httpUri="https://zenodo.org/record/5825908/files/figure.png" pageId="13" pageNumber="13" startId="13.[109,195,1048,1069]" targetBox="[111,1203,241,1014]" targetPageId="13">
|
||
<paragraph id="D91836B36D5FFF9C2750FBD1FC1FFB75" blockId="13.[109,1212,1046,1159]" pageId="13" pageNumber="13">
|
||
FIGURE 4. Map of southeastern Mexico and northwestern Central America, plotting known locality records of five species of the arboreal Neotropical
|
||
<emphasis id="EBD3EAA16D5FFF9C26C3FBFDFDE1FBBE" box="[510,601,1076,1100]" italics="true" pageId="13" pageNumber="13">
|
||
“
|
||
<taxonomicName id="1EA74D306D5FFF9C2539FBFDFDEBFBBE" baseAuthorityName="Kraepelin" baseAuthorityYear="1891" box="[516,595,1076,1100]" class="Insecta" family="Cerambycidae" genus="Centruroides" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="13" phylum="Arthropoda" rank="species" species="thorellii">thorellii</taxonomicName>
|
||
”
|
||
</emphasis>
|
||
clade of
|
||
<taxonomicName id="1EA74D306D5FFF9C2585FBFDFC0AFBBE" authority="Marx, 1890" authorityName="MARX" authorityYear="1890" box="[696,946,1076,1100]" class="Insecta" family="Cerambycidae" genus="Centruroides" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="13" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="EBD3EAA16D5FFF9C2585FBFDFC8DFBBE" box="[696,821,1076,1100]" italics="true" pageId="13" pageNumber="13">Centruroides</emphasis>
|
||
<bibRefCitation id="BD364B426D5FFF9C2400FBFDFC0AFBBE" author="Marx" box="[829,946,1076,1100]" firstAuthor="Marx" pageId="13" pageNumber="13" pagination="207 - 211" refId="ref39358" refString="Marx, G. 1890 (" 1889 "). Arachnida. In L. O. Howard (editor), Scientific results of the explorations by the U. S. Fish Commission Steamer Albatross, No. V. Annotated catalogue of the insects collected in 1887 - ' 88. Proceedings of the United States National Museum 12: 207 - 211." type="journal article" year="1890">Marx, 1890</bibRefCitation>
|
||
</taxonomicName>
|
||
, bark scorpions (
|
||
<taxonomicName id="1EA74D306D5FFF9C235DFBFDFB04FBBE" box="[1120,1212,1076,1100]" class="Arachnida" family="Buthidae" kingdom="Animalia" order="Scorpiones" pageId="13" pageNumber="13" phylum="Arthropoda" rank="family">Buthidae</taxonomicName>
|
||
C.L. Koch, 1837):
|
||
<taxonomicName id="1EA74D306D5FFF9C261EFB9BFE35FB9B" authority="Goodman & Prendini & Francke & Esposito, 2021" authorityName="Goodman & Prendini & Francke & Esposito" authorityYear="2021" box="[291,397,1105,1129]" class="Insecta" family="Cerambycidae" genus="Centruroides" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="13" phylum="Arthropoda" rank="species" species="berstoni" status="sp. nov.">
|
||
<emphasis id="EBD3EAA16D5FFF9C261EFB9BFE35FB9B" box="[291,397,1105,1129]" italics="true" pageId="13" pageNumber="13">C. berstoni</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicNameLabel id="F0E057DA6D5FFF9C26A4FB9BFE5DFB9B" box="[409,485,1106,1129]" pageId="13" pageNumber="13" rank="species">sp. nov.</taxonomicNameLabel>
|
||
(circles);
|
||
<taxonomicName id="1EA74D306D5FFF9C257AFB9BFD68FB9B" authority="Goodman & Prendini & Francke & Esposito, 2021" authorityName="Goodman & Prendini & Francke & Esposito" authorityYear="2021" box="[583,720,1105,1129]" class="Insecta" family="Cerambycidae" genus="Centruroides" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="13" phylum="Arthropoda" rank="species" species="hamadryas" status="sp. nov.">
|
||
<emphasis id="EBD3EAA16D5FFF9C257AFB9BFD68FB9B" box="[583,720,1105,1129]" italics="true" pageId="13" pageNumber="13">C. hamadryas</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicNameLabel id="F0E057DA6D5FFF9C25E0FB9BFC91FB9B" box="[733,809,1106,1129]" pageId="13" pageNumber="13" rank="species">sp. nov.</taxonomicNameLabel>
|
||
(diamonds);
|
||
<taxonomicName id="1EA74D306D5FFF9C2492FB9BFB04FB9B" authority="Armas, 1996" authorityName="Armas" authorityYear="1996" box="[943,1212,1105,1129]" class="Insecta" family="Cerambycidae" genus="Centruroides" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="13" phylum="Arthropoda" rank="species" species="hoffmanni">
|
||
<emphasis id="EBD3EAA16D5FFF9C2492FB9BFB8AFB9B" box="[943,1074,1105,1129]" italics="true" pageId="13" pageNumber="13">C. hoffmanni</emphasis>
|
||
<bibRefCitation id="BD364B426D5FFF9C2307FB98FB04FB9B" author="Armas" box="[1082,1212,1105,1129]" firstAuthor="Armas" pageId="13" pageNumber="13" pagination="21 - 33" refId="ref36038" refString="Armas, L. F. de. 1996. Presencia de Centruroides schmidti Sissom en el sureste de Mexico y descripcion de dos especies nuevas (Scorpiones: Buthidae). Revista Nicaraguense de Entomologia 36: 21 - 33." type="journal article" year="1996">Armas, 1996</bibRefCitation>
|
||
</taxonomicName>
|
||
(triangles);
|
||
<taxonomicName id="1EA74D306D5FFF9C27DFFBA6FE66FB74" authority="Sissom, 1995" authorityName=": Sissom" authorityYear="1995" box="[226,478,1134,1158]" class="Insecta" family="Cerambycidae" genus="Centruroides" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="13" phylum="Arthropoda" rank="species" species="schmidti">
|
||
<emphasis id="EBD3EAA16D5FFF9C27DFFBA6FEEBFB74" box="[226,339,1134,1158]" italics="true" pageId="13" pageNumber="13">C. schmidti</emphasis>
|
||
<bibRefCitation id="BD364B426D5FFF9C2667FBA7FE66FB74" author="Sissom, W. D." box="[346,478,1134,1158]" journalOrPublisher="Journal of Arachnology" pageId="13" pageNumber="13" pagination="91 - 99" part="23" refId="ref40441" refString="Sissom, W. D. 1995. Redescription of the scorpion Centruroides thorelli Kraepelin (Buthidae) and description of two new species. Journal of Arachnology 23: 91 - 99." title="Redescription of the scorpion Centruroides thorelli Kraepelin (Buthidae) and description of two new species" type="journal article" year="1995">Sissom, 1995</bibRefCitation>
|
||
</taxonomicName>
|
||
(squares);
|
||
<taxonomicName id="1EA74D306D5FFF9C2572FBA6FD53FB74" authority="Goodman & Prendini & Francke & Esposito, 2021" authorityName="Goodman & Prendini & Francke & Esposito" authorityYear="2021" box="[591,747,1135,1158]" class="Insecta" family="Cerambycidae" genus="Centruroides" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="13" phylum="Arthropoda" rank="species" species="yucatanensis" status="sp. nov.">
|
||
<emphasis id="EBD3EAA16D5FFF9C2572FBA6FD53FB74" box="[591,747,1135,1158]" italics="true" pageId="13" pageNumber="13">C. yucatanensis</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicNameLabel id="F0E057DA6D5FFF9C25CBFBA6FCFAFB74" box="[758,834,1135,1158]" pageId="13" pageNumber="13" rank="species">sp. nov.</taxonomicNameLabel>
|
||
(crosses).
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection id="91BD65386D5FFF9C27B0FB6CFD80FA56" pageId="13" pageNumber="13" type="biology_ecology">
|
||
<paragraph id="D91836B36D5FFF9C27B0FB6CFD80FA56" blockId="13.[108,637,1188,1708]" pageId="13" pageNumber="13">
|
||
ECOLOGY: The localities at which
|
||
<taxonomicName id="1EA74D306D5FFF9C253AFB6CFDC4FB4E" authorityName="Goodman & Prendini & Francke & Esposito" authorityYear="2021" box="[519,636,1188,1212]" class="Insecta" family="Cerambycidae" genus="Centruroides" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="13" phylum="Arthropoda" rank="species" species="berstoni">
|
||
<emphasis id="EBD3EAA16D5FFF9C253AFB6CFDC4FB4E" box="[519,636,1188,1212]" italics="true" pageId="13" pageNumber="13">C. berstoni</emphasis>
|
||
</taxonomicName>
|
||
was recorded range in altitude from
|
||
<quantity id="1E5F9B566D5FFF9C26D2FB0CFDF5FB2F" box="[495,589,1221,1245]" metricMagnitude="1" metricUnit="m" metricValue="1.6" metricValueMax="1.7" metricValueMin="1.5" pageId="13" pageNumber="13" unit="m" value="16.0" valueMax="17.0" valueMin="15.0">15–17 m</quantity>
|
||
and occur in a region of humid tropical lowland rainforest. Specimens were collected at night with UV light detection, mostly on trees, sitting on bark or branches, and large shoots of bamboo. The habitat and habitus are consistent with the arboreal, corticolous ecomorphotype (
|
||
<bibRefCitation id="BD364B426D5FFF9C2640FA45FD94FA56" author="Prendini, L." box="[381,556,1420,1444]" editor="V. Fet & P. A. Selden" journalOrPublisher="Burnham Beeches, Bucks, U. K.: British Arachnological Society" pageId="13" pageNumber="13" pagination="113 - 138" refId="ref40441" refString="Prendini, L. 2001 a. Substratum specialization and speciation in southern African scorpions: the Effect Hypothesis revisited. In V. Fet, and P. A. Selden (editors), Scorpions 2001: in memoriam Gary A. Polis: 113 - 138. Burnham Beeches, Bucks, U. K.: British Arachnological Society." title="Substratum specialization and speciation in southern African scorpions: the Effect Hypothesis revisited" type="book chapter" volumeTitle="Scorpions 2001: in memoriam Gary A. Polis" year="2001">Prendini, 2001a</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="91BD65386D5FFF9C27B1FA64FC19FB2C" pageId="13" pageNumber="13" type="discussion">
|
||
<paragraph id="D91836B36D5FFF9C27B1FA64FC19FB2C" blockId="13.[108,637,1188,1708]" lastBlockId="13.[684,1216,1188,1709]" pageId="13" pageNumber="13">
|
||
REMARKS: The
|
||
<typeStatus id="061C88116D5FFF9C2667FA64FE78FA37" box="[346,448,1453,1477]" pageId="13" pageNumber="13">paratype</typeStatus>
|
||
female of
|
||
<taxonomicName id="1EA74D306D5FFF9C255EFA64FF71FA14" authorityName=": Sissom" authorityYear="1995" class="Insecta" family="Cerambycidae" genus="Centruroides" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="13" phylum="Arthropoda" rank="species" species="schmidti">
|
||
<emphasis id="EBD3EAA16D5FFF9C255EFA64FF71FA14" italics="true" pageId="13" pageNumber="13">C. schmidti</emphasis>
|
||
</taxonomicName>
|
||
, from Morales,
|
||
<collectingCountry id="A1B076236D5FFF9C26B7FA07FDBEFA14" box="[394,518,1486,1510]" name="Guatemala" pageId="13" pageNumber="13">Guatemala</collectingCountry>
|
||
, was misidentified by
|
||
<bibRefCitation id="BD364B426D5FFF9C2618FA26FE64F9F5" author="Sissom, W. D." box="[293,476,1519,1543]" journalOrPublisher="Journal of Arachnology" pageId="13" pageNumber="13" pagination="91 - 99" part="23" refId="ref40441" refString="Sissom, W. D. 1995. Redescription of the scorpion Centruroides thorelli Kraepelin (Buthidae) and description of two new species. Journal of Arachnology 23: 91 - 99." title="Redescription of the scorpion Centruroides thorelli Kraepelin (Buthidae) and description of two new species" type="journal article" year="1995">Sissom (1995)</bibRefCitation>
|
||
. Differences between the
|
||
<typeStatus id="061C88116D5FFF9C2639F9D8FEDEF9DB" box="[260,358,1553,1577]" pageId="13" pageNumber="13">paratype</typeStatus>
|
||
and other females that species, evident from the illustrations (
|
||
<bibRefCitation id="BD364B426D5FFF9C251BF9F8FF49F998" author="Sissom, W. D." journalOrPublisher="Journal of Arachnology" pageId="13" pageNumber="13" pagination="91 - 99" part="23" refId="ref40441" refString="Sissom, W. D. 1995. Redescription of the scorpion Centruroides thorelli Kraepelin (Buthidae) and description of two new species. Journal of Arachnology 23: 91 - 99." title="Redescription of the scorpion Centruroides thorelli Kraepelin (Buthidae) and description of two new species" type="journal article" year="1995">Sissom, 1995: 94–95</bibRefCitation>
|
||
figs. 16–18), include faint reticulate infuscation on the chelicerae, less infuscation of the carapace and tergites, less granulation and setation of the metasomal segments, and a bilobed telson vesicle.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="91BD65386D5FFF9E25F1FB2EFF74FB62" lastPageId="15" lastPageNumber="15" pageId="13" pageNumber="13" type="materials_examined">
|
||
<paragraph id="D91836B36D5FFF9E25F1FB2EFF74FB62" blockId="13.[684,1216,1188,1709]" lastBlockId="15.[108,637,1011,1168]" lastPageId="15" lastPageNumber="15" pageId="13" pageNumber="13">
|
||
<materialsCitation id="69CF3CEE6D5FFF9C25F1FB2EFC5EFA71" ID-GBIF-Occurrence="3427257354" collectingDate="2019-09-25" collectionCode="MATERIAL" collectorName="A. M. Goodman" country="Guatemala" elevation="15" latitude="15.734806" location="Biotopo Chocon Machacas" longLatPrecision="1" longitude="-88.91589" municipality="Municipio Livingston" pageId="13" pageNumber="13" specimenCount="10" specimenCount-female="1" specimenCount-male="9" stateProvince="Izabal">
|
||
<collectionCode id="BFB6AE766D5FFF9C25F1FB2EFC83FB0D" box="[716,827,1255,1279]" pageId="13" pageNumber="13">MATERIAL</collectionCode>
|
||
EXAMINED:
|
||
<emphasis id="EBD3EAA16D5FFF9C24F9FB2FFBD3FB0C" bold="true" box="[964,1131,1254,1278]" pageId="13" pageNumber="13">
|
||
<collectingCountry id="A1B076236D5FFF9C24F9FB2FFBD0FB0C" box="[964,1128,1254,1278]" name="Guatemala" pageId="13" pageNumber="13">GUATEMALA</collectingCountry>
|
||
:
|
||
</emphasis>
|
||
<emphasis id="EBD3EAA16D5FFF9C2349FB2EFCE8FAD2" italics="true" pageId="13" pageNumber="13">
|
||
<collectingRegion id="1B63F8516D5FFF9C2349FB2EFCE8FAD2" country="Guatemala" name="Izabal" pageId="13" pageNumber="13">Departamento Izabal</collectingRegion>
|
||
</emphasis>
|
||
:
|
||
<collectingMunicipality id="397CACC96D5FFF9C2459FAC1FBEDFAD2" box="[868,1109,1288,1312]" pageId="13" pageNumber="13">Município Livingston</collectingMunicipality>
|
||
:
|
||
<location id="DC7860686D5FFF9C235AFAC1FCC1FAB3" LSID="urn:lsid:plazi:treatment:510E87A56D5AFF9E2436FBA1FF74FB62:DC7860686D5FFF9C235AFAC1FCC1FAB3" country="Guatemala" latitude="15.734806" longLatPrecision="1" longitude="-88.91589" municipality="Municipio Livingston" name="Biotopo Chocon Machacas" pageId="13" pageNumber="13" stateProvince="Izabal">Biotopo Chocón Machacas</location>
|
||
,
|
||
<geoCoordinate id="BC9350746D5FFF9C24B7FAE0FBA0FAB2" box="[906,1048,1320,1345]" degrees="15" direction="north" minutes="44" orientation="latitude" pageId="13" pageNumber="13" precision="1" seconds="05.3" value="15.734806">15°44′05.3″N</geoCoordinate>
|
||
<geoCoordinate id="BC9350746D5FFF9C2318FAE0FB04FAB2" box="[1061,1212,1320,1345]" degrees="88" direction="west" minutes="54" orientation="longitude" pageId="13" pageNumber="13" precision="1" seconds="57.2" value="-88.91589">88°54′57.2″W</geoCoordinate>
|
||
,
|
||
<quantity id="1E5F9B566D5FFF9C2591FA83FD50FA93" box="[684,744,1354,1378]" metricMagnitude="1" metricUnit="m" metricValue="1.5" pageId="13" pageNumber="13" unit="m" value="15.0">
|
||
<elevation id="528AD1806D5FFF9C2591FA83FD50FA93" box="[684,744,1354,1378]" metricMagnitude="1" metricUnit="m" metricValue="1.5" pageId="13" pageNumber="13" unit="m" value="15.0">15 m</elevation>
|
||
</quantity>
|
||
,
|
||
<date id="AD1910736D5FFF9C25CAFA83FCDCFA90" box="[759,868,1354,1378]" pageId="13" pageNumber="13" value="2019-09-25">
|
||
<collectingDate id="BD5DE99B6D5FFF9C25CAFA83FCDCFA90" box="[759,868,1354,1378]" pageId="13" pageNumber="13" value="2019-09-25">25.ix.2019</collectingDate>
|
||
</date>
|
||
,
|
||
<collectorName id="745253656D5FFF9C244EFA83FB9AFA90" box="[883,1058,1354,1378]" pageId="13" pageNumber="13">A.M. Goodman</collectorName>
|
||
,
|
||
<specimenCount id="CFA1FD3A6D5FFF9C230CFA83FBD8FA90" box="[1073,1120,1353,1378]" pageId="13" pageNumber="13" type="male">8 ♂</specimenCount>
|
||
,
|
||
<specimenCount id="CFA1FD3A6D5FFF9C234DFA83FB27FA90" box="[1136,1183,1353,1378]" pageId="13" pageNumber="13" type="female">1 ♀</specimenCount>
|
||
,
|
||
<specimenCount id="CFA1FD3A6D5FFF9C2392FA83FD4EFA71" pageId="13" pageNumber="13" type="male">1 juv. ♂</specimenCount>
|
||
(CASENT 9073271)
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation id="69CF3CEE6D5FFF9C24C4FAA2FB04F9F5" ID-GBIF-Occurrence="3427257379" collectingDate="2006-07-08" collectingDateMax="2019-09-25" collectingDateMin="2006-07-08" collectorName="J. Huff & C. Viquez & D. Ortiz & A. M. Goodman" country="Guatemala" elevation="49" latitude="15.734806" location="Hotel Tijax" longLatPrecision="1" longitude="-88.91589" municipality="Rio Dulce" pageId="13" pageNumber="13" specimenCount="2" specimenCount-male="2" stateProvince="Izabal">
|
||
<collectingMunicipality id="397CACC96D5FFF9C24C4FAA2FBD7FA71" box="[1017,1135,1387,1411]" pageId="13" pageNumber="13">Río Dulce</collectingMunicipality>
|
||
,
|
||
<location id="DC7860686D5FFF9C23BCFAA2FD5BFA56" LSID="urn:lsid:plazi:treatment:510E87A56D5AFF9E2436FBA1FF74FB62:DC7860686D5FFF9C23BCFAA2FD5BFA56" country="Guatemala" latitude="15.734806" longLatPrecision="1" longitude="-88.91589" municipality="Rio Dulce" name="Hotel Tijax" pageId="13" pageNumber="13" stateProvince="Izabal">Hotel Tijax</location>
|
||
,
|
||
<geoCoordinate id="BC9350746D5FFF9C25D2FA45FCC2FA51" box="[751,890,1419,1444]" degrees="15" direction="north" minutes="40" orientation="latitude" pageId="13" pageNumber="13" precision="1" seconds="12.2" value="15.670056">15°40′12.2″N</geoCoordinate>
|
||
<geoCoordinate id="BC9350746D5FFF9C24BEFA45FBB8FA51" box="[899,1024,1419,1444]" degrees="89" direction="west" minutes="00" orientation="longitude" pageId="13" pageNumber="13" precision="15" seconds="27" value="-89.0075">89°00′27″W</geoCoordinate>
|
||
,
|
||
<quantity id="1E5F9B566D5FFF9C2331FA45FBFEFA51" box="[1036,1094,1420,1444]" metricMagnitude="1" metricUnit="m" metricValue="4.9" pageId="13" pageNumber="13" unit="m" value="49.0">
|
||
<elevation id="528AD1806D5FFF9C2331FA45FBFEFA51" box="[1036,1094,1420,1444]" metricMagnitude="1" metricUnit="m" metricValue="4.9" pageId="13" pageNumber="13" unit="m" value="49.0">49 m</elevation>
|
||
</quantity>
|
||
,
|
||
<date id="AD1910736D5FFF9C236FFA45FB01FA56" box="[1106,1209,1420,1444]" pageId="13" pageNumber="13" value="2006-07-08">
|
||
<collectingDate id="BD5DE99B6D5FFF9C236FFA45FB01FA56" box="[1106,1209,1420,1444]" pageId="13" pageNumber="13" value="2006-07-08">8.vii.2006</collectingDate>
|
||
</date>
|
||
,
|
||
<collectorName id="745253656D5FFF9C2591FA67FD4FFA37" box="[684,759,1453,1477]" pageId="13" pageNumber="13">J. Huff</collectorName>
|
||
,
|
||
<collectorName id="745253656D5FFF9C2439FA64FCC9FA37" box="[772,881,1453,1477]" pageId="13" pageNumber="13">C. Víquez</collectorName>
|
||
, and
|
||
<collectorName id="745253656D5FFF9C248DFA67FBB5FA37" box="[944,1037,1453,1477]" pageId="13" pageNumber="13">D. Ortíz</collectorName>
|
||
, collected along trails through old secondary growth tropical forest using UV at night,
|
||
<specimenCount id="CFA1FD3A6D5FFF9C249BFA26FC6BF9F5" box="[934,979,1518,1543]" pageId="13" pageNumber="13" type="male">1 ♂</specimenCount>
|
||
(AMNH [LP 5984]),
|
||
</materialsCitation>
|
||
<materialsCitation id="69CF3CEE6D5FFF9C2591F9D9FB97F97E" ID-GBIF-Occurrence="3427257375" collectingDate="2006-07-08" collectingDateMax="2019-09-25" collectingDateMin="2006-07-08" collectorName="A. M. Goodman" country="Guatemala" elevation="49" latitude="15.734806" location="Hotel Tijax" longLatPrecision="1" longitude="-88.91589" municipality="Rio Dulce" pageId="13" pageNumber="13" specimenCount="2" specimenCount-male="2" stateProvince="Izabal">
|
||
<geoCoordinate id="BC9350746D5FFF9C2591F9D9FC8EF9DA" box="[684,822,1551,1576]" degrees="15" direction="north" minutes="39" orientation="latitude" pageId="13" pageNumber="13" precision="1" seconds="51.2" value="15.664222">15°39′51.2″N</geoCoordinate>
|
||
<geoCoordinate id="BC9350746D5FFF9C2406F9D9FC71F9DA" box="[827,969,1551,1576]" degrees="89" direction="west" minutes="00" orientation="longitude" pageId="13" pageNumber="13" precision="1" seconds="14.6" value="-89.00406">89°00′14.6″W</geoCoordinate>
|
||
,
|
||
<quantity id="1E5F9B566D5FFF9C24EFF9D9FBB0F9DA" box="[978,1032,1552,1576]" metricMagnitude="1" metricUnit="m" metricValue="1.7" pageId="13" pageNumber="13" unit="m" value="17.0">
|
||
<elevation id="528AD1806D5FFF9C24EFF9D9FBB0F9DA" box="[978,1032,1552,1576]" metricMagnitude="1" metricUnit="m" metricValue="1.7" pageId="13" pageNumber="13" unit="m" value="17.0">17 m</elevation>
|
||
</quantity>
|
||
,
|
||
<date id="AD1910736D5FFF9C232EF9D9FBC6F9DA" box="[1043,1150,1552,1576]" pageId="13" pageNumber="13" value="2019-09-24">
|
||
<collectingDate id="BD5DE99B6D5FFF9C232EF9D9FBC6F9DA" box="[1043,1150,1552,1576]" pageId="13" pageNumber="13" value="2019-09-24">24.ix.2019</collectingDate>
|
||
</date>
|
||
,
|
||
<collectorName id="745253656D5FFF9C23BAF9D9FCA1F9BB" pageId="13" pageNumber="13">A.M. Goodman</collectorName>
|
||
, collected along gravel road of Hacienda Tijax Parking Lot, flanked by bamboo groves and live fencing,
|
||
<specimenCount id="CFA1FD3A6D5FFF9C2416F9BDFCEBF979" box="[811,851,1650,1676]" pageId="13" pageNumber="13" type="male">1 ♂</specimenCount>
|
||
(CASENT 9073272),
|
||
</materialsCitation>
|
||
<materialsCitation id="69CF3CEE6D5FFF9E230EF9BAFDB0FBFE" ID-GBIF-Occurrence="3427257302" collectingDate="2006-07-08" collectingDateMax="2019-09-25" collectingDateMin="2006-07-08" collectorName="A. M. Goodman" country="Guatemala" elevation="49" lastPageId="15" lastPageNumber="15" latitude="15.734806" location="Hotel Tijax" longLatPrecision="1" longitude="-88.91589" municipality="Rio Dulce" pageId="13" pageNumber="13" specimenCount="2" specimenCount-male="2" stateProvince="Izabal">
|
||
<geoCoordinate id="BC9350746D5FFF9C230EF9BAFB04F979" box="[1075,1212,1650,1675]" degrees="15" direction="north" minutes="44" orientation="latitude" pageId="13" pageNumber="13" precision="1" seconds="05.3" value="15.734806">15°44′05.3″N</geoCoordinate>
|
||
<geoCoordinate id="BC9350746D5FFF9C2591F95DFC82F95E" box="[684,826,1683,1708]" degrees="88" direction="west" minutes="54" orientation="longitude" pageId="13" pageNumber="13" precision="1" seconds="57.2" value="-88.91589">88°54′57.2″W</geoCoordinate>
|
||
,
|
||
<quantity id="1E5F9B566D5FFF9C2479F95DFCC3F95E" box="[836,891,1684,1708]" metricMagnitude="1" metricUnit="m" metricValue="1.5" pageId="13" pageNumber="13" unit="m" value="15.0">
|
||
<elevation id="528AD1806D5FFF9C2479F95DFCC3F95E" box="[836,891,1684,1708]" metricMagnitude="1" metricUnit="m" metricValue="1.5" pageId="13" pageNumber="13" unit="m" value="15.0">15 m</elevation>
|
||
</quantity>
|
||
,
|
||
<date id="AD1910736D5FFF9C24BAF95DFC49F95E" box="[903,1009,1684,1708]" pageId="13" pageNumber="13" value="2019-09-25">
|
||
<collectingDate id="BD5DE99B6D5FFF9C24BAF95DFC49F95E" box="[903,1009,1684,1708]" pageId="13" pageNumber="13" value="2019-09-25">25.ix.2019</collectingDate>
|
||
</date>
|
||
,
|
||
<collectorName id="745253656D5FFF9C24C1F95CFB1DF95E" box="[1020,1189,1684,1709]" pageId="13" pageNumber="13">A.M. Goodman</collectorName>
|
||
, 8
|
||
<subSection id="B0282F586D5DFF9E2751FC3AFF74FB62" pageId="15" pageNumber="15" type="multiple">
|
||
<specimenCount id="CFA1FD3A6D5DFF9E2751FC3AFF3BFBFE" box="[108,131,1011,1036]" pageId="15" pageNumber="15" type="male">♂</specimenCount>
|
||
,
|
||
<specimenCount id="CFA1FD3A6D5DFF9E27ACFC3DFF05FBFE" box="[145,189,1011,1036]" pageId="15" pageNumber="15" type="female">1 ♀</specimenCount>
|
||
,
|
||
<specimenCount id="CFA1FD3A6D5DFF9E27F6FC3DFE9CFBFE" box="[203,292,1011,1036]" pageId="15" pageNumber="15" type="male">1 juv. ♂</specimenCount>
|
||
(CASENT 9073271).
|
||
<materialsCitation id="69CF3CEE6D5DFF9E2532FC3DFF74FB62" ID-GBIF-Occurrence="3427257368" collectingDate="2006-07-08" collectingDateMax="2019-09-25" collectingDateMin="2006-07-08" collectorName="D. Barrales & R. Monjaraz" country="Guatemala" elevation="49" latitude="15.734806" location="Hotel Tijax" longLatPrecision="1" longitude="-88.91589" municipality="Rio Dulce" pageId="15" pageNumber="15" specimenCount="2" specimenCount-male="2" stateProvince="Izabal">
|
||
Município Morales: Morales, Finca Fiymeza, Sendero Anfibio,
|
||
<geoCoordinate id="BC9350746D5DFF9E2751FBFFFF41FBBC" box="[108,249,1077,1102]" degrees="15" direction="north" minutes="24" orientation="latitude" pageId="15" pageNumber="15" precision="1" seconds="24.1" value="15.406694">15°24′24.1″N</geoCoordinate>
|
||
<geoCoordinate id="BC9350746D5DFF9E2639FBFFFE2EFBBC" box="[260,406,1077,1102]" degrees="88" direction="west" minutes="41" orientation="longitude" pageId="15" pageNumber="15" precision="1" seconds="46.8" value="-88.696335">88°41′46.8″W</geoCoordinate>
|
||
,
|
||
<quantity id="1E5F9B566D5DFF9E2698FBFFFE56FBBC" box="[421,494,1078,1102]" metricMagnitude="2" metricUnit="m" metricValue="5.95" pageId="15" pageNumber="15" unit="m" value="595.0">595 m</quantity>
|
||
,
|
||
<date id="AD1910736D5DFF9E26C0FBFFFDC1FBBC" box="[509,633,1078,1102]" pageId="15" pageNumber="15" value="2017-08-17">17.viii.2017</date>
|
||
, D. Barrales and R. Monjaraz,
|
||
<specimenCount id="CFA1FD3A6D5DFF9E2680FB9EFDA5FB9D" box="[445,541,1110,1135]" pageId="15" pageNumber="15" type="female">1 juv. ♀</specimenCount>
|
||
(CNAN SC3968).
|
||
</materialsCitation>
|
||
</subSection>
|
||
</materialsCitation>
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |