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<document id="F8443D03BBB3A18D350C163ACFD5A8D7" ID-DOI="10.5281/zenodo.5330604" ID-ISSN="0374-1036" ID-Zenodo-Dep="5330604" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1630168017471" checkinUser="felipe" docAuthor="Fikáček, Martin, Prokin, Alexander, Angus, Robert B., Pono, Alexander, Marenko, Yue, Yanli, Ren, Dong &amp; Prokop, Jakub" docDate="2012" docId="B970A055FFCEFFEBFE78264CFBE4FC4A" docLanguage="en" docName="ActaEntMusNatPra.2012.52.1.89-127.pdf" docOrigin="Acta Entomologica Musei Nationalis Pragae 52 (1)" docStyle="DocumentStyle:E9BE312A6EDF9B84FC5211AAA80D80BF.10:ActaEntMusNatPra.2005-2016.journal_article" docStyleId="E9BE312A6EDF9B84FC5211AAA80D80BF" docStyleName="ActaEntMusNatPra.2005-2016.journal_article" docStyleVersion="10" docTitle="Helophorus Fabricius 1775" docType="treatment" docVersion="5" lastPageNumber="103" masterDocId="4549D82DFFC0FFE5FF9B2756FFB9FFFB" masterDocTitle="Revision of Mesozoic fossils of the helophorid lineage of the superfamily Hydrophiloidea (Coleoptera: Polyphaga)" masterLastPageNumber="127" masterPageNumber="89" pageNumber="103" updateTime="1699171640213" updateUser="plazi" zenodo-license-document="CC-BY-4.0">
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<mods:title id="C61692D79945856AC889BC8856AFE65A">Revision of Mesozoic fossils of the helophorid lineage of the superfamily Hydrophiloidea (Coleoptera: Polyphaga)</mods:title>
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<mods:namePart id="D7A824BBD0CAFD3D3112E6591166686D">Fikáček, Martin</mods:namePart>
<mods:affiliation id="F922A460216DC16F4E161E71557A2B4D">Department of Entomology, National Museum, Kunratice 1, CZ- 148 00 Praha 4, Czech Republic; e-mail: mfikacek @ gmail. com &amp; Department of Zoology, Faculty of Science, Charles University in Prague, Viničná 7, CZ- 128 44 Praha 2, Czech Republic; e-mail: jprokop @ natur. cuni. cz</mods:affiliation>
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<mods:namePart id="FF14F0603BB0642866C0307970F363B2">Prokin, Alexander</mods:namePart>
<mods:affiliation id="3A33417DA037446442282689F4024B2E">Papanin Institute for Inland Water Biology, Russian Academy of Science, Borok, Russia; e-mail: prokina @ mail. ru &amp; Voronezh State University, Research-Educational Centre “ Venevitinovo ”, Voronezh, Russia</mods:affiliation>
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<mods:namePart id="BD91D4E7AFCBFD0B4E460D4FB956FE23">Angus, Robert B.</mods:namePart>
<mods:affiliation id="CA16B9F0A9BB44303481AB5FA101A3BA">Department of Entomology, The Natural History Museum, London, UK; e-mail: r. angus @ rhul. ac. uk</mods:affiliation>
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<mods:namePart id="A707F523229556DAC4C133C01D1DE940">Pono, Alexander</mods:namePart>
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<mods:namePart id="8AB2F555C848FC45E032A0942569B080">Marenko</mods:namePart>
<mods:affiliation id="342CD042912F482EA4E63CCEEA973B6F">Borissiak Paleontological Institute, Russian Academy of Sciences, Moscow, Russia; e-mail: aponom @ paleo. ru</mods:affiliation>
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<mods:namePart id="7BC6C5CABF4DACA205BE0E0543B73F8C">Yue, Yanli</mods:namePart>
<mods:affiliation id="DAB443D0DF275B845FFE45442C25ADE0">School of Life Science, Ningxia University, Ningxia, China; e-mail: yueyanli 0407 @ yahoo. cn</mods:affiliation>
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<mods:namePart id="9EEC47B2F0415AFFBDD14701DEE84BAC">Ren, Dong</mods:namePart>
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<mods:namePart id="C22D06439568D7A59E4520A62AAA7452">Prokop, Jakub</mods:namePart>
<mods:affiliation id="8050A0DA398CD993400BF38B27155F1C">Department of Zoology, Faculty of Science, Charles University in Prague, Viničná 7, CZ- 128 44 Praha 2, Czech Republic; e-mail: jprokop @ natur. cuni. cz</mods:affiliation>
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<mods:title id="48CCE14CEDA036EA46986726C39D848A">Acta Entomologica Musei Nationalis Pragae</mods:title>
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<mods:date id="6F1D5F5EF1E19C4F847DB042831FE507">2012</mods:date>
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<treatment id="B970A055FFCEFFEBFE78264CFBE4FC4A" ID-DOI="http://doi.org/10.5281/zenodo.5398641" ID-Zenodo-Dep="5398641" LSID="urn:lsid:plazi:treatment:B970A055FFCEFFEBFE78264CFBE4FC4A" httpUri="http://treatment.plazi.org/id/B970A055FFCEFFEBFE78264CFBE4FC4A" lastPageNumber="103" pageId="14" pageNumber="103">
<subSubSection id="79C342C8FFCEFFEBFE78264CFCA3FEC8" box="[483,794,282,307]" pageId="14" pageNumber="103" type="nomenclature">
<paragraph id="31661143FFCEFFEBFE78264CFCA3FEC8" blockId="14.[483,794,282,307]" box="[483,794,282,307]" pageId="14" pageNumber="103">
<heading id="6A2EA62FFFCEFFEBFE78264CFCA3FEC8" bold="true" box="[483,794,282,307]" centered="true" fontSize="8" level="2" pageId="14" pageNumber="103" reason="3">
<taxonomicName id="F6D96AC0FFCEFFEBFE78264CFCA3FEC8" ID-CoL="8VRQF" authority="Fabricius, 1775" authorityName="Fabricius" authorityYear="1775" box="[483,794,282,307]" class="Insecta" family="Helophoridae" genus="Helophorus" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="103" phylum="Arthropoda" rank="genus">
<emphasis id="03ADCD51FFCEFFEBFE78264CFCA3FEC8" bold="true" box="[483,794,282,307]" pageId="14" pageNumber="103">
<emphasis id="03ADCD51FFCEFFEBFE78264CFDDCFEC8" bold="true" box="[483,613,282,307]" italics="true" pageId="14" pageNumber="103">Helophorus</emphasis>
Fabricius, 1775
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<typeStatus id="EE62AFE1FFCEFFEBFF0C261CFF76FE99" box="[151,207,330,354]" pageId="14" pageNumber="103">Type</typeStatus>
species.
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<taxonomicName id="F6D96AC0FFCEFFEBFEA8261DFD31FE99" authority="Linnaeus, 1758" authorityName="Linnaeus" authorityYear="1758" box="[307,648,330,354]" class="Insecta" family="Silphidae" genus="Silpha" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="103" phylum="Arthropoda" rank="species" species="aquatica">
<emphasis id="03ADCD51FFCEFFEBFEA8261DFE65FE99" box="[307,476,331,354]" italics="true" pageId="14" pageNumber="103">Silpha aquatica</emphasis>
Linnaeus, 1758
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(=
<taxonomicName id="F6D96AC0FFCEFFEBFD34261DFF6FFE79" ID-CoL="3KHHV" authority="(Linnaeus, 1758)" baseAuthorityName="Linnaeus" baseAuthorityYear="1758" class="Insecta" family="Helophoridae" genus="Helophorus" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="103" phylum="Arthropoda" rank="species" species="aquaticus">
<emphasis id="03ADCD51FFCEFFEBFD34261DFC94FE99" box="[687,813,331,354]" italics="true" pageId="14" pageNumber="103">Helophorus</emphasis>
(s. str.)
<emphasis id="03ADCD51FFCEFFEBFC1D261DFC57FE99" box="[902,1006,331,354]" italics="true" pageId="14" pageNumber="103">aquaticus</emphasis>
(Linnaeus, 1758)
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(modern)), designated by
<bibRefCitation id="55486CB2FFCEFFEBFE6B263DFD11FE79" author="LATREILLE P. A." box="[496,680,362,386]" pageId="14" pageNumber="103" refId="ref15345" refString="LATREILLE P. A. 1810: Considerations generales sur l'Ordre naturel des Animaux composant les Classes des Crustaces, des Arachnides, et des Insectes. F. Schoell, Paris, 444 pp." type="book" year="1810">LATREILLE (1810)</bibRefCitation>
.
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<emphasis id="03ADCD51FFCEFFEBFF0C26DCFE98FE59" bold="true" box="[151,289,394,418]" pageId="14" pageNumber="103">Time range.</emphasis>
Late Jurassic (ca. 150146 mya) to recent.
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<subSubSection id="79C342C8FFCEFFEBFF0C26EFFBE4FC4A" pageId="14" pageNumber="103" type="diagnosis">
<paragraph id="31661143FFCEFFEBFF0C26EFFB90FD0A" blockId="14.[151,1127,440,946]" pageId="14" pageNumber="103">
<emphasis id="03ADCD51FFCEFFEBFF0C26EFFEE0FE2A" bold="true" box="[151,345,440,465]" pageId="14" pageNumber="103">
Diagnosis.
<emphasis id="03ADCD51FFCEFFEBFE8826EEFEE9FE2A" bold="true" box="[275,336,440,465]" italics="true" pageId="14" pageNumber="103">Adult</emphasis>
:
</emphasis>
Head and pronotum at least partly with setiferous granules in most species (granulation totally reduced in few species only); frontoclypeal suture very distinct, its median portion grooved; mentum distinctly transverse or only slightly wider than long; gula usually strongly constricted (moderately wide with well-separated gular sutures in few species only); pronotum with one to five longitudinal grooves (if less than five grooves are present, then all pronotal intervals bear large and very distinct setiferous granules); lateral margin of pronotum at least slightly crenulate; anterolateral portion of hypomeron with antennal groove; mesanepisterna not meeting mesally; mesoventrite very narrow at anterior margin, bearing a transverse ridge posteromedially; elytra of variable coloration, but never pale with dark stripes along elytral series; elytra in some species with elevated or costate alternate intervals.
</paragraph>
<paragraph id="31661143FFCEFFEBFF2E25ACFBE4FC4A" blockId="14.[151,1127,440,946]" pageId="14" pageNumber="103">
<emphasis id="03ADCD51FFCEFFEBFF2E25ACFEB9FCEA" bold="true" box="[181,256,761,785]" pageId="14" pageNumber="103">
<emphasis id="03ADCD51FFCEFFEBFF2E25ACFF4FFCEA" bold="true" box="[181,246,762,785]" italics="true" pageId="14" pageNumber="103">Larva</emphasis>
:
</emphasis>
Head prognathous; nasale simply triangular or with lobate lateral margins; epistomal lobes large, slightly overlapping nasale, bearing series of stout setae; mandible with two retinacular teeth; labium without ligula; each parietale with 6 stemmata situated in area of darker cuticle; all thoracic segments with large dorsal tergite; abdominal segments 18 each with a pair of dorsal sclerites and an additional large sclerite laterally of them on each side; tracheal system holopneustic, spiracular atrium absent; urogomphi large, three-segmented.
</paragraph>
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