210 lines
23 KiB
XML
210 lines
23 KiB
XML
<document id="55B8563CE189736C4C47D3FD858C9458" ID-DOI="10.5281/zenodo.5330604" ID-ISSN="0374-1036" ID-Zenodo-Dep="5330604" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1630168017471" checkinUser="felipe" docAuthor="Fikáček, Martin, Prokin, Alexander, Angus, Robert B., Pono, Alexander, Marenko, Yue, Yanli, Ren, Dong & Prokop, Jakub" docDate="2012" docId="B970A055FFCAFFEFFE5B278DFD9DFB48" docLanguage="en" docName="ActaEntMusNatPra.2012.52.1.89-127.pdf" docOrigin="Acta Entomologica Musei Nationalis Pragae 52 (1)" docStyle="DocumentStyle:E9BE312A6EDF9B84FC5211AAA80D80BF.10:ActaEntMusNatPra.2005-2016.journal_article" docStyleId="E9BE312A6EDF9B84FC5211AAA80D80BF" docStyleName="ActaEntMusNatPra.2005-2016.journal_article" docStyleVersion="10" docTitle="Cretotaenia Ponomarenko 1977" docType="treatment" docVersion="4" lastPageNumber="99" masterDocId="4549D82DFFC0FFE5FF9B2756FFB9FFFB" masterDocTitle="Revision of Mesozoic fossils of the helophorid lineage of the superfamily Hydrophiloidea (Coleoptera: Polyphaga)" masterLastPageNumber="127" masterPageNumber="89" pageNumber="99" updateTime="1699171640213" updateUser="plazi" zenodo-license-document="CC-BY-4.0">
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<mods:title id="FF8840FB4DE7C2C60036DAA753B890DD">Revision of Mesozoic fossils of the helophorid lineage of the superfamily Hydrophiloidea (Coleoptera: Polyphaga)</mods:title>
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<mods:namePart id="BCE72F4C1E77E26254CCAB1E80DDCDC6">Fikáček, Martin</mods:namePart>
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<mods:affiliation id="D8455C666FCFDD4B8EAFC05418E248CA">Department of Entomology, National Museum, Kunratice 1, CZ- 148 00 Praha 4, Czech Republic; e-mail: mfikacek @ gmail. com & Department of Zoology, Faculty of Science, Charles University in Prague, Viničná 7, CZ- 128 44 Praha 2, Czech Republic; e-mail: jprokop @ natur. cuni. cz</mods:affiliation>
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<mods:namePart id="576B5DDA01D714A47086676C824A2CBB">Prokin, Alexander</mods:namePart>
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<mods:affiliation id="037728596E71AB0827A038C408040EAE">Papanin Institute for Inland Water Biology, Russian Academy of Science, Borok, Russia; e-mail: prokina @ mail. ru & Voronezh State University, Research-Educational Centre “ Venevitinovo ”, Voronezh, Russia</mods:affiliation>
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</mods:name>
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<mods:name id="96ED005F1000A5A5674B6F31F268CDBA" type="personal">
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<mods:namePart id="C9E730A4E9070CE44A8B3238346F9EA1">Angus, Robert B.</mods:namePart>
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<mods:affiliation id="583A9E5C9007149EC79023CC40A77867">Department of Entomology, The Natural History Museum, London, UK; e-mail: r. angus @ rhul. ac. uk</mods:affiliation>
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<mods:namePart id="E163AB23C580F58EAFB2438F37F09C41">Pono, Alexander</mods:namePart>
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<mods:roleTerm id="F179354AF5D662CA3417E8986F534084">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="166164633CF8AC98735166822259D091">Marenko</mods:namePart>
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<mods:affiliation id="7946D044FAAC247264E279DF6CC12C63">Borissiak Paleontological Institute, Russian Academy of Sciences, Moscow, Russia; e-mail: aponom @ paleo. ru</mods:affiliation>
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</mods:name>
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<mods:name id="8B7FF118CFC66D7790FAF636E5A16A6E" type="personal">
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<mods:roleTerm id="D07E1678C43AD7FAEF918BD23FE7E290">Author</mods:roleTerm>
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<mods:namePart id="47A4F6FAFD0A483D650F42653BC92217">Yue, Yanli</mods:namePart>
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<mods:affiliation id="054DE9B39C5FD5060CCD7FA35D1A97B2">School of Life Science, Ningxia University, Ningxia, China; e-mail: yueyanli 0407 @ yahoo. cn</mods:affiliation>
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<mods:name id="350439A001C8F93543387BF150D9EB47" type="personal">
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<mods:namePart id="B436F952AF0ACB7E6D48FE8BD549819E">Ren, Dong</mods:namePart>
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<mods:namePart id="30AA0EE77E18512AA94DEDCC45F458CF">Prokop, Jakub</mods:namePart>
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<mods:affiliation id="87C80F6C8E778F4BB603844CCDE0B265">Department of Zoology, Faculty of Science, Charles University in Prague, Viničná 7, CZ- 128 44 Praha 2, Czech Republic; e-mail: jprokop @ natur. cuni. cz</mods:affiliation>
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<mods:typeOfResource id="66723D6D9008BCD4185F587A0EB3CB02">text</mods:typeOfResource>
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<mods:title id="B4997F4228037BE5D98DEF3D8E708A9A">Acta Entomologica Musei Nationalis Pragae</mods:title>
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</mods:titleInfo>
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<mods:part id="94AA4EA769F30FE2E9E2FFFBD48B829C">
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<mods:date id="E4159A514B6B8DE952966814AFE37964">2012</mods:date>
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<mods:number id="1282D64EF2601AC5D29E888CD6DC1289">2012-06-30</mods:number>
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<mods:detail id="24971B65121CB1DACB951107F186D50C" type="volume">
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<mods:number id="20DAEB3C9C357A8D322C38EB50D1416E">52</mods:number>
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<mods:number id="7D989201DE74BCA33365EAA4D21EE8DC">1</mods:number>
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<mods:start id="4A34AA1E85F38A16F354325BB02A5158">89</mods:start>
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<mods:end id="801E108760DD886C1981300978B8681D">127</mods:end>
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<mods:classification id="12AE89F66A1DA0277A7D30AA3D100DD3">journal article</mods:classification>
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<mods:identifier id="D0B51166F0514662A2BD1F1B895CEF1B" type="DOI">10.5281/zenodo.5330604</mods:identifier>
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<mods:identifier id="C433228DE77233EFE9D602B789AAB7C7" type="ISSN">0374-1036</mods:identifier>
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<mods:identifier id="1D73D091CC13A551E18713F43FFE463D" type="Zenodo-Dep">5330604</mods:identifier>
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<treatment id="B970A055FFCAFFEFFE5B278DFD9DFB48" LSID="urn:lsid:plazi:treatment:B970A055FFCAFFEFFE5B278DFD9DFB48" httpUri="http://treatment.plazi.org/id/B970A055FFCAFFEFFE5B278DFD9DFB48" lastPageNumber="99" pageId="10" pageNumber="99">
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<subSubSection id="79C342C8FFCAFFEFFE5B278DFC84FF08" box="[448,829,219,243]" pageId="10" pageNumber="99" type="nomenclature">
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<paragraph id="31661143FFCAFFEFFE5B278DFC84FF08" blockId="10.[448,829,219,243]" box="[448,829,219,243]" pageId="10" pageNumber="99">
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<heading id="6A2EA62FFFCAFFEFFE5B278DFC84FF08" bold="true" box="[448,829,219,243]" centered="true" fontSize="8" level="2" pageId="10" pageNumber="99" reason="3">
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<emphasis id="03ADCD51FFCAFFEFFE5B278DFC84FF08" bold="true" box="[448,829,219,243]" pageId="10" pageNumber="99">
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†
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<taxonomicName id="F6D96AC0FFCAFFEFFE4F278DFC84FF08" ID-CoL="8XKMJ" authority="Ponomarenko, 1977" authorityName="Ponomarenko" authorityYear="1977" box="[468,829,219,243]" class="Insecta" family="Helophoridae" genus="Cretotaenia" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="99" phylum="Arthropoda" rank="genus">
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||
<emphasis id="03ADCD51FFCAFFEFFE4F278DFDECFF08" bold="true" box="[468,597,219,243]" italics="true" pageId="10" pageNumber="99">Cretotaenia</emphasis>
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||
Ponomarenko, 1977
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</taxonomicName>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="79C342C8FFCAFFEFFF0C2654FE67FEEF" box="[151,478,257,276]" pageId="10" pageNumber="99" type="reference_group">
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<paragraph id="31661143FFCAFFEFFF0C2654FE67FEEF" blockId="10.[151,478,257,276]" box="[151,478,257,276]" pageId="10" pageNumber="99">
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<treatmentCitationGroup id="11C9366DFFCAFFEFFF0C2654FE67FEEF" box="[151,478,257,276]" pageId="10" pageNumber="99">
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<treatmentCitation id="B0783752FFCAFFEFFF0C2654FE63FEEF" author="PONOMARENKO A. G." box="[151,474,257,276]" page="92" pageId="10" pageNumber="99" year="1977">
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<taxonomicName id="F6D96AC0FFCAFFEFFF0C2654FE63FEEF" ID-CoL="8XKMJ" authority="Ponomarenko, 1977 a: 92" authorityName="Ponomarenko" authorityPageNumber="92" authorityYear="1977" box="[151,474,257,276]" class="Insecta" family="Helophoridae" genus="Cretotaenia" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="99" phylum="Arthropoda" rank="genus">
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||
<emphasis id="03ADCD51FFCAFFEFFF0C2654FF45FEEF" box="[151,252,258,276]" italics="true" pageId="10" pageNumber="99">Cretotaenia</emphasis>
|
||
<bibRefCitation id="55486CB2FFCAFFEFFE992657FE63FEEF" author="PONOMARENKO A. G." box="[258,474,257,276]" pageId="10" pageNumber="99" pagination="17 - 104" refId="ref15442" refString="PONOMARENKO A. G. 1977 a: Podotryad Adephaga. [Suborder Adephaga]. Pp. 17 - 104. In: ARNOLDI L. V., ZHERIKHIN V. V., NIKRITIN L. M. & PONOMARENKO A. G. (eds.): Mezozoiskie zhestkokrylye. [Mesozoic Coleoptera], Nauka, Moskva, 204 pp (in Russian). [for English translation see ARNOLDI et al. 1992]." type="book chapter" year="1977">Ponomarenko, 1977a: 92</bibRefCitation>
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</taxonomicName>
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||
</treatmentCitation>
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.
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||
</treatmentCitationGroup>
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||
</paragraph>
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||
</subSubSection>
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||
<subSubSection id="79C342C8FFCAFFEFFF0C267BFC7CFE9E" pageId="10" pageNumber="99" type="type_taxon">
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||
<paragraph id="31661143FFCAFFEFFF0C267BFC4CFEBF" blockId="10.[151,1013,300,357]" box="[151,1013,300,325]" pageId="10" pageNumber="99">
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||
<emphasis id="03ADCD51FFCAFFEFFF0C267BFE92FEBF" bold="true" box="[151,299,300,325]" pageId="10" pageNumber="99">
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||
<typeStatus id="EE62AFE1FFCAFFEFFF0C267BFF76FEBE" box="[151,207,301,325]" pageId="10" pageNumber="99">Type</typeStatus>
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species.
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</emphasis>
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||
<taxonomicName id="F6D96AC0FFCAFFEFFEA92678FD66FEBE" ID-CoL="ZFDZ" authority="Ponomarenko, 1977" authorityName="Ponomarenko" authorityYear="1977" box="[306,735,300,325]" class="Insecta" family="Helophoridae" genus="Cretotaenia" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="99" phylum="Arthropoda" rank="species" species="pallida">
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||
<emphasis id="03ADCD51FFCAFFEFFEA92678FDBAFEBE" box="[306,515,302,325]" italics="true" pageId="10" pageNumber="99">Cretotaenia pallida</emphasis>
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Ponomarenko, 1977
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</taxonomicName>
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(by original designation).
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||
</paragraph>
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||
<paragraph id="31661143FFCAFFEFFF0C261AFC7CFE9E" blockId="10.[151,1013,300,357]" box="[151,965,332,357]" pageId="10" pageNumber="99">
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||
<emphasis id="03ADCD51FFCAFFEFFF0C261AFE98FE9E" bold="true" box="[151,289,332,357]" pageId="10" pageNumber="99">Time range.</emphasis>
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||
Early Cretaceous, Berriasian to Hauterivian, ca. 146–135 mya.
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||
</paragraph>
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||
</subSubSection>
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||
<subSubSection id="79C342C8FFCAFFEFFF0C262DFD9DFB48" pageId="10" pageNumber="99" type="diagnosis">
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||
<paragraph id="31661143FFCAFFEFFF0C262DFC30FDC8" blockId="10.[151,1127,379,1203]" pageId="10" pageNumber="99">
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||
<emphasis id="03ADCD51FFCAFFEFFF0C262DFEE5FE6F" bold="true" box="[151,348,379,404]" pageId="10" pageNumber="99">
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Diagnosis.
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<emphasis id="03ADCD51FFCAFFEFFE89262AFEEAFE68" bold="true" box="[274,339,380,403]" italics="true" pageId="10" pageNumber="99">Larva</emphasis>
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:
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</emphasis>
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Head hyperprognathous; nasale simply triangular, epistomal lobes large, slightly overlapping nasale, bearing series of setae; mandible with two retinacular teeth; labium without ligula; each parietale with 6 stemmata situated in area of darker cuticle; all thoracic segments with large dorsal tergite; abdominal segments 1–8 each with a pair of dorsal sclerites and a pair of minute sclerites on each side above spiracle; tracheal system holopneustic, spiracular atrium absent; urogomphi large, 3-segmented.
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||
</paragraph>
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||
<paragraph id="31661143FFCAFFEFFF0C256AFBF6FC28" blockId="10.[151,1127,379,1203]" pageId="10" pageNumber="99">
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||
<emphasis id="03ADCD51FFCAFFEFFF0C256AFF6DFDAF" bold="true" box="[151,212,572,596]" pageId="10" pageNumber="99">Note.</emphasis>
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The genus was originally described as
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<taxonomicName id="F6D96AC0FFCAFFEFFD09256DFD46FDA8" box="[658,767,571,595]" class="Magnoliopsida" higherTaxonomySource="GBIF" kingdom="Plantae" order="Malvales" pageId="10" pageNumber="126" phylum="Tracheophyta" rank="subOrder" subOrder="Adephaga">Adephaga</taxonomicName>
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<emphasis id="03ADCD51FFCAFFEFFC93256BFC18FDAF" box="[776,929,573,596]" italics="true" pageId="10" pageNumber="99">incertae sedis</emphasis>
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by
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||
