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<document id="91ADE942AD7A7C6B2E356887DDF144CF" ID-DOI="10.11646/phytotaxa.614.1.1" ID-ISSN="1179-3163" ID-Zenodo-Dep="8389307" IM.bibliography_approvedBy="diego" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="diego" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="diego" IM.treatmentCitations_approvedBy="diego" IM.treatments_approvedBy="diego" checkinTime="1695979127173" checkinUser="plazi" docAuthor="Henderson, Andrew" docDate="2023" docId="03B387DAFFDC1F4AFF50FAD7FC07895E" docLanguage="en" docName="phytotaxa.614.1.1.pdf" docOrigin="Phytotaxa 614 (1)" docSource="http://dx.doi.org/10.11646/phytotaxa.614.1.1" docStyle="DocumentStyle:F08184CE06D8A97EA3E6DE35D99648B0.3:Phytotaxa.2014-.monograph" docStyleId="F08184CE06D8A97EA3E6DE35D99648B0" docStyleName="Phytotaxa.2014-.monograph" docStyleVersion="3" docTitle="Coccothrinax bonettiana Henderson, Fernandez &amp; Bacon" docType="treatment" docVersion="1" lastPageNumber="38" masterDocId="FF8AFFA2FFFF1F6FFFD8FFB9FFF9881B" masterDocTitle="A revision of Coccothrinax, Hemithrinax, Leucothrinax, Thrinax, and Zombia (Arecaceae)" masterLastPageNumber="115" masterPageNumber="1" pageNumber="36" updateTime="1696265644834" updateUser="diego">
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<mods:title id="6BD4C25BBF45666426D37C8C5677870E">A revision of Coccothrinax, Hemithrinax, Leucothrinax, Thrinax, and Zombia (Arecaceae)</mods:title>
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<mods:date id="4CE8EC67458A6CC47E635D3C9778F064">2023</mods:date>
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<mods:identifier id="C00B2118A917BB2500A90D1205DB3EB1" type="DOI">10.11646/phytotaxa.614.1.1</mods:identifier>
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<treatment id="03B387DAFFDC1F4AFF50FAD7FC07895E" LSID="urn:lsid:plazi:treatment:03B387DAFFDC1F4AFF50FAD7FC07895E" httpUri="http://treatment.plazi.org/id/03B387DAFFDC1F4AFF50FAD7FC07895E" lastPageId="37" lastPageNumber="38" pageId="35" pageNumber="36">
<subSubSection id="C3006547FFDC1F4CFF50FAD7FAB98D93" box="[136,1344,1389,1417]" pageId="35" pageNumber="36" type="nomenclature">
<paragraph id="8BA536CCFFDC1F4CFF50FAD7FAB98D93" blockId="35.[136,1452,1389,1522]" box="[136,1344,1389,1417]" pageId="35" pageNumber="36">
<emphasis id="B96EEADEFFDC1F4CFF50FAD7FAB98D93" bold="true" box="[136,1344,1389,1417]" pageId="35" pageNumber="36">
1.6.
<taxonomicName id="4C1A4D4FFFDC1F4CFF1AFAD7FC6A8D93" authority="Henderson, Fernandez &amp; Bacon" authorityName="Henderson, Fernandez &amp; Bacon" box="[194,915,1389,1417]" class="Liliopsida" family="Arecaceae" genus="Coccothrinax" kingdom="Plantae" order="Arecales" pageId="35" pageNumber="36" phylum="Tracheophyta" rank="species" species="bonettiana">
<emphasis id="B96EEADEFFDC1F4CFF1AFAD7FE108D92" bold="true" box="[194,489,1390,1417]" italics="true" pageId="35" pageNumber="36">Coccothrinax bonettiana</emphasis>
Henderson, Fernández &amp; Bacon
</taxonomicName>
in
<bibRefCitation id="EF8B4B3DFFDC1F4CFC11FAD7FAC38D93" author="Henderson, A. &amp; Fernandez, E. &amp; Montero, O. &amp; Clase, T. &amp; Gratacos, X. &amp; Bacon, C." box="[969,1338,1390,1417]" pageId="35" pageNumber="36" pagination="101 - 111" refId="ref62198" refString="Henderson, A., Fernandez, E., Montero, O., Clase, T., Gratacos, X. &amp; Bacon, C. (2023) Five new species of Coccothrinax (Cryosophileae, Coryphoideae, Arecaceae) from Hispaniola. Phytotaxa 589 (2): 101 - 111. https: // doi. org / 10.11646 / phytotaxa. 589.2.1" type="journal article" year="2023">
Henderson
<emphasis id="B96EEADEFFDC1F4CFB84FAD6FB658D92" bold="true" box="[1116,1180,1390,1417]" italics="true" pageId="35" pageNumber="36">et al.</emphasis>
(2023: 102)
</bibRefCitation>
.
