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<document id="C5BCA410E1273B3280F615F1D82F6A43" ID-CLB-Dataset="270518" ID-DOI="10.11646/phytotaxa.614.1.1" ID-GBIF-Dataset="143c8e81-3541-4298-b233-d3ce4e79d26c" ID-ISSN="1179-3163" ID-Zenodo-Dep="8389307" IM.bibliography_approvedBy="diego" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="diego" IM.treatments_approvedBy="diego" checkinTime="1695979127173" checkinUser="plazi" docAuthor="Henderson, Andrew" docDate="2023" docId="03B387DAFFE41F71FF50FADFFD748CEA" docLanguage="en" docName="phytotaxa.614.1.1.pdf" docOrigin="Phytotaxa 614 (1)" docSource="http://dx.doi.org/10.11646/phytotaxa.614.1.1" docStyle="DocumentStyle:F08184CE06D8A97EA3E6DE35D99648B0.3:Phytotaxa.2014-.monograph" docStyleId="F08184CE06D8A97EA3E6DE35D99648B0" docStyleName="Phytotaxa.2014-.monograph" docStyleVersion="3" docTitle="Coccothrinax argentea Sargent ex Schumann 1901" docType="treatment" docVersion="7" lastPageNumber="31" masterDocId="FF8AFFA2FFFF1F6FFFD8FFB9FFF9881B" masterDocTitle="A revision of Coccothrinax, Hemithrinax, Leucothrinax, Thrinax, and Zombia (Arecaceae)" masterLastPageNumber="115" masterPageNumber="1" pageNumber="28" updateTime="1699690893055" updateUser="plazi">
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<mods:title id="BC1C2D690B7F9DAE4FA8FB1548581465">A revision of Coccothrinax, Hemithrinax, Leucothrinax, Thrinax, and Zombia (Arecaceae)</mods:title>
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<treatment id="03B387DAFFE41F71FF50FADFFD748CEA" ID-DOI="http://doi.org/10.5281/zenodo.8400225" ID-GBIF-Taxon="213635818" ID-Zenodo-Dep="8400225" LSID="urn:lsid:plazi:treatment:03B387DAFFE41F71FF50FADFFD748CEA" httpUri="http://treatment.plazi.org/id/03B387DAFFE41F71FF50FADFFD748CEA" lastPageId="30" lastPageNumber="31" pageId="27" pageNumber="28">
<subSubSection id="C3006547FFE41F74FF50FADFFB1F8D9B" box="[136,1254,1382,1409]" pageId="27" pageNumber="28" type="nomenclature">
<paragraph id="8BA536CCFFE41F74FF50FADFFB1F8D9B" blockId="27.[136,1452,1382,1516]" box="[136,1254,1382,1409]" pageId="27" pageNumber="28">
<emphasis id="B96EEADEFFE41F74FF50FADFFB1F8D9B" bold="true" box="[136,1254,1382,1409]" pageId="27" pageNumber="28">
1.3.
<taxonomicName id="4C1A4D4FFFE41F74FF6FFADFFB198D9B" ID-CoL="8W8X2" ID-ENA="480894" authority="(Loddiges ex Schultes &amp; Schultes) Sargent ex Schumann (1901: 469)" authorityName="Sargent ex Schumann" authorityPageNumber="469" authorityYear="1901" baseAuthorityName="Loddiges ex Schultes &amp; Schultes" box="[183,1248,1382,1409]" class="Liliopsida" family="Arecaceae" genus="Coccothrinax" kingdom="Plantae" order="Arecales" pageId="27" pageNumber="28" phylum="Tracheophyta" rank="species" species="argentea">
<emphasis id="B96EEADEFFE41F74FF6FFADFFE478D9A" bold="true" box="[183,446,1382,1409]" italics="true" pageId="27" pageNumber="28">Coccothrinax argentea</emphasis>
(Loddiges ex Schultes &amp; Schultes) Sargent ex
<bibRefCitation id="EF8B4B3DFFE41F74FC0FFADFFB198D9B" author="Schumann, K." box="[983,1248,1382,1408]" pageId="27" pageNumber="28" pagination="449 - 545" refId="ref64771" refString="Schumann, K. (1901) Neue Arten der Siphonogamen 1899. Just's Botanischer Jahresbericht 27 (1): 449 - 545." type="journal article" year="1901">Schumann (1901: 469)</bibRefCitation>
</taxonomicName>
.
</emphasis>
</paragraph>
</subSubSection>
<subSubSection id="C3006547FFE41F74FB33FADFFF4C8DD2" pageId="27" pageNumber="28" type="reference_group">
<paragraph id="8BA536CCFFE41F74FB33FADFFD6A8DBF" blockId="27.[136,1452,1382,1516]" pageId="27" pageNumber="28">
<taxonomicName id="4C1A4D4FFFE41F74FB33FADFFD778DBF" ID-CoL="56LVL" authority="Loddiges ex Schultes &amp; Schultes (1830: 1300)" authorityName="Loddiges ex Schultes &amp; Schultes" authorityPageNumber="1300" authorityYear="1830" class="Liliopsida" family="Arecaceae" genus="Thrinax" kingdom="Plantae" order="Arecales" pageId="27" pageNumber="28" phylum="Tracheophyta" rank="species" species="argentea">
<emphasis id="B96EEADEFFE41F74FB33FADFFA558D9B" box="[1259,1452,1382,1408]" italics="true" pageId="27" pageNumber="28">Thrinax argentea</emphasis>
Loddiges ex
<bibRefCitation id="EF8B4B3DFFE41F74FECEFA33FD778DBF" author="Schultes, J. &amp; Schultes, J." box="[278,654,1418,1444]" pageId="27" pageNumber="28" pagination="725 - 1815" refId="ref64737" refString="Schultes, J. &amp; Schultes, J. (1830) Systema vegetabilium. Volume 7 part 2. J. G. Cottae, Stuttgardtiae, pp. 725 - 1815." type="book chapter" year="1830">Schultes &amp; Schultes (1830: 1300)</bibRefCitation>
</taxonomicName>
.