<bibRefCitation id="55486CB2FFCAFFEFFC4B256AFF55FD8F" author="PONOMARENKO A. G." pageId="10" pageNumber="99" pagination="17 - 104" refId="ref15442" refString="PONOMARENKO A. G. 1977 a: Podotryad Adephaga. [Suborder Adephaga]. Pp. 17 - 104. In: ARNOLDI L. V., ZHERIKHIN V. V., NIKRITIN L. M. & PONOMARENKO A. G. (eds.): Mezozoiskie zhestkokrylye. [Mesozoic Coleoptera], Nauka, Moskva, 204 pp (in Russian). [for English translation see ARNOLDI et al. 1992]." type="book chapter" year="1977">PONOMARENKO (1977a)</bibRefCitation>
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||
, but
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||
<bibRefCitation id="55486CB2FFCAFFEFFE80250AFE71FD8F" author="CROWSON R. A." box="[283,456,604,628]" pageId="10" pageNumber="99" refId="ref14380" refString="CROWSON R. A. 1981: The biology of the Coleoptera. Academic Press, London, 802 pp." type="book" year="1981">CROWSON (1981)</bibRefCitation>
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||
later noted that it may rather be related to
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<taxonomicName id="F6D96AC0FFCAFFEFFCED250BFC4BFD8F" authorityName="Fabricius" authorityYear="1775" box="[886,1010,605,628]" class="Insecta" family="Helophoridae" genus="Helophorus" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="99" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="03ADCD51FFCAFFEFFCED250BFC4BFD8F" box="[886,1010,605,628]" italics="true" pageId="10" pageNumber="99">Helophorus</emphasis>
|
||
</taxonomicName>
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||
. A detailed comparison of all preserved characters including those not mentioned in the original description confirmed this opinion:
|
||
<taxonomicName id="F6D96AC0FFCAFFEFFE0325CBFDAFFD4F" authorityName="Ponomarenko" authorityYear="1977" box="[408,534,669,692]" class="Insecta" family="Helophoridae" genus="Cretotaenia" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="99" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="03ADCD51FFCAFFEFFE0325CBFDAFFD4F" box="[408,534,669,692]" italics="true" pageId="10" pageNumber="99">Cretotaenia</emphasis>
|
||
</taxonomicName>
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||
precisely matches modern
|
||
<taxonomicName id="F6D96AC0FFCAFFEFFCA025CBFC00FD4F" authorityName="Fabricius" authorityYear="1775" box="[827,953,669,692]" class="Insecta" family="Helophoridae" genus="Helophorus" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="99" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="03ADCD51FFCAFFEFFCA025CBFC00FD4F" box="[827,953,669,692]" italics="true" pageId="10" pageNumber="99">Helophorus</emphasis>
|
||
</taxonomicName>
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||
larvae in nearly all preserved characters except the labium which does not protract as much anteriad, different arrangement of abdominal sclerites (two small sclerites are present laterally of dorsal large sclerite and dorsally of the spiracle in
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<taxonomicName id="F6D96AC0FFCAFFEFFDA325ABFD0FFCEF" authorityName="Ponomarenko" authorityYear="1977" box="[568,694,765,788]" class="Insecta" family="Helophoridae" genus="Cretotaenia" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="99" phylum="Arthropoda" rank="genus">
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||
<emphasis id="03ADCD51FFCAFFEFFDA325ABFD0FFCEF" box="[568,694,765,788]" italics="true" pageId="10" pageNumber="99">Cretotaenia</emphasis>
|
||
</taxonomicName>
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||
, whereas
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||
<taxonomicName id="F6D96AC0FFCAFFEFFCBF25ABFC1BFCEF" authorityName="Fabricius" authorityYear="1775" box="[804,930,765,788]" class="Insecta" family="Helophoridae" genus="Helophorus" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="99" phylum="Arthropoda" rank="genus">
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||
<emphasis id="03ADCD51FFCAFFEFFCBF25ABFC1BFCEF" box="[804,930,765,788]" italics="true" pageId="10" pageNumber="99">Helophorus</emphasis>
|