</emphasis>
</paragraph>
</subSubSection>
<subSubSection id="C3006547FFDC1F4CFA88FAD6FC658DE9" pageId="35" pageNumber="36" type="materials_examined">
<paragraph id="8BA536CCFFDC1F4CFA88FAD6FC658DE9" blockId="35.[136,1452,1389,1522]" pageId="35" pageNumber="36">
<materialsCitation id="3B723C91FFDC1F4CFA88FAD6FC658DE9" collectingDate="2021-12-19" collectionCode="O, T, C, E, JBSD, NY" country="Dominican Republic" county="Type" elevation="22" latitude="19.616667" location="Municipio Gaspar Hernandez" longLatPrecision="1271" longitude="-70.21667" municipality="Provincia" pageId="35" pageNumber="36" specimenCount="1" stateProvince="Espaillat" typeStatus="holotype">
<collectingCounty id="62C44E40FFDC1F4CFA88FAD6FA728D92" box="[1360,1419,1391,1417]" pageId="35" pageNumber="36">Type</collectingCounty>
:—
<collectingCountry id="F30D765CFFDC1F4CFF50FA2BFE478DB7" box="[136,446,1426,1452]" name="Dominican Republic" pageId="35" pageNumber="36">DOMINICAN REPUBLIC</collectingCountry>
.
<collectingMunicipality id="6BC1ACB6FFDC1F4CFE12FA28FDCC8DB0" box="[458,565,1425,1451]" pageId="35" pageNumber="36">Provincia</collectingMunicipality>
<collectingRegion id="49DEF82EFFDC1F4CFDE5FA28FD5B8DB0" box="[573,674,1425,1451]" country="Dominican Republic" name="Espaillat" pageId="35" pageNumber="36">Espaillat</collectingRegion>
,
<location id="8EC56017FFDC1F4CFD76FA28FC078DB0" LSID="urn:lsid:plazi:treatment:03B387DAFFDC1F4AFF50FAD7FC07895E:8EC56017FFDC1F4CFD76FA28FC078DB0" box="[686,1022,1425,1452]" country="Dominican Republic" county="Type" latitude="19.616667" longLatPrecision="1271" longitude="-70.21667" municipality="Provincia" name="Municipio Gaspar Hernandez" pageId="35" pageNumber="36" stateProvince="Espaillat">Municipio Gaspar Hernández</location>
, saliendo del
<location id="8EC56017FFDC1F4CFB42FA28FA798DB0" LSID="urn:lsid:plazi:treatment:03B387DAFFDC1F4AFF50FAD7FC07895E:8EC56017FFDC1F4CFB42FA28FA798DB0" box="[1178,1408,1425,1452]" country="Dominican Republic" county="Type" latitude="19.616667" longLatPrecision="1271" longitude="-70.21667" municipality="Provincia" name="Poblado Rio Piedras" pageId="35" pageNumber="36" stateProvince="Espaillat">
Poblado
<location id="8EC56017FFDC1F4CFB26FA2BFA798DB0" LSID="urn:lsid:plazi:treatment:03B387DAFFDC1F4AFF50FAD7FC07895E:8EC56017FFDC1F4CFB26FA2BFA798DB0" box="[1278,1408,1425,1452]" country="Dominican Republic" county="Type" latitude="19.616667" longLatPrecision="1271" longitude="-70.21667" municipality="Provincia" name="Rio Piedras" pageId="35" pageNumber="36" stateProvince="Espaillat">Río Piedras</location>
</location>
a
<quantity id="4CE29B29FFDC1F4CFA45FA2BFF558DD4" metricMagnitude="3" metricUnit="m" metricValue="2.0" pageId="35" pageNumber="36" unit="km" value="2.0">2 km</quantity>
a lo largo de la carretera, en una finca privada,
<geoCoordinate id="EE2E500BFFDC1F4CFD66FA0CFCE58DD4" box="[702,796,1461,1487]" degrees="19" direction="north" minutes="37" orientation="latitude" pageId="35" pageNumber="36" precision="925" value="19.616667">19º37N</geoCoordinate>
<geoCoordinate id="EE2E500BFFDC1F4CFCFBFA0CFC7E8DD4" box="[803,903,1461,1487]" degrees="70" direction="west" minutes="13" orientation="longitude" pageId="35" pageNumber="36" precision="925" value="-70.21667">70º13W</geoCoordinate>
,
<quantity id="4CE29B29FFDC1F4CFC4AFA0CFC348DD4" box="[914,973,1461,1487]" metricMagnitude="1" metricUnit="m" metricValue="2.2" pageId="35" pageNumber="36" unit="m" value="22.0">
<elevation id="0037D1FFFFDC1F4CFC4AFA0CFC348DD4" box="[914,973,1461,1487]" metricMagnitude="1" metricUnit="m" metricValue="2.2" pageId="35" pageNumber="36" unit="m" value="22.0">22 m</elevation>
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,
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<collectingDate id="EFE0E9E4FFDC1F4CFC00FA0CFB498DD4" box="[984,1200,1461,1487]" pageId="35" pageNumber="36" value="2021-12-19">19 December 2021</collectingDate>
</date>
,
<emphasis id="B96EEADEFFDC1F4CFB62FA0CFE258DE9" italics="true" pageId="35" pageNumber="36">
<collectionCode id="ED0BAE09FFDC1F4CFB62FA0CFB288DD4" box="[1210,1233,1461,1487]" country="Norway" lsid="urn:lsid:biocol.org:col:13093" name="Botanical Museum - University of Oslo" pageId="35" pageNumber="36" type="Herbarium">O</collectionCode>
. Montero,
<collectionCode id="ED0BAE09FFDC1F4CFA91FA0CFAA38DD5" box="[1353,1370,1461,1486]" name="Tavera, Department of Geology and Geophysics" pageId="35" pageNumber="36">T</collectionCode>
. Clase,
<collectionCode id="ED0BAE09FFDC1F4CFF50FA61FF678DE9" box="[136,158,1496,1522]" country="Denmark" name="University of Copenhagen" pageId="35" pageNumber="36" type="Herbarium">C</collectionCode>
. Bacon &amp;
<collectionCode id="ED0BAE09FFDC1F4CFECCFA60FED08DE9" box="[276,297,1497,1522]" country="United Kingdom" lsid="urn:lsid:biocol.org:col:15670" name="Royal Botanic Garden Edinburgh" pageId="35" pageNumber="36" type="Herbarium">E</collectionCode>
. Fernández 435
</emphasis>
(
<typeStatus id="54A1886EFFDC1F4CFE32FA61FDB78DE9" box="[490,590,1496,1522]" pageId="35" pageNumber="36">holotype</typeStatus>
<collectionCode id="ED0BAE09FFDC1F4CFD8DFA61FD668DE9" box="[597,671,1496,1522]" country="Dominican Republic" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15541" lsid="urn:lsid:biocol.org:col:15541" name="Jardín Botánico Nacional Dr. Rafael M. Moscoso" pageId="35" pageNumber="36" type="Herbarium">JBSD</collectionCode>
!, isotype
<collectionCode id="ED0BAE09FFDC1F4CFCDEFA61FCCE8DE9" box="[774,823,1496,1522]" country="USA" lsid="urn:lsid:biocol.org:col:15556" name="William and Lynda Steere Herbarium of the New York Botanical Garden" pageId="35" pageNumber="36" type="Herbarium">NY</collectionCode>
!).
<figureCitation id="13212A49FFDC1F4CFC97FA61FC658DE9" box="[847,924,1496,1522]" captionStart="PLATE 6" captionStartId="36.[136,214,1930,1951]" captionTargetBox="[151,1433,194,1904]" captionTargetId="figure-27@36.[150,1435,193,1906]" captionTargetPageId="36" captionText="PLATE 6. Coccothrinax bonettiana, Dominican Republic, with leaf segments pendulous at the apices, giving a three-dimensional appearance to the leaf. (Image by Eladio Fernández)." figureDoi="http://doi.org/10.5281/zenodo.8389361" httpUri="https://zenodo.org/record/8389361/files/figure.png" pageId="35" pageNumber="36">Plate 6</figureCitation>
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C3006547FFDC1F4CFF50F9A7FC1F803E" pageId="35" pageNumber="36" type="description">
<paragraph id="8BA536CCFFDC1F4CFF50F9A7FC1F803E" blockId="35.[136,1452,1566,2086]" pageId="35" pageNumber="36">
<emphasis id="B96EEADEFFDC1F4CFF50F9A7FF298E23" bold="true" box="[136,208,1566,1592]" pageId="35" pageNumber="36">Stems</emphasis>
4.1(2.86.5) m long, diameter not recorded, solitary.
<emphasis id="B96EEADEFFDC1F4CFCC5F9A7FC968E23" bold="true" box="[797,879,1566,1592]" pageId="35" pageNumber="36">Leaves</emphasis>
more or less deciduous or only leaf bases persisting on stem; leaf sheath fibers 1.2(0.82.0) mm diameter, stout, woody, loosely woven, ± joined or briefly free at the apices; petioles 8.3(6.99.9) mm diameter just below the apex; palmans 8.5(6.410.8) cm long, relatively short, with the adaxial veins prominent and terminating in a slight raised ridge and distinct pulvinus; leaf blades not wedge-shaped; segments 33(3039) per leaf, the middle ones 62.6(58.070.0) cm long and 2.9(2.33.7) cm wide; segments pendulous at the apices, giving a three-dimensional appearance to the leaf; middle leaf segments relatively long and narrow, tapering from base to apex, scarcely folded, flexible and not leathery, a shoulder or constriction absent or poorly developed, the apices thin, deeply splitting and breaking off; middle leaf segment apices attenuate; leaf segments not waxy or sometimes with a deciduous, thin layer of wax adaxially; leaf segments without indumentum, scales, or wax abaxially, with well-developed transverse veinlets.
<emphasis id="B96EEADEFFDC1F4CFD19F8E2FC938F6E" bold="true" box="[705,874,1883,1909]" pageId="35" pageNumber="36">Inflorescences</emphasis>
erect, at least initially, amongst or above the leaves, with few partial inflorescences; rachis bracts narrow, closely sheathing, sparsely tomentose, usually without hairs at the apex; partial inflorescences 4(35); proximalmost rachillae straight, 7.7(6.610.3) cm long and
<quantity id="4CE29B29FFDC1F4CFB63F81BFABB8FA7" box="[1211,1346,1954,1980]" metricMagnitude="-3" metricUnit="m" metricValue="1.25" metricValueMax="1.3" metricValueMin="1.2" pageId="35" pageNumber="36" unit="mm" value="1.25" valueMax="1.3" valueMin="1.2">1.21.3 mm</quantity>
diameter in fruit; rachillae glabrous at or near anthesis; stamens 7; fruit pedicels 0.4(0.20.7) mm long;
<emphasis id="B96EEADEFFDC1F4CFB7FF87DFB118FC5" bold="true" box="[1191,1256,1988,2014]" pageId="35" pageNumber="36">fruits</emphasis>
7.5(7.17.9) mm long and 8.2(7.58.8) mm diameter, color not recorded; fruit surfaces smooth or sometimes with projecting fibers; seed surfaces deeply lobed, the lobes running from base of seeds almost to apices.
</paragraph>
</subSubSection>
<caption id="DF656644FFDB1F4BFF50F833FD7F8FDF" ID-DOI="http://doi.org/10.5281/zenodo.8389361" ID-Zenodo-Dep="8389361" httpUri="https://zenodo.org/record/8389361/files/figure.png" pageId="36" pageNumber="37" startId="36.[136,214,1930,1951]" targetBox="[151,1433,194,1904]" targetPageId="36" targetType="plate">
<paragraph id="8BA536CCFFDB1F4BFF50F833FD7F8FDF" blockId="36.[136,1451,1930,1988]" pageId="36" pageNumber="37">
<emphasis id="B96EEADEFFDB1F4BFF50F833FF148F84" bold="true" box="[136,237,1930,1951]" pageId="36" pageNumber="37">PLATE 6</emphasis>
.
<taxonomicName id="4C1A4D4FFFDB1F4BFF25F833FE088F84" box="[253,497,1930,1951]" class="Liliopsida" family="Arecaceae" genus="Coccothrinax" kingdom="Plantae" order="Arecales" pageId="36" pageNumber="37" phylum="Tracheophyta" rank="species" species="bonettiana">
<emphasis id="B96EEADEFFDB1F4BFF25F833FE088F84" box="[253,497,1930,1951]" italics="true" pageId="36" pageNumber="37">Coccothrinax bonettiana</emphasis>
</taxonomicName>
, Dominican Republic, with leaf segments pendulous at the apices, giving a three-dimensional appearance to the leaf. (Image by Eladio Fernández).
</paragraph>
</caption>
<subSubSection id="C3006547FFDA1F4AFF65FF27FD9588C0" pageId="37" pageNumber="38" type="distribution">
<paragraph id="8BA536CCFFDA1F4AFF65FF27FD9588C0" blockId="37.[136,1452,158,326]" pageId="37" pageNumber="38">
<emphasis id="B96EEADEFFDA1F4AFF65FF27FDEB88A3" bold="true" box="[189,530,158,184]" pageId="37" pageNumber="38">Distribution and habitat:—</emphasis>
North-central
<collectingCountry id="F30D765CFFDA1F4AFD6EFF27FC5D88A3" box="[694,932,158,184]" name="Dominican Republic" pageId="37" pageNumber="38">Dominican Republic</collectingCountry>
(
<collectingRegion id="49DEF82EFFDA1F4AFC61FF27FBE788A3" box="[953,1054,158,184]" country="Dominican Republic" name="Espaillat" pageId="37" pageNumber="38">Espaillat</collectingRegion>
) (
<figureCitation id="13212A49FFDA1F4AFBE4FF27FB6A88A3" box="[1084,1171,158,184]" captionStart="FIGURE 10" captionStartId="32.[136,229,1508,1530]" captionTargetBox="[155,1429,198,1479]" captionTargetId="figure-27@32.[150,1434,193,1484]" captionTargetPageId="32" captionText="FIGURE 10. Distribution of Coccothrinax baracoensis, C. bonettiana, C. borhidiana, and C. boschiana." figureDoi="http://doi.org/10.5281/zenodo.8389355" httpUri="https://zenodo.org/record/8389355/files/figure.png" pageId="37" pageNumber="38">Fig. 10</figureCitation>
) in broadleaf forest on serpentine soils at 80(22230) m elevation.
</paragraph>
</subSubSection>
<subSubSection id="C3006547FFDA1F4AFF65FF5DFC07895E" pageId="37" pageNumber="38" type="discussion">
<paragraph id="8BA536CCFFDA1F4AFF65FF5DFC07895E" blockId="37.[136,1452,158,326]" pageId="37" pageNumber="38">
<emphasis id="B96EEADEFFDA1F4AFF65FF5DFE5588E4" bold="true" box="[189,428,228,255]" pageId="37" pageNumber="38">Taxonomic notes:—</emphasis>
As a preliminary species,
<taxonomicName id="4C1A4D4FFFDA1F4AFD17FF5CFC1E88E4" box="[719,999,229,255]" class="Liliopsida" family="Arecaceae" genus="Coccothrinax" kingdom="Plantae" order="Arecales" pageId="37" pageNumber="38" phylum="Tracheophyta" rank="species" species="bonettiana">
<emphasis id="B96EEADEFFDA1F4AFD17FF5CFC1E88E4" box="[719,999,229,255]" italics="true" pageId="37" pageNumber="38">Coccothrinax bonettiana</emphasis>
</taxonomicName>
has a unique combination of qualitative character states and is recognized as a phylogenetic species. It is notable for its stout, woody, loosely woven leaf sheath fibers and segments without any indumentum abaxially, and serpentine habitat.
</paragraph>
</subSubSection>
</treatment>
</document>