</paragraph>
<paragraph id="8BA536CCFFE41F74FD40FA33FF4C8DD2" blockId="27.[136,1452,1382,1516]" pageId="27" pageNumber="28">
<taxonomicName id="4C1A4D4FFFE41F74FD40FA33FF488DD2" authority="(Loddiges ex Schultes &amp; Schultes) Cook (1941: 50)" authorityName="Cook" authorityPageNumber="50" authorityYear="1941" baseAuthorityName="Loddiges ex Schultes &amp; Schultes" class="Liliopsida" family="Arecaceae" genus="Acanthorriza" kingdom="Plantae" order="Arecales" pageId="27" pageNumber="28" phylum="Tracheophyta" rank="species" species="argentea">
<emphasis id="B96EEADEFFE41F74FD40FA33FC6C8DBF" box="[664,917,1418,1444]" italics="true" pageId="27" pageNumber="28">Acanthorriza argentea</emphasis>
(Loddiges ex Schultes &amp; Schultes)
<bibRefCitation id="EF8B4B3DFFE41F74FAF8FA33FF488DD2" author="Cook, O." pageId="27" pageNumber="28" pagination="21 - 52" refId="ref61536" refString="Cook, O. (1941) A Haitian cactus palm adapted to Florida. National Horticultural Magazine 20: 21 - 52." type="journal article" year="1941">Cook (1941: 50)</bibRefCitation>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="C3006547FFE41F74FF62FA16FF6F8DF7" pageId="27" pageNumber="28" type="materials_examined">
<paragraph id="8BA536CCFFE41F74FF62FA16FF6F8DF7" blockId="27.[136,1452,1382,1516]" pageId="27" pageNumber="28">
<typeStatus id="54A1886EFFE41F74FF62FA16FEE08DD2" box="[186,281,1455,1481]" pageId="27" pageNumber="28" type="neotype">Neotype</typeStatus>
(designated here):—
<collectingCountry id="F30D765CFFE41F74FDDBFA17FCCE8DD3" box="[515,823,1454,1480]" name="Dominican Republic" pageId="27" pageNumber="28">DOMINICAN REPUBLIC</collectingCountry>
. Haina,
<date id="FFA4100CFFE41F74FC48FA17FBDB8DD3" box="[912,1058,1454,1480]" pageId="27" pageNumber="28" value="1939-07-26">26 July 1939</date>
,
<emphasis id="B96EEADEFFE41F74FBF3FA16FB468DD3" box="[1067,1215,1454,1480]" italics="true" pageId="27" pageNumber="28">L. Bailey 269</emphasis>
(
<typeStatus id="54A1886EFFE41F74FB15FA17FADE8DD3" box="[1229,1319,1454,1480]" pageId="27" pageNumber="28" type="neotype">neotype</typeStatus>
BH!).
<figureCitation id="13212A49FFE41F74FAABFA17FF6F8DF7" captionStart="PLATE 3" captionStartId="28.[136,214,1917,1938]" captionTargetBox="[151,1433,194,1891]" captionTargetId="figure-27@28.[150,1434,193,1893]" captionTargetPageId="28" captionText="PLATE 3. Coccothrinax argentea, Dominican Republic, with leaf segments not pendulous at the apices, giving the leaf a flat appearance." figureDoi="http://doi.org/10.5281/zenodo.8389345" httpUri="https://zenodo.org/record/8389345/files/figure.png" pageId="27" pageNumber="28">Plate 3</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="C3006547FFE41F74FF50F9A7FC2C8E43" pageId="27" pageNumber="28" type="reference_group">
<paragraph id="8BA536CCFFE41F74FF50F9A7FC2C8E43" blockId="27.[136,1452,1566,1624]" pageId="27" pageNumber="28">
<taxonomicName id="4C1A4D4FFFE41F74FF50F9A7FE708E2F" ID-CoL="56LWB" authority="Beccari" authorityName="Beccari" box="[136,393,1566,1588]" class="Liliopsida" family="Arecaceae" genus="Thrinax" kingdom="Plantae" order="Arecales" pageId="27" pageNumber="28" phylum="Tracheophyta" rank="species" species="longistyla">
<emphasis id="B96EEADEFFE41F74FF50F9A7FEC38E28" box="[136,314,1566,1587]" italics="true" pageId="27" pageNumber="28">Thrinax longistyla</emphasis>
<materialsCitation id="3B723C91FFE41F74FE98F9A7FE708E2F" ID-GBIF-Occurrence="4418891335" box="[320,393,1566,1588]" collectingDate="1910-06" collectionCode="M, GH, FI, K, B" country="Dominican Republic" location="Barahona" pageId="27" pageNumber="28" specimenCount="3" stateProvince="Barahona" typeStatus="lectotype">Beccari</materialsCitation>
</taxonomicName>
in
<bibRefCitation id="EF8B4B3DFFE41F74FE71F9A7FDA68E28" author="Urban, I." box="[425,607,1566,1588]" pageId="27" pageNumber="28" refId="ref65121" refString="Urban, I. (1912) Symbolae Antillanae seu Fundamente Florae Indiae Occidentalis. Volume 7. Fratres Borntraeger, Lipsiae, 580 pp." type="book" year="1912">Urban (1912: 170)</bibRefCitation>
.
<typeStatus id="54A1886EFFE41F74FDB1F9A7FD308E28" box="[617,713,1566,1587]" pageId="27" pageNumber="28" type="lectotype">Lectotype</typeStatus>
(designated here):—
<collectingCountry id="F30D765CFFE41F74FC49F9A7FB668E28" box="[913,1183,1566,1588]" name="Dominican Republic" pageId="27" pageNumber="28">DOMINICAN REPUBLIC</collectingCountry>
.
<collectingRegion id="49DEF82EFFE41F74FB70F9A7FAFF8E2F" box="[1192,1286,1566,1588]" country="Dominican Republic" name="Barahona" pageId="27" pageNumber="28">Barahona</collectingRegion>
, 0 m,
<date id="FFA4100CFFE41F74FA9BF9A7FA5E8E28" box="[1347,1447,1566,1587]" pageId="27" pageNumber="28" value="1910-06">
<collectingDate id="EFE0E9E4FFE41F74FA9BF9A7FA5E8E28" box="[1347,1447,1566,1587]" pageId="27" pageNumber="28" value="1910-06">June 1910</collectingDate>
</date>
,
<emphasis id="B96EEADEFFE41F74FF65F9FBFEA88E4C" box="[189,337,1602,1623]" italics="true" pageId="27" pageNumber="28">M. Fuertes 415</emphasis>
(
<typeStatus id="54A1886EFFE41F74FE85F9FBFE4A8E43" box="[349,435,1602,1624]" pageId="27" pageNumber="28" type="holotype">holotype</typeStatus>
GH!, isotypes FI!, K!, the
<typeStatus id="54A1886EFFE41F74FD6EF9FBFCF38E43" box="[694,778,1602,1624]" pageId="27" pageNumber="28" type="holotype">holotype</typeStatus>
at B was destroyed).
</paragraph>
</subSubSection>
<subSubSection id="C3006547FFE41F72FF50F93FFF0B8953" lastPageId="29" lastPageNumber="30" pageId="27" pageNumber="28" type="description">
<paragraph id="8BA536CCFFE41F72FF50F93FFF0B8953" blockId="27.[136,1452,1670,2093]" lastBlockId="29.[136,1452,158,437]" lastPageId="29" lastPageNumber="30" pageId="27" pageNumber="28">
<emphasis id="B96EEADEFFE41F74FF50F93FFF298EBB" bold="true" box="[136,208,1670,1696]" pageId="27" pageNumber="28">Stems</emphasis>
5.0(1.520.0) m long and 5.2(4.56.5) cm diameter, solitary or rarely clustered.
<emphasis id="B96EEADEFFE41F74FB8AF93FFB5D8EBB" bold="true" box="[1106,1188,1670,1696]" pageId="27" pageNumber="28">Leaves</emphasis>
more or less deciduous or only leaf bases persisting on stem; leaf sheath fibers 0.3(0.10.5) mm diameter, closely woven, forming persistent, triangular ligules at the apices; petioles 10.5(5.317.9) mm diameter just below the apex; palmans 14.8(4.332.0) cm long, relatively long, without prominent adaxial veins; leaf blades not wedge-shaped; segments 42(2558) per leaf, the middle ones 53.1(32.075.0) cm long and 2.9(1.55.5) cm wide; segments not pendulous at the apices, giving the leaf a flat appearance; middle leaf segments relatively long and narrow, tapering from base to apex, scarcely folded, flexible and not leathery, a shoulder or constriction absent or poorly developed, the apices thin, deeply splitting and breaking off; middle leaf segment apices attenuate; leaf segments not waxy or sometimes with a deciduous, thin layer of wax adaxially, densely indumentose abaxially, with irregularly shaped, semi-persistent, interlocking, fimbriate hairs without an obvious center, with well-developed transverse veinlets.
<emphasis id="B96EEADEFFE41F74FBE2F873FB1A8FFF" bold="true" box="[1082,1251,1994,2020]" pageId="27" pageNumber="28">Inflorescences</emphasis>
curving, arching, or pendulous amongst the leaves, with few partial inflorescences; rachis bracts somewhat flattened, loosely sheathing, usually tomentose with a dense tuft of erect hairs at the apex; partial inflorescences 4(25); proximalmost rachillae straight, 9.2(5.215.0) cm long and 1.3(0.91.9) mm diameter in fruit; rachillae glabrous at or near anthesis; stamens 8(610); fruit pedicels 0.8(0.32.5) mm long;
<emphasis id="B96EEADEFFE21F72FD5FFF7BFD3188C7" bold="true" box="[647,712,194,220]" pageId="29" pageNumber="30">fruits</emphasis>
7.7(5.69.1) mm long and 7.9(5.59.5) mm diameter, purple, dark purple, red-purple, reddish-purple, purple-black, red-pink, red-black, grayish-brown, blackish, or black; fruit surfaces smooth or sometimes with projecting fibers; seed surfaces deeply lobed, the lobes running from base of seeds almost to apices.
</paragraph>
</subSubSection>
<caption id="DF656644FFE31F73FF50F8C4FF028FAC" ID-DOI="http://doi.org/10.5281/zenodo.8389345" ID-Zenodo-Dep="8389345" httpUri="https://zenodo.org/record/8389345/files/figure.png" pageId="28" pageNumber="29" startId="28.[136,214,1917,1938]" targetBox="[151,1433,194,1891]" targetPageId="28" targetType="plate">
<paragraph id="8BA536CCFFE31F73FF50F8C4FF028FAC" blockId="28.[136,1451,1917,1975]" pageId="28" pageNumber="29">
<emphasis id="B96EEADEFFE31F73FF50F8C4FF178F89" bold="true" box="[136,238,1917,1938]" pageId="28" pageNumber="29">PLATE 3</emphasis>
.
<taxonomicName id="4C1A4D4FFFE31F73FED9F8C4FE1D8F88" authorityName="Sargent ex Schumann" authorityYear="1901" baseAuthorityName="Loddiges ex Schultes &amp; Schultes" box="[257,484,1917,1939]" class="Liliopsida" family="Arecaceae" genus="Coccothrinax" kingdom="Plantae" order="Arecales" pageId="28" pageNumber="29" phylum="Tracheophyta" rank="species" species="argentea">
<emphasis id="B96EEADEFFE31F73FED9F8C4FE1D8F88" box="[257,484,1917,1939]" italics="true" pageId="28" pageNumber="29">Coccothrinax argentea</emphasis>
</taxonomicName>
, Dominican Republic, with leaf segments not pendulous at the apices, giving the leaf a flat appearance.
</paragraph>
</caption>
<subSubSection id="C3006547FFE21F72FF65FEEBFC7C89AF" pageId="29" pageNumber="30" type="distribution">
<paragraph id="8BA536CCFFE21F72FF65FEEBFC7C89AF" blockId="29.[136,1452,158,437]" pageId="29" pageNumber="30">
<emphasis id="B96EEADEFFE21F72FF65FEEBFDF28977" bold="true" box="[189,523,338,364]" pageId="29" pageNumber="30">Distribution and habitat:—</emphasis>
Hispaniola (
<collectingCountry id="F30D765CFFE21F72FD4EFEEBFD2B8977" box="[662,722,338,364]" name="Haiti" pageId="29" pageNumber="30">Haiti</collectingCountry>
,
<collectingCountry id="F30D765CFFE21F72FD39FEEBFC378977" box="[737,974,338,364]" name="Dominican Republic" pageId="29" pageNumber="30">Dominican Republic</collectingCountry>
) (
<figureCitation id="13212A49FFE21F72FC3FFEEBFBD48977" box="[999,1069,338,364]" captionStart="FIGURE 9" captionStartId="29.[136,229,1815,1837]" captionTargetBox="[155,1429,505,1786]" captionTargetId="figure-121@29.[150,1434,500,1791]" captionTargetPageId="29" captionText="FIGURE 9. Distribution of Coccothrinax argentea and C. barbadensis." figureDoi="http://doi.org/10.5281/zenodo.8389347" httpUri="https://zenodo.org/record/8389347/files/figure.png" pageId="29" pageNumber="30">Fig. 9</figureCitation>
) in deciduous or semi-deciduous forest, xeromorphic forest, pine forest, broad leaf forest, dry forest, or secondary forest, and persisting in disturbed areas, on limestone or serpentine soils, at 259(01,000) m elevation.
</paragraph>
</subSubSection>
<caption id="DF656644FFE21F72FF50F8AEFCC38F36" ID-DOI="http://doi.org/10.5281/zenodo.8389347" ID-Zenodo-Dep="8389347" box="[136,826,1815,1837]" httpUri="https://zenodo.org/record/8389347/files/figure.png" pageId="29" pageNumber="30" startId="29.[136,229,1815,1837]" targetBox="[155,1429,505,1786]" targetPageId="29" targetType="figure">
<paragraph id="8BA536CCFFE21F72FF50F8AEFCC38F36" blockId="29.[136,826,1815,1837]" box="[136,826,1815,1837]" pageId="29" pageNumber="30">
<emphasis id="B96EEADEFFE21F72FF50F8AEFF0E8F37" bold="true" box="[136,247,1815,1837]" pageId="29" pageNumber="30">FIGURE 9</emphasis>
. Distribution of
<taxonomicName id="4C1A4D4FFFE21F72FE4FF8AEFD8D8F36" authorityName="Sargent ex Schumann" authorityYear="1901" baseAuthorityName="Loddiges ex Schultes &amp; Schultes" box="[407,628,1815,1837]" class="Liliopsida" family="Arecaceae" genus="Coccothrinax" kingdom="Plantae" order="Arecales" pageId="29" pageNumber="30" phylum="Tracheophyta" rank="species" species="argentea">
<emphasis id="B96EEADEFFE21F72FE4FF8AEFD8D8F36" box="[407,628,1815,1837]" italics="true" pageId="29" pageNumber="30">Coccothrinax argentea</emphasis>
</taxonomicName>
and
<taxonomicName id="4C1A4D4FFFE21F72FD7BF8AEFCCD8F37" authorityName="Beccari" authorityYear="1907" baseAuthorityName="Loddiges ex Martius" box="[675,820,1815,1836]" class="Liliopsida" family="Arecaceae" genus="Coccothrinax" kingdom="Plantae" order="Arecales" pageId="29" pageNumber="30" phylum="Tracheophyta" rank="species" species="barbadensis">
<emphasis id="B96EEADEFFE21F72FD7BF8AEFCCD8F37" box="[675,820,1815,1836]" italics="true" pageId="29" pageNumber="30">C. barbadensis</emphasis>
</taxonomicName>
.
</paragraph>
</caption>
<subSubSection id="C3006547FFE21F71FF65F8E2FD748CEA" lastPageId="30" lastPageNumber="31" pageId="29" pageNumber="30" type="discussion">
<paragraph id="8BA536CCFFE21F71FF65F8E2FE87891B" blockId="29.[136,1452,1883,2090]" lastBlockId="30.[136,1452,158,1265]" lastPageId="30" lastPageNumber="31" pageId="29" pageNumber="30">
<emphasis id="B96EEADEFFE21F72FF65F8E2FE578F6D" bold="true" box="[189,430,1883,1910]" pageId="29" pageNumber="30">Taxonomic notes:—</emphasis>
As a preliminary species
<taxonomicName id="4C1A4D4FFFE21F72FD0AF8E2FC2E8F6E" authorityName="Sargent ex Schumann" authorityYear="1901" baseAuthorityName="Loddiges ex Schultes &amp; Schultes" box="[722,983,1883,1909]" class="Liliopsida" family="Arecaceae" genus="Coccothrinax" kingdom="Plantae" order="Arecales" pageId="29" pageNumber="30" phylum="Tracheophyta" rank="species" species="argentea">
<emphasis id="B96EEADEFFE21F72FD0AF8E2FC2E8F6E" box="[722,983,1883,1909]" italics="true" pageId="29" pageNumber="30">Coccothrinax argentea</emphasis>
</taxonomicName>
was polymorphic for one variable (stem branching). However, splitting it into two species such that variables were consistent within each one led to improbable species that could not be justified morphologically or geographically. Therefore the polymorphic variable was treated as a trait and
<taxonomicName id="4C1A4D4FFFE21F72FEC7F871FE5A8FFA" authorityName="Sargent ex Schumann" authorityYear="1901" baseAuthorityName="Loddiges ex Schultes &amp; Schultes" box="[287,419,1992,2018]" class="Liliopsida" family="Arecaceae" genus="Coccothrinax" kingdom="Plantae" order="Arecales" pageId="29" pageNumber="30" phylum="Tracheophyta" rank="species" species="argentea">
<emphasis id="B96EEADEFFE21F72FEC7F871FE5A8FFA" box="[287,419,1992,2018]" italics="true" pageId="29" pageNumber="30">C. argentea</emphasis>
</taxonomicName>
is recognized as a phylogenetic species.
<bibRefCitation id="EF8B4B3DFFE21F72FCA8F87EFBE08FFA" author="Bailey, L." box="[880,1049,1991,2017]" pageId="29" pageNumber="30" pagination="247 - 259" refId="ref60742" refString="Bailey, L. (1939 b) Coccothrinax in the southern Greater Antilles. Gentes Herbarum 4: 247 - 259." type="journal article" year="1939">Bailey (1939b)</bibRefCitation>
gave a discussion of the taxonomic history of
<taxonomicName id="4C1A4D4FFFE21F72FF25F855FE78801E" authorityName="Sargent ex Schumann" authorityYear="1901" baseAuthorityName="Loddiges ex Schultes &amp; Schultes" box="[253,385,2028,2054]" class="Liliopsida" family="Arecaceae" genus="Coccothrinax" kingdom="Plantae" order="Arecales" pageId="29" pageNumber="30" phylum="Tracheophyta" rank="species" species="argentea">
<emphasis id="B96EEADEFFE21F72FF25F855FE78801E" box="[253,385,2028,2054]" italics="true" pageId="29" pageNumber="30">C. argentea</emphasis>
</taxonomicName>
, and a detailed description based on a specimen (
<emphasis id="B96EEADEFFE21F72FC6EF852FBC9801E" box="[950,1072,2027,2053]" italics="true" pageId="29" pageNumber="30">Bailey 269</emphasis>
) that he had collected near
<collectingRegion id="49DEF82EFFE21F72FAB4F855FF0B8032" country="Dominican Republic" name="Santo Domingo" pageId="29" pageNumber="30">Santo Domingo</collectingRegion>
in
<collectingCountry id="F30D765CFFE21F72FEC4F7B6FE3D8032" box="[284,452,2063,2089]" name="Dominican Republic" pageId="29" pageNumber="30">the Dominican</collectingCountry>
Republic. This specimen is designated here as the
<typeStatus id="54A1886EFFE21F72FBC2F7A9FB8F8031" box="[1050,1142,2064,2090]" pageId="29" pageNumber="30" type="neotype">neotype</typeStatus>
. One other name has been applied to specimens from this area,
<taxonomicName id="4C1A4D4FFFE11F71FDFCFF27FD0888A3" box="[548,753,158,184]" class="Liliopsida" family="Arecaceae" genus="Thrinax" kingdom="Plantae" order="Arecales" pageId="30" pageNumber="31" phylum="Tracheophyta" rank="species" species="longistyla">
<emphasis id="B96EEADEFFE11F71FDFCFF27FD0888A3" box="[548,753,158,184]" italics="true" pageId="30" pageNumber="31">Thrinax longistyla</emphasis>
</taxonomicName>
.
<bibRefCitation id="EF8B4B3DFFE11F71FCD8FF27FC7288A2" author="Read, R." box="[768,907,158,185]" pageId="30" pageNumber="31" pagination="1 - 98" refId="ref64159" refString="Read, R. (1975) The genus Thrinax (Palmae: Coryphoideae). Smithsonian Contributions to Botany 19: 1 - 98. https: // doi. org / 10.5479 / si. 0081024 x. 19" type="journal article" year="1975">Read (1975)</bibRefCitation>
considered this to be a
<taxonomicName id="4C1A4D4FFFE11F71FB4BFF27FAD288A3" authorityName="Sargent" authorityYear="1899" box="[1171,1323,158,184]" class="Liliopsida" family="Arecaceae" genus="Coccothrinax" kingdom="Plantae" order="Arecales" pageId="30" pageNumber="31" phylum="Tracheophyta" rank="genus">
<emphasis id="B96EEADEFFE11F71FB4BFF27FAD288A3" box="[1171,1323,158,184]" italics="true" pageId="30" pageNumber="31">Coccothrinax</emphasis>
</taxonomicName>
rather than a
<taxonomicName id="4C1A4D4FFFE11F71FF45FF7BFF0F88C7" authorityName="Linnaeus f. ex Swartz" authorityYear="1788" box="[157,246,194,220]" class="Liliopsida" family="Arecaceae" genus="Thrinax" kingdom="Plantae" order="Arecales" pageId="30" pageNumber="31" phylum="Tracheophyta" rank="genus">
<emphasis id="B96EEADEFFE11F71FF45FF7BFF0F88C7" box="[157,246,194,220]" italics="true" pageId="30" pageNumber="31">Thrinax</emphasis>
</taxonomicName>
, and it was included as a synonym of
<taxonomicName id="4C1A4D4FFFE11F71FD74FF7AFCC888C7" authorityName="Sargent ex Schumann" authorityYear="1901" baseAuthorityName="Loddiges ex Schultes &amp; Schultes" box="[684,817,195,221]" class="Liliopsida" family="Arecaceae" genus="Coccothrinax" kingdom="Plantae" order="Arecales" pageId="30" pageNumber="31" phylum="Tracheophyta" rank="species" species="argentea">
<emphasis id="B96EEADEFFE11F71FD74FF7AFCC888C7" box="[684,817,195,221]" italics="true" pageId="30" pageNumber="31">C. argentea</emphasis>
</taxonomicName>
by
<bibRefCitation id="EF8B4B3DFFE11F71FC85FF7BFBFA88C7" author="POWO" box="[861,1027,194,220]" pageId="30" pageNumber="31" refId="ref63984" refString="POWO (2023) Plants of the World Online. Facilitated by the Royal Botanic Gardens, Kew. Published on the Internet. Available from: http: // www. plantsoftheworldonline. org / (accessed 24 May 2023)" type="url" year="2023">POWO (2023)</bibRefCitation>
and its type share all character states with other specimens.
</paragraph>
<paragraph id="8BA536CCFFE11F71FF65FEB3FF188953" blockId="30.[136,1452,158,1265]" pageId="30" pageNumber="31">
<taxonomicName id="4C1A4D4FFFE11F71FF65FEB3FE3B893F" authorityName="Sargent ex Schumann" authorityYear="1901" baseAuthorityName="Loddiges ex Schultes &amp; Schultes" box="[189,450,266,292]" class="Liliopsida" family="Arecaceae" genus="Coccothrinax" kingdom="Plantae" order="Arecales" pageId="30" pageNumber="31" phylum="Tracheophyta" rank="species" species="argentea">
<emphasis id="B96EEADEFFE11F71FF65FEB3FE3B893F" box="[189,450,266,292]" italics="true" pageId="30" pageNumber="31">Coccothrinax argentea</emphasis>
</taxonomicName>
is widespread in Hispaniola and appears to be a complicated and confusingly variable species.
</paragraph>
<paragraph id="8BA536CCFFE11F71FF65FEEBFDED89E7" blockId="30.[136,1452,158,1265]" pageId="30" pageNumber="31">
<emphasis id="B96EEADEFFE11F71FF65FEEBFE438977" bold="true" box="[189,442,338,364]" pageId="30" pageNumber="31">Subspecific variation</emphasis>
:—Specimens from several, different areas have relatively short palmans with prominent adaxial veins, approaching those of
<taxonomicName id="4C1A4D4FFFE11F71FDFAFECEFD63898B" authorityName="Burret" authorityYear="1929" box="[546,666,374,401]" class="Liliopsida" family="Arecaceae" genus="Coccothrinax" kingdom="Plantae" order="Arecales" pageId="30" pageNumber="31" phylum="Tracheophyta" rank="species" species="gracilis">
<emphasis id="B96EEADEFFE11F71FDFAFECEFD63898B" box="[546,666,374,401]" italics="true" pageId="30" pageNumber="31">C. gracilis</emphasis>
</taxonomicName>
. For example,
<specimenCount id="9D1CFD45FFE11F71FC9EFECEFC17898B" box="[838,1006,374,401]" pageId="30" pageNumber="31" type="generic">two specimens</specimenCount>
(
<emphasis id="B96EEADEFFE11F71FC27FECEFB0C898B" box="[1023,1269,374,400]" italics="true" pageId="30" pageNumber="31">Liogier 14587, 16128</emphasis>
) from the north coast of
<collectingCountry id="F30D765CFFE11F71FF3CFE23FE7189AF" box="[228,392,410,436]" name="Dominican Republic" pageId="30" pageNumber="31">the Dominican</collectingCountry>
Republic, from limestone cliffs, have this kind of short palman. Other specimens from scattered localities, often on serpentine soils, also have short palmans, for example those from the Loma la Peguera, the site of a nickel mine with serpentine soils.
</paragraph>
<paragraph id="8BA536CCFFE11F71FF65FDBFFAA78A5F" blockId="30.[136,1452,158,1265]" pageId="30" pageNumber="31">
One specimen (
<emphasis id="B96EEADEFFE11F71FEB7FDBEFDF98A3B" box="[367,512,518,544]" italics="true" pageId="30" pageNumber="31">Montero 436</emphasis>
) has clustered stems. It is from near
<collectingRegion id="49DEF82EFFE11F71FC7DFDBEFBD78A3B" box="[933,1070,518,545]" country="Dominican Republic" name="Puerto Plata" pageId="30" pageNumber="31">Puerto Plata</collectingRegion>
in northern
<collectingCountry id="F30D765CFFE11F71FB63FDBFFA5E8A3B" box="[1211,1447,518,544]" name="Dominican Republic" pageId="30" pageNumber="31">Dominican Republic</collectingCountry>
, near an area with other unusual palms, including an unidentified
<taxonomicName id="4C1A4D4FFFE11F71FC87FD93FAA38A5F" authority="(Rodriguez-Pena et al. 2014)" baseAuthorityName="Rodriguez-Pena" baseAuthorityYear="2014" box="[863,1370,554,580]" class="Liliopsida" family="Arecaceae" genus="Pseudophoenix" kingdom="Plantae" order="Arecales" pageId="30" pageNumber="31" phylum="Tracheophyta" rank="genus">
<emphasis id="B96EEADEFFE11F71FC87FD93FBF38A5F" box="[863,1034,554,580]" italics="true" pageId="30" pageNumber="31">Pseudophoenix</emphasis>
(
<bibRefCitation id="EF8B4B3DFFE11F71FBC0FD93FAAB8A5F" author="Rodriguez-Pena, R. &amp; Jestrow, B. &amp; Meerow, A. &amp; Clase, T. &amp; Jimenez-Rodriguez, F. &amp; Griffith, P. &amp; Santiago-Valentin, E. &amp; Sustache-Sustache, J. &amp; Francisco-Ortega, J." box="[1048,1362,554,580]" pageId="30" pageNumber="31" pagination="469 - 485" refId="ref64408" refString="Rodriguez-Pena, R., Jestrow, B., Meerow, A., Clase, T., Jimenez-Rodriguez, F., Griffith, P., Santiago-Valentin, E., Sustache-Sustache, J. &amp; Francisco-Ortega, J. (2014) Genetic diversity and differentiation of Pseudophoenix (Arecaceae) in Hispaniola. Botanical Journal of the Linnean Society 176: 469 - 485. https: // doi. org / 10.1111 / boj. 12223" type="journal article" year="2014">
Rodríguez-Peña
<emphasis id="B96EEADEFFE11F71FB0EFD92FAF78A5F" box="[1238,1294,554,580]" italics="true" pageId="30" pageNumber="31">et al.</emphasis>
2014
</bibRefCitation>
)
</taxonomicName>
.
</paragraph>
<paragraph id="8BA536CCFFE11F71FF65FDF6FEF08B07" blockId="30.[136,1452,158,1265]" pageId="30" pageNumber="31">
In the south-central part of
<collectingCountry id="F30D765CFFE11F71FE20FDF7FD668A73" box="[504,671,590,616]" name="Dominican Republic" pageId="30" pageNumber="31">the Dominican</collectingCountry>
Republic specimens have fruits with projecting fibers (illustrated in
<bibRefCitation id="EF8B4B3DFFE11F71FF50FDCBFE1A8A97" author="Fernandez, E. &amp; Gottschalk, A." box="[136,483,626,652]" pageId="30" pageNumber="31" refId="ref61896" refString="Fernandez, E. &amp; Gottschalk, A. (2017) Palmas de Espanola. Grupo Sid, 159 pp." type="book" year="2017">Fernández &amp; Gottschalk 2017</bibRefCitation>
, page 102), somewhat different from fruit surfaces of other specimens. The
<typeStatus id="54A1886EFFE11F71FA8BFDCAFA558A96" box="[1363,1452,627,653]" pageId="30" pageNumber="31" type="neotype">neotype</typeStatus>
has fruits with projecting fibers.
<collectingCounty id="62C44E40FFE11F71FE20FD2FFD098AAB" box="[504,752,662,688]" pageId="30" pageNumber="31">The Botanical Garden</collectingCounty>
in
<collectingRegion id="49DEF82EFFE11F71FCCEFD2FFC3E8AAB" box="[790,967,662,688]" country="Dominican Republic" name="Santo Domingo" pageId="30" pageNumber="31">Santo Domingo</collectingRegion>
is situated within this south-central part of
<collectingCountry id="F30D765CFFE11F71FF50FD03FED78ACF" box="[136,302,698,724]" name="Dominican Republic" pageId="30" pageNumber="31">the Dominican</collectingCountry>
<collectingMunicipality id="6BC1ACB6FFE11F71FEEFFD03FE658ACF" box="[311,412,698,724]" pageId="30" pageNumber="31">Republic</collectingMunicipality>
and here, in an area presumed to be part of the original vegetation, plants have tall, slender stems that often curve or bend, and a small, compact crown of leaves.
<location id="8EC56017FFE11F71FC4FFD67FC368AE3" LSID="urn:lsid:plazi:treatment:03B387DAFFE41F71FF50FADFFD748CEA:8EC56017FFE11F71FC4FFD67FC368AE3" box="[919,975,734,760]" country="Dominican Republic" county="The Botanical Garden" municipality="Republic" name="Such" pageId="30" pageNumber="31" stateProvince="Santo Domingo">Such</location>
plants have not been seen in other parts of
<location id="8EC56017FFE11F71FF50FCBBFEFC8B07" LSID="urn:lsid:plazi:treatment:03B387DAFFE41F71FF50FADFFD748CEA:8EC56017FFE11F71FF50FCBBFEFC8B07" box="[136,261,770,796]" country="Dominican Republic" county="The Botanical Garden" municipality="Republic" name="Hispaniola" pageId="30" pageNumber="31" stateProvince="Santo Domingo">Hispaniola</location>
.
</paragraph>
<paragraph id="8BA536CCFFE11F71FF65FC9FFED38BEF" blockId="30.[136,1452,158,1265]" pageId="30" pageNumber="31">
There are two unusual populations in the peninsula part of southwestern
<collectingCountry id="F30D765CFFE11F71FBD0FC9FFBBD8B5B" box="[1032,1092,806,832]" name="Haiti" pageId="30" pageNumber="31">Haiti</collectingCountry>
. One specimen (
<emphasis id="B96EEADEFFE11F71FADDFC9EFA5B8B5B" box="[1285,1442,806,832]" italics="true" pageId="30" pageNumber="31">Zanoni 26116</emphasis>
) from the central part of the peninsula has obvious transverse veinlets and long pedicels (like
<taxonomicName id="4C1A4D4FFFE11F71FB7FFCF2FAA88B7F" authorityName="Beccari" authorityYear="1907" baseAuthorityName="Loddiges ex Martius" box="[1191,1361,842,869]" class="Liliopsida" family="Arecaceae" genus="Coccothrinax" kingdom="Plantae" order="Arecales" pageId="30" pageNumber="31" phylum="Tracheophyta" rank="species" species="barbadensis">
<emphasis id="B96EEADEFFE11F71FB7FFCF2FAA88B7F" box="[1191,1361,842,869]" italics="true" pageId="30" pageNumber="31">C. barbadensis</emphasis>
</taxonomicName>
). In the extreme southwestern part of the peninsula there are
<specimenCount id="9D1CFD45FFE11F71FD09FCD7FC828B93" box="[721,891,878,904]" pageId="30" pageNumber="31" type="generic">five specimens</specimenCount>
. Three have long pedicels but two (
<emphasis id="B96EEADEFFE11F71FADBFCD7FEBB8BB7" italics="true" pageId="30" pageNumber="31">Bartlett 17491, Henderson 1030</emphasis>
) have short pedicels like those of most specimens of
<taxonomicName id="4C1A4D4FFFE11F71FC46FC2AFBDB8BB7" authorityName="Sargent ex Schumann" authorityYear="1901" baseAuthorityName="Loddiges ex Schultes &amp; Schultes" box="[926,1058,915,941]" class="Liliopsida" family="Arecaceae" genus="Coccothrinax" kingdom="Plantae" order="Arecales" pageId="30" pageNumber="31" phylum="Tracheophyta" rank="species" species="argentea">
<emphasis id="B96EEADEFFE11F71FC46FC2AFBDB8BB7" box="[926,1058,915,941]" italics="true" pageId="30" pageNumber="31">C. argentea</emphasis>
</taxonomicName>
from Hispaniola. All specimens of this extreme southwestern population are from roadsides or disturbed, hilly areas, mostly between Cavaillon and St. Louis-du-Sud.
</paragraph>
<paragraph id="8BA536CCFFE11F71FF65FC47FD748CEA" blockId="30.[136,1452,158,1265]" pageId="30" pageNumber="31">
According to several specimen labels,
<taxonomicName id="4C1A4D4FFFE11F71FDA1FC46FD068C03" authorityName="Sargent ex Schumann" authorityYear="1901" baseAuthorityName="Loddiges ex Schultes &amp; Schultes" box="[633,767,1023,1049]" class="Liliopsida" family="Arecaceae" genus="Coccothrinax" kingdom="Plantae" order="Arecales" pageId="30" pageNumber="31" phylum="Tracheophyta" rank="species" species="argentea">
<emphasis id="B96EEADEFFE11F71FDA1FC46FD068C03" box="[633,767,1023,1049]" italics="true" pageId="30" pageNumber="31">C. argentea</emphasis>
</taxonomicName>
and
<taxonomicName id="4C1A4D4FFFE11F71FCE2FC46FC588C03" authorityName="Bailey" authorityYear="1939" box="[826,929,1023,1049]" class="Liliopsida" family="Arecaceae" genus="Coccothrinax" kingdom="Plantae" order="Arecales" pageId="30" pageNumber="31" phylum="Tracheophyta" rank="species" species="spissa">
<emphasis id="B96EEADEFFE11F71FCE2FC46FC588C03" box="[826,929,1023,1049]" italics="true" pageId="30" pageNumber="31">C. spissa</emphasis>
</taxonomicName>
occur together in mixed populations in some areas in Hispaniola. These two species can sometimes be difficult to distinguish and it seems likely that there are hybrids between them. Apart from its smaller size,
<taxonomicName id="4C1A4D4FFFE11F71FD0EFBFEFCA58C7B" authorityName="Sargent ex Schumann" authorityYear="1901" baseAuthorityName="Loddiges ex Schultes &amp; Schultes" box="[726,860,1095,1121]" class="Liliopsida" family="Arecaceae" genus="Coccothrinax" kingdom="Plantae" order="Arecales" pageId="30" pageNumber="31" phylum="Tracheophyta" rank="species" species="argentea">
<emphasis id="B96EEADEFFE11F71FD0EFBFEFCA58C7B" box="[726,860,1095,1121]" italics="true" pageId="30" pageNumber="31">C. argentea</emphasis>
</taxonomicName>
can usually be distinguished by its shorter, thinner, regularly arranged rachillae, It also usually has a persistent, triangular ligule (
<figureCitation id="13212A49FFE11F71FC06FBD3FBCE8C9F" box="[990,1079,1130,1156]" captionStart="FIGURE 2" captionStartId="10.[136,229,1917,1939]" captionTargetBox="[151,1433,194,1891]" captionTargetId="figure-27@10.[150,1434,193,1893]" captionTargetPageId="10" captionText="FIGURE 2. A. Sheath fibers thin (usually &lt;0.5 mm diameter), closely woven, forming persistent, triangular ligules at the apices (C. argentea). B. Sheath fibers thin (usually &lt;0.5 mm diameter), flimsy, loosely woven, free and greatly elongate at the apices (C. crinita). C. Sheath fibers stout (usually&gt;1 mm diameter), woody, loosely (rarely closely) woven, ± joined or briefly free at the apices (C. miraguama). D. Sheath fibers stout (usually&gt;1 mm diameter), woody, loosely woven, the inner and outer layer combining at the apices to form erect, spine-like fibers (C. salvatoris)." figureDoi="http://doi.org/10.5281/zenodo.8389315" httpUri="https://zenodo.org/record/8389315/files/figure.png" pageId="30" pageNumber="31">Fig. 2A</figureCitation>
). Specimens of
<taxonomicName id="4C1A4D4FFFE11F71FB3EFBD2FA918C9F" authorityName="Sargent ex Schumann" authorityYear="1901" baseAuthorityName="Loddiges ex Schultes &amp; Schultes" box="[1254,1384,1131,1157]" class="Liliopsida" family="Arecaceae" genus="Coccothrinax" kingdom="Plantae" order="Arecales" pageId="30" pageNumber="31" phylum="Tracheophyta" rank="species" species="argentea">
<emphasis id="B96EEADEFFE11F71FB3EFBD2FA918C9F" box="[1254,1384,1131,1157]" italics="true" pageId="30" pageNumber="31">C. argentea</emphasis>
</taxonomicName>
differ significantly from those of
<taxonomicName id="4C1A4D4FFFE11F71FE6DFB36FDE18CB3" authorityName="Bailey" authorityYear="1939" box="[437,536,1167,1193]" class="Liliopsida" family="Arecaceae" genus="Coccothrinax" kingdom="Plantae" order="Arecales" pageId="30" pageNumber="31" phylum="Tracheophyta" rank="species" species="spissa">
<emphasis id="B96EEADEFFE11F71FE6DFB36FDE18CB3" box="[437,536,1167,1193]" italics="true" pageId="30" pageNumber="31">C. spissa</emphasis>
</taxonomicName>
in 10 variables (stem diameter, petiole width, palman length, number of segments, segment length, segment width, rachilla length, rachillae width, fruit length, fruit diameter), with
<taxonomicName id="4C1A4D4FFFE11F71FB0CFB0AFAA18CD7" authorityName="Sargent ex Schumann" authorityYear="1901" baseAuthorityName="Loddiges ex Schultes &amp; Schultes" box="[1236,1368,1203,1229]" class="Liliopsida" family="Arecaceae" genus="Coccothrinax" kingdom="Plantae" order="Arecales" pageId="30" pageNumber="31" phylum="Tracheophyta" rank="species" species="argentea">
<emphasis id="B96EEADEFFE11F71FB0CFB0AFAA18CD7" box="[1236,1368,1203,1229]" italics="true" pageId="30" pageNumber="31">C. argentea</emphasis>
</taxonomicName>
having lower values for all variables (
<emphasis id="B96EEADEFFE11F71FE05FB6EFE178CEB" box="[477,494,1239,1264]" italics="true" pageId="30" pageNumber="31">t-</emphasis>
test,
<emphasis id="B96EEADEFFE11F71FDFBFB6EFDCD8CEB" box="[547,564,1239,1264]" italics="true" pageId="30" pageNumber="31">P</emphasis>
&lt;0.05).
</paragraph>
</subSubSection>
</treatment>
</document>