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</taxonomicName>
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||
has a single large sclerite in this position) and the dorsally situated occipital foramen (situated posteriorly in
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<taxonomicName id="F6D96AC0FFCAFFEFFF0C246BFEAAFCAF" authorityName="Fabricius" authorityYear="1775" box="[151,275,829,852]" class="Insecta" family="Helophoridae" genus="Helophorus" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="99" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="03ADCD51FFCAFFEFFF0C246BFEAAFCAF" box="[151,275,829,852]" italics="true" pageId="10" pageNumber="99">Helophorus</emphasis>
|
||
</taxonomicName>
|
||
). The phylogenetic analysis by
|
||
<bibRefCitation id="55486CB2FFCAFFEFFDCC246AFC92FCAF" author="FIKACEK M. & PROKIN A. & ANGUS R. B. & PONOMARENKO A. & YUE Y. & REN D. & PROKOP J." box="[599,811,827,852]" pageId="10" pageNumber="99" pagination="37" refId="ref14855" refString="FIKACEK M., PROKIN A., ANGUS R. B., PONOMARENKO A., YUE Y., REN D. & PROKOP J. 2012: Phylogeny and the fossil record of the Helophoridae reveal Jurassic origin of modern hydrophiloid lineages (Coleoptera: Polyphaga). Systematic Entomology 37: in press, DOI: 10.1111 / j. 1365 - 3113.2012.00630. x" type="book chapter" year="2012">FIKÁČEK et al. (2012)</bibRefCitation>
|
||
also confirms that
|
||
<taxonomicName id="F6D96AC0FFCAFFEFFC71246BFBDFFCAF" authorityName="Ponomarenko" authorityYear="1977" box="[1002,1126,829,852]" class="Insecta" family="Helophoridae" genus="Cretotaenia" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="99" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="03ADCD51FFCAFFEFFC71246BFBDFFCAF" box="[1002,1126,829,852]" italics="true" pageId="10" pageNumber="99">Cretotaenia</emphasis>
|
||
</taxonomicName>
|
||
is related to the
|
||
<taxonomicName id="F6D96AC0FFCAFFEFFEA7240DFE69FC88" authorityName="Leach" authorityYear="1815" box="[316,464,859,883]" class="Insecta" family="Helophoridae" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="99" phylum="Arthropoda" rank="family">Helophoridae</taxonomicName>
|
||
, and the genus is therefore included into the helophorid lineage in the present work.
|
||
<bibRefCitation id="55486CB2FFCAFFEFFEEB242AFDDBFC6F" author="ZHERIKHIN V. V. & MOSTOVSKI M. B. & VRSANSKY P. & BLAGODEROV V. A. & LUKASHEVICH E." box="[368,610,891,916]" pageId="10" pageNumber="99" pagination="185 - 191" refId="ref16251" refString="ZHERIKHIN V. V., MOSTOVSKI M. B., VRSANSKY P., BLAGODEROV V. A. & LUKASHEVICH E. 1998: The unique Lower Cretaceous locality Baissa and other contemporaneous fossil sites in North and West Transbaikalia. Pp. 185 - 191. In: Proceedings of the first international paleoentomological conference, Moscow 1998. AMBA Projects International, 121 pp." type="book chapter" year="1998">ZHERIKHIN et al. (1998)</bibRefCitation>
|
||
hypothesized that
|
||
<taxonomicName id="F6D96AC0FFCAFFEFFCB2242BFC43FC6F" authorityName="Ponomarenko" authorityYear="1977" box="[809,1018,893,916]" class="Insecta" family="Helophoridae" genus="Cretotaenia" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="99" phylum="Arthropoda" rank="species" species="pallida">
|
||
<emphasis id="03ADCD51FFCAFFEFFCB2242BFC43FC6F" box="[809,1018,893,916]" italics="true" pageId="10" pageNumber="99">Cretotaenia pallida</emphasis>
|
||
</taxonomicName>
|
||
is likely a larval form of
|
||
<taxonomicName id="F6D96AC0FFCAFFEFFEA824CBFDB6FC4F" authorityName="Fikáček & Prokin & Angus & Pono & Marenko & Yue & Ren & Prokop" authorityYear="2012" baseAuthorityName="Ponomarenko" baseAuthorityYear="1987" box="[307,527,925,948]" class="Insecta" family="Helophoridae" genus="Laetopsia" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="99" phylum="Arthropoda" rank="species" species="baissensis">
|
||
<emphasis id="03ADCD51FFCAFFEFFEA824CBFDB6FC4F" box="[307,527,925,948]" italics="true" pageId="10" pageNumber="99">Laetopsia baissensis</emphasis>
|
||
</taxonomicName>
|
||
, which cannot be confirmed or rejected by our analyses even though both genera seem to represent basal extinct clades of the helophorid lineage.
|
||
</paragraph>
|
||
<paragraph id="31661143FFCAFFEFFF2E248AFD9DFB48" blockId="10.[151,1127,379,1203]" pageId="10" pageNumber="99">
|
||
At first view, the number of stemmata seems to be five on each parietale in the specimens examined. However, the arrangement of stemmata in the fossils precisely fits that in modern
|
||
<taxonomicName id="F6D96AC0FFCAFFEFFF0C234BFEACFBCF" authorityName="Fabricius" authorityYear="1775" box="[151,277,1053,1076]" class="Insecta" family="Helophoridae" genus="Helophorus" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="99" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="03ADCD51FFCAFFEFFF0C234BFEACFBCF" box="[151,277,1053,1076]" italics="true" pageId="10" pageNumber="99">Helophorus</emphasis>
|
||
</taxonomicName>
|
||
(see
|
||
<bibRefCitation id="55486CB2FFCAFFEFFED0234AFDAFFBCF" author="FIKACEK M. & PROKIN A. & ANGUS R. B. & PONOMARENKO A. & YUE Y. & REN D. & PROKOP J." box="[331,534,1051,1076]" pageId="10" pageNumber="99" pagination="37" refId="ref14855" refString="FIKACEK M., PROKIN A., ANGUS R. B., PONOMARENKO A., YUE Y., REN D. & PROKOP J. 2012: Phylogeny and the fossil record of the Helophoridae reveal Jurassic origin of modern hydrophiloid lineages (Coleoptera: Polyphaga). Systematic Entomology 37: in press, DOI: 10.1111 / j. 1365 - 3113.2012.00630. x" type="book chapter" year="2012">FIKÁČEK et al. 2012</bibRefCitation>
|
||
,
|
||
<figureCitation id="A9E20DC6FFCAFFEFFD84234DFDC9FBCF" box="[543,624,1051,1076]" captionStart="Figs" captionStartId="4.[151,190,907,926]" captionTargetBox="[217,1122,220,883]" captionTargetId="figure-188@4.[201,1129,216,894]" captionTargetPageId="4" captionText="Figs. 1–4. Hydrophilopsia longitarsalis Ponomarenko, 1987 (holotype). 1 – piece, general view; 2 – counterpiece, general view; 3 – detail of antenna; 4 – detail of the aedeagus. Scale bar: 2 mm (corresponds to Figs. 1–2). Abbreviations: acl – antennal club; aed – aedeagus; ant – antenna; cup – cupule; ins – intermediate antennomeres; ped – pedicel; sc – scapus." figureDoi="http://doi.org/10.5281/zenodo.5330606" httpUri="https://zenodo.org/record/5330606/files/figure.png" pageId="10" pageNumber="99">Fig. 1H</figureCitation>
|
||
) in which five stemmata are situated dorsally in the area of darkened cuticle (and hence preserved in the fossils), whereas the largest stemma is rather isolated from the others, situated more ventrally outside of the area of darkened cuticle (and is therefore not preserved in the fossil). For that reason we consider
|
||
<taxonomicName id="F6D96AC0FFCAFFEFFC73232BFBDFFB6F" authorityName="Ponomarenko" authorityYear="1977" box="[1000,1126,1149,1172]" class="Insecta" family="Helophoridae" genus="Cretotaenia" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="99" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="03ADCD51FFCAFFEFFC73232BFBDFFB6F" box="[1000,1126,1149,1172]" italics="true" pageId="10" pageNumber="99">Cretotaenia</emphasis>
|
||
</taxonomicName>
|
||
having 6 stemmata on each parietale.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |