208 lines
26 KiB
XML
208 lines
26 KiB
XML
<document id="8DD4F90E6CF608955EC6241AF55F1216" ID-DOI="10.13133/2284-4880/545" ID-ISSN="2284-4880" ID-Zenodo-Dep="12764154" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1721242503917" checkinUser="felipe" docAuthor="Ballerio, Alberto" docDate="2021" docId="0F747550110BFFB0FF16AB85180890A8" docLanguage="en" docName="FragEntomol.53.2.105-282.pdf" docOrigin="Fragmenta entomologica 53 (2)" docSource="http://dx.doi.org/10.13133/2284-4880/545" docStyle="DocumentStyle:8499FF4308F512386C0E43DC290A16BD.1:FragEntomol.2021-.journal_article.0cover" docStyleId="8499FF4308F512386C0E43DC290A16BD" docStyleName="FragEntomol.2021-.journal_article.0cover" docStyleVersion="1" docTitle="Goudotostes montanellus Ballerio, 2021, sp. nov." docType="treatment" docUuid="595C4F15-CF0F-4E4E-AD49-850FEC36F509" docUuidSource="ZooBank" docVersion="3" lastPageNumber="187" masterDocId="F34D0D28115AFFE2FF9EAF31192A9224" masterDocTitle="The Ceratocanthinae of Madagascar and Comoro Islands: a revision of the genera Synarmostes and Goudotostes, and of the flightless Philharmostes, with description of 64 new species (Coleoptera: Scarabaeoidea, Hybosoridae)" masterLastPageNumber="282" masterPageNumber="105" pageNumber="186" updateTime="1721247602959" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-NC-4.0">
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<mods:titleInfo id="D9BD28581B777E3ECC4209675F643FF5">
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<mods:title id="7CDB40A6C5212385EA20DA69C083EFB9">The Ceratocanthinae of Madagascar and Comoro Islands: a revision of the genera Synarmostes and Goudotostes, and of the flightless Philharmostes, with description of 64 new species (Coleoptera: Scarabaeoidea, Hybosoridae)</mods:title>
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<mods:name id="A4ABE4063E3C4A69B27B63E26AB66908" type="personal">
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<mods:roleTerm id="F09A2881AE44A29B3C2DE87B285D9CD7">Author</mods:roleTerm>
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<mods:namePart id="B371CB68556BAE6A5FA99F63F069BBAC">Ballerio, Alberto</mods:namePart>
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<mods:typeOfResource id="E62B397B6853BB3E83E2736348960C29">text</mods:typeOfResource>
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<mods:title id="E0E4EC028DDADC00E4741FAD0DBE36AA">Fragmenta entomologica</mods:title>
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</mods:titleInfo>
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<mods:part id="DEF180566E1DDE1B453932BEC1C2B3CE">
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<mods:date id="12326402981DDB69D0E69F36861DB200">2021</mods:date>
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<mods:number id="207AA8113D60B5B805C41D43BDD4EF06">2021-11-30</mods:number>
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<mods:number id="A813BA04A1D080C655594BD6A1D744AC">53</mods:number>
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<mods:detail id="D45FC16F787E04B664A0224AC95E080B" type="issue">
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<mods:number id="8E5C1B75F4BA978509D8CF62C766764F">2</mods:number>
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<mods:start id="3504D52168F8F64DE84C53B2D78220E0">105</mods:start>
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<mods:end id="F8034E95C3306CC43B8563C38152AA8B">282</mods:end>
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<mods:url id="14124C9650147771708165FBB351F121">http://dx.doi.org/10.13133/2284-4880/545</mods:url>
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<mods:classification id="8AC3F8460C9E53C785F2A11248351389">journal article</mods:classification>
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<mods:identifier id="F3D52470F982926403CD15568954C496" type="DOI">10.13133/2284-4880/545</mods:identifier>
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<mods:identifier id="BAECB5F2A25FCFF681A9B8F9534DE01F" type="ISSN">2284-4880</mods:identifier>
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<treatment id="0F747550110BFFB0FF16AB85180890A8" ID-DOI="http://doi.org/10.5281/zenodo.12764075" ID-Zenodo-Dep="12764075" LSID="urn:lsid:zoobank.org:act:595C4F15-CF0F-4E4E-AD49-850FEC36F509" httpUri="http://treatment.plazi.org/id/0F747550110BFFB0FF16AB85180890A8" lastPageId="82" lastPageNumber="187" pageId="81" pageNumber="186">
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<subSubSection id="CFC797CD110BFFB3FF16AB8518D996E8" box="[136,499,1204,1228]" pageId="81" pageNumber="186" type="nomenclature">
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<paragraph id="8762C446110BFFB3FF16AB8518D996E8" blockId="81.[136,499,1204,1228]" box="[136,499,1204,1228]" pageId="81" pageNumber="186">
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<heading id="DC2A732A110BFFB3FF16AB8518D996E8" box="[136,499,1204,1228]" fontSize="10" level="2" pageId="81" pageNumber="186" reason="5">
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<taxonomicName id="40DDBFC5110BFFB3FF16AB8518BC96E8" ID-CoL="8SSQT" authorityName="Ballerio" authorityYear="2021" box="[136,406,1204,1228]" class="Insecta" family="Hybosoridae" genus="Goudotostes" kingdom="Animalia" order="Coleoptera" pageId="81" pageNumber="186" phylum="Arthropoda" rank="species" species="montanellus" status="sp. nov.">
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<emphasis id="B5A91854110BFFB3FF16AB8518BC96E8" box="[136,406,1204,1228]" italics="true" pageId="81" pageNumber="186">Goudotostes montanellus</emphasis>
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</taxonomicName>
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<emphasis id="B5A91854110BFFB3FE03AB8518D996E8" bold="true" box="[413,499,1204,1228]" pageId="81" pageNumber="186">
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||
<taxonomicNameLabel id="AE9AA52F110BFFB3FE03AB8518D996E8" box="[413,499,1204,1228]" pageId="81" pageNumber="186" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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||
</heading>
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||
</paragraph>
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</subSubSection>
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<subSubSection id="CFC797CD110BFFB3FF16ABE51B2C9708" pageId="81" pageNumber="186" type="description">
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<paragraph id="8762C446110BFFB3FF16ABE51B2C9708" blockId="81.[136,755,1236,1324]" pageId="81" pageNumber="186">
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(
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<figureCitation id="1FE6D8C3110BFFB3FF0EABE519CB96C8" box="[144,225,1236,1260]" captionStart="Fig" captionStartId="110.[151,182,1858,1877]" captionTargetBox="[188,1394,177,1817]" captionTargetId="figure-13@110.[161,1424,160,1823]" captionTargetPageId="110" captionText="Fig. 45 – Goudotostes montanellus sp. nov., a, holotype, dorsal view of distended specimen; b, paratype, ventral view of rolled up specimen; c, paratype, dorsal view of rolled up specimen; d, paratype, lateral view of rolled up specimen; e, paratype, pronotum of rolled up specimen; f, holotype, spiculum gastrale; g–j, holotype, four views of aedeagus." figureDoi="http://doi.org/10.5281/zenodo.12764265" httpUri="https://zenodo.org/record/12764265/files/figure.png" pageId="81" pageNumber="186">Figs 45</figureCitation>
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a-j, 77) http://zoobank.org/
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<uri id="F34CC844110BFFB3FEFBABC51B2C9708" pageId="81" pageNumber="186">
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urn:lsid:zoobank.org:act:
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<uuid id="F37BFE93110BFFB3FDE9ABC51B2C9708" pageId="81" pageNumber="186">595C4F15- CF0F-4E4E-AD49-850FEC36F509</uuid>
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</uri>
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</paragraph>
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||
</subSubSection>
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||
<subSubSection id="CFC797CD110BFFB3FF30AA651B439488" pageId="81" pageNumber="186" type="materials_examined">
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<paragraph id="8762C446110BFFB3FF30AA651B439488" blockId="81.[136,757,1364,1932]" pageId="81" pageNumber="186">
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<emphasis id="B5A91854110BFFB3FF30AA65181F9748" bold="true" box="[174,309,1364,1388]" pageId="81" pageNumber="186">Type series.</emphasis>
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<materialsCitation id="37B5CE1B110BFFB3FEA0AA651B4394A8" collectingDate="2001-01-20" collectingDateMax="2001-01-26" collectingDateMin="2001-01-20" collectorName="San Francisco, CA & Fisher & Griswold" country="Madagascar" county="California Acad. of Sciences" elevation="925" latitude="-12.534445" location="Academy of Sciences" longLatPrecision="21" longitude="49.179447" municipality="d'Antisiranana Parc National Montagne d'Ambre" pageId="81" pageNumber="186" specimenCount="1" specimenCount-male="1" stateProvince="California" typeStatus="holotype">
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<typeStatus id="58667AE4110BFFB3FEA0AA65188E9748" box="[318,420,1364,1388]" pageId="81" pageNumber="186" type="holotype">Holotype</typeStatus>
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||
,
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<specimenCount id="91DB0FCF110BFFB3FE2FAA6418ED9748" box="[433,455,1365,1388]" count="1" pageId="81" pageNumber="186" type="male">♂</specimenCount>
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||
, deposited in the collection of the
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<collectingRegion id="45190AA4110BFFB3FF43AA45186397A8" box="[221,329,1396,1420]" country="United States of America" name="California" pageId="81" pageNumber="186">California</collectingRegion>
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<location id="8202929D110BFFB3FEC9AA451B7897A8" LSID="urn:lsid:plazi:treatment:0F747550110BFFB0FF16AB85180890A8:8202929D110BFFB3FEC9AA451B7897A8" box="[343,594,1396,1420]" country="Madagascar" county="California Acad. of Sciences" latitude="-12.534445" longLatPrecision="21" longitude="49.179447" municipality="d'Antisiranana Parc National Montagne d'Ambre" name="Academy of Sciences" pageId="81" pageNumber="186" stateProvince="California">Academy of Sciences</location>
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,
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<collectorName id="2A28A190110BFFB3FDFBAA4519F09788" pageId="81" pageNumber="186">San Francisco, CA</collectorName>
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, USA, labelled as follows: / CASTYPE 20273 / CASENT 8013658 /
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<collectingCountry id="FFCA84D6110BFFB3FEF0AA8518D997E8" box="[366,499,1460,1484]" name="Madagascar" pageId="81" pageNumber="186">Madagascar</collectingCountry>
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:
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<location id="8202929D110BFFB3FE60AA85189397C9" LSID="urn:lsid:plazi:treatment:0F747550110BFFB0FF16AB85180890A8:8202929D110BFFB3FE60AA85189397C9" country="Madagascar" county="California Acad. of Sciences" latitude="-12.534445" longLatPrecision="21" longitude="49.179447" municipality="d'Antisiranana Parc National Montagne d'Ambre" name="Province d'Antisiranana Parc National Montagne" pageId="81" pageNumber="186" stateProvince="California">
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Province
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<collectingMunicipality id="67065E3C110BFFB3FDF8AA85189397C9" pageId="81" pageNumber="186">d’Antisiranana Parc National Montagne</collectingMunicipality>
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</location>
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d’Ambre,
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<locationDeviation id="96D0DD64110BFFB3FDA8AAE519DF9428" location="Joffreville" pageId="81" pageNumber="186">
|
||
<quantity id="402569A3110BFFB3FDA8AAE51BAE97C8" box="[566,644,1492,1516]" metricMagnitude="3" metricUnit="m" metricValue="3.6" pageId="81" pageNumber="186" unit="km" value="3.6">3.6 km</quantity>
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235° SW Joffreville
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||
</locationDeviation>
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elev.
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<quantity id="402569A3110BFFB3FEADAAC5185C9428" box="[307,374,1524,1548]" metricMagnitude="2" metricUnit="m" metricValue="9.25" pageId="81" pageNumber="186" unit="m" value="925.0">
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||
<elevation id="0CF02375110BFFB3FEADAAC5185C9428" box="[307,374,1524,1548]" metricMagnitude="2" metricUnit="m" metricValue="9.25" pageId="81" pageNumber="186" unit="m" value="925.0">925 m</elevation>
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||
</quantity>
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||
<date id="F363E286110BFFB3FEE2AAC51B0C9428" box="[380,550,1524,1548]" pageId="81" pageNumber="186" value="2001-01-20" valueMax="2001-01-26" valueMin="2001-01-20">
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<collectingDate id="E3271B6E110BFFB3FEE2AAC51B0C9428" box="[380,550,1524,1548]" pageId="81" pageNumber="186" value="2001-01-20" valueMax="2001-01-26" valueMin="2001-01-20">20-26 Jan. 2001</collectingDate>
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||
</date>
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||
/
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||
<geoCoordinate id="E2E9A281110BFFB3FDA4AAC51B919428" box="[570,699,1524,1548]" degrees="12" direction="south" minutes="32" orientation="latitude" pageId="81" pageNumber="186" precision="15" seconds="4" value="-12.534445">12° 32’ 4” S</geoCoordinate>
|
||
<geoCoordinate id="E2E9A281110BFFB3FD5FAAC519C59408" degrees="049" direction="east" minutes="10" orientation="longitude" pageId="81" pageNumber="186" precision="15" seconds="46" value="49.179447">049° 10’ 46” E</geoCoordinate>
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||
coll.
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||
<collectorName id="2A28A190110BFFB3FEB0A92518589408" box="[302,370,1556,1580]" pageId="81" pageNumber="186">Fisher</collectorName>
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||
,
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<collectorName id="2A28A190110BFFB3FEE0A92518CB9408" box="[382,481,1556,1580]" pageId="81" pageNumber="186">Griswold</collectorName>
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et al.
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<collectingCounty id="6E03BCCA110BFFB3FDBBA92519CF9468" pageId="81" pageNumber="186">California Acad. of Sciences</collectingCounty>
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sifted litter montane rainforest code: BLF2564 / [distended, glued on card, dissected, male genitalia glued in DMHF resin on a separate card, same pin]
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</materialsCitation>
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.
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<materialsCitation id="37B5CE1B110BFFB3FDEDA9441B4F9488" collectingDate="2001-01-20" collectingDateMax="2001-01-26" collectingDateMin="2001-01-20" collectionCode="CASC" collectorName="San Francisco, CA & Fisher & Griswold" country="Madagascar" county="California Acad. of Sciences" elevation="925" latitude="-12.534445" location="Academy of Sciences" longLatPrecision="21" longitude="49.179447" municipality="d'Antisiranana Parc National Montagne d'Ambre" pageId="81" pageNumber="186" specimenCount="1" specimenCount-male="1" stateProvince="California" typeStatus="paratype">
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<typeStatus id="58667AE4110BFFB3FDEDA9441BF594A9" box="[627,735,1653,1677]" pageId="81" pageNumber="186" type="paratype">Paratypes</typeStatus>
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||
:
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<specimenCount id="91DB0FCF110BFFB3FD76A94519B69488" count="1" pageId="81" pageNumber="186" type="male">1 ♂</specimenCount>
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||
(
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<collectionCode id="E1CC5C83110BFFB3FF34A9A519DF9488" box="[170,245,1684,1708]" pageId="81" pageNumber="186">CASC</collectionCode>
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), same collecting data as holotype
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</materialsCitation>
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.
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</paragraph>
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</subSubSection>
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<subSubSection id="CFC797CD110BFFB3FF30A9851DD99008" pageId="81" pageNumber="186" type="diagnosis">
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<paragraph id="8762C446110BFFB3FF30A9851DD99008" blockId="81.[136,757,1364,1932]" lastBlockId="81.[816,1437,180,1932]" pageId="81" pageNumber="186">
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<emphasis id="B5A91854110BFFB3FF30A985180B94E8" bold="true" box="[174,289,1716,1740]" pageId="81" pageNumber="186">Diagnosis.</emphasis>
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<taxonomicName id="40DDBFC5110BFFB3FEB2A9851B1694E8" authorityName="Ballerio" authorityYear="2021" box="[300,572,1716,1740]" class="Insecta" family="Hybosoridae" genus="Goudotostes" kingdom="Animalia" order="Coleoptera" pageId="81" pageNumber="186" phylum="Arthropoda" rank="species" species="montanellus" status="sp. nov.">
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<emphasis id="B5A91854110BFFB3FEB2A9851B1694E8" box="[300,572,1716,1740]" italics="true" pageId="81" pageNumber="186">Goudotostes montanellus</emphasis>
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</taxonomicName>
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<emphasis id="B5A91854110BFFB3FDD9A9851B8A94E8" bold="true" box="[583,672,1716,1740]" pageId="81" pageNumber="186">
|
||
<taxonomicNameLabel id="AE9AA52F110BFFB3FDD9A9851B8A94E8" box="[583,672,1716,1740]" pageId="81" pageNumber="186" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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can be differentiated from all other species of
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<taxonomicName id="40DDBFC5110BFFB3FDD6A9E51BE694C8" authorityName=", sensu Ballerio" authorityYear="2008" box="[584,716,1748,1772]" class="Insecta" family="Hybosoridae" genus="Goudotostes" kingdom="Animalia" order="Coleoptera" pageId="81" pageNumber="186" phylum="Arthropoda" rank="genus">
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<emphasis id="B5A91854110BFFB3FDD6A9E51BE694C8" box="[584,716,1748,1772]" italics="true" pageId="81" pageNumber="186">Goudotostes</emphasis>
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</taxonomicName>
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by the following combination of characters: a) large size, b) dorsal ocular area present (vestigial), c) head with a weak tubercle on clypeal disc, d) pronotal anterior carina monolobate, e) pronotal anterior carina weakly subtrapezoidal in frontal view, f) pronotal anterior carina weakly raised and without generating a longitudinal sharp carina visible in dorsal view, g) pronotal basal carina monolobate, h) pronotum weakly saddle-shaped in lateral view, i) pronotal disc with gibbosities and lacking any distinct carinae and/or tubercles (apart from the monolobate anterior and posterior carinae), j) elytra with three (including sutural stria) irregular rows, distinctly spaced out from each other, of strongly raised tubercles and relatively long longitudinal carinae, k) aedeagus with short asymmetrical parameres. The species falls within the
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<taxonomicName id="40DDBFC5110BFFB3FBCBAEE51D9493C8" baseAuthorityName="Laporte" baseAuthorityYear="1840" box="[1109,1214,468,492]" class="Insecta" family="Hybosoridae" genus="Goudotostes" kingdom="Animalia" order="Coleoptera" pageId="81" pageNumber="186" phylum="Arthropoda" rank="species" species="scabrosus">
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<emphasis id="B5A91854110BFFB3FBCBAEE51D9493C8" box="[1109,1214,468,492]" italics="true" pageId="81" pageNumber="186">scabrosus</emphasis>
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</taxonomicName>
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group of species and can be differentiated by the other species of the group by the combination of characters e), f), and j).
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</paragraph>
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</subSubSection>
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||
<subSubSection id="CFC797CD110BFFB0FCC8AD051A2993C8" lastPageId="82" lastPageNumber="187" pageId="81" pageNumber="186" type="description">
|
||
<paragraph id="8762C446110BFFB0FCC8AD051A2993C8" blockId="81.[816,1437,180,1932]" lastBlockId="82.[151,773,180,652]" lastPageId="82" lastPageNumber="187" pageId="81" pageNumber="186">
|
||
<emphasis id="B5A91854110BFFB3FCC8AD051ACB9068" bold="true" box="[854,993,564,588]" pageId="81" pageNumber="186">Description.</emphasis>
|
||
Size: HL =
|
||
<quantity id="402569A3110BFFB3FBF4AD051DEB9068" box="[1130,1217,564,588]" metricMagnitude="-3" metricUnit="m" metricValue="1.1" pageId="81" pageNumber="186" unit="mm" value="1.1">1.1 mm</quantity>
|
||
; HW =
|
||
<quantity id="402569A3110BFFB3FA84AD051C589068" box="[1306,1394,564,588]" metricMagnitude="-3" metricUnit="m" metricValue="1.5" pageId="81" pageNumber="186" unit="mm" value="1.5">1.5 mm</quantity>
|
||
; PL =
|
||
<quantity id="402569A3110BFFB3FCD6AD651AB59048" box="[840,927,596,620]" metricMagnitude="-3" metricUnit="m" metricValue="1.4" pageId="81" pageNumber="186" unit="mm" value="1.4">1.4 mm</quantity>
|
||
; PW =
|
||
<quantity id="402569A3110BFFB3FC6DAD651D609048" box="[1011,1098,596,620]" metricMagnitude="-3" metricUnit="m" metricValue="2.5" pageId="81" pageNumber="186" unit="mm" value="2.5">2.5 mm</quantity>
|
||
; EL =
|
||
<quantity id="402569A3110BFFB3FB08AD651DC79048" box="[1174,1261,596,620]" metricMagnitude="-3" metricUnit="m" metricValue="2.7" pageId="81" pageNumber="186" unit="mm" value="2.7">2.7 mm</quantity>
|
||
; EW =
|
||
<quantity id="402569A3110BFFB3FADCAD651CB39048" box="[1346,1433,596,620]" metricMagnitude="-3" metricUnit="m" metricValue="2.3" pageId="81" pageNumber="186" unit="mm" value="2.3">2.3 mm</quantity>
|
||
. Overall morphology as in generic diagnosis. Large-sized
|
||
<taxonomicName id="40DDBFC5110BFFB3FCAEADA51A9F9088" authorityName=", sensu Ballerio" authorityYear="2008" box="[816,949,660,684]" class="Insecta" family="Hybosoridae" genus="Goudotostes" kingdom="Animalia" order="Coleoptera" pageId="81" pageNumber="186" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B5A91854110BFFB3FCAEADA51A9F9088" box="[816,949,660,684]" italics="true" pageId="81" pageNumber="186">Goudotostes</emphasis>
|
||
</taxonomicName>
|
||
. Body strongly convex. Dark brown with metallic green sheen on carinae, shiny, dorsum with setigerous punctures bearing a very short seta (50×), underside, tarsi and antennae dark reddish-brown, setation yellowish-brown.
|
||
<emphasis id="B5A91854110BFFB3FC32AC251ACC9108" box="[940,998,788,812]" italics="true" pageId="81" pageNumber="186">Head</emphasis>
|
||
: W/L ratio = 1.45; clypeal apex obtuse, apical projection strongly developed and sexually dimorphic, sides slightly rounded with slightly unaligned genae; dorsal ocular area present and very small, interocular distance about 30 times the maximum width of dorsal ocular area, ventral ocular area medium-sized, genae acute, distinctly protruding outwards, genal suture indistinct; head dorsal surface with a weak tubercle, punctation made of irregular dense impressed large simple punctures, surface along clypeal fore margin covered by some transverse irregular lines. Interpunctural distance of punctures inferior than punctural diameter.
|
||
<emphasis id="B5A91854110BFFB3FBD8AB441D9A96A8" box="[1094,1200,1141,1164]" italics="true" pageId="81" pageNumber="186">Pronotum</emphasis>
|
||
: subtrapezoidal (dorsal view) (W/L ratio = 1.87), convex, fore angles gently truncate; pronotal surface with disc covered by dense impressed large simple punctures, interpunctural distance inferior than punctural diameter, sides with some horseshoe-shaped punctures with opening directed outwards; pronotal bead present but not appreciable in dorsal view; pronotal sculpturing as follows: monolobate basal carina forming a small inverted V-shaped tubercle in dorsal view, monolobate anterior carina strongly raised (profile of pronotum saddle-shaped in lateral view) generating a subtrapezoidal (in frontal view) tubercle, disc with a depression in the middle and two weak paradiscal gibbosities on each side.
|
||
<taxonomicName id="40DDBFC5110BFFB3FCF4A9251AF99408" box="[874,979,1556,1580]" class="Trilobita" family="Styginidae" genus="Scutellum" kingdom="Animalia" order="Corynexochida" pageId="81" pageNumber="186" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B5A91854110BFFB3FCF4A9251AF99408" box="[874,979,1556,1580]" italics="true" pageId="81" pageNumber="186">Scutellum</emphasis>
|
||
</taxonomicName>
|
||
: completely covered by deep large simple punctures.
|
||
<emphasis id="B5A91854110BFFB3FC39A9051AC09468" box="[935,1002,1588,1612]" italics="true" pageId="81" pageNumber="186">Elytra</emphasis>
|
||
: slightly longer than wide (W/L ratio = 0.90), subrectangular (dorsal view), convex; humeral callus indistinct; elytral surface uniformly covered by small deeply impressed comma-shaped punctures with opening directed backwards or outwards, interpunctural distance inferior than punctural diameter; striated articular area hardly visible in lateral view, relatively narrow and short; inferior sutural stria complete and deep; marginal area large and distinctly protrudent outwards; elytral articular process well developed, smooth and shiny with a furrow bordering scutellar sides. Elytral sculpturing as follows: base with one tubercle near apex of scutellum (parasutural tubercle), next to it one longer raised longitudinal carina, then another shorter and finally an oblique carina marking humeral area, remaining dorsal surface with three (including sutural one) irregular and asymmetrical rows of weakly raised relatively long carinae, sides and apical area with denser and irregular tubercles; surface of tubercles made of deep irregular large punctures.
|
||
<emphasis id="B5A918541108FFB0FD95AEA41B669388" box="[523,588,405,428]" italics="true" pageId="82" pageNumber="187">Wings</emphasis>
|
||
: flightless.
|
||
<emphasis id="B5A918541108FFB0FD52AEA519DC93E8" italics="true" pageId="82" pageNumber="187">Male genitalia</emphasis>
|
||
: aedeagus with parameres (L =
|
||
<quantity id="402569A31108FFB0FDFAAE851B9193E8" box="[612,699,436,460]" metricMagnitude="-3" metricUnit="m" metricValue="1.7" pageId="82" pageNumber="187" unit="mm" value="1.7">1.7 mm</quantity>
|
||
) as in
|
||
<figureCitation id="1FE6D8C31108FFB0FF09AEE519CC93C8" box="[151,230,468,492]" captionStart="Fig" captionStartId="110.[151,182,1858,1877]" captionTargetBox="[188,1394,177,1817]" captionTargetId="figure-13@110.[161,1424,160,1823]" captionTargetPageId="110" captionText="Fig. 45 – Goudotostes montanellus sp. nov., a, holotype, dorsal view of distended specimen; b, paratype, ventral view of rolled up specimen; c, paratype, dorsal view of rolled up specimen; d, paratype, lateral view of rolled up specimen; e, paratype, pronotum of rolled up specimen; f, holotype, spiculum gastrale; g–j, holotype, four views of aedeagus." figureDoi="http://doi.org/10.5281/zenodo.12764265" httpUri="https://zenodo.org/record/12764265/files/figure.png" pageId="82" pageNumber="187">Figs 45</figureCitation>
|
||
g-j, spiculum gastrale (L = 2.0 mm) as in
|
||
<figureCitation id="1FE6D8C31108FFB0FD3CAEE51BD593C8" box="[674,767,468,492]" captionStart="Fig" captionStartId="110.[151,182,1858,1877]" captionTargetBox="[188,1394,177,1817]" captionTargetId="figure-13@110.[161,1424,160,1823]" captionTargetPageId="110" captionText="Fig. 45 – Goudotostes montanellus sp. nov., a, holotype, dorsal view of distended specimen; b, paratype, ventral view of rolled up specimen; c, paratype, dorsal view of rolled up specimen; d, paratype, lateral view of rolled up specimen; e, paratype, pronotum of rolled up specimen; f, holotype, spiculum gastrale; g–j, holotype, four views of aedeagus." figureDoi="http://doi.org/10.5281/zenodo.12764265" httpUri="https://zenodo.org/record/12764265/files/figure.png" pageId="82" pageNumber="187">
|
||
Fig.
|
||
<specimenCount id="91DB0FCF1108FFB0FD4DAEE51BD593C8" box="[723,767,468,492]" count="45" pageId="82" pageNumber="187" type="generic">45 f</specimenCount>
|
||
</figureCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="CFC797CD1108FFB0FF23AEC518B89008" pageId="82" pageNumber="187" type="etymology">
|
||
<paragraph id="8762C4461108FFB0FF23AEC518B89008" blockId="82.[151,773,180,652]" pageId="82" pageNumber="187">
|
||
<emphasis id="B5A918541108FFB0FF23AEC518149028" bold="true" box="[189,318,500,524]" pageId="82" pageNumber="187">Etymology.</emphasis>
|
||
Latin adjective in the nominative singular, diminutive of montane.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="CFC797CD1108FFB0FF23AD05180890A8" pageId="82" pageNumber="187" type="distribution">
|
||
<paragraph id="8762C4461108FFB0FF23AD05180890A8" blockId="82.[151,773,180,652]" pageId="82" pageNumber="187">
|
||
<emphasis id="B5A918541108FFB0FF23AD0518F69068" bold="true" box="[189,476,564,588]" pageId="82" pageNumber="187">Distribution and habitat.</emphasis>
|
||
Known only from the
|
||
<typeStatus id="58667AE41108FFB0FD4BAD041A299069" box="[725,771,565,589]" pageId="82" pageNumber="187">type</typeStatus>
|
||
locality. See under
|
||
<taxonomicName id="40DDBFC51108FFB0FEF4AD641B519048" authorityName="Ballerio" authorityYear="2021" box="[362,635,597,620]" class="Insecta" family="Hybosoridae" genus="Synarmostes" kingdom="Animalia" order="Coleoptera" pageId="82" pageNumber="187" phylum="Arthropoda" rank="species" species="annamariae" status="sp. nov.">
|
||
<emphasis id="B5A918541108FFB0FEF4AD641B519048" box="[362,635,597,620]" italics="true" pageId="82" pageNumber="187">Synarmostes annamariae</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B5A918541108FFB0FD1AAD651BF19048" bold="true" box="[644,731,596,620]" pageId="82" pageNumber="187">
|
||
<taxonomicNameLabel id="AE9AA52F1108FFB0FD1AAD651BF19048" box="[644,731,596,620]" pageId="82" pageNumber="187" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
for more details.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |