306 lines
38 KiB
XML
306 lines
38 KiB
XML
<document id="3D756EC1875B9626D41353125E41B15C" ID-DOI="10.13133/2284-4880/545" ID-ISSN="2284-4880" ID-Zenodo-Dep="12764154" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1721242503917" checkinUser="felipe" docAuthor="Ballerio, Alberto" docDate="2021" docId="0F747550111DFFAAFF16AC251A2E91C8" docLanguage="en" docName="FragEntomol.53.2.105-282.pdf" docOrigin="Fragmenta entomologica 53 (2)" docSource="http://dx.doi.org/10.13133/2284-4880/545" docStyle="DocumentStyle:8499FF4308F512386C0E43DC290A16BD.1:FragEntomol.2021-.journal_article.0cover" docStyleId="8499FF4308F512386C0E43DC290A16BD" docStyleName="FragEntomol.2021-.journal_article.0cover" docStyleVersion="1" docTitle="Goudotostes elegans Ballerio, 2021, sp. nov." docType="treatment" docVersion="2" lastPageNumber="177" masterDocId="F34D0D28115AFFE2FF9EAF31192A9224" masterDocTitle="The Ceratocanthinae of Madagascar and Comoro Islands: a revision of the genera Synarmostes and Goudotostes, and of the flightless Philharmostes, with description of 64 new species (Coleoptera: Scarabaeoidea, Hybosoridae)" masterLastPageNumber="282" masterPageNumber="105" pageNumber="176" updateTime="1721247602959" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-NC-4.0">
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<mods:mods id="A89431A694D803734F3CAAD96361FC43" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="85B1508E70FC46176D8C8BCDC8BC6285">
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<mods:title id="36941A46A228155C1EB5A8789DE9369E">The Ceratocanthinae of Madagascar and Comoro Islands: a revision of the genera Synarmostes and Goudotostes, and of the flightless Philharmostes, with description of 64 new species (Coleoptera: Scarabaeoidea, Hybosoridae)</mods:title>
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</mods:titleInfo>
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<mods:name id="57E6443DD67943F04102E73C768BFA34" type="personal">
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<mods:role id="AFCADE321817380BF69822BB2FDC756C">
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<mods:roleTerm id="2195F60E2D77BA524C00DDC5B8F061EB">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="38881503CF384844BDF12DB5308A990D">Ballerio, Alberto</mods:namePart>
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<mods:typeOfResource id="75A4EC89219726342AACDF52739877DE">text</mods:typeOfResource>
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<mods:title id="4641F1D7BC9896D9E898CE67C2D17C6F">Fragmenta entomologica</mods:title>
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</mods:titleInfo>
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<mods:part id="9D5E794580036701EDE9A99876744456">
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<mods:date id="960C0418F58FF9AC4B5A4C601937484A">2021</mods:date>
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<mods:detail id="8500A65DC18B5D6839CC9391B28446CC" type="pubDate">
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<mods:number id="A0A4C07FA9E53F17B61E0B3555801ABA">2021-11-30</mods:number>
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</mods:detail>
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<mods:detail id="3AB699EB702F1FDD060E2142B12503A3" type="volume">
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<mods:number id="8F6E320AB3FE5D30F03937FB1FDF861C">53</mods:number>
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<mods:detail id="14E3BA936E11F7A359F400A31C45CF51" type="issue">
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<mods:number id="277DD36C70EC4C33D403404660B9022D">2</mods:number>
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</mods:detail>
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<mods:extent id="EAA3AD917FDDE312FD77C1CD1FFD77C3" unit="page">
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<mods:start id="3702AFBEF9A00E508A26C19FB520C7EF">105</mods:start>
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<mods:end id="7A0C14D3C81251DA288C907AE143624C">282</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:location id="7C9D4137429B75A1791384522E0D9AF0">
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<mods:url id="20D4399DF558B1F5C7048F5A53EB5193">http://dx.doi.org/10.13133/2284-4880/545</mods:url>
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</mods:location>
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<mods:classification id="4998BB02CB209D58399CFE89935DC512">journal article</mods:classification>
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<mods:identifier id="485AF0C4ECDC6253C218F23BC8E9B870" type="DOI">10.13133/2284-4880/545</mods:identifier>
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<mods:identifier id="5FFEAB21C4F24B2321AA1CC2A393C9C3" type="ISSN">2284-4880</mods:identifier>
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<mods:identifier id="CD9D2168DD91ADC236264C93096395A3" type="ZooBank">B3F75A2C-7603-4027-B8E1-F4A8B5A0CDD9</mods:identifier>
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</mods:mods>
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<treatment id="0F747550111DFFAAFF16AC251A2E91C8" LSID="urn:lsid:plazi:treatment:0F747550111DFFAAFF16AC251A2E91C8" httpUri="http://treatment.plazi.org/id/0F747550111DFFAAFF16AC251A2E91C8" lastPageId="72" lastPageNumber="177" pageId="71" pageNumber="176">
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<subSubSection id="CFC797CD111DFFA5FF16AC2518E89108" box="[136,450,788,812]" pageId="71" pageNumber="176" type="nomenclature">
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<paragraph id="8762C446111DFFA5FF16AC2518E89108" blockId="71.[136,450,788,812]" box="[136,450,788,812]" pageId="71" pageNumber="176">
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<heading id="DC2A732A111DFFA5FF16AC2518E89108" box="[136,450,788,812]" fontSize="10" level="2" pageId="71" pageNumber="176" reason="5">
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<taxonomicName id="40DDBFC5111DFFA5FF16AC25184F9108" ID-CoL="8SSSD" authorityName="Ballerio" authorityYear="2021" box="[136,357,788,812]" class="Insecta" family="Hybosoridae" genus="Goudotostes" kingdom="Animalia" order="Coleoptera" pageId="71" pageNumber="176" phylum="Arthropoda" rank="species" species="elegans" status="sp. nov.">
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||
<emphasis id="B5A91854111DFFA5FF16AC25184F9108" box="[136,357,788,812]" italics="true" pageId="71" pageNumber="176">Goudotostes elegans</emphasis>
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</taxonomicName>
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<emphasis id="B5A91854111DFFA5FEF2AC2518E89108" bold="true" box="[364,450,788,812]" pageId="71" pageNumber="176">
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<taxonomicNameLabel id="AE9AA52F111DFFA5FEF2AC2518E89108" box="[364,450,788,812]" pageId="71" pageNumber="176" rank="species">sp. nov.</taxonomicNameLabel>
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||
</emphasis>
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||
</heading>
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||
</paragraph>
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||
</subSubSection>
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||
<subSubSection id="CFC797CD111DFFA5FF16AC051B2391A8" pageId="71" pageNumber="176" type="description">
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<paragraph id="8762C446111DFFA5FF16AC051B2391A8" blockId="71.[136,756,820,908]" pageId="71" pageNumber="176">
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(
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<figureCitation id="1FE6D8C3111DFFA5FF0EAC0519CB9168" box="[144,225,820,844]" captionStart="Fig" captionStartId="128.[151,182,1858,1877]" captionTargetBox="[110,1457,225,1815]" captionTargetId="figure-13@128.[102,1486,151,1845]" captionTargetPageId="128" captionText="Fig. 63 – Goudotostes elegans sp. nov., a, holotype, dorsal view of distended specimen; b, paratype, ventral view of rolled up specimen; c, paratype, dorsal view of rolled up specimen; d, paratype, lateral view of rolled up specimen; e, specimen excluded from type series from Ranomafana, dorsal view of rolled up specimen; f, holotype, spiculum gastrale; g-j, holotype, four views of aedeagus." figureDoi="http://doi.org/10.5281/zenodo.12764307" httpUri="https://zenodo.org/record/12764307/files/figure.png" pageId="71" pageNumber="176">Figs 63</figureCitation>
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a-j, 77) http://zoobank.org/
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||
<uri id="F34CC844111DFFA5FEFFAC651B2391A8" pageId="71" pageNumber="176">
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||
urn:lsid:zoobank.org:act:
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||
<uuid id="F37BFE93111DFFA5FDF0AC651B2391A8" pageId="71" pageNumber="176">269ED34F- 5B37-4A9D-AF0A-F0B32E397794</uuid>
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||
</uri>
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||
</paragraph>
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||
</subSubSection>
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||
<subSubSection id="CFC797CD111DFFA5FF30AC8518919468" pageId="71" pageNumber="176" type="materials_examined">
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||
<paragraph id="8762C446111DFFA5FF30AC8518919468" blockId="71.[136,757,948,1932]" pageId="71" pageNumber="176">
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||
<emphasis id="B5A91854111DFFA5FF30AC85181C91E8" bold="true" box="[174,310,948,972]" pageId="71" pageNumber="176">Type series.</emphasis>
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||
<materialsCitation id="37B5CE1B111DFFA5FEA0AC8519D396C8" collectingDate="2003-02-05" collectingDateMax="2003-02-09" collectingDateMin="2003-02-05" collectorName="San Francisco, CA & Fisher & Griswold" country="Madagascar" county="Academy of Sciences" determinerName="A. Ballerio" elevation="770" latitude="-22.843334" location="Sakaraha" longLatPrecision="21" longitude="44.71" municipality="Parc National de Zombitse" pageId="71" pageNumber="176" specimenCount="1" specimenCount-male="1" stateProvince="Toliara" typeStatus="holotype">
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||
<typeStatus id="58667AE4111DFFA5FEA0AC85188C91E8" box="[318,422,948,972]" pageId="71" pageNumber="176" type="holotype">Holotype</typeStatus>
|
||
,
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||
<specimenCount id="91DB0FCF111DFFA5FE2CAC8418E291E8" box="[434,456,949,972]" count="1" pageId="71" pageNumber="176" type="male">♂</specimenCount>
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||
, deposited in the collection of the California
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||
<collectingCounty id="6E03BCCA111DFFA5FEC9ACE51B7991C8" box="[343,595,980,1004]" pageId="71" pageNumber="176">Academy of Sciences</collectingCounty>
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||
,
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||
<collectorName id="2A28A190111DFFA5FDFBACE519F09628" pageId="71" pageNumber="176">San Francisco, CA</collectorName>
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||
, USA, labelled as follows: / CASTYPE 20264 / CASENT 8135298 /
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||
<collectingCountry id="FFCA84D6111DFFA5FEF0AB2518DC9608" box="[366,502,1044,1068]" name="Madagascar" pageId="71" pageNumber="176">Madagascar</collectingCountry>
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||
:
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||
<collectingRegion id="45190AA4111DFFA5FD9EAB251BA79608" box="[512,653,1044,1068]" country="Madagascar" name="Toliara" pageId="71" pageNumber="176">Toliara Prov.</collectingRegion>
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||
<collectingMunicipality id="67065E3C111DFFA5FD0AAB24187D9668" pageId="71" pageNumber="176">Parc National de Zombitse</collectingMunicipality>
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||
<quantity id="402569A3111DFFA5FEC1AB0518939668" box="[351,441,1076,1100]" metricMagnitude="4" metricUnit="m" metricValue="1.98" pageId="71" pageNumber="176" unit="km" value="19.8">19.8 km</quantity>
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||
<geoCoordinate id="E2E9A281111DFFA5FE5FAB0518D59668" box="[449,511,1076,1100]" degrees="84" direction="east" orientation="longitude" pageId="71" pageNumber="176" precision="55555" value="84.0">84° E</geoCoordinate>
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||
<location id="8202929D111DFFA5FD96AB051B469668" LSID="urn:lsid:plazi:treatment:0F747550111DFFAAFF16AC251A2E91C8:8202929D111DFFA5FD96AB051B469668" box="[520,620,1076,1100]" country="Madagascar" county="Academy of Sciences" latitude="-22.843334" longLatPrecision="21" longitude="44.71" municipality="Parc National de Zombitse" name="Sakaraha" pageId="71" pageNumber="176" stateProvince="Toliara">Sakaraha</location>
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||
elev.
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||
<quantity id="402569A3111DFFA5FD30AB051BD99668" box="[686,755,1076,1100]" metricMagnitude="2" metricUnit="m" metricValue="7.7" pageId="71" pageNumber="176" unit="m" value="770.0">
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||
<elevation id="0CF02375111DFFA5FD30AB051BD99668" box="[686,755,1076,1100]" metricMagnitude="2" metricUnit="m" metricValue="7.7" pageId="71" pageNumber="176" unit="m" value="770.0">770 m</elevation>
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||
</quantity>
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||
<date id="F363E286111DFFA5FF17AB65180A9648" box="[137,288,1108,1132]" pageId="71" pageNumber="176" value="2003-02-05" valueMax="2003-02-09" valueMin="2003-02-05">
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||
<collectingDate id="E3271B6E111DFFA5FF17AB65180A9648" box="[137,288,1108,1132]" pageId="71" pageNumber="176" value="2003-02-05" valueMax="2003-02-09" valueMin="2003-02-05">5-9 Feb. 2003</collectingDate>
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||
</date>
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||
<geoCoordinate id="E2E9A281111DFFA5FEB9AB6518939648" box="[295,441,1108,1132]" degrees="22" direction="south" minutes="50" orientation="latitude" pageId="71" pageNumber="176" precision="15" seconds="36" value="-22.843334">22° 50’ 36” S</geoCoordinate>
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||
<geoCoordinate id="E2E9A281111DFFA5FE21AB651B4A9648" box="[447,608,1108,1132]" degrees="044" direction="east" minutes="42" orientation="longitude" pageId="71" pageNumber="176" precision="15" seconds="36" value="44.71">044° 42’ 36” E</geoCoordinate>
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||
/ coll.
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||
<collectorName id="2A28A190111DFFA5FD35AB651BDB9648" box="[683,753,1108,1132]" pageId="71" pageNumber="176">Fisher</collectorName>
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||
,
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||
<collectorName id="2A28A190111DFFA5FF16AB4519C796A8" box="[136,237,1140,1164]" pageId="71" pageNumber="176">Griswold</collectorName>
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||
et al.
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||
<location id="8202929D111DFFA5FEA8AB451B5896A8" LSID="urn:lsid:plazi:treatment:0F747550111DFFAAFF16AC251A2E91C8:8202929D111DFFA5FEA8AB451B5896A8" box="[310,626,1140,1164]" country="Madagascar" county="Academy of Sciences" latitude="-22.843334" longLatPrecision="21" longitude="44.71" municipality="Parc National de Zombitse" name="California Acad. of Sciences" pageId="71" pageNumber="176" stateProvince="Toliara">California Acad. of Sciences</location>
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sifted litter (leaf mold, rotten wood) in tropical dry forest collection code: BLF7510/ /
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||
<taxonomicName id="40DDBFC5111DFFA5FED0AB851B0596E8" authorityName="Ballerio" authorityYear="2021" box="[334,559,1204,1228]" class="Insecta" family="Hybosoridae" genus="Goudotostes" kingdom="Animalia" order="Coleoptera" pageId="71" pageNumber="176" phylum="Arthropoda" rank="species" species="elegans" status="sp. nov.">
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||
<emphasis id="B5A91854111DFFA5FED0AB851B0596E8" box="[334,559,1204,1228]" italics="true" pageId="71" pageNumber="176">Goudotostes elegans</emphasis>
|
||
</taxonomicName>
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||
<taxonomicNameLabel id="AE9AA52F111DFFA5FDA8AB841B5B96E8" box="[566,625,1205,1228]" pageId="71" pageNumber="176" rank="species">n. sp.</taxonomicNameLabel>
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||
det.
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||
<determinerName id="72BE2882111DFFA5FD3AAB85199096C8" pageId="71" pageNumber="176">A. Ballerio</determinerName>
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||
2020
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||
</materialsCitation>
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||
<typeStatus id="58667AE4111DFFA5FE9CABE5185996C8" box="[258,371,1236,1260]" pageId="71" pageNumber="176" type="holotype">Holotypus</typeStatus>
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||
<specimenCount id="91DB0FCF111DFFA5FEE2ABE418BA96C8" box="[380,400,1237,1260]" count="1" pageId="71" pageNumber="176" type="male">♂</specimenCount>
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/ [distended specimen glued on a card, dissected, male genitalia glued in DMHF resin on a separate card, same pin].
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<materialsCitation id="37B5CE1B111DFFA5FE75AA2418859748" collectingDate="2003-02-05" collectingDateMax="2003-02-09" collectingDateMin="2003-02-05" collectionCode="CASC, ABCB" collectorName="San Francisco, CA & Fisher & Griswold" country="Madagascar" county="Academy of Sciences" determinerName="A. Ballerio" elevation="770" latitude="-22.843334" location="Sakaraha" longLatPrecision="21" longitude="44.71" municipality="Parc National de Zombitse" pageId="71" pageNumber="176" specimenCount="6" specimenCount-female="2" specimenCount-male="4" stateProvince="Toliara" typeStatus="paratype">
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||
<typeStatus id="58667AE4111DFFA5FE75AA241B7F9709" box="[491,597,1301,1325]" pageId="71" pageNumber="176" type="paratype">Paratypes</typeStatus>
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||
[
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||
<specimenCount id="91DB0FCF111DFFA5FDF7AA251BBF9708" box="[617,661,1300,1324]" count="1" pageId="71" pageNumber="176" type="male">1 ♂</specimenCount>
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||
dissect- ed]:
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||
<specimenCount id="91DB0FCF111DFFA5FF24AA0519DD9768" box="[186,247,1332,1356]" count="3" pageId="71" pageNumber="176" type="male">3 ♂♂</specimenCount>
|
||
and
|
||
<specimenCount id="91DB0FCF111DFFA5FEB0AA0518419768" box="[302,363,1332,1356]" count="2" pageId="71" pageNumber="176" type="female">2 ♀♀</specimenCount>
|
||
(
|
||
<quantity id="402569A3111DFFA5FEE2AA05188C9768" box="[380,422,1332,1356]" metricMagnitude="-2" metricUnit="m" metricValue="7.62" pageId="71" pageNumber="176" unit="in" value="3.0">3 in</quantity>
|
||
<collectionCode id="E1CC5C83111DFFA5FE30AA0518DE9768" box="[430,500,1332,1356]" pageId="71" pageNumber="176">CASC</collectionCode>
|
||
and
|
||
<quantity id="402569A3111DFFA5FDB5AA051B7F9768" box="[555,597,1332,1356]" metricMagnitude="-2" metricUnit="m" metricValue="5.08" pageId="71" pageNumber="176" unit="in" value="2.0">2 in</quantity>
|
||
<collectionCode id="E1CC5C83111DFFA5FDC5AA051B839768" box="[603,681,1332,1356]" pageId="71" pageNumber="176">ABCB</collectionCode>
|
||
), same collecting data as holotype
|
||
</materialsCitation>
|
||
.
|
||
<emphasis id="B5A91854111DFFA5FE27AA6518C597A8" bold="true" pageId="71" pageNumber="176">Further material examined (excluded from the type series)</emphasis>
|
||
:
|
||
<materialsCitation id="37B5CE1B111DFFA5FE61AA45188F9468" collectingDate="2003-03-27" collectingDateMax="2003-03-31" collectingDateMin="2003-03-27" collectionCode="CASC, ABCB" collectorName="Fisher & Griswold" country="Madagascar" elevation="1100" latitude="-21.289999" location="California Acad. of Sciences" longLatPrecision="21" longitude="47.433334" municipality="Parc National de Ranomafana Vatoharanana River" pageId="71" pageNumber="176" specimenCount="3" specimenCount-female="3" stateProvince="Fianarantsoa">
|
||
<specimenCount id="91DB0FCF111DFFA5FE61AA451B1697A8" box="[511,572,1396,1420]" count="3" pageId="71" pageNumber="176" type="female">3 ♀♀</specimenCount>
|
||
(
|
||
<quantity id="402569A3111DFFA5FDD2AA451B5D97A8" box="[588,631,1396,1420]" metricMagnitude="-2" metricUnit="m" metricValue="5.08" pageId="71" pageNumber="176" unit="in" value="2.0">2 in</quantity>
|
||
<collectionCode id="E1CC5C83111DFFA5FDE1AA451BEF97A8" box="[639,709,1396,1420]" pageId="71" pageNumber="176">CASC</collectionCode>
|
||
and
|
||
<quantity id="402569A3111DFFA5FF16AAA5199F9788" box="[136,181,1428,1452]" metricMagnitude="-2" metricUnit="m" metricValue="2.54" pageId="71" pageNumber="176" unit="in" value="1.0">1 in</quantity>
|
||
<collectionCode id="E1CC5C83111DFFA5FF23AAA518269788" box="[189,268,1428,1452]" pageId="71" pageNumber="176">ABCB</collectionCode>
|
||
): /
|
||
<collectingCountry id="FFCA84D6111DFFA5FEADAAA518919788" box="[307,443,1428,1452]" name="Madagascar" pageId="71" pageNumber="176">Madagascar</collectingCountry>
|
||
:
|
||
<collectingRegion id="45190AA4111DFFA5FE59AAA51B789788" box="[455,594,1428,1452]" country="Madagascar" name="Fianarantsoa" pageId="71" pageNumber="176">Fianarantsoa</collectingRegion>
|
||
<collectingMunicipality id="67065E3C111DFFA5FDC2AAA41B0B97E8" pageId="71" pageNumber="176">Parc National de Ranomafana Vatoharanana River</collectingMunicipality>
|
||
<locationDeviation id="96D0DD64111DFFA5FDB0AA85183B97C8" location="Ranomafana" pageId="71" pageNumber="176">
|
||
<quantity id="402569A3111DFFA5FDB0AA851B5597E8" box="[558,639,1460,1484]" metricMagnitude="3" metricUnit="m" metricValue="4.1" pageId="71" pageNumber="176" unit="km" value="4.1">4.1 km</quantity>
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||
231° SW Ranomafana
|
||
</locationDeviation>
|
||
elev.
|
||
<quantity id="402569A3111DFFA5FEC3AAE5189997C8" box="[349,435,1492,1516]" metricMagnitude="3" metricUnit="m" metricValue="1.1" pageId="71" pageNumber="176" unit="m" value="1100.0">
|
||
<elevation id="0CF02375111DFFA5FEC3AAE5189997C8" box="[349,435,1492,1516]" metricMagnitude="3" metricUnit="m" metricValue="1.1" pageId="71" pageNumber="176" unit="m" value="1100.0">1100 m</elevation>
|
||
</quantity>
|
||
<date id="F363E286111DFFA5FE21AAE51BAA97C8" box="[447,640,1492,1516]" pageId="71" pageNumber="176" value="2003-03-27" valueMax="2003-03-31" valueMin="2003-03-27">
|
||
<collectingDate id="E3271B6E111DFFA5FE21AAE51BAA97C8" box="[447,640,1492,1516]" pageId="71" pageNumber="176" value="2003-03-27" valueMax="2003-03-31" valueMin="2003-03-27">27-31 Mar. 2003</collectingDate>
|
||
</date>
|
||
/
|
||
<geoCoordinate id="E2E9A281111DFFA5FD3EAAE519EC9428" degrees="21" direction="south" minutes="17" orientation="latitude" pageId="71" pageNumber="176" precision="15" seconds="24" value="-21.289999">21° 17’ 24” S</geoCoordinate>
|
||
<geoCoordinate id="E2E9A281111DFFA5FF53AAC518459428" box="[205,367,1524,1548]" degrees="047" direction="east" minutes="26" orientation="longitude" pageId="71" pageNumber="176" precision="15" seconds="00" value="47.433334">047° 26’ 00” E</geoCoordinate>
|
||
/
|
||
<location id="8202929D111DFFA5FE1BAAC51B979428" LSID="urn:lsid:plazi:treatment:0F747550111DFFAAFF16AC251A2E91C8:8202929D111DFFA5FE1BAAC51B979428" box="[389,701,1524,1548]" country="Madagascar" latitude="-21.289999" longLatPrecision="21" longitude="47.433334" municipality="Parc National de Ranomafana Vatoharanana River" name="California Acad. of Sciences" pageId="71" pageNumber="176" stateProvince="Fianarantsoa">California Acad. of Sciences</location>
|
||
coll.
|
||
<collectorName id="2A28A190111DFFA5FF16A92519E49408" box="[136,206,1556,1580]" pageId="71" pageNumber="176">Fisher</collectorName>
|
||
,
|
||
<collectorName id="2A28A190111DFFA5FF40A92518699408" box="[222,323,1556,1580]" pageId="71" pageNumber="176">Griswold</collectorName>
|
||
et al. sifted litter, montane rainforest collection code: BLF8400
|
||
</materialsCitation>
|
||
/.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="CFC797CD111DFFA5FF30A9651D3D9328" pageId="71" pageNumber="176" type="diagnosis">
|
||
<paragraph id="8762C446111DFFA5FF30A9651D3D9328" blockId="71.[136,757,948,1932]" lastBlockId="71.[816,1437,180,1932]" pageId="71" pageNumber="176">
|
||
<emphasis id="B5A91854111DFFA5FF30A96518099448" bold="true" box="[174,291,1620,1644]" pageId="71" pageNumber="176">Diagnosis.</emphasis>
|
||
<taxonomicName id="40DDBFC5111DFFA5FEB7A9651B2C9448" authorityName="Ballerio" authorityYear="2021" box="[297,518,1620,1644]" class="Insecta" family="Hybosoridae" genus="Goudotostes" kingdom="Animalia" order="Coleoptera" pageId="71" pageNumber="176" phylum="Arthropoda" rank="species" species="elegans" status="sp. nov.">
|
||
<emphasis id="B5A91854111DFFA5FEB7A9651B2C9448" box="[297,518,1620,1644]" italics="true" pageId="71" pageNumber="176">Goudotostes elegans</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B5A91854111DFFA5FD90A9651B499448" bold="true" box="[526,611,1620,1644]" pageId="71" pageNumber="176">
|
||
<taxonomicNameLabel id="AE9AA52F111DFFA5FD90A9651B499448" box="[526,611,1620,1644]" pageId="71" pageNumber="176" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
can be easily differentiated from all other species of
|
||
<taxonomicName id="40DDBFC5111DFFA5FDB8A9451B8194A8" authorityName=", sensu Ballerio" authorityYear="2008" box="[550,683,1652,1676]" class="Insecta" family="Hybosoridae" genus="Goudotostes" kingdom="Animalia" order="Coleoptera" pageId="71" pageNumber="176" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B5A91854111DFFA5FDB8A9451B8194A8" box="[550,683,1652,1676]" italics="true" pageId="71" pageNumber="176">Goudotostes</emphasis>
|
||
</taxonomicName>
|
||
by the following combination of characters: a) small size, b) dorsal ocular area absent, c) pronotal anterior carina monolobate, d) pronotal basal carina monolobate, e) each elytron with one median longitudinal continuous carina occupying about three quarters of elytral length, f) aedeagus with distinctly hooked left paramere. Because of the peculiar elytral carination, the only species with which
|
||
<taxonomicName id="40DDBFC5111DFFA5FDF1A86519F395A8" authorityName="Ballerio" authorityYear="2021" class="Insecta" family="Hybosoridae" genus="Goudotostes" kingdom="Animalia" order="Coleoptera" pageId="71" pageNumber="176" phylum="Arthropoda" rank="species" species="elegans" status="sp. nov.">
|
||
<emphasis id="B5A91854111DFFA5FDF1A86519F395A8" italics="true" pageId="71" pageNumber="176">Goudotostes elegans</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B5A91854111DFFA5FF7DA845181195A8" bold="true" box="[227,315,1908,1932]" pageId="71" pageNumber="176">
|
||
<taxonomicNameLabel id="AE9AA52F111DFFA5FF7DA845181195A8" box="[227,315,1908,1932]" pageId="71" pageNumber="176" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
can be mistaken is
|
||
<taxonomicName id="40DDBFC5111DFFA5FD85A8451BDE95A8" authorityName="Ballerio" authorityYear="2021" box="[539,756,1908,1932]" class="Insecta" family="Hybosoridae" genus="Goudotostes" kingdom="Animalia" order="Coleoptera" pageId="71" pageNumber="176" phylum="Arthropoda" rank="species" species="angelii" status="sp. nov.">
|
||
<emphasis id="B5A91854111DFFA5FD85A8451BDE95A8" box="[539,756,1908,1932]" italics="true" pageId="71" pageNumber="176">Goudotostes angelii</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B5A91854111DFFA5FCAEAF851AAD92E8" bold="true" box="[816,903,180,204]" pageId="71" pageNumber="176">
|
||
<taxonomicNameLabel id="AE9AA52F111DFFA5FCAEAF851AAD92E8" box="[816,903,180,204]" pageId="71" pageNumber="176" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, which however is larger and which has bilobate basal and anterior pronotal carina as well as a small vestigial dorsal ocular area.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="CFC797CD111DFFAAFCC8AE2518889028" lastPageId="72" lastPageNumber="177" pageId="71" pageNumber="176" type="description">
|
||
<paragraph id="8762C446111DFFAAFCC8AE2518EA9328" blockId="71.[816,1437,180,1932]" lastBlockId="72.[151,773,180,1004]" lastPageId="72" lastPageNumber="177" pageId="71" pageNumber="176">
|
||
<emphasis id="B5A91854111DFFA5FCC8AE251AF79308" bold="true" box="[854,989,276,300]" pageId="71" pageNumber="176">Description.</emphasis>
|
||
Size: HL =
|
||
<quantity id="402569A3111DFFA5FBFEAE251D9E9308" box="[1120,1204,276,300]" metricMagnitude="-4" metricUnit="m" metricValue="8.0" pageId="71" pageNumber="176" unit="mm" value="0.8">0.8 mm</quantity>
|
||
; HW =
|
||
<quantity id="402569A3111DFFA5FA97AE251C779308" box="[1289,1373,276,300]" metricMagnitude="-3" metricUnit="m" metricValue="1.1" pageId="71" pageNumber="176" unit="mm" value="1.1">1.1 mm</quantity>
|
||
; PL =
|
||
<quantity id="402569A3111DFFA5FCAEAE051AA99368" box="[816,899,308,332]" metricMagnitude="-3" metricUnit="m" metricValue="1.2" pageId="71" pageNumber="176" unit="mm" value="1.2">1.2 mm</quantity>
|
||
; PW =
|
||
<quantity id="402569A3111DFFA5FC50AE051D0B9368" box="[974,1057,308,332]" metricMagnitude="-3" metricUnit="m" metricValue="2.1" pageId="71" pageNumber="176" unit="mm" value="2.1">2.1 mm</quantity>
|
||
; EL =
|
||
<quantity id="402569A3111DFFA5FBFAAE051D9D9368" box="[1124,1207,308,332]" metricMagnitude="-3" metricUnit="m" metricValue="2.2" pageId="71" pageNumber="176" unit="mm" value="2.2">2.2 mm</quantity>
|
||
; EW = 2.0 mm. Overall morphology as in generic diagnosis. Small-sized
|
||
<taxonomicName id="40DDBFC5111DFFA5FAD2AE651A4193A8" authorityName=", sensu Ballerio" authorityYear="2008" class="Insecta" family="Hybosoridae" genus="Goudotostes" kingdom="Animalia" order="Coleoptera" pageId="71" pageNumber="176" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B5A91854111DFFA5FAD2AE651A4193A8" italics="true" pageId="71" pageNumber="176">Goudotostes</emphasis>
|
||
</taxonomicName>
|
||
. Body convex. Black, shiny, dorsum with extremely short fine setation (50×), underside, tarsi and antennae dark reddish-brown, setation yellowish-brown.
|
||
<emphasis id="B5A91854111DFFA5FB70AE851C0D93E8" box="[1262,1319,436,460]" italics="true" pageId="71" pageNumber="176">Head</emphasis>
|
||
: W/L ratio = 1.58; clypeal apex obtuse, apical projection developed and sexually dimorphic, sides relatively rounded with slightly unaligned genae; dorsal ocular area absent, genal canthus indistinct, ventral ocular area small-sized, genae acute, distinctly protruding outwards, genal suture indistinct; head dorsal surface with vertex covered by large impressed simple punctures becoming transversely comma-shaped on frons and clypeus, fore margin marked by some irregular transverse lines. Interpunctural distance of punctures equal to or inferior than punctural diameter. Frons bearing a transverse row of three weakly raised tubercles.
|
||
<emphasis id="B5A91854111DFFA5FAB3AC241CBF9108" box="[1325,1429,789,812]" italics="true" pageId="71" pageNumber="176">Pronotum</emphasis>
|
||
: subrectangular (dorsal view) (W/L ratio = 1.67), convex, fore angles broadly subtruncate; pronotal disc covered by short large impressed transverse comma-shaped punctures becoming smaller horseshoe-shaped with opening directed outwards at sides and base, each one bearing a fine setigerous pore inside, interpunctural distance about equal to punctural diameter; pronotal bead complete although not visible at base; pronotal base distinctly raised in correspondence with scutellum; pronotal anterior carina monolobate; pronotal sculpturing made of large and raised tubercles and carinae as follows: posterior carina monolobate, disc with two raised tubercles, each side with one raised tubercle and one transverse carina, anterior monolobate carina distinctly raised with truncate apex.
|
||
<taxonomicName id="40DDBFC5111DFFA5FBDDABE51D8096C8" box="[1091,1194,1236,1260]" class="Trilobita" family="Styginidae" genus="Scutellum" kingdom="Animalia" order="Corynexochida" pageId="71" pageNumber="176" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B5A91854111DFFA5FBDDABE51D8096C8" box="[1091,1194,1236,1260]" italics="true" pageId="71" pageNumber="176">Scutellum</emphasis>
|
||
</taxonomicName>
|
||
: covered by impressed transverse comma-shaped punctures, apical third and sides smooth.
|
||
<emphasis id="B5A91854111DFFA5FC12AA251AE79708" box="[908,973,1300,1324]" italics="true" pageId="71" pageNumber="176">Elytra</emphasis>
|
||
: longer than wide (W/L ratio = 0.86), subovoidal (dorsal view), convex; inferior sutural stria visible and complete; humeral callus marked by a tubercle; elytral surface uniformly covered by large horseshoe-shaped punctures with opening directed backwards and having a setigerous pore inside, interpunctural distance about equal to punctural diameter. Striated articular area hardly visible in lateral view, relatively narrow and short; marginal area medium-sized; elytral articular process well developed, smooth and shiny, isolated from remaining elytral surface by a deep furrow. Elytral sculpturing as follows: base with one raised large oblique tubercle near apex of scutellum (parasutural tubercle), next to it a humeral oblique tubercle (humeral tubercle); almost parallel to elytral suture and aligned with the tubercle near scutellar apex a slightly sinuated longitudinal carina with smooth edge, starting just after the tubercle and ending before elytral apical third; elytral sides with three irregular rows of short longitudinal carinae, outer row with four carinae, inner rows with a lesser number of carinae; elytral distal third with some weakly raised irregular tubercles.
|
||
<emphasis id="B5A918541112FFAAFE9BAF84186F92E8" box="[261,325,181,204]" italics="true" pageId="72" pageNumber="177">Wings</emphasis>
|
||
: flightless.
|
||
<emphasis id="B5A918541112FFAAFE5FAF851B7792E8" box="[449,605,180,204]" italics="true" pageId="72" pageNumber="177">Male genitalia</emphasis>
|
||
: aedeagus (L =
|
||
<quantity id="402569A31112FFAAFF07AFE519C092C8" box="[153,234,212,236]" metricMagnitude="-4" metricUnit="m" metricValue="8.0" pageId="72" pageNumber="177" unit="mm" value="0.8">0.8 mm</quantity>
|
||
) with parameres as in
|
||
<figureCitation id="1FE6D8C31112FFAAFE4CAFE51B3592C8" box="[466,543,212,236]" captionStart="Fig" captionStartId="128.[151,182,1858,1877]" captionTargetBox="[110,1457,225,1815]" captionTargetId="figure-13@128.[102,1486,151,1845]" captionTargetPageId="128" captionText="Fig. 63 – Goudotostes elegans sp. nov., a, holotype, dorsal view of distended specimen; b, paratype, ventral view of rolled up specimen; c, paratype, dorsal view of rolled up specimen; d, paratype, lateral view of rolled up specimen; e, specimen excluded from type series from Ranomafana, dorsal view of rolled up specimen; f, holotype, spiculum gastrale; g-j, holotype, four views of aedeagus." figureDoi="http://doi.org/10.5281/zenodo.12764307" httpUri="https://zenodo.org/record/12764307/files/figure.png" pageId="72" pageNumber="177">Figs 63</figureCitation>
|
||
g-j, spiculum gastrale (L =
|
||
<quantity id="402569A31112FFAAFF54AFC518369328" box="[202,284,244,268]" metricMagnitude="-4" metricUnit="m" metricValue="7.0" pageId="72" pageNumber="177" unit="mm" value="0.7">0.7 mm</quantity>
|
||
) as in
|
||
<figureCitation id="1FE6D8C31112FFAAFEFEAFC518969328" box="[352,444,244,268]" captionStart="Fig" captionStartId="128.[151,182,1858,1877]" captionTargetBox="[110,1457,225,1815]" captionTargetId="figure-13@128.[102,1486,151,1845]" captionTargetPageId="128" captionText="Fig. 63 – Goudotostes elegans sp. nov., a, holotype, dorsal view of distended specimen; b, paratype, ventral view of rolled up specimen; c, paratype, dorsal view of rolled up specimen; d, paratype, lateral view of rolled up specimen; e, specimen excluded from type series from Ranomafana, dorsal view of rolled up specimen; f, holotype, spiculum gastrale; g-j, holotype, four views of aedeagus." figureDoi="http://doi.org/10.5281/zenodo.12764307" httpUri="https://zenodo.org/record/12764307/files/figure.png" pageId="72" pageNumber="177">
|
||
Fig.
|
||
<specimenCount id="91DB0FCF1112FFAAFE11AFC518969328" box="[399,444,244,268]" count="63" pageId="72" pageNumber="177" type="generic">63 f</specimenCount>
|
||
</figureCitation>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8762C4461112FFAAFF23AE2518889028" blockId="72.[151,773,180,1004]" pageId="72" pageNumber="177">
|
||
<emphasis id="B5A918541112FFAAFF23AE2518119308" bold="true" box="[189,315,276,300]" pageId="72" pageNumber="177">Variability.</emphasis>
|
||
I attribute to this species (without including them in the
|
||
<typeStatus id="58667AE41112FFAAFEE6AE04188C9369" box="[376,422,309,333]" pageId="72" pageNumber="177">type</typeStatus>
|
||
series) also
|
||
<specimenCount id="91DB0FCF1112FFAAFDAAAE051BEC9368" box="[564,710,308,332]" count="3" pageId="72" pageNumber="177" type="female">three females</specimenCount>
|
||
from Ranomafana, some
|
||
<quantity id="402569A31112FFAAFEF6AE6518929348" box="[360,440,340,364]" metricMagnitude="5" metricUnit="m" metricValue="3.0" pageId="72" pageNumber="177" unit="km" value="300.0">300 km</quantity>
|
||
East of the
|
||
<typeStatus id="58667AE41112FFAAFDAAAE641B489349" box="[564,610,341,365]" pageId="72" pageNumber="177">type</typeStatus>
|
||
locality. Those
|
||
<specimenCount id="91DB0FCF1112FFAAFF09AE45186293A8" box="[151,328,372,396]" count="3" pageId="72" pageNumber="177" type="generic">three specimens</specimenCount>
|
||
display slightly stronger elytral sculpturing (carinae and tubercles) and slightly denser elytral punctuation (
|
||
<figureCitation id="1FE6D8C31112FFAAFEBAAE8518AC93E8" box="[292,390,436,460]" captionStart="Fig" captionStartId="128.[151,182,1858,1877]" captionTargetBox="[110,1457,225,1815]" captionTargetId="figure-13@128.[102,1486,151,1845]" captionTargetPageId="128" captionText="Fig. 63 – Goudotostes elegans sp. nov., a, holotype, dorsal view of distended specimen; b, paratype, ventral view of rolled up specimen; c, paratype, dorsal view of rolled up specimen; d, paratype, lateral view of rolled up specimen; e, specimen excluded from type series from Ranomafana, dorsal view of rolled up specimen; f, holotype, spiculum gastrale; g-j, holotype, four views of aedeagus." figureDoi="http://doi.org/10.5281/zenodo.12764307" httpUri="https://zenodo.org/record/12764307/files/figure.png" pageId="72" pageNumber="177">Fig. 63 e</figureCitation>
|
||
). Only the finding of males and the examination of the aedeagus could reveal if they represent a separate species or not.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="CFC797CD1112FFAAFF23AD2518489048" pageId="72" pageNumber="177" type="etymology">
|
||
<paragraph id="8762C4461112FFAAFF23AD2518489048" blockId="72.[151,773,180,1004]" pageId="72" pageNumber="177">
|
||
<emphasis id="B5A918541112FFAAFF23AD2518149008" bold="true" box="[189,318,532,556]" pageId="72" pageNumber="177">Etymology.</emphasis>
|
||
Latin adjective in the nominative singular, meaning “elegant” due to elegant geometric pattern of the elytral sculpturing.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="CFC797CD1112FFAAFF23AD451A2E91C8" pageId="72" pageNumber="177" type="distribution">
|
||
<paragraph id="8762C4461112FFAAFF23AD451A2E91C8" blockId="72.[151,773,180,1004]" pageId="72" pageNumber="177">
|
||
<emphasis id="B5A918541112FFAAFF23AD4518F190A8" bold="true" box="[189,475,628,652]" pageId="72" pageNumber="177">Distribution and habitat.</emphasis>
|
||
Known from two localities in southern
|
||
<collectingCountry id="FFCA84D61112FFAAFE83ADA5188A9088" box="[285,416,660,684]" name="Madagascar" pageId="72" pageNumber="177">Madagascar</collectingCountry>
|
||
. The
|
||
<typeStatus id="58667AE41112FFAAFE7FADA41B259089" box="[481,527,661,685]" pageId="72" pageNumber="177">type</typeStatus>
|
||
locality is in western
|
||
<collectingCountry id="FFCA84D61112FFAAFF09AD85183290E8" box="[151,280,692,716]" name="Madagascar" pageId="72" pageNumber="177">Madagascar</collectingCountry>
|
||
and the
|
||
<typeStatus id="58667AE41112FFAAFE1DAD84189B90E9" box="[387,433,693,717]" pageId="72" pageNumber="177">type</typeStatus>
|
||
series was there collected by sifting litter (leaf mold, rotten wood) in tropical dry forest. For more details on the
|
||
<typeStatus id="58667AE41112FFAAFE5DADC418DB9129" box="[451,497,757,781]" pageId="72" pageNumber="177">type</typeStatus>
|
||
locality see under
|
||
<taxonomicName id="40DDBFC51112FFAAFD23ADC4187E9108" authorityName="Ballerio" authorityYear="2021" class="Insecta" family="Hybosoridae" genus="Synarmostes" kingdom="Animalia" order="Coleoptera" pageId="72" pageNumber="177" phylum="Arthropoda" rank="species" species="separatus" status="sp. nov.">
|
||
<emphasis id="B5A918541112FFAAFD23ADC4187E9108" italics="true" pageId="72" pageNumber="177">Synarmostes separatus</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B5A918541112FFAAFEFEAC2518919108" bold="true" box="[352,443,788,812]" pageId="72" pageNumber="177">
|
||
<taxonomicNameLabel id="AE9AA52F1112FFAAFEFEAC2518919108" box="[352,443,788,812]" pageId="72" pageNumber="177" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
. The
|
||
<specimenCount id="91DB0FCF1112FFAAFD9CAC251BBD9108" box="[514,663,788,812]" count="3" pageId="72" pageNumber="177" type="female">three females</specimenCount>
|
||
excluded from the
|
||
<typeStatus id="58667AE41112FFAAFF62AC0418009169" box="[252,298,821,845]" pageId="72" pageNumber="177">type</typeStatus>
|
||
series from Ranomafana (the exact collecting locality falls within the protected area “Parc National de Ranomafana”), if confirmed to belog to
|
||
<taxonomicName id="40DDBFC51112FFAAFDF7AC441BF191A8" authorityName="Ballerio" authorityYear="2021" box="[617,731,884,908]" class="Insecta" family="Hybosoridae" genus="Goudotostes" kingdom="Animalia" order="Coleoptera" pageId="72" pageNumber="177" phylum="Arthropoda" rank="species" species="elegans" status="sp. nov.">
|
||
<emphasis id="B5A918541112FFAAFDF7AC441BF191A8" box="[617,731,884,908]" italics="true" pageId="72" pageNumber="177">G. elegans</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B5A918541112FFAAFD7DAC4519EC9188" bold="true" pageId="72" pageNumber="177">
|
||
<taxonomicNameLabel id="AE9AA52F1112FFAAFD7DAC4519EC9188" pageId="72" pageNumber="177" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, would extend the ecological range of the species to “medium altitude moist evergreen forest” (
|
||
<bibRefCitation id="E34CB9B71112FFAAFDC0AC8519E491C8" author="Goodman S. M. & Raherilalao M. J. & Wohlhauser S." pageId="72" pageNumber="177" refId="ref102713" refString="Goodman S. M., Raherilalao M. J., Wohlhauser S. (eds). 2018. Les aires potegees terrestres de Madagascar: leur histoire, description et biote / The terrestrial protected areas of Madagascar: their history, description and biota. Association Vahatra, Antananarivo. 1716 pp." type="book" year="2018">Goodman et al. 2018</bibRefCitation>
|
||
) also known as “humid forest” (
|
||
<bibRefCitation id="E34CB9B71112FFAAFD81ACE51BD291C8" author="Moat J. & Smith P." box="[543,760,980,1004]" pageId="72" pageNumber="177" refId="ref103114" refString="Moat J., Smith P. 2007. Atlas of the vegetation of Madagascar. Royal Botanic Garden, Kew 124 pp." type="book" year="2007">Moat & Smith 2007</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |